IndraLab

Statements


ASXL1 affects BAP1
28 | 1 181 130
ASXL1 binds BAP1.
28 | 146 120
28 | 129 113

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"The mammalian homologue of Calypso, BAP1, directly associates with ASXL1, and the mammalian BAP1 and ASXL1 complex was shown to possess deubiquitinase activity in vitro 39."

No evidence text available

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"OGT improves ASXL1 stability by inducing O-GlcNAcylation at the ASXL1 Ser199 site [127], while ASXL1 interacts with and activates BAP1 [66]."

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"BAP1 interacts with the polycomb factors ASXL1 and 2 and its substrates HCF-1 and OGT, but the precise mechanism how its loss of function bypasses growth arrest in BRAF mutated nevi remains to be elucidated."

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"8 BAP1 interacts with ASXL1 (MIM 612990) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"ASXL1 forms a complex with the deubiquitination enzyme BAP1 and removes monoubiquitin from H2AK119Ub, to derepress genes targeted by PRC1 [ xref ]."
| PMC

sparser
"The endogenous BAP1 interaction with ASXL1 was confirmed by IP and WB analysis ( Supplemental Fig. 2B )."

No evidence text available

No evidence text available

sparser
"Using the same strategy, we found that human BAP1 also forms a stable complex with the N-terminal domain of human ASXL1 (ASXL1(2–365)) but not with the human PcG proteins BMI1 or RING1A ( xref , right, lanes 8–10)."
Mutated ASXL1 binds BAP1. 10 / 11
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"This hyperactive mutant ASXL1/BAP1 complex is thought to promote impairment of myeloid differentiation through inhibition of H2AK119ub at posterior HOXA genes and IRF8, encoding an essential transcription factor in myeloid lineages [18]."

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"Cancer associated ASXL1 mutations may act as gain-of-function mutations of the ASXL1 and BAP1 complex."

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"The mutant ASXL1/BAP1 complex inhibits the differentiation of HSPCs toward multilineage except for immature monocytes, and promoted myeloid leukemogenesis induced by RUNX1-ETO, an oncogenic fusion protein frequently coexisting with ASXL1 mutations [72]."

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"This hyperactive mutant ASXL1 and BAP1 complex induces upregulation of posterior HOXA genes and IRF8 through inhibition of H2AK119ub, impairing multilineage differentiation of haematopoietic progenitors (except for that toward monocytes), and accelerates RUNX1-ETO-induced leukaemogenesis."

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"Based of the possibility that ASXL1 truncations might act as gain-of-function mutations of the ASXL1 and BAP1 complex, we examined whether hyperactive ASXL-BAP1 complexes could alter the fate of haematopoietic cells in vivo."

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"Our studies here raise the possibility that ASXL1 mutations may act as gain-of-function mutations of the ASXL1 and BAP1 complex."

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"We have examined the role of H2AK119Ub in regulating gene expression by using hyperactive mutations of the ASXL1 and BAP1 complex that allowed us to deplete ~ 90% of total H2AK119Ub."

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"Mechanistically, mutant ASXL1 interacted with and stabilized BAP1 (BRCA1 Associated Protein 1), a nuclear ubiquitin carboxy-terminal hydrolase, resulting in the deubiquitination and activation of AKT,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The overactive mutant ASXL1/BAP1 complex aberrantly de-ubiquitinated H2AK119Ub, followed by de-repression of its target genes including posterior HOXA genes and IRF8 , leading to leukemic transformati[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"[Molecular mechanisms by which the mutant ASXL1 and BAP1 complex aggravates myeloid leukemia]."
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

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"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

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"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

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"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

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"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
EED binds BAP1 and ASXL1. 4 / 4
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"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
BAP1 binds ASXL1 and ASXL2. 3 / 3
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sparser
"Although both ASXL1 and ASXL2 have been identified to bind to BAP1, recent biochemical characterization of BAP1 complexes by xref suggest that BAP1 forms two mutually exclusive complexes with ASXL1 and ASXL2."

sparser
"Proteomic analysis of BAP1 interacting partners using this system revealed that BAP1 interacts with ASXL1 and ASXL2 in vivo as well as host cell factor 1 (HCF-1), O-GlcNAc transferase (OGT), and Forkhead box protein K1/2 (FOXK1/2) ( xref )."

sparser
"Depletion of the PHD fingers did not affect the stability of ASXL1 or ASXL2, or the interaction between ASXL1, ASXL2, and BAP1 (Additional file xref : Fig. S2A, B)."
BAP1 binds ASXL1 and ASXH domain. 2 / 2
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"ASXL1 directly binds to BAP1 through the ASXH domain and its adjoining region."

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"As mentioned earlier, the interaction between BAP1 and ASXL1 occurs through the ASXH domain."
ASXL1 activates BAP1.
| 1 35 10
ASXL1 activates BAP1. 10 / 40
| 1 29 10

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"Because of the strong conservation of key elements between UCH-L5 and RPN13 DEU and BAP1 and ASXL1, we anticipate that the ASXL1 DEUBAD domain employs similar strategies to activate BAP1."

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"Increasing concentrations of TG2-179-1 were incubated with H2A-Ub nucleosomes containing the DEUBAD domain of ASXL1 (ASXL1 ), which activates BAP1 by increasing its affinity for the ubiquitin moiety of H2A-Ub ."

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"As part of the Polycomb repression machinery, BAP1 is activated by the deubiquitinase adaptor domain of ASXL1 mediating gene repression by cleaving ubiquitin (Ub) from histone H2A in nucleosomes."

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"The molecular mechanism of BAP1 activation by ASXL1 remains elusive, as no structures are available for either BAP1 or ASXL1."

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"BAP1 is activated by ASXL1 in a similar manner where the DEUBAD domain of ASXL1 increases affinity of BAP1 for the ubiquitinated substrate [98]."

sparser
"A recent study suggests that ASXL1 might activate Bap1 in a similar manner xref ."

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"Several studies reported that ASXL1 truncation enhanced the catalytic deubiquitinating activity of BAP1 towards mono-ubiquitinated H2AK119 [ 20 , 36 ]."

sparser
"The molecular mechanism of BAP1 activation by ASXL1 remains elusive, as no structures are available for either BAP1 or ASXL1."

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"A previous report showed that the deubiquitinase adapter (DEUBAD) domain of ASXL1 increases BAP1 's affinity for ubiquitin on H2A 15, and we showed that monoubiquitinated ASXL1-MT enhances nuclear retention of BAP1."

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"To further confirm that GGNBP2 loss could affect the activation of BAP1 by ASXL1 in deubiquitinating H2A, Ggnbp2KO cells were transfected with ASXL1 siRNA together with Ggnbp2 overexpression plasmids."
Mutated ASXL1 activates BAP1. 6 / 6
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"ASXL1 mutations found in AML aberrantly enhance the DUB activity of BAP-1 and thus, deplete the levels of H3K27me 3."

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"Monoubiquitination of mutant ASXL1 enhanced the catalytic function of BAP1, resulting in a profound reduction in H2AK119ub by counteracting the PRC1 complex."

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"Simultaneous overexpression of BAP1 and mutant ASXL1 promotes the deubiquitinase activity of BAP1, leading to the loss of H2AK119Ub followed by H3K27me3 reduction (Balasubramani et al., 2015)."

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"In our model, BAP1 mRNA did not change in KI mice, and physiological expression of Asxl1 mutant seemed to mildly support Bap1 function, which is not sufficient for global H3K27me3 reduction."

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"However, recent studies have reported that hyperactive catalytic function of BAP1 caused by ASXL1 mutation in haematopoietic cells disturbs normal haematopoiesis and promotes myeloid leukaemogenesis i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Mechanistically, BAP1 mono-ubiquitinated and stabilized mutant ASXL1, which in turn enhanced the DUB activity of BAP1."
BAP1 affects ASXL1
28 | 151 120
BAP1 binds ASXL1.
28 | 146 120
28 | 129 113

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"The mammalian homologue of Calypso, BAP1, directly associates with ASXL1, and the mammalian BAP1 and ASXL1 complex was shown to possess deubiquitinase activity in vitro 39."

No evidence text available

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"OGT improves ASXL1 stability by inducing O-GlcNAcylation at the ASXL1 Ser199 site [127], while ASXL1 interacts with and activates BAP1 [66]."

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"BAP1 interacts with the polycomb factors ASXL1 and 2 and its substrates HCF-1 and OGT, but the precise mechanism how its loss of function bypasses growth arrest in BRAF mutated nevi remains to be elucidated."

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"8 BAP1 interacts with ASXL1 (MIM 612990) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"ASXL1 forms a complex with the deubiquitination enzyme BAP1 and removes monoubiquitin from H2AK119Ub, to derepress genes targeted by PRC1 [ xref ]."
| PMC

sparser
"The endogenous BAP1 interaction with ASXL1 was confirmed by IP and WB analysis ( Supplemental Fig. 2B )."

No evidence text available

No evidence text available

sparser
"Using the same strategy, we found that human BAP1 also forms a stable complex with the N-terminal domain of human ASXL1 (ASXL1(2–365)) but not with the human PcG proteins BMI1 or RING1A ( xref , right, lanes 8–10)."
Mutated ASXL1 binds BAP1. 10 / 11
| 11

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"This hyperactive mutant ASXL1/BAP1 complex is thought to promote impairment of myeloid differentiation through inhibition of H2AK119ub at posterior HOXA genes and IRF8, encoding an essential transcription factor in myeloid lineages [18]."

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"Cancer associated ASXL1 mutations may act as gain-of-function mutations of the ASXL1 and BAP1 complex."

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"The mutant ASXL1/BAP1 complex inhibits the differentiation of HSPCs toward multilineage except for immature monocytes, and promoted myeloid leukemogenesis induced by RUNX1-ETO, an oncogenic fusion protein frequently coexisting with ASXL1 mutations [72]."

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"This hyperactive mutant ASXL1 and BAP1 complex induces upregulation of posterior HOXA genes and IRF8 through inhibition of H2AK119ub, impairing multilineage differentiation of haematopoietic progenitors (except for that toward monocytes), and accelerates RUNX1-ETO-induced leukaemogenesis."

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"Based of the possibility that ASXL1 truncations might act as gain-of-function mutations of the ASXL1 and BAP1 complex, we examined whether hyperactive ASXL-BAP1 complexes could alter the fate of haematopoietic cells in vivo."

reach
"Our studies here raise the possibility that ASXL1 mutations may act as gain-of-function mutations of the ASXL1 and BAP1 complex."

reach
"We have examined the role of H2AK119Ub in regulating gene expression by using hyperactive mutations of the ASXL1 and BAP1 complex that allowed us to deplete ~ 90% of total H2AK119Ub."

reach
"Mechanistically, mutant ASXL1 interacted with and stabilized BAP1 (BRCA1 Associated Protein 1), a nuclear ubiquitin carboxy-terminal hydrolase, resulting in the deubiquitination and activation of AKT,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The overactive mutant ASXL1/BAP1 complex aberrantly de-ubiquitinated H2AK119Ub, followed by de-repression of its target genes including posterior HOXA genes and IRF8 , leading to leukemic transformati[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"[Molecular mechanisms by which the mutant ASXL1 and BAP1 complex aggravates myeloid leukemia]."
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

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"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

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"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

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"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

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"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
BAP1 binds ASXL1 and ASXL2. 3 / 3
| 3

sparser
"Although both ASXL1 and ASXL2 have been identified to bind to BAP1, recent biochemical characterization of BAP1 complexes by xref suggest that BAP1 forms two mutually exclusive complexes with ASXL1 and ASXL2."

sparser
"Proteomic analysis of BAP1 interacting partners using this system revealed that BAP1 interacts with ASXL1 and ASXL2 in vivo as well as host cell factor 1 (HCF-1), O-GlcNAc transferase (OGT), and Forkhead box protein K1/2 (FOXK1/2) ( xref )."

sparser
"Depletion of the PHD fingers did not affect the stability of ASXL1 or ASXL2, or the interaction between ASXL1, ASXL2, and BAP1 (Additional file xref : Fig. S2A, B)."
BAP1 binds ASXL1 and ASXH domain. 2 / 2
| 2

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"ASXL1 directly binds to BAP1 through the ASXH domain and its adjoining region."

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"As mentioned earlier, the interaction between BAP1 and ASXL1 occurs through the ASXH domain."
BAP1 activates ASXL1.
| 5
BAP1 activates ASXL1. 5 / 5
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"Reduced BAP1 activity prevents ASXL1 truncation driven myeloid malignancy in vivo."

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"Truncated ASXL1 increases BAP1 protein stability, enhances BAP1 recruitment to chromatin and promotes the expression of a pro-leukemic transcriptional signature."

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"RNA sequencing analyses revealed that reduced BAP1 expression suppressed ASXL1 Y591X -induced upregulation of the transcription factors activator protein 1 ( AP-1 ) and early growth response 1/2 ( EGR[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Strikingly, these include both loss-of-function mutations in Bap1 and Asxl1/2 and gain-of-function mutations in which truncated ASXL1 increases BAP1 stability."

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"Guo and her colleagues found that deletion of one allele of Bap1 in Asxl1 -truncated mice prevented ASXL1 truncation-driven myeloid malignancy [ 38 ]."
BAP1 affects SLC7A11
| 7 170 4
BAP1 decreases the amount of SLC7A11.
| 102
BAP1 decreases the amount of SLC7A11. 10 / 102
| 102

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"Together, our data showed that BAP1 decreases whereas PRC1 increases H2Aub binding on the SLC7A11 promoter, but both BAP1 and PRC1 represses SLC7A11 expression, thus supporting a model that a balance between H2A ubiquitination and de-ubiquitination might be important for maintaining SLC7A11 repression (see Discussion)."

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"In addition, it 's been found that in renal cancer cell line, as a nuclear deubiquitinating enzyme, the tumour suppressor BRCA1 associated protein 1 decreases SLC7A11 expression, thereby causing lipid peroxidation and ferroptosis [XREF_BIBR]."

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"SLC7A11 is an essential BAP1 target in human cancers, and BAP1 represses SLC7A11 expression via reducing H2Aub occupancy on SLC7A11 promoter in a deubiquitinating dependent manner [XREF_BIBR, XREF_BIBR]."

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"Another study (Zhang Y. et al., 2018) suggests that the tumor suppressor BRCA1-associated protein 1 (BAP1) also promotes ferroptosis by downregulating the expression of SLC7A11."

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"Studies have shown that BAP1 inhibits the expression of solute carrier family 7 member 11 (SLC7A11) by decreasing H2Aub occupancy on it."

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"Indeed, p53 or the histone deubiquitinating enzyme BAP1 inhibiting SLC7A11 expression promotes ferroptosis (124, 125)."

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"Studies have found that SLC7A11 is a key target gene of BAP1, and BAP1 inhibits the expression of SLC7A11 by reducing H2Aub on the SLC7A11 promoter, promoting the occurrence of ferroptosis, and then inhibiting tumor growth."

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"BAP1 reduces H2Aub occupancy on the SLC7A11 promoter and represses SLC7A11 expression in a deubiquitin-dependent, but not NFE2L2- and ATF4-dependent manner [87]."

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"The mechanism may be that BAP1 suppresses SLC7A11 transcription by reducing histone 2A ubiquitination (H2Aub) on the SLC7A11 gene."

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"In this manner, BAP1 reduces SLC7A11 expression and cysteine uptake, leading lipid peroxidation upregulation and the ferroptosis induction (71)."
BAP1 inhibits SLC7A11.
| 7 30 1
| 7 30 1

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"Moreover, BAP1 partially prevents the progression of tumors via SLC7A11 as well as ferroptosis, and cancer-related BAP1 mutants end up losing their capacities to suppress SLC7A11 and to stimulate ferroptosis [25]."

eidos
"In this manner , BAP1 reduces SLC7A11 expression and cysteine uptake , leading lipid peroxidation upregulation and the ferroptosis induction ( 71 ) ."

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"Luciferase reporter assay showed that BAP1-WT could reduce the SLC7A11 promoter, while BAP1-MUT had no effect on the promoter."

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"For instance, BAP1 and p53 inhibited SLC7A11 to induce ferroptosis, while KEAP1 prevented NRF2 from activating SLC7A11 ( xref - xref )."

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"BAP1, as a tumor suppressor, promotes ferroptosis and inhibits cancer cell growth by inhibiting SLC7A11 [94]."

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"The repression of SLC7A11 by BAP1 unveils a potential ferroptosis axis implicated in placental physiology and pathology, shedding light on novel mechanisms underlying pregnancy disorders ."

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"Several studies have revealed that BAP1 can inhibit ubiquitinated histone 2A (H2Aub) occupancy on the SLC7A11 promoter (Zhang et al., 2018)."

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"It has been recently reported that BAP1 facilitates lipid peroxidation and promotes ferroptosis through repressing SLC7A11 [24]."

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"Analysis of our H2Aub ChIP-seq data revealed that restoration of BAP1 WT, but not BAP1 C91A, markedly decreased H2Aub occupancy at both the promoter and gene body of SLC7A11 (XREF_FIG), which was further confirmed by H2Aub ChIP assay on the SLC7A11 promoter and representative exons (XREF_FIG and XREF_SUPPLEMENTARY)."

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"Together, our data suggested a model that the BAP1 containing PR and DUB complex binds on the SLC7A11 promoter, where BAP1 removes ubiquitin from H2Aub, and BAP1 dependent H2Aub reduction on SLC7A11 is associated with BAP1 mediated SLC7A11 repression."
BAP1 increases the amount of SLC7A11.
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BAP1 increases the amount of SLC7A11. 10 / 18
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"BAP1 interacts with Nrf2 to modulate SLC7A11 expression and GSH levels [116,117,118]."
| PMC

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"As discussed in Boxes 2 and 3, inactivation of tumour suppressors such as p53, BAP1, kelch-like ECH associated protein 1 (KEAP1) and ARF (p14), or activation of oncogenic KRAS, upregulates SLC7A11 expression, which can be dependent or independent of nuclear factor erythroid 2-related factor 2 (NRF2), conferring ferroptosis evasion and promoting tumour growth ."

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"Another tumor suppressor, BAP1, which is often lost or mutated in various cancers, has been shown to inhibit SLC7A11 expression, thereby reducing cystine uptake and facilitating ferroptosis in cancer cells."

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"This is consistent with our finding that BAP1 represses SLC7A11 expression (therefore, lower dosages of erastin were needed to block SLC7A11 activity and to induce ferroptosis in BAP1 expressing cells, which exhibited lower expression of SLC7A11)."

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"The tumor suppressor BRCA1 associated protein 1 (BAP1) represses SLC7A11 expression via reducing H2Aub occupancy on SLC7A11 promoter in a deubiquitinating dependent manner [XREF_BIBR]."

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"Moreover, tumor suppressor factor BAP1, an epigenetic regulator, has also been shown to downregulate SLC7A11 expression to promote ferroptosis, but the extent of BAP1’s pro-ferroptotic activity in tumor suppression remains unclear (Zhang et al., 2018)."

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"For example, the tumor suppressor gene, BRCA1-associated protein 1 (BAP1), promotes lipid peroxidation by downregulating the expression level of SLC7A11, thereby promoting tumor cell ferroptosis (42)."

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"Conversely, BAP1 deficiency by CRISPR technology increased SLC7A11 expression in several BAP1-proficient renal cancer cells with low SLC7A11 levels, including 786-O, Caki1, and ACHN cells (XREF_FIG - XREF_FIG and XREF_SUPPLEMENTARY - XREF_SUPPLEMENTARY)."

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"The tumor suppressor protein BRCA1-associated protein 1 (BAP1) inhibits SLC7A11 expression in a deubiquitinating-dependent manner and induces lipid peroxidation to promote ferroptosis (53)."

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"BAP1 mediated deubiquitination of histone 2A ubiquitination (H2Aub), while monoubiquitination of histone H2B on lysine 120 (H2Bub1) induced the expression of SLC7A11 during ferroptosis and consequentl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 deubiquitinates SLC7A11.
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BAP1 deubiquitinates SLC7A11. 8 / 8
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"At the SLC7A11 promoter, BAP1 deubiquitinates H2Aub and subsequently inhibits the expression of SLC7A11."

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"BAP1 deubiquitinates histone 2A (H2Aub) on the SLC7A11 promoter and represses SLC7A11 expression in a p53-independent way (39, 40)."

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"BAP1 represses solute carrier family 7 member 11 (SLC7A11) expression by reducing H2A ubiquitination of the SLC7A11 promoter, resulting in lipid peroxidation and ferroptosis (Zhang et al., 2019)."

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"Given that miR-31-5p has been shown to reduces DOX-induced cardiac toxicity [16], we investigated the role of the targeted relationship between miR-31-5p and BAP1 in SCM.Against this background, it is clear that the regulation of SLC7A11 deubiquitination by BAP1 may have an impact on ferroptosis in SCM."

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"To determine whether BAP1 induced the deubiquitination of SLC7A11, the deubiquitination enzyme inhibitor G5 was added to examine its effect on the ubiquitination level of SLC7A11."

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"These findings imply that BAP1 induces the deubiquitination of SLC7A11, thereby inhibiting its expression."

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"As shown above, BAP1 promotes the deubiquitination of SLC7A11, leading to ferroptosis in H9C2 cells."

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"BRCA1-associated protein 1 (BAP1) decreases SLC7A11 promoter H2A ubiquitination via an H2A deubiquitinase that functions as a tumor suppressor."
BAP1 activates SLC7A11.
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BAP1 activates SLC7A11. 8 / 8
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"The FTO/BAP1 axis promoted MPP + -induced ferroptosis by suppressing SLC7A11."

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"The reduction of the tumor suppressor protein BAP1 and the ubiquitin ligase PRC1 of H2Aub can increase H2Aub uptake of SLC7A11 promoter [107]."

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"Several tumor-suppressors, such as p53 [69] , BAP1 [70] , and KEAP1 [71] , promote ferroptosis through inactivating SLC7A11, while some oncogenic signaling pathways such as PI3K-mTORC1 pathway have be[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Our data that BAP1 plays a role in regulating several metabolic pathways in osteoclasts also supports this concept.Repression of Slc7a11, the cystine/glutamate xCT amino acid antiporter that antagonizes glutamate metabolism in osteoclasts, is one mechanism by which BAP1 regulates such pathways ."

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"SLC7A11 is upregulated by BRCA1-associated protein 1 (BAP1) inactivation in cancer cells, which increases the uptake of cystine and the synthesis of GSH to develop the growth of cancer by inhibiting ferroptosis (Zhang et al. 2018)."

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"BAP1 can induce ferroptosis in a similar process to TP53 by downregulating SLC7A11 [57]."

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"In brief, according to their analyses, BAP1 inactivation leads to an up-regulation of the Solute Carrier family 7 member 11 ( SLC7A11 )."

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"It has been reported that the inactivation of BAP1 upregulates SLC7A11 and suppresses ferroptosis, resulting in tumor development."
BAP1 ubiquitinates SLC7A11.
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BAP1 ubiquitinates SLC7A11. 4 / 4
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"A recent study reported that BAP1 inhibits cystine uptake and makes cells more susceptible to ferroptosis by deubiquitinating H2Aub on SLC7A11."

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"Specifically, our study suggests a model in which the tumor suppressor BAP1 normally functions to repress the expression of SLC7A11 at least partly by deubiquitinating H2Aub on SLC7A11, thereby inhibiting cystine uptake into cells and rendering them more sensitive to ferroptosis."

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"18, NO. 8, 773-783 https://doi.org/10.1080/15384101.2019.1597506EXTRA VIEWRegulation of H2A ubiquitination and SLC7A11 expression by BAP1 and PRC1Yilei Zhang a, Pranavi Koppula a, b, and Boyi Gan a, baDepartment of Experimental Radiation Oncology, the University of Texas MD Anderson Cancer Center, Houston, TX, USA; bDepartment of Experimental Radiation Oncology, The University of Texas MD Anderson UTHealth Graduate School of Biomedical Sciences, Houston, TX, USAABSTRACTSLC7A11 (or xCT) imports extracellular cystine into cells to promote glutathione synthesis, thus inhibiting ferroptosis."

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"Cell CycleISSN : 1538-4101 (Print) 1551-4005 (Online) Journal homepage : https://www.tandfonline.com/loi/kccy20Regulation of H2A ubiquitination and SLC7A11 expression by BAP1 and PRC1Yilei Zhang, Pranavi Koppula & Boyi GanTo cite this article : Yilei Zhang, Pranavi Koppula & Boyi Gan (2019) Regulation of H2A ubiquitination and SLC7A11 expression by BAP1 and PRC1, Cell Cycle, 18:8, 773-783, DOI : 10.1080/15384101.2019.1597506 To link to this article : https://doi.org/10.1080/15384101.2019.1597506View supplementary material Accepted author version posted online : 23 Mar 2019."
BAP1 binds SLC7A11.
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"Targeting the BAP1-SLC7A11 axis may provide novel insights into the treatment of ferroptosis-related intestinal diseases."

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"Overexpressed BAP1 WT or BAP1 C91A binds directly to the Slc7a11 promoter (Fig.  xref )."

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"Notably, ChIP qPCR data using BAP1 antibody indicates BAP1 binding to the Slc7a11 promoter."
BAP1 affects BRCA1
5 1 1 | 86 58
BAP1 binds BRCA1.
5 1 | 41 58
5 1 | 39 53

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"The region of BAP1 that binds to BRCA1 spans amino acids 596–721, while the region that interacts with BARD1 is from 182 to 365 [ xref ]."

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"Hence, as BAP1 protein interacts with BRCA1 protein, PARP inhibitors might also be useful in the treatment of patients with somatic BAP1 inactivation."

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"BRCA1 also interacts with BAP-1 (BRCA1 associated protein) [28], a member of the ubiquitin C-terminal hydrolase family, that removes ubiquitin from proteins."

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"XREF_BIBR It was initially suggested that BAP1 bound BRCA1 cleaves ubiquitin, and enhances the growth-suppressive effects of BRCA1."

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"Thus, NUMB enhances Notch signaling by regulating the ubiquitinating activity of the BAP1BRCA1 complex."

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"Importantly, artificial upregulation of BAP1 restored the protein level of BRCA1 in cells, while it had little effect on the BRCA1 mRNA expression.Following the findings above, we then focused on the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"First, the interaction between BAP1 and BRCA1 in RTECs was validated by Co-IP and Western blot analysis ( Fig. 5 A)."

reach
"The interaction between BAP1 and BRCA1 and their roles in AKI were further examined in vivo ."

reach
"Therefore, we opine that the BAP1/BRCA1 interaction inactivates the NF-κB signaling to protect septic AKI.In conclusion, this study demonstrates that BAP1 protects mice from sepsis-induced AKI by deub[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The proteasome has both ubiquitin ligases and DUBs that associate with it (Crosas et al., 2006) , and several DUB-ligase pairs interact directly, including BRCC36-BRCA1, BAP1-BRCA1, USP4-Ro52, USP7-MDM2, USP8-GRAIL, USP20-pVHL, USP33-pVHL and USP44-APC (Kee and Huibregtse, 2007; Marfany and Denuc, 2008; Ventii and Wilkinson, 2008) ."
BRCA1 binds BARD1 and BAP1. 7 / 7
| 2 5

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear and needs to be studied in more detail."

reach
"Until recently, the combination of pemetrexed and cisplatin was the only approved first-line therapy, yielding a median overall survival (OS) of 12.1 months.2 Nivolumab plus ipilimumab was approved recently as initial treatment for malignant pleural mesothelioma (MPM) on the basis of a phase 3 clinical trial that resulted in a median OS of 18.1 months.3 4 There are no approved therapies for patients with tumor relapse after first-line therapies.Risk factors in developing mesothelioma include exposure to asbestos5 and other carcinogenic fibers such as erionite,6 and previous radiation therapy.7 8 Studies of familial clusters of mesothelioma have revealed genetic predisposition to develop MM.9 10 Recent work has identified germline mutations in BAP1 that can predispose to mesothelioma11 12 and other cancers such as uveal and cutaneous melanoma, basal cell carcinoma, meningioma, and renal cell cancers.13 14 We previously identified deleterious germline mutations in 12% of 241 consecutive patients with MM who enrolled on a prospective natural history study, with BAP1 being the most frequent germline mutation.15 BAP1 was also recurrently inactivated at the somatic level, suggesting BAP1 variants undergoing positive selection in the context of the classic “two-hit” model.BAP1, a nuclear deubiquitylase, was initially discovered as an interaction partner of the tumor suppressor BRCA1.16 Subsequent studies revealed that BAP1 binds to BARD1, which, in turn, binds to BRCA1, forming the BRCA1-BARD1-BAP1 complex that is crucial for the homologous recombination (HR)–mediated repair of DNA double-strand breaks (DSBs).17 PARP enzymes play a major role in DNA single-strand break repair and base excision repair pathways."

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear."

reach
"One feature of BAP1 is its association with BRCA1-associated RING domain 1 (BARD1) and BRCA1, forming the BRCA1-BARD1-BAP1 complex."

sparser
"The ring domains of BRCA1 (residues 1-109) are associated with BARD1 [ xref ] and BAP1 [ xref ]."

sparser
"BAP1 interacts with BARD1 and BRCA1 and inhibits the E3 ligase activity of BRCA1/BARD1."

sparser
"BAP1 and BRCA1-BARD1 complex coordinately regulate ubiquitination during the DNA damage response and the cell cycle [ xref , xref ]."
BAP1 activates BRCA1.
| 17
BAP1 activates BRCA1. 10 / 20
| 17

reach
"BAP1 can deubiquitylate Ub-H2AK119 at the site of UV-induced DNA damage, and promote DNA repair by regulating the accumulation of repair proteins (BRCA1, RAD51, and RPA) (7)."

reach
"To examine whether BAP1 mediates BRCA1 protein stability through deubiquitination modification, ubiquitination test was performed."

reach
"It was found that downregulation of BAP1 increased the degradation rate of BRCA1 protein in cells."

reach
"XREF_BIBR suggested that BAP1 could function for HR repair in the BRCA1 related pathway, and they hypothesized that BAP1 enables BRCA1 to readily accumulate DSB sites."

reach
"Initially identified in a yeast two-hybrid screen using the RING-finger domain of the breast cancer type 1 susceptibility protein BRCA1 as bait, BAP1 was shown to enhance BRCA1 mediated growth suppression in MCF7 cells [XREF_BIBR]."

reach
"Conceivably, BAP1 might modulate BRCA1 mediated tumor suppression by hydrolyzing ubiquitin moieties from the BRCA1 and BARD1 heterodimer, its associated proteins, or even its substrates [61]."

reach
"BRCA1-associated protein-1 (BAP1), a ubiquitin C-terminal hydrolase (UCH), was firstly identified as a DUB to mediate BRCA1 stability [15]."

reach
"Their studies revealed differential expression of the cancer-related lncRNAs Malat1, Neat1, H19, Meg3, and their associated miRNA-target pairs, regulating various hallmarks of tumorigenesis such as cell cycle and p53 signaling.BRCA1-Associated Protein 1 (BAP1) is known to enhance BRCA1 tumor suppressor potential and its inactivation is associated with various cancers, in particular clear-cell renal cell carcinoma (ccRCC)."

reach
"BRCA1-associated protein 1 (BAP1) is the translation product of BRCA1 and belongs to the family of deubiquitinating enzymes."

reach
"It is still not known whether BAP1 functions to modulate BRCA1 ubiquitynating activity and/or to regulate ubiquitin dependent degradation of BRCA1 itself."
BAP1 inhibits BRCA1.
| 15
BAP1 inhibits BRCA1. 10 / 19
| 15

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"A previous study on the expression of BAP1 tumor suppressor gene in 1222 patients with TCGA breast cancer data reported that the expression of BAP1, which enhances BRCA1-mediated suppression of cell proliferation through BRCA1 stabilization, is highly correlated with USP19 expression."
| PMC

reach
"BAP1 enhances BRCA1 mediated suppression of cell growth in colony formation assays, and this suppression by BAP1 is augmented by its UCH enzymatic domain."

reach
"BAP1, a ubiquitin carboxy-terminal hydrolase, interacts with the RING finger domain of BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1, protecting BRCA1 from proteasome-mediated degradation (37, 38)."

reach
"Furthermore, the two proteins appeared to interact in a biologically significant manner because BAP1 augmented the growth suppressive properties of BRCA1 in breast cancer cells."

reach
"BAP1 enhances BRCA1 mediated suppression of cell proliferation through stabilizing BRCA1 [XREF_BIBR]."

reach
"XREF_BIBR It was initially suggested that BAP1 bound BRCA1 cleaves ubiquitin, and enhances the growth-suppressive effects of BRCA1."

reach
"We further found that BAP1 depletion suppressed the assembly of constitutive BRCA1 foci, which are associated with homologous recombination (HR), but had minimal effect on gamma-H2AX foci and did not affect proteins associated with nonhomologous end joining, suggesting that BAP1 affects radiosensitivity in HNSCC by modifying HR."

reach
"In our study, we found that BAP1 depletion decreased the assembly of constitutive BRCA1 foci, which are associated with HR, but had minimal effect on gamma-H2AX foci."

reach
"BAP1 enhances BRCA1-mediated inhibition of breast cancer cell growth and may be a tumor suppressor gene that functions in the BRCA1 growth control pathway ( Jensen et al., 1998 )."

reach
"Early experiments found that BAP1 enhanced BRCA1 mediated inhibition of breast cancer cell growth."
BAP1 increases the amount of BRCA1.
| 10
BAP1 increases the amount of BRCA1. 10 / 10
| 10

reach
"Noticeably, we found that exogenous BAP1 significantly increased the BRCA1 protein level."

reach
"It is worth noting that BAP1 can restore the protein expression of BRCA1."

reach
"BAP1 depletion reduced the quantity of both RAD51 and BRCA1 foci without affecting the respective protein levels and exerted no effect on the formation of 53BP1 or autophosphorylated DNA-PK foci, suggesting that BAP1 functions in HR but not in NHEJ ."

reach
"Importantly, artificial upregulation of BAP1 restored the protein level of BRCA1 in cells, while it had little effect on the BRCA1 mRNA expression.Following the findings above, we then focused on the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Importantly, it was found that overexpression of BAP1 in the RTECs significantly elevated the protein level of BRCA1 and suppressed phosphorylation of NF-κB, though it did not affect the total protein[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"XREF_BIBR previously suggested that BAP1 enhanced BRCA1 mediated inhibition of cancer cell growth, although they did not directly establish the possibility that BAP1 enhances the BRCA1 protein level."

reach
"Although BRCA1 expression significantly decreased in an MM cell line with BAP1 deletion, Y-MESO-25, the transduction of the WT and even mutant forms of BAP1 induced upregulation of the BRCA1 protein level."

reach
"Indeed, previous global mRNA expression profiling suggests that BAP1 depletion can perturb BRCA1 mRNA levels [27]."

reach
"BAP1 mediated growth suppression is independent of wild-type BRCA1 expression."

reach
"BAP1, a member of the DUB family, has been shown to directly up-regulate the expression of BRCA1, a tumor suppressor in cancer cells [153] ."
BAP1 deubiquitinates BRCA1.
1 | 3
BAP1 deubiquitinates BRCA1. 4 / 4
1 | 3

"<span class="match term0">BAP1</span> also controls G1/S cell cycle progression by regulating <span class="match term1">BRCA-1</span> [111], Ying Yang 1 (YY-1), and host cell factor 1 (HCF-1)"

reach
"Therefore, we opine that the BAP1/BRCA1 interaction inactivates the NF-κB signaling to protect septic AKI.In conclusion, this study demonstrates that BAP1 protects mice from sepsis-induced AKI by deub[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We hypothesized that BAP1 might deubiquitinate BRCA1 and protect it from proteasome mediated degradation; this might explain the BRCA1 stabilization by the mutant BAP1 forms that retained the DUB activity."

reach
"As a possible mechanism of BRCA1 stabilization by BAP1 transduction in this study, we speculated that the ubiquitination of BRCA1 protein might be directly deubiquitinated by BAP1."
BAP1 is modified
10 | 1 125 23
BAP1 is phosphorylated.
10 | 63 23
BAP1 is phosphorylated. 10 / 59
| 47 12

rlimsp
"(e) Exogenous BAP1 increased both BRCA1 protein and its phosphorylation levels."

sparser
"Irradiation of MM cells induces phosphorylation of BAP1."

sparser
"Such dissociation of DUBs induced by ATM-dependent phosphorylation has been reported for another UCH family DUB, BAP1, the substrate of which is ubiquitylated H2A. Phosphorylation of BAP1 that promotes HR resulted in reduced chromatin binding of BAP1 xref ."

sparser
"Moreover, ODN-induced mitochondrial ROS generation was involved in DNA damage, BAP1 phosphorylation, upregulation of Sp1 and Bax, sensitization of anti-cancer drug-mediated apoptosis."

sparser
"Analysis of the BAP1 phosphorylation sites showed that the S292 locus of BAP1 had a higher phosphorylation level in most cancers (all p < 0.01, except for renal clear cell carcinoma), followed by locu[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"PLK-3-mediated phosphorylation of BAP1 prevents diabetic retinopathy."

sparser
"First, when MeT-5A cells, which harbor WT BAP1 , were irradiated, we observed the strongest BAP1 phosphorylation level 1 h after IR, which then gradually decreased (Fig. xref )."

sparser
"In this matter, the regulation of its ubiquitination and the activity of the kinases responsible for BAP1 phosphorylation are not well defined and could be altered in the tumoral context."

sparser
"After IR, we found that both WT and mutant BAP1 were similarly phosphorylated and phospho-BAP1 localized mainly in the nucleus."

sparser
"Q209L and BAP1 p."
BAP1 is phosphorylated on S592. 10 / 21
2 | 14 5

sparser
"The C-terminal domain of p53 (residues 293–393) induced Sp1 stabilization strongly linked to BAP1 Ser 592 phosphorylation."

sparser
"These findings indicated that cathepsin K inhibition increases Sp1 stabilization via BAP1 Ser 592 phosphorylation."

sparser
"In vivo and in vitro assays indicated that the forced expression of PLK-3 increased the phosphorylation of BAP1 at serine 592 and caspase-8 expression."

sparser
"A large-scale proteomic analysis identified BAP1 phosphorylation at Ser592, an SQ/TQ motif in ATM/ATR kinase consensus phosphorylation site, after IR xref ."

sparser
"ODN induced the phosphorylation of BAP1 at Ser 592 and enhanced the interaction of Sp1 and BAP1, resulting in Sp1 deubiquitination and destabilization."

sparser
"Phosphorylation of BAP1 at S592 is an important regulatory mechanism to dissociate BAP1 from chromatin and regulate-specific genes during DNA replication and repair xref ."

sparser
"In particular, the phosphatidylinositol 3-kinase-related kinases ATM/ATR/DNA-PK phosphorylate BAP1 at S592, which is one of the five serines in its carboxyl terminus that are modified in response to DNA damage [ xref , xref , xref , xref ]."

rlimsp
"Eletr et al.27 reported that BAP1 is phosphorylated at serine 592 following DNA damage, that phospho-BAP1 causes deubiquitination of H2AK119ub1, and that this signal promotes the HR pathway."

sparser
"Furthermore, BAP1 is phosphorylated at Ser592 in response to HU-induced replication stress xref ."

sparser
"BAP1 is also phosphorylated at serine 592 in response to DNA damage."
BAP1 is phosphorylated on S597. 4 / 4
4 |

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No evidence text available

No evidence text available

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BAP1 is phosphorylated. 4 / 4
| 4

rlimsp
"Of note, BAP1 is a substrate of the ATM/ATR checkpoint kinases, and it is phosphorylated upon several stressors, including IR, UV light and replication stress (Matsuoka et al., 2007; Eletr et al., 2013; Ismail et al., 2014; Yu et al., 2014)."

rlimsp
"Phosphorylation of BAP1 that promotes HR resulted in reduced chromatin binding of BAP1."

rlimsp
"BAP1 phosphorylation and its own catalytic activity are required for its recruitment to double-strand DNA break lesions and promotion of DNA repair by homologous recombination [35]."

rlimsp
"The central region of BAP1 is predicted to be disordered. Sites within this central region have been identified as important for binding non-ASXL complex components; for example, the HCF-binding motif and the phosphorylated threonine residue that mediates BAP1's interaction with FOXK2 have been mapped to this region [96,97]. Additionally, phosphorylation of six sites within this region has been shown to be important for BAP1's role in DNA double-strand break repair [88]."
BAP1 is phosphorylated on S583. 3 / 3
2 | 1

No evidence text available

No evidence text available

sparser
"However, several studies have shown that BAP1 Ser583 is phosphorylated in human cells after UV light exposure, providing functional significance of the serine at position 583 ( xref , xref )."
BAP1 is phosphorylated on S395. 3 / 3
2 | 1

No evidence text available

No evidence text available

sparser
"Double-stranded DNA damage resulted in prominent nuclear expression of BAP1 in DPM cells and phosphorylation of BAP1 at serine 395."
BAP1 is phosphorylated. 2 / 2
| 2

rlimsp
"We also found that both BAP1 catalytic activity and its phosphorylation are critical for promoting DNA repair and cellular recovery from DNA damage."

rlimsp
"Finally, we identified six IR-induced phosphorylation sites in BAP1 and showed that mutation of these residues inhibits BAP1 recruitment to DSB sites."
BAP1 is ubiquitinated.
| 1 32
BAP1 is ubiquitinated. 10 / 33
| 1 32

sparser
"BAP1 ubiquitination affinity is increased by DEUBAD domains (DEUBiquitinase ADaptor) of ASXL1/2/3 ."

sparser
"Moreover, we found that VCP inhibitors inhibited CCA cell growth and promoted cell apoptosis by blocking BAP1 ubiquitination degradation."

sparser
"Ubiquitylation of BAP1 near its nuclear localization sequence negatively regulates its activity by excluding BAP1 from the nucleus, where most of its targets reside [84] ."

sparser
"Investigated several E2s and DUBs demonstrated that UBE2O is specifically directed against ubiquitination of BAP1."

sparser
"The conserved region 1(CR1), conserved region 2 (CR2), and UBC domains of UBE2O are necessary for the ubiquitination of BAP1 because deletion of anyone of them abolishes the ability of UBE2O to bring ubiquitination of BAP1."

sparser
"Consistently, depletion of VCP markedly elevated the BAP1 ubiquitination level (Figure  xref )."

sparser
"Under normal growth conditions BAP1 is predominantly nuclear, whereas UBE2O is mainly cytoplasmic, suggesting that BAP1 ubiquitination is highly regulated."

sparser
"Further, deletion of N or C terminal regions of UBE2O has been found to entirely disrupt ubiquitylation of BAP1 [ xref ]."

sparser
"UBE2O could be imported into the nucleus to ubiquitinate BAP1 and promote its ubiquitin-mediated nuclear export as observed for other proteins ( Groulx and Lee, 2002; Li et al., 2003 )."

sparser
"Controlling UBE2O import itself would provide another level of regulation for the fine-tuning of BAP1 ubiquitination."
BAP1 is methylated.
| 28
BAP1 is methylated. 10 / 28
| 28

sparser
"Therefore, we investigated whether BAP1 body methylation may influence BAP1 gene expression."

sparser
"Aberrant methylation of BAP1 and SF3B1 is thought to be associated with UM metastasis ( xref )."

sparser
"Recently, also BAP1 methylation has been recognized as an important prognostic marker of UM metastasis."

sparser
"In the bone, skin, and liver, UM metastasis was found to be caused by aberrant methylation of BAP1 and SF3B1 ."

sparser
"However, despite this lack of effect, differences in BAP1 methylation between the MP41 and MP46 cell lines suggest that BAP1 expression may indeed be regulated by DNA methylation."

sparser
"Using The Cancer Genome Atlas (TCGA) database we analyzed BAP1 methylation in several tumor types, at nineteen sites spanning the entire BAP1 promoter, gene and 3′ untranslated region ( xref )."

sparser
"Similarly, no BAP1 methylation alterations were found in clear cell renal carcinoma xref ."

sparser
"We investigated BAP1 promotor methylation as a potential mechanism for epigenetic inactivation in 19 subjects with sufficient tumor and non-tumor tissues including the proband 7002 with downregulation of BAP1 in non-tumor choroid."

sparser
"Pyrosequencing showed no significant BAP1 promotor methylation in any of the 19 studied subjects including the subject with significant constitutional downregulation of BAP1 in the non-tumor tissue."

sparser
"BAP1 methylation has been recognized as a significant prognostic indicator for metastasis in UM."
BAP1 is glycosylated.
| 2
BAP1 is glycosylated. 2 / 2
| 2

sparser
"Glycosylation of BAP1 controls the self-renewal of hematopoietic stem cells and hematopoiesis [ xref ]."

sparser
"Consistent with this, Jensen et al , who first described BAP1, identified a number of putative N‐linked BAP1 glycosylation sites, suggesting that BAP1 may traverse the Golgi during maturation xref ."
BAP1 affects HCFC1
25 1 | 70 44
BAP1 binds HCFC1.
25 | 49 44
25 | 45 39

reach
"BAP1 bound to HCF-1 and BAP1 mutations did not correlate with ubiquitinated H2A (H2Aub1) levels."

reach
"Complex formation between BAP1 and the cell cycle regulator, HCF-1, bridges HCF-1 to histone-modifying complexes to regulate cell growth (2,4)."

reach
"Additionally, BAP1/HCF-1 complex can recruit either O-linked N-acetylglucosamine transferase (OGT), the Ying Yang 1 transcriptional repressor (YY1), or forkhead transcription factors (FOXK1/K2) to form distinct multiprotein complexes (6,7,49)."

No evidence text available

reach
"BAP1 interacts with the polycomb factors ASXL1 and 2 and its substrates HCF-1 and OGT, but the precise mechanism how its loss of function bypasses growth arrest in BRAF mutated nevi remains to be elucidated."

sparser
"BAP1 is believed to have gained additional functions in eukaryotes because, unlike Calypso, it contains an HCF-1 binding motif (HBM) known to mediate BAP1 binding to HCF-1 in mice and humans [ xref , xref ]."

reach
"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."

reach
"Considering the strong association between BAP1 and HCF -1 and HCF-1 and E2Fs described above, we can conclude that ubiquitination of HCF-1 prevents activation of E2F responsive promoters."

reach
"The HCFC1 and BAP1 complex contributes to cell cycle progression and BAP1 deubiquitinating activity is probably involved in HCFC1 turnover."

sparser
"To determine whether the interaction of BAP1 with HCF-1 was physiologically significant, we analyzed tumorgrafts."
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
OGT binds HCFC1 and BAP1. 2 / 2
| 2

reach
"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
BAP1 deubiquitinates HCFC1.
1 | 12
BAP1 deubiquitinates HCFC1. 10 / 13
1 | 12

reach
"Furthermore, BAP1 also deubiquitinates its main interacting protein, host cell factor 1 (HCF-1) [107] ."

reach
"Machida et al. reported that HCFC1 was deubiquitinated by the deubiquitinating enzyme BRCA1-associated protein 1 to regulate cell divisions and proliferation 39."

reach
"10 Deubiquitination of HCF1 and H2Aub by BAP1 promotes their association with EF2, YY-1, and FoxK2, which regulate cell-cycle progression, cell proliferation and cell-cycle control."

reach
"It has been proposed that BAP1 may modulate cell proliferation by deubiquitylating the transcriptional regulator HCF1."

reach
"BAP1 also promoted cell proliferation through directly interacting and deubiquitinating host cell factor-1 (HCF-1) [12] ."

reach
"HCF-1 is modified with ubiquitin in vivo, and ectopic studies suggest BAP1 deubiquitinates HCF-1."

reach
"BAP1 also interacts with and deubiquitinates the transcriptional regulator host cell factor 1 (HCF1)."

reach
"Moreover, BAP1 modulates histone H2A and HCFC1 deubiquitination and regulates transcription, cell cycle and growth, response to DNA damage and chromatin dynamics."

reach
"BAP1 can deubiquitinate HCF-1 and is crucial for HCF-1 activity ( Zhou et al., 2013 )."

reach
"It has been proposed that ubiquitylation of HCF1 blocks E2F responsive promoter activity, and that HCF1 deubiquitylation by BAP1 would remove this inhibition and promote cell proliferation 33."
BAP1 activates HCFC1.
| 4
BAP1 activates HCFC1. 4 / 5
| 4

reach
"In a mouse model, absence of one such shared binding partner, BAP1, led to a lack of HCFC1 protein (since BAP1 normally prevents HCFC1 degradation), and these mice exhibited features of myeloproliferative disease."

reach
"Among these substrates, OGT and HCF-1 levelswere found to decrease upon Bap1 deletion in mice (Dey et al., 2012), supporting a previous report on the stabilization of HCF-1 by BAP1 (Machida et al., 20[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The BAP1 nuclear deubiquitinase is known to target histones (together with ASXL1 as a Polycomb repressor subunit) and the HCF1 transcriptional co-factor, and we show that BAP1 knockdown in MPM cell lines affects E2F and Polycomb target genes."

reach
"BAP1 is a deubiquitinase targeting histones and the host cell factor-1 transcriptional co-factor, and plays a role in transcriptional regulation, chromatin modulation, cell cycle regulation, and DNA repair."
BAP1 increases the amount of HCFC1.
| 4
BAP1 increases the amount of HCFC1. 4 / 4
| 4

reach
"OGT and HCF-1 levels were decreased by Bap1 deletion, indicating a critical role for BAP1 in stabilizing these epigenetic regulators."

reach
"OGT and HCF-1 levels were decreased by Bap1 deletion, and were associated with myeloid transformation ."

reach
"BAP1 deletion has been shown to decrease the OGT and HCF1 levels, suggesting a significant role for BAP1 in stabilizing these epigenetic regulators ( Carbone et al., 2013 )."

reach
"In contrast, deletion of BAP1 decreases protein levels of both OGT and HCF-1 and induces MDS in mice [84] ."
BAP1 inhibits HCFC1.
| 1
BAP1 inhibits HCFC1. 1 / 2
| 1

reach
"BAP1 loss induces a modest over-accumulation of HCF-1, leading G1 to S phase cell cycle transition, promoting cell proliferation (4)."
BAP1 affects Neoplasms
| 3 132
BAP1 inhibits Neoplasms.
| 3 94
| 3 94

reach
"Tumor suppressors such as p53, BRCA1-associated protein 1 (BAP1), and Kelch-like ECH-associated protein 1 (KEAP1) have been shown to exert their tumor-suppressive functions via inducing ferroptosis.P53 is regarded as the most critical barrier for cancer development by its cell-cycle surveillance function."

reach
"One explanation for the limited palette of tumors promoted by BAP1 loss is that it activates a specific apoptotic program in other cell backgrounds (18)."

reach
"BAP1-inactivated melanocytoma (BIM) is a novel subgroup of melanocytic neoplasm listed in the 5th edition of WHO classification of skin tumor."

reach
"BAP1 inhibits tumor development, in part, via SLC7A11 and ferroptosis [48]."

reach
"For example, the expression of the system xc− antiporter subunit SLC7A11, required for cystine uptake and GSH synthesis, is inhibited by the tumor suppressors BAP1 , p53 , and Kelch-like ECH-associated protein 1 (KEAP1) ."

reach
"Moreover, various tumor suppressors, such as p53 and BRCA1-associated protein 1 (BAP1), have been discovered to deter cancer progression by obstructing ferroptosis [9]."

reach
"The BAP1 gene encodes BRCA1-associated protein 1, which suppresses tumors by promoting the activity of the Hippo tumor suppressor pathway[20]."

reach
"Furthermore, the genetic studies have helped to separate different entities, like BAP1-inactivated melanocytic tumors, which were initially considered a subgroup of ST since their histologic features overlap with STUMP ."

reach
"BAP1-inaktivierter melanozytärer Tumor: ein Fallbericht: BAP1 inactivated melanocytic tumor: a case report."

reach
"A deterministic role for BAP1 has been shown in mice, where Bap1 loss induces more proliferative, higher-grade tumors (Gu et al., 2017)."
BAP1 activates Neoplasms.
| 38
| 33

reach
"Through a biochemical screen, we identified BAP1 catalytic inhibitors that inhibit truncated-ASXL1-driven leukemic gene expression and impair tumor progression in vivo."

reach
"Therefore, it will be important to investigate the mechanism underlying the tumor-promoting effect of BAP1 inactivation in each cell type.In summary, this study demonstrated that Bap1 loss promotes tumorigenesis only in the tissues in which Bap1 is not engaged in pro-survival gene expression, providing an explanation for the tissue specificity of the BAP1 cancer predisposition syndrome."

reach
"Pathogenic germline variants in the BRCA1-associated protein 1 (BAP1) gene cause a dominantly inherited tumor predisposition syndrome, BAP1-TPDS, in which MM is the second most common associated cancer."

reach
"Tumor suppressor BAP1 (BRCA1-associated protein 1) promotes ferroptosis and inhibits tumor development via a similar mechanism [92]."

reach
"Germline or somatic mutations and deletions of BAP1 have been identified in various tumor types, and downregulation or inactivation of BAP1 could accelerate tumor onset, invasion, recurrence, and metastases ."

reach
"The results showed that knocked down BAP1 rescued the promoting effect of overexpressed KLF5 on volume and weight of tumor in nude mice ( Fig. 5 E)."

reach
"Significantly, it was recently revealed that xCT is involved in tumor initiation as it is directly repressed by p53 or BAP1 as a means to exert tumor suppression (27, 28)."

reach
"Pathogenic germline variants in the BRCA1-Associated Protein 1 (BAP1) tumor suppressor gene cause a rare, life-threatening tumor predisposition syndrome (BAP1-TPDS, OMIM 614327) also known as BAP1 cancer syndrome as the majority of carriers develop malignancy [1, 2]."

reach
"By contrast, BAP1-driven tumors showed higher levels of immune infiltration, especially by myeloid cells, with low levels of endothelial cells."
| PMC

reach
"Furthermore, BAP1 germline mutations cause a tumor predisposition syndrome inherited in an autosomal dominant pattern, which confers a higher risk for developing a spectrum of benign and malignant tumors , including pancreatic cancer ."
Mutated BAP1 activates Neoplasms. 5 / 5
| 5

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"Whereas BAP1 mutations drive BAP1 tumour predisposition syndrome, mesotheliomas, melanomas and other neoplasms 8–11 ; ASXL1 mutations are particularly prevalent in myeloid cancers, and are associated [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Herein, we set out to investigate whether BAP1 mutations could regulate the development in UM, and our results demonstrated that BAP1 mutations could induce the tumor immune microenvironment in UM by inactivating the NF-κB signaling pathway."

reach
"Moreover, BAP1 mutations may elevate tumor responsiveness to immunotherapy (36)."

reach
"Cutaneous melanoma, melanocytic tumors and other skin tumors:. The frequency of cutaneous melanoma (CM) development is likely to increase with germline BAP1 mutations and tumor-promoting effects of these BAP1 mutations are strongly influenced by co-occurrence of associated oncogenes such as mutant BRAF (2,43)."

reach
"For example, mesothelioma cells that carry germline BAP1 mutations have upregulated aerobic glycolysis (17), and BAP1 mutant ccRCC tumor samples display elevated glycolysis (22)."
YUH1 affects BAP1
| 132
| 130

sparser
"SEC-SAXS data of BAP1-UCH variants were collected using the BL23A beamline at the National Synchrotron Radiation Research Center (NSRRC) in Hsinchu, Taiwan as described previously."

sparser
"A stable ternary complex is formed, comprised of the BAP1-UCH, BAP1-ULD, and ASXL2-AB domains."

sparser
"This study investigates the impacts of four cancer‐associated mutations, N78S, C91W, F81V, and G128R, on the aggregation kinetics of BAP1UCH."

sparser
"Nevertheless, the restorations of the structure and DUB activity of BAP1-UCH were only partial ( Fig. 2 ) whereas its paralog UCH-L1 shows complete structural restoration and approximately 60% functio[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"These ubiquitination events are actively counteracted by BAP1 self-deubiquitination which, importantly, could only be observed when the BAP1 UCH domain interacts with the CTD ( xref B )."

sparser
"16,18 According to the COSMID (Catalogue of Somatic Mutations in Cancer) database, 60% of all cancer-associated mutations are located within BAP1-UCH."

sparser
"Of note, the BAP1-UCH loop is larger than the ones in other UCH proteins, with a high conservation of both these loop amino acids and of multiple amino acids structurally near this loop ( xref , xref , xref ), implying that larger substrates may be available to BAP1’s catalytic cleavage site than for other UCH domains, yielding a BAP1-specific recruitment of proteins/domains such as ASX and ULD for enzyme regulation ( xref )."

sparser
"This could be attributed to the fact that WT BAP1UCH is marginally stable at 37 °C in vitro."

sparser
"In contrast to primary nucleation, secondary nucleation is influenced by local environmental fluctuations and molecular noise, which can lead to variable onset times and heterogeneous spatial distribution of aggregates in cells, as reflected in the variability observed in global fitting of k 2 and n 2 parameters for BAP1UCH variants (Table  xref and S2, Supporting Information)."

sparser
"Therefore, we initiated biochemical and biophysical analyses of the BAP1-UCH, BAP1-ULD, ASXL2-AB domains and protein complex."
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
| 1 65 4

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"Depletion of BAP1 expression in cutaneous melanoma cells reduced proliferation and induced apoptosis, inhibiting tumor growth in vivo, but its overexpression in melanocytes suppressed proliferation [21]."

reach
"A previous study on the expression of BAP1 tumor suppressor gene in 1222 patients with TCGA breast cancer data reported that the expression of BAP1, which enhances BRCA1-mediated suppression of cell proliferation through BRCA1 stabilization, is highly correlated with USP19 expression."
| PMC

reach
"BaP1, a phenoxazine derivative, reduces cell proliferation, survival and cell migration through ROS generation, apoptosis promotion and angiogenesis inhibition."

reach
"Rescue expression of BAP1, but not BAP1-C91A in two neuroblastoma cells lines reduced the proliferation rate at 48 and 72h, compared with control cells."

reach
"We hypothesize that loss of BAP1 function in human splenic littoral cells causes proliferation through mechanisms that are currently unknown."

reach
"BAP1 loss promotes the proliferation, migration and invasion of PDAC cells."

sparser
"Additionally, BAP1 inhibits cell proliferation by deubiquitinating host cell factor-1 (HCF1) [ xref ]."

reach
"Wild-type BAP1 but not mutant BAP1 significantly inhibited HCC cell proliferation, invasion, epithelial-mesenchymal transition (EMT) in vitro, and tumor progression and metastasis in vivo."

reach
"Additionally, BAP1 inhibits cell proliferation by deubiquitinating host cell factor-1 (HCF1) [XREF_BIBR]."

reach
"Likewise, cell proliferation, G1-phase to S-phase cell cycle progression, and invasion of HCCC9810 cells were significantly inhibited by transfection of wild-type BAP1 but not inactive mutant BAP1C91A (Fig. 4c, d)."

reach
"Therefore, BAP1 might be serve as a potential target for high-risk NB with amplified MYCN.However, Sime et al. found that BAP1 overexpression could also induce apoptosis to retard BE2C cells proliferation by releasing Bax through 14–3–3 protein interaction and reducing the Bcl-2 expression regardless of BAP1 DUB activity [28]."

reach
"In conclusion, BAP1 is a KLF5 DUB and BAP1 promotes basal breast cell proliferation, migration, tumour growth and metastasis partially through stabilizing KLF5 (XREF_SUPPLEMENTARY)."

reach
"For example, BAP1 expression can promote pro-oncogenic pathways, including breast cancer cell proliferation and radioresistance in head and neck cancer (19, 20)."

reach
"BAP1 depletion has also been shown to reduce cell proliferation through S-phase accumulation in MPM (14), while it is suggested to function primarily as a metastasis suppressor in uveal melanoma (21)."

reach
"Downregulation of BAP1 promotes proliferation, cell cycle progression, and invasion of ICC in vitro."

reach
"Similarly, Qin et al. showed that knockdown of BAP1 in several breast cancer cell lines inhibited cell proliferation, tumorigenicity and metastasis."

reach
"Taken together, these data revealed that downregulation of BAP1 promotes proliferation, G1 to S cell cycle progression, and invasion of ICC in vitro."

reach
"BAP1 loss reduced HAP1 cell adhesion/spreading (Fig. 1D) and proliferation (Fig. 1E), considered here to be readouts of cell fitness."

reach
"In summary, despite the findings implicating BAP1 in suppression of proliferation XREF_BIBR, XREF_BIBR, some groups have reported that BAP1 promotes cell proliferation XREF_BIBR, XREF_BIBR, XREF_BIBR."

reach
"Consistently, Asada et al. reported that BAP1 depletion through CRISPR/Cas9 system substantially inhibited proliferation of several human AML cell lines with ASXL1 mutations [ 36 ]."
HCFC1 affects BAP1
25 | 49 44
25 | 45 39

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"BAP1 bound to HCF-1 and BAP1 mutations did not correlate with ubiquitinated H2A (H2Aub1) levels."

reach
"Complex formation between BAP1 and the cell cycle regulator, HCF-1, bridges HCF-1 to histone-modifying complexes to regulate cell growth (2,4)."

reach
"Additionally, BAP1/HCF-1 complex can recruit either O-linked N-acetylglucosamine transferase (OGT), the Ying Yang 1 transcriptional repressor (YY1), or forkhead transcription factors (FOXK1/K2) to form distinct multiprotein complexes (6,7,49)."

No evidence text available

reach
"BAP1 interacts with the polycomb factors ASXL1 and 2 and its substrates HCF-1 and OGT, but the precise mechanism how its loss of function bypasses growth arrest in BRAF mutated nevi remains to be elucidated."

sparser
"BAP1 is believed to have gained additional functions in eukaryotes because, unlike Calypso, it contains an HCF-1 binding motif (HBM) known to mediate BAP1 binding to HCF-1 in mice and humans [ xref , xref ]."

reach
"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."

reach
"Considering the strong association between BAP1 and HCF -1 and HCF-1 and E2Fs described above, we can conclude that ubiquitination of HCF-1 prevents activation of E2F responsive promoters."

reach
"The HCFC1 and BAP1 complex contributes to cell cycle progression and BAP1 deubiquitinating activity is probably involved in HCFC1 turnover."

sparser
"To determine whether the interaction of BAP1 with HCF-1 was physiologically significant, we analyzed tumorgrafts."
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
OGT binds HCFC1 and BAP1. 2 / 2
| 2

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"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
BRCA1 affects BAP1
5 1 | 41 58
5 1 | 39 53

sparser
"The region of BAP1 that binds to BRCA1 spans amino acids 596–721, while the region that interacts with BARD1 is from 182 to 365 [ xref ]."

sparser
"Hence, as BAP1 protein interacts with BRCA1 protein, PARP inhibitors might also be useful in the treatment of patients with somatic BAP1 inactivation."

reach
"BRCA1 also interacts with BAP-1 (BRCA1 associated protein) [28], a member of the ubiquitin C-terminal hydrolase family, that removes ubiquitin from proteins."

reach
"XREF_BIBR It was initially suggested that BAP1 bound BRCA1 cleaves ubiquitin, and enhances the growth-suppressive effects of BRCA1."

sparser
"Thus, NUMB enhances Notch signaling by regulating the ubiquitinating activity of the BAP1BRCA1 complex."

reach
"Importantly, artificial upregulation of BAP1 restored the protein level of BRCA1 in cells, while it had little effect on the BRCA1 mRNA expression.Following the findings above, we then focused on the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"First, the interaction between BAP1 and BRCA1 in RTECs was validated by Co-IP and Western blot analysis ( Fig. 5 A)."

reach
"The interaction between BAP1 and BRCA1 and their roles in AKI were further examined in vivo ."

reach
"Therefore, we opine that the BAP1/BRCA1 interaction inactivates the NF-κB signaling to protect septic AKI.In conclusion, this study demonstrates that BAP1 protects mice from sepsis-induced AKI by deub[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The proteasome has both ubiquitin ligases and DUBs that associate with it (Crosas et al., 2006) , and several DUB-ligase pairs interact directly, including BRCC36-BRCA1, BAP1-BRCA1, USP4-Ro52, USP7-MDM2, USP8-GRAIL, USP20-pVHL, USP33-pVHL and USP44-APC (Kee and Huibregtse, 2007; Marfany and Denuc, 2008; Ventii and Wilkinson, 2008) ."
BRCA1 binds BARD1 and BAP1. 7 / 7
| 2 5

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear and needs to be studied in more detail."

reach
"Until recently, the combination of pemetrexed and cisplatin was the only approved first-line therapy, yielding a median overall survival (OS) of 12.1 months.2 Nivolumab plus ipilimumab was approved recently as initial treatment for malignant pleural mesothelioma (MPM) on the basis of a phase 3 clinical trial that resulted in a median OS of 18.1 months.3 4 There are no approved therapies for patients with tumor relapse after first-line therapies.Risk factors in developing mesothelioma include exposure to asbestos5 and other carcinogenic fibers such as erionite,6 and previous radiation therapy.7 8 Studies of familial clusters of mesothelioma have revealed genetic predisposition to develop MM.9 10 Recent work has identified germline mutations in BAP1 that can predispose to mesothelioma11 12 and other cancers such as uveal and cutaneous melanoma, basal cell carcinoma, meningioma, and renal cell cancers.13 14 We previously identified deleterious germline mutations in 12% of 241 consecutive patients with MM who enrolled on a prospective natural history study, with BAP1 being the most frequent germline mutation.15 BAP1 was also recurrently inactivated at the somatic level, suggesting BAP1 variants undergoing positive selection in the context of the classic “two-hit” model.BAP1, a nuclear deubiquitylase, was initially discovered as an interaction partner of the tumor suppressor BRCA1.16 Subsequent studies revealed that BAP1 binds to BARD1, which, in turn, binds to BRCA1, forming the BRCA1-BARD1-BAP1 complex that is crucial for the homologous recombination (HR)–mediated repair of DNA double-strand breaks (DSBs).17 PARP enzymes play a major role in DNA single-strand break repair and base excision repair pathways."

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear."

reach
"One feature of BAP1 is its association with BRCA1-associated RING domain 1 (BARD1) and BRCA1, forming the BRCA1-BARD1-BAP1 complex."

sparser
"The ring domains of BRCA1 (residues 1-109) are associated with BARD1 [ xref ] and BAP1 [ xref ]."

sparser
"BAP1 interacts with BARD1 and BRCA1 and inhibits the E3 ligase activity of BRCA1/BARD1."

sparser
"BAP1 and BRCA1-BARD1 complex coordinately regulate ubiquitination during the DNA damage response and the cell cycle [ xref , xref ]."
BAP1 affects ASXL2
23 | 26 49
BAP1 binds ASXL2.
23 | 22 49
23 | 22 44

No evidence text available

No evidence text available

sparser
"ASXL2-BAP1 interactions have been shown to play a role in the suppression of solid tumorigenesis (e.g., human mesotheliomas and lung cancers), and ASXL2-BAP1 complexes, similar to ASXL1-BAP1 complexes, demonstrated to interact with BAP1-interacting proteins (e.g., YY1, FOXK1/2, OGT, and HCF1) [ xref ]."

reach
"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."

sparser
"To validate interactions between the ASXL2 and BAP1 and to show that the ASXL2-AB box can regulate BAP1 deubiquitinase activity in cells, we performed co-IP-WB assays on purified proteins from transfected HEK 293 cells."

No evidence text available

sparser
"Based on the molecular dynamics, kinetics, and stoichiometry of the intra- and inter-molecule domain-domain interactions between BAP1 and ASXL2 presented here, we draw the following conclusions."

sparser
"First, all of the single- or co-expressed and purified recombinant BAP1 and ASXL2 domain/proteins or protein complexes from both bacteria and baculovirus are well-folded in structure and are functionally active."

sparser
"Second, the interaction between BAP1 and ASXL2 is direct, specific, and stable to biochemical and biophysical manipulations."

sparser
"With the exception of KDM1B, which appears to interact specifically with BAP1-ASXL2, the BAP1 interactome is essentially identical whether the complex forms around ASXL1 or ASXL2."
BAP1 binds ASXL1 and ASXL2. 3 / 3
| 3

sparser
"Although both ASXL1 and ASXL2 have been identified to bind to BAP1, recent biochemical characterization of BAP1 complexes by xref suggest that BAP1 forms two mutually exclusive complexes with ASXL1 and ASXL2."

sparser
"Proteomic analysis of BAP1 interacting partners using this system revealed that BAP1 interacts with ASXL1 and ASXL2 in vivo as well as host cell factor 1 (HCF-1), O-GlcNAc transferase (OGT), and Forkhead box protein K1/2 (FOXK1/2) ( xref )."

sparser
"Depletion of the PHD fingers did not affect the stability of ASXL1 or ASXL2, or the interaction between ASXL1, ASXL2, and BAP1 (Additional file xref : Fig. S2A, B)."
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
BAP1 ubiquitinates ASXL2.
| 4
BAP1 leads to the ubiquitination of ASXL2. 4 / 4
| 4

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"Thus, BAP1 promotes the ubiquitination of its co-factor ASXL2 regulating its stability, and this in turn regulates the enzymatic activity and function of the DUB complex."

reach
"BAP1 promotes ASXL2 monoubiquitination on its DEUBAD domain."

reach
"As BAP1 promotes ASXL2 monoubiquitination in vivo, we evaluated whether BAP1 with E2-conjugating enzymes are sufficient to direct this event in vitro."

reach
"Our results, obtained from cell-based studies, suggest that BAP1 might either directly stimulate ASXL2 monoubiquitination or preserve monoubiquitinated ASXL2 from degradation."
ASXL2 affects BAP1
23 | 25 49
ASXL2 binds BAP1.
23 | 22 49
23 | 22 44

No evidence text available

No evidence text available

sparser
"ASXL2-BAP1 interactions have been shown to play a role in the suppression of solid tumorigenesis (e.g., human mesotheliomas and lung cancers), and ASXL2-BAP1 complexes, similar to ASXL1-BAP1 complexes, demonstrated to interact with BAP1-interacting proteins (e.g., YY1, FOXK1/2, OGT, and HCF1) [ xref ]."

reach
"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."

sparser
"To validate interactions between the ASXL2 and BAP1 and to show that the ASXL2-AB box can regulate BAP1 deubiquitinase activity in cells, we performed co-IP-WB assays on purified proteins from transfected HEK 293 cells."

No evidence text available

sparser
"Based on the molecular dynamics, kinetics, and stoichiometry of the intra- and inter-molecule domain-domain interactions between BAP1 and ASXL2 presented here, we draw the following conclusions."

sparser
"First, all of the single- or co-expressed and purified recombinant BAP1 and ASXL2 domain/proteins or protein complexes from both bacteria and baculovirus are well-folded in structure and are functionally active."

sparser
"Second, the interaction between BAP1 and ASXL2 is direct, specific, and stable to biochemical and biophysical manipulations."

sparser
"With the exception of KDM1B, which appears to interact specifically with BAP1-ASXL2, the BAP1 interactome is essentially identical whether the complex forms around ASXL1 or ASXL2."
BAP1 binds ASXL1 and ASXL2. 3 / 3
| 3

sparser
"Although both ASXL1 and ASXL2 have been identified to bind to BAP1, recent biochemical characterization of BAP1 complexes by xref suggest that BAP1 forms two mutually exclusive complexes with ASXL1 and ASXL2."

sparser
"Proteomic analysis of BAP1 interacting partners using this system revealed that BAP1 interacts with ASXL1 and ASXL2 in vivo as well as host cell factor 1 (HCF-1), O-GlcNAc transferase (OGT), and Forkhead box protein K1/2 (FOXK1/2) ( xref )."

sparser
"Depletion of the PHD fingers did not affect the stability of ASXL1 or ASXL2, or the interaction between ASXL1, ASXL2, and BAP1 (Additional file xref : Fig. S2A, B)."
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
ASXL2 activates BAP1.
| 3
ASXL2 activates BAP1. 3 / 3
| 3

reach
"The DEUBAD domains of ASXL1, ASXL2 and ASXL3 can activate BAP1 by increasing its affinity for ubiquitin."

reach
"In this paper we describe that similar to the homologous UCH-L5 and RPN13 DEU complex, the DEUBAD domains of ASXL1, ASXL2 and ASXL3 can activate BAP1 by increasing BAP1 's affinity specifically for the ubiquitin in the substrate, through a combination of mild effects."

reach
"In mesothelioma, ASXL2 enhances BAP1’s tumor-suppressive activity through interactions with BAP1 and UBE2E ."
| 2 64
BAP1 activates apoptotic process.
| 1 51
| 1 51

reach
"Therefore, BAP1 might be serve as a potential target for high-risk NB with amplified MYCN.However, Sime et al. found that BAP1 overexpression could also induce apoptosis to retard BE2C cells proliferation by releasing Bax through 14–3–3 protein interaction and reducing the Bcl-2 expression regardless of BAP1 DUB activity [28]."

reach
"Inhibition of the neuronal BAP1/P53 axis significantly reduced neurological deficits and neuronal apoptosis and improved neurological dysfunction in mice after SAH."

reach
"Cytoplasm BAP1 has been described located at the endoplasmic reticulum (ER), and promoting apoptosis through regulating Ca releasing10."

reach
"In addition to apoptosis, BAP1 also modulates the activation of cysteine-dependent cell death, ferroptosis, by downregulating the expression of SLC7A11, the major transporter for extracellular cysteine uptake."

reach
"Also, overexpression of BAP1 promoted cell apoptosis, which was reversed by overexpression SLC7A11 (Figure 6B)."

reach
"As a deubiquitinating enzyme, BAP1 contributes to gene transcription, cell differentiation, DNA damage repair, apoptosis, and cell metabolism in tumor inhibition."

reach
"In a BAP1-mutated cell line olaparib plus cisplatin increased apoptosis and senescence suggesting potentiation of the effect of olaparib alone, consistent with observations in other tumor types [49] ."

reach
"BAP1 stabilized IP3R3 and increased mitochondrial Ca 2+ concentrations and apoptosis; BAP1 (C91S) did not (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"Depletion of BAP1 expression in cutaneous melanoma cells reduced proliferation and induced apoptosis, inhibiting tumor growth in vivo, but its overexpression in melanocytes suppressed proliferation [21]."

reach
"Overexpression of BAP1 significantly reduced cell viability (Figure 9A) and promoted cell apoptosis (Figure 9B), while CsA reversed these results."
| 1 13

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"2 Specific genes have also been implicated, in particular the tumor suppressor gene BAP1 , which promotes apoptosis through binding, deubiquitylating, and stabilizing calcium release into the cytosol [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Mito-TEMPO pretreatment inhibited ODN-induced BAP1 phosphorylation, increased Bax expression, and promoted apoptosis induced by the combined treatment of ODN and oxaliplatin (Fig. 9, Fig. 10E).5 Conclusion."

reach
"BAP1 depletion increases apoptosis in colon cancer cells."

reach
"Consistent with identification of BAP1 in CRISPR/Cas9 screens for fitness genes (17) and evidence that BAP1 loss promotes apoptosis on many cell backgrounds (18), our data suggest that complete deletion of BAP1 may be incompatible with MeT5A cell survival."

reach
"Moreover, the tumor suppressor BRCA1-associated protein 1 (BAP1) which localizes at the ER, promotes ER-mitochondrial Ca 2+ transfer and induces apoptosis by stabilizing IP3R3."

reach
"In RCC cells, BAP1 inhibited glucose deprivation-induced apoptosis through interaction with the activating transcription factor 3 (ATF3) and DNA damage-inducible transcript 3 protein (DDIT3) gene promoters in a polycomb complex protein BMI-1-dependent manner ."

reach
"The Arabidopsis BAP1 and BAP2 genes are general inhibitors of programmed cell death."

reach
"The Arabidopsis BAP1 and BAP2 genes are general inhibitors of programmed cell death."

reach
"49 Moreover, BAP1 induces apoptosis by binding to the 14‐3‐3 protein or suppresses apoptosis by stabilizing liver kinase B1 (LKB1)."

reach
"In vitro , overexpression of BAP1 reduced the LPS-induced injury and apoptosis in the RTECs."
| 62
| 31

reach
"There was a low transcript number for BRCA1 associated protein 1 (BAP1), which when highly expressed suppresses metastasis [XREF_BIBR]."

reach
"Our results reveal a novel route of transendothelial migration for uveal melanoma cells, and they provide insight into the mechanism by which loss of BAP1 promotes metastasis."

reach
"26 The exact mechanism (s) by which loss of BAP1 mediates primary uveal melanoma metastasis is currently being investigated."

reach
"19 However, GNA11 mutations are also more common in tumors involving the ciliary body, which is an independent risk factor for metastasis, and in tumors with mutations in the metastasis suppressor BAP1 (Breast cancer 1, early onset (BRCA1)-associated protein 1) (see below), so it remains possible or even likely that the association between GNA11 and metastasis is not causal."

reach
"XREF_BIBR, XREF_BIBR The precise molecular explanation for why loss of BAP1 leads to metastasis in UM remains unclear."

reach
"Knockdown of BAP1 in TNBC cells inhibited tumorigenesis and metastasis, which could be partially rescued by restoration of KLF5 expression, suggesting that KLF5 mediates, at least in part, the breast cancer promoting function of BAP1 [XREF_BIBR]."

reach
"BAP1 promotes progenitor cell differentiation into melanocytes by increasing MITF expression and BAP1 loss may promote metastasis in UM by remodeling the epigenome to resemble that of stem-like migratory neural crest cells."

reach
"Differential expression of the tumor suppressor BAP-1, growth factor receptor IGF-1R, and immune checkpoints IDO and TIGIT could contribute to the relative over- and underrepresentation of cell types at different stages from normal choroidal tissue to metastasis."

reach
"XREF_BIBR, XREF_BIBR The precise reason why loss of BAP1 promotes uveal melanoma progression and metastasis remains unclear and is the subject of intense investigation."

reach
"These studies could also lead to context-selective discovery of novel therapeutic strategies and understanding whether BAP1 loss promotes metastasis."
| 31

reach
"Similarly, Qin et al. showed that knockdown of BAP1 in several breast cancer cell lines inhibited cell proliferation, tumorigenicity and metastasis."

reach
"Germline or somatic mutations and copy number alterations have been identified in several cancers and inactivation of BAP1 accelerates invasion, recurrence and metastases."

reach
"In conclusion, BAP1 is a KLF5 DUB and BAP1 promotes basal breast cell proliferation, migration, tumour growth and metastasis partially through stabilizing KLF5 (XREF_SUPPLEMENTARY)."

reach
"In this study, we discovered a novel positive regulatory mechanism for KLF5 and the functional mechanism by which BAP1 promotes breast cancer growth and metastasis."

reach
"Germline or somatic mutations and deletions of BAP1 have been identified in various tumor types, and downregulation or inactivation of BAP1 could accelerate tumor onset, invasion, recurrence, and metastases ."

reach
"Nevertheless, our results suggest that BAP1 promotes breast tumour growth and metastasis partially through stabilizing KLF5."

reach
"BAP1 deletion abrogates growth and metastasis of murine cutaneous melanoma."

reach
"BAP1 and KLF5 promote breast cancer migration, invasion and metastasis may not through p27; however, KLF5 is a transcription factor regulating numerous downstream target genes, such as FGF-BP and mPGES1."

reach
"BAP1 depletion can inhibit the growth and metastasis of hepatocellular carcinoma cells [40]."

reach
"BAP1 promotes breast cancer cell proliferation and metastasis by deubiquitinating KLF5."
Mutated BAP1 activates Neoplasm Metastasis. 4 / 4
| 4

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"BAP1 mutations increase the risk of UVM metastasis and contribute to poor prognosis, while EIF1AX mutations are often associated with good prognosis."

reach
"XREF_BIBR Therefore, both somatic mutation of BAP1 in primary UM tumors and germline mutations increase the risk of metastasis and impart low survival rate."

reach
"Germline BAP1 mutations increased risk of metastasis independent of ciliary body involvement (P =.02)."

reach
"Moreover, in uveal melanoma, somatic BAP1 mutations cause metastasis with a probability of 74%, while the same has been observed in only 36% of the patients carrying a germline mutation [104]."
BAP1 affects KLF5
7 1 | 35 11
BAP1 binds KLF5.
7 | 14 11
7 | 14 11

reach
"The deubiquitinase (DUB) BAP1 had been reported to regulate KLF5 stability in breast cancer cells, so we hypothesize that LINC00152 mediates KLF5 depending on the binding of BAP1 and KLF5."

