IndraLab

Statements


WDR48 affects USP12
14 | 142 98
WDR48 binds USP12.
14 | 107 65
14 | 81 39

reach
"(G) Superposition of the free and UAF1-bound USP12 structures focusing on the Palm domain, the catalytic cleft, BL2, and BL3."

sparser
"In the UAF1-USP12 complex structure, this aromatic residue is dislodged from the pocket but tucked underneath the Ring Finger between the side chains of Arg245 and Lys247 ( Figure 3D )."

reach
"We then show that in solution USP12 can bind to a second molecule UAF1, but with lower affinity."

reach
"Reciprocal IP-MS/MS suggests the existence of multiple distinct Dub complexes in which USP12 (or USP46) interacts with WDR48, WDR20 (or DMWD), and PHLPP (or PHLPPL) (XREF_FIG, XREF_SUPPLEMENTARY)."

sparser
"Overall Architecture of the UAF1-USP12 Complex (A and B) Ribbon diagram of the UAF1-USP12 complex in two orthogonal views."

sparser
"Overall, our two UAF1-USP12 structures not only reveal a series of structural changes of the enzyme that are connected to the binding site of UAF1 but also highlight the dynamic nature of three key loops, namely, BL2, BL3, and the CC Loop, surrounding the catalytic site of the activated enzyme."

reach
"Interestingly, the addition of WDR20 to the USP12 and UAF1 complex further increased USP12 's DUB activity."

sparser
"The crystal structures of USP1-UAF1 reveal plasticity in USP1 and key differences to USP12-UAF1 and USP46-UAF1, two related proteases."

reach
"F219 and its two interacting residues in the UAF1-bound USP12 structure are highlighted in sticks."

reach
"To confirm that these structural differences of USP12 are associated with UAF1 binding but not crystal packing or resolution limit, we crystallized the UAF1 and USP12 complex in another space group (C[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
USP12 binds WDR48 and F222. 3 / 3
| 3

reach
"Although the active site of USP12 in the UAF1-USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme ( Figure S3F ), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme, in the C2 space group structure, the USP12 Thumb [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme (XREF_SUPPLEMENTARY), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."
USP12 binds WDR48 and F219. 3 / 3
| 3

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta-strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in its pocket formed between the Fingers and Palm domains, instead of being tucked underneath the Fingers domain (XREF_FIG, XREF_FIG, XREF_FIG)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in the pocke[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
WDR48 activates USP12.
| 33 33
WDR48 activates USP12. 10 / 66
| 32 33

sparser
"Wild type UAF1 was unable to activate USP12 E190K to similar levels as compared to USP12 WT ( xref d, e) and similarly, the UAF1 3X mutant did no longer activate."

reach
"Activation of USP12 by UAF1 requires structural fine tuning at the junction of the Ub globular domain and it tail through BL1, whereas WDR20 acts independently of this site and directly affects the ca[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Activation of USP12 by UAF1 requires structural fine-tuning at the junction of the Ub globular domain and it tail through BL1, whereas WDR20 acts independently of this site and directly affects the ca[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For example, USP1, USP12, and USP46 are activated by the WD40-repeat containing UAF1, and USP7 is activated by GMPS (Cohn et al., 2007, 2009; Faesen et al., 2011; van der Knaap et al., 2005)."

sparser
"Mechanism of USP12 Activation by UAF1 and WDR20."

sparser
"Consistent with our results on USP1, activation of USP12-WDR20 by UAF1 is achieved mainly through an ~20 folds increase of k cat with little changes in K M , suggesting that substrate affinity of USP12 is not altered by UAF1 ( xref , xref )."

reach
"Together, these unique features of the Ub-free USP12 structures hint at possible involvement of the USP12 Fingers domain in enzyme regulation.To delineate the mechanism by which UAF1 activates USP12, [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"First, our results rule out the possibility that UAF1 and WDR20 activate USP12 by enhancing its substrate affinity."

sparser
"Consistent with our results on USP1, activation of USP12-WDR20 by UAF1 is achieved mainly through an 20-fold increase of k cat with little change in K M , suggesting that substrate affinity of USP12 is not altered by UAF1 ( Figures 6A and S5A-S5D) ."

sparser
"Based on our results, we propose a tenable model for USP12 activation by UAF1 and WDR20 (Figure 7) ."
WDR48 activates USP12-E190K. 1 / 1
| 1

reach
"Wild type UAF1 was unable to activate USP12 E190K to similar levels as compared to USP12 WT (XREF_FIG d, e) and similarly, the UAF1 3X mutant did no longer activate."
WDR48 decreases the amount of USP12.
| 1
WDR48 decreases the amount of USP12. 1 / 3
| 1

reach
"As Uaf-1 and WDR20 depletion reduces Usp12 levels their silencing has a comparable effect on the AR as the Usp12 depletion alone."
WDR48 deubiquitinates USP12.
| 1
WDR48 deubiquitinates USP12. 1 / 1
| 1

reach
"The E1 protein from anogenital HPV types interacts with the UAF1 associated deubiquitinating enzymes USP1, USP12, and USP46 to stimulate replication of the viral genome."
USP12 affects WDR48
14 | 109 65
USP12 binds WDR48.
14 | 107 65
14 | 81 39

reach
"(G) Superposition of the free and UAF1-bound USP12 structures focusing on the Palm domain, the catalytic cleft, BL2, and BL3."

sparser
"In the UAF1-USP12 complex structure, this aromatic residue is dislodged from the pocket but tucked underneath the Ring Finger between the side chains of Arg245 and Lys247 ( Figure 3D )."

reach
"We then show that in solution USP12 can bind to a second molecule UAF1, but with lower affinity."

reach
"Reciprocal IP-MS/MS suggests the existence of multiple distinct Dub complexes in which USP12 (or USP46) interacts with WDR48, WDR20 (or DMWD), and PHLPP (or PHLPPL) (XREF_FIG, XREF_SUPPLEMENTARY)."

sparser
"Overall Architecture of the UAF1-USP12 Complex (A and B) Ribbon diagram of the UAF1-USP12 complex in two orthogonal views."

sparser
"Overall, our two UAF1-USP12 structures not only reveal a series of structural changes of the enzyme that are connected to the binding site of UAF1 but also highlight the dynamic nature of three key loops, namely, BL2, BL3, and the CC Loop, surrounding the catalytic site of the activated enzyme."

reach
"Interestingly, the addition of WDR20 to the USP12 and UAF1 complex further increased USP12 's DUB activity."

sparser
"The crystal structures of USP1-UAF1 reveal plasticity in USP1 and key differences to USP12-UAF1 and USP46-UAF1, two related proteases."

reach
"F219 and its two interacting residues in the UAF1-bound USP12 structure are highlighted in sticks."

reach
"To confirm that these structural differences of USP12 are associated with UAF1 binding but not crystal packing or resolution limit, we crystallized the UAF1 and USP12 complex in another space group (C[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
USP12 binds WDR48 and F222. 3 / 3
| 3

reach
"Although the active site of USP12 in the UAF1-USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme ( Figure S3F ), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme, in the C2 space group structure, the USP12 Thumb [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme (XREF_SUPPLEMENTARY), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."
USP12 binds WDR48 and F219. 3 / 3
| 3

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta-strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in its pocket formed between the Fingers and Palm domains, instead of being tucked underneath the Fingers domain (XREF_FIG, XREF_FIG, XREF_FIG)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in the pocke[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 inhibits WDR48.
| 1
USP12 bound to WDR20 inhibits WDR48. 1 / 1
| 1

reach
"The partial overlap of the binding interfaces and the high affinity of WDR20 suggests that WDR20 binding to USP12 and UAF1 can either prevent binding of the second UAF1 molecule or it can actively compete out UAF1 if it is bound to the USP12 and UAF1 complex."
USP12 deubiquitinates WDR48.
| 1
USP12 deubiquitinates WDR48. 1 / 1
| 1

reach
"Usp12 and Usp46 deubiquitinate both H2A and H2B and Uaf-1 is required for this reaction [XREF_BIBR]."
WDR20 affects USP12
8 | 9 58 44
WDR20 binds USP12.
8 | 33 22
8 | 13 9

reach
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far away from its catalytic center."

sparser
"Consistent with our results on USP1, activation of USP12-WDR20 by UAF1 is achieved mainly through an 20-fold increase of k cat with little change in K M , suggesting that substrate affinity of USP12 is not altered by UAF1 ( Figures 6A and S5A-S5D) ."

No evidence text available

No evidence text available

No evidence text available

sparser
"In the two UAF1-USP12 complex structures, UAF1 binding appears to induce BL1 destabilization via a chain of conformational changes, whereas the WDR20-USP12 interaction seems to overwrite the allosteric effect of UAF1 by re-stabilizing the loop."

reach
"Results confirmed the interaction between endogenous Usp12, Uaf-1 and WDR20 in PC cells."

sparser
"Adjacent to the major USP12-WDR20 interface, the tip of USP12 BL3, which is otherwise disordered in both free and UAF1-bound forms of the enzyme, adopts a stable structure by making van der Waal contacts with WDR20."

reach
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

sparser
"The k cat and K M values are presented as means ± SD (n = 3), and the K D values were obtained through steady-state analysis (see Supplemental Experimental Procedures for technical discussion). (B) Time course of USP12 modification by the suicidal Ub-VME substrate in the absence or presence of UAF1 and WDR20. (C) Time course of LRGG-AMC (top) in contrast to Ub-AMC (bottom) hydrolysis by USP12 (green), UAF1-USP12 (orange), and USP12-WDR20 (red)."
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
WDR20 activates USP12.
| 9 24 22
WDR20 activates USP12. 10 / 58
| 9 24 22

eidos
"To investigate the mechanism by which WDR20 activates USP12 , we next determined the crystal structure of a ternary complex consisting of USP12 , UAF1 lacking its SLD , and WDR20 at a 3 ."

reach
"In agreement with our enzyme kinetics data, our binding studies revealed negligible changes in the substrate binding affinity of USP12 induced by UAF1 or WDR20 (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"In agreement with our enzyme kinetics data, our binding studies revealed negligible changes in the substrate binding affinity of USP12 induced by UAF1 or WDR20 (Figures 6A and S6) ."

reach
"Overall, our two UAF1-USP12 structures not only reveal a series of structural changes of the enzyme that are connected to the binding site of UAF1 but also highlight the dynamic nature of three key lo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Based on our result, we propose a tenable model for USP12 activation by UAF1 and WDR20 ( xref )."

sparser
"USP12 and USP46, but not USP1, can be independently activated by another WD40-repeat-containing protein, WDR20 (Burska et al., 2013; Dahlberg and Juo, 2014; Kee et al., 2010; McClurg et al., 2014) ."

sparser
"Mechanism of USP12 Activation by UAF1 and WDR20."

reach
"Both WDR48 and WDR20 stimulate USP12 and USP46 catalytic activity (k cat) without increasing substrate binding affinity, suggesting that the WDR proteins may affect DUB activity via a novel structural mechanism."

sparser
"Based on our results, we propose a tenable model for USP12 activation by UAF1 and WDR20 (Figure 7) ."

sparser
"Interestingly, USP12 and USP46 are activated by two β-propeller proteins, UAF1, and WDR20."
WDR20 decreases the amount of USP12.
| 1
WDR20 decreases the amount of USP12. 1 / 1
| 1

reach
"As Uaf-1 and WDR20 depletion reduces Usp12 levels their silencing has a comparable effect on the AR as the Usp12 depletion alone."
USP12 affects WDR20
8 | 34 22
USP12 binds WDR20.
8 | 33 22
8 | 13 9

reach
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far away from its catalytic center."

sparser
"Consistent with our results on USP1, activation of USP12-WDR20 by UAF1 is achieved mainly through an 20-fold increase of k cat with little change in K M , suggesting that substrate affinity of USP12 is not altered by UAF1 ( Figures 6A and S5A-S5D) ."