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"As shown in XREF_SUPPLEMENTARY, LINC00152 knockdown inhibits BAP1 binding with KLF5."

sparser
"Another KLF5 DUB that stood out in siRNA screening was BAP1 (BRCA1-associated protein 1), which also directly binds KLF5 and stabilizes KLF5 through deubiquitination [ xref ]."

sparser
"The deubiquitinase (DUB) BAP1 had been reported to regulate KLF5 stability in breast cancer cells ( xref , xref ), so we hypothesize that LINC00152 mediates KLF5 depending on the binding of BAP1 and KLF5."

sparser
"As shown in xref , LINC00152 knockdown inhibits BAP1 binding with KLF5."

reach
"Another KLF5 DUB that stood out in siRNA screening was BAP1 (BRCA1 associated protein 1), which also directly binds KLF5 and stabilizes KLF5 through deubiquitination [XREF_BIBR]."

sparser
"LncRNA PVT1 enhanced KLF5 protein stability by increasing the interaction between KLF5 and the deubiquitinase BAP1 [ 26 ]."

reach
"BAP1 interacts directly with KLF5 and stabilizes KLF5 via deubiquitination."

reach
"BAP1 interacts directly with KLF5."

sparser
"Recently, Jia et al. ( xref ) showed that the WWP1- BAP1-KLF5 axis is dysregulated in melanoma cancer; WWP1 was lowly expressed in melanoma cells and tissues whereas KLF5 and BAP1 were highly expressed and closely related to tumor metastasis."
BAP1 deubiquitinates KLF5.
1 | 8
BAP1 deubiquitinates KLF5. 9 / 9
1 | 8

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"Compared with GST and BAP1-C91S, BAP1 specifically decreased KLF5 polyubiquitination (XREF_FIG; XREF_SUPPLEMENTARY)."

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"For instance, BAP1 deubiquitinates and stabilizes the transcription factor KLF5, which is highly expressed in breast cancer and a potent biomarker for unfavorable prognosis in breast cancer patients, and promotes breast cancer cell proliferation, survival, and migration as well as tumor growth ."

"<span class="match term0">BAP1</span> interacts directly with <span class="match term1">KLF5</span> and stabilizes <span class="match term1">KLF5</span> via deubiquitination"

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"In breast cancer cell lines, BAP1 plays an oncogenic function by directly deubiquitinating and stabilizing KLF5 (Kruppel-like factor 5)."

reach
"BAP1 has been reported to promote breast cancer cell proliferation and metastasis by deubiquitinating KLF5 [ 18 ]."

reach
"BAP1, but not BAP1-C91S, markedly decreased KLF5 protein polyubiquitination (XREF_FIG, lanes 3 and 4), whereas another DUB, A20, did not decrease KLF5 polyubiquitination (XREF_FIG, lane 5)."

reach
"Finally, we demonstrated that knockdown of endogenous BAP1 increased the ubiquitination of endogenous KLF5 in HCC1806 (XREF_FIG)."

reach
"Second, BAP1 decreases KLF5 polyubiquitination and increases KLF5 protein stability in a DUB activity dependent manner."

reach
"Notably, KLF5 is de-ubiquitylated by BRCA1 associated protein-1 (BAP1) (Jia et al. 2021; Qin et al. 2015) and suppresses autophagy by activating the PI3K-AKT-mTOR signaling (Jia et al. 2021) that can inhibit the autophagy initiation and the autophagosome formation via modulating the activity of AMBRA1 and ULK1 complex (Xu et al. 2020; Nazio et al. 2013)."
BAP1 activates KLF5.
| 8
BAP1 activates KLF5. 8 / 8
| 8

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"However, BAP1 positively regulates KLF5 at the protein level."

reach
"For example, one study showed that BAP1 promotes the development of breast cancer by enhancing the stability of the transcription factor KLF5 (30)."

reach
"To further confirm whether BAP1 increases KLF5 protein stability, we transfected two different siRNAs targeting different regions of BAP1 into MCF10A and breast cancer cell lines (HCC1806 and HCC1937)."

reach
"To test whether BAP1 can antagonize E3 ligase mediated KLF5 degradation, we transfected KLF5 E3 ligases and BAP1 into HEK293FT cells and detected the KLF5 protein levels."

reach
"The results showed that knocked down BAP1 rescued the promoting effect of overexpressed KLF5 on volume and weight of tumor in nude mice ( Fig. 5 E)."

reach
"Collectively, these results suggest that BAP1 is a specific DUB for KLF5 and that BAP1 increases KLF5 protein stability in a DUB activity dependent manner."

reach
"These results indicate that BAP1 promotes KLF5 positive breast cell proliferation in vitro."

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"Taken together, these results demonstrate that BAP1 promotes HCC1806 breast cancer tumorigenesis partially by stabilizing KLF5."
BAP1 increases the amount of KLF5.
| 5
BAP1 increases the amount of KLF5. 5 / 5
| 5

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"Our data suggest that BAP1 promotes cell proliferation and migration, and enhances the expression of KLF5 and its downstream genes, including CyclinD1 and FGF-BP1, in the esophageal carcinoma cell line ECA109."

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"BAP1 knockdown decreased the endogenous protein levels of KLF5 and its downstream target gene FGF-BP XREF_BIBR (XREF_FIG)."

reach
"The KLF5 mRNA levels were not decreased by BAP1 knockdown in the MCF10A and HCC1806 cells (XREF_SUPPLEMENTARY)."

reach
"Subsequent investigations showed that WWP1 overexpression in melanoma cells could mediate K48-linked ubiquitination of KLF5 and induce its proteasomal degradation, whereas BAP1 overexpression reverted this modification and increased KLF5 protein expression (Jia et al. 2021)."

reach
"In addition, expression of KLF5, CyclinD1, and FGF-BP1 was increased by BAP1 overexpression and decreased by BAP1 knockdown."
IHC affects BAP1
| 51
IHC inhibits BAP1.
| 20
IHC inhibits BAP1. 10 / 20
| 20

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"Both FISH for the p16 deletion and IHC loss of BAP1 could not be shown in either case and were therefore not helpful in making the final diagnosis."

reach
"The most commonly referred to markers in malignant mesothelioma are loss of BAP1 by IHC, and the deletion of p16 by FISH."

reach
"Long-term follow-up was not available for these series [5,11]; thus, the question remains as to whether these tumors are atypical Spitz tumors or sporadic BAP1 associated melanocytic lesions.In summar[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Another issue is that most mesotheliomas of peritoneal origin do show a loss of BAP1 by IHC but do not show loss of p16 by FISH."

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"In several studies, the loss of BAP1 by IHC and the homozygous deletion of p16 by FISH is very specific for mesothelioma but still may lack sensitivity to a certain extent."

reach
"In a recent study, BAP1 loss by IHC has been demonstrated to predict superior overall survival following first-line platinum-pemetrexed combination."

reach
"Given that BAP1 loss by IHC is observed in 15% of sporadic ccRCC (11), the probability that all 3 tumors would be BAP1 negative by chance alone is 0.0034."

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"There was a significant association between BRAF mutation status and loss of BAP1 by IHC."

reach
"It is worth emphasizing that morphology alone is insufficient for a diagnosis of MIS, while demonstration of BAP1 loss by IHC or CDKN2A/p16 homozygous deletion by FISH are required [13,24]."

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"Similarly, it is worth emphasizing that morphology alone is insufficient for diagnosis of MIS, and demonstration of BAP1 loss by IHC or CDKN2A homozygous deletion by FISH is required for diagnosis [13,24]."
IHC increases the amount of BAP1.
| 17
IHC increases the amount of BAP1. 10 / 17
| 17

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"Loss of BAP1 Expression by IHC Identifies a Morphologically Distinct Subset of Tumors."

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"These data are consistent with those of XREF_BIBR) for both patient survival based on the BAP1 mutational status (32 vs 133 months for BAP1 mutation positive vs BAP1 mutation negative tumours) or BAP1 protein expression (31 vs 133 months for tumours showing negative BAP1 expression by IHC vs those with positive BAP1 expression) and thus the proposed role of BAP1 in the pathogenesis of UM with an aggressive phenotype."

reach
"Our observation that a greater number of tumors exhibited loss of BAP1 protein expression by IHC compared with BAP1 deletions or mutations indicates that BAP1 may become functionally inactivated by mechanisms other than deletions or mutations in the coding region; for example, epigenetic changes leading to silencing of the BAP1 gene or other, yet to be identified, alterations that prevent BAP1 expression."

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"2.2 PBRM1 and BAP1 Protein Expression by IHC."

reach
"However, none of these harbored deletions of p16/CDKN2A in reactive stroma, which could allow for more accurate differentiation of BM from EM with reactive stroma when in combination with BAP1 expression by IHC."

reach
"In this pilot study, we investigated whether loss of BAP1 expression by IHC is associated with prognosis in uveal melanoma patients."

reach
"As anticipated, we identified a significant association of p16 loss with poor outcomes that was independent of histologic subtype, and as previously reported, that BAP1 expression by IHC is not an ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Other malignancies instead express normal levels of BAP1 : for example BAP1 is expressed and detected by IHC in most pancreatic carcinomas [XREF_BIBR], and in most peritoneal and gynecologic serous adenocarcinomas [XREF_BIBR]."

reach
"In this pilot study, we inves- tigated whether loss of BAP1 expression by IHC is associated with prognosis in uveal melanoma patients."

reach
"In the current pilot study, we explored the hypothesis that loss of BAP1 expression by IHC is associated with prognosis in patients with uveal melanoma."
IHC decreases the amount of BAP1.
| 14
IHC decreases the amount of BAP1. 10 / 14
| 14

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"The melanomas that have been tested show biallelic BAP1 inactivation.34 One of the melanomas described in this setting arose in association with an epithelioid naevus (similar to those described above[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In this pilot study, we inves- tigated whether loss of BAP1 expression by IHC is associated with prognosis in uveal melanoma patients."

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"The loss of BAP1 expression by IHC is a reliable tool to assess this molecular alteration."

reach
"Recently, loss of BAP1 (BRCA1-associated protein-1) expression by IHC, homozygous deletion of CDKN2A (p16) by FISH, and expression of methyl-thio-adenosine phosphorylase (MTAP) by IHC were added as markers to distinguish non-neoplastic from neoplastic cells when mesothelial proliferation is confined to the serosal surface."

reach
"In the current pilot study, we explored the hypothesis that loss of BAP1 expression by IHC is associated with prognosis in patients with uveal melanoma."

reach
"XREF_BIBR, XREF_BIBR Loss of PBRM1 or BAP1 expression by IHC is associated with a worse RCC specific survival in ccRCC supporting a role for distinguishing molecular subtypes of ccRCC."

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"Both the naevus and melanoma components showed loss of BAP1 expression by IHC, XREF_BIBR suggesting that some BAP1 mutated epithelioid melanocytic tumors may progress to melanoma."

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"In absence of invasion, loss of BAP1 expression by IHC and/or the presence of a homozygous deletion of CDKN2A (p16) by FISH or by methylthioadenosine phosphorylase (MTAP) IHC allow a differential diagnosis between neoplastic and non-neoplastic cells (16-18)."

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"Here we report loss of BAP1 expression by IHC in 61% of MPM cases overall, with 77% of epithelioid cases demonstrating BAP1 loss and retention in all of the sarcomatoid cases examined."

reach
"Our observation that a greater number of tumors exhibited loss of BAP1 protein expression by IHC compared with BAP1 deletions or mutations indicates that BAP1 may become functionally inactivated by mechanisms other than deletions or mutations in the coding region; for example, epigenetic changes leading to silencing of the BAP1 gene or other, yet to be identified, alterations that prevent BAP1 expression."
BAP1 affects OGT
10 1 | 21 17
BAP1 binds OGT.
10 | 12 17
10 | 10 17

No evidence text available

sparser
"Consistent with previous reports [2,5,7] , the endogenous interaction between BAP1 and OGT was confirmed by IP with the anti-BAP1 antibody and WB analysis using the anti-OGT antibody in HEK293 cells ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

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No evidence text available

sparser
"Likewise, our finding of the BAP1OGT interaction prompted us to address whether BAP1 is O -GlcNAcylated by OGT."

reach
"Consistent with previous reports [2,5,7] , the endogenous interaction between BAP1 and OGT was confirmed by IP with the anti-BAP1 antibody and WB analysis using the anti-OGT antibody in HEK293 cells ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In this study, we confirmed that BAP1 interacts with OGT in vivo and investigated the biological significance of this interaction."

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"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."
OGT binds HCFC1 and BAP1. 2 / 2
| 2

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"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

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"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
BAP1 increases the amount of OGT.
| 5
BAP1 increases the amount of OGT. 5 / 6
| 5

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"BAP1 also increases OGT protein levels, suggesting that BAP1 is a potential deubiquitinase for OGT as well (XREF_SUPPLEMENTARY)."

reach
"BAP1 deletion has been shown to decrease the OGT and HCF1 levels, suggesting a significant role for BAP1 in stabilizing these epigenetic regulators ( Carbone et al., 2013 )."

reach
"In contrast, deletion of BAP1 decreases protein levels of both OGT and HCF-1 and induces MDS in mice [84] ."

reach
"OGT and HCF-1 levels were decreased by Bap1 deletion, and were associated with myeloid transformation ."

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"OGT and HCF-1 levels were decreased by Bap1 deletion, indicating a critical role for BAP1 in stabilizing these epigenetic regulators."
BAP1 deubiquitinates OGT.
1 | 4
BAP1 deubiquitinates OGT. 5 / 5
1 | 4

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"To investigate whether BAP1 can deubiquitylate and stabilize OGT directly, rather than decreased OGT being secondary to a reduction in HCF-1, we affinity purified ubiquitylated and Myc tagged human OGT from HEK293T cells, and then incubated it with human BAP1 and human ASXL1 residues 2-365 co-purified from Sf9 cells."

"The tumor suppressor <span class="match term0">BAP1</span> associates with ASXL1/2 to form the core Polycomb complex PR-DUB, which catalyzes the removal of mono-ubiquitin from several substrates including histone H2A. This complex also mediates the poly-deubiquitination of HCFC1, <span class="match term1">OGT</span> and PCG1-alpha, preventing them from proteasomal degradation."

reach
"OGT is a well-known protein associated with BAP1 in the BAP1 core complex, which include ASXL1, ASXL2, HCF1, OGT, and the forkhead transcription factors FOXK1 and FOXK2 [9], and BAP1 can deubiquitinate and stabilize OGT [26]."

reach
"XREF_BIBR, XREF_BIBR, XREF_BIBR BAP1 also deubiquitylates OGT and regulates its levels."

reach
"BAP1 deubiquitinates and stabilizes OGT [75], while CDK1 phosphorylates KIF18A during mitosis at S674/S684."
BAP1 affects Mesothelioma
| 47
BAP1 activates Mesothelioma.
| 25
| 18

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"Interestingly, it has been shown that germline mutations in Bap1 can induce epigenetic deregulation of the Rb protein in mice, facilitating the development of malignant mesothelioma [72]."

reach
"On the other hand, the occasionally described heterozygous deletions of these genes cannot be completely excluded in our series, because NGS procedure and mode of data analysis were not sensitive to moderate CNV (copy number variation), the detection of which was not the aim of this panel.Germline mutations in BAP1 are known to underlie sometimes melanomas, renal cell carcinomas, malignant mesotheliomas, and some other cancers [31]."

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"In addition, a mutation in the BRCA1-associated protein-1 (BAP1) gene is implicated in UM metastasis, and is also known to promote the development of cutaneous melanoma, basal cell carcinoma, malignan[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Inactivation of one BAP1 allele causes a higher incidence of peritoneal mesotheliomas than wild-type littermates, in mice exposed to asbestos fibers."

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"The authors demonstrated a possible role of the polycomb repressive complex 2 (PRC2) in Bap1-mediated mesothelioma."

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"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (frequently in the abdomen) and uveal melanoma [151,152,153]."

reach
"BAP1 PGVs accelerate mesothelioma tumorigenesis after asbestos exposure in preclinical models and may be partly predicted by clinical criteria."

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"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (often in the abdomen), and uveal melanoma [97]."

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"The data show that the top 2 mutant genes in both groups are BAP1 and NF2, which is consistent with the conclusion that the inactivation of BAP1 with loss of NF2 or CDKN2A promotes the development of malignant mesothelioma in approximately 20% of mice with double conditional knockout (CKO), as already reported in the previous literature."

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"BAP1 germline mutations underlie a rare, autosomal dominant tumour predisposition syndrome that is most commonly associated with (in order of decreasing frequency) atypical Spitz tumours, uveal melano[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Mutated BAP1 activates Mesothelioma. 7 / 7
| 7

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"The identification of BAP1 mutations in MPM has led to investigate the role of histone deacetylase inhibitors in mesothelioma patients.The phase III trial on 661 pretreated patients randomly assigned [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"High frequency of germline mutations of BAP1 were demonstrated to cause mesothelioma in 2001, when an epidemic spread of cases was reported in a village in Cappadocia (113, 118)."

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"Germline BAP1 mutations cause mesothelioma and other malignancies, namely, uveal cancer, meningioma, and melanoma, overall defined as “BAP1- related cancer syndrome”."

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"Whereas BAP1 mutations drive BAP1 tumour predisposition syndrome, mesotheliomas, melanomas and other neoplasms 8–11 ; ASXL1 mutations are particularly prevalent in myeloid cancers, and are associated [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"BAP1 mutations cause prevalently mesothelioma and melanomas."

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"BAP1 mutations can lead to mesothelioma in situ in multiple serosal cavities."

reach
"Mutation of the BAP1 gene is known to induce diffuse malignant mesothelioma [29]."
BAP1 inhibits Mesothelioma.
| 22
| 22

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"Inactivation of BAP1 sensitises tumour cells to EZH2 inhibition, which has been shown to substantially inhibit tumour growth in BAP1 -deficient malignant pleural mesothelioma mouse models."

reach
"EZH2 inhibitor tazemetostat have been proven effective in patients with relapsed or refractory, BAP1-inactivated malignant pleural mesothelioma in a multicentre, open-label, phase 2 study (Zauderer et al., 2022)."

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"Loss of BAP1 and MTAP can support the diagnosis of mesothelioma versus benign mesothelial proliferation: however, morphology (with an invasive pattern) is still the essential tool to distinguish malignant from benign disease.Molecular analyses: FISH for alterations in CDKN2A and NF2 (as IHC for BAP1 and MTAP) can be used to support the diagnosis of mesothelioma versus benign mesothelial proliferation."

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"However, TG2-179-1 kills both BAP1-insensitive and BAP1-depleted colon cancer cells as well as BAP1-null mesothelioma and ccRCC cells, indicating that TG2-179-1 targets BAP1 as well as other cellular proteins."

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"Although limited by low sample size, the latter observation suggests there may be functional crosstalk between Hippo pathway deregulation and loss of BAP1 , where a combination of both may drive an ag[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Individuals with pathogenic BAP1 variants face an elevated risk of diverse tumors, notably BAP1-inactivated melanocytic nevi/tumors, uveal melanoma, cutaneous melanoma, mesothelioma, renal cell carcinoma, and basal cell carcinoma."

reach
"Similar triple-knockout mouse models also confirmed that, although Bap1 deletion alone did not induce mesothelioma, Bap1 deletion dramatically accelerated mesothelioma development with the combined disruption of Nf2 and Cdkn2ab (Badhai et al., 2020)."

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"18 BAP-1 loss of expression might support the diagnosis of mesothelioma, both on cytological samples and on small biopsies, and can differentiate MPM from reactive mesothelial proliferations."

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"Based on its interaction with BRCA1, BAP1 has been shown to increase sensitivity to PARP inhibitors in patients with mesothelioma."

reach
"Thus, Ezh2 deletion or inhibition was shown to abrogate myeloid malignancy and mesothelioma induced by Bap1 loss (LaFave et al. 2015)."
BAP1 affects FOXK2
18 | 11 18
BAP1 binds FOXK2.
18 | 8 18
18 | 8 18

sparser
"In addition, the N-terminal portions (amino acids 8–190) including FHA domain are required for the interaction of FOXK2 with BAP1 [ 48 ]."

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

reach
"Sites within this central region have been identified as important for binding non ASXL complex components; for example, the HCF binding motif and the phosphorylated threonine residue that mediates BAP1 's interaction with FOXK2 have been mapped to this region [XREF_BIBR, XREF_BIBR]."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."

No evidence text available
BAP1 inhibits FOXK2.
| 3
BAP1 inhibits FOXK2. 3 / 3
| 3

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"Loss of BAP1 causes the increase of FoxK2 target genes, which is dependent on the Ring1B and Bmi1 complex."

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"Moreover, BAP1 is part of a ternary complex bridging FOXK2 and HCF1 and represses FOXK2 target genes, an effect requiring BAP1 DUB activity but not the interaction with HCF1 binding."

reach
"BAP1 represses FoxK2 target genes, and this effect requires BAP1 DUB activity but not interaction with HCF-1."
PBRM1 affects BAP1
| 7 39
| 7 32

sparser
"BAP1 -mutant tumors and PBRM1 -mutant tumors are also associated with different outcomes in patients ( xref )."

sparser
"Complementing the analysis of the clinical significance of mutations in PBRM1 and BAP1 in ccRCC, multiple groups, including ours, assessed the association of PBRM1 and BAP1 protein expression with clinical outcomes."

sparser
"However, how those lncRNAs are interacting with PBRM1 and BAP1 remains to be explored."

sparser
"With first-line everolimus, PBRM1 and BAP1 mutations were associated with longer (median [95% confidence interval {CI}] 12.8 [8.1, 18.4] vs 5.5 [3.1, 8.4] mo) and shorter (median [95% CI] 4.9 [2.9, 8.1] vs 10.5 [7.3, 12.9] mo) PFS1L, respectively."

sparser
"The association of a PBRM1 or BAP1 mutation with better or worse PFS1L, respectively, with everolimus is surprising because higher mTORC1 activity was reported for patients with BAP1 versus PBRM1 mutant ccRCC [ xref ]."

sparser
"BAP1 and PBRM1 mutations are associated with characteristic and non-overlapping gene expression signatures, and outcomes for patients with ccRCCs mutated in BAP1 or PBRM1 are different [ xref ]."

sparser
"These data provide evidence of negative selection of PBRM1BAP1− tumors consistent with negative genetic interaction between PBRM1 and BAP1."

sparser
"Genetically, a previous report suggests that combined loss of BAP1 and PBRM1 may be associated with rhabdoid morphology."

sparser
"We previously validated IHC assays with Sanger sequencing to evaluate PBRM1 and BAP1 protein expression in which negative nuclear staining associates with PBRM1 and BAP1 mutations. xref , xref We used background stroma and lymphocyte nuclei as a positive control, and therefore, we excluded from our analysis samples that lacked stroma/lymphocyte nuclear staining, focal negative or weak positive staining in these cells ( xref )."

sparser
"While PBRM1 and BAP1 expression have been preliminarily associated with prognostic outcome, their predictive utility remains unclear. xref However, compelling data regarding the predictive utility of KDM5C mutation have recently been presented."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
BAP1 affects ferroptosis
| 2 44
BAP1 activates ferroptosis.
| 2 35
| 2 35

reach
"Consequently, BAP1-mediated downregulation of SLC7A11 leads to elevated lipid peroxidation and ferroptosis [88]."

reach
"Interestingly, solute carrier family 7 member 11 (SLC7A11) is a crucial regulated protein of ferroptosis, which has been reported as the target gene of BAP1.10 33 Therefore, we further verified the mechanism of BAP1 and SLC7A11 in H/R-induced cardiomyocyte ferroptosis."

reach
"BAP1, as a tumor suppressor, promotes ferroptosis and inhibits cancer cell growth by inhibiting SLC7A11 [94]."

reach
"It has been recently reported that BAP1 facilitates lipid peroxidation and promotes ferroptosis through repressing SLC7A11 [24]."

reach
"For example, p53 and BRCA1-related protein 1 (BAP1) induce ferroptosis in tumor cells through multiple signaling pathways, which act as a natural barrier to cancer development (5, 6)."

reach
"BAP1 induced mPTP opening reverses the inhibiting role of BMSC-derived Exos oe-GATA-4 on H/R-induced cardiomyocytes ferroptosis by interacting with IP3R."

reach
"BAP1 can promote ferroptosis and suppress tumor tumorigenesis [39]."

reach
"BAP1 can induce ferroptosis in a similar process to TP53 by downregulating SLC7A11 [57]."

reach
"Moreover, another study suggested that BAP1 upregulation is essential for ferroptosis induced by 3,3′-diindolylmethane in BGC-823 cells, suggesting that BAP1-induced ferroptosis could be one of the potential mechanisms by which it suppresses GC progression [113]."

reach
"These results suggested that BAP1 can mediate SLC7A11 inhibition and neuronal ferroptosis by regulating H2Aub.3.3 FOXO3a regulated BAP1 expression at the transcriptional level."
BAP1 inhibits ferroptosis.
| 9
| 9

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"Moreover, BAP1 partially prevents the progression of tumors via SLC7A11 as well as ferroptosis, and cancer-related BAP1 mutants end up losing their capacities to suppress SLC7A11 and to stimulate ferroptosis [25]."

reach
"Tumor suppressor BAP1 (BRCA1-associated protein 1) promotes ferroptosis and inhibits tumor development via a similar mechanism [92]."

reach
"Zhang et al. showed that BAP1 inhibits tumor progression by promoting cellular ferroptosis (15)."

reach
"BAP1 inhibits tumor development, in part, via SLC7A11 and ferroptosis [48]."

reach
"Furthermore, the tumor suppressor BRCA1-associated protein 1 suppresses SLC7A11 transcription by decreasing H2Aub, leading to elevated lipid peroxidation and thus, increased ferroptosis (49)."

reach
"In addition, Zhang et al. discovered that another tumor suppressor BAP1 promoted ferroptosis and inhibited the growth of cancer cells by inhibiting SLC7A11 (49)."

reach
"Also, BAP1 reversed the effects of Exos oe-GATA-4 on H/R-induced cardiomyocytes' ferroptosis by downregulating SLC7A11."

reach
"It has been reported that the inactivation of BAP1 upregulates SLC7A11 and suppresses ferroptosis, resulting in tumor development."

reach
"Several tumor-suppressors, such as p53 [69] , BAP1 [70] , and KEAP1 [71] , promote ferroptosis through inactivating SLC7A11, while some oncogenic signaling pathways such as PI3K-mTORC1 pathway have be[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects PBRM1
| 7 39
| 7 32

sparser
"BAP1 -mutant tumors and PBRM1 -mutant tumors are also associated with different outcomes in patients ( xref )."

sparser
"Complementing the analysis of the clinical significance of mutations in PBRM1 and BAP1 in ccRCC, multiple groups, including ours, assessed the association of PBRM1 and BAP1 protein expression with clinical outcomes."

sparser
"However, how those lncRNAs are interacting with PBRM1 and BAP1 remains to be explored."

sparser
"With first-line everolimus, PBRM1 and BAP1 mutations were associated with longer (median [95% confidence interval {CI}] 12.8 [8.1, 18.4] vs 5.5 [3.1, 8.4] mo) and shorter (median [95% CI] 4.9 [2.9, 8.1] vs 10.5 [7.3, 12.9] mo) PFS1L, respectively."

sparser
"The association of a PBRM1 or BAP1 mutation with better or worse PFS1L, respectively, with everolimus is surprising because higher mTORC1 activity was reported for patients with BAP1 versus PBRM1 mutant ccRCC [ xref ]."

sparser
"BAP1 and PBRM1 mutations are associated with characteristic and non-overlapping gene expression signatures, and outcomes for patients with ccRCCs mutated in BAP1 or PBRM1 are different [ xref ]."

sparser
"These data provide evidence of negative selection of PBRM1BAP1− tumors consistent with negative genetic interaction between PBRM1 and BAP1."

sparser
"Genetically, a previous report suggests that combined loss of BAP1 and PBRM1 may be associated with rhabdoid morphology."

sparser
"We previously validated IHC assays with Sanger sequencing to evaluate PBRM1 and BAP1 protein expression in which negative nuclear staining associates with PBRM1 and BAP1 mutations. xref , xref We used background stroma and lymphocyte nuclei as a positive control, and therefore, we excluded from our analysis samples that lacked stroma/lymphocyte nuclear staining, focal negative or weak positive staining in these cells ( xref )."

sparser
"While PBRM1 and BAP1 expression have been preliminarily associated with prognostic outcome, their predictive utility remains unclear. xref However, compelling data regarding the predictive utility of KDM5C mutation have recently been presented."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
OGT affects BAP1
10 | 17 17
OGT binds BAP1.
10 | 12 17
10 | 10 17

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sparser
"Consistent with previous reports [2,5,7] , the endogenous interaction between BAP1 and OGT was confirmed by IP with the anti-BAP1 antibody and WB analysis using the anti-OGT antibody in HEK293 cells ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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sparser
"Likewise, our finding of the BAP1OGT interaction prompted us to address whether BAP1 is O -GlcNAcylated by OGT."

reach
"Consistent with previous reports [2,5,7] , the endogenous interaction between BAP1 and OGT was confirmed by IP with the anti-BAP1 antibody and WB analysis using the anti-OGT antibody in HEK293 cells ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In this study, we confirmed that BAP1 interacts with OGT in vivo and investigated the biological significance of this interaction."

reach
"Knock-in mice expressing BAP1 with a 3xFlag tag revealed that BAP1 interacts with HCF-1, OGT, and the polycomb group proteins ASXL1 and ASXL2 in vivo."
OGT binds HCFC1 and BAP1. 2 / 2
| 2

reach
"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
OGT activates BAP1.
| 5
OGT activates BAP1. 5 / 5
| 5

reach
"Taken together, our data suggest that OGT-mediated O -GlcNAcylation, which likely targets BAP1, prompts BAP1 to repress RAR-induced gene activation.Given the positive role of OGT in maintaining the se[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Using in vivo and in vitro approaches, we examined the OGT-mediated O -GlcNAcylation of BAP1."

reach
"Reversely, OGT mediates the O-GlcNAcylation of BAP1 by interacting with BAP1 C-terminal region [20]."

reach
"In addition, positive crosstalk between BAP1 and OGT may exist because BAP1 stabilizes OGT as a deubiquitinase [12] and OGT potentiates BAP1 as an RAR repressor via O -GlcNAcylation.Although our data [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"To determine whether BAP1 O -GlcNAcylation is directly driven by OGT, we performed glycosylation assays in vitro using purified proteins."
FOXK2 affects BAP1
18 | 8 18
18 | 8 18

sparser
"In addition, the N-terminal portions (amino acids 8–190) including FHA domain are required for the interaction of FOXK2 with BAP1 [ 48 ]."

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reach
"Sites within this central region have been identified as important for binding non ASXL complex components; for example, the HCF binding motif and the phosphorylated threonine residue that mediates BAP1 's interaction with FOXK2 have been mapped to this region [XREF_BIBR, XREF_BIBR]."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."

No evidence text available
UBE2O affects BAP1
12 | 21 6
UBE2O ubiquitinates BAP1.
| 14 5
UBE2O ubiquitinates BAP1. 10 / 19
| 14 5

reach
"Strikingly, UBE2O ubiquitinates BAP1, and modified forms were more abundant in the C91S mutant."

reach
"UBE2O can also ubiquitinate the tumor-suppressive DUB BAP1 and regulate its subcellular localization and function in chromatin remodeling."

reach
"The reaction with methylated ubiquitin was as efficient as with the unmodified ubiquitin and resulted in a similar ubiquitination pattern, indicating that UBE2O multi-monoubiquitinates BAP1."

reach
"UBE2O monoubiquitinates multiple lysines on the BAP1 NLS, thereby inducing its cytoplasmic sequestration (45)."

reach
"Thus, substrate ubiquitination by UBE2O can induce cytoplasmic localization, which might be accompanied by further proteasomal degradation as a consequence of ubiquitin chain extension by other E3 lig[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Further, deletion of N or C terminal regions of UBE2O has been found to entirely disrupt ubiquitylation of BAP1 [XREF_BIBR]."

reach
"For example, UBE2O binds to and multi-monoubiquitinates BAP1 to induce its cytoplasmic localization and promotes adipocyte differentiation [XREF_BIBR]."

reach
"For example, UBE2O mediates multiple mono-ubiquitination of BAP1, a nuclear localization signal, and leads to its cytoplasmic sequestration [XREF_BIBR]."

reach
"While ubiquitination enzyme UBE2O monoubiquitinates the NLS of BAP1 and induces translocation (inactivation) of BAP1 from the nucleus to the cytoplasm, DUB activity of the UCH-domain of BAP1 counteracts the UBE2O activity mediated through the intramolecular interaction between the UCH-domain and COOH-terminal domain of BAP1."

reach
"Ubiquitination of BAP1 by UBE2O results in sequestration of BAP1 in the cytoplasm, rendering it unable to find its nuclear targets."
| PMC
UBE2O binds BAP1.
12 | 1 1
12 | 1 1

reach
"For example, UBE2O binds to and multi-monoubiquitinates BAP1 to induce its cytoplasmic localization and promotes adipocyte differentiation [XREF_BIBR]."

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sparser
"Since BAP1 5K/R mutant still interact with UBE2O and is ubiquitinated on other sites, we also replaced the NLS region of BAP1, containing the UBE2O-interacting motif, with T-large antigen NLS which pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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UBE2O activates BAP1.
| 6
UBE2O activates BAP1. 6 / 9
| 6

reach
"Of note, overexpression of UBE2O, but not its catalytic dead form, also promoted cytoplasmic localization of endogenous BAP1."

reach
"UBE2O mediates BAP1 subcellular location."

reach
"Besides, the proteasome inhibitor MG132 cannot influence the UBE2O-mediated multimonoubiquitination pattern of BAP1 in 293T cells co-transfected with UBE2O and BAP1 or C91S mutant."

reach
"Indeed, the subcellular localization of BAP1 is coordinated by the E3 ligase UBE2O which promotes BAP1 cytoplasmic retention ."

reach
"UBE2O-mediated ubiquitylation causes BAP1 to accumulate in the cytoplasm, whereas BAP1 autodeubiquitylation promotes nuclear retention."

reach
"The ubiquitin conjugating enzyme UBE2O induces BAP1 sequestration in the cytoplasm by multi monoubiquitylation of its nuclear localization signal (NLS)."
| 3 39
| 3 21

reach
"Because BAP1 is frequently mutated in UM cells and is closely associated with metastasis ( Harbour et al., 2013; Maerker et al., 2014 ), knockdown of BAP1 was expected to promote cell invasiveness."

reach
"In functional assays, BAP1 was found to inhibit the migration and invasion of LAC cells, and promoted their apoptosis and necrosis."

reach
"Moreover, both scratching assay and Matrigel invasion assay indicated that overexpression of BAP1 in HCCC9810 cells significantly reduced cell migratory and invasion capacities, while the motility and invasion potential of RBE-shBAP1 cells were evidently enhanced compared to RBE-Mock cells (Fig. 2d, e)."

reach
"Wild-type BAP1 but not mutant BAP1 significantly inhibited HCC cell proliferation, invasion, epithelial-mesenchymal transition (EMT) in vitro, and tumor progression and metastasis in vivo."

reach
"Importantly, loss of BAP1 decreased the formation of stress fibers and membrane protrusions and induced migration and invasion defects."

reach
"Likewise, cell proliferation, G1-phase to S-phase cell cycle progression, and invasion of HCCC9810 cells were significantly inhibited by transfection of wild-type BAP1 but not inactive mutant BAP1C91A (Fig. 4c, d)."

reach
"These results indicated that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways.To further validate these findings, we used inhibitors of the ERK1/2 pathway (U0126) and JNK/c-Jun pathway (SP600125) in RBE-shBAP1 cells."

reach
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways."

reach
"Importantly, loss of BAP1 decreased the formation of stress fibers and membrane protrusions and induced migration and invasion defects."

eidos
"Taken together , these data revealed that downregulation of BAP1 promotes proliferation , G1 to S cell cycle progression , and invasion of ICC in vitro ."

reach
"To further determine how loss of BAP1 accelerates B cell lymphoma cells growth in vivo via regulating immune cell infiltration and TME, we conducted single-cell RNA-Seq (scRNA-seq) with the tumor samples from both BAP1-WT and -KO tissues (Supplemental Figure 5A)."

reach
"Interestingly, loss of BAP1 expression was found to promote T cell infiltration in uveal melanoma [79]."

reach
"Interestingly, recent research has revealed that the gain of chromosomal 8q may worsen prognosis by activating macrophage infiltration, and the loss of BAP1 expression may drive T cell infiltration in UM (Gezgin et al., 2017), suggesting that immune infiltration plays a crucial role in the prognosis of UM.Emerging evidence shows that the immune microenvironment is crucial for cancer progression and response to therapeutics (Quail and Joyce, 2013)."
| 18

reach
"BRCA1 associated protein 1 deficiency in lung adenocarcinoma predicts poor outcome and increased tumor invasion."

reach
"Consistently, silence of BAP1 or KLF5 expression significantly decreased the invasion of HCC1937 cells in transwell matrigel invasion assays (XREF_FIG)."

reach
"Although BAP1 is a tumor suppressor when overexpressed, the knockdown of BAP1 in wild-type UM cells also suppressed cell growth and invasiveness ( Matatall et al., 2013 )."

reach
"This suggests BAP1 may cooperate with other genes to mediate cell invasion and metastasis.BAP1 was recently identified as a transcriptional coregulator that binds indirectly to promoters close to tran[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Germline or somatic mutations and deletions of BAP1 have been identified in various tumor types, and downregulation or inactivation of BAP1 could accelerate tumor onset, invasion, recurrence, and metastases ."

reach
"BAP1 and BCL9L, as oncogenes, promote the growth, invasion and lung metastasis of breast cancer by interacting with β-catenin and Pygopus (39-41)."

reach
"BAP1 may inhibit the progression of LAC by attenuating tumor invasiveness, a function that was supported by the findings of the Transwell assay."

reach
"Downregulation of BAP1 promotes proliferation, cell cycle progression, and invasion of ICC in vitro."

reach
"Germline or somatic mutations and copy number alterations have been identified in several cancers and inactivation of BAP1 accelerates invasion, recurrence and metastases."

reach
"The simultaneous occurrence of SF3B1 mutations and BAP1 deficiency in Mel202 cells reduced their invasion capacity in the zebrafish xenograft model."

reach
"Loss of BAP1 reduces immune cell infiltration and accelerates B cell lymphoma growth in vivo.."

reach
"We then investigated whether and how BAP1 modulates T lymphocyte infiltration in MPM."

reach
"Loss of BAP1 leads to a dramatic reduction of MHC-II gene expression and reduced immune cell infiltration in animal models, resulting in a substantial acceleration of tumor growth."

reach
"Lack of BAP1 increased the production of chemokines to attract T-cell aggregation, resulting in greater T-cell infiltration, a negative prognostic factor according to Broggi, G et al. [18]."

reach
"Mutations in BAP1, which were the most frequent in CCRCC among all cancers (Figure 3A), were suggested to be pathogenic and promote CD8 T cell infiltration in CCRCC."
BAP1 affects cell death
| 1 37 1
BAP1 activates cell death.
| 1 25
| 1 25

reach
"BAP1 induces cell death via interaction with 14-3-3 in neuroblastoma."

reach
"In contrast, at higher concentrations, BaP1 stains lipid droplets and induces a regulated cell death process mediated by vacuolar membrane permeabilization."

reach
"P-I metalloproteinases and LAAO arrested the cells in G0/G1 phase of cell cycle and also led to cytotoxic activity in endothelial cells by inducing necrosis/late apoptotic mode of cell death."

reach
"BAP1 promotes cell death in neuroblastoma cells."

reach
"The mechanism that BAP1 induces cell death is mediated via an interaction with 14-3-3 protein."

reach
"The intersection between BAP1-mediated cell death regulation and pregnancy complications provides a fertile ground for further exploration, holding promise for improving our understanding and management of pregnancy-related disorders."

reach
"Two different strategies were applied to analyze the mechanism of BAP1 inducing cell death in neuroblastoma."

reach
"For example, BAP1 can enhance progression through the G1-S checkpoint and subsequently induce cell death by a process with similarities to both apoptosis and necrosis [XREF_BIBR]."

reach
"Under glucose starvation, BAP1 can directly bind to the promoter of ATF3 and CHOP, inhibit its transcription, and thus inhibit the UPR and cell death induced by the ER stress (XREF_FIG p)."

reach
"Although this remains to be investigated, these results, suggest that the majority of BAP1 partners regulate SLC7A11 expression and possibility impact ferroptosis.Because ferroptosis and apoptosis are two distinct programs in BAP1-mediated cell death, it will be worth to investigate whether these processes act in concert or independently during cancer development."
BAP1 inhibits cell death.
| 12 1
| 12 1

reach
"The group of flg22 induced MPK4 dependent genes encode important regulators of plant defence such as the cell death inhibitor BAP1 [XREF_BIBR], the calcium dependent protein kinase CPK5 [XREF_BIBR], the exocyst complex subunit EXO70B2 [XREF_BIBR], the cyclic nucleotide gated ion channel CNGC11 [XREF_BIBR] or the BAK1 like receptor kinase BKK1 and SERK4 [XREF_BIBR]."

reach
"Arabidopsis C2 domain proteins, BAP1, and its homologue BAP2, negatively regulate biotic and abiotic cell death."

reach
"Changes in mitochondrial permeability are required for the activation of the apoptotic process and the loss of BAP1 protects the cells from caspase-induced cell death as a consequence of IP3R3 level reduction and Ca2 + signalling decrease [45]."

reach
"Here we show that BAP1 inhibits cell death induced by unresolved metabolic stress."

reach
"Arabidopsis C2 domain proteins, BAP1 and its homologue BAP2, negatively regulate biotic and abiotic cell death [12,13] ."

reach
"Thus, BAP1 might suppress tumorigenesis by facilitating tumor cell death by activation of both these pathways."

reach
"XREF_BIBR reported that BAP1 overexpression in the BAP1-null NCI-H226 human non-small-cell lung cancer cell line suppressed growth in DUB activity- and nuclear localization dependent manners by accelerating the cell cycle G1/S transition and inducing cell death."

reach
"The powerful activity of heterozygous germline BAP1 mutations to facilitate asbestos induced transformation and cause mesothelioma has been linked to the alterations in the mechanisms that regulate DNA repair and cell death caused by decreased BAP1 protein levels."

sparser
"Here we show that BAP1 inhibits cell death induced by unresolved metabolic stress."

reach
"He et al. showed that Bap1 knockout in mouse embryonic stem (ES) cells, fibroblasts, and keratinocytes caused cell death through the intrinsic apoptosis pathway (1), which is mediated by Bax/Bak oligomerization."
BAP1 affects BARD1
| 1 10 26
BAP1 binds BARD1.
| 1 8 26
| 1 6 18

sparser
"The loss of the ring finger domain abrogates the binding of BARD1 and BAP1, which are important regulators of E3 ubiquitin ligase activity of BRCA1."

sparser
"BAP1 binding to BARD1 results in inhibition of the E3 ligase activity of BRCA1/BARD1 via the prevention of RING heterodimer formation ( xref et al. , 2009)."

sparser
"BAP1, a ubiquitin carboxy-terminal hydrolase, interacts with the RING finger domain of BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1, protecting BRCA1 from proteasome-mediated degradation ( xref , xref )."

reach
"BAP1 binds and deubiquitylates BARD1, thus modulating the E3 ligase activity of BRCA1-BARD1 28."

sparser
"BAP1 binds and deubiquitylates BARD1, thus modulating the E3 ligase activity of BRCA1–BARD1 xref ."

reach
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1 and BARD1 complex, and that BAP1 and BRCA1 and BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway."

sparser
"BAP1 binds to the BRCA1-associated RING domain protein 1 (BARD1) and inhibits E3 ubiquitin ligase activity of the BRCA1/BARD1 complex, which modulates the DNA damage response ( xref )."

sparser
"BAP1 also directly interacts with BARD1."

sparser
"Several studies later demonstrated that BAP1 mainly interacts with BARD1 perturbing BARD1/BRCA1 complex [ xref ]."

sparser
"Domains comprised by residues 182-365 of BAP1 interact with the RING finger domain of BARD1, and surface plasmon resonance spectroscopy (BIAcore) analyses showed that BAP1 interferes with the BRCA1/BARD1 association."
BRCA1 binds BARD1 and BAP1. 7 / 7
| 2 5

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear and needs to be studied in more detail."

reach
"Until recently, the combination of pemetrexed and cisplatin was the only approved first-line therapy, yielding a median overall survival (OS) of 12.1 months.2 Nivolumab plus ipilimumab was approved recently as initial treatment for malignant pleural mesothelioma (MPM) on the basis of a phase 3 clinical trial that resulted in a median OS of 18.1 months.3 4 There are no approved therapies for patients with tumor relapse after first-line therapies.Risk factors in developing mesothelioma include exposure to asbestos5 and other carcinogenic fibers such as erionite,6 and previous radiation therapy.7 8 Studies of familial clusters of mesothelioma have revealed genetic predisposition to develop MM.9 10 Recent work has identified germline mutations in BAP1 that can predispose to mesothelioma11 12 and other cancers such as uveal and cutaneous melanoma, basal cell carcinoma, meningioma, and renal cell cancers.13 14 We previously identified deleterious germline mutations in 12% of 241 consecutive patients with MM who enrolled on a prospective natural history study, with BAP1 being the most frequent germline mutation.15 BAP1 was also recurrently inactivated at the somatic level, suggesting BAP1 variants undergoing positive selection in the context of the classic “two-hit” model.BAP1, a nuclear deubiquitylase, was initially discovered as an interaction partner of the tumor suppressor BRCA1.16 Subsequent studies revealed that BAP1 binds to BARD1, which, in turn, binds to BRCA1, forming the BRCA1-BARD1-BAP1 complex that is crucial for the homologous recombination (HR)–mediated repair of DNA double-strand breaks (DSBs).17 PARP enzymes play a major role in DNA single-strand break repair and base excision repair pathways."

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear."

reach
"One feature of BAP1 is its association with BRCA1-associated RING domain 1 (BARD1) and BRCA1, forming the BRCA1-BARD1-BAP1 complex."

sparser
"The ring domains of BRCA1 (residues 1-109) are associated with BARD1 [ xref ] and BAP1 [ xref ]."

sparser
"BAP1 interacts with BARD1 and BRCA1 and inhibits the E3 ligase activity of BRCA1/BARD1."

sparser
"BAP1 and BRCA1-BARD1 complex coordinately regulate ubiquitination during the DNA damage response and the cell cycle [ xref , xref ]."

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 inhibits BARD1.
| 2
BAP1 inhibits BARD1. 2 / 2
| 2

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"BAP1, a ubiquitin carboxy-terminal hydrolase, interacts with the RING finger domain of BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1, protecting BRCA1 from proteasome-mediated degradation (37, 38)."

reach
"BAP1 acts as a tumor suppressor and inhibits E3 ligase activity in BRCA1/BARD1."
BAP1 affects cell growth
| 34
BAP1 inhibits cell growth.
| 24
| 24

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"Whether BAP1 also affects E2F1 ubiquitination to regulate its activity and the expression of E2F1 downstream target genes remains to be clarified.Previous studies have demonstrated that the overexpres[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 enhances BRCA1 mediated suppression of cell growth in colony formation assays, and this suppression by BAP1 is augmented by its UCH enzymatic domain."

reach
"BAP1 was originally discovered as a binding partner for BRCA1 [XREF_BIBR] and shown to inhibit cancer cell growth."

reach
"That study also showed that BAP1 slows cell growth through altered G1-S checkpoint regulation."

reach
"BAP1 knockout enhances anchorage-independent cell growth in human mesothelial cells."

reach
"However, in the semi-soft agar colony formation assay, the number of colonies was significantly increased in the BAP1-KO cells (Fig. S1b), suggesting that BAP1 loss might enhance anchorage-independent cell growth in human mesothelial cells."

reach
"BAP1 enhances BRCA1-mediated inhibition of breast cancer cell growth and may be a tumor suppressor gene that functions in the BRCA1 growth control pathway ( Jensen et al., 1998 )."

reach
"18 BAP1 reintroduction into two different BAP1 deficient ccRCC cell lines reduced cell growth."

reach
"26 BAP1 knockdown using small interfering RNA (siRNA) in MPM cell lines inhibited cell growth, and resulted in inactivation of HCF1 and downregulation of downstream E2F responsive genes."

reach
"In a previous report, BAP1 increased cell growth, whereas abnormal activity of BAP1 inhibited cell growth in several cancers."
BAP1 activates cell growth.
| 10
| 10

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"The slight upregulation of MYCN protein level in BPA1 overexpressed NB cells and other substrates of BAP1 which exert predominant function in the BAP1 overexpression setting might be the underlying mechanisms for BAP1 expression-induced cell growth retardation."

reach
"In mesothelioma pathogenesis, BAP1 dysfunction contributes to altered cellular growth, enhanced survival, and resistance to apoptosis."

reach
"In cutaneous melanoma cell lines, however, stable over-expression of BAP1 promoted cell growth while knockdown of BAP1 suppressed cell proliferation with concomitant decrease of survivin, an anti-apoptotic protein [17]."

reach
"In a previous report, BAP1 increased cell growth, whereas abnormal activity of BAP1 inhibited cell growth in several cancers."

reach
"Depletion of BAP1 inhibits neuroblastoma cell growth."

reach
"It was reported that depletion of BAP1 in cells with normal expression level of BAP1 could decrease the cell growth and lead to a delayed G1 to S1 transition [34]."

reach
"Although BAP1 is a tumor suppressor when overexpressed, the knockdown of BAP1 in wild-type UM cells also suppressed cell growth and invasiveness ( Matatall et al., 2013 )."

reach
"In this study, we showed that BAP1 knockdown decreased DNA synthesis, the number of S-phase cells and cell growth in HCC1806 cells."

reach
"Furthermore, BAP1 knockout in clear cell renal cell carcinoma (ccRCC) inhibits cell growth and migration and promotes the mesenchymal-epithelial transition (MET) partially by downregulating Snail and reducing Rho family GTPases activity [40]."

reach
"When KLF5 is depleted, BAP1 failed to promote cell growth (XREF_SUPPLEMENTARY)."
BAP1 affects H2Aub
| 1 34
BAP1 deubiquitinates H2Aub.
| 11
BAP1 deubiquitinates H2Aub. 10 / 11
| 11

reach
"BAP1 deubiquitinates H2Aub."

reach
"In conclusion, we found that BAP1 can deubiquitinate H2Aub and induces radioresistance in HNSCC cells via modulation of HR."

reach
"Other studies in mammalian cells have also shown that BAP1 represses the expression of some target genes with BAP1 dependent deubiquitination of H2Aub on the corresponding gene promoters XREF_BIBR, XREF_BIBR."

reach
"BAP1 deubiquitinates histone 2A (H2Aub) on the SLC7A11 promoter and represses SLC7A11 expression in a p53-independent way (39, 40)."

reach
"At the SLC7A11 promoter, BAP1 deubiquitinates H2Aub and subsequently inhibits the expression of SLC7A11."

reach
"10 Deubiquitination of HCF1 and H2Aub by BAP1 promotes their association with EF2, YY-1, and FoxK2, which regulate cell-cycle progression, cell proliferation and cell-cycle control."

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"Although USP16 is clearly demonstrated to facilitate transcriptional regulation in this manner, BRCA1-associated protein 1 (BAP1), a member of the UCH family of DUBs, can also deubiquitinate H2Aub and regulate transcription, implying a degree of redundancy [66,67]."

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"These results suggest that HG inhibits SLC7A11 function by promoting BAP1 expression, thereby inducing ferroptosis in podocytes.3.5 HG promotes ferroptosis by promoting BAP1 deubiquitination of H2Aub on SLC7A11."

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"BAP1 has been shown to induce deubiquitination of H2Aub on the SLC7A11 promoter."

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"According to a report by Zhang et al., BAP1 causes the deubiquitination of H2Aub, which prevents SLC7A11 from being expressed [14]."
BAP1 inhibits H2Aub.
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BAP1 inhibits H2Aub. 10 / 10
| 1 9

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"This scenario is possible because H2Aub levels at a DSB are inversely correlated with BAP1 recruitment , and BAP1 depletion increases H2Aub accumulation at a DSB (Fig. 1)."

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"Genomic analyses together with robust statistics have revealed that the restoration of BAP1 facilitates the formation of the polycomb repressive deubiquitinase (PR-DUB) complex and inhibits ubiquitinated histone 2A (H2Aub) occupancy on the SLC7A11 promoter."

reach
"BRCA1-associated protein 1 (BAP1) negatively regulated H2Aub to decrease SLC7A11 expression, leading to lipid peroxidation accumulation."

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"Analysis of our H2Aub ChIP-seq data revealed that restoration of BAP1 WT, but not BAP1 C91A, markedly decreased H2Aub occupancy at both the promoter and gene body of SLC7A11 (XREF_FIG), which was further confirmed by H2Aub ChIP assay on the SLC7A11 promoter and representative exons (XREF_FIG and XREF_SUPPLEMENTARY)."

eidos
"Our own findings suggest that BAP1 inhibition leads to increased level of H2Aub , which interferes with chromatin and histone modifications at double-strand breaks ."

reach
"Several studies have revealed that BAP1 can inhibit ubiquitinated histone 2A (H2Aub) occupancy on the SLC7A11 promoter (Zhang et al., 2018)."

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"Monoubiquitinated histone H2A, H2Aub (K118 in Drosophila or K119 in vertebrates) is catalyzed by the Polycomb group (PcG) repressive complex 1 (PRC1) and reversed by the PcG-repressive deubiquitinase (PR-DUB) and BAP1 complex."

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"For example, BAP1 has been found to reduce H2Aub occupancy of the SLC7A11 promoter, leading to the suppression of SLC7A11 expression and contributing to elevated ferroptosis and lipid peroxidation [60]."

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"The combination of H2Aub and SLC7A11 promoter was detected by ChIP with H2Aub antibody or lgG, and qPCR and agarose gel electrophoresis showed that H2Aub could directly bind to SLC7A11 promoter, overexpressing BAP1 could reduce H2Aub enrichment on SLC7A11 promoter (Fig. 1E–F)."

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"As a tumor suppressor, BAP1 can target SLC7A11, reduce the H2Aub occupancy on the SLC7A11 promoter, inhibit the function of the amino acid transport complex system Xc-, and induce cancer cell death by promoting lipid peroxidation in cancer cells [16]."
BAP1 decreases the amount of H2Aub.
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BAP1 decreases the amount of H2Aub. 7 / 7
| 7

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"Our analysis showed that BAP1 restoration in control UMRC6 cells decreased H2Aub and SLC7A11 levels to similar degrees to that in NRF2 or ATF4 KO UMRC6 cells, suggesting that BAP1 regulates SLC7A11 expression independent of NRF2 or ATF4.PRC1 regulates SLC7A11 expression and h2aub occupancy on the SLC7A11 promoterSince our study suggested that BAP1 represses SLC7A11 expression through regulating H2Aub CELL CYCLE777Figure 1."

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"As the expression of BAP1, but not its catalytic dead form, is known to reduce the global levels of H2Aub, this strategy has been used to capture genomic locations that exhibit higher levels of H2Aub using chromatin immunoprecipitation in combination with next-generation genome sequencing (ChIP-Seq) ."

reach
"SLC7A11 appears to be a direct target of BAP1, which reduces the levels of H2Aub on SLC7A11 promoter, but yet represses its transcriptional activity."

reach
"To date, all DUBs regulating ferroptosis through the inhibition of system Xc‐/GPX, such as OTU DUB, ubiquitin aldehyde binding 1 (OTUB1), could directly interact with and stabilize SLC7A11 to inhibit ferroptosis, and BRCA1 associated protein 1 can decrease the amount of H2Aub occupying the SLC7A11 promoter to inhibit SLC7A11 expression, thereby inhibiting cystine uptake, resulting in increased levels of lipid peroxidation."

reach
"Our own findings suggest that BAP1 inhibition leads to increased level of H2Aub, which interferes with chromatin and histone modifications at double-strand breaks."

reach
"Furthermore, BAP1 was found to inhibit the expression of SLC7A11 by reducing the H2Aub level on the promoter of SLC7A11."

reach
"To test this hypothesis, we conducted gene set enrichment analysis (GSEA) on the genes with different fold changes in BAP1 dependent H2Aub reduction (5837 genes whose H2Aub levels were reduced by BAP1 by> 1.6-fold, 1494 genes by> 2-fold, and 101 genes by> 2.5-fold)."
BAP1 increases the amount of H2Aub.
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BAP1 increases the amount of H2Aub. 4 / 4
| 4

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"Our results revealed that transfection with H2Aub K119R mutant plasmid reduced the transcription and protein expression of SLC7A11 and GPX4, and overexpression of BAP1 can promote neuronal ferroptosis by regulating H2Aub levels."

reach
"In addition, BAP1 KO in UMSCC47 cells led to increased levels of H2Aub as compared with control cells (XREF_FIG)."

reach
"Therefore, BAP1 may contribute to DNA repair by regulating H2Aub levels at damage sites."

reach
"Few thousands genomic locations have been inferred as potentially regulated by BAP1 owing to changes of H2Aub  levels."
BAP1 ubiquitinates H2Aub.
| 3
BAP1 ubiquitinates H2Aub. 3 / 3
| 3

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"Specifically, our study suggests a model in which the tumor suppressor BAP1 normally functions to repress the expression of SLC7A11 at least partly by deubiquitinating H2Aub on SLC7A11, thereby inhibiting cystine uptake into cells and rendering them more sensitive to ferroptosis."

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"Previous studies also investigated that BAP1 links ferroptosis to tumor suppression by repressing SLC7A11 expression through BAP1-mediated H2Aub deubiquitination on SLC7A11 in PDAC cells12."

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"A recent study reported that BAP1 inhibits cystine uptake and makes cells more susceptible to ferroptosis by deubiquitinating H2Aub on SLC7A11."
BAP1 affects EZH2
| 3 28 4
BAP1 inhibits EZH2.
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BAP1 inhibits EZH2. 10 / 13
| 3 10

eidos
"Indeed , we observed that loss of BAP1 results in elevated EZH2 expression and increased H3K27me3 levels , and knock-down of Ezh2 leads to decreased H3K27me3 levels in BNC cells ( Fig. 5 , B and C ; and Fig. S2 J ) ."

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"LaFave et al. [XREF_BIBR] reported that loss of BAP1 promotes MPM cell proliferation by upregulating EZH2."

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"BAP1 loss leads to reduced L3MBTL2 stability and increased EZH2 transcriptional output (XREF_FIG)."

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"Concomitant Bap1 and Ezh2 loss reduced myeloid progenitor expansion, reduced the proportion of Mac1 + Gr1 + myeloid cells, restored erythroid differentiation (CD71 + Ter119 +) and decreased proliferation of Bap1 and Ezh2 deficient progenitors (XREF_FIG and XREF_SUPPLEMENTARY)."

eidos
"BAP1 loss leads to increased expression of EZH2 ( 15 ) ."

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"BAP1 deletion promotes cell proliferation by upregulating enhancer of zeste 2 polycomb repressive complex 2 subunit (EZH2)."

eidos
"BAP1 loss also up-regulates the expression of EZH2 , a Polycomb Repressive Complex-2 ( PRC2 ) component involved in epigenetic silencing ( 59 ) and oncogenic pathways ( 60 ) , suggesting sensitivity of BAP1-deficient MPM tumors to EZH2 inhibition ."

reach
"As for the reported tumor suppressive mechanism, BAP1 can antagonize EZH2 and PRC2 or RING1B (RNF2)/PRC1 in a tissue specific manner."

reach
"Additionally, reconstitution of BAP1 into the BAP1-KO cells downregulated H3K27me3 and EZH2 (Fig. 2c), suggesting that BAP1 loss might upregulate H3K27me3 and EZH2 and activate their downstream signaling.A recent study indicated that BAP1 is localized in the endoplasmic reticulum (ER) and bound to the type 3 inositol 1, 4, and 5-triphosphate receptor (IP3R3) [22]."

reach
"To that end, a systematic literature review was performed of articles dealing with a loss of BRCA1-associated protein 1 (BAP1), methylthioadenosine (MTAP), 5-hydroxymethylcitosine (5-hmC), glucose transporter 1 (GLUT1), insulin like-growth factor II messenger RNA-binding protein 3 (IMP3), enhanced zeste homologue 2 (EZH2) staining, cyclin-dependent kinase inhibitor 2A (CDKN2A) homozygous deletion (HD) testing, soluble mesothelin, and microRNA quantification in cytological samples for the diagnosis of MPM versus reactive atypical mesothelial cells."
BAP1 decreases the amount of EZH2.
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BAP1 decreases the amount of EZH2. 7 / 7
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"BAP1 loss led to reduced L3MBTL2 stability and increased EZH2 transcription."

reach
"In contrast to the reduction of H3K27me3 seen with Asxl1 loss, Bap1 deletion leads to increased Ezh2 expression, higher levels of H3K27me3, and enhanced repression of PRC2 target genes in hematopoietic cells."

reach
"In both cell lines, replenishment of BAP1 level induced decreased EZH2 level and vice versa (Fig 3C)."

reach
"In mesothelioma, BAP1 loss promotes EZH2 expression [34], but the opposite association was seen in our data (i.e., BAP1 loss was significantly associated with <80% EZH2 expression)."

reach
"BAP1 loss leads to upregulated EZH2 expression, contributing to aberrant H3K27me3 levels and disrupted differentiation pathways, as observed in hematopoietic stem cells (HSC) and mesothelioma ."

reach
"BAP1 loss leads to increased expression of EZH2 (15)."

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"In mouse models, the loss of BAP1 leads to elevated enhancer of zeste 2 polycomb repressive complex 2 subunit (EZH2) expression and enhanced repression of polycomb repressive complex 2 (PRC2) targets [48]."
Modified BAP1 decreases the amount of EZH2. 3 / 3
| 3

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"This could represent a possible connection between BAP1 and expression of HLA in uveal melanoma, as study in mice showed that loss of BAP1 leads to increased levels of EZH2 (and PRC2) with repressed expression of its targets [XREF_BIBR], including thus CIITA, leading ultimately to less HLA class II expression."

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"Recurrent BAP1 mutations have also been observed in intrahepatic cholangiocarcinoma 25, and a recent study showed that loss of Bap1 in mice increases expression of H3K27me3 and Ezh2 while suppressing expression of PRC2 targets 26."

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"Re-expression of BAP1 in Bap1 KO bone marrow cells reduced Ezh2 mRNA expression to normal levels (XREF_SUPPLEMENTARY)."
BAP1 binds EZH2.
| 2 3
| 2 3

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"BAP-1 interactions with histone deacetylase (HDAC) and enhancer of zeste homolog 2 (EZH2) have been exploited to suppress MPM progression."

sparser
"The interaction of BAP1 and EZH2 and the efficacy of tazemetostat show great potential for the future of cancer treatment for patients harboring variants in those genes due to the more aggressive nature of those tumors and the lack of targeted treatments."

sparser
"Survival analysis revealed that BAP1 loss, but not high EZH2 expression, was associated with a better prognosis."

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"We did not identify an interaction between BAP1 and EZH2 by co-immunoprecipitation (XREF_FIG)."

sparser
"In these lung-iPSCs, the expression of EZH2 (the main component of PRC2 complex) is also very high, and the physical interactions of AXSL proteins with EZH2 and BAP1 is also reported [ xref , xref , xref ]."
BAP1 activates EZH2.
| 3 1
BAP1 activates EZH2. 4 / 4
| 3 1

sparser
"Tazemetostat, a BAP1 inhibitor, increases the level of the PCR2 complex protein EZH2, activated by BAP1, and might be a potential drug for rhabdoid meningioma caused by BAP1 loss (NCT02860286) ( xref )."

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"In line with the observations in myeloid cells, Bap1 deleted mouse MM cell proliferation is impaired upon shRNA mediated down-regulation of Ezh2."

reach
"Frequent loss-of-function mutations in epigenetic factors, such as ARID1A, SMARCA4, SMARCB1, BAP1, and KDM6A, are likely to elicit the EZH2 and PRC2 addicted situation."

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"99 In addition, RCC derived cell lines deficient in BAP1 overexpress EZH2, and are sensitive to EZH2 inhibitors in vitro."
BAP1 increases the amount of EZH2.
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BAP1 increases the amount of EZH2. 3 / 3
| 3

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"In mouse models, the loss of BAP1 leads to elevated enhancer of zeste 2 polycomb repressive complex 2 subunit (EZH2) expression and enhanced repression of polycomb repressive complex 2 (PRC2) targets [48]."

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"2 BAP1 inactivation increases expression of enhancer of zeste homolog 2 (EZH2) or histone-lysine N-methyltransferase EZH2, 3 a component of the histone methyltransferase Polycomb repressive complex 2 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Tazemetostat, a BAP1 inhibitor, increases the level of the PCR2 complex protein EZH2, activated by BAP1, and might be a potential drug for rhabdoid meningioma caused by BAP1 loss (NCT02860286) (50)."
BAP1 affects Melanoma
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BAP1 inhibits Melanoma.
| 20
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"Harbour et al. do not demonstrate that BAP1 re-expression is sufficient to suppress the metastatic potential of uveal melanomas; however, if it does as one would predict from this study, substrates critical in migration, anoikis, and colonization should be considered."

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"In melanoma, BAP1 deficiency significantly elevated anaerobic glycolysis levels, leading to more lactate production in tumor cells [11]."

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"We present a unique case of BAP1-inactivated melanoma with anaplastic lymphoma kinase (ALK) fusion arising in a pre-existing BAP1-inactivated nevus in a 47-year-old female patient who presented with a dome-shaped red papule on the superior crus of the right antihelix."

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"BRCA1-Associated Protein-1 Inactivated Melanoma Arising in a Pre-existing Nevus With ALK Fusion and Low Tumor Mutational Burden."

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"Individuals with pathogenic BAP1 variants face an elevated risk of diverse tumors, notably BAP1-inactivated melanocytic nevi/tumors, uveal melanoma, cutaneous melanoma, mesothelioma, renal cell carcinoma, and basal cell carcinoma."

reach
"The BAP1 gene is located on chromosome 3, and loss of BAP1 has been suggested to drive T cell infiltration and help develop an inflammatory microenvironment in uveal melanoma [ 15 ]."

reach
"Here we report the clinical features, histopathologic findings, and chromosomal copy number data of 19 BAP1-inactivated melanocytomas (BIMs) and compare their features to those of five BAP1-inactivated melanomas."

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"Although not fully diagnostic, a mitotic count of ≥3/mm 2 and ≥6 CNAs in the appropriate context is supportive of a diagnosis of BAP1-inactivated melanoma."

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"BAP1-Inactivated Melanoma Arising From BAP1-Inactivated Melanocytic Tumor in a Patient With BAP1 Germline Mutation: A Case Report and Review of the Literature."

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"The addition of BAP1 loss increased tumor proliferation and melanoma size."
BAP1 activates Melanoma.
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BAP1 activates Melanoma. 8 / 8
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"In addition, a mutation in the BRCA1-associated protein-1 (BAP1) gene is implicated in UM metastasis, and is also known to promote the development of cutaneous melanoma, basal cell carcinoma, malignan[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"On the other hand, the occasionally described heterozygous deletions of these genes cannot be completely excluded in our series, because NGS procedure and mode of data analysis were not sensitive to moderate CNV (copy number variation), the detection of which was not the aim of this panel.Germline mutations in BAP1 are known to underlie sometimes melanomas, renal cell carcinomas, malignant mesotheliomas, and some other cancers [31]."

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"On the transcriptomic level, uveal melanoma progression and aggressiveness is driven by inactivation of both BAP1, a subunit of the polycomb-repressive deubiquitylase complex (8) and polycomb repressive complex 1 (PRC1) activity, leading to derepression of PRC1 target genes (12)."

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"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (often in the abdomen), and uveal melanoma [97]."

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"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (frequently in the abdomen) and uveal melanoma [151,152,153]."

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"BAP1 germline mutations underlie a rare, autosomal dominant tumour predisposition syndrome that is most commonly associated with (in order of decreasing frequency) atypical Spitz tumours, uveal melano[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"This pathway is exploited by immune checkpoint inhibitors in the treatment of melanoma.12 Coordination of the immune response with therapies targeting oncogenes is a possible treatment strategy.12 89CDKN2A, CDK4, MCR1, BAP1, and TERT promoter germline mutations increase melanoma risk within families.15 96 97 No specific germline mutation is associated with acral melanoma yet; in fact, MCR1 variants were less common in acral lentiginous melanoma patients in a Swedish study."

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"Melanoma predisposition syndromes caused by pathogenic variants in POT1 and BAP1 are more recently described, and both are associated with Spitzoid tumors."
Mutated BAP1 activates Melanoma. 5 / 5
| 5

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"In melanoma, BAP1 mutations promote tumor cell glycolysis, leading to increased lactate production."

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"Whereas BAP1 mutations drive BAP1 tumour predisposition syndrome, mesotheliomas, melanomas and other neoplasms 8–11 ; ASXL1 mutations are particularly prevalent in myeloid cancers, and are associated [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Germline BAP1 mutations cause mesothelioma and other malignancies, namely, uveal cancer, meningioma, and melanoma, overall defined as “BAP1- related cancer syndrome”."

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"BAP1 mutations cause prevalently mesothelioma and melanomas."

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"Moreover, in uveal melanoma, somatic BAP1 mutations cause metastasis with a probability of 74%, while the same has been observed in only 36% of the patients carrying a germline mutation [104]."
KLF5 affects BAP1
7 | 14 11
7 | 14 11

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"The deubiquitinase (DUB) BAP1 had been reported to regulate KLF5 stability in breast cancer cells, so we hypothesize that LINC00152 mediates KLF5 depending on the binding of BAP1 and KLF5."

reach
"As shown in XREF_SUPPLEMENTARY, LINC00152 knockdown inhibits BAP1 binding with KLF5."

sparser
"Another KLF5 DUB that stood out in siRNA screening was BAP1 (BRCA1-associated protein 1), which also directly binds KLF5 and stabilizes KLF5 through deubiquitination [ xref ]."

sparser
"The deubiquitinase (DUB) BAP1 had been reported to regulate KLF5 stability in breast cancer cells ( xref , xref ), so we hypothesize that LINC00152 mediates KLF5 depending on the binding of BAP1 and KLF5."

sparser
"As shown in xref , LINC00152 knockdown inhibits BAP1 binding with KLF5."

reach
"Another KLF5 DUB that stood out in siRNA screening was BAP1 (BRCA1 associated protein 1), which also directly binds KLF5 and stabilizes KLF5 through deubiquitination [XREF_BIBR]."

sparser
"LncRNA PVT1 enhanced KLF5 protein stability by increasing the interaction between KLF5 and the deubiquitinase BAP1 [ 26 ]."

reach
"BAP1 interacts directly with KLF5 and stabilizes KLF5 via deubiquitination."

reach
"BAP1 interacts directly with KLF5."

sparser
"Recently, Jia et al. ( xref ) showed that the WWP1- BAP1-KLF5 axis is dysregulated in melanoma cancer; WWP1 was lowly expressed in melanoma cells and tissues whereas KLF5 and BAP1 were highly expressed and closely related to tumor metastasis."
| 32
| 17

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"In functional studies, depletion of BAP1 protein in the 92.1 UM cell line using siRNAs led to a change in cell morphology from spindle to a more epithelioid phenotype, loss of melanocytic differentiation and transition from a ' class 1 ' to a ' class 2 ' gene expression profile."

reach
"BAP1 loss also leads to the failure of the H3K27ac histone mark to accumulate at the promoter sites of key lineage commitment genes, highlighting its role in the broader regulation of transcription and cell differentiation (19)."

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"The recent finding that BAP1 loss leads to defective differentiation and an arrested primitive phenotype in vertebrate development and uveal melanoma suggests that BAP1 loss can promote tumor progression by inducing cell dedifferentiation and stem like behavior."

reach
"Studies suggest that down regulated expression of BAP1 led to impaired differentiation in UM cells."

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"However, as previously reported, depletion of BAP1 in UM cells induced loss of melanocytic differentiation together with acquisition of a gene expression profile seen in advanced metastasizing tumors."

reach
"Consistent with those results, treatment of BAP1 deficient uveal melanoma cells with an HDAC inhibitor restored the expression of the melanocyte differentiation markers, which were down-regulated by BAP1 depletion, in a dose dependent manner."

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"Depletion of BAP1 in cultured class 1 UM cells induced a loss of melanocytic differentiation and acquisition of a class 2 gene expression profile, suggesting that the loss of BAP1 may be mechanistically linked to metastasis."

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"Instead, prenatal Bap1 loss profoundly impairs postnatal enteric neuron differentiation and induces postnatal enteric neuron loss."

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"ASXL1-MT and BAP1 impairs multilineage differentiation of HSPCs."

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"ASXL1-MT alone modestly, and coexpression of ASXL1-MT and BAP1 dramatically blocked G-CSF-induced granulocytic differentiation of 32Dcl3."
BAP1 bound to ASXL1-MT inhibits cell differentiation. 2 / 2
| 2

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"The ASXL1-MT and BAP1 complex is not a simple growth-accelerator, but rather inhibits terminal differentiation of HSPCs toward granulocytes, macrophages, mast cells, and erythrocytes."

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"Together, these results indicate that the ASXL1-MT and BAP1 complex impairs terminal differentiation of HSPCs, except for differentiation toward monocytes."
| 15

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"Interestingly, ASXL1-MT and BAP1 promotes monocyte differentiation, while it inhibits terminal differentiation to macrophage, which may account for the frequent (40-50%) detection of ASXL1 mutations in CMML patients."

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"6 In line with these findings, several reports have indicated that truncated ASXL1 enhances BAP1 complex activity, thereby promoting depletion of the H2AK119Ub mark and aberrant myeloid differentiation."

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"BAP1 reduction significantly decreased the colony-forming capacity of ASXL1 mutant primary AML cells, and partially rescued the differentiation block in myeloid cells."

reach
"Paradoxically, recent findings have confirmed that BAP1 also promotes growth and differentiation even in uveal melanomas where loss of BAP1 expression is a common event."

reach
"As a deubiquitinating enzyme, BAP1 contributes to gene transcription, cell differentiation, DNA damage repair, apoptosis, and cell metabolism in tumor inhibition."

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"Moreover, BAP1’s modulation of cell differentiation through HDACs and EZH2 expression levels holds promise for understanding and potentially addressing pregnancy complications characterized by aberrant epigenetic modifications."

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"BAP1 depletion caused a reduction in mRNA levels of neural crest migration genes (ROBO1), melanocyte differentiation genes (CTNNB1, EDNRB and SOX10) and other genes that are down-regulated in class 2 tumors (LMCD1 and LTA4H)."

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"Both ASXL1 and BAP1 were downregulated during RA induced P19 cell differentiation with concomitant increase of ubiquitinated H2B, leading to activation of Hox genes."

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"In the monocyte and macrophage differentiation assay, coexpression of ASXL1-MT and BAP1 substantially promoted differentiation toward CD115 + monocytes while inhibiting their terminal differentiation to CD115 + F4/80 + macrophages."

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"Downregulation of ASXL1 and BAP1 Promotes Hox Gene Exp (A) Downregulation of ASXL1 and BAP1 during RA induced differentiation."
BAP1 affects BAP1
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BAP1 inhibits BAP1.
| 11
BAP1 inhibits BAP1. 10 / 13
| 11

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"BAP1-inactivated melanocytoma is associated with BRAF or NRAS and BRCA1-associated protein 1 (BAP1) mutations."

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"Tumors with both miR-31 and BAP1 overexpression exhibited significantly higher levels of BAP1 compared to tumors with miR-31 overexpression, suggesting that BAP1 overexpression could rescue the BAP1 suppression caused by miR-31."

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"There were no chromosomal losses in either the Bap1 or Nf2 regions."

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"Overexpression of wild-type BAP1 reversed the sensitivity of BAP1 knockout cells to RITA, while overexpression of the mutant BAP1 (C91S) with inactivated ubiquitinase did not produce any rescue effect."

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"Ubiquitylation of BAP1 near its nuclear localization sequence negatively regulates its activity by excluding BAP1 from the nucleus, where most of its targets reside [84] ."

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"BAP1-inactivated melanocytic tumours (BIMTs) occur sporadically and in association with a familial tumour predisposition syndrome involving germline mutations in the BRCA1-associated protein-1 (BAP1) gene."

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"Furthermore, although CDKN2A/p16 homozygous deletion was not confirmed by FISH, there was loss of BAP-1 based on IHC (Fig. 2G)."

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"Although there was no loss of MTAP based on IHC and homozygous deletion of CDKN2A/p16 based on FISH (Fig. 5E and F), there was loss of BAP-1 based on IHC (Fig. 5D)."

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"To further investigate whether BAP1 overexpression induced the opposite effect of BAP1 depletion on MSC migration, we performed wound healing, transwell migration, and invasion assays under BAP1 overexpression conditions."

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"A loss of BAP1 (BRCA1-associated protein-1) expression, measured by immunohistochemistry, was also associated with this phenotype (r = −0.38 and q value = 4.61 × 10 ; Supplementary Fig. 19)."
BAP1 increases the amount of BAP1.
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BAP1 increases the amount of BAP1. 7 / 7
| 7

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"Mechanically, we show that BAP1 represses metabolic stress induced UPR and cell death through activating transcription factor 3 (ATF3) and C/EBP homologous protein (CHOP), and reveal that BAP1 binds to ATF3 and CHOP promoters and inhibits their transcription."

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"Deletion of chromosome 3 and BAP1 inactivating mutations cause loss of BAP1 expression and function in cancer."

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"DIM promotes proliferation and osteogenic differentiation of MC3T3-E1 cells in vitro, and also plays a bone promoting role by increasing the interaction between BRCA1-Associated Protein 1(BAP1) and Inositol 1,4,5-Trisphosphate Receptor(IP3R), up-regulating the expression of BAP1 and IP3R and downstream storage operation calcium entry (SOCE) related protein Recombinant Stromal Interaction Molecule 1(STIM1)."

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"We also detect the mRNA and protein expression level of BAP1, tumors with BAP1 plasmid transfected group exhibited higher BAP1 mRNA and protein expression compared with the control group, suggesting that BAP1 plasmid overexpression could rescue the expression of BAP1 suppressed by miR-31."

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"Genetic mutations in BAP1 could modulate BAP1 protein expression or could alter BAP1 function in the absence of alterations in protein expression."

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"BAP1-IP3R3 CoIP was barely detectable in BAP1 +/- fibroblasts; BAP1 rescue significantly increased the amount of BAP1 that CoIP with IP3R3 (XREF_FIG)."

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"Furthermore, qRT-PCR and western blotting revealed that BAP1 mRNA and protein levels were notably enhanced in HCT116 and RKO cells by miR-593-3p inhibitor transfection, whereas miR-593-3p mimic apparently restrained BAP1 expression (Fig. 4E and F)."
BAP1 activates BAP1.
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BAP1 activates BAP1. 7 / 7
| 7

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"While ubiquitination enzyme UBE2O monoubiquitinates the NLS of BAP1 and induces translocation (inactivation) of BAP1 from the nucleus to the cytoplasm, DUB activity of the UCH-domain of BAP1 counteracts the UBE2O activity mediated through the intramolecular interaction between the UCH-domain and COOH-terminal domain of BAP1."

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"Identified in 2011, the inherited cancer predisposition syndrome caused by germline mutations in the tumor suppressor gene BAP1 (BRCA1 Associated Protein 1) has shed light for the first time on monoge[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The BAP1 cancer syndrome (Mendelian Inheritance in Man Tumor Predisposition Syndrome # 614327) is caused by heterozygous germline mutation in the BRCA1 associated protein-1 gene (BAP1) located at chromosome 3p21.1."

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"Heterozygous loss of BAP1 would reduce overall levels of BAP1 protein, disfavouring bidentate complex assembly and decreasing specific activity."

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"Reduced levels of BAP1 in the genetically predisposed carriers of the mutated BAP1 forms are responsible for the reduction both of IP3R3 levels and Ca flux, preventing BAP1 cells that accumulate DNA damage from executing apoptosis."

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"Germline pathogenic variants in the BRCA1 associated protein-1 (BAP1) gene cause the BAP1 tumor predisposition syndrome (TPDS)."

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"Pathogenic germline variants in the BRCA1 associated protein 1 (BAP1) gene cause the BAP1 tumor predisposition syndrome (BAP1-TPDS) with increased risk of several cancers, the most frequent of which is uveal melanoma (UM)."
BAP1 decreases the amount of BAP1.
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BAP1 decreases the amount of BAP1. 4 / 5
| 4

reach
"Using patient derived lymphoblastoid cell lines, we found that carriers of pathogenic BAP1 germline variants (c. 852_del and c. 1358_1359del) had reduced expression of full-length BAP1, Vimentin and Snail, as compared to controls with wild-type BAP1 (BAP1 WT), whereas BAP1 germline VUS (c. 299T> C and c. 551A> G) or likely benign carriers were not significantly different from wild-type BAP1 WT."

reach
"Furthermore, the BAP1 expression pattern observed in the Human Protein Atlas supports an inverse correlation between BAP1 expression and cell motility, with blood cells, known for their high motility, showing notably lower levels of BAP1 expression (Supplementary Fig. 2 of the SI)."

reach
"These results are consistent with no observed change in cell cycle profile in skin fibroblasts taken from BAP1 heterozygous individuals that had reduced BAP1 protein levels [18], although different responses in other cell types cannot be excluded."

reach
"This result is perhaps surprising given that BAP1 inactivation or loss of BAP1 expression has been detected using IHC and whole-exome sequencing in approximately 25% of the intrahepatic CCAs [XREF_BIBR, XREF_BIBR, XREF_BIBR]."
BAP1 affects ITPR3
4 | 2 18 5
BAP1 binds ITPR3.
4 | 4 5
4 | 4 5

sparser
"Cytoplasmic BAP1 binds to IP3R3 and promotes calcium ion release from the endoplasmic reticulum into the cytosol via IP3R3 deubiquitination, thereby inducing apoptosis."

reach
"Similarly, IP3R3 is bound and deubiquitinated by BRCA1-associated protein 1 (BAP1) at the ER [33] ."

reach
"For example, BAP1 binds to IP3R3 to deubiquitinate and promote apoptosis."

No evidence text available

reach
"Overexpression of the N-terminus of the IP3R3 binds and sequesters BAP1 leading to decreased IP3R3 levels, an effect counter balanced by BAP1 co-overexpression (XREF_FIG)."

No evidence text available

sparser
"Recently, a novel BAP1 cytoplasmic function was introduced, showing that ER-localized BAP1 can bind and stabilize IP3R3 to modulate Ca2+ release resulting in apoptosis [ xref ]."

sparser
"We confirmed the specificity of the BAP1 interaction with IP3R3 in HEK-293 cells expressing FLAG-BAP1 ( xref )."

No evidence text available

reach
"The data clearly indicated that BAP1 physically interacts with the type-3 inositol-1,4,5-trisphosphate-receptor (IP3R3), and BAP1 expression level is directly correlated with IP3R3 in wild-type and mutant fibroblasts."
BAP1 deubiquitinates ITPR3.
| 7
BAP1 deubiquitinates ITPR3. 7 / 7
| 7

reach
"At the ER, BAP1-dependent deubiquitination of the type 3 inositol-1,4,5-trisphosphate receptor (IP3R3) is necessary to sustain the IP3R3 ER channel integrity (11,55)."

reach
"We hypothesized that BAP1 might deubiquitylate and stabilize IP3R3."

reach
"For example, BAP1 promotes apoptosis by deubiquitinating and stabilizing IP3R3 at the endoplasmic reticulum (ER), which stimulates Ca release from the ER into the cytosol and thereby increases the mitochondrial Ca concentration and cytochrome C release ."

reach
"BAP1 can regulate Ca release from the ER to the cytoplasm and mitochondria to promote cell apoptosis by binding, deubiquitylating and stabilizing IP3R3 [45]."

reach
"Along with maintenance of IP3R3 ER channel integrity, BAP1-dependent deubiquitination of IP3R3 may change the threshold for IP3R3 activation by Ca itself or IP3."

reach
"BAP1 regulates Ca release from endoplasmic reticulum and mitochondria by deubiquitinating and stabilizing inositol 1,4,5-trisphosphate receptor type 3 (IP3R3), promoting apoptosis after genotoxic stress (14)."

reach
"The endoplasmic reticulum (ER) fraction is the primary location of cytoplasmic BAP1, where it deubiquitylates and stabilizes the type 3 inositol-1,4,5-trisphosphate receptor (IP3R3)."
BAP1 increases the amount of ITPR3.
| 6
Modified BAP1 increases the amount of ITPR3. 3 / 3
| 3

reach
"Loss of BAP1 therefore leads to loss of ITPR3, which in turn reduces apoptotic mitochondrial calcium signals [XREF_BIBR]."

reach
"Reduced levels of BAP1 in BAP1 +/- carriers caused reduction of both IP3R3 levels and Ca 2+ flux, preventing BAP1 +/- cells that had accumulated DNA damage from executing apoptosis."

reach
"Consequently, heterozygous loss of BAP1 leads to decreased IP3R3 levels and resistance to apoptotic stimuli, as demonstrated in human fibroblasts or mesothelial cells derived from mutant BAP1 +/- carriers and in mesothelioma cell lines 174."
BAP1 increases the amount of ITPR3. 3 / 3
| 3

reach
"Remarkably, depletion of BAP1 or IP3R3 promoted crocidolite asbestos and TNF-alpha-induced transformation of primary mesothelial cells, and re-expression of IP3R3 in BAP1 depleted cells partially reverted the phenotype."

reach
"Silencing of BAP1 in BAP1 WT fibroblasts consistently caused a reduction of IP3R3 protein levels (XREF_FIG), with concomitant reduction of mitochondrial Ca 2+ responses (XREF_FIG, and XREF_SUPPLEMENTARY), and protection from apoptosis (XREF_FIG)."

reach
"Transduction of BAP1 +/- fibroblasts with adenoviruses encoding BAP1, but not the catalytic inactive mutant BAP1 (C91S) XREF_BIBR, XREF_BIBR, rescued IP3R3 protein levels (XREF_FIG), mitochondrial Ca 2+ uptake (XREF_FIG, XREF_FIG, XREF_SUPPLEMENTARY) and resulted in enhanced apoptosis (XREF_FIG)."
BAP1 activates ITPR3.
| 2 1
BAP1 activates ITPR3. 3 / 3
| 2 1

reach
"In the cytoplasm, BAP1 modulates the stability of the IP3R3 channel, which allows the flux of Ca + from the endoplasmic reticulum into the mitochondria, where Ca + is required for the Krebs cycle and, at higher doses, to execute apoptosis."

eidos
"Thus , loss of BAP1 leads to decreased ITPR3 expression , decreased mitochondrial Ca2 + and reduced sensitivity of cancer cells to apoptosis ."

eidos
"Loss of BAP1 therefore leads to loss of ITPR3 , which in turn reduces apoptotic mitochondrial calcium signals [ 6 ] ."
IRAK1 affects BAP1
| 10 17
| 10 17

reach
"Mechanistically, BAP1 directly binds to IRAK1, competitively inhibits the interaction between IRAK1 and IRAK4, significantly prevents phosphorylation at Thr209 and Thr378 of IRAK1, as well as the autophosphorylation of IRAK1, impeding the dissociation of IRAK1 from the Myddosome complex and the sequential activation NF-κB signaling.Previous studies have demonstrated that IRAK1 undergoes phosphorylation and K63-linked ubiquitination during activation, both of which are essential for the initiation of downstream signaling pathways30, 41."

sparser
"Considering that BAP1 can inhibit the above process without requiring enzymatic activity, we hypothesize that BAP1 likely binds to IRAK1 in a competitive manner with IRAK4."

sparser
"Furthermore, ectopically overexpressed Flag-IRAK1 interacted solely with BAP1 and not with other members of the PR-DUB complex, ASXL1-3 (Fig. xref E-F), indicating a specific interaction between BAP1 and IRAK1."

sparser
"The BAP1-IRAK1 interaction primarily takes place in the cytoplasm (Fig. xref H), where BAP1 is enzymatically inactive in the absence of ASXLs."

sparser
"Together, we identified the protein-protein interaction between BAP1 and IRAK1 in cytoplasm and elucidated that BAP1 might repress NF-κB through IRAK1."

sparser
"To prove this, we silenced/overexpressed IRAK4 and detected whether the binding of BAP1 and IRAK1 was affected."

sparser
"We then tested if the phosphorylation status of IRAK1(T209) would affect BAP1-IRAK1 interaction."

sparser
"Lipopolysaccharide (LPS) treatment significantly increased the phosphorylation at IRAK1(T209) as expected, but also substantially inhibited the interaction between BAP1 and IRAK1 (Fig. xref D)."

sparser
"Consistently, λ-phosphatase treatment prominently enhanced BAP1-IRAK1 interaction (Fig. xref E)."

sparser
"The interaction between BAP1 and IRAK1 is closely related to the phosphorylation status of IRAK1."
YY1 affects BAP1
6 | 8 9
6 | 6 4

sparser
"Instead, association of Yin Yang 1 with BAP1 could be the remnant of an ancient association between Pleohomeotic and Calypso, which would have allowed for the recognition of Polycomb response elements by the Drosophila PR-DUB complex."

reach
"However, BAP1 can also form complexes with many other proteins, including the tumor suppressor BRCA1, host cell factor-1 (HCF-1), N-acetylglucosamine transferase, the forkhead box transcription factors FOXK1/2, MBD family proteins MBD5/6, transcription factor YY1 and ubiquitin conjugating enzyme UBE20 [XREF_BIBR, XREF_BIBR - XREF_BIBR]."

reach
"may be important for the growth effects of HCF-1, which are partly mediated by modulation, at E2F responsive promoters, of progression of cells through the G1/S-phase boundary and transcription of gen[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Given BAP1’s crucial role in early placentation xref , it is plausible that BAP1 may interact with YY1 to regulate trophoblast proliferation and invasion, offering a novel avenue for exploring the mechanisms underlying pregnancy disorders."

reach
"BAP1 interacts with BARD1 (182aeuro " 365), HCF-1 (365aeuro " 385), BRCA1 (596aeuro " 721) and YY1 (642- 686).10,88 Numbers refer to amino acid positions."

No evidence text available

No evidence text available

sparser
"Thus, targeting the BAP1YY1 axis may represent a novel therapeutic approach in PM."

reach
"– YY1 (Ying Yang 1) transcription factor that binds to a zinc finger domain BAP1 and HCF1."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
KMT2C affects BAP1
4 | 4 18
4 | 4 18

No evidence text available

sparser
"For instance, the BAP1 H2A deubiquitinase recruits H3K4 monomethylase MLL3 to monomethylate gene enhancers, while disruption of the interaction between BAP1 and MLL3 contributes to the pathogenesis of multiple cancers [ xref ]."

No evidence text available

No evidence text available

reach
"Recently identified missense mutations within the PHD fingers 1-3 have been shown to disrupt the interaction between KMT2C and BAP1 leading to reduced recruitment of KMT2C to gene enhancers 1."

No evidence text available

sparser
"In addition, a mutation hotspot within the PHD cluster of the KMT2C gene was described, the mutations of which disrupted the interaction of KMT2C with the BAP1 (BRCA1 associated protein-1) tumor suppressor, resulting in poor patient survival in many types of cancer (liver, lung, bladder, and breast)."

sparser
"This unique domain could mediate the association of BAP1 (a histone deubiquitinase) with MLL3, but not with, other SET1/MLL family of KMTs, including MLL4."

sparser
"Cancer-associated MLL3 PHD mutations disrupt the interaction between MLL3 and BAP1 and correlate with poor patient survival."

sparser
"MLL3 COMPASS uniquely associates with the BAP1 complex."
BAP1 affects YY1
6 | 8 9
6 | 6 4

sparser
"Instead, association of Yin Yang 1 with BAP1 could be the remnant of an ancient association between Pleohomeotic and Calypso, which would have allowed for the recognition of Polycomb response elements by the Drosophila PR-DUB complex."

reach
"However, BAP1 can also form complexes with many other proteins, including the tumor suppressor BRCA1, host cell factor-1 (HCF-1), N-acetylglucosamine transferase, the forkhead box transcription factors FOXK1/2, MBD family proteins MBD5/6, transcription factor YY1 and ubiquitin conjugating enzyme UBE20 [XREF_BIBR, XREF_BIBR - XREF_BIBR]."

reach
"may be important for the growth effects of HCF-1, which are partly mediated by modulation, at E2F responsive promoters, of progression of cells through the G1/S-phase boundary and transcription of gen[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Given BAP1’s crucial role in early placentation xref , it is plausible that BAP1 may interact with YY1 to regulate trophoblast proliferation and invasion, offering a novel avenue for exploring the mechanisms underlying pregnancy disorders."

reach
"BAP1 interacts with BARD1 (182aeuro " 365), HCF-1 (365aeuro " 385), BRCA1 (596aeuro " 721) and YY1 (642- 686).10,88 Numbers refer to amino acid positions."

No evidence text available

No evidence text available

sparser
"Thus, targeting the BAP1YY1 axis may represent a novel therapeutic approach in PM."

reach
"– YY1 (Ying Yang 1) transcription factor that binds to a zinc finger domain BAP1 and HCF1."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
BAP1 affects KMT2C
4 | 4 18
4 | 4 18

No evidence text available

sparser
"For instance, the BAP1 H2A deubiquitinase recruits H3K4 monomethylase MLL3 to monomethylate gene enhancers, while disruption of the interaction between BAP1 and MLL3 contributes to the pathogenesis of multiple cancers [ xref ]."

No evidence text available

No evidence text available

reach
"Recently identified missense mutations within the PHD fingers 1-3 have been shown to disrupt the interaction between KMT2C and BAP1 leading to reduced recruitment of KMT2C to gene enhancers 1."

No evidence text available

sparser
"In addition, a mutation hotspot within the PHD cluster of the KMT2C gene was described, the mutations of which disrupted the interaction of KMT2C with the BAP1 (BRCA1 associated protein-1) tumor suppressor, resulting in poor patient survival in many types of cancer (liver, lung, bladder, and breast)."

sparser
"This unique domain could mediate the association of BAP1 (a histone deubiquitinase) with MLL3, but not with, other SET1/MLL family of KMTs, including MLL4."

sparser
"Cancer-associated MLL3 PHD mutations disrupt the interaction between MLL3 and BAP1 and correlate with poor patient survival."

sparser
"MLL3 COMPASS uniquely associates with the BAP1 complex."
| 26
| 15

reach
"Of note, UM is driven by activating mutations in Gαq pathway, which are recurrently associated with a second mutation in BAP1 (BRCA1-associated protein 1), SF3B1 or EIF1AX (eukaryotic translation initiation factor 1A X-linked)."

reach
"BAP1 depletion appears to promote UM cell migration underneath the monolayer."

reach
"The results suggest that knowledge of mutations in BAP1 and EIF1AX can enhance prognostication of UM beyond that determined by chromosome 3 and tumor characteristics."

reach
"However, the molecular mechanisms through which BAP1 promotes UM metastasis is still unclear.An important open question is whether the sunlight exposure could cooperate with BAP1 inactivation in UM development and progression."

reach
"XREF_BIBR UM in BAP1 germ line mutants is usually diagnosed between the ages of 30 and 59 years, and is driven by inactivating mutations in the lone functional BAP1 gene, analogous to the frequent loss of chromosome 3 observed in high-risk sporadic disease."

reach
"In the bone, skin, and liver, UM metastasis was found to be caused by aberrant methylation of BAP1 and SF3B1."

reach
"BAP1 loss-of-function mutations promote the metastatic spread of UM cancers in patients XREF_BIBR, and TEM should be a rate limiting step of metastasis."

reach
"These studies demonstrate that BAP1 deficiency slows the proliferation of UM cells through regulation of S6 phosphorylation."

reach
"Nevertheless, the key node of BAP1 pathway in driving or mediating UM metastasis remains unexplored.LaFave LM et al. reported that knockout of BAP1 results in elevated transcription of EZH2 gene in acute myeloid leukemia mouse bone marrow cells and human mesothelioma cells [48]."

reach
"Using BAP1‐competent UM cell lines engineered to reduce BAP1 expression through doxycycline‐induced shRNA expression, we found that reduction of BAP1 correlated with increased histone H2A ubiquitination, suggesting that in UM, BAP1 contributes to epigenetic regulation (Figure 4A)."
Mutated BAP1 activates 2'-O-methyluridine. 4 / 4
| 4

reach
"Regarding tumor mutation, we discovered that BAP1 was more frequently mutated in the Cluster1 subtype, which was consistent with prior evidence suggesting BAP1 mutations enhance the possibility of UM patients developing metastases [32]."

reach
"BAP1 mutations enhanced the malignant phenotypes of UM cells."

reach
"Herein, we set out to investigate whether BAP1 mutations could regulate the development in UM, and our results demonstrated that BAP1 mutations could induce the tumor immune microenvironment in UM by inactivating the NF-κB signaling pathway."

reach
"BAP1 is a high-risk locus in UM and germline BAP1 mutations increase the risk of developing UM and other tumours such as melanocytic skin tumours, mesothelioma, and renal cell carcinoma (known as the BAP1-predisposition syndrome) [8,174]."
| 11

reach
"Early changes resulting in gain of chromosome 8q may activate macrophage infiltration, while sequential loss of BAP1 expression seems to drive T cell infiltration in UM."

reach
"Kaler et al. analyzed scRNA-seq data from 11 UM tumor samples and confirmed that BAP1 loss can lead to an increase in PROS1 expression in class 2 UM (characterized by poor prognosis and high metastatic risk) cells while increasing MERTK expression in CD163+ macrophages (25)."

reach
"It has been found that mutations in BAP1, SF3B1, and EIF1AX in UM with different prognoses exhibit different types of methylation cluster status, and hypermethylation of chromosome 3 in UM is also associated with downregulation of BAP1 gene expression, which was further confirmed in vitro experiments that knockdown of BAP1 gene or deletion of the protein induces effects on methylation status in UM cells, causing UM cells to exhibit a low metastatic risk phenotype [20–22]."

reach
"Interestingly, recent research has revealed that the gain of chromosomal 8q may worsen prognosis by activating macrophage infiltration, and the loss of BAP1 expression may drive T cell infiltration in UM (Gezgin et al., 2017), suggesting that immune infiltration plays a crucial role in the prognosis of UM.Emerging evidence shows that the immune microenvironment is crucial for cancer progression and response to therapeutics (Quail and Joyce, 2013)."

reach
"A reduction of BAP1 activity causes UM cells to acquire characteristics reminiscent of stem cells, leading to upregulation of genes with stem cell-like properties and developmental processes."

reach
"Uveal melanoma (UM) with BAP1 inactivating mutations has a high risk of metastasis, but the mechanism behind BAP1 deficiency driving UM metastasis is unknown."

reach
"Loss of BAP1 enhances UM transmigration."

reach
"Therefore, we hypothesized that decreasing BAP1 might increase the ability of UM cells to perform TEM in our model system."

reach
"BAP1 promotes progenitor cell differentiation into melanocytes by increasing MITF expression and BAP1 loss may promote metastasis in UM by remodeling the epigenome to resemble that of stem-like migratory neural crest cells."

reach
"Next, we used movie analysis of living cells to examine BAP1 depleted UM cells interacting with EC monolayers."
FOXO3 affects BAP1
| 11 14
FOXO3 binds BAP1.
| 2 14
| 2 14

sparser
"Both endogenous and exogenous CoIP demonstrated that BAP1 protein could directly bind to FOXO3a protein in cells."

sparser
"Moreover, the binding of the transcription factor FOXO3a with the BAP1 promoter can modulate neuronal lipid peroxidation and ferroptosis by enhancing BAP1 expression."

sparser
"Consequently, the FOXO3a-BAP1 axis might represent a promising therapeutic strategy for addressing neuronal ferroptosis in patients with SAH."

sparser
"Our findings elucidate the impact of FOXO3a and BAP1 on SLC7A11-related ferroptosis following SAH both in vivo and in vitro , and the inhibition of the FOXO3a-BAP1 axis can significantly attenuate neuronal damage and ferroptosis in EBI after SAH."

sparser
"Preliminary experiments have revealed that FOXO3a can bind to the BAP1 promoter and regulate the expression of BAP1."

sparser
"Consequently, this study aimed to investigate the potential mechanism by which FOXO3a-BAP1 influences neuronal ferroptosis in the pathogenesis of EBI after SAH."

sparser
"BAP1 directly interacted with FOXO3a."

sparser
"To explore whether DUB BAP1 directly regulates the above proteins through deubiquitination in neurons after SAH, the binding of BAP1 with FOXO3a proteins was detected by prediction of protein-protein interactions and CoIP assay."

sparser
"Endogenous CoIP also revealed that BAP1 protein could directly bind to FOXO3a protein in HT22 cells ( xref B)."

reach
"To explore whether DUB BAP1 directly regulates the above proteins through deubiquitination in neurons after SAH, the binding of BAP1 with FOXO3a proteins was detected by prediction of protein-protein interactions and CoIP assay."
FOXO3 increases the amount of BAP1.
| 9
FOXO3 increases the amount of BAP1. 9 / 9
| 9

reach
"To further clarify the regulatory effect of FOXO3a on BAP1, western blotting showed that silencing FOXO3a significantly inhibited the expression level of BAP1 protein in neurons after SAH, and increased the expression of H2Aub and SLC7A11 (Fig. 4J–L)."

reach
"qPCR also found that silencing FOXO3a significantly inhibited BAP1 mRNA levels and increased SLC7A11 mRNA levels after SAH (Fig. 4M–N)."

reach
"Immunofluorescence also appeared that silencing FOXO3a significantly inhibited the expression level of BAP1 protein in neurons after SAH (Fig. 4O)."

reach
"Immunofluorescence also revealed that FOXO3a overexpression further increased the protein expression level of BAP1 in HT22 cells (Fig. 5C)."

reach
"These results indicated that FOXO3a was activated in hemin-treated HT22 cells, and silencing FOXO3a effectively inhibited the expression of BAP1 and increased the expression level of SLC7A11.3.5 Overexpressed FOXO3a aggravated neuronal ferroptosis after treatment with hemin."

reach
"The FOXO3a overexpression plasmid was also used to transfect HT22 cells, qPCR showed that overexpression of FOXO3a significantly increased the mRNA transcription of BAP1 and inhibited the mRNA level of SLC7A11 after treatment with hemin (Fig. 5A–B)."

reach
"After treatment with hemin, FOXO3a overexpression significantly increased the expression of FOXO3a and BAP1 proteins, and inhibited the protein expressions of SLC7A11 and GPX4 in HT22 cells (Fig. 5D–F)."

reach
"Overexpression of FOXO3a could increase the expression of BAP1 and aggravate ferroptosis in neurons."

reach
"Silencing FOXO3a significantly suppressed the expression of BAP1, modulated the levels of SLC7A11 and GPX4, and attenuated the level of lipid peroxidation and mitochondrial damage in neuron."
FOXK1 affects BAP1
12 | 3 9
12 | 3 9

No evidence text available

sparser
"Thus, O-GlcNacylated FOXK1 associates with BAP1 and promote E2F pathway and oncogenesis."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."

sparser
"Here the authors find that the O-GlcNAcylated FOXK1 associates with the deubiquitinase BAP1 to upregulate transcription of E2F target genes and promote cancer progression."

No evidence text available

No evidence text available

reach
"By comparing BAP1 recruitment in IMR90 cells expressing FOXK1 with those expressing FOXK1 , we identified 2,130 regions showing reduced recruitment of BAP1 in FOXK1 (Fig. 6e)."

sparser
"For example, analysis of 407 pleural mesothelioma cases and 389 controls with a comprehensive history of asbestos exposure revealed an increased risk of abnormalities in chromosomal region 7p22.2, which includes the gene encoding the Forkhead box protein K1 (FOXK1) that interacts with BAP1 [ xref ]."
| PMC

No evidence text available

No evidence text available
BAP1 affects SP1
2 | 12 10
BAP1 binds SP1.
2 | 4 10
2 | 4 10

sparser
"Therefore, ODN enhanced oxaliplatin-induced apoptosis by interacting with BAP1 and Sp1 to induce Sp1 stabilization-mediated Bax upregulation ( xref )."

sparser
"ODN induces BAP1 interaction with Sp1."

sparser
"We postulated that ODN may influence the interaction between BAP1 and Sp1."

sparser
"Interestingly, ODN and cathepsin K siRNA treatment markedly increased Sp1 and BAP1 interaction ( xref A)."

sparser
"Sp1-BAP1 interaction was increased by ODN treatment ( xref F)."

sparser
"ODN induced the phosphorylation of BAP1 at Ser 592 and enhanced the interaction of Sp1 and BAP1, resulting in Sp1 deubiquitination and destabilization."

reach
"We postulated that ODN may influence the interaction between BAP1 and Sp1."

sparser
"Mechanistically, ODN induced BAP1 phosphorylation and enhanced Sp1-BAP1 interaction, resulting in Sp1 ubiquitination and degradation."

sparser
"To investigate the functional role of p53, we tested the interaction between Sp1 and BAP1 after p53 immunoprecipitation in ODN- and cathepsin K siRNA-treated cells and identified more p53, Sp1, and BAP1 protein complexes in the ODN- and cathepsin K siRNA-treated cells ( xref B)."

sparser
"These data demonstrated that the C-terminal domain of p53 can effectively enhance Sp1 stability by increasing the interaction with BAP1 and Sp1."
BAP1 decreases the amount of SP1.
| 4
BAP1 decreases the amount of SP1. 4 / 4
| 4

reach
"WT BAP1 reduced the ubiquitination levels of Sp1, whereas C91A BAP1 did not reduce ubiquitination levels (Fig. 6D)."

reach
"The knockdown or catalytic mutant form of BAP1 abolished the ODN-induced upregulation of Sp1 and Bax expression."

reach
"In addition, the ectopic expression of BAP1 greatly decreased the ubiquitination level of Sp1 and the upregulation of Sp1 and Bax, whereas the catalytic mutant of BAP1 showed no apparent effect (Fig. 6B–D)."

reach
"OTUB1, OTUD7A, and BAP1 downregulated Sp1 and Bax expression (Fig. 5)."
BAP1 activates SP1.
| 4
BAP1 activates SP1. 4 / 4
| 4

reach
"Therefore, BAP1 was involved in ODN-mediated Sp1 stabilization and might be a potential target for cancer therapy in drug development."

reach
"Mitochondrial ROS scavengers prevented DNA damage, BAP1-mediated Sp1 stabilization, and Bax upregulation by ODN."

reach
"BAP1 phosphorylation-mediated Sp1 stabilization plays a critical role in cathepsin K inhibition-induced C-terminal p53-dependent Bax upregulation."

reach
"ODN-induced Bax upregulation was regulated by Sp1 stabilization, which was modulated by BAP1-dependent deubiquitination mediated by the C-terminal domain of p53."
BAP1 affects INO80
2 | 18 3
BAP1 activates INO80.
| 9
BAP1 activates INO80. 9 / 10
| 9

reach
"Importantly, Ino80 levels are often reduced in BAP1-null mesothelioma cells, which lack a BAP1 mediated Ino80 stabilization mechanism, and downregulated in BAP1 defective cancer cells in mesothelioma patients [XREF_BIBR]."

reach
"Importantly, a correlation was found between BAP1 and INO80 expression in mesothelioma, and re-expression of BAP1 in H226 cells exhibiting low levels of INO80 fully rescued the INO80 levels, showing that the low INO80 expression level in H226 cells was due to a lack of BAP1-mediated INO80 stabilization ."

reach
"Ino80 is downregulated in BAP1 defective mesothelioma, owing to the lack of a BAP1 mediated Ino80 stabilization mechanism, thus raising the possibility of a tumorigenic role for INO80 in this cancer."

reach
"While individual expression of CHIP and BAP1 increased Ino80, as expected, coexpression of these proteins increased endogenous Ino80 (XREF_FIG) and transfected Flag-Ino80 beyond the levels achieved by individual expression of either protein (XREF_FIG)."

reach
"Therefore, BAP1 promoted replication fork progression via dual mechanisms: stabilization via deubiquitination and recruitment of INO80 to the replication fork."

reach
"Conversely, knockdown of CHIP, BAP1, or both decreased Ino80 preferentially in the chromatin fractions (XREF_FIG)."

reach
"One reason for this could be that the simultaneous knockdown of CHIP and BAP1 decreased Ino80 only marginally compared to the individual knockdown, which is likely due to a limitation of siRNA knockdown."

reach
"By recruiting INO80 to the stalled forks, BAP1 allows stress induced stalled replication forks to restart, a mechanism that suppresses genome instability and thus cancer development."

reach
"Linked to this is the role of BAP1 in replication-coupled repair, where BAP1 aids to recruit and stabilise the chromatin remodeller INO80 to sites of DNA damage or replication stress, possibly to promote resection through its remodelling activity [116] (Figures 2A and 3A)."
BAP1 binds INO80.
2 | 3 3
2 | 3 3

sparser
"BAP1 and INO80 bound to replication forks in response to replication stress induced by nucleotide depletion after hydroxyurea (HU) treatment."

sparser
"Both BAP1 and INO80 bind chromatin at replication forks upon HU treatment."

reach
"Both BAP1 and INO80 bind chromatin at replication forks upon HU treatment."

No evidence text available

reach
"A recent study demonstrates that Ino80 interacts with BRCA1 associated protein-1 (BAP1), a tumor suppressor that also stabilizes Ino80, in normal DNA replication [XREF_BIBR]."

No evidence text available

reach
"BAP1 interacts with and deubiquitylates INO80, playing a critical role in stabilizing and targeting the INO80 chromatin remodeling complex to the replication forks."

sparser
"Both BAP1 and INO80 bound to replication forks during S phase, and BAP1 depletion inhibited both INO80 recruitment to the fork and fork progression."
BAP1 deubiquitinates INO80.
| 6
BAP1 deubiquitinates INO80. 6 / 6
| 6

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"BAP1 overexpression also abolished basal Ino80 ubiquitination (XREF_FIG and XREF_FIG, rightmost lanes)."

reach
"BAP1 deubiquitylates INO80 and thereby protects the protein from ubiquitin mediated proteolysis."

reach
"BAP1 interacts with and deubiquitylates INO80, playing a critical role in stabilizing and targeting the INO80 chromatin remodeling complex to the replication forks."

reach
"In line with this notion, BAP1 depletion resulted in reduced localization of RPA, RAD51 and BRCA1 to IRIF or microlaser irradiation site [100,101], although it remains ambiguous whether either histone[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Since BAP1 deubiquitinates and stabilizes Ino80, we investigated the relationship between CHIP and BAP1 in Ino80 ubiquitination."

reach
"In addition to MCRS1, a recent study revealed that BAP1 deubiquitinates and stabilizes Ino80, a key subunit of the INO80 chromatin remodeling complex implicated in DNA replication during stress."
BAP1 affects HSF1
| 14 10
BAP1 binds HSF1.
| 5 10
| 5 10

sparser
"BAP1-HSF1 interaction inhibits the deacetylation of HSF1-K80 mediated by SIRT1, thereby promoting HSP70 mediated monomerization of HSF1 and its dissociation from target gene loci."

reach
"This may suggest that the binding between BAP1 and HSF1 could inhibit the ubiquitination of BAP1 mediated by UBE2O [24], or there may be other unexplored mechanisms at play."

sparser
"Co-immunoprecipitation assays indicated that treatment with the SIRT1 inhibitor, EX-527, notably augmented the level of acetylated lysine of HFS1, as well as the interaction between HSF1 and BAP1 (Fig.  xref A)."

sparser
"The BAP1-HSF1 interaction preserves the acetylation of HSF1-K80 and promotes HSF1-HSP70 interaction, facilitating HSF1 oligomerization and detachment from the chromatin."

sparser
"The BAP1-HSF1 interaction safeguards the acetylation of HSF1 at the K80 site, promotes the monomerization of HSF1, and facilitates its dissociation from DNA."

reach
"More importantly, both wild-type (WT) and C91A mutated BAP1 attenuated the HSF1-SIRT1 interaction (Fig. 5C, Supplementary Fig. S3G), and SIRT1 knockdown resulted in an enhanced binding between HSF1 and BAP1 (Fig. 5D, Supplementary Fig. S3H), indicating that BAP1 and SIRT1 might compete with each other for binding to HSF1."

sparser
"BAP1 interacts with the N-terminal of HSF1."

sparser
"The subsequent Co-IP assay revealed that the N-terminal of HSF1 was pivotal for the formation of the BAP1-HSF1 complex (Fig.  xref F)."

reach
"Co-immunoprecipitation assays indicated that treatment with the SIRT1 inhibitor, EX-527, notably augmented the level of acetylated lysine of HFS1, as well as the interaction between HSF1 and BAP1 (Fig. 6A)."

sparser
"Interestingly, the interaction between BAP1 and HSF1 exclusively occurs in the nucleus (Fig.  xref B)."
BAP1 inhibits HSF1.
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BAP1 inhibits HSF1. 6 / 6
| 6

reach
"BAP1 represses the transcriptional activity of HSF1."

reach
"As the data above has confirmed that BAP1 inhibits the transcriptional activity of HSF1, we selected genes that are downregulated by HSF1 KD for enrichment analysis."

reach
"Furthermore, RT-qPCR and HSF1 ChIP-qPCR analysis with several classical genes (ATF4 and CREB5) of TNF signaling pathway indicated that BAP1 represses their expression via HSF1 (Supplementary Fig. S2C-D)."

reach
"To delve deeper into how BAP1 suppresses HSF1 activity through protein-protein interaction (PPI) with HSF1, we proceeded to construct truncations of HSF1 based on its major functional domains (Fig. 3E)."

reach
"Additionally, EX-527 treatment attenuated the upregulation of HSF1 activity induced by BAP1 deletion (Fig. 6B)."

reach
"However, these interrogations do not undermine our conclusion that BAP1 inhibits HSF1 transcriptional activity independently of its enzymatic activity.As previously stated, the significance of BAP1 in maintenance of genomic integrity, DNA damage repair, cellular metabolism, and cell death is relatively well-established [27–30]."
BAP1 activates HSF1.
| 3
BAP1 activates HSF1. 3 / 3
| 3

reach
"These findings imply that the absence of BAP1 can augment cellular sensitivity to specific therapeutic interventions, thereby furnishing a theoretical rationale for therapies aimed at BAP1-associated cancers.Additionally, our prior research has unveiled that BAP1 can modulate the activity of HSF1 through a non-enzymatic mechanism, ultimately impacting the responsiveness to immune checkpoint inhibitors33."

reach
"Given that BAP1 modulates the transcriptional activity of HSF1 without impacting its expression, we sought to investigate whether BAP1 exerts its influence on HSF1 activity through post-transcriptional mechanisms."

reach
"However, the luciferase report gene assay indicated that the knock-out of BAP1 notably upregulates the transcriptional activity of HSF1 (Fig. 2F)."
BAP1 affects FOXK1
12 | 3 9
12 | 3 9

No evidence text available

sparser
"Thus, O-GlcNacylated FOXK1 associates with BAP1 and promote E2F pathway and oncogenesis."

reach
"BAP1 also associates with transcription factors, including FOXK1, FOXK2, and Yin Yang1 (YY1), which ensure the recruitment of this DUB to chromatin (133, 142)."

sparser
"Here the authors find that the O-GlcNAcylated FOXK1 associates with the deubiquitinase BAP1 to upregulate transcription of E2F target genes and promote cancer progression."

No evidence text available

No evidence text available

reach
"By comparing BAP1 recruitment in IMR90 cells expressing FOXK1 with those expressing FOXK1 , we identified 2,130 regions showing reduced recruitment of BAP1 in FOXK1 (Fig. 6e)."

sparser
"For example, analysis of 407 pleural mesothelioma cases and 389 controls with a comprehensive history of asbestos exposure revealed an increased risk of abnormalities in chromosomal region 7p22.2, which includes the gene encoding the Forkhead box protein K1 (FOXK1) that interacts with BAP1 [ xref ]."
| PMC

No evidence text available

No evidence text available
BAP1 affects DNA repair
| 4 20
BAP1 activates DNA repair.
| 4 15
| 4 15

eidos
"BAP1 is recruited to double strand breaks ( DSB ) and promotes DNA repair and survival after DNA damage induction [ 18,19,20 ] ."

reach
"BAP1 can deubiquitylate Ub-H2AK119 at the site of UV-induced DNA damage, and promote DNA repair by regulating the accumulation of repair proteins (BRCA1, RAD51, and RPA) (7)."

eidos
"BAP1 can deubiquitylate Ub-H2AK119 at the site of UV-induced DNA damage , and promote DNA repair by regulating the accumulation of repair proteins ( BRCA1 , RAD51 , and RPA ) ( 7 ) ."

reach
"In the nucleus, BAP1 modulates the transcriptional regulation of several gene programs and promotes DNA repair by facilitating homologous recombination ."

reach
"Indeed, by using a green fluorescent protein (GFP)-based reporter assay employing a recognition site for the rare-cutting I-SceI endonuclease for induction of DSBs , we found that loss of BAP1 caused a substantial reduction of GFP-positive cells, indicating loss of proper DNA repair (Supplementary Fig. 5k)."

reach
"Interestingly, BRCA1 proteins were decreased in the MM cells with BAP1 deletion, and transduction of the mutants as well as WT BAP1 increased BRCA1 proteins, suggesting that BAP1 may promote DNA repair partly through stabilizing BRCA1."

reach
"Although LOH of additional genes within Chr 3p21.1 may contribute to tumor suppression, and loss of BAP1 in pancreatic cancer may be secondary—rather than the cause of genomic instability—our mouse model suggests Bap1 ablation suffices to cause defective DNA repair, pancreatitis, and cooperates with oncogenic Kras to promote pancreatic cancer."

eidos
"BAP1 promotes DNA repair , regulates transcription , cell differentiation , and cell death 23 ."

reach
"Reduced BAP1 levels impair DNA repair as well as different forms of cell death and induce metabolic alterations that together favor cancer development and growth."

reach
"Additionally, BAP1 modulates transcription, DNA repair, and replication by regulating chromatin remodeling via histone H2AK119 deubiquitination (12)."
BAP1 inhibits DNA repair.
| 5
| 5

reach
"Specifically, BAP1 suppresses genome instability via its DNA repair activity and eliminates cancer-prone cells that exhibit genome instability by promoting apoptosis."

reach
"The powerful activity of heterozygous germline BAP1 mutations to facilitate asbestos induced transformation and cause mesothelioma has been linked to the alterations in the mechanisms that regulate DNA repair and cell death caused by decreased BAP1 protein levels."

reach
"Functional studies were not conducted because cells from these patients were not available.Our findings revealed that both BAP1 and BARD1 inactivate similar pathways: DNA repair, calcium signaling, and cell death, although through different mechanisms."

reach
"Defects in BAP1 disrupt DNA replication, DNA repair, calcium homeostasis, cell proliferation and differentiation, and cell metabolism [34]."

reach
"BAP-1 deficiency causes damage to DNA as well as defects in the DNA repair mechanism [103]."
ASXL3 affects BAP1
1 6 | 8 8
ASXL3 binds BAP1.
1 6 | 6 7
1 6 | 6 7

No evidence text available

sparser
"Our in vitro analysis indicates that also ASXL3 can bind and activate BAP1, which is not surprising in the light of the high sequence similarity between the DEUBAD domains in the ASXL family."

No evidence text available

reach
"Mechanistically, the chromatin bound ASXL3 functions as an adaptor protein, which directly connects BRD4 to the BAP1/ASXL3 complex and maintains chromatin occupancy of the functional BAP1/ASXL3/BRD4 epigenetic axis at active enhancers (20)."

No evidence text available

"We report a critical link between BAP1 complex and BRD4, which is bridged by the physical interaction between ASXL3 and BRD4 in an SCLC subtype (SCLC-A), which expresses a high level of ASCL1. We further showed that ASXL3 functions as an adaptor protein, which directly interacts with BRD4's extra-terminal (ET) domain via a novel BRD4 binding motif (BBM), and maintains chromatin occupancy of BRD4 to active enhancers."

No evidence text available

No evidence text available

reach
"ASXL3 interacts with BAP1, a ubiquitin terminal hydroxylase and removes the mono-ubiquitin (Ub1) from the H2AK119Ub1 [XREF_BIBR]."

reach
"We found that ASXL3 interacts with BAP1, a hydrolase that removes mono-ubiquitin from histone H2A lysine 119 (H2AK119Ub1) as a component of the Polycomb repressive deubiquitination (PR-DUB) complex."
ASXL3 activates BAP1.
| 2 1
ASXL3 activates BAP1. 3 / 3
| 2 1

reach
"The DEUBAD domains of ASXL1, ASXL2 and ASXL3 can activate BAP1 by increasing its affinity for ubiquitin."

reach
"In this paper we describe that similar to the homologous UCH-L5 and RPN13 DEU complex, the DEUBAD domains of ASXL1, ASXL2 and ASXL3 can activate BAP1 by increasing BAP1 's affinity specifically for the ubiquitin in the substrate, through a combination of mild effects."

sparser
"Our in vitro analysis indicates that also ASXL3 can bind and activate BAP1, which is not surprising in the light of the high sequence similarity between the DEUBAD domains in the ASXL family."
BAP1 affects Syndrome
| 23
| 21

reach
"BAP1 tumor predisposition syndrome is caused by pathogenic germline variants in the BAP1 tumor suppressor gene and characterized by a predisposition to various tumors including meningioma."

reach
"Pathogenic germline variants in the BRCA1-associated protein 1 (BAP1) gene cause a dominantly inherited tumor predisposition syndrome, BAP1-TPDS, in which MM is the second most common associated cancer."

reach
"BAP1-Tumor Predisposition Syndrome (TPDS) is caused by germline variants in BAP1 and predisposes to solid tumors."

reach
"Pathogenic germline variants in the BRCA1-Associated Protein 1 (BAP1) tumor suppressor gene cause a rare, life-threatening tumor predisposition syndrome (BAP1-TPDS, OMIM 614327) also known as BAP1 cancer syndrome as the majority of carriers develop malignancy [1, 2]."

reach
"Furthermore, BAP1 germline mutations cause a tumor predisposition syndrome inherited in an autosomal dominant pattern, which confers a higher risk for developing a spectrum of benign and malignant tumors , including pancreatic cancer ."

reach
"Additionally, melanoma risk is elevated in mixed cancer syndromes caused by mutations in PTEN, BRCA1, BRCA2, RB1, BAP1 and TP53 [56]."

reach
"(3)In previous studies, we found that heterozygous germline mutations in the BAP1 gene cause the BAP1 Cancer Syndrome, characterized by a high incidence of mesothelioma (4–9)."

reach
"Melanoma predisposition syndromes caused by pathogenic variants in POT1 and BAP1 are more recently described, and both are associated with Spitzoid tumors."

reach
"Altogether, these results define a clinical syndrome caused by rare germline missense BAP1 variants that alter chromatin remodeling through abnormal histone ubiquitination and lead to transcriptional dysregulation of developmental genes."

reach
"The presence of multiple BAP1-negative melanocytic neoplasms is a hallmark of familial cancer susceptibility syndrome caused by germline mutations in BAP1."
Mutated BAP1 activates Syndrome. 2 / 2
| 2

reach
"Whereas BAP1 mutations drive BAP1 tumour predisposition syndrome, mesotheliomas, melanomas and other neoplasms 8–11 ; ASXL1 mutations are particularly prevalent in myeloid cancers, and are associated [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 mutation can increase the sensitivity of HNSCC to radiotherapy and lead to tumor susceptibility syndrome, which is not related to the status of high-risk human papillomavirus (HPV) and p53 tumor suppressor protein."
BAP1 affects FOXO3
| 9 14
BAP1 binds FOXO3.
| 2 14
| 2 14

sparser
"Both endogenous and exogenous CoIP demonstrated that BAP1 protein could directly bind to FOXO3a protein in cells."

sparser
"Moreover, the binding of the transcription factor FOXO3a with the BAP1 promoter can modulate neuronal lipid peroxidation and ferroptosis by enhancing BAP1 expression."

sparser
"Consequently, the FOXO3a-BAP1 axis might represent a promising therapeutic strategy for addressing neuronal ferroptosis in patients with SAH."

sparser
"Our findings elucidate the impact of FOXO3a and BAP1 on SLC7A11-related ferroptosis following SAH both in vivo and in vitro , and the inhibition of the FOXO3a-BAP1 axis can significantly attenuate neuronal damage and ferroptosis in EBI after SAH."

sparser
"Preliminary experiments have revealed that FOXO3a can bind to the BAP1 promoter and regulate the expression of BAP1."

sparser
"Consequently, this study aimed to investigate the potential mechanism by which FOXO3a-BAP1 influences neuronal ferroptosis in the pathogenesis of EBI after SAH."

sparser
"BAP1 directly interacted with FOXO3a."

sparser
"To explore whether DUB BAP1 directly regulates the above proteins through deubiquitination in neurons after SAH, the binding of BAP1 with FOXO3a proteins was detected by prediction of protein-protein interactions and CoIP assay."

sparser
"Endogenous CoIP also revealed that BAP1 protein could directly bind to FOXO3a protein in HT22 cells ( xref B)."

reach
"To explore whether DUB BAP1 directly regulates the above proteins through deubiquitination in neurons after SAH, the binding of BAP1 with FOXO3a proteins was detected by prediction of protein-protein interactions and CoIP assay."
BAP1 activates FOXO3.
| 4
BAP1 activates FOXO3. 4 / 4
| 4

reach
"These results suggested that BAP1 can mediate SLC7A11 inhibition and neuronal ferroptosis by regulating H2Aub.3.3 FOXO3a regulated BAP1 expression at the transcriptional level."

reach
"This indicated that the increase in BAP1 after SAH could significantly reduce the degradation of FOXO3a protein for increasing the stability of FOXO3a protein, and inhibiting BAP1 can promote the degradation of FOXO3a protein."

reach
"Furthermore, BAP1 protein appears to inhibit FOXO3a degradation via the proteasome-dependent pathway, thereby preserving the activity and stability of the FOXO3a protein."

reach
"These results indicated that FOXO3a was activated in hemin-treated HT22 cells, and silencing FOXO3a effectively inhibited the expression of BAP1 and increased the expression level of SLC7A11.3.5 Overexpressed FOXO3a aggravated neuronal ferroptosis after treatment with hemin."
BAP1 inhibits FOXO3.
| 3
BAP1 inhibits FOXO3. 3 / 3
| 3

reach
"This indicated that the increase in BAP1 after SAH could significantly reduce the degradation of FOXO3a protein for increasing the stability of FOXO3a protein, and inhibiting BAP1 can promote the degradation of FOXO3a protein."

reach
"Furthermore, BAP1 protein appears to inhibit FOXO3a degradation via the proteasome-dependent pathway, thereby preserving the activity and stability of the FOXO3a protein."

reach
"After transfection of Flag-FOXO3a, HA-BAP1 and various ubiquitin mutants (K48, K63, K48R and K63R) into 293T cells, it was observed that BAP1 predominantly reduced the ubiquitin modification of the FOXO3a protein at the K48 position, thereby verified the experimental findings regarding the half-life of FOXO3a."
| 23
BAP1 activates Carcinogenesis.
| 14
| 11

reach
"The strongest evidence implicating BAP1 (and PBRM1) in promoting ccRCC tumorigenesis and regulating aggressiveness is provided by genetically engineered mouse models (GEMM)."

reach
"PVT1 binding with KLF5 causes KLF5 to stabilise and this elevates the BAP1 levels, which in turn promotes beta-catenin signalling, thus enhancing TNBC carcinogenesis (Li et al., 2018)."

reach
"The fact that we determined that highly penetrant, heterozygous pathogenic BAP1 null variants by themselves, as studied by Kadariya et al. , can initiate and drive MM tumorigenesis in mice absent asbestos exposure supports their conclusions, in agreement with Chanock ."

reach
"While RNF2, through H2A monoubiquitination, silences the BCL2 and MCL1 prosurvival genes and thereby induces apoptosis in some cell types, this does not occur in melanocytes, suggesting that BAP1 promotes tumorigenesis in cells that do not engage such RNF2 apoptotic programme.Moving to the p53 tumor suppressor, Karen Vousden (London, UK), who was awarded the CDD Award 2018, retraced the long road leading to her key discoveries of p53 functions."

reach
"It is expected that BAP1 promotes tumorigenesis of CCA by modulating DNA repair, gene transcription, and post-translational modification, such as glycosylation."

reach
"Besides, a study of families with BAP1 germline mutations shows that mutated BAP1 may promote oncogenesis and increase the risk of KIRC by 8-fold through altering cellular metabolism and the tumor microenvironment (25)."

reach
"Deficiency of BAP1 inhibits neuroblastoma tumorigenesis through destabilization of MYCN."

reach
"BAP1 PGVs accelerate mesothelioma tumorigenesis after asbestos exposure in preclinical models and may be partly predicted by clinical criteria."

reach
"42 Other research has speculated that SF3B1 and BAP1 variants can independently lead to oncogenesis and/or malignant progression, reducing the need of their co-occurrence in the natural course of the disease."

reach
"For instance, in the ~40% of tumors that are seemingly wild-type for both PBRM1 and BAP1, what other events cooperate with VHL loss to promote oncogenesis?"
Mutated BAP1 activates Carcinogenesis. 3 / 3
| 3

reach
"There has been increased interest in the mechanisms by which BAP1 mutations lead to carcinogenesis."

reach
"We aim to give a concise review of the carcinogenesis caused by BAP1 mutations, and discuss clinical implications including suggested screening guidelines for patients with germline BAP1 mutations, and potential targeted therapeutic options in patients with mutated BAP1."

reach
"With respect to cancer biology, BAP1 loss confers metabolic advantages to cancer cells that might partially account for how BAP1 mutations increase the susceptibility of tumorigenesis (62,65)."
BAP1 inhibits Carcinogenesis.
| 9

reach
"Therefore, it will be important to investigate the mechanism underlying the tumor-promoting effect of BAP1 inactivation in each cell type.In summary, this study demonstrated that Bap1 loss promotes tumorigenesis only in the tissues in which Bap1 is not engaged in pro-survival gene expression, providing an explanation for the tissue specificity of the BAP1 cancer predisposition syndrome."

reach
"If the loss of either NF2 or BAP1 mediates oncogenesis via the perturbation of Hippo signalling, the failure to take into account the additional Hippo kinase module inactivating events present in PM may have confounded previous efforts to stratify patients according to mutation status."

reach
"It remains to be established as to how the oxidation-mediated functional loss and misfolding of BAP1 may lead to oncogenesis in a similar mechanism."

reach
"A subsequent study reported that BAP1 modulated gene expression in a cell-type specific manner, and loss of BAP1 promoted tumorigenesis in cells that did not engage an RNF2-dependent apoptotic program ."

reach
"BAP1 can promote ferroptosis and suppress tumor tumorigenesis [39]."

reach
"Loss of one BAP1 allele either inherited or acquired during life has been associated with environmental stress-induced carcinogenesis, like UV light for uveal melanoma and asbestos for mesothelioma."

reach
"In mice, BAP1 loss in the germ line is embryonic lethal (10), and BAP1 loss in the developing kidney causes differentiation failure and tumorigenesis (11)."

reach
"Paradoxically, despite the fact that BAP1 loss promotes tumorigenesis, BAP1 inactivation causes cell death in many cell types."

reach
"p53 and BRCA1 Associated Protein 1 (BAP1) inhibit carcinogenesis in part by antagonizing SLC7A11 and induce ferroptosis (Zhang Y. et al., 2019)."
| 18 4
| 11

reach
"BAP1 stabilizes BRCA1 protein and restores cell survival rates of MM cells with BAP1 deletion after IR."

reach
"While most human MMs exhibit somatic alterations of BAP1, NF2 and/or CDKN2A, currently it is not known if inactivation of BAP1 cooperates with loss of NF2 and/or CDKN2A to drive a more aggressive MM phenotype."

reach
"Importantly, however, our data suggest that dysregulation of the Rb path way may be required for Bap1 driven MM pathogenesis, at least in an experimental model."

reach
"Recently, Bap1 +/- knockout mice were found to have increased susceptibility to asbestos induced MM compared to wild type litter mates (Xu et al., 2014)."

reach
"Rather, the presence of a wild type allele in most studied cases, suggests that BAP1 haplo-insufficiency may lead to peritoneal MM."

reach
"While homozygous CKO of Bap1, Cdkn2a, or Nf2 alone gave rise to few or no MMs, inactivation of Bap1 cooperated with loss of either Nf2 or Cdkn2a to drive development of MM in ~ 20% of double-CKO mice, and a high incidence (22/26, 85%) of MMs was observed in Bap1; Nf2; Cdkn2a (triple)-CKO mice."

reach
"There was no association between total SNV and survival; ever versus never smokers; or patients with epithelioid versus non epithelioid histologies.The three archetypical somatic drivers of MM - BAP1,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 4

reach
"We are investigating how germline BAP1 mutations on one hand promote MM development, while on the other hand they are associated to reduce disease aggressiveness."

reach
"A possible explanation is that germline BAP1 mutations alone can cause MM and many other tumor types."

reach
"BAP1 is the most promising driver gene in MM pathogenesis, because more than 60% of MM biopsies contain BAP1 mutations and inactivation [XREF_BIBR, XREF_BIBR] and germline BAP1 mutations cause a cancer syndrome with high incidence of MM [XREF_BIBR, XREF_BIBR]."

reach
"We recently reported in vivo genetic evidence that germline heterozygous mutation of Bap1 accelerates development of asbestos induced MM."

reach
"BAP1 suppresses MM cell proliferation."

reach
"In effusions with mesotheliomatous cells or atypical mesothelial cells of uncertain significance, negative BAP1 IHC strongly supports a diagnosis of MM."

reach
"In this study, we observed BAP1 expression loss in five of eight epithelioid-type MM cases (62.5%), while also observing loss in both sarcomatoid and epithelioid components in five of 13 biphasic-type MM cases (38.5%)."

reach
"Our results suggested that, whereas WT BAP1 suppresses MM cell proliferation and restores cell survival after IR damage, some mutant BAP1 may also moderately retain these functions."

reach
"Loss of BAP1 is frequently observed in MM and has been proven to increase the risk of MM [XREF_BIBR - XREF_BIBR]."

reach
"Transduction of the WT BAP1 vector inhibited cell proliferation of the Y-MESO-25 MM cell line with BAP1 deletion by approximately 50% compared to the cells with control vector (GFP)."

reach
"With the colony formation assay, we also found that WT BAP1 suppressed the anchorage independent cell growth of the MM cells with BAP1 deletion, whereas mutant BAP1 showed little or no reduced effect."

sparser
"The demonstration that BAP1 mutations are strongly associated with MM has provided a real breakthrough in the study of genetic predisposition in MM, that may explain why only a fraction of asbestos-exposed individuals go on to develop MM."

sparser
"BAP1 knockdown in MM cell lines has been paradoxically associated to a decreased proliferation, with an accumulation of cells in the S phase of the cell cycle xref , suggesting that BAP1 loss might promote tumorigenesis inducing a delayed, but more permissive, G1/S checkpoint xref ."

sparser
"Here, we tested the hypothesis that MM associated with germline BAP1 mutations has a better prognosis compared with sporadic MM."

sparser
"The association of germline BAP1 mutations with familial MM has been confirmed in a series of recent reports ( xref - xref )."
| 1 14 1

reach
"For example, one study showed that BAP1 promotes the development of breast cancer by enhancing the stability of the transcription factor KLF5 (30)."

reach
"p53 binds to the SLC7A11 promoter, directly suppressing its transcription; the nuclear deubiquitinase (DUB) BAP1 represses SLC7A11 transcription by removing histone 2A ubiquitination from the SLC7A11 promoter; and KEAP1 represses SLC7A11 transcription through degrading NRF2, a master transcription factor of antioxidant response and regulator of SLC7A11."

reach
"Previously, we found that hypoxia-inducible factor 2α (HIF2α) and BAP1 activate interferon-stimulated gene factor 3 (ISGF3), a transcription factor activated by type I interferons and a tumor suppressor in ccRCC xenograft models."

reach
"BAP1 complex promotes transcription by opposing PRC1 mediated H2A ubiquitylation."

reach
"As a deubiquitinating enzyme, BAP1 contributes to gene transcription, cell differentiation, DNA damage repair, apoptosis, and cell metabolism in tumor inhibition."

eidos
"Taken together , these results establish that PRC1 is epistatic to BAP1.com and that BAP1 promotes transcription by counteracting PRC1-mediated H2A ubiquitination ( Fig. 7 ) ."

reach
"These included the SETD1A, KMT2C and MLL3, KMT2E and MLL5, and KMT2A and MLL1 methyltransferases that install activating histone H3-lysine 4 trimethylation (H3K4me3) marks; OGT (O-GlcNac transferase), a protein that modulates multiple cell pathways but more recently has been implicated in promoting promoter occupancy of RNAPII; and BAP1 (BRCA1 Associated Protein 1), a deubiquitinase that stimulates transcription, at least in part, by removing repressive histone H2A-K119 monoubiquitin."

reach
"Using RNAi, we found that certain deubiquitinases, BAP1, USP14 and OTUD5 each promote proviral transcription."

reach
"Luchini et al. also reported that BAP1 mutated clear cell renal carcinomas were frequently observed in females and were associated with high tumor grade (p < 0.0001), increased all-cause mortality, cancer-specific mortality, and risk of recurrence.NCOR1P1, Nuclear Receptor Corepressor 1 Pseudogene 1, is predicted to enable transcription corepressor activity, and to be involved in the negative regulation of transcription by RNA polymerase II ."

reach
"Additionally, BAP1 modulates transcription, DNA repair, and replication by regulating chromatin remodeling via histone H2AK119 deubiquitination (12)."

eidos
"Similarly , BAP1 , an epigenetic regulator and tumor suppressor mutated in various types of cancer , can downregulate the transcription of SLC7A11 and thus potentiate ferroptosis ( Zhang et al ., 2018 ) ."

eidos
"Moreover , ZEB1 promoted down-regulation of crucial genes involved in melanocyte differentiation , including BAP1 , melanocyte inducing transcription factor ( MITF9 ) , tyrosinase ( TYR ) , and tyrosinase-related protein 1 ( TYRP ) ."

eidos
"Under glucose starvation , BAP1 can directly bind to the promoter of ATF3 and CHOP , inhibit its transcription , and thus inhibit the UPR and cell death induced by the ER stress ( Figure 2p ) ."

reach
"In general, various molecules regulate SLC7A11, such as tumor suppressor P53, tumor suppressor BRCA1-associated protein 1 (BAP1), activating transcription factor 3/4 (ATF3/ATF4), beclin 1 (BECN1), and nuclear factor erythroid 2-related factor 2 (NRF2) (Figure 3)."

reach
"BAP1 also represses or activates transcription by associating with or regulating other transcription factors (36)."
BAP1 affects ASXL3
1 6 | 6 7
1 6 | 6 7

No evidence text available

sparser
"Our in vitro analysis indicates that also ASXL3 can bind and activate BAP1, which is not surprising in the light of the high sequence similarity between the DEUBAD domains in the ASXL family."

No evidence text available

reach
"Mechanistically, the chromatin bound ASXL3 functions as an adaptor protein, which directly connects BRD4 to the BAP1/ASXL3 complex and maintains chromatin occupancy of the functional BAP1/ASXL3/BRD4 epigenetic axis at active enhancers (20)."

No evidence text available

"We report a critical link between BAP1 complex and BRD4, which is bridged by the physical interaction between ASXL3 and BRD4 in an SCLC subtype (SCLC-A), which expresses a high level of ASCL1. We further showed that ASXL3 functions as an adaptor protein, which directly interacts with BRD4's extra-terminal (ET) domain via a novel BRD4 binding motif (BBM), and maintains chromatin occupancy of BRD4 to active enhancers."

No evidence text available

No evidence text available

reach
"ASXL3 interacts with BAP1, a ubiquitin terminal hydroxylase and removes the mono-ubiquitin (Ub1) from the H2AK119Ub1 [XREF_BIBR]."

reach
"We found that ASXL3 interacts with BAP1, a hydrolase that removes mono-ubiquitin from histone H2A lysine 119 (H2AK119Ub1) as a component of the Polycomb repressive deubiquitination (PR-DUB) complex."
SETD2 affects BAP1
| 1 19
| 1 10

sparser
"Across clear cell- and papillary-associated RCC subtypes in TCGA cohort, worse survival was associated with SETD2 or BAP1 mutation, with their related gene transcriptional signatures involving greater numbers of RCC cases and also being predictive of worse outcome ( xref , xref )."

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"BAP1 and SETD2 mutations (6–12%) are associated with worse CSS, suggesting their roles in disease progression."

sparser
"Furthermore, PBRM1 mutations have been found to confer a slight increase of death risk, while SETD2 or BAP1 mutations were associated with serious death risks in ccRCC patients [ xref – xref ]."

sparser
"Similarly, a study of 609 patients with ccRCC from two distinct cohorts revealed that mutations of 3p21 epigenetic regulators BAP1 and SETD2 were associated with worse cancer-specific survival (CSS), suggesting that they play key roles in disease progression[ xref ]."

sparser
"In pleural and peritoneal mesothelioma, co-occurring alterations occurred on the same chromosome or in close proximity for PBRM1–BAP1 (both 3p21) and SETD2BAP1 (both 3p21), which was observed in the TCGA cohort but not in the recent publication by Zauderer and colleagues [ xref ]."

reach
"The association between VHL, PBRM1, BAP1, SETD2 AND KDM5C mutation status and overall survival was evaluated in 167 patients with metastatic ccRCC who underwent genomic sequencing of their primary tumor."

sparser
"Here, we focus on the impact of individual mutations of PBRM1 , BAP1 , and SETD2 on cancer specific survival in an expanded version of our original cohort (3 additional patients and several more months of followup and events) and further demonstrate the association of BAP1 and SETD2 mutations with worse cancer specific survival (the TCGA cohort n=421), providing a molecular link between gene mutations and cancer specific outcomes in ccRCC."

sparser
"BAP1 and SETD2 mutations are associated with worse cancer specific survival and likely take place during the disease progression phase."

sparser
"Mutations in other genes like SETD2 and BAP1 which have been associated with inferior clinical outcomes in other studies were not found to be statistically significant in our analysis after adjustment."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
HIF1A affects BAP1
4 | 1 15
4 | 1 15

sparser
"We demonstrated that during hypoxia, BAP1 binds, deubiquitylates, and stabilizes HIF-1α, the master regulator of the hypoxia response and tumor cell invasion."

reach
"HIF1alpha-dependent metabolism:. Bioinformatics interrogation of The Cancer Genome Atlas (TCGA) data have associated BAP1 alterations with changes in hypoxia-inducible factor-1 alpha (HIFα) expression (25,76,77)."

sparser
"BAP1 is a key positive regulator of HIF-1α in hypoxia and binds to the N-terminal region of HIF-1α, where HIF-1α recognizes DNA and dimerizes with HIF-1β to form the heterodimeric transactivating complex HIF."

sparser
"We found that BAP1 binds to the N-terminal region of HIF-1α, where HIF-1α binds DNA and dimerizes with HIF-1β forming the heterodimeric transactivating complex HIF."

sparser
"BAP1 Interacts with HIF-1α."

sparser
"Co-immunoprecipitation (CoIP) and proximity ligation assay (PLA) experiments revealed that 1) HIF-1α and BAP1 bind to each other and co-precipitate ( xref ), and 2) the nuclei of BAP1 WT cells contained significantly more PLA positive signals—evidence of BAP1 and HIF-1α interaction—than BAP1 +/− cells ( xref )."

sparser
"We further investigated the specificity of the BAP1 interaction with HIF-1α in HEK-293 cells expressing Myc- BAP1 and HA-HIF-1α, using HA-Tag as bait."

sparser
"We established a computational model of the binding complex of BAP1 and HIF-1α."

sparser
"The NLS domain of BAP1 binds to HIF-1α-r73 and the thermodynamic stability of the binding complex increases through electrostatic interactions with DNA."

sparser
"CoIP experiments in cells co-transfected with full-length Myc-tagged BAP1 (Myc- BAP1 ) and HA-tagged full-length HIF-1α, or HIF-1α fragments covering residues 74 to 826 [HIF-1α(74-826)], 2-400 [HIF-1α(2-400)], 401-826 [HIF-1α(401-826)] ( xref ), confirmed that BAP1 binds to the N terminus region of HIF-1α [HIF-1α(2-400)] ( xref )."
BAP1 affects SETD2
| 1 19
| 1 10

sparser
"Across clear cell- and papillary-associated RCC subtypes in TCGA cohort, worse survival was associated with SETD2 or BAP1 mutation, with their related gene transcriptional signatures involving greater numbers of RCC cases and also being predictive of worse outcome ( xref , xref )."

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"BAP1 and SETD2 mutations (6–12%) are associated with worse CSS, suggesting their roles in disease progression."

sparser
"Furthermore, PBRM1 mutations have been found to confer a slight increase of death risk, while SETD2 or BAP1 mutations were associated with serious death risks in ccRCC patients [ xref – xref ]."

sparser
"Similarly, a study of 609 patients with ccRCC from two distinct cohorts revealed that mutations of 3p21 epigenetic regulators BAP1 and SETD2 were associated with worse cancer-specific survival (CSS), suggesting that they play key roles in disease progression[ xref ]."

sparser
"In pleural and peritoneal mesothelioma, co-occurring alterations occurred on the same chromosome or in close proximity for PBRM1–BAP1 (both 3p21) and SETD2BAP1 (both 3p21), which was observed in the TCGA cohort but not in the recent publication by Zauderer and colleagues [ xref ]."

reach
"The association between VHL, PBRM1, BAP1, SETD2 AND KDM5C mutation status and overall survival was evaluated in 167 patients with metastatic ccRCC who underwent genomic sequencing of their primary tumor."

sparser
"Here, we focus on the impact of individual mutations of PBRM1 , BAP1 , and SETD2 on cancer specific survival in an expanded version of our original cohort (3 additional patients and several more months of followup and events) and further demonstrate the association of BAP1 and SETD2 mutations with worse cancer specific survival (the TCGA cohort n=421), providing a molecular link between gene mutations and cancer specific outcomes in ccRCC."

sparser
"BAP1 and SETD2 mutations are associated with worse cancer specific survival and likely take place during the disease progression phase."

sparser
"Mutations in other genes like SETD2 and BAP1 which have been associated with inferior clinical outcomes in other studies were not found to be statistically significant in our analysis after adjustment."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
BAP1 affects MCRS1
3 | 8 7
BAP1 binds MCRS1.
3 | 2 7
3 | 2 7

sparser
"Since BAP1MCRS1 interaction has not been reported in the literature, we aimed to investigate the potential functional relationship between BAP1 and MCRS1."

sparser
"We next determined the region of BAP1 that binds to MCRS1."

reach
"Since BAP1-MCRS1 interaction has not been reported in the literature, we aimed to investigate the potential functional relationship between BAP1 and MCRS1.To verify that MCRS1 is a bona fide BAP1 inte[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To verify that MCRS1 is a BAP1 interactor, we first examined whether BAP1 can interact with MCRS1 in cells."

sparser
"Next, we investigated whether endogenous BAP1 and MCRS1 can interact with each other."

sparser
"Taken together, these results indicated that BAP1 forms a complex with MCRS1 ."

sparser
"To gain more insight into the BAP1MCRS1 interaction, we determined which region of MCRS1 mediated its interaction with BAP1."

reach
"Taken together, these results indicated that BAP1 forms a complex with MCRS1 in vivo."

sparser
"BAP1 interacted with MCRS1 and DID01 and stabilized these centrosome proteins via deubiquitination, which partially contributed to the suppression of multipolar spindle formation and chromosome aberrations in HK-2 human kidney cells."