No evidence text available

No evidence text available

No evidence text available

sparser
"In the two UAF1-USP12 complex structures, UAF1 binding appears to induce BL1 destabilization via a chain of conformational changes, whereas the WDR20-USP12 interaction seems to overwrite the allosteric effect of UAF1 by re-stabilizing the loop."

reach
"Results confirmed the interaction between endogenous Usp12, Uaf-1 and WDR20 in PC cells."

sparser
"Adjacent to the major USP12-WDR20 interface, the tip of USP12 BL3, which is otherwise disordered in both free and UAF1-bound forms of the enzyme, adopts a stable structure by making van der Waal contacts with WDR20."

reach
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

sparser
"The k cat and K M values are presented as means ± SD (n = 3), and the K D values were obtained through steady-state analysis (see Supplemental Experimental Procedures for technical discussion). (B) Time course of USP12 modification by the suicidal Ub-VME substrate in the absence or presence of UAF1 and WDR20. (C) Time course of LRGG-AMC (top) in contrast to Ub-AMC (bottom) hydrolysis by USP12 (green), UAF1-USP12 (orange), and USP12-WDR20 (red)."
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 inhibits WDR20.
| 1
USP12 inhibits WDR20. 1 / 1
| 1

reach
"Consistent with an important role in binding USP12, mutations of W306 and F262 effectively abolish the ability of WDR20 to interact with and activate USP12 ( Figures 4C and S4D )."
USP12 affects USP46
2 | 15 34
USP12 binds USP46.
2 | 12 34
2 | 12 11

reach
"The more distantly related USP1 is also regulated by WDR48, but WDR20 is unique to USP12 and USP46 complexes."

sparser
"BIAcore SPR studies demonstrated a dose dependent binding of galeterone to USP12 and USP46 (Figure xref )."

reach
"Conversely, in the CWR22Rv1 cell line silencing AR FL alone decreased galeterone efficiency by 4 fold compared to SCR control and silencing USP12 or USP12 and USP46 complex members had no significant effect on galeterone efficacy."

sparser
"The unique binding mode elucidated here thus provides a framework to guide further exploration into the role of the larger USP1 insertions and the particular interactions of USP12 and USP46 in mediati[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Similarly galeterone inhibition did not have a significant effect on transcript levels of USP12 and USP46 complex members UAF1 (adj. p = 0.4) and WDR20 (adj. p = 0.25)."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"Biochemical and proteomic studies showed that two WDR proteins, WDR48 (also known as USP1-associated factor, UAF1) and WDR20, interact with USP46 and USP12 (Cohn et al., 2009; Sowa et al., 2009; Kee et al., 2010)."

reach
"Our data demonstrates a proof-of-principle for USP12 and USP46 complex targeting in PC and uncovers additional mechanisms of galeterone activity."

sparser
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis."

reach
"We therefore tested C527 for its ability to inhibit the purified USP12 and USP46 complex or other DUBs."
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
TBK1 binds USP46 and USP12. 1 / 1
| 1

sparser
"In addition, USP12 and USP46 were not associated with TBK1 ( xref )."
USP12 activates USP46.
| 2
USP12 activates USP46. 1 / 1
| 1

reach
"Identification of Usp12 as a modifier of HD suggested that Usp12 could rescue neurodegeneration in related disorders, such as amyotrophic lateral sclerosis (ALS) and Parkinson 's disease (PD) that are characterized by proteotoxic stress."
USP12 bound to WDR48 activates USP46. 1 / 1
| 1

reach
"In C. elegans, USP46 and USP12 both bind to USP1 associated factor 1 (UAF1 and WDR48) and the binding dramatically enhances the activity of USP12 and USP46."
USP12 inhibits USP46.
| 1
USP12 inhibits USP46. 1 / 1
| 1

reach
"We propose a mechanism by which Usp12 suppresses HD related neurodegeneration via induction of autophagy."
USP46 affects USP12
2 | 13 34
USP46 binds USP12.
2 | 12 34
2 | 12 11

reach
"The more distantly related USP1 is also regulated by WDR48, but WDR20 is unique to USP12 and USP46 complexes."

sparser
"BIAcore SPR studies demonstrated a dose dependent binding of galeterone to USP12 and USP46 (Figure xref )."

reach
"Conversely, in the CWR22Rv1 cell line silencing AR FL alone decreased galeterone efficiency by 4 fold compared to SCR control and silencing USP12 or USP12 and USP46 complex members had no significant effect on galeterone efficacy."

sparser
"The unique binding mode elucidated here thus provides a framework to guide further exploration into the role of the larger USP1 insertions and the particular interactions of USP12 and USP46 in mediati[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Similarly galeterone inhibition did not have a significant effect on transcript levels of USP12 and USP46 complex members UAF1 (adj. p = 0.4) and WDR20 (adj. p = 0.25)."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"Biochemical and proteomic studies showed that two WDR proteins, WDR48 (also known as USP1-associated factor, UAF1) and WDR20, interact with USP46 and USP12 (Cohn et al., 2009; Sowa et al., 2009; Kee et al., 2010)."

reach
"Our data demonstrates a proof-of-principle for USP12 and USP46 complex targeting in PC and uncovers additional mechanisms of galeterone activity."

sparser
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis."

reach
"We therefore tested C527 for its ability to inhibit the purified USP12 and USP46 complex or other DUBs."
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
TBK1 binds USP46 and USP12. 1 / 1
| 1

sparser
"In addition, USP12 and USP46 were not associated with TBK1 ( xref )."
USP46 activates USP12.
| 1
USP46 bound to WDR48 activates USP12. 1 / 1
| 1

reach
"In C. elegans, USP46 and USP12 both bind to USP1 associated factor 1 (UAF1 and WDR48) and the binding dramatically enhances the activity of USP12 and USP46."
USP12 affects AR
3 1 | 23 6
USP12 binds AR.
3 | 3 5
3 | 3 5

sparser
"Although we were able to confirm the USP12- AR FL interaction we could not detect any interaction between USP12 and AR V7 (Figure xref )."

sparser
"To validate this result we overexpressed USP12 with AR FL, N-terminus of AR (AR ND) and AR N-terminus with DNA binding domain (AR ND-DNA BD) to localise the USP12-AR interaction."

reach
"The requirement of the AR ligand binding domain to ensure an interaction between USP12 and AR is supported by our results in Figure XREF_FIG showing that AR FL is stabilised following USP12 overexpression whereas the AR V7 proteins levels are unaffected."

sparser
"The requirement of the AR ligand binding domain to ensure an interaction between USP12 and AR is supported by our results in Figure xref showing that AR FL is stabilised following USP12 overexpression whereas the AR V7 proteins levels are unaffected (Figure xref )."

sparser
"Usp12 was capable of interacting with both AR S213A and AR S213A/S791A with comparable efficiency to wild-type confirming that these sites are not required for this interaction (Figure xref )."

No evidence text available

No evidence text available

reach
"To investigate if this was caused by the lack of interaction between AR and Usp12 in the absence of the specific serine residues, we overexpressed Usp12 alongside wild-type and mutant ARs followed by immunoprecipitation."

reach
"We have recently reported that Usp12, in complex with Uaf-1 and WDR20, can directly bind and deubiquitinate the AR resulting in increased receptor stability and transcriptional activity [XREF_BIBR]."

No evidence text available
USP12 activates AR.
| 7 1
USP12 activates AR. 8 / 10
| 7 1

reach
"Cell lines used in this study express low levels of endogenous Usp12 and Uaf-1, consequently even without the addition of Uaf-1 overexpression of WDR20 alone was enough to significantly enhance the AR activity."

reach
"Deubiquitinating enzyme Usp12 is a novel co-activator of the androgen receptor."

reach
"Our previous observations for the LNCaP cell line that express AR FL but not AR Vs have demonstrated that USP12 silencing causes a decrease in transcript expression of all the AR target genes [XREF_BIBR] supporting our data that AR Vs are not targeted by USP12 in the AR V expressing CWR22Rv1 cell line."

reach
"Interestingly, Usp12 and WDR20 alone significantly increased AR activity but this effect was greatest when Usp12, Uaf-1 and WDR20 were overexpressed simultaneously."

reach
"Also, USP12 co-localizes with AR in the cytoplasm and promotes AR transcriptional activity by confronting the ubiquitin dependent degradation of AR [XREF_BIBR]."

sparser
"This data suggests that AR co-activation by Usp12 and Uaf-1 is largely driven through negating AR phosphorylation by Akt inactivation."

reach
"This further confirms that each complex member alone has a limited effect on AR activity but in combination the Usp12 complex significantly increases AR transcriptional activity implying that targeting any of the binding partners should offer the same efficacy as targeting Usp12 alone."

reach
"We have recently reported that Usp12, in combination with Uaf-1, can increase the transcriptional activity of AR [XREF_BIBR], and hence we wanted to analyse the importance of Akt phosphorylation sites in this process."
USP12 deubiquitinates AR.
1 | 5
USP12 deubiquitinates AR. 6 / 6
1 | 5

reach
"USP12 in the presence of its cofactors Uaf-1 and WDR20 deubiquitinates AR to enhance AR protein stability and transcriptional activity [71] USP12 in complex with Uaf-1 and WDR20 also abrogates phospho[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

"The AR is known to be regulated by a number of post-translational modifications and we have recently identified the deubiquitinating enzyme Usp12 as a positive regulator of AR."

reach
"We determined that Usp12 deubiquitinates the AR resulting in elevated receptor stability and activity."

reach
"We have recently reported that AR is deubiquitinated and stabilised by Usp12 resulting in increased transcriptional activity."

reach
"We find that USP12 deubiquitinates MDM2 and AR, which in turn controls the levels of the TP53 tumour suppressor and AR oncogene in prostate cancer."

reach
"In this report we demonstrate that Usp12, in complex with Uaf-1 and WDR20, deubiquitinates the AR to enhance receptor stability and transcriptional activity."
USP12 inhibits AR.
| 2
USP12 inhibits AR. 2 / 3
| 2

reach
"Also, USP12 co-localizes with AR in the cytoplasm and promotes AR transcriptional activity by confronting the ubiquitin dependent degradation of AR [XREF_BIBR]."

reach
"USP12 inhibition in AR V positive lines drives the AR V expression profile."
USP12 dephosphorylates AR.
| 3
USP12 leads to the dephosphorylation of AR on S215. 3 / 3
| 3

reach
"In agreement with the above result, knockdown of Usp12 in LNCaP cells increased both AR S213 phosphorylation and AR turnover."

reach
"As such, overexpressing Usp12 inhibits AR S213 phosphorylation resulting in increased AR stability and transcriptional activity."

reach
"Our data demonstrates that Usp12 increases the levels of PHLPP and PHLPPL that in turn decreases the active pAkt pool and inhibits AR S213 phosphorylation by Akt."
USP12 increases the amount of AR.
| 2
USP12 increases the amount of AR. 2 / 2
| 2

reach
"Our data suggests that USP12 is unable to modulate AR V levels or activity but is capable of targeting AR FL."

reach
"Our previous observations for the LNCaP cell line that express AR FL but not AR Vs have demonstrated that USP12 silencing causes a decrease in transcript expression of all the AR target genes [XREF_BIBR] supporting our data that AR Vs are not targeted by USP12 in the AR V expressing CWR22Rv1 cell line."
USP12 ubiquitinates AR.
| 1
USP12 leads to the ubiquitination of AR. 1 / 1
| 1

reach
"Prior studies suggested a role of USP12 in prostate carcinomas, wherein USP12 promoted AR signalling and tumour cell proliferation, as well as therapy resistance, by regulating AR ubiquitination and MDM2-P53 signalling ."
WDR48 affects WDR20
| 20 9
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 affects PHLPP1
5 1 | 14 4
USP12 binds PHLPP1.
5 | 6 4
5 | 6 2

reach
"We demonstrated that Usp12, and two Usp12 interacting proteins, Uaf-1 and WDR20 [XREF_BIBR], [XREF_BIBR], form a complex with PHLPP and PHLPPL."

reach
"Additionally, Usp12 was predicted to interact with both of these phosphatases and a more recent report confirmed the interaction between Usp12 and PHLPP in colorectal cells [XREF_BIBR]."