No evidence text available
BAP1 activates MCRS1.
| 3
BAP1 activates MCRS1. 3 / 4
| 3

reach
"BAP1 also contributes to chromosome stability by binding the microspherule protein 1 (MCRS1) which plays an essential role in spindle assembly XREF_BIBR."

reach
"To determine whether BAP1 increases MCRS1 by extending its half-life, the protein level of MCRS1 was monitored after treatment with the protein synthesis inhibitor cycloheximide (CHX)."

reach
"In the absence of de novo protein synthesis, the half-life of endogenous MCRS1 protein was much shorter in the BAP1 depleted cells than that in control cells, further suggesting that BAP1 regulates MC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 deubiquitinates MCRS1.
| 3
BAP1 deubiquitinates MCRS1. 3 / 3
| 3

reach
"Moreover, de-ubiquitination of MCRS1 by BAP1 plays important roles in regulating proper mitotic progression and prevents chromosome instability.The yeast two-hybrid screen was performed with full leng[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Since BAP1 is a de-ubiquitinating enzyme, we explored whether BAP1 de-ubiquitinates and stabilizes MCRS1."

reach
"BAP1 can deubiquitinate γ-tubulin [32], as well as microspherule protein 1 (MCRS1), a protein that caps the minus ends of mitotic microtubules to prevent their depolymerization [33, 34]."
BAP1 affects HIF1A
4 | 1 15
4 | 1 15

sparser
"We demonstrated that during hypoxia, BAP1 binds, deubiquitylates, and stabilizes HIF-1α, the master regulator of the hypoxia response and tumor cell invasion."

reach
"HIF1alpha-dependent metabolism:. Bioinformatics interrogation of The Cancer Genome Atlas (TCGA) data have associated BAP1 alterations with changes in hypoxia-inducible factor-1 alpha (HIFα) expression (25,76,77)."

sparser
"BAP1 is a key positive regulator of HIF-1α in hypoxia and binds to the N-terminal region of HIF-1α, where HIF-1α recognizes DNA and dimerizes with HIF-1β to form the heterodimeric transactivating complex HIF."

sparser
"We found that BAP1 binds to the N-terminal region of HIF-1α, where HIF-1α binds DNA and dimerizes with HIF-1β forming the heterodimeric transactivating complex HIF."

sparser
"BAP1 Interacts with HIF-1α."

sparser
"Co-immunoprecipitation (CoIP) and proximity ligation assay (PLA) experiments revealed that 1) HIF-1α and BAP1 bind to each other and co-precipitate ( xref ), and 2) the nuclei of BAP1 WT cells contained significantly more PLA positive signals—evidence of BAP1 and HIF-1α interaction—than BAP1 +/− cells ( xref )."

sparser
"We further investigated the specificity of the BAP1 interaction with HIF-1α in HEK-293 cells expressing Myc- BAP1 and HA-HIF-1α, using HA-Tag as bait."

sparser
"We established a computational model of the binding complex of BAP1 and HIF-1α."

sparser
"The NLS domain of BAP1 binds to HIF-1α-r73 and the thermodynamic stability of the binding complex increases through electrostatic interactions with DNA."

sparser
"CoIP experiments in cells co-transfected with full-length Myc-tagged BAP1 (Myc- BAP1 ) and HA-tagged full-length HIF-1α, or HIF-1α fragments covering residues 74 to 826 [HIF-1α(74-826)], 2-400 [HIF-1α(2-400)], 401-826 [HIF-1α(401-826)] ( xref ), confirmed that BAP1 binds to the N terminus region of HIF-1α [HIF-1α(2-400)] ( xref )."
BAP1 affects Ubiquitin
| 6 12
BAP1 binds Ubiquitin.
| 2 12
| 2 12

sparser
"Thus it seems likely that ASXL1 DEU also uses this ubiquitin interface to stabilize ubiquitin binding of BAP1 and stimulate activity."

sparser
"The cryo-EM map revealed BAP1 bound to the nucleosome in a catalytic conformation, with clearly resolved BAP1-nucleosome, ASXL1-nucleosome, BAP1-Ub, and ASXL1-Ub interfaces ( xref and fig."

sparser
"In this complex, BAP1 is the catalytic subunit and its activity is strongly dependent on the interaction with conserved DEUBiquitinase Adaptor (DEUBAD) domain of ASXLs that induces conformational changes able to increase BAP1 affinity for ubiquitin [ xref ]."

reach
"BAP1 Ub binding mutants, Y33A/I226A/F228A, L8A/F168A/N229A, and L35A/I214A/F228A/L230A were generated using gene synthesis (BioBasic) and then subcloned into a modified pENTR D-Topo plasmid (Life Technologies)."

sparser
"We have undertaken extensive mutational approaches to gain mechanistic insight into BAP1ubiquitin interaction."

sparser
"Our comparative thermodynamic analysis reveals that BAP1ubiquitin interaction is majorly driven by entropy factor which is unique amongst UCHs."

sparser
"Our study sheds light on BAP1 interaction with ubiquitin, which will be useful in understanding its enzymatic function."

sparser
"The cryo-EM map revealed BAP1 bound to the nucleosome in a catalytic conformation, with clearly resolved BAP1-nucleosome, ASXL1-nucleosome, BAP1-Ub, and ASXL1-Ub interfaces ( xref and xref )."

sparser
"However, mechanistic insights into BAP1ubiquitin interaction remain enigmatic."

sparser
"The thermodynamic analysis reveals that BAP1ubiquitin interaction is entropically driven."
BAP1 activates Ubiquitin.
| 4
| 4

reach
"The deubiquitinase (DUB) and tumor suppressor BAP1 catalyzes ubiquitin removal from histone H2A Lys 119 and coordinates cell proliferation, but how BAP1 partners modulate its function remains poorly understood."

reach
"The BAP1 gene produces a 729 AA protein containing several functional domains: a ubiquitin C-terminal hydrolase (UCH) domain, a BRCA1-associated RING domain protein 1 (BARD1) binding domain, an HCF-C1 binding motif (HBM) and a nuclear localization signal ."

reach
"BAP1 and the BRCA1 and BARD1 complex BRCA1 participates in several multiprotein complexes, and plays roles in : repair of double stranded DNA (dsDNA) breaks by homologous recombination; chromatin remo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"As part of the Polycomb repression machinery, BAP1 is activated by the deubiquitinase adaptor domain of ASXL1 mediating gene repression by cleaving ubiquitin (Ub) from histone H2A in nucleosomes."
BAP1 affects PTEN
7 1 | 3 8
BAP1 binds PTEN.
7 | 1 8
7 | 1 8

sparser
"In analyses adjusted only for cohort, mutations in BAP1 (median: 8 vs 5.2 cm; p = 0.001) and PTEN (median: 8 vs 5.9 cm; p = 0.026) were significantly associated with larger median tumor diameter."

sparser
"BAP1 can physically bind to and deubiquitinate PTEN, which inhibits the ubiquitination-mediated degradation of PTEN and thus stabilizes PTEN protein."

sparser
"These findings demonstrate that BAP1 is an important deubiquitinase of PTEN for its stability and the BAP1-PTEN signaling axis plays a crucial role in tumor suppression."

sparser
"Mechanistically, BAP1 interacts with PTEN and stabilizes PTEN through deubiquitination, and further negatively regulates HCC cell EMT by inactivating the AKT/GSK-3β/Snail pathway."

sparser
"In our cohort, BAP1 and PTEN are significantly associated with a higher tumor grade and DNM2 is significantly associated with a lower tumor grade."

sparser
"MiR-200c-3p in M2 macrophage exosomes can influence the PI3K/AKT signaling pathway via obstructing the interaction between BAP1 gene and PTEN gene in hepatocytes, which could strengthen the resistance to sorafenib ( xref )."

sparser
"In PUCH- ccRCC cohort, BAP1 and PTEN are significantly associated with a higher tumor grade (G3&G4, P  = 0.004; P  = 0.030)."

sparser
"Though BAP1 and PTEN is not significantly associated with a higher tumor grade ( P  > 0.05), the percentage of G3&G4 is higher in both BAP1 and PTEN mutation groups."

No evidence text available

No evidence text available
BAP1 deubiquitinates PTEN.
1 | 2
BAP1 deubiquitinates PTEN. 3 / 3
1 | 2

reach
"However, another study suggests that BAP1 may deubiquitinate and stabilize PTEN, leading to Akt inhibition [133]."

"Mechanistically, <span class="match term0">BAP1</span> complexed with <span class="match term1">PTEN</span> and stabilized <span class="match term1">PTEN</span> via deubiquitination and, furthermore, negatively regulated HCC cell EMT by deactivating the AKT/GSK-3beta/Snail pathway"

reach
"BAP1 can physically bind to and deubiquitinate PTEN, which inhibits the ubiquitination mediated degradation of PTEN and thus stabilizes PTEN protein."
VHL affects BAP1
| 3 15
| 3 13

sparser
"Additionally, in vitro binding assays demonstrated that purified glutathione S -transferase (GST)–BAP1, but not GST alone, could directly interact with recombinant His-VHL, confirming a direct interaction between BAP1 and pVHL ( xref )."

sparser
"Given the direct interaction between BAP1 and pVHL, thus, we further investigated whether BAP1 could suppress CRC progression by regulating pVHL stability and function."

sparser
"While a single knockout of Vhl and Pbrm1 failed to develop kidney cancer in mice, both Bap1-Vhl and Bap1-Pbrm1 double-knockout mice developed ccRCC [ xref , xref , xref ]."

sparser
"We also confirmed that the BAP1-pVHL axis is applicable in pancreatic and ovarian cancer (fig."

sparser
"First, BAP1 directly interacts with pVHL in cells and in vitro ( xref )."

sparser
"In keeping with their previous work demonstrating that BAP1 loss results in mTOR activation, the authors found that Vhl-Bap1 –deficient tumors showed strong phospho-S6 (pS6) expression."

sparser
"In addition, we examined the potential functions of the BAP1-pVHL axis in other tumor types."

sparser
"The faithful Vhl-Bap1 and Vhl-Pbrm1 GEM models developed by Gu and colleagues add to the recent salvo of GEM models of ccRCC based upon concurrent deletion of Vhl, p53 , and Rb1 by Frew and colleagues or the inactivation of Vhl and Cdkn2a with concurrent activation of MYC by our group ( xref , xref )."

sparser
"The endogenous interaction between BAP1 and pVHL was further validated through co-immunoprecipitation assays in HCT116 cells ( xref )."

reach
"Mice deficient for either VHL or VHL together with one allele of BAP1 developed multiple lesions spanning from benign cysts to cystic and solid ccRCC XREF_BIBR suggesting a tumor suppressive cooperation between BAP1 and pVHL."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
Ubiquitin affects BAP1
| 5 12
Ubiquitin binds BAP1.
| 2 12
| 2 12

sparser
"Thus it seems likely that ASXL1 DEU also uses this ubiquitin interface to stabilize ubiquitin binding of BAP1 and stimulate activity."

sparser
"The cryo-EM map revealed BAP1 bound to the nucleosome in a catalytic conformation, with clearly resolved BAP1-nucleosome, ASXL1-nucleosome, BAP1-Ub, and ASXL1-Ub interfaces ( xref and fig."

sparser
"In this complex, BAP1 is the catalytic subunit and its activity is strongly dependent on the interaction with conserved DEUBiquitinase Adaptor (DEUBAD) domain of ASXLs that induces conformational changes able to increase BAP1 affinity for ubiquitin [ xref ]."

reach
"BAP1 Ub binding mutants, Y33A/I226A/F228A, L8A/F168A/N229A, and L35A/I214A/F228A/L230A were generated using gene synthesis (BioBasic) and then subcloned into a modified pENTR D-Topo plasmid (Life Technologies)."

sparser
"We have undertaken extensive mutational approaches to gain mechanistic insight into BAP1ubiquitin interaction."

sparser
"Our comparative thermodynamic analysis reveals that BAP1ubiquitin interaction is majorly driven by entropy factor which is unique amongst UCHs."

sparser
"Our study sheds light on BAP1 interaction with ubiquitin, which will be useful in understanding its enzymatic function."

sparser
"The cryo-EM map revealed BAP1 bound to the nucleosome in a catalytic conformation, with clearly resolved BAP1-nucleosome, ASXL1-nucleosome, BAP1-Ub, and ASXL1-Ub interfaces ( xref and xref )."

sparser
"However, mechanistic insights into BAP1ubiquitin interaction remain enigmatic."

sparser
"The thermodynamic analysis reveals that BAP1ubiquitin interaction is entropically driven."
Ubiquitin inhibits BAP1.
| 3
| 3

reach
"CoIP of Flag-HIF-1α showed reduced ubiquitin levels when cells overexpressed BAP1, but not in cells overexpressing the catalytic inactive BAP1(C91S), compared to mock control (Fig. 5A)."

reach
"The fact that an ubiquitin conjugate can inhibit BAP1 to a greater extent than ubiquitin alone, illustrates how this DUB prefers binding its target (ubiquitin conjugate) over its product (ubiquitin alone)."

reach
"ASXL1 WT could induce H2AK119 ubiquitination (Ub) through negatively regulating BAP1 which removes ubiquitination of H2AK119 as a deubiquitinase (Fig. 2b)."
ULD affects BAP1
| 18
| 16

sparser
"Mutations in BAP1-ULD , which occurs in several tumors, abolish the interaction with ASXL2 resulting in a missed stimulation on BAP1 activity."

sparser
"His- or GST-tagged full-length BAP1, BAP1-UCH and BAP1-ULD domains were expressed in bacteria (Bac-) or baculovirus (Bv-), respectively ( xref )."

sparser
"Biophysical and biochemical characterization of BAP1-UCH, BAP1-ULD, ASXL2-AB, and the UCH/ULD/AB complex."

sparser
"Therefore, we initiated biochemical and biophysical analyses of the BAP1-UCH, BAP1-ULD, ASXL2-AB domains and protein complex."

sparser
"The establishment of Ub-AMC assay and SPR for investigating protein-protein interactions would make it possible to validate small molecule hits in future screens for drug compounds that recover loss-of-function in cells with BAP1-ULD mutations/deletions ( xref )."

sparser
"We next tested the affinity and kinetics of BAP1-ULD and ASXL2-AB using SPR."

sparser
"Biochemical analyses of purified recombinant protein complexes from bacteria reveal a direct interaction between the BAP1-ULD domain and ASXL2-AB box."

sparser
"Furthermore, our biochemical work demonstrated that the ASXL2-AB box binds to the BAP1-ULD domain, but not to BAP1-UCH."

sparser
"Using computer molecular modeling of UCHL5 structures, we predicted that the BAP1-ULD domain folds back to the BAP1-UCH catalytic domain and that the ASXL2-AB box stabilizes the UCH catalytic loop via a unique BAP1 mechanism not seen in other UCH proteins, allowing for ubiquitin to fit into the active site ( xref , xref )."

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
BAP1 affects VHL
| 3 15
| 3 13

sparser
"Additionally, in vitro binding assays demonstrated that purified glutathione S -transferase (GST)–BAP1, but not GST alone, could directly interact with recombinant His-VHL, confirming a direct interaction between BAP1 and pVHL ( xref )."

sparser
"Given the direct interaction between BAP1 and pVHL, thus, we further investigated whether BAP1 could suppress CRC progression by regulating pVHL stability and function."

sparser
"While a single knockout of Vhl and Pbrm1 failed to develop kidney cancer in mice, both Bap1-Vhl and Bap1-Pbrm1 double-knockout mice developed ccRCC [ xref , xref , xref ]."

sparser
"We also confirmed that the BAP1-pVHL axis is applicable in pancreatic and ovarian cancer (fig."

sparser
"First, BAP1 directly interacts with pVHL in cells and in vitro ( xref )."

sparser
"In keeping with their previous work demonstrating that BAP1 loss results in mTOR activation, the authors found that Vhl-Bap1 –deficient tumors showed strong phospho-S6 (pS6) expression."

sparser
"In addition, we examined the potential functions of the BAP1-pVHL axis in other tumor types."

sparser
"The faithful Vhl-Bap1 and Vhl-Pbrm1 GEM models developed by Gu and colleagues add to the recent salvo of GEM models of ccRCC based upon concurrent deletion of Vhl, p53 , and Rb1 by Frew and colleagues or the inactivation of Vhl and Cdkn2a with concurrent activation of MYC by our group ( xref , xref )."

sparser
"The endogenous interaction between BAP1 and pVHL was further validated through co-immunoprecipitation assays in HCT116 cells ( xref )."

reach
"Mice deficient for either VHL or VHL together with one allele of BAP1 developed multiple lesions spanning from benign cysts to cystic and solid ccRCC XREF_BIBR suggesting a tumor suppressive cooperation between BAP1 and pVHL."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
BAP1 affects ULD
| 18
| 16

sparser
"Mutations in BAP1-ULD , which occurs in several tumors, abolish the interaction with ASXL2 resulting in a missed stimulation on BAP1 activity."

sparser
"His- or GST-tagged full-length BAP1, BAP1-UCH and BAP1-ULD domains were expressed in bacteria (Bac-) or baculovirus (Bv-), respectively ( xref )."

sparser
"Biophysical and biochemical characterization of BAP1-UCH, BAP1-ULD, ASXL2-AB, and the UCH/ULD/AB complex."

sparser
"Therefore, we initiated biochemical and biophysical analyses of the BAP1-UCH, BAP1-ULD, ASXL2-AB domains and protein complex."

sparser
"The establishment of Ub-AMC assay and SPR for investigating protein-protein interactions would make it possible to validate small molecule hits in future screens for drug compounds that recover loss-of-function in cells with BAP1-ULD mutations/deletions ( xref )."

sparser
"We next tested the affinity and kinetics of BAP1-ULD and ASXL2-AB using SPR."

sparser
"Biochemical analyses of purified recombinant protein complexes from bacteria reveal a direct interaction between the BAP1-ULD domain and ASXL2-AB box."

sparser
"Furthermore, our biochemical work demonstrated that the ASXL2-AB box binds to the BAP1-ULD domain, but not to BAP1-UCH."

sparser
"Using computer molecular modeling of UCHL5 structures, we predicted that the BAP1-ULD domain folds back to the BAP1-UCH catalytic domain and that the ASXL2-AB box stabilizes the UCH catalytic loop via a unique BAP1 mechanism not seen in other UCH proteins, allowing for ubiquitin to fit into the active site ( xref , xref )."

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
SUFU affects BAP1
| 17
| 17

sparser
"Furthermore, transcriptomic analyses and rescue experiments were performed to investigate the downstream regulatory mechanisms of the BAP1-SUFU axis in CRC cell proliferation."

sparser
"Together, these findings highlight the therapeutic relevance of the BAP1-SUFU axis in CRC and suggest that targeting BAP1 may impair tumor growth by destabilizing SUFU."

sparser
"Targeting the BAP1-SUFU regulatory axis as a therapeutic strategy in colorectal cancer."

sparser
"Collectively, these findings demonstrate that the BAP1-SUFU axis drives colorectal cancer progression in vivo and establish this regulatory pathway as a promising therapeutic target, with BAP1 inhibition exerting strong anti-tumor efficacy."

sparser
"Since canonical Hh signaling did not account for the phenotypes induced by SUFU manipulation, we next sought to identify the downstream mechanisms through which the BAP1-SUFU axis drives oncogenic behavior in CRC."

sparser
"Together, these results indicate that BAP1 interacts with SUFU in the cytoplasm and stabilizes it by reducing ubiquitination."

sparser
"To further investigate the role of the BAP1-SUFU axis impact in regulating proliferation-related genes, we conducted transcriptomic analysis following BAP1 knockdown combined with SUFU overexpression."

sparser
"Transcriptomic and mechanistic analyses revealed that disruption of the BAP1-SUFU axis alters cell-cycle transcriptional programs through a Hh-independent mechanism and implicates E2F1 as a downstream regulator, which was confirmed by Western blot and rescue experiments demonstrating its essential role in driving the proliferative phenotype."

sparser
"Having established the cellular and molecular roles of the BAP1-SUFU axis, we next interrogated its tumorigenic potential in vivo."

sparser
"Overall, these findings suggest that the BAP1-SUFU axis regulates CRC transcriptional output through cell-cycle and replication pathways, independent of Hedgehog signaling."
PT33 affects BAP1
| 11 6
PT33 binds BAP1. 10 / 17
| 11 6

reach
"Here, we described one example of such a mechanism in which PT33 covalently bound BAP1 functions as a molecular linker between BRCA1 and its E3 ligase-HERC2."

sparser
"We showed that HERC2 can be recruited by the PT33-bound BAP1 and competes with BARD1 to bind to the RING domain of BRCA1, ultimately leading to BRCA1 estrangement from DSB sites."

reach
"We showed that HERC2 can be recruited by the PT33-bound BAP1 and competes with BARD1 to bind to the RING domain of BRCA1, ultimately leading to BRCA1 estrangement from DSB sites."

sparser
"On the basis of these findings, we next examined whether the covalently interaction of BAP1 with PT33 may alter BAP1 sub-cellular location, and found that after IR treatment, PT33 had no apparent effect on the nuclear translocation of BAP1 in HCT116 cells ( xref H)."

reach
"The deubiquitinating activities were calculated as the fluorescence increment (folds) per minute, indicating the deubiquitination rate of substrate Ub-AMC.2.4 Determine the kinetic parameters of covalent binding of PT33 towards BAP1."

sparser
"These results indicate that PT33 treatment had no apparent impact on the protein level of BRCA1 in presence or absence of BAP1, and suggest that BAP1 inhibition with PT33 (or PT33-bound BAP1) is radically different from BAP1 depletion in HR suppression."

reach
"We observed that PT33 resulted in the increase of the thermal stability of BAP1, thus confirming that PT33 can bind to BAP1."

sparser
"BAP1 binding with PT33 recruits HERC2 to compete with BARD1 for BRCA1 interaction."

reach
"We then performed molecular docking to predict the binding mode and investigate the PT33BAP1 interaction interface."

sparser
"Therefore, we carried out co-IP/WB assays with ectopic expression of Myc- HERC2 (3559–4834 aa 18 ), and found that after IR, PT33-bound BAP1 showed markedly stronger interaction with BRCA1 than uncombined BAP1; only PT33-bound BAP1 could recruit HERC2 to target BRCA1, but not uncombined BAP1."
PARP1 affects BAP1
6 | 8 2
PARP1 binds BAP1.
6 | 2 2
6 | 2 2

No evidence text available

No evidence text available

No evidence text available

reach
"PARylation was not required for the interaction between BAP1 and PARP1 since treatment with PARP inhibitors did not interfere with this interaction ."

sparser
"PARylation was not required for the interaction between BAP1 and PARP1 since treatment with PARP inhibitors did not interfere with this interaction xref ."

No evidence text available

reach
"Poly(ADP-ribose) polymerase 1 (PARP1) interacts with and recruits BAP1 to damage sites, with BAP1 recruitment peaking after the DDB2 and XPC damage sensors."

No evidence text available

No evidence text available

sparser
"PARP1 interacted with BAP1 after UV irradiation, and depletion of PARP1 or treatment with PARP inhibitors reduced BAP1 chromatin binding and BAP1 colocalization with CPDs."
PARP1 activates BAP1.
| 6
PARP1 activates BAP1. 6 / 6
| 6

reach
"PARP1 also promotes BAP1 stability via crosstalk between PARylation and ubiquitination."

reach
"A series of in vitro studies showed that PARP1 stimulated BAP1 activity toward Ub-AMC artificial substrates and that PARylation further enhanced this activity."

reach
"Interestingly, PARP1 stimulated BAP1 activity toward the physiological substrates of H2Aub that had assembled into a nucleosome, but PARylation completely inhibited this activity and was accompanied by strong BAP1 binding to the nucleosomes."

reach
"It was proposed that although PARP1 stimulates BAP1 both intrinsically and via PARylation, probably via an allosteric mechanism, PARylated BAP1 formed an unproductive complex with H2A-ub nucleosomes, rendering H2Aub untargetable ."

reach
"These questions include (1) Does PARP1 stimulate BAP1 within cells, and how is this activity regulated during CPD repair?"

reach
"(4) Does PARP1-mediated BAP1 PARylation exert an effect on DSB repair in which PARP1 plays an important role ."
BAP1 affects cell cycle
| 2 15
BAP1 activates cell cycle.
| 9
| 9

reach
"In functional experiments, BAP1 has been shown to suppress lung cancer tumorigenesis in athymic nude mice, promote the cell cycle and induce both apoptosis and necrosis."

reach
"The E2F1 targets pathway was involved in the BAP1-mediated cell cycle progression of uveal melanoma cells [3]."

reach
"Downregulation of BAP1 promotes proliferation, cell cycle progression, and invasion of ICC in vitro."

reach
"Compared with controls, BAP1 loss was associated with a decreased number of smaller colonies ( p < 0.01) ( Fig. 3 B and D), indicating that BAP1 knockdown indeed inhibits anchorage-independent growth [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 loss causes transient cell cycle inhibition."

reach
"Moreover, RNA mediated depletion of BAP1 promoted cell cycle G1-S progression XREF_BIBR."

reach
"In sharp contrast, RNA interference mediated depletion of BAP1 induced cell cycle progression defects and inhibited cell proliferation in vitro XREF_BIBR XREF_BIBR, suggesting that BAP1 may have dual roles in cancer development."

reach
"A previous study has shown that BAP1 accelerates the G(1)-S checkpoint process by influencing the cell cycle and induces cell death through processes characterized by apoptosis and necrosis (19)."

reach
"In the spleen, Bap1 deletion reduced cell cycle proteins (p = 6.2e-20), consistent with pancytopenia often observed in myelodysplastic disorders (XREF_FIG)."
BAP1 inhibits cell cycle.
| 2 6
| 2 6

eidos
"These results indicated that BAP1 suppressed ICC cell proliferation , cell cycle progression , and invasion via inhibiting ERK1 / 2 and JNK / c-Jun signaling pathways ."

reach
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways."

reach
"Interestingly, there was no substantial difference in cell viability, BrdU incorporation or cell cycle profile between BAP1 deficient and control cells after stable expression of the shRNA constructs for at least 14days, indicating that the initial cell cycle inhibition caused by BAP1 depletion was transient."

reach
"XREF_BIBR reported that BAP1 overexpression in the BAP1-null NCI-H226 human non-small-cell lung cancer cell line suppressed growth in DUB activity- and nuclear localization dependent manners by accelerating the cell cycle G1/S transition and inducing cell death."

eidos
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation , cell cycle progression , and invasion via inhibiting ERK1 / 2 and JNK / c-Jun signaling pathways ."

reach
"For example, while re-expression of BAP1 in BAP1-null H226 cells retarded cell cycle progression leading to an accumulation of cells in the S phase , depletion of BAP1 in some BAP1-proficient cells led to reduced cell proliferation with delayed G1-to-S progression ."

reach
"These results indicated that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways.To further validate these findings, we used inhibitors of the ERK1/2 pathway (U0126) and JNK/c-Jun pathway (SP600125) in RBE-shBAP1 cells."

reach
"Silencing BAP1 could cause retarded cell proliferation in multiple mesothelioma cell lines through arresting S-phase of cell cycle, which was remitted by reintroducing BAP1 [13] ."
BAP1 affects SUFU
| 17
| 17

sparser
"Furthermore, transcriptomic analyses and rescue experiments were performed to investigate the downstream regulatory mechanisms of the BAP1-SUFU axis in CRC cell proliferation."

sparser
"Together, these findings highlight the therapeutic relevance of the BAP1-SUFU axis in CRC and suggest that targeting BAP1 may impair tumor growth by destabilizing SUFU."

sparser
"Targeting the BAP1-SUFU regulatory axis as a therapeutic strategy in colorectal cancer."

sparser
"Collectively, these findings demonstrate that the BAP1-SUFU axis drives colorectal cancer progression in vivo and establish this regulatory pathway as a promising therapeutic target, with BAP1 inhibition exerting strong anti-tumor efficacy."

sparser
"Since canonical Hh signaling did not account for the phenotypes induced by SUFU manipulation, we next sought to identify the downstream mechanisms through which the BAP1-SUFU axis drives oncogenic behavior in CRC."

sparser
"Together, these results indicate that BAP1 interacts with SUFU in the cytoplasm and stabilizes it by reducing ubiquitination."

sparser
"To further investigate the role of the BAP1-SUFU axis impact in regulating proliferation-related genes, we conducted transcriptomic analysis following BAP1 knockdown combined with SUFU overexpression."

sparser
"Transcriptomic and mechanistic analyses revealed that disruption of the BAP1-SUFU axis alters cell-cycle transcriptional programs through a Hh-independent mechanism and implicates E2F1 as a downstream regulator, which was confirmed by Western blot and rescue experiments demonstrating its essential role in driving the proliferative phenotype."

sparser
"Having established the cellular and molecular roles of the BAP1-SUFU axis, we next interrogated its tumorigenic potential in vivo."

sparser
"Overall, these findings suggest that the BAP1-SUFU axis regulates CRC transcriptional output through cell-cycle and replication pathways, independent of Hedgehog signaling."
BAP1 affects PT33
| 11 6
PT33 binds BAP1. 10 / 17
| 11 6

reach
"Here, we described one example of such a mechanism in which PT33 covalently bound BAP1 functions as a molecular linker between BRCA1 and its E3 ligase-HERC2."

sparser
"We showed that HERC2 can be recruited by the PT33-bound BAP1 and competes with BARD1 to bind to the RING domain of BRCA1, ultimately leading to BRCA1 estrangement from DSB sites."

reach
"We showed that HERC2 can be recruited by the PT33-bound BAP1 and competes with BARD1 to bind to the RING domain of BRCA1, ultimately leading to BRCA1 estrangement from DSB sites."

sparser
"On the basis of these findings, we next examined whether the covalently interaction of BAP1 with PT33 may alter BAP1 sub-cellular location, and found that after IR treatment, PT33 had no apparent effect on the nuclear translocation of BAP1 in HCT116 cells ( xref H)."

reach
"The deubiquitinating activities were calculated as the fluorescence increment (folds) per minute, indicating the deubiquitination rate of substrate Ub-AMC.2.4 Determine the kinetic parameters of covalent binding of PT33 towards BAP1."

sparser
"These results indicate that PT33 treatment had no apparent impact on the protein level of BRCA1 in presence or absence of BAP1, and suggest that BAP1 inhibition with PT33 (or PT33-bound BAP1) is radically different from BAP1 depletion in HR suppression."

reach
"We observed that PT33 resulted in the increase of the thermal stability of BAP1, thus confirming that PT33 can bind to BAP1."

sparser
"BAP1 binding with PT33 recruits HERC2 to compete with BARD1 for BRCA1 interaction."

reach
"We then performed molecular docking to predict the binding mode and investigate the PT33BAP1 interaction interface."

sparser
"Therefore, we carried out co-IP/WB assays with ectopic expression of Myc- HERC2 (3559–4834 aa 18 ), and found that after IR, PT33-bound BAP1 showed markedly stronger interaction with BRCA1 than uncombined BAP1; only PT33-bound BAP1 could recruit HERC2 to target BRCA1, but not uncombined BAP1."
BAP1 affects MYCN
2 | 12 3
BAP1 increases the amount of MYCN.
| 7
BAP1 increases the amount of MYCN. 7 / 7
| 7

reach
"Inhibiting BAP1 by shRNA-mediated knockdown decreases the expression of MYCN largely through reducing the BAP1-mediated deubiquitination of MYCN."

reach
"Moreover, BAP1, but not OTUD4, upregulated the MYCN protein level in a dose-dependent manner (Supplementary Fig. 1A)."

reach
"However, further experiment indicated that only BAP1 and USP7 could elevate the MYCN protein level in a dose-dependent manner, but BAP1 have the best effect on the elevation of MYCN protein levels (Fig. 1F)."

reach
"Therefore, given the better effect of BAP1 in elevating MYCN protein levels in our experimental settings, the BAP1 will be our major focus in the remainder of the study.Next, we performed the protein half-life assay and ubiquitination assay to further confirm whether BAP1 is a bona fide DUB for MYCN to plays a vital role in positively regulating MYCN protein abundance."

reach
"The results showed that BAP1-WT, but not BAP1-C91S, dramatically elevated the MYCN protein levels (Fig. 1J)."

reach
"The results indicated that FBXW7 could efficiently promote the degradation of MYCN, but BAP1 could antagonize the degradation effect of FBXW7 and markedly rescued the protein levels of MYCN in a dose-dependent manner (Supplementary Fig. 1C, D)."

reach
"Hence, our results suggest that the decreased protein levels of MYCN caused by depletion of BAP1 might contribute the resistance to BET inhibitors and Aurora Kinase A inhibitors in NB cells."
BAP1 binds MYCN.
2 | 1 3
2 | 1 3

reach
"Here, we demonstrate that BAP1 could bind and deubiquitinate MYCN to increase the stability of MYCN."

sparser
"In support of the notion that BAP1 might be an upstream DUB for MYCN, Flag-BAP1 and HA-MYCN were co-transfected into 293T cells and the immunoprecipitation (IP) assay was performed to examine whether BAP1 will interact with MYCN."

No evidence text available

sparser
"Furthermore, endogenous BAP1 could specifically interacts with MYCN in 293T cells (Fig. xref )."

sparser
"Here, we demonstrate that BAP1 could bind and deubiquitinate MYCN to increase the stability of MYCN."

No evidence text available
BAP1 deubiquitinates MYCN.
| 4
BAP1 deubiquitinates MYCN. 4 / 4
| 4

reach
"Taken together, these data suggest that the BAP1 is likely an upstream DUB of MYCN to deubiquitinate and stabilize MYCN protein."

reach
"E3 ubiquitin ligases such as FBXW7 could promote MYCN ubiquitination and degradation, however, BAP1 deubiquitinates MYCN and increases its stability."

reach
"Inhibiting BAP1 by shRNA-mediated knockdown decreases the expression of MYCN largely through reducing the BAP1-mediated deubiquitination of MYCN."

reach
"Notably, BAP1 could reduce the polyubiquitination of MYCN (Fig. 1I), which might be the underlying mechanism for the effect of BAP1 on extend half-life of MYCN."
| 1 14
BAP1 inhibits Cell Survival.
| 1 7
| 1 7

reach
"Furthermore, our results reveled that BaP1 reduced cell survival and cell motility, two desirable properties for an anticancer drug.Interestingly, when we treated RKO cells with higher doses of BaP1, 2 × IC and 4 × IC , concentrations that still have a reduced inhibitory effect in the noncancerous colon cell line NCM460, we observed well-established and correlated phenotypes."

reach
"Crucially, concomitant BAP1 and HDAC2 depletion did not additively reduce cell viability in the presence of vorinostat."

eidos
"We found that BaP1 is selective for CRC cells and reduces cell proliferation , cell survival , and cell migration ."

reach
"Overexpression of BAP1 significantly reduced cell viability (Figure 9A) and promoted cell apoptosis (Figure 9B), while CsA reversed these results."

reach
"BAP1 suppresses cell viability and growth."

reach
"BaP1 Inhibits Proliferation, Reduces Cell Survival and Motility, and Leads to ROS Generation in RKO Cells."

reach
"In summary, it seems that loss of BAP1 foster genomic instability in tumor pathogenesis, however, the activity of BAP1 promotes tumor cell survival and contributes to therapeutic resistance during irradiation."

reach
"These results indicate that the reduction in colon cancer cell viability induced by BAP1 depletion was attributed mainly to increased apoptosis."
BAP1 activates Cell Survival.
| 7
| 7

reach
"BAP1 stabilizes BRCA1 protein and restores cell survival rates of MM cells with BAP1 deletion after IR."

reach
"BAP1 promotes stalled fork restart and cell survival via INO80 in response to replication stress."

reach
"Further, downregulation of BAP1 in SK-N-FI cells by using small interfering RNA (siRNA) enhanced cell survival compared to siRNA treated control cells."

reach
"BAP1 depletion greatly reduced cell viability, as observed previously (Fig. 7A)."

reach
"Our results suggested that, whereas WT BAP1 suppresses MM cell proliferation and restores cell survival after IR damage, some mutant BAP1 may also moderately retain these functions."

reach
"To confirm the results of the colony formation assay and determine the mechanisms by which BAP1 activity supports colon cancer cell viability, we analyzed apoptosis by Annexin-V/PI double staining."

reach
"While BAP1 deletion suppresses RCC cell survival in vitro, it does not impede tumor growth in immunocompetent murine models."
SP1 affects BAP1
2 | 4 10
2 | 4 10

sparser
"Therefore, ODN enhanced oxaliplatin-induced apoptosis by interacting with BAP1 and Sp1 to induce Sp1 stabilization-mediated Bax upregulation ( xref )."

sparser
"ODN induces BAP1 interaction with Sp1."

sparser
"We postulated that ODN may influence the interaction between BAP1 and Sp1."

sparser
"Interestingly, ODN and cathepsin K siRNA treatment markedly increased Sp1 and BAP1 interaction ( xref A)."

sparser
"Sp1-BAP1 interaction was increased by ODN treatment ( xref F)."

sparser
"ODN induced the phosphorylation of BAP1 at Ser 592 and enhanced the interaction of Sp1 and BAP1, resulting in Sp1 deubiquitination and destabilization."

reach
"We postulated that ODN may influence the interaction between BAP1 and Sp1."

sparser
"Mechanistically, ODN induced BAP1 phosphorylation and enhanced Sp1-BAP1 interaction, resulting in Sp1 ubiquitination and degradation."

sparser
"To investigate the functional role of p53, we tested the interaction between Sp1 and BAP1 after p53 immunoprecipitation in ODN- and cathepsin K siRNA-treated cells and identified more p53, Sp1, and BAP1 protein complexes in the ODN- and cathepsin K siRNA-treated cells ( xref B)."

sparser
"These data demonstrated that the C-terminal domain of p53 can effectively enhance Sp1 stability by increasing the interaction with BAP1 and Sp1."
PTEN affects BAP1
7 | 1 8
7 | 1 8

sparser
"In analyses adjusted only for cohort, mutations in BAP1 (median: 8 vs 5.2 cm; p = 0.001) and PTEN (median: 8 vs 5.9 cm; p = 0.026) were significantly associated with larger median tumor diameter."

sparser
"BAP1 can physically bind to and deubiquitinate PTEN, which inhibits the ubiquitination-mediated degradation of PTEN and thus stabilizes PTEN protein."

sparser
"These findings demonstrate that BAP1 is an important deubiquitinase of PTEN for its stability and the BAP1-PTEN signaling axis plays a crucial role in tumor suppression."

sparser
"Mechanistically, BAP1 interacts with PTEN and stabilizes PTEN through deubiquitination, and further negatively regulates HCC cell EMT by inactivating the AKT/GSK-3β/Snail pathway."

sparser
"In our cohort, BAP1 and PTEN are significantly associated with a higher tumor grade and DNM2 is significantly associated with a lower tumor grade."

sparser
"MiR-200c-3p in M2 macrophage exosomes can influence the PI3K/AKT signaling pathway via obstructing the interaction between BAP1 gene and PTEN gene in hepatocytes, which could strengthen the resistance to sorafenib ( xref )."

sparser
"In PUCH- ccRCC cohort, BAP1 and PTEN are significantly associated with a higher tumor grade (G3&G4, P  = 0.004; P  = 0.030)."

sparser
"Though BAP1 and PTEN is not significantly associated with a higher tumor grade ( P  > 0.05), the percentage of G3&G4 is higher in both BAP1 and PTEN mutation groups."

No evidence text available

No evidence text available
LDHA affects BAP1
1 | 6 9
1 | 6 9

No evidence text available

sparser
"By elucidating the interaction between BAP1 and LDHA and the subsequent effects on lactate production in melanoma cells, this work provides insights into the mechanism of BAP1-mediated metabolic regulation."

sparser
"We then reasoned that the binding of BAP1 to LDHA was critical to mediate the translocation of LDHA to the nucleus since BAP1 but not LDHA harbored the NLS sequence."

reach
"By elucidating the interaction between BAP1 and LDHA and the subsequent effects on lactate production in melanoma cells, this work provides insights into the mechanism of BAP1-mediated metabolic regulation."

reach
"We also performed immunoprecipitation experiments in cell lines supplemented with wild-type and enzymatically inactive mutant BAP1, and interactions between LDHA and BAP1 were detected in each group (Fig. 2F)."

sparser
"Notably, these BAP1 mutants cannot rescue the phenotype of lactate production (Fig. xref ), suggesting that full-length BAP1 was required to maintain the interaction between BAP1 and LDHA."

sparser
"Therefore, enzyme active site mutants of BAP1 (C91G, G185R) that did not affect BAP1-LDHA interaction and BAP1 nuclear translocation were still capable of regulating the subcellular localization of LDHA."

reach
"To investigate whether BAP1 directly inhibits the enzymatic activity of LDHA by binding to LDHA, we generated full-length BAP1 and LDHA expression constructs, then expressed them in Escherichia coli system to obtain purified proteins by affinity chromatography (Fig. 3A)."

sparser
"In this study, we have found that BAP1 interacts with LDHA."

sparser
"We further performed immunoprecipitation experiments on several candidates and found that LDHA can interact with BAP1 (Fig. xref , Supplementary Fig. xref )."
BAP1 affects ASXL1-MT
| 16
BAP1 binds ASXL1-MT.
| 13
ASXL1-MT binds BAP1. 10 / 13
| 13

reach
"This hyperactive ASXL1-MT and BAP1 complex promotes aberrant myeloid differentiation of haematopoietic progenitor cells and accelerates RUNX1-ETO-driven leukaemogenesis."

reach
"In immunoprecipitation assays, BAP1 interacted with ASXL1-MT and induced its monoubiquitination."

reach
"Together, these results indicate that the ASXL1-MT and BAP1 complex impairs terminal differentiation of HSPCs, except for differentiation toward monocytes."

reach
"Cells expressing ASXL1-MT and BAP1 did not produce colonies beyond the fourth passage, suggesting that the ASXL1-MT and BAP1 complex itself is not sufficient to transform murine HSPCs."

reach
"To examine the cooperation between ASXL1-MT and BAP1 in human RUNX1-ETO cells, we transduced ASXL1-MT or ASXL1-MT-K351R (coexpressing GFP) together with BAP1 or BAP1-C91S (coexpressing NGFR), and monitored changes of the frequency of the GFP + NGFR + double positive fraction in culture."

reach
"Thus, Irf8 is also a downstream target of the ASXL1-MT and BAP1 complex, which is likely to underlie the enhanced monocytic differentiation of HSPCs coexpressing ASXL1-MT and BAP1."

reach
"These data indicate that the ASXL1-MT and BAP1 complex induces upregulation of posterior HOXA genes and IRF8 via removal of H2AK119ub."

reach
"In addition to these reports implicating BAP1 in myeloid transformation, the present results identify the underlying molecular mechanisms by which ASXL1-MT and BAP1 complex induce myeloid transformation; BAP1 plays a necessary role in maintaining aberrant posterior HOXA expression in ASXL1-mutant leukemia and in sustaining their leukemic proliferation."

reach
"The ASXL1-MT and BAP1 complex enhances transient myeloid proliferation, but does not increase the long-term proliferation potential of murine and human HSPCs."

reach
"The ASXL1-MT and BAP1 complex is not a simple growth-accelerator, but rather inhibits terminal differentiation of HSPCs toward granulocytes, macrophages, mast cells, and erythrocytes."
BAP1 ubiquitinates ASXL1-MT.
| 3
BAP1 ubiquitinates ASXL1-MT. 3 / 3
| 3

reach
"Thus, BAP1 enhances monoubiquitination of ASXL1-MT probably through the recruitment of ubiquitin ligases and the concurrent removal of the polyubiquitin chain from it."

reach
"BAP1 expression results in increased monoubiquitination of ASXL1-MT, which in turn increases the catalytic function of BAP1."

reach
"We have shown here that BAP1 recruits ubiquitin ligases to induce both mono- and polyubiquitination of ASXL1-MT."
MBD5 affects BAP1
5 | 1 9
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
5 | 1 1

No evidence text available
| DOI

No evidence text available

reach
"The relative proportion of MBD5 that associates with BAP1 (PR-DUB) is higher in differentiated NPCs relative to mESCs [27]."

sparser
"The relative proportion of MBD5 that associates with BAP1 (PR-DUB) is higher in differentiated NPCs relative to mESCs [ xref ]."

No evidence text available

No evidence text available

No evidence text available
HSF1 affects BAP1
| 5 10
| 5 10

sparser
"BAP1-HSF1 interaction inhibits the deacetylation of HSF1-K80 mediated by SIRT1, thereby promoting HSP70 mediated monomerization of HSF1 and its dissociation from target gene loci."

reach
"This may suggest that the binding between BAP1 and HSF1 could inhibit the ubiquitination of BAP1 mediated by UBE2O [24], or there may be other unexplored mechanisms at play."

sparser
"Co-immunoprecipitation assays indicated that treatment with the SIRT1 inhibitor, EX-527, notably augmented the level of acetylated lysine of HFS1, as well as the interaction between HSF1 and BAP1 (Fig.  xref A)."

sparser
"The BAP1-HSF1 interaction preserves the acetylation of HSF1-K80 and promotes HSF1-HSP70 interaction, facilitating HSF1 oligomerization and detachment from the chromatin."

sparser
"The BAP1-HSF1 interaction safeguards the acetylation of HSF1 at the K80 site, promotes the monomerization of HSF1, and facilitates its dissociation from DNA."

reach
"More importantly, both wild-type (WT) and C91A mutated BAP1 attenuated the HSF1-SIRT1 interaction (Fig. 5C, Supplementary Fig. S3G), and SIRT1 knockdown resulted in an enhanced binding between HSF1 and BAP1 (Fig. 5D, Supplementary Fig. S3H), indicating that BAP1 and SIRT1 might compete with each other for binding to HSF1."

sparser
"BAP1 interacts with the N-terminal of HSF1."

sparser
"The subsequent Co-IP assay revealed that the N-terminal of HSF1 was pivotal for the formation of the BAP1-HSF1 complex (Fig.  xref F)."

reach
"Co-immunoprecipitation assays indicated that treatment with the SIRT1 inhibitor, EX-527, notably augmented the level of acetylated lysine of HFS1, as well as the interaction between HSF1 and BAP1 (Fig. 6A)."

sparser
"Interestingly, the interaction between BAP1 and HSF1 exclusively occurs in the nucleus (Fig.  xref B)."
| 12

reach
"Phosphorylation of BAP1 that promotes HR resulted in reduced chromatin binding of BAP1 ."

reach
"BAP1 deubiquitylates Ub-H2AK119 at sites of DNA damage, and the loss of BAP1 has been shown to impair homologous recombination (HR) repair (10)."

reach
"However, another study demonstrated that the E3 activity of BRCA1–BARD1 could be inhibited by BAP1, which mediates HR repair and confer radioresistance17."

reach
"This suggests that deubiquitination of H2A by BAP1 promotes HR which is complementary to the finding that ubiquitination of K119 in response to DNA damage favors NHEJ [95]."

reach
"BAP1 loss impairs HR and double-strand break repair, promoting error-prone nonhomologous end joining, with consequent genomic instability (XREF_FIG)."

reach
"BAP1 deficiency inhibits efficient HR repair and increases radiation sensitivity, suggesting that the deubiquitination of H2AK119ub promotes HR [78]."

reach
"The function of UCHL1 in DSB repair is still unclear, but it has been established that UCHL5 promotes repair by enhancing DNA-end resection, whereas BAP1 promotes HR via an unknown mechanism ."

reach
"Then, by using a GFP-based reporter assay, the authors showed that BAP1 promoted HR in a BRCA1-related pathway without exerting a significant impact on NHEJ."

reach
"Notably, the oncometabolite 2-hydroxyglutarate produced by mutant IDH1/2 enzymes, and loss of function of ARID1A, BAP1, or BRCA2 can impair the HR pathway, rendering cancer cells sensitive to PARPi treatment (18–21, 47, 48)."

reach
"In the nucleus, BAP1 modulates the transcriptional regulation of several gene programs and promotes DNA repair by facilitating homologous recombination ."

reach
"BAP1 deubiquitylates Ub-H2AK119 at sites of DNA damage, and the loss of BAP1 has been shown to impair homologous recombination (HR) repair (10)."

reach
"We reasoned that if BAP1 interacts with BARD1 to modulate homologous recombination and possibly other cellular pathways, germline BARD1 mutations might also predispose to mesothelioma, and that these patients might have a better prognosis, similar to carriers of germline BAP1 mutations."

reach
"As BAP1 loss causes deficient HR, BAP1-mutant tumors were reported to be particularly addicted to alternative DNA repair pathways, e.g., PARP1 mediated ones."

reach
"BAP1 is rapidly recruited to DSB sites, and BAP1 depletion leads to reduced BRCA1, RAD51 and RPA foci and has been suggested to decrease HR."

reach
"BAP1 loss specifically reduces HR but not NHEJ [68,69]."
BAP1 affects calcium(2+)
| 15
BAP1 increases the amount of calcium(2+).
| 10
BAP1 increases the amount of calcium(2+). 10 / 10
| 10

reach
"In this study, we observed that BAP1 increased cardiomyocytes’ Ca level, causing mPTP opening and mitochondrial dysfunction through interactions with IP3R, and reversed the function of Exos inhibited H/R-induced cardiomyocytes ferroptosis."

reach
"Although the exact mechanism underpinning the metabolic alterations associated with BAP1 loss are currently unknown, a study from 2017 reported that BAP1 promotes calcium (Ca ) release from the ER to the mitochondria ."

reach
"For example, BAP1 promotes apoptosis by deubiquitinating and stabilizing IP3R3 at the endoplasmic reticulum (ER), which stimulates Ca release from the ER into the cytosol and thereby increases the mitochondrial Ca concentration and cytochrome C release ."

reach
"Mechanistically, BAP1 interacted with IP3R and increased cardiomyocytes' Ca 2+ level, causing mPTP opening and mitochondrial dysfunction, promoting H/R-induced cardiomyocytes' ferroptosis."

reach
"BAP1 can also modulate the ubiquitination and stability of IP3R3, regulating Ca release from endoplasmic reticulum and cell viability (38)."

reach
"A recent factor that deserves attention is the tumour suppressor BAP1, which has been shown to increase the amount of Ca 2+ released from the ER into the cytosol and mitochondria, promoting apoptosis XREF_BIBR and increasing aerobic glycolysis XREF_BIBR."

reach
"Additionally, BAP1 promotes tumor cell apoptosis by regulating ER-mitochondrial calcium release (19)."

reach
"In another report, BAP1 was found to modulate endoplasmic reticulum (ER)-mitochondria calcium release (4)."

reach
"These results indicated that BAP1 increases cardiomyocytes’ Ca level, inducing mPTP opening and mitochondrial dysfunction through interactions with IP3R."

reach
"BAP1 stabilizes type 3 inositol-1,4,5-triphosphate receptor, promoting Ca release from the endoplasmic reticulum (ER) [42]."
BAP1 activates calcium(2+).
| 5
| 5

reach
"In a paper in press in Nature, 19 we report that BAP1 modulates ER-to-mitochondria calcium (Ca 2+) release."

reach
"(ii) The tumor suppressor BRCA1 associated protein 1 (BAP1), a deubiquitylating enzyme, promotes ER-mitochondrial Ca 2+ transfer by stabilizing the IP 3 R3 114."

reach
"In a paper in press in Nature, 19 we report that BAP1 modulates ER-to-mitochondria calcium (Ca 2+) release."

reach
"Abnormalities in the function of BAP1 can induce inappropriate Ca influx into mitochondria, which may affect the regulation of apoptosis and lead to carcinogenesis ."

reach
"Loss of BAP1 or its DUB activity down-regulates IP 3 R3 and calcium flux, thereby preventing apoptosis of cells with DNA damages."
BAP1 affects TP53
2 | 4 9
BAP1 binds TP53.
2 | 9
2 | 7

sparser
"Mutual exclusivity of genomic alterations was also shown for TP53:IDH1 , IDH1: Kirsten rat sarcoma virus ( KRAS ), TP53:BAP1 , and IDH1:FGFR2 ."

sparser
"They also reported a tendency toward mutual exclusivity between multiple genes including TP53:IDH1, IDH1:KRAS, TP53:BAP1, and IDH1:FGFR2."

sparser
"Concurrent mutations in BAP1 and TP53 were associated with adverse outcome, with similar effects observed for acquired alterations in either of the two genes alone ( xref )."

sparser
"Mutations in BAP1 ( q = 0.004) and TP53 ( q = 0.001) were associated with decreased CSS in a multivariable model; only TP53 ( q = 0.005) remained significant when SSIGN score was included."

sparser
"Only three of the total 39 interactions— NF2-TP53 , NF2-SMAD4 , and BAP1-TP53 —commonly occurred in all three networks; 12 interactions were shared by only two of the networks and 24 were unique to specific networks ( xref )."

sparser
"On analyses adjusted only for cohort, we found that mutations in BAP1 ( n = 1049 for analysis; hazard ratio [HR]: 2.10; 95% confidence interval [CI]: 1.44–3.04; q < 0.001) and TP53 ( n = 784 for analysis; HR: 2.63; 95% CI: 1.36–5.08; q = 0.028) were significantly associated with increased risk of death from cancer after adjustment for multiple comparisons using competing risks regression ( xref )."

No evidence text available

sparser
"Only BAP1 and TP53 were significantly associated with an inferior OS ( P  = 0.0002; P  = 0.0082)."

No evidence text available
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
BAP1 activates TP53.
| 4
BAP1 activates TP53. 4 / 4
| 4

reach
"For example, p53 and BRCA1-related protein 1 (BAP1) induce ferroptosis in tumor cells through multiple signaling pathways, which act as a natural barrier to cancer development (5, 6)."

reach
"The upregulation of BAP1 expression can additionally augment the activity and stability of P53, thereby facilitating neuronal apoptosis and exacerbating neurological dysfunction after SAH ."

reach
"However, restoring BAP1 in p53 deficient cells still inhibited SLC7A11 expression (XREF_SUPPLEMENTARY - XREF_SUPPLEMENTARY), and the fold change in SLC7A11 expression by BAP1 restoration in p53 deficient cells was similar to that in p53-proficient cells (XREF_SUPPLEMENTARY), suggesting that BAP1 represses SLC7A11 expression independent of p53."

reach
"In addition, BAP1 overexpression enhanced P53 activity and stability by reducing P53 proteasome-mediated degradation."
BAP1 affects STK11
5 1 | 6 3
BAP1 binds STK11.
5 | 2 3
5 | 2 3

No evidence text available

reach
"BAP1 deficiency enhanced cancer cell proliferation and lipid synthesis via the LKB1-AMPK-mTOR signalling pathway.In conclusion, we suggest that BAP1 interacts with and deubiquitinates LKB1, thereby af[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The positive colony could survive in SD-4 (-Ade, -His, -Leu, -Trp) medium (Clontech) developing a blue colour in the presence of X-Gal (Sigma), thereby indicating an interaction between BAP1 and LKB1."

sparser
"These results validated a direct interaction between BAP1 and LKB1."

sparser
"We demonstrated a direct interaction between LKB1 and BAP1 ( Fig. 1 E)."

No evidence text available

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reach
"These results validated a direct interaction between BAP1 and LKB1."
BAP1 deubiquitinates STK11.
1 | 2
BAP1 deubiquitinates STK11. 3 / 3
1 | 2

reach
"BAP1 deficiency enhanced cancer cell proliferation and lipid synthesis via the LKB1-AMPK-mTOR signalling pathway.In conclusion, we suggest that BAP1 interacts with and deubiquitinates LKB1, thereby af[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Altogether, this work has revealed a previously unknown mechanism for how BAP1 suppresses tumour through deubiquitinating LKB1."

"<span class="match term0">BAP1</span> regulates AMPK-mTOR signalling pathway through deubiquitinating and stabilizing tumour-suppressor <span class="match term1">LKB1</span>"
BAP1 inhibits STK11.
| 2
BAP1 inhibits STK11. 2 / 2
| 2

reach
"49 Moreover, BAP1 induces apoptosis by binding to the 14‐3‐3 protein or suppresses apoptosis by stabilizing liver kinase B1 (LKB1)."

reach
"Ectopic expression of wild-type BAP1, instead of the BAP1-C91S mutant, significantly prolonged the half-life of LKB1 protein in HEK 293T cells, suggesting that BAP1 inhibits the degradation of LKB1 an[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects MBD5
5 | 1 9
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
5 | 1 1

No evidence text available
| DOI

No evidence text available

reach
"The relative proportion of MBD5 that associates with BAP1 (PR-DUB) is higher in differentiated NPCs relative to mESCs [27]."

sparser
"The relative proportion of MBD5 that associates with BAP1 (PR-DUB) is higher in differentiated NPCs relative to mESCs [ xref ]."

No evidence text available

No evidence text available

No evidence text available
ASXL1-MT affects BAP1
| 15
ASXL1-MT binds BAP1.
| 13
ASXL1-MT binds BAP1. 10 / 13
| 13

reach
"This hyperactive ASXL1-MT and BAP1 complex promotes aberrant myeloid differentiation of haematopoietic progenitor cells and accelerates RUNX1-ETO-driven leukaemogenesis."

reach
"In immunoprecipitation assays, BAP1 interacted with ASXL1-MT and induced its monoubiquitination."

reach
"Together, these results indicate that the ASXL1-MT and BAP1 complex impairs terminal differentiation of HSPCs, except for differentiation toward monocytes."

reach
"Cells expressing ASXL1-MT and BAP1 did not produce colonies beyond the fourth passage, suggesting that the ASXL1-MT and BAP1 complex itself is not sufficient to transform murine HSPCs."

reach
"To examine the cooperation between ASXL1-MT and BAP1 in human RUNX1-ETO cells, we transduced ASXL1-MT or ASXL1-MT-K351R (coexpressing GFP) together with BAP1 or BAP1-C91S (coexpressing NGFR), and monitored changes of the frequency of the GFP + NGFR + double positive fraction in culture."

reach
"Thus, Irf8 is also a downstream target of the ASXL1-MT and BAP1 complex, which is likely to underlie the enhanced monocytic differentiation of HSPCs coexpressing ASXL1-MT and BAP1."

reach
"These data indicate that the ASXL1-MT and BAP1 complex induces upregulation of posterior HOXA genes and IRF8 via removal of H2AK119ub."

reach
"In addition to these reports implicating BAP1 in myeloid transformation, the present results identify the underlying molecular mechanisms by which ASXL1-MT and BAP1 complex induce myeloid transformation; BAP1 plays a necessary role in maintaining aberrant posterior HOXA expression in ASXL1-mutant leukemia and in sustaining their leukemic proliferation."

reach
"The ASXL1-MT and BAP1 complex enhances transient myeloid proliferation, but does not increase the long-term proliferation potential of murine and human HSPCs."

reach
"The ASXL1-MT and BAP1 complex is not a simple growth-accelerator, but rather inhibits terminal differentiation of HSPCs toward granulocytes, macrophages, mast cells, and erythrocytes."
ASXL1-MT activates BAP1.
| 2
ASXL1-MT activates BAP1. 2 / 2
| 2

reach
"Overexpression of ASXL1-MT promoted nuclear import of BAP1, which was observed only weakly when we used the ubiquitination deficient ASXL1-MT-K351R."

reach
"Together, these data suggest that ASXL1-MT, especially the monoubiquitinated form of ASXL1-MT, increases catalytic function of BAP1."
BAP1 affects UBE2O
12 | 1 1
12 | 1 1

reach
"For example, UBE2O binds to and multi-monoubiquitinates BAP1 to induce its cytoplasmic localization and promotes adipocyte differentiation [XREF_BIBR]."

No evidence text available

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sparser
"Since BAP1 5K/R mutant still interact with UBE2O and is ubiquitinated on other sites, we also replaced the NLS region of BAP1, containing the UBE2O-interacting motif, with T-large antigen NLS which pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

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ATM affects BAP1
| 7 7
ATM phosphorylates BAP1.
| 6 6
ATM phosphorylates BAP1. 10 / 12
| 6 6

reach
"The nuclear localized deubiquitylating enzyme BAP1 XREF_BIBR, XREF_BIBR, which is responsible for histone modification in transcription regulation XREF_BIBR, XREF_BIBR, can be phosphorylated by Ataxia telangiectasia mutated (ATM) and participates in DNA double-strand break (DSB) repair through regulating histone 2A and H2AX ubiquitylation XREF_BIBR, XREF_BIBR."

reach
"The DUB BAP1 is also recruited to DNA damage sites and is phosphorylated by ATM (Ismail et al. 2014; Yu et al. 2014)."

sparser
"BAP1 is phosphorylated by ATM, which promotes its recruitment to the DSB site."

sparser
"It was further demonstrated that both of the catalytic activity and the phosphorylation of BAP1 by ATM is required for efficient assembly of the homologous recombination (HR) factors BRCA1 and RAD51 at the DSB loci (Fig. xref ) xref ."

sparser
"The role of BAP1 phosphorylation by ATM and possibly by ATR under replication stress conditions is still unknown and needs to be addressed in future studies."

reach
"The role of BAP1 phosphorylation by ATM and possibly by ATR under replication stress conditions is still unknown and needs to be addressed in future studies."

reach
"Intriguingly, BAP1 is directly phosphorylated by ATM upon DNA damage and is required for efficient assembly of homologous recombination (HR) repair factors BRCA1 and RAD51 at IR-induced foci to promote repair of DNA double-strand breaks (DSBs) ."

reach
"BAP1 is phosphorylated by ATM kinase, and promotes repair of DSBs through PARP dependent recruitment of the polycomb DUB complex to DNA damage sites [XREF_BIBR]."

sparser
"BAP1 is phosphorylated by ATM kinase, and promotes repair of DSBs through PARP-dependent recruitment of the polycomb DUB complex to DNA damage sites [ xref ]."

reach
"BAP1 was shown to mediate poly (ADP-ribose)-dependent recruitment of the Polycomb-repressive deubiquitinase complex (PR-DUB) to sites of DNA damage, was phosphorylated by ATM, and promoted repair of DNA double-strand breaks ."
ATM binds BAP1.
| 1 1
| 1 1

reach
"8 BAP1 also interacts with ataxia telangiectasia mutated and ataxia telangiectasia and Rad3 related proteins, which regulate the DNA damage induced by UV radiation."

sparser
"Further large germline studies of biliary tract cancers are warranted to better assess the association of ATM and BAP1 with this rare cancer type."
MCRS1 affects BAP1
3 | 2 7
3 | 2 7

sparser
"Since BAP1MCRS1 interaction has not been reported in the literature, we aimed to investigate the potential functional relationship between BAP1 and MCRS1."

sparser
"We next determined the region of BAP1 that binds to MCRS1."

reach
"Since BAP1-MCRS1 interaction has not been reported in the literature, we aimed to investigate the potential functional relationship between BAP1 and MCRS1.To verify that MCRS1 is a bona fide BAP1 inte[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To verify that MCRS1 is a BAP1 interactor, we first examined whether BAP1 can interact with MCRS1 in cells."

sparser
"Next, we investigated whether endogenous BAP1 and MCRS1 can interact with each other."

sparser
"Taken together, these results indicated that BAP1 forms a complex with MCRS1 ."

sparser
"To gain more insight into the BAP1MCRS1 interaction, we determined which region of MCRS1 mediated its interaction with BAP1."

reach
"Taken together, these results indicated that BAP1 forms a complex with MCRS1 in vivo."

sparser
"BAP1 interacted with MCRS1 and DID01 and stabilized these centrosome proteins via deubiquitination, which partially contributed to the suppression of multipolar spindle formation and chromosome aberrations in HK-2 human kidney cells."

No evidence text available
ITPR3 affects BAP1
4 | 4 5
4 | 4 5

sparser
"Cytoplasmic BAP1 binds to IP3R3 and promotes calcium ion release from the endoplasmic reticulum into the cytosol via IP3R3 deubiquitination, thereby inducing apoptosis."

reach
"Similarly, IP3R3 is bound and deubiquitinated by BRCA1-associated protein 1 (BAP1) at the ER [33] ."

reach
"For example, BAP1 binds to IP3R3 to deubiquitinate and promote apoptosis."

No evidence text available

reach
"Overexpression of the N-terminus of the IP3R3 binds and sequesters BAP1 leading to decreased IP3R3 levels, an effect counter balanced by BAP1 co-overexpression (XREF_FIG)."

No evidence text available

sparser
"Recently, a novel BAP1 cytoplasmic function was introduced, showing that ER-localized BAP1 can bind and stabilize IP3R3 to modulate Ca2+ release resulting in apoptosis [ xref ]."

sparser
"We confirmed the specificity of the BAP1 interaction with IP3R3 in HEK-293 cells expressing FLAG-BAP1 ( xref )."

No evidence text available

reach
"The data clearly indicated that BAP1 physically interacts with the type-3 inositol-1,4,5-trisphosphate-receptor (IP3R3), and BAP1 expression level is directly correlated with IP3R3 in wild-type and mutant fibroblasts."
EED affects BAP1
| 13
| 9

sparser
"The molecular functions of ASXL1 are still incompletely elucidated: ASXL1 forms the polycomb-repressive deubiquitinase (PR-DUB) complex with the H2AK119 deubiquitinase BAP1 [73] , which is a myeloid t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Deubiquitination of nucleosomal H2A is performed using a minimal Polycomb Repressive Deubiquitinase (PR-DUB) complex formed by the human deubiquitinase BAP1 and the DEUBiquitinase ADaptor (DEUBAD) domain of its co-factor ASXL2."

sparser
"In this study, we have identified a critical link between Polycomb repressive deubiquitinase-BAP1 (PR-DUB) complex and BRD4, which is bridged by the physical interaction between additional sex combs-like protein 3 (ASXL3) in SCLC."

sparser
"Numerous studies have demonstrated that truncated ASXL1 mutants, including the ASXL1 fragment containing amino acids 1–587, promote myeloid transformation by forming a stable polycomb-repressive deubiquitinase (PR-DUB) complex with BAP1, enhancing BAP1 deubiquitinase (DUB) activity [ xref , xref , xref ]."

sparser
"The interaction with BRCA1 and BARD1 form a tumor suppressor heterodimeric complex. [ xref ] BAP1 regulates cell proliferation and interacts with histone-modifying complexes during cell division. [ xref , xref ] BAP1 also forms the Polycomb group repressive deubiquitinase complex (PR-DUB) through interaction with ASXL1, which is involved several developmental processes. [ xref ] Because of this functional complexity, BAP1 germline mutations predispose individuals to the aggressive tumor phenotypes."

sparser
"BAP1 interacts with ASXL family members to form the polycomb group repressive deubiquitinase complex involved in stem cell pluripotency and other developmental processes ( xref , xref )."

sparser
"ASXL1 also forms another chromatin modifying complex, the polycomb-repressive deubiquitinase (PR-DUB) complex with the H2AK119 deubiquitinase BAP1 [46] ."

sparser
"BAP1 forms the polycomb group repressive deubiquitinase complex (PR-DUB) with ASXL1/2 in the nucleus."

sparser
"BAP1 and its related proteins form the polycomb inhibitory deubiquitinase (PR-DUB) complex, whose main role is to remove mono-ubiquitin from ubiquitinated histone 2A at lysine 119 (H2Aub)."
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
E2F1 affects BAP1
2 | 5 6
2 | 5 6

sparser
"Chromatin immunoprecipitation (ChIP) assays revealed that BAP1 binds to E2F1 gene promoters, and knockdown of BAP1 leads to increased H2AK119ub1 enrichment, which may lead to transcriptional inactivat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"It showed that BAP1 was greatly reduced at the promoters ( p < 0.05; Fig. 4 E), but no effects on E2F1 occupancy were observed ( Fig. 4 C), suggesting that HCF-1 served as a scaffold for BAP1 to bind [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 binds to E2F1-responsive gene promoters through HCF-1 and upregulates their expression, and BAP1 KO inhibits the growth of uveal melanoma cells ."

No evidence text available

No evidence text available

sparser
"HCF-1 knockdown abolished BAP1 binding to E2F1-responsive promoters, whereas BAP1 knockdown had no effects on HCF-1 enrichment, indicating that HCF-1 possibly acts as a scaffold to recruit BAP1 to spe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Pan et al. have indicated that BAP1 may interact with promoters regulated by E2F1 and affect the cell cycle progression genes controlled by E2F. xref "

sparser
"Our findings suggest that SUFU’s role in CRC is determined not only by its interaction with key molecules like BAP1 and E2F1 but also through Hh-independent mechanisms."

reach
"Chromatin immunoprecipitation (ChIP) assays revealed that BAP1 binds to E2F1 gene promoters, and knockdown of BAP1 leads to increased H2AK119ub1 enrichment, which may lead to transcriptional inactivat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Hence, ChIP-reChIP was performed and the result revealed that BAP1 binds with E2F1 as well as HCF-1 on the E2F1 responsive gene promoter ( Fig. 4 G and H), suggesting that these proteins co-occupy the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects Histone
| 10 1
BAP1 deubiquitinates Histone.
| 6
BAP1 leads to the deubiquitination of Histone on A2. 3 / 3
| 3

reach
"p53 binds to the SLC7A11 promoter, directly suppressing its transcription; the nuclear deubiquitinase (DUB) BAP1 represses SLC7A11 transcription by removing histone 2A ubiquitination from the SLC7A11 promoter; and KEAP1 represses SLC7A11 transcription through degrading NRF2, a master transcription factor of antioxidant response and regulator of SLC7A11."

reach
"Recently, it has been found that BAP1 can inhibit the expression of SLC7A11 to suppress the uptake of cystine by deubiquitinating histone 2A on the SLC7A11 promoter."

reach
"Further mechanistic studies demonstrated that BAP1 suppresses SLC7A11 expression partly by deubiquitinating histone 2A ubiquitination to promote ferroptosis."
BAP1 deubiquitinates Histone. 3 / 3
| 3

reach
"BRCA1 associated protein 1 (BAP1) is a ubiquitin C-terminal hydrolase that deubiquitinates histone H2AK119ub and other proteins and regulates the expression of multiple genes."

reach
"Mechanistically, BAP1 deubiquitinates histone 2AK119-ub1 (H2AK119-ub1), an epigenetic mark that represses gene expression and can have long-lasting effects on transcription."

reach
"Recent studies indicated epigenetic modifications as a result of deubiquitinating the mono-ubiquitinated H2AK119 histone by the hyperactivated BAP1/ASXL1 is one of the main drivers of the transcriptio[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 binds Histone.
| 2 1
| 2 1

reach
"Complex formation between BAP1 and the cell cycle regulator, HCF-1, bridges HCF-1 to histone-modifying complexes to regulate cell growth (2,4)."

sparser
"ASXL3 is less well studied than its paralogs but was shown to promote BAP1 binding to its histone substrate like its paralogs [ xref ]."
| PMC

reach
"We find that ASXL1 directs the binding of the positively charged C-terminal extension of BAP1 to nucleosomal DNA and histones H3-H4 near the dyad, an addition to its role in forming the ubiquitin-binding cleft."
BAP1 ubiquitinates Histone.
| 2
BAP1 leads to the ubiquitination of Histone on A2. 2 / 2
| 2

reach
"Moreover, BAP1 inhibited the expression of solute carrier family 7 member 11 (SLC7A11) by deubiquitinating histone 2 A (H2Aub) on the promoter of SLC7A11."

reach
"Further mechanistic studies demonstrated that BAP1 suppresses SLC7A11 expression partly by deubiquitinating histone 2A ubiquitination to promote ferroptosis."
BAP1 affects EED
| 13
| 9

sparser
"The molecular functions of ASXL1 are still incompletely elucidated: ASXL1 forms the polycomb-repressive deubiquitinase (PR-DUB) complex with the H2AK119 deubiquitinase BAP1 [73] , which is a myeloid t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Deubiquitination of nucleosomal H2A is performed using a minimal Polycomb Repressive Deubiquitinase (PR-DUB) complex formed by the human deubiquitinase BAP1 and the DEUBiquitinase ADaptor (DEUBAD) domain of its co-factor ASXL2."

sparser
"In this study, we have identified a critical link between Polycomb repressive deubiquitinase-BAP1 (PR-DUB) complex and BRD4, which is bridged by the physical interaction between additional sex combs-like protein 3 (ASXL3) in SCLC."

sparser
"Numerous studies have demonstrated that truncated ASXL1 mutants, including the ASXL1 fragment containing amino acids 1–587, promote myeloid transformation by forming a stable polycomb-repressive deubiquitinase (PR-DUB) complex with BAP1, enhancing BAP1 deubiquitinase (DUB) activity [ xref , xref , xref ]."

sparser
"The interaction with BRCA1 and BARD1 form a tumor suppressor heterodimeric complex. [ xref ] BAP1 regulates cell proliferation and interacts with histone-modifying complexes during cell division. [ xref , xref ] BAP1 also forms the Polycomb group repressive deubiquitinase complex (PR-DUB) through interaction with ASXL1, which is involved several developmental processes. [ xref ] Because of this functional complexity, BAP1 germline mutations predispose individuals to the aggressive tumor phenotypes."

sparser
"BAP1 interacts with ASXL family members to form the polycomb group repressive deubiquitinase complex involved in stem cell pluripotency and other developmental processes ( xref , xref )."

sparser
"ASXL1 also forms another chromatin modifying complex, the polycomb-repressive deubiquitinase (PR-DUB) complex with the H2AK119 deubiquitinase BAP1 [46] ."

sparser
"BAP1 forms the polycomb group repressive deubiquitinase complex (PR-DUB) with ASXL1/2 in the nucleus."

sparser
"BAP1 and its related proteins form the polycomb inhibitory deubiquitinase (PR-DUB) complex, whose main role is to remove mono-ubiquitin from ubiquitinated histone 2A at lysine 119 (H2Aub)."
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
BAP1 affects E2F1
2 | 5 6
2 | 5 6

sparser
"Chromatin immunoprecipitation (ChIP) assays revealed that BAP1 binds to E2F1 gene promoters, and knockdown of BAP1 leads to increased H2AK119ub1 enrichment, which may lead to transcriptional inactivat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"It showed that BAP1 was greatly reduced at the promoters ( p < 0.05; Fig. 4 E), but no effects on E2F1 occupancy were observed ( Fig. 4 C), suggesting that HCF-1 served as a scaffold for BAP1 to bind [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 binds to E2F1-responsive gene promoters through HCF-1 and upregulates their expression, and BAP1 KO inhibits the growth of uveal melanoma cells ."