No evidence text available

reach
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs (XREF_FIG C)."

No evidence text available

No evidence text available

sparser
"Additionally, Usp12 was predicted to interact with both of these phosphatases and a more recent report confirmed the interaction between Usp12 and PHLPP in colorectal cells [ xref ]."

reach
"In agreement with the previous study, we found that both PHLPP and PHLPPL interacted with Usp12."

sparser
"In CRC, a mutation in the protein WD repeat domain 48, required for interaction between PHLPP1 and USP12, resulted in PHLPP1 downregulation and consequent increase in AKT signaling ( Gangula and Maddi[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Here we report that Usp12, in complex with Uaf-1 and WDR20, interacts with PHLPP and PHLPPL in PC cells resulting in their deubiquitination and protein stabilisation."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
USP12 deubiquitinates PHLPP1.
1 | 3
USP12 deubiquitinates PHLPP1. 4 / 4
1 | 3

reach
"Usp12 deubiquitinates and stabilises PHLPP and PHLPPL and controls Akt activation status."

reach
"Further analysis using denaturing immunoprecipitation experiments revealed that Usp12 can also deubiquitinate and stabilise both PHLPP and PHLPPL."

"USP11 stabilizes IKK-α?to stabilize and activate p53"

reach
"Secondly, Usp12 deubiquitinates and stabilises PHLPP and PHLPPL proteins which enhances dephosphorylation and deactivation of Akt."
USP12 increases the amount of PHLPP1.
| 2
USP12 increases the amount of PHLPP1. 1 / 2
| 1

reach
"Our data demonstrates that Usp12 increases the levels of PHLPP and PHLPPL that in turn decreases the active pAkt pool and inhibits AR S213 phosphorylation by Akt."
Modified USP12-C48A increases the amount of PHLPP1. 1 / 1
| 1

reach
"Importantly, overexpression of Usp12 deubiquitinated both PHLPP and PHLPPL and this elevated the steady-state levels of the enzymes, while overexpression of an enzymatically inactive Usp12 C48A mutant failed to elevate PHLPP and PHLPPL levels suggesting the importance of Usp12 enzymatic activity for phosphatase regulation."
USP12 inhibits PHLPP1.
| 2
USP12 inhibits PHLPP1. 2 / 2
| 2

reach
"However, USP12 has also been shown to inhibit AKT-dependent cell survival by stabilizing the pro-apoptotic phosphatases PHLPP1 and/or PHLPPL ."

reach
"Because PHLPP1 is a potent negative regulator of the PI3-kinase and Akt pathway, we hypothesized that WDR48 and USP12 might suppress Akt signaling by stabilizing PHLPP1."
USP12 activates PHLPP1.
| 1
USP12 activates PHLPP1. 1 / 2
| 1

reach
"In fact, overexpression of WDR48 and USP12 induced apoptosis similarly to PHLPP1."
USP12 affects WDR20, and WDR48
| 20 6
| 20 6

reach
"During revision of this manuscript, a structure for the USP12, UAF1, and WDR20 complex was published."

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."

sparser
"Our structures reveal that UAF1 and WDR20 interact with USP12 at two distinct sites far from its catalytic center."

sparser
"Binding of WDR20 to the UAF1-USP12 complex induces multiple structural changes in the enzyme, converting it into a compact form similar to Ub-bound USP46."

reach
"Our data suggest that the USP12, UAF1, and WDR20 complex dynamically shuttles between the plasma membrane, cytoplasm and nucleus."

sparser
"Structural Comparisons of Activator-Bound USP12 and Ub-Conjugated USP46(A) Superposition of UAF1-bound USP12 and UAF1-WDR20-bound USP12 highlighting stabilization of BL1 and BL3 in the ternary complex.(B and C) A comparison between the UAF1-WDR20-bound USP12 and the Ub-conjugated USP46 structures, focusing on the stabilized PK Loop, the regularized PK Helix, and the F219 residue at the Fingers-Palm domain junction. (D) A close-up view of BL1 and the catalytic cleft of USP12 in complex with UAF1 and WDR20."

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"WDR20 plays a crucial role in as a " targeting subunit " that modulates CRM1 dependent shuttling of the USP12, UAF1, and WDR20 complex between the plasma membrane, cytoplasm and nucleus."

reach
"Firstly, we confirm the presence of the Usp12, Uaf-1, and WDR20 complex in PC cells and demonstrate the importance of Uaf-1 and WDR20 for Usp12 stabilisation."

reach
"Using site directed mutagenesis, we generated a large set of USP12 and WDR20 mutants to characterize in detail the mechanisms and sequence determinants that modulate the subcellular localization of the USP12, UAF1, and WDR20 complex."
USP12 affects PHLPP2
4 1 | 9 4
USP12 binds PHLPP2.
4 | 3 4
4 | 3 2

reach
"We demonstrated that Usp12, and two Usp12 interacting proteins, Uaf-1 and WDR20 [XREF_BIBR], [XREF_BIBR], form a complex with PHLPP and PHLPPL."

No evidence text available

No evidence text available

No evidence text available

sparser
"In agreement with the previous study, we found that both PHLPP (Figure xref ) and PHLPPL (Figure xref ) interacted with Usp12."

reach
"In agreement with the previous study, we found that both PHLPP and PHLPPL interacted with Usp12."

sparser
"The Usp12 complex interacts with PHLPP and PHLPPL."

reach
"Here we report that Usp12, in complex with Uaf-1 and WDR20, interacts with PHLPP and PHLPPL in PC cells resulting in their deubiquitination and protein stabilisation."

No evidence text available
| 1

sparser
"To establish if this was the case in PC cell lines, we firstly investigated potential Usp12-PHLPP/L interactions in androgen-dependent LNCaP cells by immunoprecipitation."
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
USP12 deubiquitinates PHLPP2.
1 | 3
USP12 deubiquitinates PHLPP2. 4 / 4
1 | 3

"we demonstrate that Usp12, in complex with Uaf-1 and WDR20, directly deubiquitinates and stabilises the Akt phosphatases PHLPP and PHLPPL resulting in decreased levels of active pAkt."

reach
"Further analysis using denaturing immunoprecipitation experiments revealed that Usp12 can also deubiquitinate and stabilise both PHLPP and PHLPPL."

reach
"Secondly, Usp12 deubiquitinates and stabilises PHLPP and PHLPPL proteins which enhances dephosphorylation and deactivation of Akt."

reach
"Usp12 deubiquitinates and stabilises PHLPP and PHLPPL and controls Akt activation status."
USP12 increases the amount of PHLPP2.
| 2
USP12 increases the amount of PHLPP2. 1 / 2
| 1

reach
"Our data demonstrates that Usp12 increases the levels of PHLPP and PHLPPL that in turn decreases the active pAkt pool and inhibits AR S213 phosphorylation by Akt."
Modified USP12-C48A increases the amount of PHLPP2. 1 / 1
| 1

reach
"Importantly, overexpression of Usp12 deubiquitinated both PHLPP and PHLPPL and this elevated the steady-state levels of the enzymes, while overexpression of an enzymatically inactive Usp12 C48A mutant failed to elevate PHLPP and PHLPPL levels suggesting the importance of Usp12 enzymatic activity for phosphatase regulation."
USP12 inhibits PHLPP2.
| 1
USP12 inhibits PHLPP2. 1 / 1
| 1

reach
"However, USP12 has also been shown to inhibit AKT-dependent cell survival by stabilizing the pro-apoptotic phosphatases PHLPP1 and/or PHLPPL ."
WDR48 affects USP46
| 17
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USP12 affects CREBBP
| 15 1
| 9

sparser
"USP12 interacts with CBP and blocks the acetyltransferase activity of CBP."

sparser
"Moreover, we demonstrated that in IFN signaling, USP12 interacted with CBP but not with either p-STAT1 or TCPTP ( xref ), suggesting that USP12 affects the p-STAT1 and TCPTP interaction through targeting CBP."

sparser
"In our mass spectroscopy analysis, we noticed an interaction between USP12 and CBP."

sparser
"Together, these findings demonstrated that USP12 constitutively binds with CBP in cells."

sparser
"We also demonstrated that USP12 interacted with CBP, but not either p-STAT1 or TCPTP, suggesting that the regulation of USP12 on both TCPTP binding and p-STAT1 levels is through CBP."

sparser
"Thus, we further confirmed the interaction between USP12 and CBP."

sparser
"In bone marrow-derived macrophages (BMDMs) isolated from mice and RAW264.7 cells, endogenous USP12 also interacted with endogenous CBP ( xref )."

sparser
"The results showed that endogenous USP12 interacted with endogenous CBP in intact cells, and IFNα treatment promoted USP12 nuclear accumulation ( xref and xref )."

sparser
"Importantly, USP12 constitutively binds with CBP in mouse primary lung and liver cells ( xref )."
| 6

sparser
"The aforementioned results demonstrated that USP12 binds to the HAT domain of CBP and blocks CBP’s acetylation action."

sparser
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP’s acetyltransferase activity."

sparser
"Taken together, these findings suggest that USP12 binds to the HAT domain of CBP and blocks CBP’s acetyltransferase activity in cells."

sparser
"In vitro binding assay further demonstrated that USP12 binds with the HAT domain of CBP ( xref )."

sparser
"Interestingly, USP12 can interact with the HAT domain of CBP."

sparser
"In this study, we demonstrated that USP12 can bind with the HAT domain of CBP in vivo and in vitro ."
CREBBP binds USP12. 1 / 1
| 1

isi
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP's acetyltransferase activity."
PHLPP1 affects USP12
5 | 6 4
5 | 6 2

reach
"We demonstrated that Usp12, and two Usp12 interacting proteins, Uaf-1 and WDR20 [XREF_BIBR], [XREF_BIBR], form a complex with PHLPP and PHLPPL."

reach
"Additionally, Usp12 was predicted to interact with both of these phosphatases and a more recent report confirmed the interaction between Usp12 and PHLPP in colorectal cells [XREF_BIBR]."