No evidence text available

No evidence text available

sparser
"HCF-1 knockdown abolished BAP1 binding to E2F1-responsive promoters, whereas BAP1 knockdown had no effects on HCF-1 enrichment, indicating that HCF-1 possibly acts as a scaffold to recruit BAP1 to spe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Pan et al. have indicated that BAP1 may interact with promoters regulated by E2F1 and affect the cell cycle progression genes controlled by E2F. xref "

sparser
"Our findings suggest that SUFU’s role in CRC is determined not only by its interaction with key molecules like BAP1 and E2F1 but also through Hh-independent mechanisms."

reach
"Chromatin immunoprecipitation (ChIP) assays revealed that BAP1 binds to E2F1 gene promoters, and knockdown of BAP1 leads to increased H2AK119ub1 enrichment, which may lead to transcriptional inactivat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Hence, ChIP-reChIP was performed and the result revealed that BAP1 binds with E2F1 as well as HCF-1 on the E2F1 responsive gene promoter ( Fig. 4 G and H), suggesting that these proteins co-occupy the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects DNA Damage
| 13
| 10

reach
"Molecularly, Bap1 null melanoma cell lines have increased DNA damage measured by gammaH2aX and hyperubiquitination of histone H2a."

reach
"In heterozygous BAP1 conditions, as in the individuals of the families with the BAP1 cancer syndrome, the reduced BAP1 dosage impairs both the DNA repair, accumulating DNA damage, and the apoptotic response."

reach
"In summary, reduced BAP1 levels cause cells to accumulate DNA damage, a process that normally triggers apoptosis."

reach
"Here, we show that BAP1 promotes the repair of ultraviolet (UV)-induced DNA damage via its deubiquitination activity in various cell types, including primary melanocytes."

reach
"BAP1 can deubiquitylate Ub-H2AK119 at the site of UV-induced DNA damage, and promote DNA repair by regulating the accumulation of repair proteins (BRCA1, RAD51, and RPA) (7)."

reach
"These results demonstrated that PARP1 recruits BAP1 to sites of UV-induced DNA damage."

reach
"BAP1, a ubiquitin C-terminal hydrolase domain, can promote the repair of DNA damage induced by ultraviolet light (UV) through deubiquitination activity."

reach
"The study showed that BAP1 is recruited to sites of UV-induced DNA damage via H2A-K119-Ub and PARP1 to promote DNA repair and that both DUB activity and PARP1-mediated poly(ADP-ribosyl)ation are critical for this function of BAP1 ."

reach
"As already said, BAP1 modulates DNA damage repair mechanism."

reach
"BAP1 modulates double-strand DNA damage repair."
Mutated BAP1 activates DNA Damage. 3 / 3
| 3

reach
"Our findings suggested that BAP1 mutation, as a loss of function, upregulated the TCA cycle and initiated DNA damage through ROS accumulation, resulting in PARP1 hyperactivation–induced cell apoptosis (Figure 4P)."

reach
"BAP1 mutations enable mesothelial cells to evade apoptosis and accumulate further DNA damage, leading to carcinogenesis."

reach
"However, the spontaneous development of mesotheliomas in Bap1 +/- mice not exposed to asbestos, and the development of multiple cancer types in carriers of BAP1 mutations (XREF_FIG), including tumor types that have not been associated with known carcinogens, suggests that BAP1 mutations also drive tumor growth independently of genotoxic stress, perhaps by favoring the accumulation of age related DNA damage."
VPS affects BAP1
| 2 10
| 2 10

reach
"For example, binding between LecB and CdrA to the Psl polysaccharide in P. aeruginosa biofilms contribute to cell–cell junction formation, and binding of RbmA, RbmC, and Bap1 to VPS contribute to biofilm formation in Vibrio cholerae."

reach
"These observations indicate that Bap1 and RbmC could bind VPS, and these interactions may influence the spatial localization of these matrix proteins."

sparser
"Importantly, vesicles prepared from the strain unable to produce RbmA/RbmC/Bap1/VPS still affected microcolony formation within the biofilm although not in the same way as OMVs isolated from the parent strain."

sparser
"These observations indicate that Bap1 and RbmC could bind VPS, and these interactions may influence the spatial localization of these matrix proteins."

sparser
"Next, we investigated how the RbmC and Bap1 matrix proteins interact with VPS during biofilm formation."

sparser
"We combine genetics, microscopy, simulations, and biochemical analyses to show that V. cholerae cells are not attracted to the main matrix component ( Vibrio polysaccharide, VPS), but can be attached to each other and to the VPS network through surface-associated VPS and crosslinks formed by the protein Bap1."

sparser
"When we added Bap1 to a growing Δ ABC culture, we found disordered cell aggregates loosely connected by the VPS-Bap1 network (Fig.  xref and Fig.  xref )."

sparser
"Crucially, the recognition of community membership is closely tied to the active secretion of VPS: cells coated by VPS – which only occurs when the exopolysaccharide is actively being produced – are recognized by the existing VPS-Bap1 network or other VPS-coated cells and consequently included within the biofilm through bridging-aggregation."

sparser
"Because the matrix proteins RbmA, RbmC, and Bap1 all directly interact with VPS, biofilms lacking VPS are thought not to retain the three primary matrix proteins, either."

sparser
"Interestingly, two adhesins — Bap1 ( B iofilm- a ssociated p rotein 1) and its homologue RbmC ( R ugosity and b iofilm m odulator C) directly bind to VPS [ xref ], thus resulting in a cross-linking of cells with the matrix similar to CdrA in P. aeruginosa ."
VCP affects BAP1
2 | 1 9
2 | 1 9

sparser
"To further confirm the protein–protein interaction of BAP1 with VCP, we conducted co‐immunoprecipitation analysis by co‐transfecting 293T cells with plasmids encoding Flag‐tagged VCP and Myc‐tagged BAP1."

sparser
"Mechanically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway, thus promoting CCA progression."

No evidence text available

sparser
"It identifies VCP as a key upstream regulator mediating BAP1 homeostasis: VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway."

sparser
"In summary, these data illustrated a physical interaction between BAP1 and VCP, suggesting that VCP may act as a novel regulator of BAP1 in CCA."

sparser
"Mechanistically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway, thus promoting cell proliferation and inhibiting cell apoptosis."

No evidence text available

sparser
"The results showed that exogenous VCP physically interacted with BAP1 (Figure  xref )."

sparser
"We also observed interactions between Flag‐tagged VCP and endogenous BAP1 (Figure  xref ), as well as Myc‐tagged BAP1 and endogenous VCP (Figure  xref )."

reach
"Mechanistically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin-proteasome pathway, thus promoting cell proliferation and inhibiting cell apoptosis."
PGR affects BAP1
| 12
| 12

sparser
"Together, our data suggested a model that the BAP1-containing PR-DUB complex binds on the S LC7A11 promoter, where BAP1 removes ubiquitin from H2Aub, and BAP1-dependent H2Aub reduction on SLC7A11 is associated with BAP1-mediated SLC7A11 repression."

sparser
"Our results reveal that the physiological modulation of BAP1 determines the invasive properties of the trophoblast, delineating a new role of the BAP1 PR-DUB complex in regulating early placentation."

sparser
"These BAP1 mutants did not significantly affect BAP1’s interaction with other components of the PR-DUB complex ( xref )."

sparser
"The N-terminal PHD region of MLL3/KMT2C has been shown to bind the BAP1-containing PR-DUB complex, which is also a tumor suppressor and acts as a deubiquitinase for histone H2A."

sparser
"Similar to ASXL1, ASXL2 forms a stable and distinct PR-DUB complex with BAP1, promoting ubiquitin removal from histone H2A. However, unlike ASXL1, ASXL2 is stabilized by BAP1 [ xref , xref ], indicating the existence of additional regulatory mechanisms."

sparser
"In human cells, ASXL3 protein was shown to interact with BAP1 to form the PR-DUB complex ( xref )."

sparser
"The BAP1-PR-DUB interaction was confirmed in PLC cells by coimmunoprecipitation (fig."

sparser
"Truncated ASXL1 mutants, specifically the ASXL1 fragment containing amino acids 1–587, drive myeloid transformation by forming a stable PR-DUB complex with BAP1, thereby enhancing BAP1 deubiquitinase activity [ xref , xref , xref ]."

sparser
"Truncated ASXL1 mutants promote myeloid transformation by creating a potent PR-DUB complex with BAP1 [ xref , xref , xref ]."

sparser
"BAP1 PR-DUB complex is also regulated during human trophoblast differentiation."
MBD6 affects BAP1
4 | 8
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
Flag affects BAP1
| 1 11
| 1 11

sparser
"In support of the notion that BAP1 might be an upstream DUB for MYCN, Flag-BAP1 and HA-MYCN were co-transfected into 293T cells and the immunoprecipitation (IP) assay was performed to examine whether BAP1 will interact with MYCN."

sparser
"Immunofluorescence (IF) staining of HEK293T cells co-transfected with Flag-BAP1 and Myc-SUFU revealed strong cytoplasmic co-localization of the two proteins, with minimal nuclear overlap (Figs.  xref B and S1A), further supporting their protein-protein interaction in cells."

sparser
"HeLa cells stably expressing Flag-BAP1 were selected and lysed with IP150 buffer (200 mM Tris-HCl, pH 7.9, 15% glycerol, 1 mM EDTA, 1 mM DTT, 0.2 mM PMSF, 0.5% NP-40, and 150 mM KCl)."

sparser
"To identify the O -GlcNAcylated region of BAP1, Flag-BAP1 deletion fragments (N-terminal, amino acids [aa] 1–239; C-terminal, aa 240–729) were generated and transfected into HCT116 cells."

sparser
"Whole-cell lysates from HEK293 cells transfected with Flag-BAP1 were pulled down using WGA agarose beads."

sparser
"To investigate the biochemical function of BAP1, we performed affinity purification of the BAP1-associated protein complex using HeLa cells stably expressing Flag-BAP1."

sparser
"Moreover, we observed that Flag-ASXL1 interacted with endogenous HCF-1, FOXK1 and FOXK2 in BAP1 knockdown HEK293T cells (Fig.  xref G), and that Flag-BAP1 interacted with endogenous HCF-1, FOXK1, and FOXK2 in ASXL1 knockdown HEK293T cells (Fig.  xref H)."

sparser
"Both Flag-BAP1 and GlcNAc were detected in the elutes of WGA beads but not in control agarose (A/G) beads, further supporting that BAP1 is modified by O- GlcNAc ( Fig. 2 B)."

reach
"Full-length BAP1 (1-729), the BAP1 N terminus (1-240) and the BAP1 C terminus (597-729) interacted with KLF5-3 x Flag (XREF_FIG)."

sparser
"To directly check whether MYCN protein stability is regulated by BAP1-dependent deubiquitination, the Flag-BAP1, His-Ub and HA-MYCN were co-transfected into 293T cells."
FlaG affects BAP1
| 12
| 12

sparser
"However, FLAG-BAP1 failed to attenuate a decrease in the level of HA-KEAP1ΔBTB induced by treatment of CHX in 293T-HA-KEAP1ΔBTB cells ( xref C, right panel), suggesting that the BTB domain is responsible for maintaining the protein stability of KEAP1 by BAP1."

sparser
"Our results show that transduced FLAG-BAP1 is mostly located in the cytosol ( xref A)."

sparser
"Similar transverse distributions of each protein were observed in biofilms formed by a Δbap1 mutant expressing Bap1-FLAG from a plasmid alone and by a ΔrbmA mutant expressing RbmA-FLAG from a plasmid alone (data not shown)."

sparser
"In addition, lentiviral transduction of FLAG-BAP1 suppressed the migration ( xref E) and the invasion ( xref F) of A549 cells, and potentiated cisplatin-induced apoptosis in A549 cells ( xref G,H)."

sparser
"Real-time RT-PCR analysis demonstrates that lentiviral transduction of FLAG-BAP1 significantly decreased the mRNA level of NRF2 target genes such as heme oxygenase-1 (Hmox1), NADPH:quinone oxidoreductase-1 (Nqo1), glutamate cysteine ligase catalytic subunit (Gclc), and aldo-keto reductase family 1 member B10 (Akr1b10) in A549 cells ( xref D)."

sparser
"We then conducted ChIP-qPCR assays in UMRC-6 cells reconstituted with FLAG-BAP1 or empty vector."

sparser
"Immunoprecipitation revealed that transiently overexpressed FLAG-BAP1 bound to both transiently overexpressed GFP-ERK1 and GFP-ERK2 ( xref )."

sparser
"We observed that FLAG-BAP1 ( xref B, upper panel), but not FLAG-UCHL3 ( xref B, lower panel), directly binds to HA-KEAP1."

sparser
"However, FLAG-BAP1 did not bind to HA-NRF2 ( xref C)."

sparser
"Our result shows that FLAG-BAP1 does not selectively bind to HA-KEAP1ΔBTB ( xref A, right panel), suggesting that BAP1 binds to the BTB domain of KEAP1."
EZH2 affects BAP1
| 9 3
EZH2 binds BAP1.
| 2 3
| 2 3

reach
"BAP-1 interactions with histone deacetylase (HDAC) and enhancer of zeste homolog 2 (EZH2) have been exploited to suppress MPM progression."

sparser
"The interaction of BAP1 and EZH2 and the efficacy of tazemetostat show great potential for the future of cancer treatment for patients harboring variants in those genes due to the more aggressive nature of those tumors and the lack of targeted treatments."

sparser
"Survival analysis revealed that BAP1 loss, but not high EZH2 expression, was associated with a better prognosis."

reach
"We did not identify an interaction between BAP1 and EZH2 by co-immunoprecipitation (XREF_FIG)."

sparser
"In these lung-iPSCs, the expression of EZH2 (the main component of PRC2 complex) is also very high, and the physical interactions of AXSL proteins with EZH2 and BAP1 is also reported [ xref , xref , xref ]."
EZH2 activates BAP1.
| 4
EZH2 activates BAP1. 4 / 4
| 4

reach
"In BAP1-deficient human mesothelioma cells, EZH2 inhibition also produced synthetic lethality ."

reach
"Inactivation of BAP1 sensitises tumour cells to EZH2 inhibition, which has been shown to substantially inhibit tumour growth in BAP1 -deficient malignant pleural mesothelioma mouse models."

reach
"Interestingly, EZH2 depletion by both genetic and pharmacologic methods abrogated the oncogenic effect of BAP1 loss."

reach
"EZH2 inhibition in combination with ZA limits tumor growth and prolongs survival of Bap1-deficient mesothelioma mice."
EZH2 inhibits BAP1.
| 3
EZH2 inhibits BAP1. 3 / 3
| 3

reach
"Concomitant Bap1 and Ezh2 loss reduced myeloid progenitor expansion, reduced the proportion of Mac1 + Gr1 + myeloid cells, restored erythroid differentiation (CD71 + Ter119 +) and decreased proliferation of Bap1 and Ezh2 deficient progenitors (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"35 This is in line with our observations that EZH2 inhibition upregulates genes of the cholesterol and mevalonate biosynthesis pathways in BAP1-mutated settings with concomitant addiction to these pathways."

reach
"Ezh2 haploinsufficiency reduced, but did not abrogate, Bap1 deficient myeloproliferation (XREF_SUPPLEMENTARY) consistent with a dose dependent requirement for Ezh2."
| 1 11
BAP1 activates cell migration.
| 7
| 7

reach
"It was observed that overexpression of BAP1 WT slightly decreased invasion ability but increased cell migration."

reach
"Upregulation of miR-31 and Knockdown of BAP1 Promoted Cell Migration In Vitro."

reach
"BAP1 probably accelerates ECA109 cell migration and proliferation by increasing KLF5 expression [20]."

reach
"We found that dual knockdown of BAP1 and miR-31 significantly reversed cell migration ability of HeLa and C33A cells induced by single-knockdown of miR-31."

reach
"BAP1 promotes breast cancer cell proliferation and metastasis through deubiquitination of KLF5.33 High BBS1 gene expression facilitates survival of malignant pleural mesothelioma.34 Crosstalk between macrophages and cancer cells via CCR2 and CX3CR1 is the underlying mechanism driving lung cancer.35 RIPK4 overexpression promotes pancreatic cancer cell migration and invasion through PEBP1 degradation-induced activation of the RAF1/MEK/ERK pathway."

reach
"BAP1 depletion appears to promote UM cell migration underneath the monolayer."

reach
"BaP1 IC and 2 × IC triggered a decrease in cell migration, compared with untreated cells, after 12 h of treatment (Figure 3e,f)."
BAP1 inhibits cell migration.
| 1 4
| 1 4

reach
"Knockdown of BAP1 and KLF5 significantly decreased cell migration compared with the Lucsh negative control in HCC1806 and HCC1937 cells using wound healing and transwell assays (XREF_FIG; XREF_SUPPLEMENTARY)."

reach
"The overexpression of CAST rescued the cell migration phenotype caused by BAP1 loss."

reach
"BaP1, a phenoxazine derivative, reduces cell proliferation, survival and cell migration through ROS generation, apoptosis promotion and angiogenesis inhibition."

reach
"However, BAP1 is closely related to lymph node status, suggesting that BAP1 also mediates suppression of cell migration in LAC which is in keeping with the findings of the wound healing assay."

eidos
"We found that BaP1 is selective for CRC cells and reduces cell proliferation , cell survival , and cell migration ."
BAP1 affects VPS
| 2 10
| 2 10

reach
"For example, binding between LecB and CdrA to the Psl polysaccharide in P. aeruginosa biofilms contribute to cell–cell junction formation, and binding of RbmA, RbmC, and Bap1 to VPS contribute to biofilm formation in Vibrio cholerae."

reach
"These observations indicate that Bap1 and RbmC could bind VPS, and these interactions may influence the spatial localization of these matrix proteins."

sparser
"Importantly, vesicles prepared from the strain unable to produce RbmA/RbmC/Bap1/VPS still affected microcolony formation within the biofilm although not in the same way as OMVs isolated from the parent strain."

sparser
"These observations indicate that Bap1 and RbmC could bind VPS, and these interactions may influence the spatial localization of these matrix proteins."

sparser
"Next, we investigated how the RbmC and Bap1 matrix proteins interact with VPS during biofilm formation."

sparser
"We combine genetics, microscopy, simulations, and biochemical analyses to show that V. cholerae cells are not attracted to the main matrix component ( Vibrio polysaccharide, VPS), but can be attached to each other and to the VPS network through surface-associated VPS and crosslinks formed by the protein Bap1."

sparser
"When we added Bap1 to a growing Δ ABC culture, we found disordered cell aggregates loosely connected by the VPS-Bap1 network (Fig.  xref and Fig.  xref )."

sparser
"Crucially, the recognition of community membership is closely tied to the active secretion of VPS: cells coated by VPS – which only occurs when the exopolysaccharide is actively being produced – are recognized by the existing VPS-Bap1 network or other VPS-coated cells and consequently included within the biofilm through bridging-aggregation."

sparser
"Because the matrix proteins RbmA, RbmC, and Bap1 all directly interact with VPS, biofilms lacking VPS are thought not to retain the three primary matrix proteins, either."

sparser
"Interestingly, two adhesins — Bap1 ( B iofilm- a ssociated p rotein 1) and its homologue RbmC ( R ugosity and b iofilm m odulator C) directly bind to VPS [ xref ], thus resulting in a cross-linking of cells with the matrix similar to CdrA in P. aeruginosa ."
BAP1 affects VCP
2 | 1 9
2 | 1 9

sparser
"To further confirm the protein–protein interaction of BAP1 with VCP, we conducted co‐immunoprecipitation analysis by co‐transfecting 293T cells with plasmids encoding Flag‐tagged VCP and Myc‐tagged BAP1."

sparser
"Mechanically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway, thus promoting CCA progression."

No evidence text available

sparser
"It identifies VCP as a key upstream regulator mediating BAP1 homeostasis: VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway."

sparser
"In summary, these data illustrated a physical interaction between BAP1 and VCP, suggesting that VCP may act as a novel regulator of BAP1 in CCA."

sparser
"Mechanistically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin‐proteasome pathway, thus promoting cell proliferation and inhibiting cell apoptosis."

No evidence text available

sparser
"The results showed that exogenous VCP physically interacted with BAP1 (Figure  xref )."

sparser
"We also observed interactions between Flag‐tagged VCP and endogenous BAP1 (Figure  xref ), as well as Myc‐tagged BAP1 and endogenous VCP (Figure  xref )."

reach
"Mechanistically, VCP binds to BAP1 and promotes the latter's ubiquitination degradation via the ubiquitin-proteasome pathway, thus promoting cell proliferation and inhibiting cell apoptosis."
BAP1 affects PGR
| 12
| 12

sparser
"Together, our data suggested a model that the BAP1-containing PR-DUB complex binds on the S LC7A11 promoter, where BAP1 removes ubiquitin from H2Aub, and BAP1-dependent H2Aub reduction on SLC7A11 is associated with BAP1-mediated SLC7A11 repression."

sparser
"Our results reveal that the physiological modulation of BAP1 determines the invasive properties of the trophoblast, delineating a new role of the BAP1 PR-DUB complex in regulating early placentation."

sparser
"These BAP1 mutants did not significantly affect BAP1’s interaction with other components of the PR-DUB complex ( xref )."

sparser
"The N-terminal PHD region of MLL3/KMT2C has been shown to bind the BAP1-containing PR-DUB complex, which is also a tumor suppressor and acts as a deubiquitinase for histone H2A."

sparser
"Similar to ASXL1, ASXL2 forms a stable and distinct PR-DUB complex with BAP1, promoting ubiquitin removal from histone H2A. However, unlike ASXL1, ASXL2 is stabilized by BAP1 [ xref , xref ], indicating the existence of additional regulatory mechanisms."

sparser
"In human cells, ASXL3 protein was shown to interact with BAP1 to form the PR-DUB complex ( xref )."

sparser
"The BAP1-PR-DUB interaction was confirmed in PLC cells by coimmunoprecipitation (fig."

sparser
"Truncated ASXL1 mutants, specifically the ASXL1 fragment containing amino acids 1–587, drive myeloid transformation by forming a stable PR-DUB complex with BAP1, thereby enhancing BAP1 deubiquitinase activity [ xref , xref , xref ]."

sparser
"Truncated ASXL1 mutants promote myeloid transformation by creating a potent PR-DUB complex with BAP1 [ xref , xref , xref ]."

sparser
"BAP1 PR-DUB complex is also regulated during human trophoblast differentiation."
BAP1 affects MBD6
4 | 8
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
BAP1 affects KEAP1
1 2 | 4 5
BAP1 binds KEAP1.
1 2 | 1 5
1 2 | 1 5

sparser
"Alternatively, it can be speculated that BAP1 might competitively bind to KEAP1 at the BTB domain with unknown protein(s) affecting the stability of KEAP1."

sparser
"In order to identify the domains of BAP1 that interact with KEAP1, we generated FLAG-BAP1 mutant plasmids lacking individual domains (FLAG-BAP1ΔCD, FLAG-BAP1ΔCC1, FLAG-BAP1ΔCL, FLAG-BAP1ΔCC2, FLAG-BAP1ΔULD, and FLAG-BAP1ΔCTE)."

reach
"Tumor suppressors, such as p53, BAP1, Kelch-like ECH associated protein 1 (KEAP1), function through induction of ferroptosis mainly via regulating SLC7A11 expression [4], [5], [6]."

sparser
"Immunoprecipitation of endogenous BAP1 demonstrates that BAP1 interacts with endogenous KEAP1 ( xref D)."

sparser
"In order to identify the domain of KEAP1 that interacts with BAP1, we generated pcDNA3-HA-KEAP1 mutant plasmids lacking individual domains (HA-KEAP1ΔBTB, HA-KEAP1ΔBACK, and HA-KEAP1ΔKELCH)."

"BAP1 directly binds to and deubiquitinates KEAP1. BAP1 stabilizes KEAP1 by binding to the BTB domain."

sparser
"Our result shows that FLAG-BAP1 does not selectively bind to HA-KEAP1ΔBTB ( xref A, right panel), suggesting that BAP1 binds to the BTB domain of KEAP1."

No evidence text available

No evidence text available
BAP1 deubiquitinates KEAP1.
| 3
BAP1 deubiquitinates KEAP1. 3 / 3
| 3

reach
"In the present study, we have identified that BAP1 deubiquitinates KEAP1 by binding to the BTB domain."

reach
"Together, our study demonstrates that KEAP1 deubiquitination by BAP1 is novel tumor suppressive mechanism of LUAD."

reach
"BAP1 Downregulates NRF2 Target Genes and Exerts Anti-Tumorigenic Effects by Deubiquitinating KEAP1 in Lung Adenocarcinoma."
BAP1 affects Histone_H2A
| 12
BAP1 ubiquitinates Histone_H2A.
| 7
BAP1 ubiquitinates Histone_H2A. 7 / 7
| 7

reach
"Genomic analyses together with robust statistics have revealed that the restoration of BAP1 facilitates the formation of the polycomb repressive deubiquitinase (PR-DUB) complex and inhibits ubiquitinated histone 2A (H2Aub) occupancy on the SLC7A11 promoter."

reach
"BRCA1-associated protein 1 (BAP1) [18], the crucial constituent of the deubiquitinase complex, attenuated the initiation and extension of SLC7A11 transcription by deubiquitinating histone 2A, while the steady-state levels and half-life of SLC7A11 were improved by combination with the ovarian tumor family member deubiquitinase (OTUB1) [19]."

reach
"Several studies have revealed that BAP1 can inhibit ubiquitinated histone 2A (H2Aub) occupancy on the SLC7A11 promoter (Zhang et al., 2018)."

reach
"The mechanism may be that BAP1 suppresses SLC7A11 transcription by reducing histone 2A ubiquitination (H2Aub) on the SLC7A11 gene."

reach
"Moreover, the tumor suppressor gene, BRCA1-associated protein 1 (BAP1), was found to reduce SLC7A11 expression by decreasing histone 2A ubiquitination occupancy on its promoter in human cancer, which consequently resulted in increased ferroptosis (Zhang et al., 2018)."

reach
"The tumour suppressor BRCA1-associated protein 1 (BAP1) is a nuclear deubiquitinating enzyme and can reduce histone 2A ubiquitination (H2Aub) on chromatin."

reach
"BAP1 mediated deubiquitination of histone 2A ubiquitination (H2Aub), while monoubiquitination of histone H2B on lysine 120 (H2Bub1) induced the expression of SLC7A11 during ferroptosis and consequentl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 deubiquitinates Histone_H2A.
| 5
BAP1 deubiquitinates Histone_H2A. 5 / 5
| 5

reach
"BRCA1-associated protein 1 (BAP1) [18], the crucial constituent of the deubiquitinase complex, attenuated the initiation and extension of SLC7A11 transcription by deubiquitinating histone 2A, while the steady-state levels and half-life of SLC7A11 were improved by combination with the ovarian tumor family member deubiquitinase (OTUB1) [19]."

reach
"BAP1 deubiquitinates histone 2A (H2Aub) on the SLC7A11 promoter and represses SLC7A11 expression in a p53-independent way (39, 40)."

reach
"The mechanism may be that BAP1 represses the transcription of SLC7A11 by reducing histone 2A ubiquitination (H2Aub) on the SLC7A11 gene (49)."

reach
"BAP1 deubiquitinates histone 2A in the regulatory chromatin of the of SLC7A11 gene, which encodes, a key transporter of extracellular cysteine."

reach
"Specially, previous studies demonstrate that BAP1 downregulates SLC7A11 expression by reducing histone 2A ubiquitination occupancy on its promoter (Zhang et al., 2018b)."
BAP1 affects HIF1
| 12
BAP1 binds HIF1.
| 10
| 10

reach
"Co-immunoprecipitation (CoIP) and proximity ligation assay (PLA) experiments revealed that 1) HIF-1α and BAP1 bind to each other and co-precipitate (Fig. 2A), and 2) the nuclei of BAP1 cells contained significantly more PLA positive signals—evidence of BAP1 and HIF-1α interaction—than BAP1 cells (Fig. 2 B and C)."

reach
"The crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) (49) shows that without BAP1, HIF-1α, and HIF-1β bind to DNA (49), thus BAP1 is not required for HIF-1α-HIF-1β complex formation."

reach
"Aligning the crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) (49) to our structural model for the binding complex of BAP1-HIF-1α showed that both BAP1 and HIF-1β bind to the same residues of HIF-1α (1-73) on the DNA; however, in Fig. 3B, we demonstrate that BAP1, HIF-1α and the DNA form a complex without HIF-1β."

reach
"Therefore, our data indicate that BAP1 is not required for HIF-1α-HIF-1β complex formation to functionally bind to DNA, that HIF-1β is not required for BAP1-HIF-1α complex formation to functional binding to DNA, and that although DNA facilitates the binding of BAP1 and HIF-1α, it is not required to maintain the binding of both BAP1-HIF-1α and BAP1-HIF-1β."

reach
"In summary, our data suggest that BAP1 directly binds and stabilizes both HIF-1α and HIF-1β increasing their intra-nuclear availability for dimer formation, thus fine-tuning HIF activities required to support malignant cell growth."

reach
"So far, the pathogenic variants reported in ClinVar for both HIF-1α and HIF-1β are not located among the crucial residues of HIF-1α-r73 where HIF-1α can bind to BAP1, HIF-1β and DNA, or of HIF-1β (2-470) where HIF-1β can bind to BAP1, HIF-1α, and DNA."

reach
"We further investigated the specificity of the BAP1 interaction with HIF-1α in HEK-293 cells expressing Myc-BAP1 and HA-HIF-1α, using HA-Tag as bait."

reach
"To study the binding between BAP1 and HIF-1α, first, a rigid docking protocol was applied to model the binding complex of the CTD of BAP1 (the NLS domain is removed due to its flexibility) and HIF-1α by using the ClusPro server (43–45) (SI Appendix, Fig. S2B)."

reach
"Notably, BAP1 binding to HIF-1α-r73-DNA does not require HIF-1β for the interaction."

reach
"We tested the hypothesis that although DNA facilitates the binding between BAP1 and HIF-1α, this binding complex still holds in the absence of DNA."
BAP1 decreases the amount of HIF1.
| 2
BAP1 decreases the amount of HIF1. 2 / 2
| 2

reach
"We found the opposite to be true: BAP1 primary cells had reduced HIF-1α levels in hypoxia, and mesothelioma cells carrying biallelic BAP1 mutations almost constantly lost nuclear HIF-1α."

reach
"Mutations in BAP1 that disrupt its function result in decreased levels of HIF-1α, implicating BAP1 as a key regulator of HIF-1α stability in response to low oxygen levels ."
| 12
| 6

reach
"Specifically, BAP1 suppresses genome instability via its DNA repair activity and eliminates cancer-prone cells that exhibit genome instability by promoting apoptosis."

reach
"Here, we were particularly interested in potential roles of BAP1 in controlling mitotic progression and chromosome segregation and hence how loss of BAP1 contributes to genome instability."

reach
"BAP1 loss induces BRCA1 dependent and independent defective spindle organization, checkpoint dysfunction and genomic instability."

reach
"This indicates that BAP1 loss did not significantly augment genomic instability in our BNC mouse model."

reach
"The loss of BAP1 leads to genomic instability, which contribute to chronic pancreatitis and predisposes to cancer development11."

reach
"Loss of Bap1 causes genomic instability."
| 6

reach
"The lack of BAP1 was shown to increase genome instability and alter the response to microtubule targeting agents [67]."

reach
"Importantly though, we observed a number of extra mitotic phenotypes upon depletion of BAP1 that did not occur upon BRCA1 depletion indicating the presence of additional mechanisms through which BAP1 may contribute to mitotic progression and genome stability (Fig. 7)."

reach
"Deletion or heterozygous loss of Bap1 in murine pancreata causes genomic instability, tissue damage, and pancreatitis with full penetrance."

reach
"The loss of BAP1 also impairs the DNA damage response and increases genomic instability."

reach
"Low BAP1 expression promotes genomic instability by failing to regulate DNA repair pathways or by altering the expression of other tumor suppressor genes, contributing to an aggressive cancer phenotype."

reach
"The complement factor H-related (CFHR) protein family induces the ubiquitination of polo-like kinase 1 (PLK1) at lysine 209 (Zhu et al., 2021a), BRCA1-associated protein 1 (BAP1) enhances genomic stability (Perkail et al., 2020), and USP9X is found to restrain tumor growth in the patient-derived tumor xenograft (PDX) model (Pal et al., 2018)."
BAP1 affects Flag
| 1 11
| 1 11

sparser
"In support of the notion that BAP1 might be an upstream DUB for MYCN, Flag-BAP1 and HA-MYCN were co-transfected into 293T cells and the immunoprecipitation (IP) assay was performed to examine whether BAP1 will interact with MYCN."

sparser
"Immunofluorescence (IF) staining of HEK293T cells co-transfected with Flag-BAP1 and Myc-SUFU revealed strong cytoplasmic co-localization of the two proteins, with minimal nuclear overlap (Figs.  xref B and S1A), further supporting their protein-protein interaction in cells."

sparser
"HeLa cells stably expressing Flag-BAP1 were selected and lysed with IP150 buffer (200 mM Tris-HCl, pH 7.9, 15% glycerol, 1 mM EDTA, 1 mM DTT, 0.2 mM PMSF, 0.5% NP-40, and 150 mM KCl)."

sparser
"To identify the O -GlcNAcylated region of BAP1, Flag-BAP1 deletion fragments (N-terminal, amino acids [aa] 1–239; C-terminal, aa 240–729) were generated and transfected into HCT116 cells."

sparser
"Whole-cell lysates from HEK293 cells transfected with Flag-BAP1 were pulled down using WGA agarose beads."

sparser
"To investigate the biochemical function of BAP1, we performed affinity purification of the BAP1-associated protein complex using HeLa cells stably expressing Flag-BAP1."

sparser
"Moreover, we observed that Flag-ASXL1 interacted with endogenous HCF-1, FOXK1 and FOXK2 in BAP1 knockdown HEK293T cells (Fig.  xref G), and that Flag-BAP1 interacted with endogenous HCF-1, FOXK1, and FOXK2 in ASXL1 knockdown HEK293T cells (Fig.  xref H)."

sparser
"Both Flag-BAP1 and GlcNAc were detected in the elutes of WGA beads but not in control agarose (A/G) beads, further supporting that BAP1 is modified by O- GlcNAc ( Fig. 2 B)."

reach
"Full-length BAP1 (1-729), the BAP1 N terminus (1-240) and the BAP1 C terminus (597-729) interacted with KLF5-3 x Flag (XREF_FIG)."

sparser
"To directly check whether MYCN protein stability is regulated by BAP1-dependent deubiquitination, the Flag-BAP1, His-Ub and HA-MYCN were co-transfected into 293T cells."
BAP1 affects FlaG
| 12
| 12

sparser
"However, FLAG-BAP1 failed to attenuate a decrease in the level of HA-KEAP1ΔBTB induced by treatment of CHX in 293T-HA-KEAP1ΔBTB cells ( xref C, right panel), suggesting that the BTB domain is responsible for maintaining the protein stability of KEAP1 by BAP1."

sparser
"Our results show that transduced FLAG-BAP1 is mostly located in the cytosol ( xref A)."

sparser
"Similar transverse distributions of each protein were observed in biofilms formed by a Δbap1 mutant expressing Bap1-FLAG from a plasmid alone and by a ΔrbmA mutant expressing RbmA-FLAG from a plasmid alone (data not shown)."

sparser
"In addition, lentiviral transduction of FLAG-BAP1 suppressed the migration ( xref E) and the invasion ( xref F) of A549 cells, and potentiated cisplatin-induced apoptosis in A549 cells ( xref G,H)."

sparser
"Real-time RT-PCR analysis demonstrates that lentiviral transduction of FLAG-BAP1 significantly decreased the mRNA level of NRF2 target genes such as heme oxygenase-1 (Hmox1), NADPH:quinone oxidoreductase-1 (Nqo1), glutamate cysteine ligase catalytic subunit (Gclc), and aldo-keto reductase family 1 member B10 (Akr1b10) in A549 cells ( xref D)."

sparser
"We then conducted ChIP-qPCR assays in UMRC-6 cells reconstituted with FLAG-BAP1 or empty vector."

sparser
"Immunoprecipitation revealed that transiently overexpressed FLAG-BAP1 bound to both transiently overexpressed GFP-ERK1 and GFP-ERK2 ( xref )."

sparser
"We observed that FLAG-BAP1 ( xref B, upper panel), but not FLAG-UCHL3 ( xref B, lower panel), directly binds to HA-KEAP1."

sparser
"However, FLAG-BAP1 did not bind to HA-NRF2 ( xref C)."

sparser
"Our result shows that FLAG-BAP1 does not selectively bind to HA-KEAP1ΔBTB ( xref A, right panel), suggesting that BAP1 binds to the BTB domain of KEAP1."
BAP1 affects E3_Ub_ligase
| 6 6
BAP1 inhibits E3_Ub_ligase.
| 6 1
| 6 1

reach
"XREF_BIBR suggested that BAP1 inhibits the E3 ligase activity of BRCA1 in a manner independent of its deubiquitination activity."

sparser
"In this regard, however, Nishikawa et al . xref suggested that BAP1 inhibits the E3 ligase activity of BRCA1 in a manner independent of its deubiquitination activity."

reach
"However, another study demonstrated that the E3 activity of BRCA1–BARD1 could be inhibited by BAP1, which mediates HR repair and confer radioresistance17."

reach
"The E3 ligase activity of the BRCA1-BARD1 tumor-suppressor complex is suppressed by the interaction of BAP1 with BARD1 through its deubiquitylation."

reach
"BAP1 acts as a tumor suppressor and inhibits E3 ligase activity in BRCA1/BARD1."

reach
"The BRCA1 associated protein 1 (BAP1), a de-ubiquitinating enzyme (DUB), also inhibits the E3 ligase activity of BRCA1-BARD1 (Nishikawa et al., 2009)."

reach
"BAP1, a ubiquitin carboxy-terminal hydrolase, interacts with the RING finger domain of BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1, protecting BRCA1 from proteasome-mediated degradation (37, 38)."
| 5

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"One of these is BAP1 which binds the BRCA1 ubiquitin ligase and functions as a tumor suppressor ( Ventii et al., 2008 )."

sparser
"In addition, ubiquitin ligase, which is closely associated with BAP1 protein, is a good candidate therapeutic target."
MiR-31 affects BAP1
| 11
MiR-31 decreases the amount of BAP1.
| 6
Modified miR-31 decreases the amount of BAP1. 3 / 3
| 3

reach
"Inhibition of miR-31 expression enhanced the level of BAP1 in the two cell lines."

reach
"Increased expression of miR-31 reduced the BAP1 expression, leading to cell proliferation and suppressed apoptosis [XREF_BIBR]."

reach
"Thus, the 1.5-2 fold increases of miR-31 levels in lung cancer tissues are sufficient to reduce the expression of BAP1 protein in vivo."
MiR-31 decreases the amount of BAP1. 3 / 3
| 3

reach
"In this study, we identified BAP1 as a direct target gene of miR-31 and showed that miR-31 inhibited BAP1 expression."

reach
"We also detect the mRNA and protein expression level of BAP1, tumors with BAP1 plasmid transfected group exhibited higher BAP1 mRNA and protein expression compared with the control group, suggesting that BAP1 plasmid overexpression could rescue the expression of BAP1 suppressed by miR-31."

reach
"As anticipated, the expression of the BAP1 protein was significantly reduced by the introduction of miR-31, whereas the miR-31 inhibitor significantly increased the BAP1 protein levels in A549 cells."
MiR-31 inhibits BAP1.
| 3
MiR-31 inhibits BAP1. 3 / 3
| 3

reach
"Finally, the cells transfected with miR-31 and the BAP1 overexpression plasmid exhibited significantly lower proliferation rates compared with the cells transfected with miR-31 and the control plasmid, suggesting that miR-31-resistant BAP1 could rescue the suppression of BAP1 by miR-31 and attenuate the proliferative effect of miR-31."

reach
"BAP1 suppresses lung cancer progression and is inhibited by miR-31."

reach
"Tumors with both miR-31 and BAP1 overexpression exhibited significantly higher levels of BAP1 compared to tumors with miR-31 overexpression, suggesting that BAP1 overexpression could rescue the BAP1 suppression caused by miR-31."
MiR-31 binds BAP1.
| 2
BAP1 binds miR-31. 2 / 2
| 2

reach
"The aim of this study was to evaluate the interaction between BAP1 and miR-31 in MPM and their prognostic role in MPM."

reach
"These results indicate that miR-31 binds to BAP1 3 '-UTR directly and negatively regulates BAP1 expression in cervical cancer cells."
TP53 affects BAP1
2 | 9
2 | 7

sparser
"Mutual exclusivity of genomic alterations was also shown for TP53:IDH1 , IDH1: Kirsten rat sarcoma virus ( KRAS ), TP53:BAP1 , and IDH1:FGFR2 ."

sparser
"They also reported a tendency toward mutual exclusivity between multiple genes including TP53:IDH1, IDH1:KRAS, TP53:BAP1, and IDH1:FGFR2."

sparser
"Concurrent mutations in BAP1 and TP53 were associated with adverse outcome, with similar effects observed for acquired alterations in either of the two genes alone ( xref )."

sparser
"Mutations in BAP1 ( q = 0.004) and TP53 ( q = 0.001) were associated with decreased CSS in a multivariable model; only TP53 ( q = 0.005) remained significant when SSIGN score was included."

sparser
"Only three of the total 39 interactions— NF2-TP53 , NF2-SMAD4 , and BAP1-TP53 —commonly occurred in all three networks; 12 interactions were shared by only two of the networks and 24 were unique to specific networks ( xref )."

sparser
"On analyses adjusted only for cohort, we found that mutations in BAP1 ( n = 1049 for analysis; hazard ratio [HR]: 2.10; 95% confidence interval [CI]: 1.44–3.04; q < 0.001) and TP53 ( n = 784 for analysis; HR: 2.63; 95% CI: 1.36–5.08; q = 0.028) were significantly associated with increased risk of death from cancer after adjustment for multiple comparisons using competing risks regression ( xref )."

No evidence text available

sparser
"Only BAP1 and TP53 were significantly associated with an inferior OS ( P  = 0.0002; P  = 0.0082)."

No evidence text available
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
MP46 affects BAP1
| 11
BAP1 binds MP46. 10 / 11
| 11

sparser
"As expected, neither WZB117 nor 6-AN had any significant effect on the MP46 and MP46-BAP1 growth ( xref )."

sparser
"We then compared FAO-oxygen consumption rate (OCR) levels between MP46 and MP46-BAP1 by Seahorse analysis."

sparser
"First, we measured FA utilization levels between MP46 and MP46-BAP1 by incubating cells with palmitate (PAL) as the only oxidizable energy substrate."

sparser
"We did not observe significant differences in glycolysis, glycolytic capacity or OXPHOS parameters between MP46 and MP46-BAP1 ( xref ), although SRC levels were lower in MP46 than MP46-BAP1 ( xref )."

sparser
"Following PAL treatment, OCR levels increased significantly in MP46, but were unchanged in MP46-BAP1 ( xref ), suggesting that MP46 utilized or metabolized FA more efficiently than MP46-BAP1."

sparser
"We also tested for FAO-dependent OCR between MP46 and MP46-BAP1 by treatment with the FAO inhibitor, perhexiline (PHX)."

sparser
"Next, we treated MP46 and MP46-BAP1 with IACS or ETO."

sparser
"In comparison to MP46-BAP1, the predominant (56%) metabolites increased in MP46 were related to nucleotide biosynthesis pathways, while most metabolite decreases were associated with amino acid and FA metabolism ( xref and xref )."

sparser
"We found that MP65 (OXPHOS high ) were more resistant to both metabolic stress environments than MP65-BAP1, whereas MP46 (OXPHOS low ) were more vulnerable to stressful conditions compared to MP46-BAP1 ( xref and xref )."

sparser
"We further investigated the metabolite levels related to FA metabolism and observed MP46 had significantly lower levels of FAs and palmitoylcarnitine, which is involved in FA transport into mitochondria for oxidation compared to MP46-BAP1, but higher acetyl-carnitine levels, a byproduct of FAO ( xref )."
| 11
| 7

reach
"Moreover, BAP1 deficiency drives the reprogramming of cell metabolism, promoting anaerobic glycolysis for energy production rather than mitochondrial respiration and increasing extracellular lactate secretion which induces immune evasion, tumour growth and cell malignant transformation.BAP1 emerges as a highly tissue-specific and context-specific tumour suppressor participating to the biology of the tumour with multiple mechanisms and different levels (summary in Table 1)."

reach
"This resulted in a reduced list revealing that BAP1-downregulated genes were enriched in cell metabolism-associated genes."

reach
"In this study, we found that overexpression of miR-31-5p or inhibition of BAP1 expression decreased the levels of intracellular ROS, Fe , and iron metabolism markers, while the GSH content was increased."

reach
"Diminished Pgc1alpha signaling likely contributes to some of the observed phenotypes in Bap1ko mice, but Bap1 likely modulates metabolism by additional means."

reach
"A reduction in BAP1 protein levels supports the reprogramming of cancer cell metabolism, with a switch from TCA/OXPHOS to aerobic glycolysis [140]."

reach
"Defects in BAP1 disrupt DNA replication, DNA repair, calcium homeostasis, cell proliferation and differentiation, and cell metabolism [34]."

reach
"Gene ontology (GO) analysis of 354 BAP1 upregulated and 187 BAP1 downregulated genes revealed that, while BAP1 upregulated genes were enriched in diverse cellular processes (XREF_SUPPLEMENTARY and XREF_SUPPLEMENTARY), the genes that were downregulated by BAP1 showed striking enrichment in metabolism related biological processes, among which " response to oxidative stress " was the most significantly enriched (XREF_FIG and XREF_SUPPLEMENTARY)."
BAP1 activates metabolic process.
| 4

reach
"BAP1 expression mediates cell metabolism."

reach
"As a deubiquitinating enzyme, BAP1 contributes to gene transcription, cell differentiation, DNA damage repair, apoptosis, and cell metabolism in tumor inhibition."

reach
"BAP1 Modulates Cellular Metabolism."

reach
"In non-tumoral cells, reduced BAP1 levels induced the Warburg effect, a shift of cell metabolism from oxidative phosphorylation (Krebs cycle) to aerobic glycolysis, originally identified as typical of cancer cells."
| 11
| 7
Mutated BAP1 activates glycolytic process. 4 / 4
| 4

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"For example, mesothelioma cells that carry germline BAP1 mutations have upregulated aerobic glycolysis (17), and BAP1 mutant ccRCC tumor samples display elevated glycolysis (22)."

reach
"These data suggest that BAP1 mutant UM cells have increased aerobic glycolysis compared to BAP1 wild type UM cells."

reach
"In melanoma, BAP1 mutations promote tumor cell glycolysis, leading to increased lactate production."

reach
"BAP1 mutations cause a switch from oxidative phosphorylation to aerobic glycolysis [52]."

reach
"Intact BAP1 catalytic activity could increase aerobic glycolysis and lactate secretion and reduce mitochondrial respiration and ATP production [49]."

reach
"In non-tumoral cells, reduced BAP1 levels induced the Warburg effect, a shift of cell metabolism from oxidative phosphorylation (Krebs cycle) to aerobic glycolysis, originally identified as typical of cancer cells."

reach
"Moreover, BAP1 fibroblasts enhance aerobic glycolysis and lactate secretion, but reduce mitochondrial respiration and ATP production, when compared with wild-type BAP1."
| 4

reach
"XREF_FIG shows that BAP1 +/- fibroblasts transduced with AdBAP1 had reduced glycolysis and glycolytic capacity; in contrast, AdC91S was ineffective at reducing glycolysis (rate of glycolysis in BAP1 +/- transduced with : AdGFP 24.94 +/-1.77 mpH and min, AdBAP1 6.92 +/-0.24 mpH and min, AdBAP1 (C91S) 20.33 +/-2.92 mpH and min; glycolytic capacity : AdGFP 30.33 +/-1.71 mpH and min, AdBAP1 11.50 +/-0.43 mpH and min, AdC91S 25.08 +/-3.25 mpH and min)."

reach
"The oxygen-dependent mechanisms that cause HIF-1α degradation and the genes that suppress HIF-1α in hypoxia have been studied in detail (10, 34), while the gene products that by facilitating HIF-1α expression and activity in hypoxia influence tumor invasion and metastases, remain largely unknown.We reported that reduced BAP1 levels increase aerobic glycolysis (5)."

reach
"In melanoma, BAP1 deficiency significantly elevated anaerobic glycolysis levels, leading to more lactate production in tumor cells [11]."

reach
"Additionally, reduced levels of BAP1 induce aerobic glycolysis, which favors tumor growth (64)."
| 11
| 11

reach
"As shown in XREF_FIG and quantified in XREF_FIG, Bap1 increased surface adhesion of wild-type V. cholerae, a Deltabap1DeltarbmC mutant, and a DeltavpsL mutant in monolayer minimal medium, while a control protein, bovine serum albumin (BSA), had no effect."

reach
"This suggests that communal Bap1 secreted by nearby biofilm cells may also increase surface adhesion of bystanders that have not yet been reprogrammed for biofilm matrix synthesis."

reach
"Exogenous Bap1 increases surface adhesion of planktonic bystanders as well."

reach
"The loss of function in the defective mutants is unlikely due to changes in the production or secretion level of the mutant protein (Supplementary Fig. 4a–d), and in all defective strains the adhesion defects can be rescued by WT Bap1/RbmC expressed from a plasmid (Supplementary Fig. 4e)."

reach
"Two of the proteins, RbmC and Bap1, are seemingly redundant paralogs that coat substrate surfaces and promotes cell adhesion, while another protein RbmA coats cell surfaces and promotes cell-cell cohesion."

reach
"Among the accessory proteins, RbmA has been shown to adhere biofilm-residing cells to each other , whereas Bap1 and RbmC contribute to cell-to-surface adhesion as well as to the formation of envelope structures surrounding Vc biofilm clusters ."

reach
"BAP1 loss reduced HAP1 cell adhesion/spreading (Fig. 1D) and proliferation (Fig. 1E), considered here to be readouts of cell fitness."

reach
"Bulk adhesion measurements for Bap1 and RbmC, which showed that Bap1 and RbmC contribute to surface adhesion in a surface-dependent manner, are largely consistent with our findings (31)."

reach
"Because exogenously provided Bap1 restored biofilm surface adhesion to a Deltabap1DeltarbmC mutant, we questioned whether Bap1 synthesis could be a joint venture in the biofilm community."

reach
"Bap1 mediates adhesion of bystander cells."
BAP1 affects PARP1
6 | 2 2
6 | 2 2

No evidence text available

No evidence text available

No evidence text available

reach
"PARylation was not required for the interaction between BAP1 and PARP1 since treatment with PARP inhibitors did not interfere with this interaction ."

sparser
"PARylation was not required for the interaction between BAP1 and PARP1 since treatment with PARP inhibitors did not interfere with this interaction xref ."

No evidence text available

reach
"Poly(ADP-ribose) polymerase 1 (PARP1) interacts with and recruits BAP1 to damage sites, with BAP1 recruitment peaking after the DDB2 and XPC damage sensors."

No evidence text available

No evidence text available

sparser
"PARP1 interacted with BAP1 after UV irradiation, and depletion of PARP1 or treatment with PARP inhibitors reduced BAP1 chromatin binding and BAP1 colocalization with CPDs."
BAP1 affects MP46
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BAP1 binds MP46. 10 / 11
| 11

sparser
"As expected, neither WZB117 nor 6-AN had any significant effect on the MP46 and MP46-BAP1 growth ( xref )."

sparser
"We then compared FAO-oxygen consumption rate (OCR) levels between MP46 and MP46-BAP1 by Seahorse analysis."

sparser
"First, we measured FA utilization levels between MP46 and MP46-BAP1 by incubating cells with palmitate (PAL) as the only oxidizable energy substrate."

sparser
"We did not observe significant differences in glycolysis, glycolytic capacity or OXPHOS parameters between MP46 and MP46-BAP1 ( xref ), although SRC levels were lower in MP46 than MP46-BAP1 ( xref )."

sparser
"Following PAL treatment, OCR levels increased significantly in MP46, but were unchanged in MP46-BAP1 ( xref ), suggesting that MP46 utilized or metabolized FA more efficiently than MP46-BAP1."

sparser
"We also tested for FAO-dependent OCR between MP46 and MP46-BAP1 by treatment with the FAO inhibitor, perhexiline (PHX)."

sparser
"Next, we treated MP46 and MP46-BAP1 with IACS or ETO."

sparser
"In comparison to MP46-BAP1, the predominant (56%) metabolites increased in MP46 were related to nucleotide biosynthesis pathways, while most metabolite decreases were associated with amino acid and FA metabolism ( xref and xref )."

sparser
"We found that MP65 (OXPHOS high ) were more resistant to both metabolic stress environments than MP65-BAP1, whereas MP46 (OXPHOS low ) were more vulnerable to stressful conditions compared to MP46-BAP1 ( xref and xref )."

sparser
"We further investigated the metabolite levels related to FA metabolism and observed MP46 had significantly lower levels of FAs and palmitoylcarnitine, which is involved in FA transport into mitochondria for oxidation compared to MP46-BAP1, but higher acetyl-carnitine levels, a byproduct of FAO ( xref )."
BAP1 affects ERK
| 9 2
BAP1 inhibits ERK.
| 4
BAP1 inhibits ERK. 4 / 4
| 4

reach
"Taken together, our data suggested that BAP1 negatively regulated the ERK1/2 and JNK/c-Jun signaling pathways to exert its tumor-suppressive functions in ICC."

reach
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways."

reach
"The observed inverse relationship between BAP1 expression and ERK1/2 polyubiquitination suggests that BAP1 inhibits the ERK signaling pathway by deubiquitinating the K63-linked polyubiquitin of ERK1/2."

reach
"BAP1 restricted the growth cycle of intrahepatic cholangiocarcinoma by regulating ERK1/2 and JNK/c-Jun axis (Chen et al. 2018)."
BAP1 dephosphorylates ERK.
| 4
BAP1 leads to the dephosphorylation of ERK. 4 / 4
| 4

reach
"Our study of the signaling pathways revealed that BAP1 depletion induced ERK1/2 phosphorylation without affecting other upstream signaling molecules (Fig. 3A, B)."

reach
"BAP1 depletion in MSCs upregulates ERK phosphorylation, thereby increasing the expression of the migration factor, osteopontin."

reach
"More importantly, our results also confirmed that BAP1 deficiency increased the phosphorylation and activity of ERK1/2 and JNK/c-Jun signaling to promote ICC progression."

reach
"BAP1 depletion increased ERK1/2 phosphorylation, indicating a negative regulatory effect of BAP1 on ERK1/2 activation.Recently, it was discovered that the K63-linked polyubiquitination of ERK1/2 is closely related to the phosphorylation status of ERK1/2 (28)."
BAP1 binds ERK.
| 1 2
| 1 2

sparser
"Further examination revealed that BAP1 interacts with phosphorylated ERK1/2, deubiquitinating their ubiquitins, and thus attenuating the ERK signaling pathway."

sparser
"Collectively, these results suggest that the binding of BAP1 to phosphorylated-ERK1/2 may lead to a reduction in its phosphorylation."

reach
"Surprisingly, we found that neither ERK1/2 nor any other signaling molecules upstream of ERK bound to BAP1; only phosphorylated-ERK1/2 co-precipitated with endogenous BAP1 (Fig. 4B)."
TG2-179-1 affects BAP1
| 10
TG2-179-1 inhibits BAP1.
| 7
TG2-179-1 inhibits BAP1. 7 / 7
| 7

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"TG2-179-1 inhibits the DUB activity of BAP1 by binding to its active site."

reach
"We then determined whether TG2-179-1 inhibits BAP1 using purified recombinant BAP1 proteins and the fluorogenic artificial DUB substrate Ub-AMC as a BAP1 substrate."

reach
"In support of this prediction, TG2-179-1 inhibited BAP1 even after immunoprecipitation of TG2-179-1-bound BAP1 with extensive washing."

reach
"Incubation of BAP1 with increasing concentrations of TG2-179-1 reduced the emission of fluorescence from Ub-AMC in a dose-dependent manner (Fig. 5C), indicating that TG2-179-1 inhibited the DUB activity of BAP1."

reach
"TG2-179-1 inhibited the DUB activity of BAP1 toward nucleosomal H2A-Ub substrates in a dose-dependent manner (Fig. 5D)."

reach
"Having verified that TG2-179-1 inhibits BAP1, we investigated its impact on the viability of colon cancer cells."

reach
"However, TG2-179-1 kills both BAP1-insensitive and BAP1-depleted colon cancer cells as well as BAP1-null mesothelioma and ccRCC cells, indicating that TG2-179-1 targets BAP1 as well as other cellular proteins."
TG2-179-1 binds BAP1.
| 3
BAP1 binds TG2-179-1. 3 / 3
| 3

reach
"We therefore propose that the covalent interaction between BAP1 and TG2-179-1 is reversible."

reach
"These data clearly demonstrated that TG2-179-1 inhibits the DUB activity of BAP1.Analysis of the structural model of the BAP1TG2-179-1 complex using the CavityPlus server showed that Cys91 was predicted to form a covalent bond with an electron-deficient atom in TG2-179-1 ."

reach
"Our in silico docking model predicted that TG2-179-1 binds to BAP1 with its fluorobenzene and thiolane rings deeply embedded into the active site pocket, which sterically allows TG2-179-1 to interact covalently with Cys91 in the active site via its electrophilic nitrile carbon atom."
STK11 affects BAP1
5 | 2 3
5 | 2 3

No evidence text available

reach
"BAP1 deficiency enhanced cancer cell proliferation and lipid synthesis via the LKB1-AMPK-mTOR signalling pathway.In conclusion, we suggest that BAP1 interacts with and deubiquitinates LKB1, thereby af[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The positive colony could survive in SD-4 (-Ade, -His, -Leu, -Trp) medium (Clontech) developing a blue colour in the presence of X-Gal (Sigma), thereby indicating an interaction between BAP1 and LKB1."

sparser
"These results validated a direct interaction between BAP1 and LKB1."

sparser
"We demonstrated a direct interaction between LKB1 and BAP1 ( Fig. 1 E)."

No evidence text available

No evidence text available

No evidence text available

No evidence text available

reach
"These results validated a direct interaction between BAP1 and LKB1."
HIF1 affects BAP1
| 10
| 10

reach
"Co-immunoprecipitation (CoIP) and proximity ligation assay (PLA) experiments revealed that 1) HIF-1α and BAP1 bind to each other and co-precipitate (Fig. 2A), and 2) the nuclei of BAP1 cells contained significantly more PLA positive signals—evidence of BAP1 and HIF-1α interaction—than BAP1 cells (Fig. 2 B and C)."

reach
"The crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) (49) shows that without BAP1, HIF-1α, and HIF-1β bind to DNA (49), thus BAP1 is not required for HIF-1α-HIF-1β complex formation."

reach
"Aligning the crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) (49) to our structural model for the binding complex of BAP1-HIF-1α showed that both BAP1 and HIF-1β bind to the same residues of HIF-1α (1-73) on the DNA; however, in Fig. 3B, we demonstrate that BAP1, HIF-1α and the DNA form a complex without HIF-1β."

reach
"Therefore, our data indicate that BAP1 is not required for HIF-1α-HIF-1β complex formation to functionally bind to DNA, that HIF-1β is not required for BAP1-HIF-1α complex formation to functional binding to DNA, and that although DNA facilitates the binding of BAP1 and HIF-1α, it is not required to maintain the binding of both BAP1-HIF-1α and BAP1-HIF-1β."

reach
"In summary, our data suggest that BAP1 directly binds and stabilizes both HIF-1α and HIF-1β increasing their intra-nuclear availability for dimer formation, thus fine-tuning HIF activities required to support malignant cell growth."

reach
"So far, the pathogenic variants reported in ClinVar for both HIF-1α and HIF-1β are not located among the crucial residues of HIF-1α-r73 where HIF-1α can bind to BAP1, HIF-1β and DNA, or of HIF-1β (2-470) where HIF-1β can bind to BAP1, HIF-1α, and DNA."

reach
"We further investigated the specificity of the BAP1 interaction with HIF-1α in HEK-293 cells expressing Myc-BAP1 and HA-HIF-1α, using HA-Tag as bait."

reach
"To study the binding between BAP1 and HIF-1α, first, a rigid docking protocol was applied to model the binding complex of the CTD of BAP1 (the NLS domain is removed due to its flexibility) and HIF-1α by using the ClusPro server (43–45) (SI Appendix, Fig. S2B)."

reach
"Notably, BAP1 binding to HIF-1α-r73-DNA does not require HIF-1β for the interaction."

reach
"We tested the hypothesis that although DNA facilitates the binding between BAP1 and HIF-1α, this binding complex still holds in the absence of DNA."
| 10
BAP1 inhibits immune response.
| 6

reach
"These results indicate that BAP1 inactivation in MPM correlates with an increase in DCs and inflammatory macrophages and a decrease in immunosuppressive MDSCs, further supporting the notion that BAP1 deficiency promotes the immune response in MPM."

reach
"We fortuitously found that deletion of Bap1 in cancer cells can trigger an anti-tumor immunity, regardless of the expression of PD-L1."

reach
"Loss of histone deubiquitinase Bap1 triggers anti-tumor immunity."

reach
"We found that loss of histone deubiquitinase Bap1 in cancer cells activates a cDC1-CD8 + T cell-dependent anti-tumor immunity."

reach
"Consequently, BAP1 loss may promote metastasis at least in part by allowing tumor cells to evade the patient’s immune response."

reach
"To understand how the loss of Bap1 activates anti-tumor immunity, we conducted a comparison of the immune subsets present in the tumor microenvironment of Bap1-KO and control WT (MC38-Cas9) tumors."
BAP1 activates immune response.
| 4

reach
"However, beyond this experimental model, it is important to consider that BAP1 activities in other cell types may also contribute to the regulation of antibody mediated immunity in vivo."

reach
"These results indicate that BAP1 inactivation in MPM correlates with an increase in DCs and inflammatory macrophages and a decrease in immunosuppressive MDSCs, further supporting the notion that BAP1 deficiency promotes the immune response in MPM."

reach
"Here, our molecular profiling data revealed that BAP1-deficient MPM is enriched for immune-related pathways (e.g., IFN-α/γ response) and strongly correlates with the gene signature of DCs, a professional antigen-presenting cell type that induces the activation and differentiation of naive T lymphocytes,35 suggesting that BAP1 contributes to antitumor immunity in MPM beyond its function intrinsic to cancer cells.First, we observed that BAP1-deficient MPM exhibited increased expression of several inhibitory immune checkpoints, such as PD-L1, PD-1, and LAG3."

reach
"B-cell intrinsic regulation of antibody mediated immunity by histone H2A deubiquitinase BAP1."
BAP1 affects PRC2_complex
| 6 4
BAP1 inhibits PRC2_complex.
| 6
| 6

reach
"BAP1 loss induces PRC2-mediated transcriptional repression in mouse and human mesothelioma."

reach
"It has been proposed that the loss of BAP1 promotes PRC2 activity and that the cells that lack BAP1 are sensitive to EZH2 inhibition."

reach
"In mouse models, the loss of BAP1 leads to elevated enhancer of zeste 2 polycomb repressive complex 2 subunit (EZH2) expression and enhanced repression of polycomb repressive complex 2 (PRC2) targets [48]."

reach
"In MPM cell lines, BAP1 loss was observed to be related to increased EZH2 expression and enhanced repression of PRC2-regulated targets."

reach
"As for the reported tumor suppressive mechanism, BAP1 can antagonize EZH2 and PRC2 or RING1B (RNF2)/PRC1 in a tissue specific manner."

reach
"Loss of BAP1 has been shown to be synthetically lethal with inhibition of EZH2 or PRC2."
| 4

sparser
"Thus, our analysis and experimental validation across mouse and human mesothelioma cells clearly identify a set of genes regulated via the BAP1-PRC2 axis."

sparser
"BAP1 does not interact with PRC2 proteins ( xref )."

sparser
"ASXL1 functions in transcriptional repression through its interaction with PRC2 and BAP1. xref BCORL1 is a transcriptional co-repressor that interacts with PCGF1, a core complex of the PRC1.1 complex. xref The RUNX1-CBF-b heterodimer mediates transcription by binding to RUNX sites, but also represses transcription by interacting and recruiting BMI1 of the PRC1 complex to target sites. xref IKZF1 regulates transcription by interacting with repressive epigenetic complexes such as HDAC1, HDAC2, CHD3, CHD4, and SWI/SNF complex, and also recruits PRC2 to target gene loci in T cells. xref Thus, while the commonly mutated genes in CML have their own exclusive roles in transcriptional regulation, they also share a striking commonality as modulators of the PRC."

sparser
"In addition, BAP1 mutations in ccRCC tumors were associated with downregulated expression of target genes of polycomb repressive complex 2 (PRC2) [ xref ] ( xref )."
BAP1 affects H2AK119Ub
| 10
BAP1 deubiquitinates H2AK119Ub.
| 6
BAP1 deubiquitinates H2AK119Ub. 6 / 6
| 6

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"cBioPortal functional prediction and our structural, biochemical, and cellular studies indicate all these mutations lead to a loss of function where BAP1/ASXL1 fails to effectively deubiquitinate H2AK119Ub, and thus causes aberrant accumulation of this chromatin modification."

reach
"Our data support a model where BAP1/ASXL1 deubiquitinates H2AK119Ub chromatin by anchoring to the nucleosomal acidic patch, DNA dyad, and the DNA exit regions using conserved arginine and lysine tracts."

reach
"These results further provide a molecular explanation for how >50 mutations in BAP1 and ASXL1 found in cancer can dysregulate H2AK119Ub deubiquitination, providing insight into understanding cancer etiology."

reach
"BAP1 deubiquitinates H2AK119Ub, regulating chromatin packing and gene expression (Scheuermann et al., 2010; Foglizzo et al., 2018)."

reach
"Our data support a model where BAP1/ASXL1 deubiquitinates H2AK119Ub chromatin by anchoring to the nucleosomal acidic patch, DNA dyad, and the DNA exit regions using conserved arginine and lysine tracts."

reach
"As is the case with the loss-of-function model, these data suggest that mutant ASXL1 inhibits PRC2 function, followed by upregulation of its target genes in a dominant-negative manner, which contribut[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 binds H2AK119Ub.
| 4
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

reach
"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

reach
"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

reach
"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

reach
"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
OmpT affects BAP1
| 3 6
| 3 6

sparser
"We investigated Bap1 binding to OmpT, a major outer membrane porin of V. cholerae containing an integrin-binding domain (LDV peptide)."

sparser
"In presence of PmB, Bap1 was not able to bind to the OmpT protein from the Δ ompT /pBR- ompT * (LDV mutant) mutant to the same extent as to the wild type OmpT protein ( xref ), suggesting a role for the LDV domain in Bap1 binding to OmpT on OMVs ( xref )."

sparser
"Taken together, our results strongly support the hypothesis that Bap1 binds to OmpT on OMVs of V. cholerae A1552 through the LDV tripeptide integrin-binding domain of OmpT."

sparser
"Bap1-OmpT interaction requires the presence of the integrin-binding domain (leucine-aspartic acid-valine LDV peptide) of OmpT and the FG-GAP domains of Bap1 ( xref )."

reach
"It has been reported that Bap1 could associate with outer membrane vesicles (OMVs) by binding to OmpT."

reach
"We investigated Bap1 binding to OmpT, a major outer membrane porin of V. cholerae containing an integrin binding domain (LDV peptide)."

reach
"Taken together, our results strongly support the hypothesis that Bap1 binds to OmpT on OMVs of V. cholerae A1552 through the LDV tripeptide integrin binding domain of OmpT."

sparser
"It was also reported that RbmC does not interact with OmpT in the OMVs, although it is currently unclear why Bap1, but not RbmC, binds to OmpT."

sparser
"Under polymyxin B-induced stress conditions, Vibrio cholerae O1 El Tor produces BMVs containing the protein OmpT that binds the secreted protein Bap1, which in turn binds the AMP LL-37 ( xref )."
KEAP1 affects BAP1
1 2 | 1 5
1 2 | 1 5

sparser
"Alternatively, it can be speculated that BAP1 might competitively bind to KEAP1 at the BTB domain with unknown protein(s) affecting the stability of KEAP1."

sparser
"In order to identify the domains of BAP1 that interact with KEAP1, we generated FLAG-BAP1 mutant plasmids lacking individual domains (FLAG-BAP1ΔCD, FLAG-BAP1ΔCC1, FLAG-BAP1ΔCL, FLAG-BAP1ΔCC2, FLAG-BAP1ΔULD, and FLAG-BAP1ΔCTE)."

reach
"Tumor suppressors, such as p53, BAP1, Kelch-like ECH associated protein 1 (KEAP1), function through induction of ferroptosis mainly via regulating SLC7A11 expression [4], [5], [6]."

sparser
"Immunoprecipitation of endogenous BAP1 demonstrates that BAP1 interacts with endogenous KEAP1 ( xref D)."

sparser
"In order to identify the domain of KEAP1 that interacts with BAP1, we generated pcDNA3-HA-KEAP1 mutant plasmids lacking individual domains (HA-KEAP1ΔBTB, HA-KEAP1ΔBACK, and HA-KEAP1ΔKELCH)."

"BAP1 directly binds to and deubiquitinates KEAP1. BAP1 stabilizes KEAP1 by binding to the BTB domain."

sparser
"Our result shows that FLAG-BAP1 does not selectively bind to HA-KEAP1ΔBTB ( xref A, right panel), suggesting that BAP1 binds to the BTB domain of KEAP1."

No evidence text available

No evidence text available
BAP1 affects ompT
| 3 6
| 3 6

sparser
"We investigated Bap1 binding to OmpT, a major outer membrane porin of V. cholerae containing an integrin-binding domain (LDV peptide)."

sparser
"In presence of PmB, Bap1 was not able to bind to the OmpT protein from the Δ ompT /pBR- ompT * (LDV mutant) mutant to the same extent as to the wild type OmpT protein ( xref ), suggesting a role for the LDV domain in Bap1 binding to OmpT on OMVs ( xref )."

sparser
"Taken together, our results strongly support the hypothesis that Bap1 binds to OmpT on OMVs of V. cholerae A1552 through the LDV tripeptide integrin-binding domain of OmpT."

sparser
"Bap1-OmpT interaction requires the presence of the integrin-binding domain (leucine-aspartic acid-valine LDV peptide) of OmpT and the FG-GAP domains of Bap1 ( xref )."

reach
"It has been reported that Bap1 could associate with outer membrane vesicles (OMVs) by binding to OmpT."

reach
"We investigated Bap1 binding to OmpT, a major outer membrane porin of V. cholerae containing an integrin binding domain (LDV peptide)."

reach
"Taken together, our results strongly support the hypothesis that Bap1 binds to OmpT on OMVs of V. cholerae A1552 through the LDV tripeptide integrin binding domain of OmpT."

sparser
"It was also reported that RbmC does not interact with OmpT in the OMVs, although it is currently unclear why Bap1, but not RbmC, binds to OmpT."

sparser
"Under polymyxin B-induced stress conditions, Vibrio cholerae O1 El Tor produces BMVs containing the protein OmpT that binds the secreted protein Bap1, which in turn binds the AMP LL-37 ( xref )."
| 9

reach
"Through a biochemical screen, we identified BAP1 catalytic inhibitors that inhibit truncated-ASXL1-driven leukemic gene expression and impair tumor progression in vivo."

reach
"In the prevailing model, nuclear BAP1 activates gene expression by deubiquitinating H2AK119Ub and safeguards Polycomb-mediated repression by removing any misplaced H2AK119Ub (5-8)."

reach
"In this study, we identified a key signaling pathway involving the histone H2AK119 deubiquitinase BRCA1 associated protein 1 (BAP1), the interferon regulatory factor interferon regulatory factor 1 (IRF1), and the MHC-II transactivator class II transactivator (CIITA), which directly activates MHC-II gene expression."

reach
"Meta-analyses have indicated that mutations in PBRM1 and SETD2 tend to occur together, but mutations in PBRM1 and BAP1 tend to be mutually exclusive and are thought to underlie different pathologic fe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Collectively, these results indicated that BAP1 may directly regulate MHC-II cluster gene expression, and this regulatory effect is independent of cell types."

reach
"Based on our independent analysis, BAP1 deficiency in myeloid progenitors led to a dramatic increase of H3K27me3 levels at the MHC-II loci (Supplemental Figure 2G), and a substantial reduction in MHC-II gene expression (Supplemental Figure 2, H and I), which is consistent with our conclusions."

reach
"Although BAP1 can directly regulate gene expression, this pattern suggests critical epigenetic roles for BAP1 during ENS cell lineage specification."

reach
"Furthermore, our findings that BaP1 induced gene expression of COX-2 and mPGES-1 demonstrate that BaP1 can trigger transcriptional mechanisms in FLSs that result in increased protein expression."

reach
"BAP1 deubiquitinating enzymatic activity is an important modulator of gene expression."

reach
"In addition to a potentially critical role for BAP1 inactivation in epigenetic deregulation in ccRCC, BAP1 has been found to promote DNA double-strand break repair by homologous recombination, thereby adding to its tumor suppressor function [XREF_BIBR]."

reach
"Although BAP1 stimulates DNA double-strand break repair, whether it functions in nucleotide excision repair (NER) is unknown."

reach
"BAP1 protein was reported to form a complex with host cell factor-1 (HCF-1) and promotes DNA double-strand break repair."

reach
"BAP1 loss impairs HR and double-strand break repair, promoting error-prone nonhomologous end joining, with consequent genomic instability (XREF_FIG)."

reach
"The literature suggests that BAP1 promotes DNA double-strand break repair and that its loss leads to homologous recombination repair defects, reminiscent of BRCA1/2 deficiency."

reach
"Tumor suppressor and deubiquitinase BAP1 promotes DNA double-strand break repair."

reach
"Tumor suppressor and deubiquitinase BAP1 promotes DNA double-strand break repair."

reach
"Tumor suppressor and deubiquitinase BAP1 promotes DNA double-strand break repair."

reach
"Tumor suppressor and deubiquitinase BAP1 promotes DNA double-strand break repair."
BAP1 affects angiogenesis
| 1 8
| 1 8

reach
"BaP1 Reduces Tumor Growth, Angiogenesis, and Tumor Proliferation."

reach
"Furthermore, the BaP1 treatments reduced the percentage of blood vessel (angiogenesis) around the tumor."

reach
"P-I metalloproteinases and L-amino acid oxidases from Bothrops species inhibit angiogenesis."

reach
"Furthermore, we observed that BaP1 decreased angiogenesis, characterized by a significant decrease in the vascularization formed around the tumors, as well as reduced tumor proliferation evidenced by the reduced levels of Ki67 expression in the tumors treated with the compound.Overall, our data highlights BaP1 as a molecule with a potent and selective anticancer activity against CRC cells and as a very interesting agent to disturb and counteract the important roles of lysosome in cancer."

reach
"In vivo results using a chicken embryo choriollantoic membrane (CAM) assay showed that BaP1 inhibits tumor growth, angiogenesis, and tumor proliferation."

reach
"P-I metalloproteinases and LAAO from Bothrops spp. inhibit angiogenesis ."

eidos
"In vivo results using a chicken embryo choriollantoic membrane ( CAM ) assay showed that BaP1 inhibits tumor growth , angiogenesis , and tumor proliferation ."

reach
"Furthermore, our in vivo results, using CAM assay, sustained the anticancer activity of BaP1, as it was found that BaP1 inhibits tumor growth, angiogenesis and tumor proliferation, relevant hallmarks of cancer."

reach
"This compound reduced the survival, proliferation, and migration of human colorectal cancer cells, and BaP1 blocked in vivo tumor growth and proliferation as well as angiogenesis."
BAP1 affects XRCC1
| 9
BAP1 inhibits XRCC1.
| 5
BAP1 inhibits XRCC1. 5 / 5
| 5

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"In contrast to BAP1, which inhibits RCC by upregulation of STING activity and activation of interferon, UCHL5 promotes RCC by regulating immune infiltration and antigen presentation of B cells, and its increased level in RCC blood samples suggest its potential as a prognostic marker."

reach
"BAP1-TPDS is associated with an increased risk of a specific skin lesion, BAP1-inactivated melanocytic tumour (BIMT) (34), cutaneous melanoma (CM) (34), renal cell carcinoma (RCC) (35), and basal cell carcinoma (BCC) (36)."

reach
"Truncation mutations in BAP1, disrupting its peptidase region, were the predominant characteristics of kidney cancer.Additionally, we discovered the underlying biochemical mechanism by which BAP1 suppresses RCC development."

reach
"In conclusion, our findings suggest that BAP1 suppresses RCC by regulating a series of biological processes, including the STING -involved IFN-I pathway."

reach
"BAP1 knockout in mice leads to MM growth, larger tumors and increased number of metastases and BAP1 protein expression is also correlated with poor prognosis and shorter survival in RCC and cholangiocarcinoma, as well as UM (22–25)."
BAP1 activates XRCC1.
| 4
BAP1 activates XRCC1. 4 / 4
| 4

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"Meta-analyses have indicated that mutations in PBRM1 and SETD2 tend to occur together, but mutations in PBRM1 and BAP1 tend to be mutually exclusive and are thought to underlie different pathologic fe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"While BAP1 deletion suppresses RCC cell survival in vitro, it does not impede tumor growth in immunocompetent murine models."

reach
"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (frequently in the abdomen) and uveal melanoma [151,152,153]."

reach
"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (often in the abdomen), and uveal melanoma [97]."
BAP1 affects PTGS2
| 9
BAP1 increases the amount of PTGS2. 9 / 9
| 9

reach
"Having shown that the inducible COX-2 isoform participates in the signaling pathway triggered by BaP1 that leads to PGE production, we next investigated whether BaP1 is able to induce protein expression of COX-2 in FLSs."

reach
"Accordingly, our data show that BaP1 induced a marked increase in protein expression of both COX-2 and mPGES-1, the increase in COX-2 expression occurring before that of mPGES-1."

reach
"Furthermore, our findings that BaP1 induced gene expression of COX-2 and mPGES-1 demonstrate that BaP1 can trigger transcriptional mechanisms in FLSs that result in increased protein expression."

reach
"We found that the EP4 receptor positively regulates BaP1-triggered PGE release and expression of COX-2 and mPGES-1, which are critically involved in the biosynthesis of PGE induced by this metalloprotease."

reach
"Intriguingly, BAP1 was unfolded to increase the COX-2 and mPGES-1 expression by activating the NF-κB pathway (Viana et al. 2020)."

reach
"In view of previous reports that IL-1β induces PGE production via up-regulation of COX-2 and PGESm-1 expression in several cell types, including synovial fibroblasts , it is plausible to suggest that, under the present experimental conditions, IL-1β up-regulates BaP1-stimulated production of PGE by inducing expression of COX-2 and/or mPGES-1 in FLSs."

reach
"Gene and protein expression of both COX-2 and mPGES-1 are induced by BaP1 via activation of distinct subunits of the NF-κB transcription factor."

reach
"Partially consistent with our results, BAP1 was unfolded to increase the COX-2 and mPGES-1 expression by activating the NF-κB signaling pathway and inducing the release of IL-1β (Viana et al. 2020), but this study failed to explore the regulatory role of BAP1 mutations on the NF-κB signaling pathway."

reach
"Expression of COX-2 and mPGES-1 are induced by BaP1 via activation of NF-kappaB pathway."
3p21.1 losses affects BAP1
| 9
3p21.1 losses activates BAP1. 9 / 9
| 9

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"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."

reach
"The nuclear deubiquitinase BAP1 is commonly inactivated by somatic mutations and 3p21.1 losses in malignant pleural mesothelioma."
P14_3_3 affects BAP1
| 8
| 8

sparser
"Next, we tested the hypothesis that the BAP1 interaction with 14-3-3 releases the binding between 14-3-3 and Bax, which further facilitates cell death."

sparser
"Another study in 2018, showed that BAP1 interacts with the protein 14-3-3 in neuroblastoma cells causing the release of Bax and the decrease of pro-survival factors such as Bcl-2, to promote apoptosis via the intrinsic pathway.[ xref ]"

sparser
"These results suggest that BAP1 interaction with 14-3-3 prevents cell survival signaling, which is essential for growth of the neuroblastoma cells (Fig.  xref )."

sparser
"BAP1 binding to 14-3-3 facilitate the apoptotic signaling in neuroblastoma."

sparser
"The interaction between BAP1 and 14-3-3 protein was confirmed by performing the co-IP of FLAG-tagged BAP1 and pull-down of endogenous 14-3-3 in neuroblastoma cells (Fig.  xref )."

sparser
"The reciprocal IP using 14-3-3 antibodies also indicated the formation of the complex between BAP1 and 14-3-3 (Fig.  xref )."

sparser
"In addition, fluorescence resonance energy transfer (FRET) assay demonstrated a direct interaction between BAP1 and 14-3-3 (Fig.  xref )."

sparser
"We could also confirm that the short fragment of BAP1-ULD domain and 14-3-3-alpha helices 7-9 regions facilitate interaction between these two proteins (Fig.  xref ), while the interaction between BAP1 and 14-3-3-σ inactive mutants were reduced compared to the full-length 14-3-3-σ (Fig.  xref )."
TMED2 affects BAP1
| 8
| 8

sparser
"Validation of the BAP1 COPI interaction."
| PMC

sparser
"In order to confirm the interaction between BAP1 and the heptameric COPI complex we performed AP-immunoblot experiments ( xref )."
| PMC

sparser
"To exclude that the BAP1 interaction with the COPI complex is an artifact of overexpression of the exogenous BAP1 construct, we immunoprecipitated endogenous BAP1 using different antibodies against BAP1 in cell lysate from HeLa FRT parental cells ( xref )."
| PMC

sparser
"AP-immunoblotting of GFP-COPE shows interaction of COPA as expected, however BAP1 cannot be observed which is probably due to the low levels of BAP1 that bind to the COPI complex ( xref )."
| PMC

sparser
"Although the absolute and relative amounts of COPI that interacts with BAP1 in the AP-MS is low ( xref and xref ), the interaction can still be biologically relevant and is validated with endogenous protein as seen in the immunoblot experiments ( xref )."
| PMC

sparser
"It was intriguing that BAP1 has such a sequence in its C-terminus, but our mutational analysis suggests that the interaction between BAP1 and the COPI complex is not mediated through the C-terminal KxKxx sequence in BAP1."
| PMC

sparser
"We must be absolutely sure that the BAP1-COPI interaction is not an artifact."
| PMC

sparser
"We agree with reviewer 2 that the interaction between BAP1 and the COPI complex may be a substrate interaction as evidenced by the low protein stoichiometry observed in the affinity purification (Fig 5)."
| PMC
Neoplasms affects BAP1
| 8
| 8

reach
"Subsequent experiments by Ventii et al. (2008) suggested tumor growth suppression of BAP1-restored lung cancers in animal models."

reach
"Xenografted Ren-02 BAP1 knockdown tumors were excised from mice treated with vehicle (40% PEG-400, 0.9% NaCl, 10% DMSO, filtered) or diABZI (3 mg/kg) twice weekly for 3.5‒5 weeks via intraperitoneal injection."

reach
"The sPD-L1 level in PE of MPM was lower in BAP1 loss tumors (208.0 ± 167.4 pg/ml) than in BAP1-retained tumors (338.7 ± 457.4 pg/ml, p = 0.085), although the difference was not significant (Supplementary Fig. S4a)."

reach
"TGF-β in PE of MPM was significantly lower in cases with BAP1 loss tumors (9218 ± 6776 pg/ml) than in cases with BAP1-retained tumors (15,474 ± 15,944 pg/ml, p = 0.023)."

reach
"However, a limitation of our study is that eye pigmentation is deduced from genotypes, which are also risk SNPs for UM, making it challenging to derive causal statements.Status of chromosome 3 and BAP1 delineates 2 UM subtypes, M3 and BAP1-inactivated high-risk tumors and D3 and wild-type BAP1 low-risk tumors (2-4,8)."

reach
"Besides, several other transcription factors or tumor suppressors, such as activating transcription factor 3 (ATF3), BRCA1 associated protein-1 (BAP1), ADP-ribosylation factor (ARF), and Kelch-like ECH-associated protein 1 (KEAP1), diminish the expression or suppress the activity of SLC7A11 to induce ferroptosis in cancer cells (Fan et al. 2017; Roh et al. 2017; Wang et al. 2020; Zhang et al. 2018b)."

reach
"The compound iBAP (36) inhibited truncated-ASXL1-driven leukemic gene expression and hindered tumor progression in vivo , which led to small-molecular catalytic inhibitors of BAP1 as a potential anti-[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Moreover, fibrotic tumors with histological features of both HCC and FLC have been observed in patients with mutations inactivating BAP1, with a chromosomal gain of PRKACA or loss of PRKAR2A (another inhibitory regulatory subunit) ."
MLL2 affects BAP1
| 8
MLL2 binds BAP1. 8 / 8
| 8

sparser
"The MLL4 and BAP1 co-occupancy depends on ASXL proteins that directly interact with both MLL4 and BAP1 and bring together two enzymatic activities to specific genomic loci."

sparser
"MLL4 occupancy on MLL4 + BAP1 + genomic regions depends on functional ASXLs that interact with both MLL4 and BAP1, and CRC-relevant mutations attenuate the formation of the MLL4-ASXL complex."

sparser
"MBH of ASXL1/2 is required for the association of BAP1 with MLL4 in ESCs."

sparser
"Although ASXLs have been shown to form complexes with both the deubiquitinase BAP1 and the methyltransferase MLL4(12,15), the importance of ASXLs in genomic distribution of BAP1 and MLL4 in CRC remains unclear."

sparser
"These results indicated that BAP1 associates with MLL4 in ESCs in an ASXL1/2 MBH-dependent manner."

sparser
"To assess whether ASXLs play a role in regulating MLL4 and BAP1 binding at specific chromatin sites, we generated MLL4-, ASXL2-or BAP1-deficient human colon cancer HCT116 cells using the CRISPR/Cas9 approach."

sparser
"Interdependent genomic binding of MLL4 and BAP1 on enhancers."

sparser
"The fact that MLL4 and BAP1 do not directly interact and are linked through their concomitant binding to ASXLs suggested a possible role of ASXLs in this regulation."
MBD6 affects MBD5
| 8
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
KDM5C affects BAP1
| 1 7
| 1 2

sparser
"Mutations of KDM5C and BAP1 were significantly associated with evidence of renal vein invasion (p=0.022 and 0.046, respectively)."

sparser
"Mutations of the KDM5C and BAP1 genes were significantly associated with evidence of renal vein invasion on contrast-enhanced CT of ccRCC (p=0.022 and 0.046, respectively) The genotype of solid ccRCC differed significantly from the one of multicystic ccRCC."

reach
"S4G), indicating an SL interaction between BAP1 and KDM5C in UMRC6 cells.To determine whether the abovementioned SL interactions exist in patients with cancer, we checked The Cancer Genome Atlas (TCGA) data for somatic mutations targeting TSC2/LKB1 in non–small cell lung cancer and BAP1/KDM5C in ccRCC."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
KDM1B affects BAP1
6 | 2
6 | 2

No evidence text available

No evidence text available

sparser
"In addition, the mass spectrometric analysis by xref also identified a physical interaction between BAP1 and the H3K4 demethylase KDM1B (also known as LSD1B), an intriguing interaction that might implicate BAP1/ASXL in KDM1B function and will need to be validated further."

No evidence text available

No evidence text available

No evidence text available

sparser
"Mechanistically, we found that BAP1 interacts with KDM1B and stabilizes it through deubiquitination."

No evidence text available
INO80 affects BAP1
2 | 3 3
2 | 3 3

sparser
"BAP1 and INO80 bound to replication forks in response to replication stress induced by nucleotide depletion after hydroxyurea (HU) treatment."

sparser
"Both BAP1 and INO80 bind chromatin at replication forks upon HU treatment."

reach
"Both BAP1 and INO80 bind chromatin at replication forks upon HU treatment."

No evidence text available

reach
"A recent study demonstrates that Ino80 interacts with BRCA1 associated protein-1 (BAP1), a tumor suppressor that also stabilizes Ino80, in normal DNA replication [XREF_BIBR]."

No evidence text available

reach
"BAP1 interacts with and deubiquitylates INO80, playing a critical role in stabilizing and targeting the INO80 chromatin remodeling complex to the replication forks."

sparser
"Both BAP1 and INO80 bound to replication forks during S phase, and BAP1 depletion inhibited both INO80 recruitment to the fork and fork progression."
H2AC8 affects BAP1
8 |
8 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

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No evidence text available
COPA affects BAP1
4 | 2 2
4 | 2 2

No evidence text available

No evidence text available

sparser
"We note that only a small fraction of total COPA binds to BAP1 as input levels are much higher than COPA levels in the coIP, correlating with the stoichiometry observed in the AP-MS experiment ( xref and xref )."
| PMC

sparser
"These data confirm the interaction between BAP1 and COPA on endogenous protein and indicate that this interaction is not due to overexpression artefacts."
| PMC

reach
"In all BAP1 IP samples, but not the negative control normal IgG samples endogenous COPA coimmunoprecipitates with endogenous BAP1.We note that only a small fraction of total COPA binds to BAP1 as input levels are much higher than COPA levels in the coIP, correlating with the stoichiometry observed in the AP-MS experiment (Fig 5A and S2 File)."
| PMC

reach
"These data confirm the interaction between BAP1 and COPA on endogenous protein and indicate that this interaction is not due to overexpression artefacts."
| PMC

No evidence text available

No evidence text available
CDKN2A affects BAP1
1 | 6
CDKN2A inhibits BAP1.
| 5
CDKN2A inhibits BAP1. 5 / 5
| 5

reach
"To that end, a systematic literature review was performed of articles dealing with a loss of BRCA1-associated protein 1 (BAP1), methylthioadenosine (MTAP), 5-hydroxymethylcitosine (5-hmC), glucose transporter 1 (GLUT1), insulin like-growth factor II messenger RNA-binding protein 3 (IMP3), enhanced zeste homologue 2 (EZH2) staining, cyclin-dependent kinase inhibitor 2A (CDKN2A) homozygous deletion (HD) testing, soluble mesothelin, and microRNA quantification in cytological samples for the diagnosis of MPM versus reactive atypical mesothelial cells."

reach
"It is worth emphasizing that morphology alone is insufficient for a diagnosis of MIS, while demonstration of BAP1 loss by IHC or CDKN2A/p16 homozygous deletion by FISH are required [13,24]."

reach
"Loss of BAP-1 protein on IHC and homozygous deletion of the cyclin-dependent kinase inhibitor 2A (CDKN2A) gene (p16) on chromosome 9p21 by FISH on cell blocks/biopsy establishes the diagnosis of MM an[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The 2 peritoneal mesothelioma cases demonstrated loss of BAP1 but not p16."

reach
"Contribution of BAP1 loss and p16 (CDKN2A) deletion analysis to the definitive diagnosis of mesothelioma in effusion cytology."
CDKN2A binds BAP1.
1 | 1
1 | 1

reach
"χ 2 test or Fisher's exact test (in case of low event rates [n<5]) were used to investigate the association between BRCA1 and BAP1, p16ink4A, and PD-L1 expression and response outcome."

No evidence text available
BAP1 affects p14_3_3
| 8
| 8

sparser
"Next, we tested the hypothesis that the BAP1 interaction with 14-3-3 releases the binding between 14-3-3 and Bax, which further facilitates cell death."

sparser
"Another study in 2018, showed that BAP1 interacts with the protein 14-3-3 in neuroblastoma cells causing the release of Bax and the decrease of pro-survival factors such as Bcl-2, to promote apoptosis via the intrinsic pathway.[ xref ]"

sparser
"These results suggest that BAP1 interaction with 14-3-3 prevents cell survival signaling, which is essential for growth of the neuroblastoma cells (Fig.  xref )."

sparser
"BAP1 binding to 14-3-3 facilitate the apoptotic signaling in neuroblastoma."

sparser
"The interaction between BAP1 and 14-3-3 protein was confirmed by performing the co-IP of FLAG-tagged BAP1 and pull-down of endogenous 14-3-3 in neuroblastoma cells (Fig.  xref )."

sparser
"The reciprocal IP using 14-3-3 antibodies also indicated the formation of the complex between BAP1 and 14-3-3 (Fig.  xref )."

sparser
"In addition, fluorescence resonance energy transfer (FRET) assay demonstrated a direct interaction between BAP1 and 14-3-3 (Fig.  xref )."

sparser
"We could also confirm that the short fragment of BAP1-ULD domain and 14-3-3-alpha helices 7-9 regions facilitate interaction between these two proteins (Fig.  xref ), while the interaction between BAP1 and 14-3-3-σ inactive mutants were reduced compared to the full-length 14-3-3-σ (Fig.  xref )."
BAP1 affects miR-31
| 8
BAP1 inhibits miR-31.
| 3
BAP1 inhibits miR-31. 3 / 3
| 3

reach
"Xenografts with both miR-31 and BAP1 overexpression exhibited reduced cell mitosis compared to xenografts with miR-31 overexpression, suggesting that BAP1 overexpression could attenuate the pro proliferative effect of miR-31."

reach
"Finally, the cells transfected with miR-31 and the BAP1 overexpression plasmid exhibited significantly lower proliferation rates compared with the cells transfected with miR-31 and the control plasmid, suggesting that miR-31-resistant BAP1 could rescue the suppression of BAP1 by miR-31 and attenuate the proliferative effect of miR-31."

reach
"Additionally, BAP1 overexpression attenuated the promotive effect of miR-31 on tumor growth, suggesting that miR-31 might promote tumor growth by silencing BAP1."
BAP1 activates miR-31.
| 3
BAP1 activates miR-31. 3 / 3
| 3

reach
"We further examined whether BAP1 was a direct target of miR-31 in cervical cancer cells using dual-luciferase reporter assay."

reach
"In this study, we showed that BAP1 was a direct target of miR-31 and an inverse correlation between miR-31 and BAP1 expression in cervical cancer cells and tissues."

reach
"Bioinformatic and dual-luciferase reporter gene assay showed that BAP1 was the direct target of miR-31."
BAP1 binds miR-31.
| 2
BAP1 binds miR-31. 2 / 2
| 2

reach
"The aim of this study was to evaluate the interaction between BAP1 and miR-31 in MPM and their prognostic role in MPM."

reach
"These results indicate that miR-31 binds to BAP1 3 '-UTR directly and negatively regulates BAP1 expression in cervical cancer cells."

reach
"Additionally, BAP1 regulates EMT during trophoblast differentiation and invasion, while decreased BAP1 expression triggers EMT and enhances migration and invasion (27)."

reach
"In vitro, BAP1 inhibited proliferation and EMT."

reach
"This indicates that BAP1 function loss inhibits TGF-β secretion in PE, thus implying that BAP1 alterations affect the EMT and tumor immunoescape , although its clinical significance remains unknown."

reach
"Interestingly, focal adhesion was enriched in shBAP1#1 and shBAP1#2 samples (Figure 5), suggesting BAP1 might inhibit EMT in osteosarcoma."

reach
"BAP1 suppressed osteosarcoma cell growth and EMT."

reach
"One study on cervical cancer cell lines reported that BAP1 knockdown induced increases in N-cadherin and vimentin and a decrease in E-cadherin (which are hallmarks of EMT) and increased cell migration [91]."

reach
"In summary, we hypothesised that UMs with low MITF had more invasive phenotypes and less favourable EMT profiles (high TWIST1, SNAI1) than UMs with high MITF (which have high ZEB2 and SNAI2), and that this EMT shift may be initiated by BAP1 inactivation."

reach
"We also validated that BAP1 could promote cervical carcinogenesis and EMT progression was regulated by miR-31."
BAP1 affects TUBG
| 8
BAP1 deubiquitinates TUBG.
| 6
BAP1 deubiquitinates TUBG. 6 / 6
| 6

reach
"Another study showed de-ubiquitination of gamma-tubulin by BAP1 prevents chromosome instability in breast cancer cells [15]."

reach
"20 Additionally, loss of BAP1 or its DUB activity reduces deubiquitination of Gamma-tubulin in breast cancer cell lines, which may be attributed to the perturbation of mitotic spindle fiber formation [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 can deubiquitinate γ-tubulin [32], as well as microspherule protein 1 (MCRS1), a protein that caps the minus ends of mitotic microtubules to prevent their depolymerization [33, 34]."

reach
"While the mechanism remains to be fully elucidated, it seems that deubiquitylation of gamma-tubulin by BAP1 during mitosis allows proper spindle organisation and function [XREF_BIBR]."

reach
"Deubiquitination of gamma-tubulin by BAP1 prevents chromosome instability in breast cancer cells."

reach
"A recent study has linked the deubiquitination of gamma-tubulin by BAP1 to breast cancer."
BAP1 binds TUBG.
| 2
| 2

reach
"A study reported that BAP1 interacts directly with and localizes to γ-tubulin during mitosis and that BAP1 stabilizes γ-tubulin via deubiquitination to support microtubule nucleation and mitotic spindle assembly, thereby ensuring chromosome segregation and preventing chromosome abnormalities."

reach
"Interaction between BAP1 and γ-tubulin occurs during metaphase, anaphase and telophase, but not during prophase, during which γ-tubulin is ubiquitylated."
BAP1 affects TMED2
| 8
| 8

sparser
"Validation of the BAP1 COPI interaction."
| PMC

sparser
"In order to confirm the interaction between BAP1 and the heptameric COPI complex we performed AP-immunoblot experiments ( xref )."
| PMC

sparser
"To exclude that the BAP1 interaction with the COPI complex is an artifact of overexpression of the exogenous BAP1 construct, we immunoprecipitated endogenous BAP1 using different antibodies against BAP1 in cell lysate from HeLa FRT parental cells ( xref )."
| PMC

sparser
"AP-immunoblotting of GFP-COPE shows interaction of COPA as expected, however BAP1 cannot be observed which is probably due to the low levels of BAP1 that bind to the COPI complex ( xref )."
| PMC

sparser
"Although the absolute and relative amounts of COPI that interacts with BAP1 in the AP-MS is low ( xref and xref ), the interaction can still be biologically relevant and is validated with endogenous protein as seen in the immunoblot experiments ( xref )."
| PMC

sparser
"It was intriguing that BAP1 has such a sequence in its C-terminus, but our mutational analysis suggests that the interaction between BAP1 and the COPI complex is not mediated through the C-terminal KxKxx sequence in BAP1."
| PMC

sparser
"We must be absolutely sure that the BAP1-COPI interaction is not an artifact."
| PMC

sparser
"We agree with reviewer 2 that the interaction between BAP1 and the COPI complex may be a substrate interaction as evidenced by the low protein stoichiometry observed in the affinity purification (Fig 5)."
| PMC
BAP1 affects TERT
| 3 5
BAP1 binds TERT.
| 5
| 5

sparser
"Shorter overall survival was associated with TERT mutations ( p  = 0.03, median 29.6 months for MT versus 52.6 months for WT) or BAP1 mutations ( p  = 0.02, median 28.7 months for MT versus not reached for WT)."

sparser
"The presence of BAP1 or TERT promoter mutations was associated with a significantly worse OS."

sparser
"An exploratory analysis was performed to investigate if acquired mutations in BAP1 or TERT were associated with MSKCC prognostic risk group or the presence of sarcomatoid features on histopathologic review."

sparser
"This was also the case for mutually exclusive alterations such as SETD2–CDKN2A (3p21–9p21) and TERTBAP1 (5p15–3p21) (Fig.  xref and Supplementary Tables  xref – xref )."

sparser
"Emerging evidence (particularly for hepatocellular carcinoma) suggests that TERT could interact with BAP1 via the WNT/β-catenin signaling pathway, which is often activated in tumors with TERT overexpression."
BAP1 decreases the amount of TERT.
| 3
BAP1 decreases the amount of TERT. 3 / 3
| 3

reach
"Chromosome 3p21.1-21.3 encoded sequences BAP1, but not SETD2 PARP-3 and PBRM1, repress hTERT transcription within 21NT cells."

reach
"Interestingly, literature data suggest that BAP1 could downregulate TERT expression (Linne et al., 2017)."

reach
"In conclusion, the results from the present study provide evidence to suggest that BAP1 and possibly PARP-3 repress hTERT transcription within breast cancer cells, which supports the hypothesis that multiple sequences on human chromosome 3p may be responsible for regulating hTERT transcription."
BAP1 affects PTGES
| 8
BAP1 increases the amount of PTGES. 8 / 8
| 8

reach
"We found that the EP4 receptor positively regulates BaP1-triggered PGE release and expression of COX-2 and mPGES-1, which are critically involved in the biosynthesis of PGE induced by this metalloprotease."

reach
"Partially consistent with our results, BAP1 was unfolded to increase the COX-2 and mPGES-1 expression by activating the NF-κB signaling pathway and inducing the release of IL-1β (Viana et al. 2020), but this study failed to explore the regulatory role of BAP1 mutations on the NF-κB signaling pathway."

reach
"Expression of COX-2 and mPGES-1 are induced by BaP1 via activation of NF-kappaB pathway."

reach
"As shown in Fig. 5, pretreatment of cells with SN50 inhibited BaP1-induced COX-2 protein expression (Fig. 5C,D) and BaP1-induced mPGES-1 protein expression (Fig. 5G,H)."

reach
"Accordingly, our data show that BaP1 induced a marked increase in protein expression of both COX-2 and mPGES-1, the increase in COX-2 expression occurring before that of mPGES-1."

reach
"Furthermore, our findings that BaP1 induced gene expression of COX-2 and mPGES-1 demonstrate that BaP1 can trigger transcriptional mechanisms in FLSs that result in increased protein expression."

reach
"Intriguingly, BAP1 was unfolded to increase the COX-2 and mPGES-1 expression by activating the NF-κB pathway (Viana et al. 2020)."

reach
"Gene and protein expression of both COX-2 and mPGES-1 are induced by BaP1 via activation of distinct subunits of the NF-κB transcription factor."
BAP1 affects PROS1
| 1 7
BAP1 inhibits PROS1.
| 1 4
BAP1 inhibits PROS1. 5 / 5
| 1 4

eidos
"Inactivation of BAP1 promotes the upregulation of vitamin K-dependent protein S ( PROS1 ) , which interacts with the tyrosine-protein kinase Mer ( MERTK ) receptor on M2 macrophages to induce an immunosuppressive environment ."

reach
"Loss of BAP1 could promote the immunosuppressive microenvironment in Class 2 μm by regulating PROS1 (Kaler et al. 2022)."

reach
"Deletion of BAP1 in UMC026 cells and knockdown of BAP1 in Mel202 (class 1) cells resulted in upregulation of PROS1, whereas ectopic expression of BAP1 in BAP1-deficient MP46 (class 2) cells resulted in downregulation of PROS1 (Figure 1B and Supplementary Figures S1 and S2)."

reach
"These findings suggest that BAP1 represses PROS1 at least in part by epigenetic mechanisms, and that BAP1 loss leads to increased PROS1 expression."

reach
"However, it remains unknown how BAP1 mutations may mechanistically lead to immune suppression.As a potential explanation, we found here that BAP1 loss results in upregulation of PROS1 in UM cells through epigenetic mechanisms involving H3K27ac accumulation at the PROS1 locus, consistent with our previous findings [18]."
BAP1 decreases the amount of PROS1.
| 3
BAP1 decreases the amount of PROS1. 3 / 3
| 3

reach
"Kaler et al. found that BAP1 deficiency leads to increased PROS1 expression in melanoma cells, polarizes macrophages toward the M2 state, activates the suppressive tumor immune microenvironment, and ultimately promotes immune escape [29]."

reach
"These findings suggest that BAP1 represses PROS1 at least in part by epigenetic mechanisms, and that BAP1 loss leads to increased PROS1 expression."

reach
"Here, we show that BAP1 loss leads to increased expression of PROS1 in uveal melanocytes and melanoma cells, which in turn leads to phosphorylation and activation of the receptor tyrosine kinase MERTK on adjacent macrophages, driving them into a suppressive M2-polarized state."
BAP1 affects Necrosis
| 6
BAP1 activates Necrosis. 6 / 8
| 6

reach
"Functionally, BAP1 promotes apoptosis and necrosis, and down-regulates the migration and invasion abilities of LAC cells."

reach
"In functional assays, BAP1 was found to inhibit the migration and invasion of LAC cells, and promoted their apoptosis and necrosis."

reach
"P-I metalloproteinases and LAAO arrested the cells in G0/G1 phase of cell cycle and also led to cytotoxic activity in endothelial cells by inducing necrosis/late apoptotic mode of cell death."

reach
"These results indicated that P-I metalloproteinases and LAAO induced both necrosis and late apoptotic mode of cells death (Fig. 3A, Additional file 3B)."

reach
"In functional experiments, BAP1 has been shown to suppress lung cancer tumorigenesis in athymic nude mice, promote the cell cycle and induce both apoptosis and necrosis."

reach
"It is also prudent to note that expression of BAP1 increased the population of cells that stain positive for both AnnexinV and PI, an indication of late apoptosis and/or necrosis."
BAP1 affects Mice
| 8
BAP1 inhibits Mice.
| 4
BAP1 inhibits Mice. 4 / 4
| 4

reach
"For example, differentiation to mature neurons and glia could require antagonism of PRC-mediated transcriptional repression.We were thus intrigued by our serendipitous discovery that Bap1 deletion in ENS lineages caused profound bowel dysfunction and early death in mice, mimicking human nCIPO or HSCR physiology."

reach
"Thus, it is not surprising that human BAP1 mutations predispose to cancer (18) and global Bap1 knockout causes early embryonic lethality in mice (38)."

reach
"We discovered that fetal loss of the epigenetic regulator Bap1 in the ENS lineage caused severe postnatal bowel dysfunction and early death in Tyrosinase-Cre Bap1 fl/fl mice."

reach
"We found that loss of BAP1 leads to the development of chronic pancreatitis in mice and is associated with poor prognosis in PDA patients."
BAP1 activates Mice.
| 4
BAP1 activates Mice. 4 / 4
| 4

reach
"These results further confirm that the eradication of MC38 cells triggered by Bap1 deletion is independent of PD-L1 expression.Moreover, the overexpression of human WT BAP1 in Bap1-KO MC38 cells promoted their growth in host mice (Supplementary Fig. 3g–j)."

reach
"The fact that we determined that highly penetrant, heterozygous pathogenic BAP1 null variants by themselves, as studied by Kadariya et al. , can initiate and drive MM tumorigenesis in mice absent asbestos exposure supports their conclusions, in agreement with Chanock ."

reach
"In this study, we identified a decreased expression pattern of BAP1 in septic mice, and its artificial upregulation of BAP1 enhanced survival of the model mice and ameliorated the pathological changes[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 depletion inhibits the growth of colon tumors in xenograft mice."
BAP1 affects MLL2
| 8
MLL2 binds BAP1. 8 / 8
| 8

sparser
"The MLL4 and BAP1 co-occupancy depends on ASXL proteins that directly interact with both MLL4 and BAP1 and bring together two enzymatic activities to specific genomic loci."

sparser
"MLL4 occupancy on MLL4 + BAP1 + genomic regions depends on functional ASXLs that interact with both MLL4 and BAP1, and CRC-relevant mutations attenuate the formation of the MLL4-ASXL complex."

sparser
"MBH of ASXL1/2 is required for the association of BAP1 with MLL4 in ESCs."

sparser
"Although ASXLs have been shown to form complexes with both the deubiquitinase BAP1 and the methyltransferase MLL4(12,15), the importance of ASXLs in genomic distribution of BAP1 and MLL4 in CRC remains unclear."

sparser
"These results indicated that BAP1 associates with MLL4 in ESCs in an ASXL1/2 MBH-dependent manner."

sparser
"To assess whether ASXLs play a role in regulating MLL4 and BAP1 binding at specific chromatin sites, we generated MLL4-, ASXL2-or BAP1-deficient human colon cancer HCT116 cells using the CRISPR/Cas9 approach."

sparser
"Interdependent genomic binding of MLL4 and BAP1 on enhancers."

sparser
"The fact that MLL4 and BAP1 do not directly interact and are linked through their concomitant binding to ASXLs suggested a possible role of ASXLs in this regulation."
BAP1 affects MBD5, and MBD6
| 8
MBD5 binds MBD6 and BAP1. 8 / 8
| 8

sparser
"Furthermore, we have demonstrated that MBD5 or MBD6 do not directly interact with BAP1; instead, they bind to the C-terminal PHD fingers of all three ASXL subunits (ASXL1-3)."

sparser
"This result indicates an indirect interaction between MBD5, MBD6, and BAP1."

sparser
"To determine whether MBD5 and MBD6 can also bind to chromatin similar to other BAP1 subunits, we conducted Chromatin Immunopreciptiation Sequencing (ChIP-seq) and determined the chromatin binding profiles of MBD5 and MBD6 with our validated homemade antibodies (Additional file xref : Fig. S1F)."

sparser
"To investigate how MBD5 and MBD6 subunits are incorporated into the BAP1 complex, we first hypothesized that MBD5 and MBD6 may directly interact with BAP1 itself, similar to how other subunits function (e.g., ASXL1-3 and FOXK1/2)."

sparser
"Therefore, to determine whether ASXLs play a role in mediating the interaction between MBD5, MBD6, and BAP1, we knocked out all three ASXL genes (TKO) by CRISPR in HEK293T cells."

sparser
"Surprisingly, as shown in Fig. xref B, the protein-protein interaction between MBD5, MBD6, and BAP1 is completely abolished in ASXL-TKO cells (Fig. xref B)."

sparser
"Therefore, MBD5 and MBD6 have been excluded from the BAP1 complex due to a lack of evidence on endogenous protein-protein interaction between MBD5, MBD6, and BAP1 [ xref , xref – xref ]."

sparser
"After testing the specificity of the antibodies, we performed immunoprecipitation in HEK293T cells and demonstrated a stable protein-protein interaction between endogenous MBD5, MBD6, and BAP1 (Fig. xref C, D)."
BAP1 affects KDM5C
| 1 7
| 1 2

sparser
"Mutations of KDM5C and BAP1 were significantly associated with evidence of renal vein invasion (p=0.022 and 0.046, respectively)."

sparser
"Mutations of the KDM5C and BAP1 genes were significantly associated with evidence of renal vein invasion on contrast-enhanced CT of ccRCC (p=0.022 and 0.046, respectively) The genotype of solid ccRCC differed significantly from the one of multicystic ccRCC."

reach
"S4G), indicating an SL interaction between BAP1 and KDM5C in UMRC6 cells.To determine whether the abovementioned SL interactions exist in patients with cancer, we checked The Cancer Genome Atlas (TCGA) data for somatic mutations targeting TSC2/LKB1 in non–small cell lung cancer and BAP1/KDM5C in ccRCC."
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
BAP1 affects KDM1B
6 | 2
6 | 2

No evidence text available

No evidence text available

sparser
"In addition, the mass spectrometric analysis by xref also identified a physical interaction between BAP1 and the H3K4 demethylase KDM1B (also known as LSD1B), an intriguing interaction that might implicate BAP1/ASXL in KDM1B function and will need to be validated further."

No evidence text available

No evidence text available

No evidence text available

sparser
"Mechanistically, we found that BAP1 interacts with KDM1B and stabilizes it through deubiquitination."

No evidence text available
BAP1 affects H2AK119ub
| 8
BAP1 activates H2AK119ub.
| 6
BAP1 activates H2AK119ub. 6 / 6
| 6

reach
"Such as, the mutant ASXL1 (ASXL1-MT)/BAP1 complex can impair the multidirectional differentiation of HSPCs (except to monocytes) and promote RUNX1-ETO-induced murine c-Kit+ progenitor cell leukemogenesis by removing H2AK119ub, which drives the upregulation of the HOXA gene and interferon regulatory factor 8 (IRF8) [135]."

reach
"In line with our results using murine 32Dcl3 cells, ASXL1-MT was associated with global reduction of H2AK119ub, which was dramatically enhanced by the coexpression of BAP1."

reach
"32Dcl3 cells expressing ASXL1-MT showed global reduction of H2AK119ub, which was dramatically enhanced by the coexpression of BAP1, but not by that of BAP1-C91S."

reach
"Expression of ASXL1-WT showed no effect on global reduction of H2AK119ub, which was only slightly enhanced by the coexpression of BAP1."

reach
"Moreover, BAP1 depletion also abrogated the global reduction of H2AK119ub induced by ASXL1-MT."

reach
"H2AK119ub is associated with chromatin compaction and gene repression.61 Truncated ASXL1 strengthens its interaction with BAP1, which enhances removal of H2AK119ub."
BAP1 inhibits H2AK119ub.
| 2
BAP1 inhibits H2AK119ub. 2 / 2
| 2

reach
"Yet, on the other hand, the loss of BAP1 in mouse ESCs leads to spreading and genome-wide accumulation of H2AK119ub, resulting in reductions in global gene expression [95,96]."

reach
"The authors suggest that the function of BAP1 is to limit the spread of H2AK119ub to relevant target sites."
BAP1 affects H2AC8
8 |
8 |

No evidence text available

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BAP1 affects COPA
4 | 2 2
4 | 2 2

No evidence text available

No evidence text available

sparser
"We note that only a small fraction of total COPA binds to BAP1 as input levels are much higher than COPA levels in the coIP, correlating with the stoichiometry observed in the AP-MS experiment ( xref and xref )."
| PMC

sparser
"These data confirm the interaction between BAP1 and COPA on endogenous protein and indicate that this interaction is not due to overexpression artefacts."
| PMC

reach
"In all BAP1 IP samples, but not the negative control normal IgG samples endogenous COPA coimmunoprecipitates with endogenous BAP1.We note that only a small fraction of total COPA binds to BAP1 as input levels are much higher than COPA levels in the coIP, correlating with the stoichiometry observed in the AP-MS experiment (Fig 5A and S2 File)."
| PMC

reach
"These data confirm the interaction between BAP1 and COPA on endogenous protein and indicate that this interaction is not due to overexpression artefacts."
| PMC

No evidence text available

No evidence text available
BAP1 affects ASXL
| 8
ASXL binds BAP1. 8 / 8
| 8

reach
"Although H2Aub was previously involved in cell proliferation , it is not clear, at this time, whether the observed effects of ASXL2/BAP1 defective in monoubiquitination are solely due to disruption of H2A deubiquitination or the effect of BAP1/ASXL complexes on other substrates.Our data suggest that the signaling pathway we characterized here is important for tumor suppression."

reach
"This was in congruence with the fact that BAP1 binds to ASXL to form the Polycomb repressive deubiquitinase complex (PR-DUB) that deubiquitinates histone H2A [XREF_BIBR]."

reach
"BAP1 serves a key role in stabilizing epigenetic regulators, including O-linked N-acetylglucosamine (GlcNAc) transferase, host cell factor C1 and ASXL1, and thus an ASXL/BAP1 complex may inhibit chronic myelomonocytic leukemia, in which ASXL1 is frequently mutated (53)."

reach
"To be noticed, both leukemias and UMs associated with MBD4 inactivation share a consistent inactivation of the BAP1 and ASXL complex."

reach
"Our data raise the possibility that ASXL1 truncation mutations confer gain-of-function on the ASXL and BAP1 complex."

reach
"Human ASXL1 is mutated frequently in chronic myelomonocytic leukemia (CMML) so an ASXL and BAP1 complex may suppress CMML."

reach
"BAP1 and ASXL complex modulation regulates epithelial-mesenchymal transition during trophoblast differentiation and invasion."

reach
"The EMT is associated with upregulation of BAP1/ASXL complex modulation [9] , expression of TGFB1, as well as expression of PAPPA and PAPPA2 , which encode metalloproteinases that are known to be invo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
YY1 affects HCFC1
| 2 5
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
TNPO1 affects BAP1
5 | 2
5 | 2

No evidence text available

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sparser
"Having established the multi-mUb of BAP1 699–729 by UBE2O, we asked whether TNPO1 binding to BAP1 699–729 would down-regulate the ubiquitin ligase activity of UBE2O. Preincubation of the FITC-labeled BAP1 699–729 with one or two molar ratios of TNPO1 for 1 h before initiating the mUb by UBE2O resulted in a substantially reduced level of multi-mUb of BAP1 699–729 in a TNPO1 dose–dependent manner ( xref ; and xref )."

sparser
"Our structural and biochemical findings collectively demonstrated that TNPO1 binding to the NLS of BAP1 renders the mUb by UBE2O, which has direct implications in promoting BAP1 nuclear localization."
SETD2 affects PBRM1
| 7
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
| 7

reach
"BAP1 was originally found to interact with the RING finger domain of BRCA1 and to perform the cell growth-suppressive function."

reach
"BAP1 direct physical interaction with BRCA1-RING finger domain was associated to enhancement of BRCA1 tumour suppressor activity in breast cancer [11] (Fig. 1a)."

reach
"BAP1 interacts with the BRCA1 RING finger domain, which mediates BRCA1 's E3 ubiquitin ligase activity [XREF_BIBR]."

reach
"It has recently been reported that the RING finger domain of the Breast and Ovarian Cancer Susceptibility Gene product, BCRA-1, binds to the nuclear localized ubiquitin carboxy-terminal hydrolase BAP1[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For example, the N-terminal RING finger domain of BRCA-1, the hereditary breast and ovarian cancer tumor suppresser gene, interacts with BAP-1, a novel ubiquitin hydrolase that enhances BRCA-1-mediate[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 is mainly localized in the nucleus and exerts tumor suppressive effects by binding to the RING finger domain of BRCA1 [1]."

reach
"Especially, BAP1 binds to the RING finger domain of breast cancer type 1 susceptibility protein 1 (BRCA1), which plays a key role in tumor suppression and DNA repair [1] ."
RBBP7 affects BAP1
5 | 2
5 | 2

No evidence text available

No evidence text available

No evidence text available

sparser
"Mass spectrum analysis of human kidney cell line 786-O indicated that BAP1 can interact with RBBP7 [ xref ]."

sparser
"Herein, our results indicate that RBBP7 can bind to BAP1."

No evidence text available

No evidence text available
PBRM1 affects SETD2
| 7
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
MBP affects BAP1
| 7
| 7

sparser
"The results illustrated the ability of TNPO1 to dissociate homodimeric MBP-BAP1 CTD to form a heterodimeric TNPO1:MBP-BAP1 CTD complex with a 1:1 binding stoichiometry."

sparser
"Preincubation of MBP-BAP1 CTD with TNPO1 showed an increased apparent mol wt of 136.2 ± 8.1 kD for the main elution peak, which corresponded to a 1:1 binary complex ( xref )."

sparser
"Recombinant importin-αs and TNPO1 were expressed in Escherichia coli and purified to homogeneity to investigate their interactions with the CTD of BAP1 (residues 596–729) fused with a maltose-binding protein (MBP) tag (hereafter MBP-BAP1 CTD ) by size-exclusion chromatography (SEC; xref )."

sparser
"In the presence of two-fold excess karyopherins, MBP-BAP1 CTD formed stable complexes with individual karyopherins and coeluted at earlier elution volumes than did isolated proteins ( xref )."

sparser
"The formation of the binary complexes was confirmed by SDS-PAGE, which showed distinct protein bands of the early elution peaks, corresponding to MBP-BAP1 CTD and individual karyopherins ( xref )."

sparser
"MBP-BAP1 CTD by itself displayed protein concentration–dependent oligomerization with an apparent mol wt of 92.0–109.8 kD on increasing protein concentration ≤60 µM ( xref )."

sparser
"As the theoretical mol wt of an MBP-BAP1 CTD monomer is 59 kD, the results suggested a dynamic monomer-dimer equilibrium for MBP-BAP1 CTD ."
ITPR1 affects BAP1
| 7
| 7

sparser
"These results further confirmed a close interaction between BAP1 and IP3R. Overexpression of BAP1 significantly increased the expression of IP3R, while the knock down of BAP1 reduced the expression ( xref C and 7D)."

sparser
"Taken together, these results indicated that DIM could induce ferroptosis to exert anti-cancer effects via BAP1-IP3R axis, suggesting its effective therapeutic potential in gastric cancer."

sparser
"Ye et al .'s study [ xref ] showed that 3,3'-diindolyl methane can induce ferroptosis in BCC-823 GC cells and can induce ferroptosis through BAP1-IP3R axis to play an anticancer role, suggesting its effective therapeutic potential in GC."

sparser
"Mechanistically, BAP1 interacted with IP3R and increased cardiomyocytes’ Ca 2+ level, causing mPTP opening and mitochondrial dysfunction, promoting H/R-induced cardiomyocytes’ ferroptosis."

sparser
"In gastric cancer, DIM exerts antitumor effects through multiple mechanisms: Induction of ferroptosis via the BAP1-IP3R axis [ xref ]; STIM1-mediated store-operated calcium entry leading to cell death [ xref ]; inhibition of proliferation and induction of apoptosis via the TRAF2-p38 signaling pathway [ xref ]; potentiation of paclitaxel’s antitumor effects through the Akt/FOXM1 signaling cascade [ xref ]; suppression of cell growth via Hippo signaling pathway activation [ xref ]; activation of the aryl hydrocarbon receptor pathway, promoting its nuclear translocation, inducing apoptosis, delaying cell cycle progression, and inhibiting proliferation [ xref ]."

sparser
"Studies have shown that DIM plays an anticancer role by inducing ferroptosis in gastric cancer cells through the BAP1IP3R axis [ xref ]."

sparser
"3,3'-Diindolylmethane induces ferroptosis by BAP1-IP3R axis in BGC-823 gastric cancer cells."
HCFC1 affects YY1
| 2 5
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
BRCA1 affects BARD1
| 2 5
BRCA1 binds BARD1 and BAP1. 7 / 7
| 2 5

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear and needs to be studied in more detail."

reach
"Until recently, the combination of pemetrexed and cisplatin was the only approved first-line therapy, yielding a median overall survival (OS) of 12.1 months.2 Nivolumab plus ipilimumab was approved recently as initial treatment for malignant pleural mesothelioma (MPM) on the basis of a phase 3 clinical trial that resulted in a median OS of 18.1 months.3 4 There are no approved therapies for patients with tumor relapse after first-line therapies.Risk factors in developing mesothelioma include exposure to asbestos5 and other carcinogenic fibers such as erionite,6 and previous radiation therapy.7 8 Studies of familial clusters of mesothelioma have revealed genetic predisposition to develop MM.9 10 Recent work has identified germline mutations in BAP1 that can predispose to mesothelioma11 12 and other cancers such as uveal and cutaneous melanoma, basal cell carcinoma, meningioma, and renal cell cancers.13 14 We previously identified deleterious germline mutations in 12% of 241 consecutive patients with MM who enrolled on a prospective natural history study, with BAP1 being the most frequent germline mutation.15 BAP1 was also recurrently inactivated at the somatic level, suggesting BAP1 variants undergoing positive selection in the context of the classic “two-hit” model.BAP1, a nuclear deubiquitylase, was initially discovered as an interaction partner of the tumor suppressor BRCA1.16 Subsequent studies revealed that BAP1 binds to BARD1, which, in turn, binds to BRCA1, forming the BRCA1-BARD1-BAP1 complex that is crucial for the homologous recombination (HR)–mediated repair of DNA double-strand breaks (DSBs).17 PARP enzymes play a major role in DNA single-strand break repair and base excision repair pathways."

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear."

reach
"One feature of BAP1 is its association with BRCA1-associated RING domain 1 (BARD1) and BRCA1, forming the BRCA1-BARD1-BAP1 complex."

sparser
"The ring domains of BRCA1 (residues 1-109) are associated with BARD1 [ xref ] and BAP1 [ xref ]."

sparser
"BAP1 interacts with BARD1 and BRCA1 and inhibits the E3 ligase activity of BRCA1/BARD1."

sparser
"BAP1 and BRCA1-BARD1 complex coordinately regulate ubiquitination during the DNA damage response and the cell cycle [ xref , xref ]."
BAP1 affects localization
| 1 6
BAP1 activates localization.
| 4
| 4

reach
"Nuclear BAP1 is also associated in chromatin remodeling (49), DNA double-strand repair (50,51), and auto-deubiquitylation promoting its own nuclear localization (52)."

reach
"BAP1 inactivation promotes lactate production by leveraging the subcellular localization of LDHA in melanoma."

reach
"It was recently shown that the ubiquitin conjugating enzyme UBE2O multi-monoubiquitinates tumor suppressor BAP1 on its NLS bp to promote cytoplasmic localization."

reach
"For example, UBE2O binds to and multi-monoubiquitinates BAP1 to induce its cytoplasmic localization and promotes adipocyte differentiation [XREF_BIBR]."
BAP1 inhibits localization.
| 1 2
| 1 2

reach
"Both variants are found in several families with BAP1-TPDS and they cause loss of function (LOF) at the protein level by nearly eliminating BAP1 deubiquitinating activity, abolishing nuclear localization, or both (Fig. 5)."

reach
"Thus, Bap1 appears to inhibit the epigenetic function of Hdac4, at least in part, by restricting its localization to the nucleus."

eidos
"BAP1 ( BRCA1-associated protein-1 ) is a more recently discovered deubiquitinase member of the DRED complex and knockdown of BAP1 led to increased ubiquitination of NCoR1 and reduced its localization to the beta globin locus [ 32 ] ."
BAP1 affects TNPO1
5 | 2
5 | 2

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

sparser
"Having established the multi-mUb of BAP1 699–729 by UBE2O, we asked whether TNPO1 binding to BAP1 699–729 would down-regulate the ubiquitin ligase activity of UBE2O. Preincubation of the FITC-labeled BAP1 699–729 with one or two molar ratios of TNPO1 for 1 h before initiating the mUb by UBE2O resulted in a substantially reduced level of multi-mUb of BAP1 699–729 in a TNPO1 dose–dependent manner ( xref ; and xref )."

sparser
"Our structural and biochemical findings collectively demonstrated that TNPO1 binding to the NLS of BAP1 renders the mUb by UBE2O, which has direct implications in promoting BAP1 nuclear localization."
BAP1 affects STMN1
1 | 6
BAP1 binds STMN1.
1 | 3
1 | 3

reach
"Semi-endogenous co-immunoprecipitation of SMN with intact or truncated Bap1 revealed that the ubiquitin C-terminal hydrolase domain of Bap1 is essential for the binding of Bap1 to SMN (Supplemental Figure 7F)."

reach
"Taken together, we concluded that Bap1 directly binds SMN and prevents SMN degradation by posttranslational deubiquitination of SMN.Since a lysine residue is one of the predominant targets for ubiquitination, we examined which of the lysine residues in SMN is critical for its ubiquitin-dependent degradation by introducing a non-lysine mutation on phylogenetically conserved lysine residues (Supplemental Figure 8A)."

No evidence text available

reach
"These results indicate that Bap1 posttranscriptionally regulates SMN levels and thus affects the function of SMN in FAPs.To test the direct interaction between Bap1 and SMN, we performed endogenous reciprocal immunoprecipitation experiments using freshly isolated FAPs and revealed that Bap1 directly binds SMN in FAPs (Figure 7C)."
BAP1 deubiquitinates STMN1.
| 3
BAP1 deubiquitinates STMN1. 3 / 3
| 3

reach
"Deubiquitination of SMN by Bap1.."

reach
"Notably, overexpression of Bap1 reversed the ubiquitination of SMN (Figure 7D), and this result indicated that Bap1 could regulate SMN levels by acting as a deubiquitinating enzyme."

reach
"While Bap1 could prevent the accumulation of ubiquitinated SMN in the UPS-inhibited cells, Bap1 was unable to deubiquitinate SMN (Figure 7E)."
BAP1 affects S phase
| 7
BAP1 inhibits S phase.
| 4
BAP1 inhibits S phase. 4 / 4
| 4

reach
"Further cell cycle analysis revealed that overexpression of BAP1 in HCCC9810 cells significantly reduced cells in the S phase and arrested cells in the G0/G1 phase, which suggested that overexpression of BAP1 resulted in decreased G1-phase to S-phase cell cycle progression."

reach
"Silencing BAP1 could cause retarded cell proliferation in multiple mesothelioma cell lines through arresting S-phase of cell cycle, which was remitted by reintroducing BAP1 [13] ."

reach
"Although paradoxical for a tumor suppressor gene and contrary to in vivo observations, these data are consistent across cell lines of different origin [31,32] , suggesting that BAP1 suppression causes[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 regulates replication fork formation, disrupts S phase arrest, and maintains G1max S phase fidelity under normal cellular conditions [29]."
BAP1 activates S phase.
| 3
BAP1 activates S phase. 3 / 3
| 3

reach
"Similarly, downregulation of BAP1 markedly increased cells in the S phase and reduced cells in the G0/G1 phase, consistent with the results of a recent study that depletion of BAP1 resulted in a modest accumulation of host cell factor 1, which promoted the transition from G1 to S phase ."

reach
"BAP1 knockdown downregulated E2F1 target genes essential for S phase initiation and subsequent G1/S transition (shown in Fig. 2 )."

reach
"Lost expression of BAP1 seems to promote the survival of H28 cells, reducing effects of transfection on cell cycle block, while normal levels of BAP1 in wild type cells H2052 caused a S-phase retardat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 7

reach
"BAP1 was originally found to interact with the RING finger domain of BRCA1 and to perform the cell growth-suppressive function."

reach
"BAP1 direct physical interaction with BRCA1-RING finger domain was associated to enhancement of BRCA1 tumour suppressor activity in breast cancer [11] (Fig. 1a)."

reach
"BAP1 interacts with the BRCA1 RING finger domain, which mediates BRCA1 's E3 ubiquitin ligase activity [XREF_BIBR]."

reach
"It has recently been reported that the RING finger domain of the Breast and Ovarian Cancer Susceptibility Gene product, BCRA-1, binds to the nuclear localized ubiquitin carboxy-terminal hydrolase BAP1[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For example, the N-terminal RING finger domain of BRCA-1, the hereditary breast and ovarian cancer tumor suppresser gene, interacts with BAP-1, a novel ubiquitin hydrolase that enhances BRCA-1-mediate[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 is mainly localized in the nucleus and exerts tumor suppressive effects by binding to the RING finger domain of BRCA1 [1]."

reach
"Especially, BAP1 binds to the RING finger domain of breast cancer type 1 susceptibility protein 1 (BRCA1), which plays a key role in tumor suppression and DNA repair [1] ."
BAP1 affects RBBP7
5 | 2
5 | 2

No evidence text available

No evidence text available

No evidence text available

sparser
"Mass spectrum analysis of human kidney cell line 786-O indicated that BAP1 can interact with RBBP7 [ xref ]."

sparser
"Herein, our results indicate that RBBP7 can bind to BAP1."

No evidence text available

No evidence text available
BAP1 affects MBP
| 7
| 7

sparser
"The results illustrated the ability of TNPO1 to dissociate homodimeric MBP-BAP1 CTD to form a heterodimeric TNPO1:MBP-BAP1 CTD complex with a 1:1 binding stoichiometry."

sparser
"Preincubation of MBP-BAP1 CTD with TNPO1 showed an increased apparent mol wt of 136.2 ± 8.1 kD for the main elution peak, which corresponded to a 1:1 binary complex ( xref )."

sparser
"Recombinant importin-αs and TNPO1 were expressed in Escherichia coli and purified to homogeneity to investigate their interactions with the CTD of BAP1 (residues 596–729) fused with a maltose-binding protein (MBP) tag (hereafter MBP-BAP1 CTD ) by size-exclusion chromatography (SEC; xref )."

sparser
"In the presence of two-fold excess karyopherins, MBP-BAP1 CTD formed stable complexes with individual karyopherins and coeluted at earlier elution volumes than did isolated proteins ( xref )."

sparser
"The formation of the binary complexes was confirmed by SDS-PAGE, which showed distinct protein bands of the early elution peaks, corresponding to MBP-BAP1 CTD and individual karyopherins ( xref )."

sparser
"MBP-BAP1 CTD by itself displayed protein concentration–dependent oligomerization with an apparent mol wt of 92.0–109.8 kD on increasing protein concentration ≤60 µM ( xref )."

sparser
"As the theoretical mol wt of an MBP-BAP1 CTD monomer is 59 kD, the results suggested a dynamic monomer-dimer equilibrium for MBP-BAP1 CTD ."
BAP1 affects MBD4
| 6 1
BAP1 binds MBD4.
| 3 1
| 3 1

reach
"In addition, an examination of the interaction between Med1 and BAP1 proteins showed that Med1 and BAP1 proteins are binding and are involved in the development and metastasis of cancer (Figure 3)."

reach
"Although the binding of Med1 and BAP1 involved in the cancer growth or metastasis, mechanisms are still unclear."

sparser
"Epidemiological studies have also shown an increased risk of prostate cancer, thyroid cancer, and leukemia among UM patients, which is not currently associated with BAP1 or MBD4 ( xref , xref )."

reach
"However, in the future, the results of this study will contribute to various research fields, such as the mechanism of regulating the expression of target genes by the binding of Med1 and BAP1 and the growth and metastasis of cancer by Med1 and BAP1."
BAP1 increases the amount of MBD4.
| 3
BAP1 increases the amount of MBD4. 3 / 3
| 3

reach
"Med1 protein expression was increased by BAP1 wild-type overexpression in MCF7 cells, whereas it was decreased by BAP1 C91A mutant overexpression (Figure 2b)."

reach
"These results addressed that the regulation of Med1 protein expression by BAP1 is regulated at the protein level and is regulated by different mechanisms from growth factors."

reach
"Med1 protein expression was highly increased by BAP1 overexpression, whereas it did not affect USP21 and USP35 (Figure 2a)."
BAP1 affects ITPR1
| 7
| 7

sparser
"These results further confirmed a close interaction between BAP1 and IP3R. Overexpression of BAP1 significantly increased the expression of IP3R, while the knock down of BAP1 reduced the expression ( xref C and 7D)."

sparser
"Taken together, these results indicated that DIM could induce ferroptosis to exert anti-cancer effects via BAP1-IP3R axis, suggesting its effective therapeutic potential in gastric cancer."

sparser
"Ye et al .'s study [ xref ] showed that 3,3'-diindolyl methane can induce ferroptosis in BCC-823 GC cells and can induce ferroptosis through BAP1-IP3R axis to play an anticancer role, suggesting its effective therapeutic potential in GC."

sparser
"Mechanistically, BAP1 interacted with IP3R and increased cardiomyocytes’ Ca 2+ level, causing mPTP opening and mitochondrial dysfunction, promoting H/R-induced cardiomyocytes’ ferroptosis."

sparser
"In gastric cancer, DIM exerts antitumor effects through multiple mechanisms: Induction of ferroptosis via the BAP1-IP3R axis [ xref ]; STIM1-mediated store-operated calcium entry leading to cell death [ xref ]; inhibition of proliferation and induction of apoptosis via the TRAF2-p38 signaling pathway [ xref ]; potentiation of paclitaxel’s antitumor effects through the Akt/FOXM1 signaling cascade [ xref ]; suppression of cell growth via Hippo signaling pathway activation [ xref ]; activation of the aryl hydrocarbon receptor pathway, promoting its nuclear translocation, inducing apoptosis, delaying cell cycle progression, and inhibiting proliferation [ xref ]."

sparser
"Studies have shown that DIM plays an anticancer role by inducing ferroptosis in gastric cancer cells through the BAP1IP3R axis [ xref ]."

sparser
"3,3'-Diindolylmethane induces ferroptosis by BAP1-IP3R axis in BGC-823 gastric cancer cells."
BAP1 affects IRF8
| 7
BAP1 activates IRF8. 7 / 7
| 7

reach
"Additionally, BAP1-inactivation notably increases IRF8 activity, which is involved in DC differentiation."

reach
"RNA sequencing analyses revealed that reduced BAP1 expression suppressed ASXL1 Y591X -induced upregulation of the transcription factors activator protein 1 ( AP-1 ) and early growth response 1/2 ( EGR[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We also examined whether ASXL1-MT and BAP1 upregulated posterior HOXA genes and IRF8 in human RUNX1-ETO cells by qRT-PCR."

reach
"ASXL1-MT and BAP1 induces upregulation of HOXA genes and IRF8."

reach
"In addition to posterior Hoxa genes, we found that coexpression of ASXL1-MT and BAP1 in c-Kit + cells led to upregulation of Irf8, a master regulator of monocyte differentiation, at both mRNA and protein levels 37."

reach
"Moreover, the mutant ASXL1 (ASXL1-MT)/BAP1 complex promotes MLL rearrangement leukemia progression by reducing the upregulation of HOXA5, HOXA7, HOXA9, and IRF8 induced by H2AK119ub1 [136]."

reach
"Further chromatin immunoprecipitation quantitative PCR (ChIP-qPCR) assays confirmed that BAP1 reduction rescued the decreased occupancy of H2AK119Ub around the transcription start site (TSS) region of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects HCFC1, and YY1
| 2 5
BAP1 binds HCFC1 and YY1. 7 / 7
| 2 5

sparser
"31 Further, BAP1 forms a protein complex with the host cell factor 1 and Yin and Yang 1."

sparser
"3,10,12 BAP1 and HCF-1 can also form a complex with the transcription factor YY1, which recruits BAP1 and HCF-1 to various promoters to regulate gene expression."

sparser
"Finally, BAP1 forms complexes with the transcription factors Yin Yang 1 (YY1) and host cell factor 1 (HCF-1), and controls cell-cycle progression at G1/S by co-regulating transcription from HCF-1/E2F-governed promoters [ xref , xref ]."

sparser
"BAP1 forms a ternary complex with HCF1 and YY1, which acts as transcriptional repressor of genes involved in cell proliferation ( xref )."

reach
"Prominent among the positively enriched gene sets were those related to writers of histone H3K27Me3, including PRC2 and SUZ12 (XREF_FIG), as well as genes participating in cell proliferation and transcription regulation involving ubiquitin signaling mediated by the Bap1, HCF1, and YY1 complex, EED related stem cell pluripotency, cytokine signaling, and cell adhesion; 100 of the top pathways differentially affected in MMs from TKO versus DKO mice are presented in XREF_SUPPLEMENTARY."

reach
"Our RNA-seq analysis revealed that MMs from Bap1; Nf2; Cdkn2a CKO mice exhibit positively enriched gene sets consistent with cellular processes previously implicated in Bap1 function, such as genes mediated by the Bap1, HCF1, and YY1 complex, which play a role in cellular proliferation and transcription regulation involving ubiquitin signaling."

sparser
"Yu H, Mashtalir N, Daou S, et al. The ubiquitin carboxyl hydrolase BAP1 forms a ternary complex with YY1 and HCF-1 and is a critical regulator of gene expression."
BAP1 affects H2AK119ub1
| 7
BAP1 deubiquitinates H2AK119ub1.
| 3
BAP1 deubiquitinates H2AK119ub1. 3 / 3
| 3

reach
"Mammalian BAP1 deubiquitylates H2AK119ub1, 87 and BAP1 reintroduction into BAP1 deficient ccRCC cell lines affected global levels of H2AK119ub1."

reach
"XREF_BIBR reported that BAP1 is phosphorylated at serine 592 following DNA damage, that phospho-BAP1 causes deubiquitination of H2AK119ub1, and that this signal promotes the HR pathway."

reach
"87 As Drosophila Calypso, human BAP1 deubiquitylates H2AK119ub1."
BAP1 decreases the amount of H2AK119ub1.
| 2
BAP1 decreases the amount of H2AK119ub1. 2 / 2
| 2

reach
"Moreover, the reintroduction of wild-type BAP1 into 769-P cells (BAP1 mutant cells) downregulates H2AK119ub1 levels in renal cancer cells ( Pena-Llopis et al., 2012 )."

reach
"Previous studies showed that BAP1 knockdown led to increased global H2AK119ub1 levels in UM and renal carcinoma cell lines ( Landreville et al., 2012; Pena-Llopis et al., 2012 )."
BAP1 activates H2AK119ub1.
| 2
BAP1 activates H2AK119ub1. 2 / 2
| 2

reach
"In human tumors, H2AK119ub1 deposition is enhanced by frequent inactivating mutations of the H2AK119ub1-specific deubiquitinase BAP1 (Carbone et al., 2013)."

reach
"BAP1 enhances Polycomb repression by counteracting widespread H2AK119ub1 deposition and chromatin condensation."
BAP1 affects BARD1, and BRCA1
| 2 5
BRCA1 binds BARD1 and BAP1. 7 / 7
| 2 5

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear and needs to be studied in more detail."

reach
"Until recently, the combination of pemetrexed and cisplatin was the only approved first-line therapy, yielding a median overall survival (OS) of 12.1 months.2 Nivolumab plus ipilimumab was approved recently as initial treatment for malignant pleural mesothelioma (MPM) on the basis of a phase 3 clinical trial that resulted in a median OS of 18.1 months.3 4 There are no approved therapies for patients with tumor relapse after first-line therapies.Risk factors in developing mesothelioma include exposure to asbestos5 and other carcinogenic fibers such as erionite,6 and previous radiation therapy.7 8 Studies of familial clusters of mesothelioma have revealed genetic predisposition to develop MM.9 10 Recent work has identified germline mutations in BAP1 that can predispose to mesothelioma11 12 and other cancers such as uveal and cutaneous melanoma, basal cell carcinoma, meningioma, and renal cell cancers.13 14 We previously identified deleterious germline mutations in 12% of 241 consecutive patients with MM who enrolled on a prospective natural history study, with BAP1 being the most frequent germline mutation.15 BAP1 was also recurrently inactivated at the somatic level, suggesting BAP1 variants undergoing positive selection in the context of the classic “two-hit” model.BAP1, a nuclear deubiquitylase, was initially discovered as an interaction partner of the tumor suppressor BRCA1.16 Subsequent studies revealed that BAP1 binds to BARD1, which, in turn, binds to BRCA1, forming the BRCA1-BARD1-BAP1 complex that is crucial for the homologous recombination (HR)–mediated repair of DNA double-strand breaks (DSBs).17 PARP enzymes play a major role in DNA single-strand break repair and base excision repair pathways."

sparser
"Therefore, the association of BAP1 with BRCA1 and BARD1 is currently unclear."

reach
"One feature of BAP1 is its association with BRCA1-associated RING domain 1 (BARD1) and BRCA1, forming the BRCA1-BARD1-BAP1 complex."

sparser
"The ring domains of BRCA1 (residues 1-109) are associated with BARD1 [ xref ] and BAP1 [ xref ]."

sparser
"BAP1 interacts with BARD1 and BRCA1 and inhibits the E3 ligase activity of BRCA1/BARD1."

sparser
"BAP1 and BRCA1-BARD1 complex coordinately regulate ubiquitination during the DNA damage response and the cell cycle [ xref , xref ]."
SF3B1 affects BAP1
1 | 5
1 | 2

sparser
"That study highlighted the role of coding and non-coding genes that were differentially expressed between SCNA subtypes of monosomy 3 uveal melanomas that were associated with metastasis. xref Five genes have been reported to be frequently mutated in uveal melanoma. xref Mutations in GNAQ and GNA11 occur early in tumor formation while BAP1 , SF3B1 , and EIF1AX mutations likely occur later in tumor progression. xref Mutation in EIF1AX is an indicator of good prognosis, whereas mutations in SF3B1 and BAP1 are associated with intermediate and poor prognosis."

sparser
"To further investigate the genetic interaction between BAP1 and SF3B1 mutations and to exclude that other alterations in Mel202 might contribute to the incompatibility with BAP1 deletion, we asked whether deletion of the SF3B1 ‐mutant allele in Mel202 cells could reverse BAP1 KO‐induced growth arrest."

No evidence text available
| 3

sparser
"The Rotterdam Ocular Melanoma Study Group investigated the association of EIF1AX , SF3B1 and BAP1 mutation with disease-free survival and metastatic risk of patients with uveal melanoma. xref In their recent article, they published a Kaplan-Meier survival probability curve regarding these mutations."

sparser
"Uveal melanoma (UM) is a relative rare disease and has a high mortality rate due to metastasis in about half of all patients within 15 y after diagnosis. xref , xref , xref It is the most common primary intra‐ocular malignancy in adults in the Western world. xref UM specific mutations in the alpha subunit genes GNAQ and GNA11 are described as well as mutations in BAP1 , SF3B1 , and EIF1AX . xref , xref , xref Mutations in the latter 3 genes are found in ∼75% of all UM and are useful for prognostication of patients. xref , xref , xref BAP1 ‐mutated UM gives rise to early‐onset metastasis whereas SF3B1 ‐mutated UM gives rise to late‐onset metastasis and EIF1AX ‐mutated UM hardly metastasizes. xref Mutations in PLCB4 and CYSLTR2 are described in UM in a mutually exclusive manner to GNAQ or GNA11 mutations but so far have not been associated with prognosis. xref , xref Copy number alterations in chromosomes 1, 3, 6, and 8 are correlated with prognosis of the UM patient. xref , xref UM with EIF1AX , SF3B1 and BAP1 mutations are associated with unique chromosomal patterns, suggesting distinct UM subclasses."

sparser
"Other significantly mutated genes in UM are BAP1 , EIF1AX and SF3B1 , which also form a second mutually‐exclusive subgroup xref (Table  xref )."
N1ICD affects BAP1
| 6
BAP1 binds N1ICD. 6 / 6
| 6

sparser
"In a more stringent assay, the interaction between N1ICD and endogenous BAP1 was validated by FLAG-bound IP in HeLa N1ICD-FLAG cells ( xref )."

sparser
"Parallelly, protein levels of activated NOTCH1 was found to be highly expressed in cortical NPCs ( xref ), and BAP1 associates with endogenous activated N1ICD in cortical NPCs ( xref )."

sparser
"Interestingly, we found that the RAM–ANK and PEST domains of N1ICD, regions required for NUMB association, mediate N1ICD interaction with BAP1 as well ( xref ), and BAP1 regulates N1ICD stability through the PEST domain ( xref )."

sparser
"Moreover, the presence of full-length NUMB, but not the AB-deleted NUMB, significantly enhances the interaction between BAP1 and N1ICD."

sparser
"Finally, NUMB overexpression enhanced the interaction between BAP1 and N1ICD ( xref ), but co-expression of non-N1ICD-bound NUMB-∆2–15 could not promote BAP1N1ICD association, thus failing to boost stabilization of N1ICD ( xref )."

sparser
"Next, we asked whether BAP1 is associated with N1ICD."
MYCN affects BAP1
2 | 1 3
2 | 1 3

reach
"Here, we demonstrate that BAP1 could bind and deubiquitinate MYCN to increase the stability of MYCN."

sparser
"In support of the notion that BAP1 might be an upstream DUB for MYCN, Flag-BAP1 and HA-MYCN were co-transfected into 293T cells and the immunoprecipitation (IP) assay was performed to examine whether BAP1 will interact with MYCN."

No evidence text available

sparser
"Furthermore, endogenous BAP1 could specifically interacts with MYCN in 293T cells (Fig. xref )."

sparser
"Here, we demonstrate that BAP1 could bind and deubiquitinate MYCN to increase the stability of MYCN."

No evidence text available
MED1 affects BAP1
| 6
MED1 binds BAP1. 6 / 6
| 6

sparser
"Med1 is associated with BAP1."

sparser
"To investigate the interaction between Med1 and BAP1 protein, co-immunoprecipitation was performed in the BAP1 overexpressing cell line, BAP1 WT and BAP1 C91A mutant were overexpressed, and IP was performed using an anti-Med1 antibody in BAP1 or BAP1 C91A overexpressed cells."

sparser
"These results demonstrate that BAP1 is associated with Med1, but not with the BAP1 catalytic mutant."

sparser
"In addition, an examination of the interaction between Med1 and BAP1 proteins showed that Med1 and BAP1 proteins are binding and are involved in the development and metastasis of cancer ( xref )."

sparser
"Although the binding of Med1 and BAP1 involved in the cancer growth or metastasis, mechanisms are still unclear."

sparser
"However, in the future, the results of this study will contribute to various research fields, such as the mechanism of regulating the expression of target genes by the binding of Med1 and BAP1 and the growth and metastasis of cancer by Med1 and BAP1."
HDAC1 affects BAP1
4 | 1 1
4 | 1 1

No evidence text available

No evidence text available

No evidence text available

sparser
"We recently discovered that in the nucleus BAP1 binds and deubiquitylates HDAC1, which in turn binds and deacetylates HMGB1, keeping HMGB1 in the nucleus."

No evidence text available

reach
"We recently discovered that in the nucleus BAP1 binds and deubiquitylates HDAC1, which in turn binds and deacetylates HMGB1, keeping HMGB1 in the nucleus."
HAT1 affects BAP1
5 | 1
5 | 1

No evidence text available

No evidence text available

No evidence text available

reach
"HAT1 and COPI interaction with BAP1 is observed in the cytoplasm and not in the nuclear fractions (Figs 1A and 1B and 4)."
| PMC

No evidence text available

No evidence text available
2 | 1 2
BAP1 translocates.
| 1 2
BAP1 translocates to the cytoplasm. 3 / 3
| 1 2

sparser
"Our data do not exclude the possibility that, under specific conditions, BAP1 might be exported to the cytoplasm prior to its ubiquitination, although we did not identify any export signal in BAP1."

reach
"Our data do not exclude the possibility that, under specific conditions, BAP1 might be exported to the cytoplasm prior to its ubiquitination, although we did not identify any export signal in BAP1."

sparser
"The putative E2/E3 hybrid ubiquitin ligase, UBE2O, has been reported to mediate cytoplasmic translocation of BAP1 by multi‐monoubiquitylation of the NLS."
BAP1 binds.
2 |
2 |

No evidence text available

No evidence text available
| 3

reach
"Recent findings by Souri et al., concluded that loss of chromosome 3 and BAP1 expression correlated with the upregulation of the NF-κB (NF-κB1, NF-κB2, and RELB) signaling pathway and consequently promotes inflammation in UM tumors ."

reach
"For example, loss of the CDKN2A locus has been linked to immune evasion and reduced immunotherapy responses [54], while loss of BAP1 promotes TME inflammation, suggesting its potential as a biomarker for improved immunotherapy response [55]."

reach
"Loss of Bap1 increases tumor inflammation."
| 3

reach
"The BaP1-induced inflammatory response is characterized by formation of the complement component C5a and increase of neutrophil chemotaxis [ 23 ]."

reach
"These results indicated that BAP1 affects the expression of chemokines and cytokines, T and B cell migration, and activated inflammation associating with innate immunity."

reach
"It is concluded that MT-III and BaP1 induce a local inflammatory response associated with the synthesis of IL-1beta, IL-6 and MMPs."
BAP1 affects gemcitabine
| 6
| 6

reach
"Parasramka et al [158] found that BAP1 expression could modulate CCA cell sensitivity to gemcitabine, or more specifically, sensitivity to gemcitabine was increased in low BAP1 expressing CCA cells than in high BAP1 expressing CCA cells."

reach
"Preclinical evidence suggests that BAP1 wild-type status increases sensitivity to gemcitabine."

reach
"BAP1 dependent expression of long non coding RNA NEAT-1 contributes to sensitivity to gemcitabine in cholangiocarcinoma."

reach
"Inversely BAP1 expression was found to increase the effect of gemcitabine in mesothelioma cell lines, showing that the expression of BAP1 may affect outcomes differently according to type of chemotherapy (Guazzelli et al. 2019)."

reach
"BAP1 modulates sensitivity to gemcitabine in CCA cells."

reach
"Taken together these studies indicate that BAP1 expression modulates sensitivity to gemcitabine."
BAP1 affects SF3B1
1 | 5
1 | 2

sparser
"That study highlighted the role of coding and non-coding genes that were differentially expressed between SCNA subtypes of monosomy 3 uveal melanomas that were associated with metastasis. xref Five genes have been reported to be frequently mutated in uveal melanoma. xref Mutations in GNAQ and GNA11 occur early in tumor formation while BAP1 , SF3B1 , and EIF1AX mutations likely occur later in tumor progression. xref Mutation in EIF1AX is an indicator of good prognosis, whereas mutations in SF3B1 and BAP1 are associated with intermediate and poor prognosis."

sparser
"To further investigate the genetic interaction between BAP1 and SF3B1 mutations and to exclude that other alterations in Mel202 might contribute to the incompatibility with BAP1 deletion, we asked whether deletion of the SF3B1 ‐mutant allele in Mel202 cells could reverse BAP1 KO‐induced growth arrest."