No evidence text available

reach
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs (XREF_FIG C)."

No evidence text available

No evidence text available

sparser
"Additionally, Usp12 was predicted to interact with both of these phosphatases and a more recent report confirmed the interaction between Usp12 and PHLPP in colorectal cells [ xref ]."

reach
"In agreement with the previous study, we found that both PHLPP and PHLPPL interacted with Usp12."

sparser
"In CRC, a mutation in the protein WD repeat domain 48, required for interaction between PHLPP1 and USP12, resulted in PHLPP1 downregulation and consequent increase in AKT signaling ( Gangula and Maddi[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Here we report that Usp12, in complex with Uaf-1 and WDR20, interacts with PHLPP and PHLPPL in PC cells resulting in their deubiquitination and protein stabilisation."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
CREBBP affects USP12
| 15 1
| 9

sparser
"USP12 interacts with CBP and blocks the acetyltransferase activity of CBP."

sparser
"Moreover, we demonstrated that in IFN signaling, USP12 interacted with CBP but not with either p-STAT1 or TCPTP ( xref ), suggesting that USP12 affects the p-STAT1 and TCPTP interaction through targeting CBP."

sparser
"In our mass spectroscopy analysis, we noticed an interaction between USP12 and CBP."

sparser
"Together, these findings demonstrated that USP12 constitutively binds with CBP in cells."

sparser
"We also demonstrated that USP12 interacted with CBP, but not either p-STAT1 or TCPTP, suggesting that the regulation of USP12 on both TCPTP binding and p-STAT1 levels is through CBP."

sparser
"Thus, we further confirmed the interaction between USP12 and CBP."

sparser
"In bone marrow-derived macrophages (BMDMs) isolated from mice and RAW264.7 cells, endogenous USP12 also interacted with endogenous CBP ( xref )."

sparser
"The results showed that endogenous USP12 interacted with endogenous CBP in intact cells, and IFNα treatment promoted USP12 nuclear accumulation ( xref and xref )."

sparser
"Importantly, USP12 constitutively binds with CBP in mouse primary lung and liver cells ( xref )."
| 6

sparser
"The aforementioned results demonstrated that USP12 binds to the HAT domain of CBP and blocks CBP’s acetylation action."

sparser
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP’s acetyltransferase activity."

sparser
"Taken together, these findings suggest that USP12 binds to the HAT domain of CBP and blocks CBP’s acetyltransferase activity in cells."

sparser
"In vitro binding assay further demonstrated that USP12 binds with the HAT domain of CBP ( xref )."

sparser
"Interestingly, USP12 can interact with the HAT domain of CBP."

sparser
"In this study, we demonstrated that USP12 can bind with the HAT domain of CBP in vivo and in vitro ."
CREBBP binds USP12. 1 / 1
| 1

isi
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP's acetyltransferase activity."
USP12 affects autophagy
| 2 12
| 2 12

reach
"Usp12 stimulation of autophagy, in the context of mHTT, could lead to more efficient cargo loading in autophagosomes, and thereby reduce mHTT associated toxicity."

reach
"USP12 also non-catalytically induces cell autophagy in Huntington’s disease ."

reach
"Another possibility is that Usp12 stimulation of autophagy confers neuroprotection primarily by clearing cargo other than mHTT."

reach
"Both alpha-synuclein and TDP-43 are substrates for autophagic degradation XREF_BIBR, XREF_BIBR, XREF_BIBR, and so, we might predict that stimulation of autophagy by Usp12 overexpression would also be neuroprotective in the related PD and ALS neuron models."

reach
"Deubiquitinase Usp12 functions noncatalytically to induce autophagy and confer neuroprotection in models of Huntington 's disease."

reach
"Surprisingly, the catalytic activity of Usp12 is not required to protect against neurodegeneration or induce autophagy."

reach
"Depletion of USP12 suppressed cell growth and clonogenicity and inhibited autophagy."

reach
"Publisher Correction : Deubiquitinase Usp12 functions noncatalytically to induce autophagy and confer neuroprotection in models of Huntington 's disease."

reach
"Deubiquitylase USP12 induces pro-survival autophagy and bortezomib resistance in multiple myeloma by stabilizing HMGB1."

eidos
"Depletion of USP12 suppressed cell growth and clonogenicity and inhibited autophagy ."
| 4 9
USP12 translocates from the cytoplasm to the nucleus. 5 / 5
| 1 4

reach
"Furthermore, we found that in IFN signaling USP12 translocates from the cytoplasm to the nucleus."

sparser
"USP12 translocates from the cytoplasm to the nucleus in IFN signaling."

sparser
"Furthermore, we found that in IFN signaling USP12 translocates from the cytoplasm to the nucleus."

sparser
"To our surprise, in IFN signaling USP12 translocates from the cytoplasm to the nucleus."

sparser
"To our surprise, we observed that during mIFNβ treatment, cytoplasmic USP12 protein levels dramatically decreased and nuclear USP12 protein levels increased accordingly ( xref ), suggesting that USP12 translocates from the cytoplasm to the nucleus in IFN signaling."
USP12 translocates to the nucleus. 3 / 3
| 1 2

reach
"Lehoux et al., found that USP12 and USP46 fused to red fluorescent protein were predominantly cytoplasmic in C33A cervical carcinoma cells, but were recruited to the nucleus via an interaction with the HPV E1 helicase [XREF_BIBR]."

sparser
"Given that we have demonstrated that USP12 translocates into the nucleus in IFN signaling, we next sought to determine whether USP12 regulates p-STAT1 levels through CBP."

sparser
"USP12 translocates into the nucleus in IFN signaling, which critically regulates nuclear p-STAT1 levels and IFN antiviral activity by inhibiting CBP’s acetyltransferase activity."
USP12 translocates from the nucleus to the cytosol. 2 / 2
| 1 1

sparser
"Stimulation with anti-CD3 resulted in phosphorylation and time-dependent translocation of Usp12 from the nucleus to the cytosol."

reach
"Stimulation with anti-CD3 resulted in phosphorylation and time dependent translocation of Usp12 from the nucleus to the cytosol."
USP12 translocates from the nucleus to the cytoplasm. 2 / 2
| 2

sparser
"It has been reported that in TCR signaling, USP12 translocates from the nucleus to the cytoplasm to deubiquitinate LAT and Trat1 proteins [ xref ]."

sparser
"A recent report showed that in TCR signaling, USP12 translocates from the nucleus to the cytoplasm, where USP12 deubiquitinates and stabilizes LAT and Trat1 and prevents their lysosome-dependent degradation in T cells."
USP12 translocates from the cytoplasm. 1 / 1
| 1

reach
"XREF_BIBR Liu and colleagues demonstrate that Type 1 IFN signaling stimulates the translocation of USP12 from cytoplasm to the nucleus, that blocks the acetylation effect of CREB binding protein (CBP) in nucleus."
AR affects USP12
3 | 5 5
AR binds USP12.
3 | 3 5
3 | 3 5

sparser
"Although we were able to confirm the USP12- AR FL interaction we could not detect any interaction between USP12 and AR V7 (Figure xref )."

sparser
"To validate this result we overexpressed USP12 with AR FL, N-terminus of AR (AR ND) and AR N-terminus with DNA binding domain (AR ND-DNA BD) to localise the USP12-AR interaction."

reach
"The requirement of the AR ligand binding domain to ensure an interaction between USP12 and AR is supported by our results in Figure XREF_FIG showing that AR FL is stabilised following USP12 overexpression whereas the AR V7 proteins levels are unaffected."

sparser
"The requirement of the AR ligand binding domain to ensure an interaction between USP12 and AR is supported by our results in Figure xref showing that AR FL is stabilised following USP12 overexpression whereas the AR V7 proteins levels are unaffected (Figure xref )."

sparser
"Usp12 was capable of interacting with both AR S213A and AR S213A/S791A with comparable efficiency to wild-type confirming that these sites are not required for this interaction (Figure xref )."

No evidence text available

No evidence text available

reach
"To investigate if this was caused by the lack of interaction between AR and Usp12 in the absence of the specific serine residues, we overexpressed Usp12 alongside wild-type and mutant ARs followed by immunoprecipitation."

reach
"We have recently reported that Usp12, in complex with Uaf-1 and WDR20, can directly bind and deubiquitinate the AR resulting in increased receptor stability and transcriptional activity [XREF_BIBR]."

No evidence text available
AR activates USP12.
| 2
AR activates USP12. 2 / 2
| 2

reach
"USP12 in the presence of its cofactors Uaf-1 and WDR20 deubiquitinates AR to enhance AR protein stability and transcriptional activity [71] USP12 in complex with Uaf-1 and WDR20 also abrogates phospho[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Akt inhibition abrogated S213 phosphorylation of AR and decreased the ability of Usp12 to stabilise AR protein levels."
PHLPP2 affects USP12
4 | 3 4
4 | 3 2

reach
"We demonstrated that Usp12, and two Usp12 interacting proteins, Uaf-1 and WDR20 [XREF_BIBR], [XREF_BIBR], form a complex with PHLPP and PHLPPL."

No evidence text available

No evidence text available

No evidence text available

sparser
"In agreement with the previous study, we found that both PHLPP (Figure xref ) and PHLPPL (Figure xref ) interacted with Usp12."

reach
"In agreement with the previous study, we found that both PHLPP and PHLPPL interacted with Usp12."

sparser
"The Usp12 complex interacts with PHLPP and PHLPPL."

reach
"Here we report that Usp12, in complex with Uaf-1 and WDR20, interacts with PHLPP and PHLPPL in PC cells resulting in their deubiquitination and protein stabilisation."

No evidence text available
| 1

sparser
"To establish if this was the case in PC cell lines, we firstly investigated potential Usp12-PHLPP/L interactions in androgen-dependent LNCaP cells by immunoprecipitation."
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
UAF1 affects USP12
| 10
UAF1 activates USP12. 10 / 10
| 10

eidos
"Based on our results , we propose a tenable model for USP12 activation by UAF1 and WDR20 ( Figure 7 ) ."

eidos
"First , our results rule out the possibility that UAF1 and WDR20 activate USP12 by enhancing its substrate affinity ."

eidos
"With a mutation introduced to the BL1 residue F262 , USP12 could no longer be activated by UAF1 but retained full WDR20-stimulated activity ( Figures 6D and 6E ) ."

eidos
"To gain deeper mechanistic insights into USP12 activation by UAF1 and WDR20 , we first determined the steady-state kinetics of substrate hydrolysis by USP12 in different forms ."

eidos
"To delineate the mechanism by which UAF1 activates USP12 , we next determined the crystal structure of USP12 in a complex with full-length human UAF1 in the F222 space group at 3 ."

eidos
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases ( USPs ) constitute the largest family of deubiquitinating enzymes , whose catalytic competency is often modulated by their binding partners through unknown mechanisms ."

eidos
"Activation of USP12 by UAF1 requires structural fine-tuning at the junction of the Ub globular domain and it tail through BL1 , whereas WDR20 acts independently of this site and directly affects the catalytic cleft ."

eidos
"This result clearly indicates that UAF1 and WDR20 allosterically activate USP12 through two distinct pathways ."

eidos
"UAF1 activates USP12 by binding to the tip of its Fingers domain and transmitting a long-range allosteric signal to BL1 ."

eidos
"Nevertheless , the free USP12 structure alone is insufficient to link these predicted changes to USP12 activation by UAF1 or WDR20 ."
USP12 affects USP46, and WDR48
| 9
| 9

sparser
"In addition to USP1, two other DUBs, USP12 and USP46 interact with UAF1 [ xref ], but the UAF1-binding sites in these DUBs have not yet been mapped."

sparser
"In C. elegans , USP46 and USP12 both bind to USP1-associated factor 1 (UAF1/WDR48) and the binding dramatically enhances the activity of USP12 and USP46 ( xref , xref ; xref )."

sparser
"UAF1 also binds and activates two other DUBs, USP12 and USP46 ( xref ), and studies that reveal how UAF1 binds and activates USP12 and USP46 suggest that UAF1 will bind to USP1 in an analogous manner ( xref , xref )."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"UAF1 also interacts with DUBs USP12 and USP46 ( García-Santisteban et al., 2013 ) and, thus, both genes also could be involved in senescence."

sparser
"Recent crystallographic studies demonstrate that UAF1 binds to a distinct site on USP12 or USP46 and, through an allosteric interaction, stimulates DUB activity ( xref ; xref )."

sparser
"The mode of interaction of USP12 and USP46 with UAF1 is still unclear."

sparser
"Recent structural work revealed how the WD-repeat protein UAF1 interacted with the catalytic domain of USP46 and USP12, which is facilitated predominantly through the “fingers” subdomain of the USP core, mediating long-range allosteric interactions eventually leading to enhanced enzyme efficiency ( xref , xref )."

sparser
"P65 interacted with USP12, USP46 and UAF1 in both resting and LPS-stimulated macrophages, but exhibited no association with USP1 (Fig.  xref )."

reach
"As a result, depleting Usp12 decreased PC cellular proliferation and increased cellular apoptosis suggesting it may be a potential target for CRPC therapy [XREF_BIBR]."

reach
"As shown in XREF_FIG B, knockdown of WDR48 and USP12 by shRNA accelerated the rate of cell proliferation compared with control shRNA transfected cells."

reach
"Functionally, we show that WDR48 and USP12 suppress proliferation of tumor cells."

reach
"In contrast, overexpression of WDR48 and USP12 (XREF_FIG C) suppressed cell proliferation similarly to PHLPP1 (XREF_FIG D)."

reach
"We further established that Usp12 depletion reduced PC cell proliferation and induced apoptosis, furthermore Usp12 levels are increased in PC tissue [XREF_BIBR]."

reach
"Furthermore, Usp12 silencing was shown to reduce proliferation of PC cells."
USP12 affects PPM1B
| 8
USP12 deubiquitinates PPM1B.
| 4
USP12 deubiquitinates PPM1B. 4 / 4
| 4

reach
"The interaction between PPM1B and USP12 was then validated (Fig. 4c), raising the question of whether USP12 deubiquitinates PPM1B."

reach
"USP12 regulates chemokine production by deubiquitinating PPM1B."

reach
"We found that USP12 downregulation caused insufficient deubiquitination of PPM1B and thereby facilitated tumour growth by promoting the generation of the protumourigenic milieu."

reach
"The ubiquitination experiments showed that ectopic expression of USP12 but not the C48S mutant decreased the levels of PPM1B polyubiquitination (Fig. 4d)."
USP12 ubiquitinates PPM1B.
| 1
USP12 leads to the ubiquitination of PPM1B. 1 / 1
| 1

reach
"USP12 downregulation in tumour cells accelerates PPM1B ubiquitination and degradation and therefore promotes NF-κB activity in orchestrating the TME."
USP12 increases the amount of PPM1B.
| 1
USP12 increases the amount of PPM1B. 1 / 1
| 1

reach
"As expected, USP12 knockdown decreased the levels of both PPM1B and IκB (Fig. 4e)."
USP12 decreases the amount of PPM1B.
| 1
USP12 decreases the amount of PPM1B. 1 / 1
| 1

reach
"The ubiquitination experiments showed that ectopic expression of USP12 but not the C48S mutant decreased the levels of PPM1B polyubiquitination (Fig. 4d)."
USP12 binds PPM1B.
| 1
| 1

reach
"The interaction between PPM1B and USP12 was then validated (Fig. 4c), raising the question of whether USP12 deubiquitinates PPM1B."
USP1 affects USP12
| 1 7
USP1 binds USP12.
| 7
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
USP1 activates USP12.
| 1
USP1 activates USP12. 1 / 1
| 1

reach
"The WD-repeat protein WDR48 (USP1 associated factor UAF-1) stimulates activity of ubiquitin specific proteases USP1, USP12, and USP46."
| 1 6
| 1 5

reach
"WDR48 and USP12 Negatively Regulate Akt Signaling and Promote Apoptosis."

reach
"In fact, overexpression of WDR48 and USP12 induced apoptosis similarly to PHLPP1."

reach
"Thus, these results suggest that WDR48 and USP12 negatively regulate Akt signaling and thereby induce apoptosis in conjunction with PHLPP1."

reach
"As a result, depleting Usp12 decreased PC cellular proliferation and increased cellular apoptosis suggesting it may be a potential target for CRPC therapy [XREF_BIBR]."

eidos
"In vivo experiments showed that USP12-knockdown could suppress tumor growth in mice , and immuno-blotting revealed that USP12 could induce G2 / M arrest through the cyclin dependent kinase 1 / cyclinB1 axis , and trigger apoptosis via the p38 / mitogen-activated protein kinase pathway ."

reach
"Invivo experiments showed that USP12-knockdown could suppress tumor growth in mice, and immuno blotting revealed that USP12 could induce G2/M arrest through the cyclin dependent kinase1 and cyclinB1 axis, and trigger apoptosis via the p38 and mitogen activated protein kinase pathway."
| 1

reach
"We further established that Usp12 depletion reduced PC cell proliferation and induced apoptosis, furthermore Usp12 levels are increased in PC tissue [XREF_BIBR]."
USP12 affects USP1
| 7
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
USP12 affects TGFB
| 7
USP12 activates TGFB. 7 / 7
| 7

reach
"The dauer larva-constitutive C. elegans phenotype caused by defective DAF-7 and TGF-beta signaling was enhanced and suppressed, respectively, by ubh-1 deletion and overexpression in the loss-of-function genetic backgrounds of daf7, daf-1 and TGF-betaRI, and daf4 and R-SMAD, but not of daf-8 and R-SMAD."

reach
"This suggested that UBH-1 may stimulate DAF-7 and TGF-beta signaling via DAF-8 and R-SMAD."

reach
"This suggested that UBH-1 may stimulate DAF-7 and TGF-beta signaling via DAF-8 and R-SMAD."

reach
"The dauer larva-constitutive C. elegans phenotype caused by defective DAF-7 and TGF-beta signaling was enhanced and suppressed, respectively, by ubh-1 deletion and overexpression in the loss-of-function genetic backgrounds of daf7, daf-1 and TGF-betaRI, and daf4 and R-SMAD, but not of daf-8 and R-SMAD."

reach
"The authors showed that UBH-1 in Caenorhabditis elegans and its human homolog UCHL1 promotes DAF-7/ TGF-β signalling, indicating that the mechanism of TGF-β signalling is conserved among animal species [67]."

reach
"Here, we show that the deubiquitinating enzyme UBH-1 in Caenorhabditis elegans and its human homolog, ubiquitin C-terminal hydrolase-L1 (UCH-L1), stimulate DAF-7 and TGF-beta signaling, suggesting that this mode of regulation of TGF-beta signaling is conserved across animal species."

reach
"Here, we show that the deubiquitinating enzyme UBH-1 in Caenorhabditis elegans and its human homolog, ubiquitin C-terminal hydrolase-L1 (UCH-L1), stimulate DAF-7 and TGF-beta signaling, suggesting that this mode of regulation of TGF-beta signaling is conserved across animal species."
USP12 affects AKT
| 7
USP12 inhibits AKT.
| 4
USP12 inhibits AKT. 4 / 4
| 4

reach
"WDR48 and USP12 Negatively Regulate Akt Signaling and Promote Apoptosis."

reach
"As knockdown of WDR48 and USP12 increased Akt activation, we next tested whether WDR48 and USP12 could potentiate the suppression of Akt signaling by PHLPP1."

reach
"Thus, these results suggest that WDR48 and USP12 negatively regulate Akt signaling and thereby induce apoptosis in conjunction with PHLPP1."

reach
"Because PHLPP1 is a potent negative regulator of the PI3-kinase and Akt pathway, we hypothesized that WDR48 and USP12 might suppress Akt signaling by stabilizing PHLPP1."
USP12 phosphorylates AKT.
| 2
USP12 leads to the phosphorylation of AKT. 2 / 2
| 2

reach
"Secondly, Usp12 deubiquitinates and stabilises PHLPP and PHLPPL proteins which enhances dephosphorylation and deactivation of Akt."

reach
"Knockdown of Usp12, or its complex members Uaf-1 and WDR20, increased Akt phosphorylation without any change to the total Akt protein in PC cells."
USP12 dephosphorylates AKT.
| 1
USP12 dephosphorylates AKT. 1 / 1
| 1

reach
"XREF_BIBR Notably, a recent study that screened a panel with 22 deubiquitinating enzymes in vitro showed that galeterone inhibits the enzymatic activity of USP12 and also binds directly to the USP12 and USP12 and UAF1 complexes, which dephosphorylate AKT."
USP1 affects USP46
| 7
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
DMWD affects USP12
3 | 2 2
DMWD binds USP12.
3 | 1 2
3 | 1

No evidence text available

reach
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."

No evidence text available

No evidence text available
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
DMWD activates USP12.
| 1
DMWD activates USP12. 1 / 1
| 1

reach
"Finally, we show that both DMWD and WDR20 promote USP12 enzymatic activity, but they differentially modulate the subcellular localization of the DUB."
USP12 affects Ubiquitin
| 1 5
USP12 binds Ubiquitin.
| 5
| 3

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( xref , xref )."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D, 6E, and S5F) ."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Interestingly, it has been shown that USP12-Ub and UAF1 have two possible binding sites: The Finger domain and the back of the ubiquitin-binding cleft [ xref ]."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 inhibits Ubiquitin.
| 1
| 1

reach
"Also, USP12 co-localizes with AR in the cytoplasm and promotes AR transcriptional activity by confronting the ubiquitin dependent degradation of AR [XREF_BIBR]."
USP12 affects TMPRSS11D
| 6
| 6

sparser
"The aforementioned results demonstrated that USP12 binds to the HAT domain of CBP and blocks CBP’s acetylation action."

sparser
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP’s acetyltransferase activity."

sparser
"Taken together, these findings suggest that USP12 binds to the HAT domain of CBP and blocks CBP’s acetyltransferase activity in cells."

sparser
"In vitro binding assay further demonstrated that USP12 binds with the HAT domain of CBP ( xref )."

sparser
"Interestingly, USP12 can interact with the HAT domain of CBP."

sparser
"In this study, we demonstrated that USP12 can bind with the HAT domain of CBP in vivo and in vitro ."
USP12 affects DMWD
3 | 1 2
3 | 1

No evidence text available

reach
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."