No evidence text available
| 3

sparser
"The Rotterdam Ocular Melanoma Study Group investigated the association of EIF1AX , SF3B1 and BAP1 mutation with disease-free survival and metastatic risk of patients with uveal melanoma. xref In their recent article, they published a Kaplan-Meier survival probability curve regarding these mutations."

sparser
"Uveal melanoma (UM) is a relative rare disease and has a high mortality rate due to metastasis in about half of all patients within 15 y after diagnosis. xref , xref , xref It is the most common primary intra‐ocular malignancy in adults in the Western world. xref UM specific mutations in the alpha subunit genes GNAQ and GNA11 are described as well as mutations in BAP1 , SF3B1 , and EIF1AX . xref , xref , xref Mutations in the latter 3 genes are found in ∼75% of all UM and are useful for prognostication of patients. xref , xref , xref BAP1 ‐mutated UM gives rise to early‐onset metastasis whereas SF3B1 ‐mutated UM gives rise to late‐onset metastasis and EIF1AX ‐mutated UM hardly metastasizes. xref Mutations in PLCB4 and CYSLTR2 are described in UM in a mutually exclusive manner to GNAQ or GNA11 mutations but so far have not been associated with prognosis. xref , xref Copy number alterations in chromosomes 1, 3, 6, and 8 are correlated with prognosis of the UM patient. xref , xref UM with EIF1AX , SF3B1 and BAP1 mutations are associated with unique chromosomal patterns, suggesting distinct UM subclasses."

sparser
"Other significantly mutated genes in UM are BAP1 , EIF1AX and SF3B1 , which also form a second mutually‐exclusive subgroup xref (Table  xref )."
BAP1 affects RAR
| 6
BAP1 inhibits RAR. 6 / 6
| 6

reach
"Co-expression of OGT led to a considerable increase in BAP1-mediated RAR repression ( Fig. 3 A), which was impaired following treatment with the OGT inhibitor alloxan ( Fig. 3 B)."

reach
"See also Figures equirement of BAP1 for ASXL1 mediated RAR repression."

reach
"BAP1 Is a Histone H2B Specific DUB In Vivo and In Vitro To gain insight into themechanisms of RAR repressionmediated by ASXL1 and BAP1, we first determined whether the deubiquiti- nase (DUB) activity [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Based on these data, we propose that BAP1 O -GlcNAcylation promotes RAR repression required for maintaining pluripotency, while its inhibition allows RAR activation to initiate ES cell differentiation[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"(B) R ASXL1 for BAP1 mediated RAR repression."

reach
"Similar to ASXL1, BAP1 overexpression repressed RAR transcriptional activity ( Supplemental Fig. 2C )."
BAP1 affects PPARGC1A
| 4 2
BAP1 binds PPARGC1A.
| 2 2
| 2 2

sparser
"Conversely, O -GlcNAcylation of the transcriptional regulator PGC1α facilitates the binding of the deubiquitinase BAP1 to PGC1α, leading thus to its stabilization by blocking its ubiquitination ( xref )."

reach
"PUGNAc treatment promotes the interaction between PGC-1alpha and BAP1, while it has no effect on the S333A mutant (XREF_FIG), suggesting that O GlcNAcylation of PGC-1alpha facilitates BAP1 binding."

reach
"38 Furthermore, the authors found that high-glucose conditions potentiated PGC1alpha glycosylation, suggesting that this O-GlcNAc-induced PGC1alpha and BAP1 complex may provide a functional explanation for the aberrant induction of hepatic gluconeogenesis in hyperglycemic patients."

sparser
"PGC-1α can also be regulated by BAP1, which directly binds to O-GlcNAcylated PGC-1α to prevent its degradation ( xref )."
BAP1 activates PPARGC1A.
| 2
BAP1 activates PPARGC1A. 2 / 2
| 2

reach
"Bap1 overexpression in the liver slightly enhances gluconeogenesis, possibly by stabilizing Pgc1alpha."

reach
"Conversely, knockdown of BAP1 in cells (XREF_SUPPLEMENTARY) decreases the stability of PGC-1alpha (XREF_FIG)."
BAP1 affects N1ICD
| 6
BAP1 binds N1ICD. 6 / 6
| 6

sparser
"In a more stringent assay, the interaction between N1ICD and endogenous BAP1 was validated by FLAG-bound IP in HeLa N1ICD-FLAG cells ( xref )."

sparser
"Parallelly, protein levels of activated NOTCH1 was found to be highly expressed in cortical NPCs ( xref ), and BAP1 associates with endogenous activated N1ICD in cortical NPCs ( xref )."

sparser
"Interestingly, we found that the RAM–ANK and PEST domains of N1ICD, regions required for NUMB association, mediate N1ICD interaction with BAP1 as well ( xref ), and BAP1 regulates N1ICD stability through the PEST domain ( xref )."

sparser
"Moreover, the presence of full-length NUMB, but not the AB-deleted NUMB, significantly enhances the interaction between BAP1 and N1ICD."

sparser
"Finally, NUMB overexpression enhanced the interaction between BAP1 and N1ICD ( xref ), but co-expression of non-N1ICD-bound NUMB-∆2–15 could not promote BAP1N1ICD association, thus failing to boost stabilization of N1ICD ( xref )."

sparser
"Next, we asked whether BAP1 is associated with N1ICD."

reach
"Using deubiquitinase siRNA library screening based on an in vitro wound healing assay, we found that silencing BAP1 significantly augmented MSC migration."

reach
"BAP1 deficiency enhances MSC migration ."

reach
"BAP1 overexpression reduces MSC migration."

reach
"Transwell migration and invasion assays revealed that BAP1 overexpression inhibited MSC migration and invasion (Fig. 2E, F)."

reach
"We observed an increase in ERK1/2 phosphorylation in the BAP1-depleted MSC, whereas other signaling molecules (AKT serine 473, focal adhesion kinase tyrosine 925, p38 mitogen-activated protein kinases, and vimentin) remained unchanged (Fig. 3A)."

reach
"BAP1 depletion enhanced MSC migration, whereas BAP1 overexpression reduced MSC migration."
BAP1 affects MED1
| 6
MED1 binds BAP1. 6 / 6
| 6

sparser
"Med1 is associated with BAP1."

sparser
"To investigate the interaction between Med1 and BAP1 protein, co-immunoprecipitation was performed in the BAP1 overexpressing cell line, BAP1 WT and BAP1 C91A mutant were overexpressed, and IP was performed using an anti-Med1 antibody in BAP1 or BAP1 C91A overexpressed cells."

sparser
"These results demonstrate that BAP1 is associated with Med1, but not with the BAP1 catalytic mutant."

sparser
"In addition, an examination of the interaction between Med1 and BAP1 proteins showed that Med1 and BAP1 proteins are binding and are involved in the development and metastasis of cancer ( xref )."

sparser
"Although the binding of Med1 and BAP1 involved in the cancer growth or metastasis, mechanisms are still unclear."

sparser
"However, in the future, the results of this study will contribute to various research fields, such as the mechanism of regulating the expression of target genes by the binding of Med1 and BAP1 and the growth and metastasis of cancer by Med1 and BAP1."
BAP1 affects HDAC1
4 | 1 1
4 | 1 1

No evidence text available

No evidence text available

No evidence text available

sparser
"We recently discovered that in the nucleus BAP1 binds and deubiquitylates HDAC1, which in turn binds and deacetylates HMGB1, keeping HMGB1 in the nucleus."

No evidence text available

reach
"We recently discovered that in the nucleus BAP1 binds and deubiquitylates HDAC1, which in turn binds and deacetylates HMGB1, keeping HMGB1 in the nucleus."
BAP1 affects HAT1
5 | 1
5 | 1

No evidence text available

No evidence text available

No evidence text available

reach
"HAT1 and COPI interaction with BAP1 is observed in the cytoplasm and not in the nuclear fractions (Figs 1A and 1B and 4)."
| PMC

No evidence text available

No evidence text available
| 6
BAP1 deubiquitinates Deubiquitinase.
| 3
BAP1 deubiquitinates Deubiquitinase. 3 / 3
| 3

reach
"Functionally, BAP1 is a nuclear deubiquitinating (DUB) enzyme belonging to the ubiquitin C-terminal hydrolase DUB subfamily."

reach
"BAP1 can encode a nuclear deubiquitinating (DUB) enzyme that participates in the composition of the polycomb repressive deubiquitinase (PR-DUB) complex."

reach
"BAP1 belongs to the deubiquitinate (DUB) family of proteins and acts as a ubiquitin carboxy-terminal hydrolase (UCH)."
BAP1 activates Deubiquitinase.
| 3
| 3

reach
"We confirmed that re-expression of BAP1 WT, but not its C91A mutant, in UMRC6 cells decreased SLC7A11 expression (XREF_FIG), suggesting that BAP1 mediated repression of SLC7A11 expression requires BAP1 's DUB activity."

reach
"In addition to TFs, SLC7A11 transcription is also repressed by BRCA1-associated protein 1 (BAP1) – a major component of a deubiquitinase (DUB) complex which catalyzes deubiquitination of histone 2 A (H2A) associated with the SLC7A11 promoter.96 Not surprisingly, BAP1 promotes ferroptosis in a DUB-dependent manner."

reach
"Incubation of BAP1 with increasing concentrations of TG2-179-1 reduced the emission of fluorescence from Ub-AMC in a dose-dependent manner (Fig. 5C), indicating that TG2-179-1 inhibited the DUB activity of BAP1."
BAP1 affects DDR
| 2 4
BAP1 activates DDR. 6 / 6
| 2 4

reach
"Thus, BAP1 is thought to modulate DDR through H2Aub."

eidos
"Thus , BAP1 is thought to modulate DDR through H2Aub ."

reach
"Indeed, inhibition of BAP1 by short hairpin RNAs (shRNAs) impaired the DDR and caused HeLa cells to become hypersensitive to ionizing radiation resulting in S-phase retardation, a phenotype similar to BRCA1 deficiency."

reach
"In such an environment, an impaired DDR caused by BAP1 deletion, might be expected to favour the accumulation of genetic damage that eventually gives rise to a malignant clone."

reach
"BAP1 can modulate the DDR via several different interactions, including deubiquitinylation of histone H2A."

eidos
"BAP1 can modulate the DDR via several different interactions , including deubiquitinylation of histone H2A ( 6 , 7 ) ."
| 4

reach
"Individuals with pathogenic BAP1 variants face an elevated risk of diverse tumors, notably BAP1-inactivated melanocytic nevi/tumors, uveal melanoma, cutaneous melanoma, mesothelioma, renal cell carcinoma, and basal cell carcinoma."

reach
"In human uveal melanoma and renal cell carcinoma, BAP1 loss leads to a dedifferentiated stem-like phenotype associated with aggressive cancer behavior and poor patient outcome (2, 3)."

reach
"BAP1-TPDS is associated with an increased risk of a specific skin lesion, BAP1-inactivated melanocytic tumour (BIMT) (34), cutaneous melanoma (CM) (34), renal cell carcinoma (RCC) (35), and basal cell carcinoma (BCC) (36)."

reach
"Germline mutations in BAP1 predispose carriers to the BAP1 tumor predisposition syndrome (BAP1-TPDS), characterized by BAP1-inactivated nevi (BINs), uveal melanoma, cutaneous melanoma, mesothelioma, renal cell carcinoma, and other tumors."
| 2

reach
"On the other hand, the occasionally described heterozygous deletions of these genes cannot be completely excluded in our series, because NGS procedure and mode of data analysis were not sensitive to moderate CNV (copy number variation), the detection of which was not the aim of this panel.Germline mutations in BAP1 are known to underlie sometimes melanomas, renal cell carcinomas, malignant mesotheliomas, and some other cancers [31]."

reach
"BAP1 germline mutations underlie a rare, autosomal dominant tumour predisposition syndrome that is most commonly associated with (in order of decreasing frequency) atypical Spitz tumours, uveal melano[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
WWP1 affects BAP1
1 | 1 3
1 | 1 3

sparser
"Besides, there is an interaction between WWP1 and BAP1 ( Figure 4 N)."

reach
"Besides, there is an interaction between WWP1 and BAP1 ( Figure 4 N)."

sparser
"The effects of the BAP1-WWP1 axis on KLF5 protein function were then investigated."

No evidence text available

sparser
"These evidence indicated that the BAP1-WWP1 axis in melanoma played an important role in regulating the function of KLF5."
UFL1 affects BAP1
| 5
| 5

sparser
"Additionally, the structural characterization of the UFL1-BAP1 complex will be instrumental in guiding such efforts and may uncover specific binding pockets or allosteric sites that could be targeted to modulate this interaction with higher specificity."

sparser
"Considering the frequent mutations and deletions of the TP53 gene in human cancers, the stabilization of pVHL by the UFL1-BAP1 axis might be particularly important in cancer cells harboring TP53 mutant or loss, such as CRC cells (SW620 and HCT116 with p53 KO) and pancreatic cancer cells (AsPC-1) (fig."

sparser
"However, several features of the BAP1-UFL1 interaction identified in our study offer pathways to address specificity."

sparser
"Second, a more selective strategy may involve enhancing the specific interaction between UFL1 and BAP1."

sparser
"For instance, high-throughput screening of Food and Drug Administration–approved drugs or other chemical libraries could identify small molecules that promote the UFL1-BAP1 interaction, thereby selectively enhancing BAP1 UFMylation."
UBC affects BAP1
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

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No evidence text available
TERT affects BAP1
| 5
| 5

sparser
"Shorter overall survival was associated with TERT mutations ( p  = 0.03, median 29.6 months for MT versus 52.6 months for WT) or BAP1 mutations ( p  = 0.02, median 28.7 months for MT versus not reached for WT)."

sparser
"The presence of BAP1 or TERT promoter mutations was associated with a significantly worse OS."

sparser
"An exploratory analysis was performed to investigate if acquired mutations in BAP1 or TERT were associated with MSKCC prognostic risk group or the presence of sarcomatoid features on histopathologic review."

sparser
"This was also the case for mutually exclusive alterations such as SETD2–CDKN2A (3p21–9p21) and TERTBAP1 (5p15–3p21) (Fig.  xref and Supplementary Tables  xref – xref )."

sparser
"Emerging evidence (particularly for hepatocellular carcinoma) suggests that TERT could interact with BAP1 via the WNT/β-catenin signaling pathway, which is often activated in tumors with TERT overexpression."
SLC7A11 affects BAP1
| 2 3
SLC7A11 binds BAP1.
| 3
| 3

sparser
"Targeting the BAP1-SLC7A11 axis may provide novel insights into the treatment of ferroptosis-related intestinal diseases."

sparser
"Overexpressed BAP1 WT or BAP1 C91A binds directly to the Slc7a11 promoter (Fig.  xref )."

sparser
"Notably, ChIP qPCR data using BAP1 antibody indicates BAP1 binding to the Slc7a11 promoter."
SLC7A11 increases the amount of BAP1.
| 2
SLC7A11 increases the amount of BAP1. 2 / 2
| 2

reach
"p53 binds to the SLC7A11 promoter, directly suppressing its transcription; the nuclear deubiquitinase (DUB) BAP1 represses SLC7A11 transcription by removing histone 2A ubiquitination from the SLC7A11 promoter; and KEAP1 represses SLC7A11 transcription through degrading NRF2, a master transcription factor of antioxidant response and regulator of SLC7A11."

reach
"A luciferase assay with the SLC7A11 promoter showed that overexpression of BAP1 WT, but not its C91A mutant, decreased luciferase activity, whereas BAP1 knockdown increased it, indicating that BAP1 regulates SLC7A11 transcription (XREF_SUPPLEMENTARY - XREF_SUPPLEMENTARY)."
SETD2 affects KDM5C
| 5
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
NUMB affects BAP1
| 5
| 5

sparser
"We next examined whether BAP1 binds to NUMB and whether this association regulates N1ICD stability."

sparser
"First, both exogenously expressed and endogenous BAP1 could bind to endogenous NUMB ( xref )."

sparser
"Our results demonstrate that Jagged1 not only induces Notch1 activation in retinal endothelial cells but also regulates its deubiquitylation and stabilization via its interaction with Numb and BAP1."

sparser
"Third, similar to N1ICD, NUMB could bind to BAP1 N-terminal or C-terminal part ( xref )."

sparser
"IL-33 enhances Notch1 intracellular domain deubiquitination via its interaction with Numb and BAP1."
MYC affects BAP1
2 | 3
2 | 3

No evidence text available

sparser
"The fusion protein plasmid HA-BRCA1, Flag-Ub, and myc-BAP1 used for cell transfection during Co-IP were designed and synthesized by Lianmai Bioengineering Co., Ltd. (Shanghai, China)."

sparser
"To investigate the possibility of PT33 covalently binding to BAP1, we synthesized a biotinylated affinity probe-PT33–Biotin by modifying the hydroxy of PT33 with biotin (Supporting Information xref ), and purified recombinant the WT and mutant (C91A) BAP1-myc tagged protein."

sparser
"Exogenously expressed Myc-BAP1 and N1ICD could be found in the same protein complex in 293T cells ( xref )."

No evidence text available
KN-93 affects BAP1
| 5
KN-93 inhibits BAP1. 5 / 5
| 5

reach
"This result suggests that KN-93 selectively suppresses the proliferation of BAP1 cells with CAMK2D-positive expression."

reach
"KN-93 induces significant apoptosis in BAP1-deficient cells."

reach
"Similarly, the cleaved PARP level was markedly increased in the KN-93-treated BAP1 cells (Fig. 5b), suggesting that KN-93 suppresses the proliferation of BAP1 cells by promoting apoptosis."

reach
"Hence, we propose that KN-93 promotes apoptosis in BAP1-negative MMe cells by increasing intracellular Ca concentrations, most likely by inhibiting CAMK2D activity."

reach
"CAMKII inhibitor KN-93 selectively suppresses the proliferation of BAP1-deficient mesothelioma cells."
KDM5C affects SETD2
| 5
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
Hypoxia affects BAP1
| 5
Hypoxia activates BAP1. 5 / 5
| 5

reach
"In this study, we found that hypoxia-induced BAP1 enhances erastin-induced ferroptosis in NPC by stabilizing H2A.Studies have shown that many tumor cells under a hypoxic microenvironment are less sensitive to chemotherapeutic agents because these agents typically require oxygen for maximum activity [12]."

reach
"Hypoxia-induced BAP1 enhances erastin-induced ferroptosis in nasopharyngeal carcinoma by stabilizing H2A."

reach
"Hypoxia-induced BAP1 enhances erastin-induced ferroptosis in NPC by stabilizing H2A."

reach
"Ferroptosis inducers targeting BAP1 may be an effective way to reduce drug resistance in NPC, especially in the hypoxic microenvironment.In conclusion, our study initially validated that hypoxia-induced BAP1 enhanced erastin-induced ferroptosis in NPC by stabilizing H2A (Fig. 7)."

reach
"We found that hypoxia significantly increased the mRNA and protein expression levels of BAP1 in both HNE1 and 6-10B cells after exposed to hypoxia for 24 h, whereas other 5 ferroptosis-related genes showed no significant difference (Fig. 3A and B)."
Histone affects BAP1
| 2 1
| 2 1

reach
"Complex formation between BAP1 and the cell cycle regulator, HCF-1, bridges HCF-1 to histone-modifying complexes to regulate cell growth (2,4)."

sparser
"ASXL3 is less well studied than its paralogs but was shown to promote BAP1 binding to its histone substrate like its paralogs [ xref ]."
| PMC

reach
"We find that ASXL1 directs the binding of the positively charged C-terminal extension of BAP1 to nucleosomal DNA and histones H3-H4 near the dyad, an addition to its role in forming the ubiquitin-binding cleft."
HMGB1 affects BAP1
4 | 1
4 | 1

No evidence text available

sparser
"We linked these findings to our discovery that BAP1 forms a trimeric protein complex with HMGB1 and with histone deacetylase 1 (HDAC1) that modulates HMGB1 acetylation and its release."

No evidence text available

No evidence text available

No evidence text available
GST affects BAP1
| 5
| 5

sparser
"Of these 30 human proteins detected, 8 proteins showed enrichment upon incubation with the nuclear protein (Fig. xref , right panel), while 22 proteins had higher levels in the GST-BAP1 fusion protein alone group, which likely represented background noise."

sparser
"Additionally, purified GST-BAP1, but not GST control, was able to bind to His-UFL1 in vitro ( xref )."

sparser
"To identify the domains responsible for LKB1 interaction, we constructed and purified GST-BAP1 deletion constructs."

sparser
"Additionally, incubation with purified GST-BAP1 WT in vitro, but not the C91S mutant, caused a notable reduction in polyubiquitinated pVHL (fig."

sparser
"We observed some differential bands above 95 kDa in the group of GST-BAP1 fusion protein incubated with PLC/PRF/5 nuclear protein, as compared with the group of GST-BAP1 fusion protein only (Fig. xref , left panel)."
E3_Ub_ligase affects BAP1
| 5

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"One of these is BAP1 which binds the BRCA1 ubiquitin ligase and functions as a tumor suppressor ( Ventii et al., 2008 )."

sparser
"In addition, ubiquitin ligase, which is closely associated with BAP1 protein, is a good candidate therapeutic target."
DIDO1 affects BAP1
3 | 1 1
3 | 1 1

No evidence text available

sparser
"BAP1 binds to DIDO1 and stabilizes DIDO1 through de-ubiquitination."

reach
"BAP1 binds to DIDO1 and stabilizes DIDO1 through de-ubiquitination."

No evidence text available

No evidence text available
CRISPR affects BAP1
| 5
CRISPR inhibits BAP1. 5 / 5
| 5

reach
"CRISPR/Cas9-mediated knockout of BAP1.."

reach
"We also identified an association of clinical BAP1 mutations with high B7H4/3 expression, and validated it as a causal relationship between CRISPR-mediated BAP1 loss and B7H4 upregulation in iCCA cell line models."

reach
"B. Artegiani et al. described that liver organoids with four common cholangiocarcinoma mutations (TP53, PTEN, SMAD4, and NF1) gained malignancy after loss-of-function of BAP1 by CRISPR/Cas9, because BAP1 could control the expression of junctional and cytoskeleton components by regulating chromatin accessibility (44)."

reach
"Artegiani et al. found that after loss-of-function of BAP1 by CRISPR/Cas9 in normal human cholangiocyte organoids, the cells became more motile and fused with other organoids, features that resemble the metastatic invasion of cancer."
| PMC

reach
"CRISPR/Cas9-mediated knockout of BAP1 in PANC1 murine pancreatic cancer cells and HEK 293 T cells induced chromosome abnormalities, including chromosome breaks, shattering and aneuploidy, and these effects were exacerbated by cell exposure to IR ."
CBX3 affects BAP1
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
CAMK2D affects BAP1
3 | 2
3 | 2

No evidence text available

No evidence text available

sparser
"Moreover, BAP1 -KO had a negligible effect on cell proliferation, whereas CAMK2D / BAP1 -DKO reduced cell proliferation, suggesting that an emerging BAP1-CAMK2D axis might play a critical role in the tumor growth of BAP1 -deficient MMe cells."

sparser
"Therefore, targeting the BAP1-CAMK2D (CAMKII delta) signaling axis is a promising therapeutic approach for most MMe cases and BAP1 tumor predisposition syndrome."

No evidence text available
Mutated BAP1 inhibits signal transduction. 5 / 5
| 5

reach
"These previous studies can support our finding that BAP1 mutations causing loss of BAP1 expression could induce the immunosuppressive microenvironment in UM.Mechanistically, we demonstrated in the current study that BAP1 mutations could inhibit the NF-κB signaling pathway, thereby repressing the cytokine secretion and antigen presentation by macrophages and inducing immunosuppressive microenvironment in UM."

reach
"A preliminary conclusion can be drawn in the current study that BAP1 mutations might repress the NF-κB signaling pathway, which repressed the cytokine secretion and antigen presentation by macrophages, inducing the immunosuppressive microenvironment, augmenting the malignant phenotypes of UM cells and ultimately promoting the growth and metastasis of UM (Fig. 10)."

reach
"BAP1 mutations inhibit the NF-kappaB signaling pathway to induce an immunosuppressive microenvironment in uveal melanoma."

reach
"BAP1 mutations inhibited the NF-kappaB signaling pathway, suppressing the cytokine secretion and antigen presentation by macrophages."

reach
"BAP1 mutations may inhibit the NF-kappaB signaling pathway, repressing the cytokine secretion and antigen presentation by macrophages, which induces the immunosuppressive microenvironment, enhances the malignant phenotypes of UM cells and ultimately promotes the growth and metastasis of UM."
BAP1 affects lactose
| 5
BAP1 inhibits lactose. 5 / 5
| 5

reach
"Functionally, BAP1 promotes apoptosis and necrosis, and down-regulates the migration and invasion abilities of LAC cells."

reach
"In functional assays, BAP1 was found to inhibit the migration and invasion of LAC cells, and promoted their apoptosis and necrosis."

reach
"BAP1 may inhibit the progression of LAC by attenuating tumor invasiveness, a function that was supported by the findings of the Transwell assay."

reach
"However, BAP1 is closely related to lymph node status, suggesting that BAP1 also mediates suppression of cell migration in LAC which is in keeping with the findings of the wound healing assay."

reach
"Finally, to elucidate the mechanism by which BAP1 suppresses invasion and metastasis in LAC cells, we analyzed the expression levels of metastasis related proteins in transfected H1299 cells and H1650 cells using IB, respectively."
BAP1 affects WWP1
1 | 1 3
1 | 1 3

sparser
"Besides, there is an interaction between WWP1 and BAP1 ( Figure 4 N)."

reach
"Besides, there is an interaction between WWP1 and BAP1 ( Figure 4 N)."

sparser
"The effects of the BAP1-WWP1 axis on KLF5 protein function were then investigated."

No evidence text available

sparser
"These evidence indicated that the BAP1-WWP1 axis in melanoma played an important role in regulating the function of KLF5."
BAP1 affects UFL1
| 5
| 5

sparser
"Additionally, the structural characterization of the UFL1-BAP1 complex will be instrumental in guiding such efforts and may uncover specific binding pockets or allosteric sites that could be targeted to modulate this interaction with higher specificity."

sparser
"Considering the frequent mutations and deletions of the TP53 gene in human cancers, the stabilization of pVHL by the UFL1-BAP1 axis might be particularly important in cancer cells harboring TP53 mutant or loss, such as CRC cells (SW620 and HCT116 with p53 KO) and pancreatic cancer cells (AsPC-1) (fig."

sparser
"However, several features of the BAP1-UFL1 interaction identified in our study offer pathways to address specificity."

sparser
"Second, a more selective strategy may involve enhancing the specific interaction between UFL1 and BAP1."

sparser
"For instance, high-throughput screening of Food and Drug Administration–approved drugs or other chemical libraries could identify small molecules that promote the UFL1-BAP1 interaction, thereby selectively enhancing BAP1 UFMylation."
BAP1 affects UBC
5 |
5 |

No evidence text available

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BAP1 affects RAD51
| 5
BAP1 activates RAD51. 5 / 5
| 5

reach
"BAP1 can deubiquitylate Ub-H2AK119 at the site of UV-induced DNA damage, and promote DNA repair by regulating the accumulation of repair proteins (BRCA1, RAD51, and RPA) (7)."

reach
"BAP1 depletion abrogates the binding of INO80 to replication forks and increases the formation of RAD51 foci following HU treatment."

reach
"BAP1 also promotes the assembly of Rad51 foci to facilitate DNA repair and replication fork restart during replication stress ."

reach
"BAP1 depletion prevented RAD51 from binding to chromatin and forming replication foci after HU treatment."

reach
"Depletion of BAP1 leads to reduced BRCA1 foci, and reduced RAD51 loading, with reduced HR repair and increased IR sensitivity coinciding with elevated H2A and H2AX ubiquitylation."
BAP1 affects NUMB
| 5
| 5

sparser
"We next examined whether BAP1 binds to NUMB and whether this association regulates N1ICD stability."

sparser
"First, both exogenously expressed and endogenous BAP1 could bind to endogenous NUMB ( xref )."

sparser
"Our results demonstrate that Jagged1 not only induces Notch1 activation in retinal endothelial cells but also regulates its deubiquitylation and stabilization via its interaction with Numb and BAP1."

sparser
"Third, similar to N1ICD, NUMB could bind to BAP1 N-terminal or C-terminal part ( xref )."

sparser
"IL-33 enhances Notch1 intracellular domain deubiquitination via its interaction with Numb and BAP1."
BAP1 affects MYC
2 | 3
2 | 3

No evidence text available

sparser
"The fusion protein plasmid HA-BRCA1, Flag-Ub, and myc-BAP1 used for cell transfection during Co-IP were designed and synthesized by Lianmai Bioengineering Co., Ltd. (Shanghai, China)."

sparser
"To investigate the possibility of PT33 covalently binding to BAP1, we synthesized a biotinylated affinity probe-PT33–Biotin by modifying the hydroxy of PT33 with biotin (Supporting Information xref ), and purified recombinant the WT and mutant (C91A) BAP1-myc tagged protein."

sparser
"Exogenously expressed Myc-BAP1 and N1ICD could be found in the same protein complex in 293T cells ( xref )."

No evidence text available
BAP1 affects HMGB1
4 | 1
4 | 1

No evidence text available

sparser
"We linked these findings to our discovery that BAP1 forms a trimeric protein complex with HMGB1 and with histone deacetylase 1 (HDAC1) that modulates HMGB1 acetylation and its release."

No evidence text available

No evidence text available

No evidence text available
BAP1 affects GST
| 5
| 5

sparser
"Of these 30 human proteins detected, 8 proteins showed enrichment upon incubation with the nuclear protein (Fig. xref , right panel), while 22 proteins had higher levels in the GST-BAP1 fusion protein alone group, which likely represented background noise."

sparser
"Additionally, purified GST-BAP1, but not GST control, was able to bind to His-UFL1 in vitro ( xref )."

sparser
"To identify the domains responsible for LKB1 interaction, we constructed and purified GST-BAP1 deletion constructs."

sparser
"Additionally, incubation with purified GST-BAP1 WT in vitro, but not the C91S mutant, caused a notable reduction in polyubiquitinated pVHL (fig."

sparser
"We observed some differential bands above 95 kDa in the group of GST-BAP1 fusion protein incubated with PLC/PRF/5 nuclear protein, as compared with the group of GST-BAP1 fusion protein only (Fig. xref , left panel)."
BAP1 affects GPX4
| 5
BAP1 decreases the amount of GPX4. 5 / 5
| 5

reach
"In this study, the overexpression of BAP1 significantly inhibited GPX4 expression and exacerbated the degree of lipid peroxidation and ferroptosis in neurons after SAH."

reach
"Furthermore, FOXO3a inhibition also inhibited the mRNA and protein expression levels of BAP1, while simultaneously increasing the levels of SLC7A11 and GPX4 after SAH in mice (Fig. 8C–F)."

reach
"Moreover, as revealed by western blot assay, knockdown of BAP1 also upregulated the expression of GPX4 reduced by LPS (Fig. 5H and I; CON vs. LPS, p = 0.002; NC(-) vs. BAP1(-), p = 0.004) and downregulated the expression of ASCL4 elevated by LPS (Fig. 5H and J; CON vs. LPS, p = 0.002; NC(-) vs. BAP1(-), p = 0.01), which were the marker proteins of ferroptosis."

reach
"In addition, overexpression of BAP1 reversed the effects of Exos , decreased the expression levels of SLC7A11 and GPX4, and increased the ACSL4, which was analyzed by RT-qPCR and western blot (Figures 6F and 6G)."

reach
"Our results revealed that transfection with H2Aub K119R mutant plasmid reduced the transcription and protein expression of SLC7A11 and GPX4, and overexpression of BAP1 can promote neuronal ferroptosis by regulating H2Aub levels."
BAP1 affects Death
| 4
BAP1 inhibits Death.
| 3
BAP1 inhibits Death. 3 / 3
| 3

reach
"Recent evidence highlights that BAP1 suppresses metabolic stress-induced death in cancer cells (70,71)."

reach
"We discovered that fetal loss of the epigenetic regulator Bap1 in the ENS lineage caused severe postnatal bowel dysfunction and early death in Tyrosinase-Cre Bap1 fl/fl mice."

reach
"For example, differentiation to mature neurons and glia could require antagonism of PRC-mediated transcriptional repression.We were thus intrigued by our serendipitous discovery that Bap1 deletion in ENS lineages caused profound bowel dysfunction and early death in mice, mimicking human nCIPO or HSCR physiology."
BAP1 activates Death.
| 1
BAP1 activates Death. 1 / 2
| 1

reach
"As a tumor suppressor, BAP1 can target SLC7A11, reduce the H2Aub occupancy on the SLC7A11 promoter, inhibit the function of the amino acid transport complex system Xc-, and induce cancer cell death by promoting lipid peroxidation in cancer cells [16]."
| 5

reach
"BAP1 contributes to colon cancer cell proliferation by accelerating DNA replication and suppressing replication stress and concomitant apoptosis."

reach
"BAP1 depletion reduces colon cancer cell proliferation concomitant with increased apoptosis and defective DNA replication."

reach
"However, our previous work showed that the stability of Ino80 is regulated by the ubiquitin-proteasome system and that BAP1 functions as a deubiquitinase that stabilizes Ino80 and thus promotes DNA replication."

reach
"Therefore, it is unlikely that BAP1 contributes to the survival and proliferation of colon cancer cells via the Apc/β-catenin pathway.BAP1 functions in both normal DNA replication and replication stress recovery."

reach
"Our data instead showed that BAP1 depletion suppressed replication fork progression with concomitant induction of replication stress and apoptosis, suggesting that BAP1 promotes colon cancer cell proliferation by increasing the rate of DNA replication."
BAP1 affects DIDO1
3 | 1 1
3 | 1 1

No evidence text available

sparser
"BAP1 binds to DIDO1 and stabilizes DIDO1 through de-ubiquitination."

reach
"BAP1 binds to DIDO1 and stabilizes DIDO1 through de-ubiquitination."

No evidence text available

No evidence text available
BAP1 affects CDKN1B
| 5
BAP1 increases the amount of CDKN1B. 5 / 5
| 5

reach
"Depletion of KLF5, BAP1 or HCF-1 also upregulated p27 mRNA levels (XREF_FIG)."

reach
"Indeed, depletion of KLF5, BAP1, HCF-1 or OGT upregulated p27 protein expression and blocked cell cycle progression (XREF_FIG)."

reach
"Knockdown of either KLF5 or BAP1 upregulated p27 protein levels in these breast cell lines as well (XREF_FIG; XREF_SUPPLEMENTARY)."

reach
"Given that depletion of BAP1 or KLF5 markedly increased p27 protein levels, we speculated that the protein complex might regulate the expression of p27 and FGF-BP (a KLF5 target gene)."

reach
"Knockdown of KLF5, BAP1 or HCF-1, but not OGT, significantly increased p27 mRNA levels (XREF_FIG)."
BAP1 affects CBX3
5 |
5 |

No evidence text available

No evidence text available

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BAP1 affects CAMK2D
3 | 2
3 | 2

No evidence text available

No evidence text available

sparser
"Moreover, BAP1 -KO had a negligible effect on cell proliferation, whereas CAMK2D / BAP1 -DKO reduced cell proliferation, suggesting that an emerging BAP1-CAMK2D axis might play a critical role in the tumor growth of BAP1 -deficient MMe cells."

sparser
"Therefore, targeting the BAP1-CAMK2D (CAMKII delta) signaling axis is a promising therapeutic approach for most MMe cases and BAP1 tumor predisposition syndrome."

No evidence text available
BAP1 affects ASXL1/2
| 4 1
BAP1 binds ASXL1/2. 5 / 5
| 4 1

reach
"The KMT2C/D-COMPASS-like complex is recruited to enhancers by the BAP1-ASXL1/2 complex, which facilitates enhancer priming (17,18)."

reach
"The tumor suppressor BAP1 associates with ASXL1/2 to form the core Polycomb complex PR and DUB, which catalyzes the removal of mono-ubiquitin from several substrates including histone H2A."

reach
"Cancer-associated BAP1 mutations lead loss of DUB activity and impair function of the BAP1-ASXL1/2 complex, causing an elevation in H2A ubiquitination levels, and subsequent cell cycle progression dysregulation (3)."

reach
"The core BAP1 and ASXL1/2 complex interacts with host cell factor-1 (HCF-1) and histone modifying complexes during cell division."

sparser
"This work provides a structural basis upon which to interpret previous studies showing that the ASXL1/2 Deubad domains bind tightly to BAP1, and activate the complex by forming a composite binding site for ubiquitin xref , xref ."
Glucose affects BAP1
| 4
Glucose decreases the amount of BAP1. 4 / 4
| 4

reach
"HG inhibits cystine uptake in podocytes by promoting the expression of BAP1 and inhibiting H2Aub deubiquitination on SLC7A11, a functional protein targeting cystine transporter system Xc-on the cell membrane, leading to the deposition of lipid peroxides and the occurrence of ferroptosis in podocytes."

reach
"HG inhibits cystine uptake in podocytes by promoting the expression of BAP1 and inhibiting H2Aub deubiquitination on SLC7A11, leading to lipid peroxide accumulation and ferroptosis in podocytes."

reach
"These findings suggest that HG inhibits cystine uptake by inhibiting SLC7A11 expression on the cell membrane, thereby influencing the regulation of ferroptosis.3.4 HG reduced SLC7A11 function by up-regulating BAP1 expression."

reach
"These results suggest that HG inhibits SLC7A11 function by promoting BAP1 expression, thereby inducing ferroptosis in podocytes.3.5 HG promotes ferroptosis by promoting BAP1 deubiquitination of H2Aub on SLC7A11."
YWHAZ affects BAP1
4 |
4 |

No evidence text available

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STMN1 affects BAP1
1 | 3
1 | 3

reach
"Semi-endogenous co-immunoprecipitation of SMN with intact or truncated Bap1 revealed that the ubiquitin C-terminal hydrolase domain of Bap1 is essential for the binding of Bap1 to SMN (Supplemental Figure 7F)."

reach
"Taken together, we concluded that Bap1 directly binds SMN and prevents SMN degradation by posttranslational deubiquitination of SMN.Since a lysine residue is one of the predominant targets for ubiquitination, we examined which of the lysine residues in SMN is critical for its ubiquitin-dependent degradation by introducing a non-lysine mutation on phylogenetically conserved lysine residues (Supplemental Figure 8A)."

No evidence text available

reach
"These results indicate that Bap1 posttranscriptionally regulates SMN levels and thus affects the function of SMN in FAPs.To test the direct interaction between Bap1 and SMN, we performed endogenous reciprocal immunoprecipitation experiments using freshly isolated FAPs and revealed that Bap1 directly binds SMN in FAPs (Figure 7C)."
PSME3 affects BAP1
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PRKDC affects BAP1
1 | 1 2
1 | 1 2

sparser
"Our results suggest reciprocal positive interactions between BAP1 and DNA-PKcs, based on phosphorylation of BAP1 by the latter and deubiquitination of DNA-PKcs by BAP1."

sparser
"Thus, functional interaction of BAP1 with DNA-PKcs supports a role for BAP1 in NHEJ DNA repair and may provide the basis for new therapeutic strategies and new insights into its role as a tumor suppressor."

reach
"Our results suggest reciprocal positive interactions between BAP1 and DNA-PKcs, based on phosphorylation of BAP1 by the latter and deubiquitination of DNA-PKcs by BAP1."

No evidence text available
PRC2_complex affects BAP1
| 4

sparser
"Thus, our analysis and experimental validation across mouse and human mesothelioma cells clearly identify a set of genes regulated via the BAP1-PRC2 axis."

sparser
"BAP1 does not interact with PRC2 proteins ( xref )."

sparser
"ASXL1 functions in transcriptional repression through its interaction with PRC2 and BAP1. xref BCORL1 is a transcriptional co-repressor that interacts with PCGF1, a core complex of the PRC1.1 complex. xref The RUNX1-CBF-b heterodimer mediates transcription by binding to RUNX sites, but also represses transcription by interacting and recruiting BMI1 of the PRC1 complex to target sites. xref IKZF1 regulates transcription by interacting with repressive epigenetic complexes such as HDAC1, HDAC2, CHD3, CHD4, and SWI/SNF complex, and also recruits PRC2 to target gene loci in T cells. xref Thus, while the commonly mutated genes in CML have their own exclusive roles in transcriptional regulation, they also share a striking commonality as modulators of the PRC."

sparser
"In addition, BAP1 mutations in ccRCC tumors were associated with downregulated expression of target genes of polycomb repressive complex 2 (PRC2) [ xref ] ( xref )."
PRC1 affects BAP1
| 1 3
| 1 3

sparser
"Given that the composition and expression of PRC1 and BAP1 complexes changes extensively during development ( xref ; xref , xref ; xref ; xref ), in future work, it will be interesting to investigate how the balance between these two opposing activities is regulated at different developmental stages and how cell type-specific H2AK119ub1 levels influence the transcriptional potential of the genome."

sparser
"Recent CRISPR screening studies by Dixit’s group further identified that depletion of PRC1 subunit Ring1B (but not Ring1A) could rescue the cell death induced by loss of BAP1 xref , indicating that there is a robust and direct epigenetic dynamic occurring between the PRC1 and BAP1 complexes that require a balanced state to properly regulate gene expression and determine cell fate."

reach
"lines used in our study identified all these PR-DUB components but failed to identify any PRC1 component as BAP1 interacting proteins, arguing against a physical interaction between BAP1 and PRC1."

sparser
"PRC1 regulates SLC7A11 expression and H2Aub occupancy on the SLC7A11 promoter. (a) ChIP–qPCR showing the binding of BMI-1 and H2Aub on the SLC7A11 promoter, and decreased binding of H2Aub on the SLC7A11 promoter upon BMI-1 knockdown. (b) ChIP– qPCR showing the binding of RNF2 and H2Aub on the SLC7A11 promoter, and decreased binding of H2Aub on the SLC7A11 promoter upon RNF-2 knockdown. (c-f) BMI-1 or RNF2 deficiency increased SLC7A11 mRNA (c, d) and protein levels (e, f) in UMRC6 cells. (g-h) BMI-1 (g) or RNF2 (h) deficiency in 786-O cells increased SLC7A11 mRNA levels. *, p < 0.05; **, p < 0.01; ***, p < 0.001; ****, p < 0.0001. lines used in our study identified all these PR-DUB components but failed to identify any PRC1 component as BAP1-interacting proteins, arguing against a physical interaction between BAP1 and PRC1."
PPARGC1A affects BAP1
| 2 2
| 2 2

sparser
"Conversely, O -GlcNAcylation of the transcriptional regulator PGC1α facilitates the binding of the deubiquitinase BAP1 to PGC1α, leading thus to its stabilization by blocking its ubiquitination ( xref )."

reach
"PUGNAc treatment promotes the interaction between PGC-1alpha and BAP1, while it has no effect on the S333A mutant (XREF_FIG), suggesting that O GlcNAcylation of PGC-1alpha facilitates BAP1 binding."

reach
"38 Furthermore, the authors found that high-glucose conditions potentiated PGC1alpha glycosylation, suggesting that this O-GlcNAc-induced PGC1alpha and BAP1 complex may provide a functional explanation for the aberrant induction of hepatic gluconeogenesis in hyperglycemic patients."

sparser
"PGC-1α can also be regulated by BAP1, which directly binds to O-GlcNAcylated PGC-1α to prevent its degradation ( xref )."
PDH affects BAP1
| 4
PDH activates mutated BAP1. 4 / 4
| 4

reach
"These results indicate that inhibition of PDH phosphorylation by PDHK1 inhibitions decreases BAP1 mutant UM cell growth via cell cycle arrest."

reach
"Moreover, we shown that inhibition of PDH phosphorylation reduces BAP1 mutant UM cell growth by induction of cell cycle arrest."

reach
"Suppression of PDH phosphorylation significantly decreases UM cell growth and glycolytic abilities of BAP1 mutant UM cells, but not BAP1 wild type UM cells."

reach
"Our results suggest that PDH inactivation causes increased glucose utilization in BAP1 mutant UM."
PARP3 affects BAP1
| 4
| 4

sparser
"Several studies and clinical trials [ xref ], have shown that PARP inhibitors influence cancers in which mutations in BRCA1 or BRCA2 are observed, which led us to assume that the cancerous growth-inhibiting interaction of BAP1 with BRCA1 may already be perturbed in this case, and that PARP inhibitors may actually be blocking the novel interaction of BAP1 with PARP3 which enhances cancer growth."

sparser
"Thus, we suspect that the novel interaction of BAP1 and PARP3 may also be perturbed by PARP inhibitors, leading to inhibition of cancer growth."

sparser
"In this scenario, it is possible that the novel interaction of BAP1 with PARP3 in cancerous cells may be promoting cancerous growth, possibly through regulation of DNA repair and apoptosis."

sparser
"Perturbation of the interaction of BAP1 with PARP3 , using PARP3 inhibitors, may then suppress cancerous growth, at least in sarcomatoid MPM."

sparser
"The demonstration that BAP1 mutations are strongly associated with MM has provided a real breakthrough in the study of genetic predisposition in MM, that may explain why only a fraction of asbestos-exposed individuals go on to develop MM."

sparser
"BAP1 knockdown in MM cell lines has been paradoxically associated to a decreased proliferation, with an accumulation of cells in the S phase of the cell cycle xref , suggesting that BAP1 loss might promote tumorigenesis inducing a delayed, but more permissive, G1/S checkpoint xref ."

sparser
"Here, we tested the hypothesis that MM associated with germline BAP1 mutations has a better prognosis compared with sporadic MM."

sparser
"The association of germline BAP1 mutations with familial MM has been confirmed in a series of recent reports ( xref - xref )."
MP65 affects BAP1
| 4
BAP1 binds MP65. 4 / 4
| 4

sparser
"Since we have demonstrated MP65 have higher glycolysis, glycolytic capacity and increased nucleotide biosynthesis metabolites compared to MP65-BAP1, we examined key enzymes involved in glycolysis and the PPP."

sparser
"Both inhibitors diminished MP65 and MP65-BAP1 growth; however, MP65 were more sensitive to these compounds compared to MP65-BAP1 ( xref )."

sparser
"In addition, these inhibitors did not alter MP65 and MP65-BAP1 growth ( xref )."

sparser
"We found that MP65 (OXPHOS high ) were more resistant to both metabolic stress environments than MP65-BAP1, whereas MP46 (OXPHOS low ) were more vulnerable to stressful conditions compared to MP46-BAP1 ( xref and xref )."
MBD4 affects BAP1
| 3 1
| 3 1

reach
"In addition, an examination of the interaction between Med1 and BAP1 proteins showed that Med1 and BAP1 proteins are binding and are involved in the development and metastasis of cancer (Figure 3)."

reach
"Although the binding of Med1 and BAP1 involved in the cancer growth or metastasis, mechanisms are still unclear."

sparser
"Epidemiological studies have also shown an increased risk of prostate cancer, thyroid cancer, and leukemia among UM patients, which is not currently associated with BAP1 or MBD4 ( xref , xref )."

reach
"However, in the future, the results of this study will contribute to various research fields, such as the mechanism of regulating the expression of target genes by the binding of Med1 and BAP1 and the growth and metastasis of cancer by Med1 and BAP1."
L3MBTL2 affects BAP1
2 | 1 1
2 | 1 1

No evidence text available

reach
"By targeting atypical polycomb protein L3MBTL2, BAP1 interacts with and stabilizes L3MBTL2, co-occupying and maintaining H4K20me1 at target gene loci, such as EZH2 locus."

sparser
"By targeting atypical polycomb protein L3MBTL2, BAP1 interacts with and stabilizes L3MBTL2, co-occupying and maintaining H4K20me1 at target gene loci, such as EZH2 locus."

No evidence text available
HDAC affects BAP1
| 4
HDAC inhibits BAP1. 4 / 4
| 4

reach
"We then investigated whether HDAC inhibition could reverse the phenotypic consequences of BAP1 loss : loss of melanocytic differentiation and acquisition of the class 2 gene expression profile."

reach
"Histone deacetylase (HDAC) inhibitors, such as the readily available valproic acid, can reverse this effect of BAP1 loss and may play a role in treated patients with high-risk class 2 uveal melanomas in an adjuvant setting prior to the emergence of overt metastatic disease [XREF_BIBR]."

reach
"Pharmacologic inhibition of histone deacetylase activity rescues Bap1-deficient gastrulation phenotype."

reach
"Histone deacetylase (HDAC) inhibitors have been shown to reverse the phenotypic effects of BAP1 loss by inducing morphologic differentiation and transition from a high-risk to a low-risk gene expression profile in uveal melanoma cells."
HCFC2 affects BAP1
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
H2AK119Ub affects BAP1
| 4
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

reach
"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

reach
"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

reach
"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

reach
"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
H2AC20 affects BAP1
3 | 1
3 | 1

sparser
"Broader applicability of PRC2 inhibition to the treatment of other cancers is supported by the general prevalence of mutations in other chromatin-associated proteins that perturb the balance of activities regulating H3K27me3, including components of the SWI/SNF chromatin remodeling complex and the BAP1 H2A deubiquitinase complex ( xref )."

No evidence text available

No evidence text available

No evidence text available
EED affects ASXL1
| 4
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
E2F affects BAP1
| 4
| 4

sparser
"Presently, however, direct evidence of BAP1 interacting with ubiquitinated E2F transcription factors is still lacking."

sparser
"In addition, co-immunoprecipitation experiments have shown that BAP1 directly associates with E2F family members, suggesting a possible additional direct control of these transcriptional factors by BAP1 xref ."

sparser
"The results here demonstrated that BAP1 binds to E2F-responsive promoters, indicating a role in epigenetic regulation."

sparser
"14 At present, it is not known whether BAP1 directly interacts with ubiquitinated E2F transcription factors."
COPE affects BAP1
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
CCDC6 affects BAP1
2 | 2
2 | 2

sparser
"The interaction between CCDC6 and BAP1 was confirmed in MPM cells, by co-immunoprecipitation."

No evidence text available

sparser
"Coiled-coil domain containing 6 (CCDC6) interacts with BAP1 and has been reported to regulate both homologous recombination and PARP inhibitor sensitivity."

No evidence text available
CAMP affects BAP1
| 4
| 4

sparser
"However, the estimated charge of the Bap1 protein monomer is −12.9 at pH 7.0 (), providing a possible explanation for the binding of the LL-37 peptide to the negatively charged target protein, the Bap1 protein, by an electrostatic interaction."

sparser
"The mechanism(s) of interaction between the LL-37 and the Bap1 protein is under investigation in our laboratory."

sparser
"Following LL-37 binding by Bap1, the concentration of free LL-37 is reduced to sub-lethal concentrations, leading to the apparent resistance and survival of V. cholerae ."

sparser
"We suggest that due to PmB treatment of the bacterial strain, the OmpT LDV residues become exposed during biogenesis on the surface of the OMVs, allowing for binding of the Bap1 protein and thereby promoting LL-37 binding to Bap1."
BAP1 affects radioresistance
| 4
BAP1 activates radioresistance. 4 / 4
| 4

eidos
"In conclusion , we found that BAP1 can deubiquitinate H2Aub and induces radioresistance in HNSCC cells via modulation of HR ."

eidos
"We observed that 2 Gy x 5 days of radiation treatment had minimal effect on control tumors ( Fig. 5 ) but led to a significant decrease in growth of BAP1-KD tumors ( Fig. 5 ; P < 0.05 ) , suggesting that BAP1 mediated radioresistance in this xenograft model ."

eidos
"Thus , BAP1 seems to regulate H2Aub in HNSCC cells and may be a mechanism by which BAP1 induces radioresistance via facilitating DNA repair ."

eidos
"BAP1 was found to mediate radioresistance in an in vivo xenograft model and HNSCC cell lines via the deubiquitination of H2A and modulation of HR ."
BAP1 affects mutations
| 4
BAP1 inhibits mutations. 4 / 4
| 4

sparser
"All carriers of inherited heterozygous germline BAP1 inactivating mutations (BAP1 +/- )developed one and often severalBAP1 -/- malignancies in their lifetime xref , mostly malignant mesothelioma (MM), uveal melanoma (UVM) xref , xref , etc xref - xref ."

sparser
"Moreover, 2 of the 3 KDM1B mutations in LUSC and LUAD are inactivating, and these are also exclusive of BAP1 inactivating mutations, suggesting that KDM1B mutations might play a role in cancer."

sparser
"BAP1 has inactivating mutations in approximately 47% of primary UMs and 84% of metastatic UMs, consistent with the association of BAP1 mutations with inferior outcomes. xref SF3B1 encodes for splicing factor 3B subunit 1, which is involved in pre-messenger RNA splicing."

sparser
"That being said, it is clear that many metastatic UM cases demonstrate chromosome 3 loss, polysomy 8q, and have BAP1 inactivating mutations."

reach
"Inactivation of BAP1 in cancer cells results in evaluated SLC7A11 expression, increased cystine uptake and GSH synthesis, and enhanced ferroptosis resistance, which contributes to tumor development."

reach
"The large number of genome editing tools and techniques available for E. coli [60,61] allow quick and precise manipulation of this organism for biotechnological applications.In the context of NP biosynthesis in E. coli, metabolic engineering has been used to build chassis strains such as HM0079 and BAP1, that are optimized to support the synthesis of complex NPs by expressing phosphopantetheine transferase, an enzyme crucial for the function of the polyketide synthases and nonribosomal peptide synthetases [9,10]."

reach
"These results highlight a mechanism by which BaP1 induces PGE biosynthesis and suggest that activation of mPGES-1 gene expression is the primary mechanism leading to an increase in this protein."

reach
"As activation of this receptor by PGE can induce synthesis of other inflammatory proteins, including COX-2 and mPGES-1, we investigated the involvement of EP4 in BaP1-induced effects."
BAP1 affects asbestos
| 4
BAP1 activates asbestos. 4 / 4
| 4

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"In mouse models, inactivation of only one BAP1 allele increases asbestos tumorigenicity."

reach
"Currently, the available human data leans toward the notion that germline mutations (BAP1 and others) may induce mesothelioma independent of asbestos exposure (Attanoos et al., 2018)."

reach
"Mutations in BAP1, detected in 30-60% of MPM cases, accelerate asbestos-induced MPM in mice [1]."

reach
"The fact that we determined that highly penetrant, heterozygous pathogenic BAP1 null variants by themselves, as studied by Kadariya et al. , can initiate and drive MM tumorigenesis in mice absent asbestos exposure supports their conclusions, in agreement with Chanock ."
BAP1 affects YWHAZ
4 |
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BAP1 affects TNFSF10
| 4
BAP1 activates TNFSF10. 4 / 4
| 4

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"While we highlight the extrinsic apoptotic pathway and proteins as being significantly altered by BAP1 status, identifying a single factor to explain BAP1 induced TRAIL resistance is extremely challenging."

reach
"BAP1 modulates TRAIL sensitivity through PR-DUB activity."

reach
"To elucidate the mechanism by which BAP1 modulates sensitivity to TRAIL we generated expression vectors containing wild-type or mutant forms of BAP1, each with an inactive functional site or protein binding domain."

reach
"rTRAIL sensitivity of the parental BAP1-null H226 MM line was significantly diminished following expression of wild-type BAP1 and each of the mutant constructs except those with an inactive deubiquitinating or ASXL protein binding site (XREF_FIG), implicating the function of these sites in BAP1 induced TRAIL resistance."
BAP1 affects PSME3
4 |
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BAP1 affects PRKDC
1 | 1 2
1 | 1 2

sparser
"Our results suggest reciprocal positive interactions between BAP1 and DNA-PKcs, based on phosphorylation of BAP1 by the latter and deubiquitination of DNA-PKcs by BAP1."

sparser
"Thus, functional interaction of BAP1 with DNA-PKcs supports a role for BAP1 in NHEJ DNA repair and may provide the basis for new therapeutic strategies and new insights into its role as a tumor suppressor."

reach
"Our results suggest reciprocal positive interactions between BAP1 and DNA-PKcs, based on phosphorylation of BAP1 by the latter and deubiquitination of DNA-PKcs by BAP1."

No evidence text available
BAP1 affects PRC1
| 1 3
| 1 3

sparser
"Given that the composition and expression of PRC1 and BAP1 complexes changes extensively during development ( xref ; xref , xref ; xref ; xref ), in future work, it will be interesting to investigate how the balance between these two opposing activities is regulated at different developmental stages and how cell type-specific H2AK119ub1 levels influence the transcriptional potential of the genome."

sparser
"Recent CRISPR screening studies by Dixit’s group further identified that depletion of PRC1 subunit Ring1B (but not Ring1A) could rescue the cell death induced by loss of BAP1 xref , indicating that there is a robust and direct epigenetic dynamic occurring between the PRC1 and BAP1 complexes that require a balanced state to properly regulate gene expression and determine cell fate."

reach
"lines used in our study identified all these PR-DUB components but failed to identify any PRC1 component as BAP1 interacting proteins, arguing against a physical interaction between BAP1 and PRC1."

sparser
"PRC1 regulates SLC7A11 expression and H2Aub occupancy on the SLC7A11 promoter. (a) ChIP–qPCR showing the binding of BMI-1 and H2Aub on the SLC7A11 promoter, and decreased binding of H2Aub on the SLC7A11 promoter upon BMI-1 knockdown. (b) ChIP– qPCR showing the binding of RNF2 and H2Aub on the SLC7A11 promoter, and decreased binding of H2Aub on the SLC7A11 promoter upon RNF-2 knockdown. (c-f) BMI-1 or RNF2 deficiency increased SLC7A11 mRNA (c, d) and protein levels (e, f) in UMRC6 cells. (g-h) BMI-1 (g) or RNF2 (h) deficiency in 786-O cells increased SLC7A11 mRNA levels. *, p < 0.05; **, p < 0.01; ***, p < 0.001; ****, p < 0.0001. lines used in our study identified all these PR-DUB components but failed to identify any PRC1 component as BAP1-interacting proteins, arguing against a physical interaction between BAP1 and PRC1."
BAP1 affects PARP3
| 4
| 4

sparser
"Several studies and clinical trials [ xref ], have shown that PARP inhibitors influence cancers in which mutations in BRCA1 or BRCA2 are observed, which led us to assume that the cancerous growth-inhibiting interaction of BAP1 with BRCA1 may already be perturbed in this case, and that PARP inhibitors may actually be blocking the novel interaction of BAP1 with PARP3 which enhances cancer growth."

sparser
"Thus, we suspect that the novel interaction of BAP1 and PARP3 may also be perturbed by PARP inhibitors, leading to inhibition of cancer growth."

sparser
"In this scenario, it is possible that the novel interaction of BAP1 with PARP3 in cancerous cells may be promoting cancerous growth, possibly through regulation of DNA repair and apoptosis."

sparser
"Perturbation of the interaction of BAP1 with PARP3 , using PARP3 inhibitors, may then suppress cancerous growth, at least in sarcomatoid MPM."
BAP1 affects Osteosarcoma
| 4
| 4

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"In this study, the roles of BAP1 in potential targets, biological functions, signaling pathways, and immune infiltration were comprehensively explored by mining the osteosarcoma datasets from GEO and TARGET databases using the rapidly developing bioinformatics in recent years.In summary, through bioinformatics and in vitro assays, this study demonstrated that BAP1 was a tumor suppressor in osteosarcoma and provided new clues for osteosarcoma treatment such as BAP1-targeted therapy."

reach
"BAP1 suppressed osteosarcoma cell growth and EMT."

reach
"In vitro, BAP1 could inhibit osteosarcoma cells proliferation and EMT.BAP1 belongs to the ubiquitin C-terminal hydrolase subfamily of deubiquitinases (27)."

reach
"BAP1 might be a tumor suppressor in osteosarcoma and a promising therapeutic target."
BAP1 affects Nevus
| 4
BAP1 inhibits Nevus. 4 / 4
| 4

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"BINs are highly penetrant, present in up to 90% of mutation carriers, and typically present as multiple, skin colored or reddish-brown, dome-shaped melanocytic tumors (also or previously called BAP1-inactivlated melanocytic tumors, Wiesner nevi, BAPomas, nevoid melanoma-like melanocytic proliferations, BAP1 mutant Spitz nevi, and melanocytic BAP1-mutated atypical intradermal tumors)."

reach
"Pathologists can enable identification of patients at risk by recognizing BAP1-inactivated nevi."

reach
"Morphology of this secondary clone strictly depends on the type of second genetic hit: inactivation of the BAP1 (BRCA1-associated protein) gene is the hallmark of BAP1-inactivated nevus (BIN) (33, 34); gain-of-function mutations of CTNNB1 or loss of APC is found in deep penetrating nevus (DPN) (35, 36); loss-of-function of PRKAR1A is typical of pigmented epithelioid melanocytoma (PEM) (37, 38)."

reach
"Affected individuals initially present with multiple skin-colored or light red dome-shaped papules, so-called BAP1-inactivated nevi/melanocytoma of the skin and develop meningiomas by the time they reach 50 years [57]."
BAP1 affects Meningioma
| 4
Mutated BAP1 activates Meningioma. 4 / 4
| 4

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"Germline BAP1 mutations have also been identified, increasing the hereditary risk of meningiomas and other cancers including uveal melanoma, cutaneous melanoma, and renal cell carcinoma [53, 54]."

reach
"Initially thought to be sporadic tumors, germline BAP1 mutations were identified in some of these patients, further supporting meningiomas as part of the BAP1 tumor predisposition syndrome spectrum."

reach
"Germline BAP1 mutations cause mesothelioma and other malignancies, namely, uveal cancer, meningioma, and melanoma, overall defined as “BAP1- related cancer syndrome”."

reach
"Certain patients with meningiomas caused by BAP1 mutations inherit BAP1 mutations and are diagnosed with BAP1 tumor predisposition syndrome (50,51)."
BAP1 affects MP65
| 4
BAP1 binds MP65. 4 / 4
| 4

sparser
"Since we have demonstrated MP65 have higher glycolysis, glycolytic capacity and increased nucleotide biosynthesis metabolites compared to MP65-BAP1, we examined key enzymes involved in glycolysis and the PPP."

sparser
"Both inhibitors diminished MP65 and MP65-BAP1 growth; however, MP65 were more sensitive to these compounds compared to MP65-BAP1 ( xref )."

sparser
"In addition, these inhibitors did not alter MP65 and MP65-BAP1 growth ( xref )."

sparser
"We found that MP65 (OXPHOS high ) were more resistant to both metabolic stress environments than MP65-BAP1, whereas MP46 (OXPHOS low ) were more vulnerable to stressful conditions compared to MP46-BAP1 ( xref and xref )."
BAP1 affects L3MBTL2
2 | 1 1
2 | 1 1

No evidence text available

reach
"By targeting atypical polycomb protein L3MBTL2, BAP1 interacts with and stabilizes L3MBTL2, co-occupying and maintaining H4K20me1 at target gene loci, such as EZH2 locus."

sparser
"By targeting atypical polycomb protein L3MBTL2, BAP1 interacts with and stabilizes L3MBTL2, co-occupying and maintaining H4K20me1 at target gene loci, such as EZH2 locus."

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BAP1 affects JUN
| 4
BAP1 activates JUN. 4 / 4
| 4

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"RNA sequencing analyses revealed that reduced BAP1 expression suppressed ASXL1 Y591X -induced upregulation of the transcription factors activator protein 1 ( AP-1 ) and early growth response 1/2 ( EGR[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways."

reach
"Furthermore, we also demonstrated that hyperactivity of ERK1/2 and JNK/c-Jun signaling induced by BAP1 downregulation was necessary for ICC cell proliferation, cell cycle progression, and invasion in vitro and in vivo using the inhibitors of the ERK1/2 pathway (U0126) and JNK/c-Jun pathway (SP600125)."

reach
"These data clearly suggested that hyperactivity of ERK1/2 and JNK/c-Jun signaling induced by BAP1 downregulation is necessary for ICC cell proliferation, cell cycle progression, and invasion."
BAP1 affects IL1B
| 4
BAP1 increases the amount of IL1B. 4 / 4
| 4

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"Rucavado et al. [14] have demonstrated that the intramuscular injection of the SVMP, BaP1, from B. asper venom produced notable tissue alterations and increases in the protein levels of IL-1β, IL-6, a[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"This study showed that the snake venom metalloprotease (SVMP) BaP1 activates FLSs to produce PGE 2 by a mechanism dependent on COX-2, mPGES-1 and iPLA 2 s. BaP1 also induces IL-1beta release, which up-regulates the production of PGE 2 at a late stage of the stimulation."

reach
"Partially consistent with our results, BAP1 was unfolded to increase the COX-2 and mPGES-1 expression by activating the NF-κB signaling pathway and inducing the release of IL-1β (Viana et al. 2020), but this study failed to explore the regulatory role of BAP1 mutations on the NF-κB signaling pathway."

reach
"Taken together, our findings show that the metalloprotease BaP1 directly activates FLSs to produce PGE by a COX-2-dependent mechanism involving the coupling of the inducible enzymes COX-2 and mPGES-1 and intracellular PLA s. BaP1 also induced release of IL-1β, which up-regulates the production of PGE at a later stage of the stimulation."
BAP1 affects HCFC2
4 |
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BAP1 affects H2AC20
3 | 1
3 | 1

sparser
"Broader applicability of PRC2 inhibition to the treatment of other cancers is supported by the general prevalence of mutations in other chromatin-associated proteins that perturb the balance of activities regulating H3K27me3, including components of the SWI/SNF chromatin remodeling complex and the BAP1 H2A deubiquitinase complex ( xref )."

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BAP1 affects E2F
| 4
| 4

sparser
"Presently, however, direct evidence of BAP1 interacting with ubiquitinated E2F transcription factors is still lacking."

sparser
"In addition, co-immunoprecipitation experiments have shown that BAP1 directly associates with E2F family members, suggesting a possible additional direct control of these transcriptional factors by BAP1 xref ."

sparser
"The results here demonstrated that BAP1 binds to E2F-responsive promoters, indicating a role in epigenetic regulation."

sparser
"14 At present, it is not known whether BAP1 directly interacts with ubiquitinated E2F transcription factors."

reach
"In addition, a mutation in the BRCA1-associated protein-1 (BAP1) gene is implicated in UM metastasis, and is also known to promote the development of cutaneous melanoma, basal cell carcinoma, malignan[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (often in the abdomen), and uveal melanoma [97]."

reach
"BAP1 germline mutations underlie a rare, autosomal dominant tumour predisposition syndrome that is most commonly associated with (in order of decreasing frequency) atypical Spitz tumours, uveal melano[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 syndrome increases the risk of aggressive RCC, cutaneous melanocytic lesions, basal cell carcinoma, mesothelioma (frequently in the abdomen) and uveal melanoma [151,152,153]."
BAP1 affects CTNNB1
| 4
BAP1 activates CTNNB1. 4 / 4
| 4

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"BAP1 depletion caused a reduction in mRNA levels of neural crest migration genes (ROBO1), melanocyte differentiation genes (CTNNB1, EDNRB and SOX10) and other genes that are down-regulated in class 2 tumors (LMCD1 and LTA4H)."

reach
"Studies have reported that BAP1 upregulates β-catenin, which further promotes TNBC tumorigenesis [29]."

reach
"Mechanistically, PVT1 binds with KLF5 and increases its stability via BAP1, which upregulates beta-catenin signaling, resulting in enhanced TNBC tumorigenesis."

reach
"PVT1 binding with KLF5 causes KLF5 to stabilise and this elevates the BAP1 levels, which in turn promotes beta-catenin signalling, thus enhancing TNBC carcinogenesis (Li et al., 2018)."
BAP1 affects COPE
4 |
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BAP1 affects CD274
| 3 1
BAP1 increases the amount of CD274. 4 / 4
| 3 1

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"We found an obvious increase in the total extent of H2AK119Ub in Bap1-KO MC38 cells compared to WT cells (Fig. 1f) and re-expression of WT human BAP1 but not a catalytical inactive BAP1 Cys91-Ala (C91A) variant [18] in Bap1-KO cells rescued both cell-surface PD-L1 accumulation levels and Cd274 mRNA expression (Fig. 1j, k), indicating that Bap1-mediated regulation of PD-L1 expression is dependent on its enzymatic activity."

reach
"BAP1 overexpression significantly increased the expression of SP1, YAP, PD-L1, CAS6, AXL, and downstream signaling proteins (PI3K, STAT3, ERK½, MMP-2, and MMP-9), while BAP1 knockdown reduced their levels."

sparser
"The results of our study showed no significant impact of BAP1 loss and expression of PD-L1 ≥ 1% on patient survival ( p = 0.541) or on time to disease progression ( p = 0.589)."

reach
"Moreover, BAP1 inactivation significantly increased the mRNA expression of the immune checkpoint receptors CD274 (PD-L1), PDCD1 (PD-1), and LAG3 (LAG3) in primary TCGA tumors, which are critical mediators of T-cell exhaustion40 (Fig. 4D, E)."
BAP1 affects CCDC6
2 | 2
2 | 2

sparser
"The interaction between CCDC6 and BAP1 was confirmed in MPM cells, by co-immunoprecipitation."

No evidence text available

sparser
"Coiled-coil domain containing 6 (CCDC6) interacts with BAP1 and has been reported to regulate both homologous recombination and PARP inhibitor sensitivity."

No evidence text available
BAP1 affects CAMP
| 4
| 4

sparser
"However, the estimated charge of the Bap1 protein monomer is −12.9 at pH 7.0 (), providing a possible explanation for the binding of the LL-37 peptide to the negatively charged target protein, the Bap1 protein, by an electrostatic interaction."

sparser
"The mechanism(s) of interaction between the LL-37 and the Bap1 protein is under investigation in our laboratory."

sparser
"Following LL-37 binding by Bap1, the concentration of free LL-37 is reduced to sub-lethal concentrations, leading to the apparent resistance and survival of V. cholerae ."

sparser
"We suggest that due to PmB treatment of the bacterial strain, the OmpT LDV residues become exposed during biogenesis on the surface of the OMVs, allowing for binding of the Bap1 protein and thereby promoting LL-37 binding to Bap1."
BAP1 affects ATP
| 4
BAP1 inhibits ATP. 4 / 4
| 4

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"Moreover, BAP1 fibroblasts enhance aerobic glycolysis and lactate secretion, but reduce mitochondrial respiration and ATP production, when compared with wild-type BAP1."

reach
"BAP1 inhibits glucose deprivation induced reactive oxygen species and ATP depletion, two cellular events contributing to the ER stress induced cell death."

reach
"BAP1 +/- fibroblasts displayed increased aerobic glycolysis and lactate secretion, and reduced mitochondrial respiration and ATP production compared with BAP1 WT."

reach
"Intact BAP1 catalytic activity could increase aerobic glycolysis and lactate secretion and reduce mitochondrial respiration and ATP production [49]."
BAP1 affects ARNT
| 4
| 4

sparser
"Therefore, we checked whether BAP1 could also bind to HIF-1β."

sparser
"CoIP experiments in cells co-transfected with full length Myc-tagged BAP1 (Myc- BAP1 ) and Flag-tagged full-length HIF-1β, or HIF-1β fragments covering residues 2 to 470 [HIF-1β(2-470)], 142-470 [HIF-1β(142-470)], 471-789 [HIF-1β(471-789)], 582-789 [HIF-1β(592-789)] ( xref ), showed that BAP1 binds to the N terminus region of HIF-1β, specifically to the DNA binding and dimerization region [HIF-1β(2-470) and HIF-1β (142-470)] ( xref )."

sparser
"We found that BAP1 also binds to the N terminus region of HIF-1β, specifically to the DNA binding and dimerization region ( xref )."

sparser
"Aligning the crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) ( xref ) to our structural model for the binding complex of BAP1 -HIF-1α reveals the significance of residue 1 to 73 of HIF-1α, as this region binds to both BAP1 and HIF-1β ( xref )."
BAP1 affects 6-dEB
| 4
BAP1 activates 6-dEB. 4 / 4
| 4

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"It has been found that BAP1 could produce 6-dEB in the absence of propionate and methylmalonate."

reach
"BAP1 produced 6-dEB at a titer of 0.32 +/- 0.11 mg l -1 from 20 mM methylmalonate (XREF_FIG)."

reach
"In addition to the plasmid-based production platform mentioned above, the three DEBS (∼30 kb in length) and PCC genes ( accA1 and pccB ) were also introduced into the E. coli BAP1 chromosome, creating[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"To rebuild the biosynthesis of 3-O-α-mycarosylerythronolide B (MEB) in E. coli, the previously reported BAP1 harboring pBP130 and pBP144 was used as the starting strain (WT) to produce 6-dEB."
ASXL1 affects H2AK119Ub
| 4
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

reach
"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

reach
"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

reach
"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

reach
"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
ASXL1 affects EED
| 4
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
ASXL1 affects BAP1, and H2AK119Ub
| 4
BAP1 binds ASXL1 and H2AK119Ub. 4 / 4
| 4

reach
"The images were collected with an exposure of 0.04 seconds per frame; the total exposure time was 2 seconds for the BAP1/ASXL1-H2AK119Ub complex, for a total of 50 frames."

reach
"In the BAP1/ASXL1-H2AK119Ub complex, we centered a spherical mask with a 60 Å pixels radius on the BAP1/ASXL1 moiety, which resulted in a clearly resolved BAP1/ASXL1 on the nucleosome at 5."

reach
"Using the best-scoring particles, we ran Non-Uniform Refinement, which yielded the BAP1/ASXL1-H2AK119Ub complex at 3."

reach
"For the BAP1/ASXL1-H2AK119Ub complex, available X-ray crystal structures were used for initial rigid body fit into our cryo-EM maps."
ASXL1 affects BAP1, and EED
| 4
EED binds BAP1 and ASXL1. 4 / 4
| 4

sparser
"In flies and humans, the Polycomb repressive deubiquitinase (PR-DUB) complex is formed through interactions of BAP1 and ASXL1 [ xref ]."

sparser
"BAP1 interacts with ASXL1 polycomb chromatin binding protein forming a polycomb repressive deubiquitinase complex that regulates histone H2A ubiquitination xref ."

sparser
"8 BAP1 interacts with ASXL1 (MIM 612990 ) to form the Polycomb group Repressive Deubiquitinase complex (PR-DUB)."

sparser
"BAP1 also interacts with ASXL1 to form the Polycomb group repressive deubiquitinase complex (PR-DUB), which is involved in stem cell pluripotency and other developmental processes ( xref , xref )."
ARNT affects BAP1
| 4
| 4

sparser
"Therefore, we checked whether BAP1 could also bind to HIF-1β."

sparser
"CoIP experiments in cells co-transfected with full length Myc-tagged BAP1 (Myc- BAP1 ) and Flag-tagged full-length HIF-1β, or HIF-1β fragments covering residues 2 to 470 [HIF-1β(2-470)], 142-470 [HIF-1β(142-470)], 471-789 [HIF-1β(471-789)], 582-789 [HIF-1β(592-789)] ( xref ), showed that BAP1 binds to the N terminus region of HIF-1β, specifically to the DNA binding and dimerization region [HIF-1β(2-470) and HIF-1β (142-470)] ( xref )."

sparser
"We found that BAP1 also binds to the N terminus region of HIF-1β, specifically to the DNA binding and dimerization region ( xref )."

sparser
"Aligning the crystal structure of HIF-1α-HIF-1β complex (PDB ID: 4zpr) ( xref ) to our structural model for the binding complex of BAP1 -HIF-1α reveals the significance of residue 1 to 73 of HIF-1α, as this region binds to both BAP1 and HIF-1β ( xref )."

reach
"BAP1 binds and deubiquitinates the transcriptional regulator host cell factor 1 (HCF-1), which interacts with histone modifying complexes [8,9]."

reach
"In fact, BAP1 binds and deubiquitinates the transcriptional regulator host cell factor 1 that interacts with histone modifying complexes [19]."

reach
"Although BAP1 interacts with a transcriptional regulator, HCF-1, and transcription factors FoxK1 and FoxK2, how BAP1 controls gene expression remains unclear."
Cre affects BAP1
| 3
Cre activates BAP1. 3 / 3
| 3

reach
"As already discussed, a strong reduction in the expression of the Bap1 exons was demonstrated in the RNA-seq datasets from Bap1 Cγ1-cre relative to control GC B cells ( Supplementary Figures S3C, D ), confirming an effective Cre-mediated Bap1 gene inactivation in the cells captured for the RNA-seq."

reach
"At the same time, the effective Cre-mediated Bap1 to Bap1 allele conversion was demonstrated by PCR-genotyping of the genomic DNA from ex vivo stimulated Bap1 Cγ1-cre and control Bap1 Cγ1-cre B cells ( Supplementary Figures S3A, B )."

reach
"Again, Bap1 Cγ1-Cre B cells demonstrated normal levels of class switching over 4 days in culture ( Supplementary Figures S6A, B ), though Cre-mediated Bap1 to Bap1 allele conversion was observed as early as days 3, albeit without full loss of the Bap1 allele ( Supplementary Figures S6C, D )."
Bisphenol A affects BAP1
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Bisphenol A increases the amount of BAP1. 3 / 3
3 |

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YWHAE affects BAP1
3 |
3 |

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YWHAB affects BAP1
3 |
3 |

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USP21 affects BAP1
1 | 2
1 | 2

reach
"Importantly, PRC1 and H2AK119ub are known to be engaged in transcriptional regulation of ESC differentiation and early embryogenesis ( Voncken et al., 2003; Stock et al., 2007; Liang et al., 2013; Alo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In particular, we report strong co-localization in the genomic binding of BAP1, USP16, and to a lesser extent USP22, conserved across different cell-types."

No evidence text available
UCHL5 affects BAP1
1 | 1 1
1 | 1 1

reach
"This suggests that these residues play functionally important roles that have yet to be defined but may include binding of specific substrates or regions of partner proteins within their respective UCH37 and BAP1 complexes."

sparser
"These differences can rationalize why GK13S does not bind to UCHL5 and BAP1."