No evidence text available

No evidence text available
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
TMPRSS11D affects USP12
| 6
| 6

sparser
"The aforementioned results demonstrated that USP12 binds to the HAT domain of CBP and blocks CBP’s acetylation action."

sparser
"Here, we found that cellular USP12 interacts with the HAT domain of CBP and inhibits CBP’s acetyltransferase activity."

sparser
"Taken together, these findings suggest that USP12 binds to the HAT domain of CBP and blocks CBP’s acetyltransferase activity in cells."

sparser
"In vitro binding assay further demonstrated that USP12 binds with the HAT domain of CBP ( xref )."

sparser
"Interestingly, USP12 can interact with the HAT domain of CBP."

sparser
"In this study, we demonstrated that USP12 can bind with the HAT domain of CBP in vivo and in vitro ."
WDR48 affects USP1
| 5
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
Ubiquitin affects USP12
| 5
| 3

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( xref , xref )."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D, 6E, and S5F) ."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Interestingly, it has been shown that USP12-Ub and UAF1 have two possible binding sites: The Finger domain and the back of the ubiquitin-binding cleft [ xref ]."
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 affects daf-7
| 5
USP12 activates daf-7. 5 / 5
| 5

reach
"This suggested that UBH-1 may stimulate DAF-7 and TGF-beta signaling via DAF-8 and R-SMAD."

reach
"This suggested that UBH-1 may stimulate DAF-7 and TGF-beta signaling via DAF-8 and R-SMAD."

reach
"Here, we show that the deubiquitinating enzyme UBH-1 in Caenorhabditis elegans and its human homolog, ubiquitin C-terminal hydrolase-L1 (UCH-L1), stimulate DAF-7 and TGF-beta signaling, suggesting that this mode of regulation of TGF-beta signaling is conserved across animal species."

reach
"Here, we show that the deubiquitinating enzyme UBH-1 in Caenorhabditis elegans and its human homolog, ubiquitin C-terminal hydrolase-L1 (UCH-L1), stimulate DAF-7 and TGF-beta signaling, suggesting that this mode of regulation of TGF-beta signaling is conserved across animal species."

reach
"The authors showed that UBH-1 in Caenorhabditis elegans and its human homolog UCHL1 promotes DAF-7/ TGF-β signalling, indicating that the mechanism of TGF-β signalling is conserved among animal species [67]."
USP12 affects Histone_H2B
| 5
USP12 deubiquitinates Histone_H2B. 5 / 5
| 5

reach
"Both USP12 and USP46 deubiquitinate histone H2A and H2B, affecting Xenopus development (Joo et al., 2011), and, unlike USP1, these smaller USPs can simultaneously bind WDR20 in addition to WDR48 (Joo [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"It has also been reported that USP12 and USP46 deubiquitinate histone H2A and H2B thereby playing a role in Xenopus development."

reach
"USP12 and USP46 prefer nucleosomal substrates and deubiquitinate both histone H2A and H2B in vitro and in vivo."

reach
"Usp12 and Usp46 deubiquitinate both H2A and H2B and Uaf-1 is required for this reaction [XREF_BIBR]."

reach
"Joo et al. demonstrated that USP46 and USP12 containing fractions from HeLa cells can deubiquitinate histones H2A and H2B to control cell fate and gastrulation during Xenopus development [XREF_BIBR]."
USP12 affects Chemokine
| 5
USP12 activates Chemokine.
| 3
| 3

reach
"Given the relevance of the USP12-mediated chemokine, such as CXCL8, CXCL1 and CCL2, in tumour development and progression, we examined the mechanism underlying the USP12-mediated regulation."

reach
"Of note, the altered production of the chemokines caused by USP12 downregulation in tumour cells may also regulate T cell infiltration and function."

reach
"This study demonstrated that AKT-mTOR hyperactivation resulted in USP12 downregulation, which in turn increased the production of protumourigenic chemokine in NSCLC, indicating that USP12 is a regulatory node in oncogenic mutation-driven TME development.The tumour suppressive function conveyed by USP12 is virtually related to the control of protumourigenic chemokine expression."
USP12 increases the amount of Chemokine.
| 1
USP12 increases the amount of Chemokine. 1 / 1
| 1

reach
"Collectively, these data indicate that dysregulated chemokine expression caused by USP12 downregulation, as exemplified by CXCL1 and CCL2, is important for the development of immune-suppressive TME.To validate whether the role of USP12 on TME reprogramming has pathological relevance in human NSCLC, we investigated the relationship between TAMs and USP12 expression in human NSCLC cohorts."
USP12 decreases the amount of Chemokine.
| 1
USP12 decreases the amount of Chemokine. 1 / 1
| 1

reach
"Given that USP12 negatively regulated protumourigenic chemokine expression and exerted a tumour-suppressive effect in immune-competent mice but not in immune-deficient mice, it seems intuitive to speculate that USP12 downregulation in tumours may help establish a favourable microenvironment for tumour growth."
USP1 affects WDR48
| 5
| 5

sparser
"For example, WDR48 interacts with USP1, USP12, and USP46 to activate their DUB activity ( Cohn et al., 2009; Yin et al., 2015 )."

sparser
"HPV E1 helicase, which possesses DNA unwinding activity and enhances viral DNA replication, could form a complex with UAF1 and any one of USP1, USP12 and USP46."

sparser
"For example, WDR48 interacts with USP1, USP12 and USP46 to activate their DUB activity ( xref ; xref )."

sparser
"UAF1 constitutively binds to USP1, USP12, and USP46, and this binding greatly enhances their deubiquitinase activity."

sparser
"For example, USP1, USP12 and USP46 can interact with WD40-repeat protein USP1-associated factor 1 (UAF1), and UAF1 stimulates the catalytic activity of human USP1 through the formation of a tight complex [ xref ]."
WDR20 affects USP46
| 4
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
USP12 affects Notch
| 4
USP12 inhibits Notch.
| 2
USP12 inhibits Notch. 2 / 2
| 2

reach
"CG7023 and USP12 was shown to negatively regulate Notch signaling in flies."

reach
"CG7023/USP12 was shown to negatively regulate Notch signaling in flies (Moretti et al., 2012)."
USP12 deubiquitinates Notch.
| 1
USP12 deubiquitinates Notch. 1 / 1
| 1

reach
"At the biochemical level, USP12 with its activator UAF1 deubiquitinate the nonactivated form of Notch in cell culture and in vitro."
USP12 activates Notch.
| 1
USP12 activates Notch. 1 / 1
| 1

reach
"The Ubiquitin specific Protease 12 (USP12) is a negative regulator of Notch signaling as USP12 directly targets Notch and directs it to lysosomal degradation."
USP12 affects METTL3
| 1 3
USP12 increases the amount of METTL3.
| 3
USP12 increases the amount of METTL3. 2 / 2
| 2

reach
"Furthermore, we discover that USP12 promote the expression of METTL3 via upregulating p300."

reach
"Finally, our data show that USP12 is partially dependent on the stabilization of p300 to activate METTL3 expression and promote myocardial hypertrophy."
USP12 bound to EP300 increases the amount of METTL3. 1 / 1
| 1

reach
"Mechanistically, USP12 binds and stabilizes p300, thereby activating the transcription of its downstream gene METTL3."
USP12 activates METTL3.
| 1
USP12 activates METTL3. 1 / 1
| 1

eidos
"Furthermore , we discover that USP12 promote the expression of METTL3 via upregulating p300 ."
USP12 affects EP300
| 1 2 1
USP12 deubiquitinates EP300.
| 1 1
USP12 deubiquitinates EP300. 2 / 2
| 1 1

reach
"De-ubiquitination of p300 by USP12 Critically Enhances METTL3 Expression and Ang II induced Cardiac Hypertrophy."

trips
"De-ubiquitination of p300 by USP12 Critically Enhances METTL3 Expression and Ang II-induced Cardiac Hypertrophy."
USP12 binds EP300.
| 1 1
| 1 1

reach
"Mechanistically, USP12 binds and stabilizes p300, thereby activating the transcription of its downstream gene METTL3."

sparser
"Mechanistically, USP12 binds and stabilizes p300, thereby activating the transcription of its downstream gene METTL3."
3 |
Valproic acid decreases the amount of USP12.
2 |
Valproic acid decreases the amount of USP12. 2 / 2
2 |

No evidence text available

No evidence text available
Valproic acid methylates USP12.
1 |
1 |

No evidence text available
RplA affects Ubiquitin
| 3
| 3

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( xref , xref )."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D, 6E, and S5F) ."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
WDR48 affects F222
| 3
USP12 binds WDR48 and F222. 3 / 3
| 3

reach
"Although the active site of USP12 in the UAF1-USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme ( Figure S3F ), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme, in the C2 space group structure, the USP12 Thumb [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme (XREF_SUPPLEMENTARY), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."
WDR48 affects F219
| 3
USP12 binds WDR48 and F219. 3 / 3
| 3

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta-strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in its pocket formed between the Fingers and Palm domains, instead of being tucked underneath the Fingers domain (XREF_FIG, XREF_FIG, XREF_FIG)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in the pocke[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
WDR proteins affects USP12
| 3
USP12 binds WDR proteins. 3 / 3
| 3

reach
"Biochemical and proteomic studies showed that two WDR proteins, WDR48 (also known as USP1 associated factor, UAF1) and WDR20, interact with USP46 and USP12."

reach
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis."

reach
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis.These recent structural and biochemical studies complement in vivo studies of the USP-46/WDR-48/WDR-20 complex in C. elegans."
Ubiquitin affects rplA
| 3
| 3

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( xref , xref )."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D, 6E, and S5F) ."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP12 affects cell cycle
| 2 1
USP12 activates cell cycle.
| 1 1
| 1 1

reach
"Flow cytometry analysis also showed that USP12 could induce cell cycle arrest at the G2/Mstage."

eidos
"In addition , USP12 promotes cell survival , proliferation , tumorigenesis , and cell cycle progression by upregulating BMI-1 , c-Myc , and cyclin D2 [ 24,25,26,27 ] ."
USP12 inhibits cell cycle.
| 1
| 1

eidos
"Flow cytometry analysis also showed that USP12 could induce cell cycle arrest at the G2 / M stage ."
USP12 affects WDR proteins
| 3
USP12 binds WDR proteins. 3 / 3
| 3

reach
"Biochemical and proteomic studies showed that two WDR proteins, WDR48 (also known as USP1 associated factor, UAF1) and WDR20, interact with USP46 and USP12."

reach
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis."

reach
"Together, these structural studies suggest that binding of the WDR proteins to USP12 or USP46 relatively far from the active site results in the rearrangement of several structural elements, which propagates to the catalytic triad increasing enzyme catalysis.These recent structural and biochemical studies complement in vivo studies of the USP-46/WDR-48/WDR-20 complex in C. elegans."
USP12 affects Ubiquitin, and rplA
| 3
| 3

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( xref , xref )."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D, 6E, and S5F) ."

sparser
"Finally, consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms, mutation of a Ub globular domain-binding residue on BL1, Y264, compromised the ability of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP12 affects SQSTM1
| 1 2
| 1 1

sparser
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation (Fig.  xref )."

reach
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation."
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
USP12 affects PC
| 3
USP12 activates PC.
| 2
USP12 activates PC. 2 / 2
| 2

reach
"As a result Usp12 promotes PC development with protein levels increased in PC patients compared to benign controls."

reach
"Furthermore, Usp12 silencing was shown to reduce proliferation of PC cells."
USP12 inhibits PC.
| 1
USP12 inhibits PC. 1 / 1
| 1

reach
"As a result, depleting Usp12 decreased PC cellular proliferation and increased cellular apoptosis suggesting it may be a potential target for CRPC therapy [XREF_BIBR]."
USP12 affects OPTN
| 1 2
| 1 1

reach
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn."

sparser
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn (Fig.  xref )."
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
USP12 affects NFKBIA
| 2 1
USP12 dephosphorylates NFKBIA. 3 / 3
| 2 1

reach
"Mechanistic analysis demonstrates that USP12 dephosphorylates IkappaBalpha, which is a well characterized inhibitor of NF-kappaB nuclear translocation, inducing proinflammatory responses in macrophages."

sparser
"Mechanistic analysis demonstrates that USP12 dephosphorylates IκBα, which is a well-characterized inhibitor of NF-κB nuclear translocation, inducing proinflammatory responses in macrophages."

reach
"Mechanistic analysis suggest that USP12 may be required for the activation of NFkappaB pathway as knockdown of USP12 reduced the inhibitory phosphorylation of IkappaBalpha, a well characterized inhibitor of NFkappaB nuclear translocation."
USP12 affects MDK
| 1 2
USP12 binds MDK.
| 2
| 2

sparser
"Therefore, the USP12-MDK axis could serve as a potential target for the therapeutic treatment of breast cancer metastasis."

sparser
"USP12 could directly interact with MDK, decrease its polyubiquitination and increase its protein stability in cells."
USP12 deubiquitinates MDK.
| 1
USP12 deubiquitinates MDK. 1 / 1
| 1

reach
"Collectively, our study identified that USP12 is responsible for deubiquitinating and stabilizing MDK and leads to metastasis by promoting angiogenesis."
USP12 affects MAP1LC3
| 3
USP12 inhibits MAP1LC3.
| 2
| 2

reach
"Conversely, Usp12 knockdown increased the mean half-life of LC3, while also significantly right shifting the distribution of LC3 half-lives in individual neurons."

reach
"Usp12 and Usp12-C48S overexpression significantly decreased the mean half-life of LC3, but did not affect the half-life of the control protein Dendra2."
USP12 activates MAP1LC3.
| 1
Mutated USP12 activates MAP1LC3. 1 / 1
| 1

reach
"Since a non catalytic mutant of Usp12, Usp12-C48S, stimulated LC3 clearance as efficiently as Usp12, we additionally conclude that this function of Usp12 is independent of its deubiquitinating activity."
USP12 affects ITCH
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
USP12 affects HMGB1
| 1 2
USP12 activates HMGB1.
| 1 1
USP12 activates HMGB1. 2 / 2
| 1 1

reach
"Deubiquitylase USP12 induces pro-survival autophagy and bortezomib resistance in multiple myeloma by stabilizing HMGB1."

eidos
"Knockdown of USP12 decreased the level of HMGB1 and suppressed HMGB1-mediated autophagy in MM ."
USP12 increases the amount of HMGB1.
| 1
USP12 increases the amount of HMGB1. 1 / 1
| 1

reach
"Knockdown of USP12 decreased the level of HMGB1 and suppressed HMGB1-mediated autophagy in MM."
USP12 affects FH
| 3
| 3

sparser
"We further found that FH-USP12 can interact with HA-p300 in cells ( xref )."

sparser
"Flag-HA tagged USP12 (FH-USP12) can bind with HA-CBP ( xref ) or endogenous CBP ( xref )."

sparser
"The results showed that CBP induced Flag-STAT1 acetylation in vitro , which was gradually inhibited by increasing amounts of FH-USP12 ( xref ), suggesting that USP12 blocks the acetylation effect of CBP on STAT1."
USP12 affects F222
| 3
USP12 binds WDR48 and F222. 3 / 3
| 3

reach
"Although the active site of USP12 in the UAF1-USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme ( Figure S3F ), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme, in the C2 space group structure, the USP12 Thumb [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme (XREF_SUPPLEMENTARY), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."
USP12 affects F219
| 3
USP12 binds WDR48 and F219. 3 / 3
| 3

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta-strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in its pocket formed between the Fingers and Palm domains, instead of being tucked underneath the Fingers domain (XREF_FIG, XREF_FIG, XREF_FIG)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in the pocke[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"USP12 promotes nonsmall cell lung cancer progression through deubiquitinating and stabilizing RRM2."

eidos
"USP12 promotes nonsmall cell lung cancer progression through deubiquitinating and stabilizing RRM2 ."

reach
"Here, we show that the deubiquitinase USP12 is commonly downregulated in the Kras G12D -driven mouse lung tumour and human non-small cell lung cancer owing to the activation of AKT-mTOR signalling."
SQSTM1 affects USP12
| 1 2
| 1 1

sparser
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation (Fig.  xref )."

reach
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation."
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
OPTN affects USP12
| 1 2
| 1 1

reach
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn."

sparser
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn (Fig.  xref )."
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
USP12 is modified
| 1 1 1
USP12 is phosphorylated.
| 1
USP12 is phosphorylated. 1 / 1
| 1

sparser
"09; p ‐value USP12  = ."
USP12 is produced.
| 1
USP12 is produced. 1 / 1
| 1

reach
"Further analysis showed that LPS reduced the levels of Sp1 which enhanced the transcriptional levels of USP12."
USP12 is deubiquitinated.
| 1
USP12 is deubiquitinated. 1 / 1
| 1

trips
"Deubiquitinating enzyme Usp12 is a novel co-activator of the androgen receptor."
ITCH affects USP12
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
FH affects USP12
| 3
| 3

sparser
"We further found that FH-USP12 can interact with HA-p300 in cells ( xref )."

sparser
"Flag-HA tagged USP12 (FH-USP12) can bind with HA-CBP ( xref ) or endogenous CBP ( xref )."

sparser
"The results showed that CBP induced Flag-STAT1 acetylation in vitro , which was gradually inhibited by increasing amounts of FH-USP12 ( xref ), suggesting that USP12 blocks the acetylation effect of CBP on STAT1."
F222 affects WDR48
| 3
USP12 binds WDR48 and F222. 3 / 3
| 3

reach
"Although the active site of USP12 in the UAF1-USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme ( Figure S3F ), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme, in the C2 space group structure, the USP12 Thumb [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although the active site of USP12 in the UAF1 and USP12 complex crystallized in F222 space group adopts the same topology as found in the free enzyme (XREF_SUPPLEMENTARY), in the C2 space group structure, the USP12 Thumb domain features a series of major structural rearrangements at the catalytic cleft."
F219 affects WDR48
| 3
USP12 binds WDR48 and F219. 3 / 3
| 3

reach
"While the Pinky Finger of USP12 in the UAF1-USP12-WDR20 complex adopts the same b strand conformation as seen in the UAF1-USP12 complex, F219 on the PK Helix becomes embedded in the pocket formed between the Fingers and the Palm domains, instead of being tucked underneath the Fingers domain ( Figures 3D, 5B , and 5C)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta-strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in its pocket formed between the Fingers and Palm domains, instead of being tucked underneath the Fingers domain (XREF_FIG, XREF_FIG, XREF_FIG)."

reach
"While the Pinky Finger of USP12 in the UAF1, USP12, and WDR20 complex adopts the same beta strand conformation as seen in the UAF1 and USP12 complex, F219 on the PK Helix becomes embedded in the pocke[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
2 |
Pirinixic acid decreases the amount of USP12. 2 / 2
2 |

No evidence text available

No evidence text available
Bisphenol A affects USP12
2 |
Bisphenol A methylates USP12.
1 |
1 |

No evidence text available
Bisphenol A decreases the amount of USP12.
1 |
Bisphenol A decreases the amount of USP12. 1 / 1
1 |

No evidence text available
WDR48 affects USPs
| 2
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
WDR48 affects PHLPP1
| 2
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
2 |
2 |

No evidence text available

No evidence text available
VSX2 affects USP12
2 |
2 |

No evidence text available

No evidence text available
Usp12-C48S affects USP12
| 2
USP12 binds Usp12-C48S. 2 / 2
| 2

reach
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn."

reach
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation."
USPs affects WDR48
| 2
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
USP12 affects mHTT-N586
| 2
USP12 inhibits mHTT-N586. 2 / 2
| 2

reach
"Usp12 knockdown with Usp12-siRNA r did not affect HTT-N586 Q17 survival but increased mHTT-N586 Q138 toxicity."

reach
"Usp12 nor Usp12-C48S suppressed mHTT-N586 Q138 neurotoxicity during Atg7 knockdown."
USP12 affects mHTT
| 2
USP12 inhibits mHTT.
| 1
USP12 inhibits mHTT. 1 / 1
| 1

reach
"Catalytically inactive Usp12 suppresses mHTT toxicity."
USP12 binds mHTT.
| 1
USP12 binds mHTT. 1 / 1
| 1

reach
"Using the available association data for Usp12 and mHTT interactions, we mapped a protein protein interaction network between Usp12 and mHTT, and identified the autophagy receptor Optineurin (Optn) as a potential link."

reach
"Deubiquitinase USP12 promotes LPS induced macrophage responses through inhibition of IkappaBalpha."

reach
"Most of the deubiquitinases, including A20 and CYLD, can inhibit NF-kappaB activation [XREF_BIBR], while USP12 positively regulates the LPS signal [XREF_BIBR]."
USP12 affects cell growth
| 1 1
| 1 1

eidos
"Depletion of USP12 suppressed cell growth and clonogenicity and inhibited autophagy ."

reach
"Depletion of USP12 suppressed cell growth and clonogenicity and inhibited autophagy."
| 2
USP12 inhibits angiogenesis.
| 1
| 1

reach
"Overexpression of MDK rescued the loss of angiogenesis ability mediated by knockdown of USP12 in breast cancer cells in vitro and in vivo."
USP12 activates angiogenesis.
| 1
| 1

reach
"USP12 promotes breast cancer angiogenesis by maintaining midkine stability."
USP12 affects VSX2
2 |
2 |

No evidence text available

No evidence text available
USP12 affects Usp12-C48S
| 2
USP12 binds Usp12-C48S. 2 / 2
| 2

reach
"Co-immunoprecipitation confirmed an interaction between Usp12 or Usp12-C48S and Optn."

reach
"An interaction between Usp12 or Usp12-C48S and p62 was also confirmed by co-immunoprecipitation."
USP12 affects USPs
| 2
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
USP12 affects USP46, WDR20, and WDR48
| 2
| 2

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., 2009; Kee et al., 2010) and further enhances their catalytic activity in vitro (Kee et al., 2010; Faesen et al., 2011)."

sparser
"WDR20 forms a unique ternary complex with WDR48 and either USP12 or USP46 (Sowa et al., xref ; Kee et al., xref ) and further enhances their catalytic activity in vitro (Kee et al., xref ; Faesen et al., xref )."
USP12 affects USP46, and WDR20
| 2
| 2

sparser
"Further analysis showed that WDR20 associates exclusively with USP12 and USP46, not with USP1."