No evidence text available
TBCB affects BAP1
| 3
| 3

sparser
"BAP1 is positively involved in the regulation of chromatin and also binds to some transcription factors and cofactors, thereby bridging chromatin-remodeling complexes and chromatin [ xref , xref ]."

sparser
"Here we show that BAP1 assembles multiprotein complexes containing numerous transcription factors and cofactors, including HCF-1 and the transcription factor Yin Yang 1 (YY1)."

sparser
"BAP1 is associated with transcriptionally active chromatin and interacts with several transcription factors and cofactors, thus acting as a chromatin scaffold for chromatin-remodeling complexes ( xref , xref )."
SNCA affects BAP1
| 3
| 3

sparser
"Α-Synuclein also interacts with a protein BAP1 (BRCA1 associated protein 1) possessing ubiquitin C-terminal hydrolase activity (UCH) [ xref ]."

sparser
"The interaction of α-synuclein with BAP1 and other DUBs is a prognostic indicator of unfavorable outcome in several types of cancer, and may also affect neurodegenerative processes [ xref ]."

sparser
"Further studies are necessary to better understand the regulatory role of α-synuclein-BAP1 interaction and its effect on specific gene regulation."
SFN affects BAP1
3 |
3 |

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SF3B1 affects EIF1AX
| 3
| 3

sparser
"The Rotterdam Ocular Melanoma Study Group investigated the association of EIF1AX , SF3B1 and BAP1 mutation with disease-free survival and metastatic risk of patients with uveal melanoma. xref In their recent article, they published a Kaplan-Meier survival probability curve regarding these mutations."

sparser
"Uveal melanoma (UM) is a relative rare disease and has a high mortality rate due to metastasis in about half of all patients within 15 y after diagnosis. xref , xref , xref It is the most common primary intra‐ocular malignancy in adults in the Western world. xref UM specific mutations in the alpha subunit genes GNAQ and GNA11 are described as well as mutations in BAP1 , SF3B1 , and EIF1AX . xref , xref , xref Mutations in the latter 3 genes are found in ∼75% of all UM and are useful for prognostication of patients. xref , xref , xref BAP1 ‐mutated UM gives rise to early‐onset metastasis whereas SF3B1 ‐mutated UM gives rise to late‐onset metastasis and EIF1AX ‐mutated UM hardly metastasizes. xref Mutations in PLCB4 and CYSLTR2 are described in UM in a mutually exclusive manner to GNAQ or GNA11 mutations but so far have not been associated with prognosis. xref , xref Copy number alterations in chromosomes 1, 3, 6, and 8 are correlated with prognosis of the UM patient. xref , xref UM with EIF1AX , SF3B1 and BAP1 mutations are associated with unique chromosomal patterns, suggesting distinct UM subclasses."

sparser
"Other significantly mutated genes in UM are BAP1 , EIF1AX and SF3B1 , which also form a second mutually‐exclusive subgroup xref (Table  xref )."
Porins affects BAP1
| 1 2
| 1 2

reach
"Bap1 binds to the OmpT porin on OMVs by means of the LDV domain of OmpT."

sparser
"We demonstrated that Bap1 binds to the OmpT porin on the OMVs through the LDV domain of OmpT."

sparser
"Bap1 binds to the OmpT porin on OMVs by means of the LDV domain of OmpT."
POLR1G affects BAP1
1 | 1 1
1 | 1 1

sparser
"BAP1 interacts with CAST in UM cells, and CAST and its subsequent calpain pathway may mediate BAP1-related cell migration regulation."

No evidence text available

reach
"BAP1 interacts with CAST in UM cells, and CAST and its subsequent calpain pathway may mediate BAP1 related cell migration regulation."
ODN affects BAP1
| 3
ODN leads to the phosphorylation of BAP1. 3 / 3
| 3

reach
"We investigated whether ODN modulates BAP1 phosphorylation."

reach
"Moreover, ODN-induced mitochondrial ROS generation was involved in DNA damage, BAP1 phosphorylation, upregulation of Sp1 and Bax, sensitization of anti-cancer drug-mediated apoptosis."

reach
"Interestingly, ODN-mediated mitochondrial ROS production induced BAP1 phosphorylation, DNA damage, and Bax upregulation."
NCIT:C94600 affects BAP1
| 3
| 3

sparser
"In conclusion, the present work points out the significance of BAP1 inhibition by BCR-ABL in the CML."

sparser
"We next asked whether the inhibition of BAP1 by BCR-ABL was dependent on the tyrosine kinase activity of the latter."

sparser
"Therefore, our results strongly suggest that BCR-ABL-induced inhibition of BAP1 (at mRNA and protein levels) is a tyrosine kinase-dependent event."
MYB affects BAP1
3 |
MYB decreases the amount of BAP1. 3 / 3
3 |

No evidence text available

No evidence text available

No evidence text available
MAFF affects BAP1
| 1 2
| 1 2

sparser
"This study describes a novel BAP1-MAFF signaling axis which is crucial for CRC growth, potentially serving as a therapeutic target and a promising prognostic biomarker for CRC."

reach
"BAP1 as a nuclear deubiquitinating enzyme (DUB), bound to and deubiquitinated MAFF, thereby stabilizing it."

sparser
"Xie et al. [ xref ], in their analysis, revealed a novel BAP1-MAFF signaling axis that may be fundamental for CRC growth and could be regarded as either a prognostic marker or a potential therapeutic target."
KPNB1 affects BAP1
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3 |

No evidence text available

No evidence text available

No evidence text available
KPNA2 affects BAP1
3 |
3 |

No evidence text available

No evidence text available

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KMT2D affects BAP1
| 3
| 3

reach
"These results revealed a substantial co-localization of BAP1 and MLL4 on promoters and active enhancers.We next examined the recruitment of BAP1 and MLL4, as well as the distribution of H3K4me1 and H3K27ac, on BAP1 MLL4 promoters and active enhancers (Fig. 7c and Suppl."

reach
"To assess whether ASXLs play a role in regulating MLL4 and BAP1 binding at specific chromatin sites, we generated MLL4-, ASXL2-or BAP1-deficient human colon cancer HCT116 cells using the CRISPR/Cas9 approach."

reach
"Interdependent genomic binding of MLL4 and BAP1 on enhancers."
KDM6A affects BAP1
2 | 1
2 | 1

No evidence text available

sparser
"Lastly, depletion of MLL3 but not MLL4 affects the interactions between UTX and BAP1 ( xref )."

No evidence text available
IPO5 affects BAP1
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
IPO4 affects BAP1
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
IGF1R affects BAP1
1 | 1 1
1 | 1 1

reach
"Interaction between IGF1R and BAP1 was confirmed in ARPE-19 cells and IGF1R silencing protected cells from HG-induced injury by regulating BAP1 expression."

sparser
"Interaction between IGF1R and BAP1 was confirmed in ARPE-19 cells and IGF1R silencing protected cells from HG-induced injury by regulating BAP1 expression."

No evidence text available
HTT affects BAP1
| 3
| 3

sparser
"Moreover, we confirmed the interaction of HTT and BAP1 in the KB1P-G3 cells using coimmunoprecipitation of endogenous proteins ( xref )."

sparser
"Concurrently, our study elucidates an interaction between HTT and BAP1, illuminating the opposing effects of HTT and BAP1 in establishing bipolarity of mitotic divisions through regulation of spindle pole number, spindle length, and pole clustering."

sparser
"The entirety of the predicted interaction domain on BAP1 (aa 528–669) is necessary for the HTTBAP1 interaction, as no alternative fragments validated successfully ( xref )."
HSPA9 affects BAP1
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
GZ17-6.02 affects BAP1
| 3
GZ17-6.02 increases the amount of BAP1. 3 / 3
| 3

reach
"As a single agent, GZ17-6.02 significantly enhanced BAP1 expression (Table 6)."

reach
"We found that GZ17-6.02 as a single agent increased BAP1 expression in uveal melanoma cells that was not altered by ERBB receptor inhibitors."

reach
"Hence, our discovery that GZ17-6.02 is enhancing BAP1 expression favors increased expression from the wild type unmutated BAP1 allele in uveal melanoma cells.GZ17-6.02 changed the methylation and acetylation of Histone H3 in uveal melanoma cells, most notably, the mono-methylation of lysine 9 and lysine 4 was reduced and this correlated with a trend for increased di-methylation of lysine 4."
GNAS affects BAP1
2 | 1
2 | 1

No evidence text available

No evidence text available

sparser
"The GNAS protein interacted with the GNAQ/11 and BAP1 proteins (online suppl."
GEP affects BAP1
| 3
BAP1 binds GEP. 3 / 3
| 3

sparser
"Inactivating mutations in the tumor suppressor BAP1 are associated with the class 2 GEP and high metastatic risk xref , whereas single nucleotide substitutions in SF3B1 and EIF1AX are found mainly in class 1 tumors and are associated with intermediate and low metastatic risk, respectively xref , xref ."

sparser
"BAP1 mutations were associated with class 2 GEP ( P < ."

sparser
"Mutations in SF3B1 and EIF1AX are associated with Class 1 GEP and have been reported to be good prognostic factors in UM, whereas BAP1 mutations are associated with Class 2 GEP and poor prognosis ( xref , xref )."
ERK affects BAP1
| 1 2
| 1 2

sparser
"Further examination revealed that BAP1 interacts with phosphorylated ERK1/2, deubiquitinating their ubiquitins, and thus attenuating the ERK signaling pathway."

sparser
"Collectively, these results suggest that the binding of BAP1 to phosphorylated-ERK1/2 may lead to a reduction in its phosphorylation."

reach
"Surprisingly, we found that neither ERK1/2 nor any other signaling molecules upstream of ERK bound to BAP1; only phosphorylated-ERK1/2 co-precipitated with endogenous BAP1 (Fig. 4B)."
EIF1AX affects SF3B1
| 3
| 3

sparser
"The Rotterdam Ocular Melanoma Study Group investigated the association of EIF1AX , SF3B1 and BAP1 mutation with disease-free survival and metastatic risk of patients with uveal melanoma. xref In their recent article, they published a Kaplan-Meier survival probability curve regarding these mutations."

sparser
"Uveal melanoma (UM) is a relative rare disease and has a high mortality rate due to metastasis in about half of all patients within 15 y after diagnosis. xref , xref , xref It is the most common primary intra‐ocular malignancy in adults in the Western world. xref UM specific mutations in the alpha subunit genes GNAQ and GNA11 are described as well as mutations in BAP1 , SF3B1 , and EIF1AX . xref , xref , xref Mutations in the latter 3 genes are found in ∼75% of all UM and are useful for prognostication of patients. xref , xref , xref BAP1 ‐mutated UM gives rise to early‐onset metastasis whereas SF3B1 ‐mutated UM gives rise to late‐onset metastasis and EIF1AX ‐mutated UM hardly metastasizes. xref Mutations in PLCB4 and CYSLTR2 are described in UM in a mutually exclusive manner to GNAQ or GNA11 mutations but so far have not been associated with prognosis. xref , xref Copy number alterations in chromosomes 1, 3, 6, and 8 are correlated with prognosis of the UM patient. xref , xref UM with EIF1AX , SF3B1 and BAP1 mutations are associated with unique chromosomal patterns, suggesting distinct UM subclasses."

sparser
"Other significantly mutated genes in UM are BAP1 , EIF1AX and SF3B1 , which also form a second mutually‐exclusive subgroup xref (Table  xref )."
| 3

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Dbi affects BAP1
| 3
Dbi phosphorylates BAP1. 3 / 3
| 3

sparser
"Mechanistically, ODN induced BAP1 phosphorylation and enhanced Sp1-BAP1 interaction, resulting in Sp1 ubiquitination and degradation."

sparser
"Interestingly, ODN-induced BAP1 phosphorylation and DNA damage were modulated by the production of mitochondrial reactive oxygen species (ROS)."

sparser
"Mito-TEMPO pretreatment inhibited ODN-induced BAP1 phosphorylation, increased Bax expression, and promoted apoptosis induced by the combined treatment of ODN and oxaliplatin ( xref , xref E)."
COPI complex affects BAP1
| 3
BAP1 binds COPI complex. 3 / 3
| 3

reach
"AP-immunoblotting of GFP-COPE shows interaction of COPA as expected, however BAP1 cannot be observed which is probably due to the low levels of BAP1 that bind to the COPI complex (Fig 5E)."
| PMC

reach
"Overall, the loss of mass of COPA on blot could not be tied to the loss of ubiquitin and additional experiments are required to explain this observation.To exclude that the BAP1 interaction with the COPI complex is an artifact of overexpression of the exogenous BAP1 construct, we immunoprecipitated endogenous BAP1 using different antibodies against BAP1 in cell lysate from HeLa FRT parental cells (Fig 5D)."
| PMC

reach
"It was intriguing that BAP1 has such a sequence in its C-terminus, but our mutational analysis suggests that the interaction between BAP1 and the COPI complex is not mediated through the C-terminal KxKxx sequence in BAP1."
| PMC
BRCA affects BAP1
| 3
| 3

reach
"We assessed whether tumour BRCA associated protein 1 (BAP1) status is a predictive biomarker for survival in patients receiving first-line combination platinum/pemetrexed therapy.introduction METHODS: PM cases (n=114) from Aalborg, Denmark were stained for BAP1 on tissue microarrays."

reach
"其他危险因素包括猿猴病毒40感染 、遗传因素及基因突变如BRCA相关蛋白1(BRCA associated protein 1, BAP1)等 。"

reach
"Recent work has shown that BRCA-associated protein 1 (BAP1), which functions as part of the polycomb repressive deubiquitinase complex that removes monoubiquitination on H2AK119, regulates SLC7A11, a subunit of the heterodimeric system x cystine-glutamate antiporter."
BIRC5 affects BAP1
| 1 2
| 1 2

sparser
"In untreated A375 and C918 cells, the loss of BAP1 was associated with a dramatic reduction in survivin levels ( xref , control “C” lanes), consistent with the prior analysis."

sparser
"Furthermore, co-immunoprecipitation experiments were done, but did not demonstrate direct protein-protein interaction between BAP1 and survivin (data not shown)."

reach
"Furthermore, co-immunoprecipitation experiments were done, but did not demonstrate direct protein protein interaction between BAP1 and survivin (data not shown)."
| 3

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects ubiquitins
| 3
BAP1 deubiquitinates ubiquitins. 3 / 3
| 3

reach
"BAP1 expression reduced the ubiquitination of GFP-ERK2, indicating that BAP1 deubiquitinates ubiquitins on GFP-ERK2 (Fig. 4D)."

reach
"Moreover, BAP1 was found to control MSC migration through the ERK signaling pathway, with BAP1 deubiquitinating ubiquitins on phosphorylated-ERK1/2."

reach
"BAP1 deubiquitinates ubiquitins on phosphorylated-ERK1/2 to suppress MSC migration ."

reach
"BAP1 binds and deubiquitinates the transcriptional regulator host cell factor 1 (HCF-1), which interacts with histone modifying complexes [8,9]."

reach
"In fact, BAP1 binds and deubiquitinates the transcriptional regulator host cell factor 1 that interacts with histone modifying complexes [19]."

reach
"Although BAP1 interacts with a transcriptional regulator, HCF-1, and transcription factors FoxK1 and FoxK2, how BAP1 controls gene expression remains unclear."
| 3

reach
"BAP1 inhibits the ER stress gene regulatory network and modulates metabolic stress response."

reach
"BAP1 loss in UM leads to ASXL2 reduction and lipid homeostasis dysregulation, conferring lipotoxicity resistance and an enhanced oxidative stress response—an adaptation that can be exploited with atorvastatin, which induces lipid peroxidation and cell death specifically in BAP1‐mutant liver metastases (Figure 6)."

reach
"BAP1 inhibits the ER stress gene regulatory network and modulates metabolic stress response."

reach
"Considering the changes from asynchronous cells may be the indirect effect due to cell cycle arrest caused by BAP1 loss, we synchronized cells at the G1/S boundary and found that BAP1 depletion also r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The BAP1 gene encodes BRCA1-associated protein 1, which suppresses tumors by promoting the activity of the Hippo tumor suppressor pathway[20]."

reach
"Their studies revealed differential expression of the cancer-related lncRNAs Malat1, Neat1, H19, Meg3, and their associated miRNA-target pairs, regulating various hallmarks of tumorigenesis such as cell cycle and p53 signaling.BRCA1-Associated Protein 1 (BAP1) is known to enhance BRCA1 tumor suppressor potential and its inactivation is associated with various cancers, in particular clear-cell renal cell carcinoma (ccRCC)."

reach
"It seems that the progression of UM is dependent on the histone H2a posttranslational modification induced by BAP1/EZH2 and PCR1."

reach
"Given the deubiquitination enzyme activity of BAP1, we tested the BAP1-mediated post-translational modification of LKB1."

reach
"It is expected that BAP1 promotes tumorigenesis of CCA by modulating DNA repair, gene transcription, and post-translational modification, such as glycosylation."
BAP1 affects mastoparan-M
| 3
| 3

sparser
"Based on this model, EZH2 inhibition was tested in BAP1 inactivated MPMs [ xref ]."

sparser
"The EZH2 inhibitor tazemetostat was administered to patients with BAP1 inactivated MPM (loss of nuclear expression)."

sparser
"In patients with BAP1-inactivated MPM, the disease control rate was 54% (95% CI: 42–67%; 33 of 61 patients) at week 12 after a median follow-up of 35.9 week; no patients had a confirmed complete response and two patients had a confirmed partial response."
BAP1 affects laminin
| 3
BAP1 inhibits laminin. 3 / 3
| 3

reach
"However, BaP1 was able to degrade laminin in PCV, evidencing its ability to reach the BM of this vessel type."

reach
"The ability of BaP1 to reduce the immunostaining of laminin to a greater extent than type IV collagen might be due to the fact that type IV collagen constitutes a more stable covalently linked network, in contrast to laminin [XREF_BIBR - XREF_BIBR]."

reach
"In the ex vivo model, BaP1 was able to degrade type IV collagen and laminin from the BM of microvessels."

reach
"Modulation of Schwann Cell Homeostasis by the BAP1 Deubiquitinase."

reach
"Overall, our results indicate Bap1 is required to restrict the spread of polycomb-associated histone modifications in Schwann cells and to promote myelin homeostasis in peripheral nerve."

reach
"Loss of BAP1 led to impaired myelin homeostasis with tomacula and premature activation of the injury program, suggesting that writers and erasers of polycomb repression play mutually reinforcing roles in gene regulation and myelin homeostasis."
BAP1 affects erastin
| 3
BAP1 activates erastin. 3 / 3
| 3

reach
"While the knockdown of BAP1 alone had no significant anti-tumor effects, BAP1 knockdown attenuated the effects of erastin on tumor regression (Fig. 6A-C)."

reach
"All the above results indicated that knockdown of BAP1 attenuated tumoricidal capacity of erastin in vivo."

reach
"Knockdown of BAP1 reduced tumoricidal capacity of erastin in vivo."
Mutated BAP1 inhibits cytokine production. 3 / 3
| 3

reach
"BAP1 mutations inhibited the NF-kappaB signaling pathway, suppressing the cytokine secretion and antigen presentation by macrophages."

reach
"BAP1 mutations inhibited cytokine secretion and antigen presentation by macrophages."

reach
"BAP1 mutations may inhibit the NF-kappaB signaling pathway, repressing the cytokine secretion and antigen presentation by macrophages, which induces the immunosuppressive microenvironment, enhances the malignant phenotypes of UM cells and ultimately promotes the growth and metastasis of UM."
BAP1 affects YWHAE
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
BAP1 affects YWHAB
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
BAP1 affects USP21
1 | 2
1 | 2

reach
"Importantly, PRC1 and H2AK119ub are known to be engaged in transcriptional regulation of ESC differentiation and early embryogenesis ( Voncken et al., 2003; Stock et al., 2007; Liang et al., 2013; Alo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In particular, we report strong co-localization in the genomic binding of BAP1, USP16, and to a lesser extent USP22, conserved across different cell-types."

No evidence text available
BAP1 affects UCHL5
1 | 1 1
1 | 1 1

reach
"This suggests that these residues play functionally important roles that have yet to be defined but may include binding of specific substrates or regions of partner proteins within their respective UCH37 and BAP1 complexes."

sparser
"These differences can rationalize why GK13S does not bind to UCHL5 and BAP1."

No evidence text available
BAP1 affects TG2-179-1
| 3
BAP1 binds TG2-179-1. 3 / 3
| 3

reach
"We therefore propose that the covalent interaction between BAP1 and TG2-179-1 is reversible."

reach
"These data clearly demonstrated that TG2-179-1 inhibits the DUB activity of BAP1.Analysis of the structural model of the BAP1TG2-179-1 complex using the CavityPlus server showed that Cys91 was predicted to form a covalent bond with an electron-deficient atom in TG2-179-1 ."

reach
"Our in silico docking model predicted that TG2-179-1 binds to BAP1 with its fluorobenzene and thiolane rings deeply embedded into the active site pocket, which sterically allows TG2-179-1 to interact covalently with Cys91 in the active site via its electrophilic nitrile carbon atom."
BAP1 affects TBCB
| 3
| 3

sparser
"BAP1 is positively involved in the regulation of chromatin and also binds to some transcription factors and cofactors, thereby bridging chromatin-remodeling complexes and chromatin [ xref , xref ]."

sparser
"Here we show that BAP1 assembles multiprotein complexes containing numerous transcription factors and cofactors, including HCF-1 and the transcription factor Yin Yang 1 (YY1)."

sparser
"BAP1 is associated with transcriptionally active chromatin and interacts with several transcription factors and cofactors, thus acting as a chromatin scaffold for chromatin-remodeling complexes ( xref , xref )."
BAP1 affects SNCA
| 3
| 3

sparser
"Α-Synuclein also interacts with a protein BAP1 (BRCA1 associated protein 1) possessing ubiquitin C-terminal hydrolase activity (UCH) [ xref ]."

sparser
"The interaction of α-synuclein with BAP1 and other DUBs is a prognostic indicator of unfavorable outcome in several types of cancer, and may also affect neurodegenerative processes [ xref ]."

sparser
"Further studies are necessary to better understand the regulatory role of α-synuclein-BAP1 interaction and its effect on specific gene regulation."
BAP1 affects SNAI1
| 3
BAP1 increases the amount of SNAI1. 3 / 3
| 3

reach
"BAP1 knockout in ccRCC cells also downregulated the expression of transcriptional repressor protein Snail and decreased the activity of Rho family GTPases, promoting the cells to undergo mesenchymal-epithelial transition."

reach
"Loss of BAP1 has been reported to downregulate the expression of Snail, promoting ccRCC cells towards mesenchymal-epithelial transition 25."

reach
"BAP1 knockout in ccRCC cells also downregulated the expression of transcriptional repressor protein Snail and decreased the activity of Rho family GTPases, promoting the cells to undergo mesenchymal-epithelial transition."
BAP1 affects SFN
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
BAP1 affects Recurrence
| 3
| 3

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"Germline or somatic mutations and copy number alterations have been identified in several cancers and inactivation of BAP1 accelerates invasion, recurrence and metastases."

reach
"Germline or somatic mutations and deletions of BAP1 have been identified in various tumor types, and downregulation or inactivation of BAP1 could accelerate tumor onset, invasion, recurrence, and metastases ."

reach
"Furthermore, BAP1 has been demonstrated to increase the risk of disease recurrence in patients with resected localized disease: in a cohort of 1479 patients, those with BAP1 loss of function (10.3%) had worse relapse-free survival, even after statistical adjustment for the UISS risk score [49]."
BAP1 affects RCC
| 3
BAP1 inhibits RCC. 3 / 3
| 3

reach
"XREF_BIBR Immunohistochemistry for BAP1 was performed on tissue microarray sections from 559 non metastatic RCC cases treated with nephrectomy : BAP1 was negative in 82 of 559 tumors (14.7%), and Cox regression indicated a significantly worse disease-free and overall survival for patients negative for BAP1 protein compared to patients with BAP1 positive tumors."

reach
"Taken together these data show that BAP1 binding to HCF-1 is likely to be important for BAP1 suppression of RCC development."

reach
"Thus, BAP1 mutation in the germline may predispose to RCC, in which case, RCC development may also be initiated by loss of BAP1."
BAP1 affects Proteasome
| 3
| 3

reach
"Following depletion of BAP1 for 72 h, treatment with MG132 for 6 h led to a substantial rescue of BRCA1 expression in MSTO-211H and NCI-H2452 cells indicating that BAP1 inhibits the proteasome-mediated degradation of BRCA1 (Fig. 1B)."

reach
"Depletion of BAP1 protein induced proteasome-mediated degradation of BRCA1 in mesothelioma cells while loss of BAP1 correlated with BRCA1 loss in mesothelioma patient tumour samples."

reach
"Furthermore, BAP1 protein appears to inhibit FOXO3a degradation via the proteasome-dependent pathway, thereby preserving the activity and stability of the FOXO3a protein."
BAP1 affects Porins
| 1 2
| 1 2

reach
"Bap1 binds to the OmpT porin on OMVs by means of the LDV domain of OmpT."

sparser
"We demonstrated that Bap1 binds to the OmpT porin on the OMVs through the LDV domain of OmpT."

sparser
"Bap1 binds to the OmpT porin on OMVs by means of the LDV domain of OmpT."
BAP1 affects PTGER4
| 3
BAP1 increases the amount of PTGER4. 3 / 3
| 3

reach
"In addition, BaP1 up-regulates EP4 receptor expression."

reach
"BaP1 also up-regulates EP4 receptor expression."

reach
"These findings are supported by the increased EP4 protein expression following BaP1 stimulation in our experimental model, confirming the ability of this receptor to activate FLSs."
BAP1 affects POLR1G
1 | 1 1
1 | 1 1

sparser
"BAP1 interacts with CAST in UM cells, and CAST and its subsequent calpain pathway may mediate BAP1-related cell migration regulation."

No evidence text available

reach
"BAP1 interacts with CAST in UM cells, and CAST and its subsequent calpain pathway may mediate BAP1 related cell migration regulation."
BAP1 affects PAFAH1B1
| 3
| 3

reach
"BAP1 was considered as a tumour suppressor in multiple solid tumours with its deletion or inactivating mutations, and Dey et al. showed that conditional loss of BAP1 in haematopoietic cells led to an [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Dey et al. also reported that conditional loss of Bap1 in haematopoietic cells led to an MDS/chronic myelomonocytic leukaemia (CMML)-like disease in mice [ 35 ], indicating that the physiological cata[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Experimentally, it has been shown that Bap1 deletion in adult mice causes MDS like diseases 46."
| 3
| 3

reach
"On the other hand, BAP1 restoration bas been found to retard cell cycle progression and promote apoptosis of neuroblastoma cells by interacting with 14-3-3 protein [ 34 ]."

reach
"Deficiency of BAP1 inhibits neuroblastoma tumorigenesis through destabilization of MYCN."

reach
"Depletion of BAP1 inhibits neuroblastoma cell growth."
BAP1 affects NPC
| 3
BAP1 activates NPC. 3 / 3
| 3

reach
"In this study, we found that hypoxia-induced BAP1 enhances erastin-induced ferroptosis in NPC by stabilizing H2A.Studies have shown that many tumor cells under a hypoxic microenvironment are less sensitive to chemotherapeutic agents because these agents typically require oxygen for maximum activity [12]."

reach
"Hypoxia-induced BAP1 enhances erastin-induced ferroptosis in NPC by stabilizing H2A."

reach
"Ferroptosis inducers targeting BAP1 may be an effective way to reduce drug resistance in NPC, especially in the hypoxic microenvironment.In conclusion, our study initially validated that hypoxia-induced BAP1 enhanced erastin-induced ferroptosis in NPC by stabilizing H2A (Fig. 7)."
BAP1 affects NFkappaB
| 3
Mutated BAP1 inhibits NFkappaB. 3 / 3
| 3

reach
"BAP1 mutations may inhibit the NF-kappaB signaling pathway, repressing the cytokine secretion and antigen presentation by macrophages, which induces the immunosuppressive microenvironment, enhances the malignant phenotypes of UM cells and ultimately promotes the growth and metastasis of UM."

reach
"BAP1 mutations inhibited the NF-kappaB signaling pathway, suppressing the cytokine secretion and antigen presentation by macrophages."

reach
"BAP1 mutations inhibit the NF-kappaB signaling pathway to induce an immunosuppressive microenvironment in uveal melanoma."
BAP1 affects MAFF
| 1 2
| 1 2

sparser
"This study describes a novel BAP1-MAFF signaling axis which is crucial for CRC growth, potentially serving as a therapeutic target and a promising prognostic biomarker for CRC."

reach
"BAP1 as a nuclear deubiquitinating enzyme (DUB), bound to and deubiquitinated MAFF, thereby stabilizing it."

sparser
"Xie et al. [ xref ], in their analysis, revealed a novel BAP1-MAFF signaling axis that may be fundamental for CRC growth and could be regarded as either a prognostic marker or a potential therapeutic target."
| 3

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"To further determine how loss of BAP1 accelerates B cell lymphoma cells growth in vivo via regulating immune cell infiltration and TME, we conducted single-cell RNA-Seq (scRNA-seq) with the tumor samples from both BAP1-WT and -KO tissues (Supplemental Figure 5A)."

reach
"However, the loss of BAP1 leads to a substantial acceleration of B cell lymphoma cell growth in immunocompetent mice, indicating that BAP1 may function as a tumor suppressor via regulating TME and tumor immune response in this context.Decrease or loss of MHC-II expression in B cell lymphoma has been associated with genetic alterations, such as CIITA deficiency, and is associated with poor survival in human B cell lymphoma."

reach
"In our current studies, we demonstrated that PRC1 is responsible for the transcriptional repression of MHC-II genes upon BAP1 depletion, as inhibition of PRC1 could largely rescue the gene-expression defect in BAP1-depleted B cell lymphoma cells."
BAP1 affects KPNB1
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BAP1 affects KPNA2
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BAP1 affects KMT2D
| 3
| 3

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"These results revealed a substantial co-localization of BAP1 and MLL4 on promoters and active enhancers.We next examined the recruitment of BAP1 and MLL4, as well as the distribution of H3K4me1 and H3K27ac, on BAP1 MLL4 promoters and active enhancers (Fig. 7c and Suppl."

reach
"To assess whether ASXLs play a role in regulating MLL4 and BAP1 binding at specific chromatin sites, we generated MLL4-, ASXL2-or BAP1-deficient human colon cancer HCT116 cells using the CRISPR/Cas9 approach."

reach
"Interdependent genomic binding of MLL4 and BAP1 on enhancers."
BAP1 affects KDM6A
2 | 1
2 | 1

No evidence text available

sparser
"Lastly, depletion of MLL3 but not MLL4 affects the interactions between UTX and BAP1 ( xref )."

No evidence text available
BAP1 affects KDM5C, PBRM1, and SETD2
| 3
| 3

sparser
"We and others have reported the associations of PBRM1 , SETD2 , BAP1 , and KDM5C mutations with advanced stage, grade, and tumor invasiveness [ xref , xref ], and discovered that SETD2 and BAP1 mutations are associated with lower cancer-specific survival rates [ xref , xref ]."

sparser
"Mutations in PBRM1, SETD2, BAP1 , and KDM5C are associated with tumors of advanced stage (odds ratio range 2.34 to 6.40, 2.5, 2.62 to 4.57, 7.6, respectively)."

sparser
"Our study indicates that mutations of PBRM1 , SETD2 , BAP1 and/or KDM5C in kidney cancers are associated with advanced stage, grade and tumor invasiveness."
BAP1 affects JNK
| 3
BAP1 inhibits JNK. 3 / 3
| 3

reach
"BAP1 restricted the growth cycle of intrahepatic cholangiocarcinoma by regulating ERK1/2 and JNK/c-Jun axis (Chen et al. 2018)."

reach
"All of these results further confirmed that BAP1 suppressed ICC cell proliferation, cell cycle progression, and invasion via inhibiting ERK1/2 and JNK/c-Jun signaling pathways."

reach
"Taken together, our data suggested that BAP1 negatively regulated the ERK1/2 and JNK/c-Jun signaling pathways to exert its tumor-suppressive functions in ICC."
BAP1 affects IPO5
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BAP1 affects IPO4
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BAP1 affects IGF1R
1 | 1 1
1 | 1 1

reach
"Interaction between IGF1R and BAP1 was confirmed in ARPE-19 cells and IGF1R silencing protected cells from HG-induced injury by regulating BAP1 expression."

sparser
"Interaction between IGF1R and BAP1 was confirmed in ARPE-19 cells and IGF1R silencing protected cells from HG-induced injury by regulating BAP1 expression."

No evidence text available
BAP1 affects IFNB1
| 2 1
BAP1 increases the amount of IFNB1. 3 / 3
| 2 1

sparser
"In UMRC6 cells, in which BAP1-WT and BAP1-C91G were expressed to a similar extent ( xref ), BAP1-WT induced IFNB1 expression, whereas BAP1-C91G did not ( xref )."

reach
"BAP1 enhanced the expression of IFN-β and stimulator of interferon genes (STING), both of which activate ISGF3."

reach
"Our previous data suggest BAP1 enhances IFNB1 expression to activate ISGF3 in ccRCC cells [1]."
BAP1 affects Hemorrhage
| 3
| 3

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"BaP1 induces hemorrhage, as well as other local pathological effects, upon intramuscular injection in mice ( Gutiérrez et al., 1995; Rucavado et al., 1995 )."

reach
"Thirty micrograms of BaP1 and 5 μg of jararhagin induced prominent hemorrhage in the injected muscle; 2 μg BaP1 did not induce hemorrhage and was selected in order to compare the activity of BaP1 and [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Three of the produced clones were able to completely neutralize BaP1-induced hemorrhage."
BAP1 affects HTT
| 3
| 3

sparser
"Moreover, we confirmed the interaction of HTT and BAP1 in the KB1P-G3 cells using coimmunoprecipitation of endogenous proteins ( xref )."

sparser
"Concurrently, our study elucidates an interaction between HTT and BAP1, illuminating the opposing effects of HTT and BAP1 in establishing bipolarity of mitotic divisions through regulation of spindle pole number, spindle length, and pole clustering."

sparser
"The entirety of the predicted interaction domain on BAP1 (aa 528–669) is necessary for the HTTBAP1 interaction, as no alternative fragments validated successfully ( xref )."
BAP1 affects HSPA9
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BAP1 affects HDAC2
| 1
BAP1 inhibits HDAC2. 1 / 3
| 1

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"In addition, we found that BAP1 depletion did not reduce the stability of residual HDAC2 under conditions where translation was blocked."
BAP1 affects HDAC
| 1 2
BAP1 inhibits HDAC. 3 / 3
| 1 2

reach
"Interestingly, HDAC4 is the only HDAC that is significantly upregulated by the loss of BAP1 in human uveal melanoma cells."

reach
"Since SAHA inhibits numerous HDACs, we questioned whether there may be a specific HDAC that is up-regulated by Bap1 loss and mediates the Bap1-deficient phenotype."

eidos
"Loss of BAP1 increases histone deacetylase levels , and a pre-clinical drug screen identified that pan-histone deacetylase inhibitors ( HDACi ) , such as quisinostat , are effective at inhibiting UM growth ( Kuznetsoff et al ., 2021 ) ."
BAP1 affects H3K27Me3
| 3
| 3

reach
"Additionally, reconstitution of BAP1 into the BAP1-KO cells downregulated H3K27me3 and EZH2 (Fig. 2c), suggesting that BAP1 loss might upregulate H3K27me3 and EZH2 and activate their downstream signaling.A recent study indicated that BAP1 is localized in the endoplasmic reticulum (ER) and bound to the type 3 inositol 1, 4, and 5-triphosphate receptor (IP3R3) [22]."

reach
"Loss of BAP1 increased the intensity of both H2AK119Ub and H3K27me3, and repressed the majority of DEG (66.2% downregulated vs. 33.8% upregulated, Fig. 6d)."

reach
"LaFave et al. demonstrated that a loss of BAP1 increases the trimethylated histone H3 lysine 27 (H3K27me3) and enhancer of zeste 2 polycomb repressive complex 2 subunits (EZH2) levels [21]."
BAP1 affects GNAS
2 | 1
2 | 1

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No evidence text available

sparser
"The GNAS protein interacted with the GNAQ/11 and BAP1 proteins (online suppl."
BAP1 affects GEP
| 3
BAP1 binds GEP. 3 / 3
| 3

sparser
"Inactivating mutations in the tumor suppressor BAP1 are associated with the class 2 GEP and high metastatic risk xref , whereas single nucleotide substitutions in SF3B1 and EIF1AX are found mainly in class 1 tumors and are associated with intermediate and low metastatic risk, respectively xref , xref ."

sparser
"BAP1 mutations were associated with class 2 GEP ( P < ."

sparser
"Mutations in SF3B1 and EIF1AX are associated with Class 1 GEP and have been reported to be good prognostic factors in UM, whereas BAP1 mutations are associated with Class 2 GEP and poor prognosis ( xref , xref )."
BAP1 affects FGFBP1
| 3
BAP1 increases the amount of FGFBP1. 3 / 3
| 3

reach
"WT BAP1, but not mutant BAP1 C91S, obviously elevated the KLF5 and FGF-BP protein levels and inhibited the p27 protein levels in both cell lines (XREF_FIG)."

reach
"As expected, knockdown of BAP1, HCF-1, OGT or KLF5 markedly increased p27 and decreased FGF-BP protein levels in HCC1806 cells (XREF_FIG)."

reach
"In addition, expression of KLF5, CyclinD1, and FGF-BP1 was increased by BAP1 overexpression and decreased by BAP1 knockdown."
BAP1 affects EIF1AX, and SF3B1
| 3
| 3

sparser
"The Rotterdam Ocular Melanoma Study Group investigated the association of EIF1AX , SF3B1 and BAP1 mutation with disease-free survival and metastatic risk of patients with uveal melanoma. xref In their recent article, they published a Kaplan-Meier survival probability curve regarding these mutations."

sparser
"Uveal melanoma (UM) is a relative rare disease and has a high mortality rate due to metastasis in about half of all patients within 15 y after diagnosis. xref , xref , xref It is the most common primary intra‐ocular malignancy in adults in the Western world. xref UM specific mutations in the alpha subunit genes GNAQ and GNA11 are described as well as mutations in BAP1 , SF3B1 , and EIF1AX . xref , xref , xref Mutations in the latter 3 genes are found in ∼75% of all UM and are useful for prognostication of patients. xref , xref , xref BAP1 ‐mutated UM gives rise to early‐onset metastasis whereas SF3B1 ‐mutated UM gives rise to late‐onset metastasis and EIF1AX ‐mutated UM hardly metastasizes. xref Mutations in PLCB4 and CYSLTR2 are described in UM in a mutually exclusive manner to GNAQ or GNA11 mutations but so far have not been associated with prognosis. xref , xref Copy number alterations in chromosomes 1, 3, 6, and 8 are correlated with prognosis of the UM patient. xref , xref UM with EIF1AX , SF3B1 and BAP1 mutations are associated with unique chromosomal patterns, suggesting distinct UM subclasses."

sparser
"Other significantly mutated genes in UM are BAP1 , EIF1AX and SF3B1 , which also form a second mutually‐exclusive subgroup xref (Table  xref )."
| 3

reach
"In addition to other factors, the deubiquitinating enzyme BAP1 is a tumor suppressor in cholangiocarcinoma."

reach
"BAP1 knockout in mice leads to MM growth, larger tumors and increased number of metastases and BAP1 protein expression is also correlated with poor prognosis and shorter survival in RCC and cholangiocarcinoma, as well as UM (22–25)."

reach
"BAP1 restricted the growth cycle of intrahepatic cholangiocarcinoma by regulating ERK1/2 and JNK/c-Jun axis (Chen et al. 2018)."
BAP1 affects COPI complex
| 3
BAP1 binds COPI complex. 3 / 3
| 3

reach
"AP-immunoblotting of GFP-COPE shows interaction of COPA as expected, however BAP1 cannot be observed which is probably due to the low levels of BAP1 that bind to the COPI complex (Fig 5E)."
| PMC

reach
"Overall, the loss of mass of COPA on blot could not be tied to the loss of ubiquitin and additional experiments are required to explain this observation.To exclude that the BAP1 interaction with the COPI complex is an artifact of overexpression of the exogenous BAP1 construct, we immunoprecipitated endogenous BAP1 using different antibodies against BAP1 in cell lysate from HeLa FRT parental cells (Fig 5D)."
| PMC

reach
"It was intriguing that BAP1 has such a sequence in its C-terminus, but our mutational analysis suggests that the interaction between BAP1 and the COPI complex is not mediated through the C-terminal KxKxx sequence in BAP1."
| PMC
BAP1 affects CDKN2A
1 | 1
1 | 1

reach
"χ 2 test or Fisher's exact test (in case of low event rates [n<5]) were used to investigate the association between BRCA1 and BAP1, p16ink4A, and PD-L1 expression and response outcome."

No evidence text available
BAP1 affects BRCA
| 3
| 3

reach
"We assessed whether tumour BRCA associated protein 1 (BAP1) status is a predictive biomarker for survival in patients receiving first-line combination platinum/pemetrexed therapy.introduction METHODS: PM cases (n=114) from Aalborg, Denmark were stained for BAP1 on tissue microarrays."

reach
"其他危险因素包括猿猴病毒40感染 、遗传因素及基因突变如BRCA相关蛋白1(BRCA associated protein 1, BAP1)等 。"

reach
"Recent work has shown that BRCA-associated protein 1 (BAP1), which functions as part of the polycomb repressive deubiquitinase complex that removes monoubiquitination on H2AK119, regulates SLC7A11, a subunit of the heterodimeric system x cystine-glutamate antiporter."
BAP1 affects BIRC5
| 1 2
| 1 2

sparser
"In untreated A375 and C918 cells, the loss of BAP1 was associated with a dramatic reduction in survivin levels ( xref , control “C” lanes), consistent with the prior analysis."

sparser
"Furthermore, co-immunoprecipitation experiments were done, but did not demonstrate direct protein-protein interaction between BAP1 and survivin (data not shown)."

reach
"Furthermore, co-immunoprecipitation experiments were done, but did not demonstrate direct protein protein interaction between BAP1 and survivin (data not shown)."
BAP1 affects BCL2L11
| 3
BAP1 inhibits BCL2L11. 3 / 3
| 3

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"The tumour was histopathologically atypical because of the presence of confluent pleomorphism, solid sheets of cells and grouped mitotic figures : these features were consistent with a melanocytic neoplasm with intermediate morphology (' BAP1 inactivated melanocytoma '; BIM) between a BAP1 inactivated melanocytic naevus and a BAP1 inactivated melanoma."

reach
"Here we report the clinical features, histopathologic findings, and chromosomal copy number data of 19 BAP1-inactivated melanocytomas (BIMs) and compare their features to those of five BAP1-inactivated melanomas."

reach
"None of the stained tumors in our study exhibited the concurrent BAP1 loss and BRAF V600E positivity that are characteristic of adult BIM, suggesting that this is a low-frequency mutation among banal tumors in the pediatric population."
BAP1 affects BARD1, and E3_Ub_ligase
| 3

sparser
"According to this hypothesis, Nishikawa et al showed that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of the BRCA1/BARD1 complex, and that BAP1 and BRCA1/BARD1 mutually regulate ubiquitination in the cell cycle and in the DNA damage response pathway. xref According to the present state of knowledge, it is reasonable to speculate that BAP1 may exert a regulatory role in the DNA damage response."

sparser
"Here, we report that BAP1 interacts with BARD1 to inhibit the E3 ligase activity of BRCA1/BARD1."

sparser
"They found that BAP1 did not appear to be involved in the deubiquitination of the BRCA1/BARD1 complex,29 and BAP1-mediated growth suppression in vitro was shown to be independent of wild-type BRCA1.4 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
BAP1 affects ATF3
| 3
BAP1 decreases the amount of ATF3. 3 / 3
| 3

reach
"Hence, it is not surprising that BAP1 exhibited a prosurvival role and attenuated the metabolic stress-induced cell death by repressing the transcription of the activating transcription factor 3 (ATF3) and C/EBP homologous protein (CHOP) in a DUB activity-dependent manner [39]."

reach
"Considering that BAP1 inhibits transcription of ATF3 and CHOP genes by binding to their corresponding promoters , BAP1 downregulation in normal trophoblast differentiation may release ATF3 promotors from inhibition, thus contributing to trophoblast invasion."

reach
"BAP1 repressed glucose deprivation-induced ER stress cell death by inhibiting transcription of two major UPR-TFs, activating transcription factor 3 (ATF3) and C/EBP homologous protein (CHOP) (70)."
BAP1 affects ASXL proteins
| 3
ASXL proteins binds BAP1. 3 / 3
| 3

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"The interaction between BAP1 and the ASXL proteins is mediated by the ~ 120-residue DEUBAD (DEUBiquitinase ADaptor) domain of the ASXL proteins [XREF_BIBR], and this region is sufficient for activation of BAP1 in in vitro H2AK119ub1 nucleosome deubiquitination assays [XREF_BIBR]."

reach
"The ASXL proteins directly interact with BAP1, KDM1A, NHRs and NCOA1 to target specific genomic loci using histone modifications."

reach
"ASXL proteins interact with BAP1 to form the polycomb repressive deubiquitinase (PR-DUB) complex, which removes monoubiquitin from histone 2A at lysine 119 (H2AK119) [XREF_BIBR, XREF_BIBR]."
BAP1 affects ANKRD17
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BAP1 affects ANKHD1
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BAP1 affects ADGRB1
| 1 2
| 1 2

reach
"Interaction between BAI1 and BAP1 was mediated by a QTEV motif in the carboxy-terminal region of BAI1 and PDZ domains of BAP1."

sparser
"Interaction between BAI1 and BAP1 was mediated by a QTEV motif in the carboxy-terminal region of BAI1 and PDZ domains of BAP1."

sparser
"Using a two-hybrid system, we isolated a cDNA that encodes a protein, named BAP1 (BAI1-associated protein), which interacts with the cytoplasmic region of BAI1."
ASXL2 affects ASXL1
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BAP1 binds ASXL1 and ASXL2. 3 / 3
| 3

sparser
"Although both ASXL1 and ASXL2 have been identified to bind to BAP1, recent biochemical characterization of BAP1 complexes by xref suggest that BAP1 forms two mutually exclusive complexes with ASXL1 and ASXL2."

sparser
"Proteomic analysis of BAP1 interacting partners using this system revealed that BAP1 interacts with ASXL1 and ASXL2 in vivo as well as host cell factor 1 (HCF-1), O-GlcNAc transferase (OGT), and Forkhead box protein K1/2 (FOXK1/2) ( xref )."

sparser
"Depletion of the PHD fingers did not affect the stability of ASXL1 or ASXL2, or the interaction between ASXL1, ASXL2, and BAP1 (Additional file xref : Fig. S2A, B)."
ASXL proteins affects BAP1
| 3
ASXL proteins binds BAP1. 3 / 3
| 3

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"The interaction between BAP1 and the ASXL proteins is mediated by the ~ 120-residue DEUBAD (DEUBiquitinase ADaptor) domain of the ASXL proteins [XREF_BIBR], and this region is sufficient for activation of BAP1 in in vitro H2AK119ub1 nucleosome deubiquitination assays [XREF_BIBR]."

reach
"The ASXL proteins directly interact with BAP1, KDM1A, NHRs and NCOA1 to target specific genomic loci using histone modifications."

reach
"ASXL proteins interact with BAP1 to form the polycomb repressive deubiquitinase (PR-DUB) complex, which removes monoubiquitin from histone 2A at lysine 119 (H2AK119) [XREF_BIBR, XREF_BIBR]."
ANKRD17 affects BAP1
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ANKHD1 affects BAP1
3 |
3 |

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ADGRB1 affects BAP1
| 1 2
| 1 2

reach
"Interaction between BAI1 and BAP1 was mediated by a QTEV motif in the carboxy-terminal region of BAI1 and PDZ domains of BAP1."

sparser
"Interaction between BAI1 and BAP1 was mediated by a QTEV motif in the carboxy-terminal region of BAI1 and PDZ domains of BAP1."

sparser
"Using a two-hybrid system, we isolated a cDNA that encodes a protein, named BAP1 (BAI1-associated protein), which interacts with the cytoplasmic region of BAI1."
| 2

sparser
"Consistently, our findings showed that an ATM inhibitor (KU-55933) and a DNA-PKcs inhibitor (NU7441) markedly inhibited ODN- and cathepsin K siRNA-induced BAP1 phosphorylation and Sp1 and Bax upregulation ( xref E)."

sparser
"Interestingly, the Mito-TEMPO and NAC markedly inhibited ODN- and cathepsin K siRNA-induced BAP1 phosphorylation and Sp1 and Bax upregulation ( xref D)."
MiR-31-5p affects BAP1
| 2
MiR-31-5p decreases the amount of BAP1. 2 / 2
| 2

reach
"These results suggest that miR-31-5p inhibits LPS-induced H9C2 cell injury by downregulating BAP1 expression."

reach
"Exogenous overexpression of miR-31-5p could inhibit the expression of BAP1 in LPS treated H9C2 cells."
2 |
Hsa-miR-200c-3p decreases the amount of BAP1. 2 / 2
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No evidence text available

No evidence text available
Fluoranthene affects BAP1
| 2
| 2

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"BAP-1 induced by fluoranthene."

reach
"BAP-1 induced by fluoranthene comparative proteomic analysis was performed on proteins extracted from fluoranthene exposed cells on 1 d, 3 d, 6 d and 8 d compared with control cells using isobaric tags for relative and absolute quantization (iTRAQ) labeling and LC-MS/MS analysis to access differentially expressed proteins."
Ferroptosis affects BAP1
| 2
| 2

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"These results suggested that hypoxia enhanced erastin-induced ferroptosis in NPC cells by upregulating BAP1."

reach
"BMSC-derived Exos overexpressing GATA-4 inhibits H/R-induced ferroptosis by upregulating miR-330-3p target-regulating BAP1."
Circ_0087851 affects BAP1
| 2
Circ_0087851 increases the amount of BAP1. 2 / 2
| 2

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"The findings disclosed that circ_0087851 upregulation elevated the expression of BAP1 by sponging miR-593-3p, thereby facilitating ferroptosis and hindering CRC progression."

reach
"As shown in Fig. 5A, circ_0087851 overexpression notably elevated the BAP1 protein levels, whereas miR-593-3p introduction or silencing of BAP1 reversed this effect."
Ab affects YUH1
| 2
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
VHL affects SETD2
| 2
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
ULD affects ab
| 2
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
TUBG affects BAP1
| 2
| 2

reach
"A study reported that BAP1 interacts directly with and localizes to γ-tubulin during mitosis and that BAP1 stabilizes γ-tubulin via deubiquitination to support microtubule nucleation and mitotic spindle assembly, thereby ensuring chromosome segregation and preventing chromosome abnormalities."

reach
"Interaction between BAP1 and γ-tubulin occurs during metaphase, anaphase and telophase, but not during prophase, during which γ-tubulin is ubiquitylated."
TP53 affects SETD2
| 2
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
TCEA1 affects BAP1
1 | 1
1 | 1

No evidence text available

sparser
"In contrast, some studies have shown that BAP1 can directly bind TFs to exert its metabolic roles independent of its DUB activity ( xref )."
SRC affects BAP1
| 2
| 2

sparser
"To further validate these findings, we designed primers for the human SRC promoter that target the area of binding of BAP1 to mouse Src promoter in ChIP-Seq."

sparser
"The obtained data indicate that BAP1 binds to the SRC promoter when BAP1 is expressed in cells ( xref ), demonstrating a direct regulation of SRC mRNA levels by BAP1."
SNRPA1 affects BAP1
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SETD2 affects VHL
| 2
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
SETD2 affects TP53
| 2
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
Reactive Oxygen Species increases the amount of BAP1. 2 / 2
| 2

reach
"Modulating BAP1 expression affects ROS homeostasis, cell motility and mitochondrial function."

reach
"inhibiting expression of BAP1, the ROS content was significantly decreased (Fig. 5A and B; CON vs. LPS, p = 0.0003; NC(-) vs. BAP1(-), p = 0.0024)."
RUVBL1 affects BAP1
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RPS27A affects BAP1
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RPS23 affects BAP1
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RPS12 affects BAP1
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RBBP4 affects BAP1
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RAD21 affects BAP1
| 2
| 2

sparser
"The somatically mutated RAD21 and BAP1 are associated with PD-L1 expression in ccRCC tumor cells and might serve as a potential predictor of the response to immunotherapy with PD-1/PD-L1 inhibitors in ccRCC patients."

sparser
"Nevertheless, only mutated RAD21 and BAP1 were associated with PD-L1 expression in tumor cells."
PSMD3 affects BAP1
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No evidence text available
PSMC5 affects BAP1
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PBRM1 affects VHL
| 2
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
OGT affects HCFC1
| 2
OGT binds HCFC1 and BAP1. 2 / 2
| 2

reach
"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
ODN- affects BAP1
| 2
ODN- leads to the phosphorylation of BAP1. 2 / 2
| 2

reach
"Consistently, our findings showed that an ATM inhibitor (KU-55933) and a DNA-PKcs inhibitor (NU7441) markedly inhibited ODN- and cathepsin K siRNA-induced BAP1 phosphorylation and Sp1 and Bax upregulation (Fig. 9E)."

reach
"Interestingly, the Mito-TEMPO and NAC markedly inhibited ODN- and cathepsin K siRNA-induced BAP1 phosphorylation and Sp1 and Bax upregulation (Fig. 9D)."
OCM affects BAP1
| 2
| 2

sparser
"Germline BAP1 mutations are associated with a more aggressive OM phenotype and a recurrent phenotypic complex of c utaneous/ o cular m elanoma, atypical m elanocytic proliferations and o ther internal n eoplasms (ie."

sparser
"These findings suggest that BAP1 alterations may be associated with a more aggressive OM phenotype."
NFE2L1 affects BAP1
| 2
NFE2L1 increases the amount of BAP1. 2 / 2
| 2

reach
"However, whether BAP1 can positively regulate the transcription of RHOJ via removing the monoubiquitination of histone H2A remains to be further delineated.Regarding the regulation of BAP1 at the transcriptional level in HCC, we identified two transcription factors, CTCF and NRF1, which positively regulated BAP1 transcription by direct binding to the promoter region of BAP1."

reach
"Furthermore, we identified two transcription factors, CTCF and NRF1, which activated BAP1 transcription by binding to BAP1 promoter region."
Mito-TEMPO affects BAP1
| 2
Mito-TEMPO leads to the dephosphorylation of BAP1. 2 / 2
| 2

reach
"Interestingly, the Mito-TEMPO and NAC markedly inhibited ODN- and cathepsin K siRNA-induced BAP1 phosphorylation and Sp1 and Bax upregulation (Fig. 9D)."

reach
"Mito-TEMPO pretreatment inhibited ODN-induced BAP1 phosphorylation, increased Bax expression, and promoted apoptosis induced by the combined treatment of ODN and oxaliplatin (Fig. 9, Fig. 10E).5 Conclusion."
MAP4 affects BAP1
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2 |

No evidence text available

No evidence text available
LINC00598 affects BAP1
2 |

No evidence text available

No evidence text available
KDM5C affects VHL
| 2
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
IDH1 affects BAP1
1 | 1
1 | 1

No evidence text available

sparser
"Fig. S7), and IDH1 interacted with the BAP1 protein (online suppl."
HUWE1 affects BAP1
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2 |

No evidence text available

No evidence text available
HSPA1A affects BAP1
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2 |

No evidence text available

No evidence text available
HCFC1 affects OGT
| 2
OGT binds HCFC1 and BAP1. 2 / 2
| 2

reach
"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
ELF2 affects BAP1
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2 |

No evidence text available

No evidence text available
EEF2 affects BAP1
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2 |

No evidence text available

No evidence text available
CTCF affects BAP1
| 2
CTCF increases the amount of BAP1. 2 / 2
| 2

reach
"However, whether BAP1 can positively regulate the transcription of RHOJ via removing the monoubiquitination of histone H2A remains to be further delineated.Regarding the regulation of BAP1 at the transcriptional level in HCC, we identified two transcription factors, CTCF and NRF1, which positively regulated BAP1 transcription by direct binding to the promoter region of BAP1."

reach
"Furthermore, we identified two transcription factors, CTCF and NRF1, which activated BAP1 transcription by binding to BAP1 promoter region."
CHD2 affects BAP1
| 2
CHD2 decreases the amount of BAP1. 2 / 2
| 2

reach
"In the current study, we demonstrated that disruption of G4 structures in the BAP1 promoter by mutagenesis reduced expression of the reporter gene, while silencing CHD2 and CHD7, helicases that likely[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These data suggest that CHD2 and CHD7 likely inhibits BAP1 expression through resolving G-quadruplex structures in its promoter.As a deubiquitinating enzyme, the regulatory role of BAP1 in multiple on[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
CBX5 affects BAP1
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No evidence text available

No evidence text available
BPS affects BAP1
| 2
BPS activates BAP1. 2 / 2
| 2

reach
"These results confirmed that the BPS construct efficiently induces genome edits in Bap1 and Pbrm1, but not in Setd2, in a dox-dependent manner.Upon confirming the genome editing we injected these ESCs into mouse blastocysts for tetraploid complementation and obtained nine founder BPS (Bap1-Pbrm1-Setd2) mice."

reach
"Collectively, these results demonstrate that the introduction of the BPS and ggt-tTA transgenes in the TRACK model induces a low frequency of somatic mutations in Bap1 and Pbrm1 in the kidneys and testes of these mice."
BON1 affects BAP1
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2 |

No evidence text available

No evidence text available
BAP1 affects γ-globin
| 2
BAP1 inhibits γ-globin. 2 / 2
| 2

reach
"To delve more deeply into the underlying mechanism by which BAP1 mediates repression of γ-globin synthesis, it would be of great significance to determine the identity of additional BAP1 target proteins in erythroid cells in future studies."

reach
"We also showed that a novel DRED repressor subunit, deubiquitinase BAP1, regulates site-specific NCoR1 recruitment within the β-globin locus and that BAP1 reduction leads to enormously increased γ-globin mRNA and protein induction (Fig. 5D)."
BAP1 affects vorinostat
| 2
| 2

reach
"The BAP1 deficient developmental phenotype could be rescued using SAHA or specific depletion of HDAC4."

reach
"BAP1 downregulation increases the sensitivity to HDACi in mesothelioma cell lines, and therefore, it would be important to see whether BAP1 aberrations modulate response to vorinostat in VANTAGE 014 study."
BAP1 affects phosphorylated-ERK1/2
| 2
BAP1 deubiquitinates phosphorylated-ERK1/2. 2 / 2
| 2

reach
"Moreover, BAP1 was found to control MSC migration through the ERK signaling pathway, with BAP1 deubiquitinating ubiquitins on phosphorylated-ERK1/2."

reach
"BAP1 deubiquitinates ubiquitins on phosphorylated-ERK1/2 to suppress MSC migration ."
BAP1 affects pathogenesis
| 2
| 2

reach
"In mesothelioma pathogenesis, BAP1 dysfunction contributes to altered cellular growth, enhanced survival, and resistance to apoptosis."

reach
"However, it has not yet been elucidated whether BAP1 modulates the pathogenesis and progression of human cancers through some common cellular and molecular mechanisms, and a pan-cancer analysis for the roles of BAP1 has not yet been conducted."

reach
"P-I metalloproteases, as BaP1 from B. asper venom and C-SVMP from B. pirajai venom, have also been described to act on the complement system: BaP1 decreased the hemolytic activity of NHS and promoted [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In fact, neutrophil chemotaxis induced by the metalloprotease BaP1 in vivo was shown to be dependent on C5a, since the migration of leukocytes was significantly decreased in C5-deficient animals ( Far[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

eidos
"Knockdown of BAP1 in PC3 or DU145 cells induced mesenchymal-to-epithelial transition ( MET ) ."

reach
"Loss of BAP1 promoted mesenchymal-epithelial transition biological function in renal caner."
BAP1 affects lactate
| 2
BAP1 activates lactate. 2 / 2
| 2

reach
"BAP1 perturbation leads to the accumulation of LDHA in cytoplasm to promote the production of lactate and NAD+."

reach
"BAP1 inactivation promotes lactate production by leveraging the subcellular localization of LDHA in melanoma."
BAP1 affects glutathione
| 2
BAP1 increases the amount of glutathione. 2 / 2
| 2

reach
"BAP1 interacts with Nrf2 to modulate SLC7A11 expression and GSH levels [116,117,118]."
| PMC

reach
"The expression or activity of System X is affected by epigenetics, transcription, and posttranscriptional and posttranslational regulators, such as a tumor protein p53 (p53), the nuclear factor, erythroid 2-like 2 (NFE2L2), BRCA1-associated protein 1 (BAP1), or mucin 1 cell surface-associated (MUC1), leading to complex feedback mechanisms to control GSH levels in ferroptosis (Hayano et al., 2016; Chen et al., 2017; Song et al., 2018; Zhang Y. et al., 2018)."
BAP1 affects glucose
| 1 1
BAP1 inhibits glucose. 2 / 2
| 1 1

reach
"BAP1 inhibits glucose deprivation induced reactive oxygen species and ATP depletion, two cellular events contributing to the ER stress induced cell death."

sparser
"In RCC cells, BAP1 inhibited glucose deprivation-induced apoptosis through interaction with the activating transcription factor 3 ( ATF3 ) and DNA damage-inducible transcript 3 protein ( DDIT3 ) gene promoters in a polycomb complex protein BMI-1-dependent manner xref ."
| 1 1
| 1 1

eidos
"O-GlcNacylated HCF-1 recruits OGT and BAP1 to activate peroxisome proliferator activator receptor gamma coactivator 1alpha ( PGC1alpha ) , stimulating gluconeogenesis ( 49 ) ."

reach
"Consequently, BAP1 stabilizes PGC-1α and promotes gluconeogenesis [7] ."
BAP1 affects cell
| 2
BAP1 inhibits cell. 2 / 2
| 2

eidos
"Similarly , downregulation of BAP1 markedly increased cells in the S phase and reduced cells in the G0 / G1 phase , consistent with the results of a recent study that depletion of BAP1 resulted in a modest accumulation of host cell factor 1 , which promoted the transition from G1 to S phase41 ."

eidos
"These results indicated that BAP1 suppressed ICC cell proliferation , cell cycle progression , and invasion via inhibiting ERK1 / 2 and JNK / c-Jun signaling pathways ."

reach
"BAP1 mutations may inhibit the NF-kappaB signaling pathway, repressing the cytokine secretion and antigen presentation by macrophages, which induces the immunosuppressive microenvironment, enhances the malignant phenotypes of UM cells and ultimately promotes the growth and metastasis of UM."

reach
"BAP1 mutations inhibited the NF-kappaB signaling pathway, suppressing the cytokine secretion and antigen presentation by macrophages."
BAP1 affects ab
| 2
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
BAP1 affects TCEA1
1 | 1
1 | 1

No evidence text available

sparser
"In contrast, some studies have shown that BAP1 can directly bind TFs to exert its metabolic roles independent of its DUB activity ( xref )."
BAP1 affects SRC
| 2
| 2

sparser
"To further validate these findings, we designed primers for the human SRC promoter that target the area of binding of BAP1 to mouse Src promoter in ChIP-Seq."

sparser
"The obtained data indicate that BAP1 binds to the SRC promoter when BAP1 is expressed in cells ( xref ), demonstrating a direct regulation of SRC mRNA levels by BAP1."
BAP1 affects SNRPA1
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects SETD2, and TP53
| 2
BAP1 binds SETD2 and TP53. 2 / 2
| 2

sparser
"Mutations in BAP1 , SETD2, and TP53 have been associated with inferior clinical results in those patients who undergo primary treatment with surgical resection [ xref ]."

sparser
"Among the prevalent non- VHL alterations, BAP1, SETD2 , and TP53 mutations are associated with a poor prognosis [ xref , xref , xref ]."
BAP1 affects RUVBL1
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2 |

No evidence text available

No evidence text available
BAP1 affects RPS27A
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects RPS23
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects RPS12
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects RBBP4
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects RAD21
| 2
| 2

sparser
"The somatically mutated RAD21 and BAP1 are associated with PD-L1 expression in ccRCC tumor cells and might serve as a potential predictor of the response to immunotherapy with PD-1/PD-L1 inhibitors in ccRCC patients."

sparser
"Nevertheless, only mutated RAD21 and BAP1 were associated with PD-L1 expression in tumor cells."
BAP1 affects Pancreatitis
| 2
| 2

reach
"Although LOH of additional genes within Chr 3p21.1 may contribute to tumor suppression, and loss of BAP1 in pancreatic cancer may be secondary—rather than the cause of genomic instability—our mouse model suggests Bap1 ablation suffices to cause defective DNA repair, pancreatitis, and cooperates with oncogenic Kras to promote pancreatic cancer."

reach
"Deletion or heterozygous loss of Bap1 in murine pancreata causes genomic instability, tissue damage, and pancreatitis with full penetrance."
BAP1 affects PSMD3
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2 |

No evidence text available

No evidence text available
BAP1 affects PSMC5
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects PDCD1
| 2
BAP1 increases the amount of PDCD1. 2 / 2
| 2

reach
"Moreover, BAP1 inactivation significantly increased the mRNA expression of the immune checkpoint receptors CD274 (PD-L1), PDCD1 (PD-1), and LAG3 (LAG3) in primary TCGA tumors, which are critical mediators of T-cell exhaustion40 (Fig. 4D, E)."

reach
"Moreover, BAP1 inactivation significantly increased the mRNA expression of the immune checkpoint receptors CD274 (PD-L1), PDCD1 (PD-1), and LAG3 (LAG3) in primary TCGA tumors, which are critical mediators of T-cell exhaustion40 (Fig. 4D, E)."
BAP1 affects PBRM1, and SETD2
| 2
PBRM1 binds BAP1 and SETD2. 2 / 2
| 2

sparser
"In fact this association with chromosome 3p loss has recently been strengthened by several publications demonstrating that 4 out of five ( VHL , PBRM1 , SETD2 and BAP1 ) of the most frequently mutated genes associated with CCRCC are on chromosome 3p xref – xref and that the mutation of either SETD2 , BAP1 or PBRM1 associates with poorer prognosis or progression in those patients xref , xref , xref ."

sparser
"The mutations in the chromatin regulators PBRM1 , BAP1 , and SETD2 were differentially associated with altered expression patterns of large numbers of genes when compared to samples bearing a background of VHL mutation ( xref , xref )."
BAP1 affects OCM
| 2
| 2

sparser
"Germline BAP1 mutations are associated with a more aggressive OM phenotype and a recurrent phenotypic complex of c utaneous/ o cular m elanoma, atypical m elanocytic proliferations and o ther internal n eoplasms (ie."

sparser
"These findings suggest that BAP1 alterations may be associated with a more aggressive OM phenotype."

eidos
"In addition , loss of BAP1 has also been described to deplete mitochondrial proteins in pancreas cells137 ."

eidos
"In the pancreas , knockdown of Bap1 diminished mitochondrial protein levels ( e.g ."
BAP1 affects MAP4
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects Leukemia
| 2
BAP1 activates Leukemia. 2 / 2
| 2

reach
"Moreover, the mutant ASXL1 (ASXL1-MT)/BAP1 complex promotes MLL rearrangement leukemia progression by reducing the upregulation of HOXA5, HOXA7, HOXA9, and IRF8 induced by H2AK119ub1 [136]."

reach
"A mutant ASXL1-BAP1-EHMT complex contributes to heterochromatin dysfunction in clonal hematopoiesis and chronic monomyelocytic leukemia."
BAP1 affects LINC00598
2 |

No evidence text available

No evidence text available
BAP1 affects KDM5C, PBRM1, SETD2, and VHL
| 2
| 2

sparser
"The objective of this hypotheses-generating Radiogenomics study was to investigate associations between CT features of ccRCC and mutations of the genes VHL, PBRM1, SETD2, KDM5C , and BAP1."

sparser
"The identification of genetic mutations is clinically relevant because both an advanced stage and poor survival have been associated with VHL, PBRM1, BAP1, SETD2, and KDM5C mutations [ xref , xref – xref ]."
BAP1 affects IDH1
1 | 1
1 | 1

No evidence text available

sparser
"Fig. S7), and IDH1 interacted with the BAP1 protein (online suppl."
BAP1 affects Histone_H2B
| 2
BAP1 deubiquitinates Histone_H2B. 2 / 2
| 2

reach
"Loss of Asx in flies led to an increase in Ub-H2A levels without influencing other chromatin marks (H3K4 me3, H3K27me3), and assays using purified proteins found Asx stimulates Calypso activity towards Ub-AMC, and that Asx and Calypso and the human orthologs BAP1 and ASXL1 deubiquitinate H2A but not H2B in reconstituted nucleosomes [XREF_BIBR]."

reach
"Subsequent WB using anti-HA and anti- Flag antibodies showed that the BAP1 WT, but not a MT, deubiquitinated H2B in vivo."
BAP1 affects HUWE1
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects HSPA1A
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2 |

No evidence text available

No evidence text available
BAP1 affects HOXA genes
| 2
BAP1 activates HOXA genes. 2 / 2
| 2

reach
"We also examined whether ASXL1-MT and BAP1 upregulated posterior HOXA genes and IRF8 in human RUNX1-ETO cells by qRT-PCR."

reach
"ASXL1-MT and BAP1 induces upregulation of HOXA genes and IRF8."
BAP1 affects HCFC1, and OGT
| 2
OGT binds HCFC1 and BAP1. 2 / 2
| 2

reach
"The fact that HCF-1 acts as an important modulator of cellular circadian rhythm [XREF_BIBR] raises the possibility that the OGT, HCF-1, and BAP1 complex interacts with the BMAL1, CLOCK, and PER complex to modulate circadian rhythm."

reach
"Within the BAP1/HCF-1/OGT complex, HCF1 is modified by O-GlcNacylation, the addition of single O-linked N-acetylglucosamine moiety to specific serine or threonine residues (50,51)."
BAP1 affects H3K27me3
| 2
Modified BAP1 decreases the amount of H3K27me3. 2 / 2
| 2

reach
"Recurrent BAP1 mutations have also been observed in intrahepatic cholangiocarcinoma 25, and a recent study showed that loss of Bap1 in mice increases expression of H3K27me3 and Ezh2 while suppressing expression of PRC2 targets 26."

reach
"Coexpression of ASXL1-MT and BAP1 also reduced bulk H3K27me3 levels, which is probably a direct consequence of loss of H2AK119Ub 24 in 32Dcl3 cells."
BAP1 affects H2AK119
| 2
BAP1 deubiquitinates H2AK119. 2 / 2
| 2

reach
"28 ASXL1 may also collaborate with BAP1, loss of which led to a myelodysplastic syndrome like syndrome in a mouse model, to deubiquitinate H2AK119 at polycomb group targets."

reach
"BAP1 is activated by ASXL1 and deubiquitylates mono-ubiquitylation of H2AK119 (Daou et al. 2015; Sahtoe et al. 2016)."
BAP1 affects ELF2
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects EEF2
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects DNMT1
| 2
BAP1 decreases the amount of DNMT1. 2 / 2
| 2

reach
"Indeed, treatment of BAP1-deficient cells with proteasome inhibitor restored DNMT1 protein level ( Figure 5 D)."

reach
"Furthermore, the expression of full-length BAP1, but not a mutant that lacks deubiquitinase activity, reduced the ubiquitination level of DNMT1 ( Figure 5 F)."

reach
"This scenario is possible because H2Aub levels at a DSB are inversely correlated with BAP1 recruitment , and BAP1 depletion increases H2Aub accumulation at a DSB (Fig. 1)."

reach
"siRNA-mediated knockdown of BAP1 causes DSB repair defects as evidenced by decreased IR-induced BRCA1, RAD51, and RPA foci [71,107,109]."
BAP1 affects DEUBAD
| 2
BAP1 leads to the ubiquitination of DEUBAD. 2 / 2
| 2

reach
"BAP1 promotes DEUBAD monoubiquitination resulting in an increased stability of ASXL2, which in turn stimulates BAP1 DUB activity."

reach
"However, monoubiquitination of DEUBAD was induced to a similar extent by WT BAP1 or a catalytically inactive (C91S) mutant (Fig. 2d)."
BAP1 affects DEGs
| 2
BAP1 inhibits DEGs. 2 / 2
| 2

reach
"RNA-Seq and subsequent GSEA and IPA revealed that reduced BAP1 expression rescued the majority of DEGs affected by ASXL1 Y591X expression, including the transcription factors AP-1 and EGR1/2 , as well[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"BAP1 knockdown restored the expression of 75% of the upregulated DEGs and 67% of downregulated DEGs altered by ASXL1 Y591X expression ( Fig. 6 A-B) to levels comparable to those in the control group."

eidos
"BAP1 participates in homologous recombination ( HR ) repair and prevents chromosomal instability through modulation of various substrates , including MCRS1 and INO80 ."

reach
"BAP1 participates in homologous recombination (HR) repair and prevents chromosomal instability through modulation of various substrates, including MCRS1 and INO80."
BAP1 affects CCR5
| 2
Mutated BAP1 increases the amount of CCR5. 2 / 2
| 2

reach
"Additionally, the mechanism by which BAP1 mutation leads to increased expression of CCR5 and its ligands remains to be investigated."

reach
"We demonstrated that BAP1 mutation led to increased expression of CCR5 on Tregs and tumor cells."
BAP1 affects CBX5
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects BON1
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects BAX
| 2
BAP1 decreases the amount of BAX. 2 / 2
| 2

reach
"The knockdown or catalytic mutant form of BAP1 abolished the ODN-induced upregulation of Sp1 and Bax expression."

reach
"OTUB1, OTUD7A, and BAP1 downregulated Sp1 and Bax expression (Fig. 5)."
BAP1 affects ATM
| 1 1
| 1 1

reach
"8 BAP1 also interacts with ataxia telangiectasia mutated and ataxia telangiectasia and Rad3 related proteins, which regulate the DNA damage induced by UV radiation."

sparser
"Further large germline studies of biliary tract cancers are warranted to better assess the association of ATM and BAP1 with this rare cancer type."
BAP1 affects ASXH domain
| 2
BAP1 binds ASXL1 and ASXH domain. 2 / 2
| 2

reach
"ASXL1 directly binds to BAP1 through the ASXH domain and its adjoining region."

reach
"As mentioned earlier, the interaction between BAP1 and ASXL1 occurs through the ASXH domain."
BAP1 affects APEX1
| 2
BAP1 inhibits APEX1. 2 / 2
| 2

reach
"BAP1 loss reduced HAP1 cell adhesion/spreading (Fig. 1D) and proliferation (Fig. 1E), considered here to be readouts of cell fitness."

reach
"BAP1-TPDS-associated BAP1 variants reduce HAP1 fitness."
BAP1 affects ANAPC7
2 |
2 |

No evidence text available

No evidence text available
BAP1 affects ADAR2
| 2
ADAR2 binds BAP1. 2 / 2
| 2

sparser
"Meanwhile, since ADAR2 is associated with wt BAP1, this may help in defining the landscape where mutated BAP1 and low ADAR2 are associated with increased type‐1 IFN signaling [ xref ]."

sparser
"In addition, the association of ADAR2 with wt BAP1 is possibly going to help explore some pending questions about BAP1 and type‐1 IFN signaling."
ASXL2 affects ab
| 2
YUH1 binds ULD, BAP1, ab, and ASXL2. 2 / 2
| 2

sparser
"Instead, the BAP1 ULD domain binds to the AB box, which leads to the formation of a stable ternary complex consisting of the BAP1-UCH, BAP1-ULD and ASXL2-AB domains ( xref – xref )."

sparser
"The peak fraction of the AB box in the complex shifted dramatically from #17 to #15 compared to AB alone, supporting our hypothesis that BAP1-UCH, BAP1-ULD and ASXL2-AB form a ternary stable complex in vitro and that ULD recruits ASXL2-AB to access the BAP1-UCH catalytic motif."
ASXL1 affects ASXH domain
| 2
BAP1 binds ASXL1 and ASXH domain. 2 / 2
| 2

reach
"ASXL1 directly binds to BAP1 through the ASXH domain and its adjoining region."

reach
"As mentioned earlier, the interaction between BAP1 and ASXL1 occurs through the ASXH domain."
ANAPC7 affects BAP1
2 |
2 |

No evidence text available

No evidence text available
ADAR2 affects BAP1
| 2
ADAR2 binds BAP1. 2 / 2
| 2

sparser
"Meanwhile, since ADAR2 is associated with wt BAP1, this may help in defining the landscape where mutated BAP1 and low ADAR2 are associated with increased type‐1 IFN signaling [ xref ]."

sparser
"In addition, the association of ADAR2 with wt BAP1 is possibly going to help explore some pending questions about BAP1 and type‐1 IFN signaling."
ABL1 affects BAP1
| 1 1
ABL1 phosphorylates BAP1 on tyrosine. 2 / 2
| 1 1

reach
"Bap-1 is a substrate for the Bcr serine threonine kinase and is also phosphorylated on tyrosine by Bcr-Abl but not by c-Abl."

rlimsp
"Bap-1 is a substrate for the Bcr serine-threonine kinase and is also phosphorylated on tyrosine by Bcr-Abl but not by c-Abl."