sparser
"WDR20 binding to USP12 and USP46 showed a significant increase in the catalytic activity in vitro [ xref , xref ]."
USP12 affects TCR
| 2
USP12 deubiquitinates TCR.
| 1
USP12 deubiquitinates TCR. 1 / 1
| 1

reach
"Similarly, the ubiquitin specific peptidase 12 (USP12) can deubiquitinate LAT at the proximal TCR (Figure 2), to protect it from ubiquitin-dependent lysosomal degradation [146]."
| PMC
USP12 activates TCR.
| 1
USP12 activates TCR. 1 / 1
| 1

reach
"In contrast, USP12 has been found to stabilize the TCR complex and promote TCR signaling through deubiquitylating TCR adaptor proteins LAT and Trat1 in primary mouse T lymphocytes."
USP12 affects STAT1
| 2
USP12 inhibits STAT1.
| 1
USP12 inhibits STAT1. 1 / 1
| 1

reach
"Other studies have shown that USP12 suppresses the PH domain and leucine rich repeat protein phosphatases, which restrains the activation of macrophages through the dephosphorylation of transcription factor STAT1 [XREF_BIBR]."
USP12 deacetylates STAT1.
| 1
USP12 leads to the deacetylation of STAT1. 1 / 1
| 1

reach
"Upon IFN treatment, USP12 accumulates in the nucleus, blocks the CREB-binding protein-induced acetylation of p-STAT1, and thus inhibits the dephosphorylation effects of TCPTP on p-STAT1, which ultimately maintains the nuclear p-STAT1 levels and IFN antiviral efficacy (114)."
USP12 affects PK
| 2
| 2

reach
"(C and D) A close-up comparison of the Pinky Finger, PK Helix, and PK Loop between free and UAF1-bound USP12 structures, with the two free USP12 structures aligned in (C)."

reach
"The dashed line indicates the disordered PK Loop of UAF1-bound USP12."
| 1 1

reach
"Knockdown of USP12 decreased the lung metastasis ability of 4T1 cells, while USP12 overexpression increased the lung metastasis ability of these cells in vivo."

eidos
"Collectively , our study identified that USP12 is responsible for deubiquitinating and stabilizing MDK and leads to metastasis by promoting angiogenesis ."

reach
"In the present study, we reported that USP12 deficiency decreased infiltration and impaired the suppressor function of monocytic (M)-MDSCs, resulting in increased CD8 + T cell response and decelerated tumor growth."

eidos
"In the present study , we reported that USP12 deficiency decreased infiltration and impaired the suppressor function of monocytic ( M ) - MDSCs , resulting in increased CD8 + T cell response and decelerated tumor growth ."
USP12 affects MYC
| 1 1
USP12 increases the amount of MYC.
| 1
USP12 increases the amount of MYC. 1 / 1
| 1

reach
"In this study, we showed that knockdown of USP12 effectively induced cell cycle arrest in HeLa cells and decreased BMI-1, c-Myc and cyclin D2 transcription levels."
USP12 binds MYC.
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
USP12 affects MDM2
1 | 1
USP12 deubiquitinates MDM2. 2 / 2
1 | 1

reach
"We find that USP12 deubiquitinates MDM2 and AR, which in turn controls the levels of the TP53 tumour suppressor and AR oncogene in prostate cancer."

"We find that USP12 deubiquitinates MDM2 and AR, which in turn controls the levels of the TP53 tumour suppressor and AR oncogene in prostate cancer."
USP12 affects LAT
1 | 1
USP12 deubiquitinates LAT.
| 1
USP12 deubiquitinates LAT. 1 / 1
| 1

reach
"Similarly, the ubiquitin specific peptidase 12 (USP12) can deubiquitinate LAT at the proximal TCR (Figure 2), to protect it from ubiquitin-dependent lysosomal degradation [146]."
| PMC
USP12 binds LAT.
1 |
1 |

No evidence text available
USP12 affects Interferon
| 2
| 2

reach
"Interestingly, USP12 positively regulates interferon (IFN) antiviral signaling independently of its deubiquitinase activity."

reach
"Additionally, USP12 positively regulates IFN antiviral signaling independently of its deubiquitinase activity."
USP12 affects IFI16
| 2
USP12 inhibits IFI16.
| 1
USP12 inhibits IFI16. 1 / 1
| 1

reach
"Mechanistically, USP12 inhibited the proteasome-dependent degradation of IFI16 through its deubiquitinase activity, thereby maintaining IFI16 stability and promoting IFI16-STING-IRF3- and p65-mediated antiviral signaling."
USP12 deubiquitinates IFI16.
| 1
USP12 deubiquitinates IFI16. 1 / 1
| 1

reach
"USP12 promotes antiviral responses by deubiquitinating and stabilizing IFI16."
USP12 affects H2BC21
1 1 |
USP12 deubiquitinates H2BC21.
1 |
USP12 deubiquitinates H2BC21. 1 / 1
1 |

"USP12 and USP46 prefer nucleosomal substrates and deubiquitinate both histone H2A and H2B in vitro and in vivo."
USP12 binds H2BC21.
1 |
1 |

No evidence text available
USP12 affects H2AC20
1 1 |
USP12 deubiquitinates H2AC20.
1 |
USP12 deubiquitinates H2AC20. 1 / 1
1 |

"USP12 and USP46 prefer nucleosomal substrates and deubiquitinate both histone H2A and H2B in vitro and in vivo."
USP12 binds H2AC20.
1 |
1 |

No evidence text available
USP12 affects GRIA1
| 2
USP12 decreases the amount of GRIA1. 2 / 2
| 2

reach
"Knock-down of USP46, but not USP12, results in increased levels of ubiquitinated GluA1, decreased surface and total levels of GluA1, and reduced mEPSC amplitudes, consistent with a role for USP46 in deubiquitinating mammalian AMPARs."

reach
"Knock-down of USP46, but not USP12, results in increased levels of ubiquitinated GluA1, decreased surface and total levels of GluA1, and reduced mEPSC amplitudes, consistent with a role for USP46 in deubiquitinating mammalian AMPARs (Huo et al., 2015)."
| 1 1
USP12 inhibits Cell Survival.
| 1

reach
"However, USP12 has also been shown to inhibit AKT-dependent cell survival by stabilizing the pro-apoptotic phosphatases PHLPP1 and/or PHLPPL ."
USP12 activates Cell Survival.
| 1
| 1

eidos
"In addition , USP12 promotes cell survival , proliferation , tumorigenesis , and cell cycle progression by upregulating BMI-1 , c-Myc , and cyclin D2 [ 24,25,26,27 ] ."
USP12 affects BAX
| 1 1
| 1 1

sparser
"The binding of USP12 to Bax shows the interaction as a DUB, which regulates ubiquitination on Bax."

reach
"The binding of USP12 to Bax shows the interaction as a DUB, which regulates ubiquitination on Bax."
USP12 affects Ang II-induced cardiac hypertrophy
| 2
USP12 activates Ang II-induced cardiac hypertrophy. 2 / 2
| 2

eidos
"Moreover , we demonstrate that USP12 aggravate Ang II-induced cardiac hypertrophy by enhancing METTL3 , a methyltransferase which catalyze N 6 - methyladenosine ( m 6 A ) modification on messenger RNA and acts as a harmful factor in pathological cardiac hypertrophy ."

eidos
"USP12 deficiency also significantly attenuate the phenotype of Ang II-induced cardiac hypertrophy in vivo ."
USP1 affects USP12, and USP46
| 2
USP1 binds USP46 and USP12. 2 / 2
| 2

sparser
"UAF1 is a WD40 repeat protein, which binds and activates three important DUBs, USP1, USP12 and USP46."

sparser
"Although the exact function of p80/Uaf1 in viral DNA replication remains unclear, this WD40 repeats-containing protein is known to interact with the de-ubiquitinating enzymes Usp1, Usp12 and Usp46, whose substrates include PCNA, histones H2A and H2B and Fanconi anaemia complementation group D type 2, a component of the Fanconi anaemia pathway [ xref , xref ]."
RFX1 affects USP12
2 |
RFX1 decreases the amount of USP12. 2 / 2
2 |

No evidence text available

No evidence text available
PK affects USP12
| 2
| 2

reach
"(C and D) A close-up comparison of the Pinky Finger, PK Helix, and PK Loop between free and UAF1-bound USP12 structures, with the two free USP12 structures aligned in (C)."

reach
"The dashed line indicates the disordered PK Loop of UAF1-bound USP12."
PHLPP1 affects WDR48
| 2
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
MDK affects USP12
| 2
| 2

sparser
"Therefore, the USP12-MDK axis could serve as a potential target for the therapeutic treatment of breast cancer metastasis."

sparser
"USP12 could directly interact with MDK, decrease its polyubiquitination and increase its protein stability in cells."
EP300 affects USP12
| 1 1
| 1 1

reach
"Mechanistically, USP12 binds and stabilizes p300, thereby activating the transcription of its downstream gene METTL3."

sparser
"Mechanistically, USP12 binds and stabilizes p300, thereby activating the transcription of its downstream gene METTL3."
CXCL1 affects USP12
| 2
CXCL1 inhibits USP12.
| 1
CXCL1 inhibits USP12. 1 / 1
| 1

reach
"Moreover, coexpression of Cxcl1 and Ccl2 efficiently attenuated activation of CD8 and CD4 T cells in tumour cells ectopically expressing USP12, as evidenced by the changes in the frequencies of TNF-α IFN-γ cells in the populations (Fig. 5g)."
CXCL1 increases the amount of USP12.
| 1
CXCL1 increases the amount of USP12. 1 / 1
| 1

reach
"Similarly, co-expression of Cxcl1 and Ccl2 in 889-S1 tumour cells substantially increased tumour incidence and growth in the presence of USP12 overexpression (Fig. 3g), suggesting the importance of the tumour-extrinsic effects controlled by USP12."
CCL2 affects USP12
| 2
CCL2 inhibits USP12.
| 1
CCL2 inhibits USP12. 1 / 1
| 1

reach
"Moreover, coexpression of Cxcl1 and Ccl2 efficiently attenuated activation of CD8 and CD4 T cells in tumour cells ectopically expressing USP12, as evidenced by the changes in the frequencies of TNF-α IFN-γ cells in the populations (Fig. 5g)."
CCL2 increases the amount of USP12.
| 1
CCL2 increases the amount of USP12. 1 / 1
| 1

reach
"Similarly, co-expression of Cxcl1 and Ccl2 in 889-S1 tumour cells substantially increased tumour incidence and growth in the presence of USP12 overexpression (Fig. 3g), suggesting the importance of the tumour-extrinsic effects controlled by USP12."
BAX affects USP12
| 1 1
| 1 1

sparser
"The binding of USP12 to Bax shows the interaction as a DUB, which regulates ubiquitination on Bax."

reach
"The binding of USP12 to Bax shows the interaction as a DUB, which regulates ubiquitination on Bax."
AKT affects USP12
| 2
AKT inhibits USP12.
| 1
AKT inhibits USP12. 1 / 1
| 1

reach
"Given that USP12 is negatively regulated by AKT signalling (Fig. 1i–m), we then tested whether restoring the expression of USP12 could counterbalance the activity of AKT in the regulation of chemokine expression."
AKT decreases the amount of USP12.
| 1
AKT decreases the amount of USP12. 1 / 1
| 1

reach
"Consistently, ectopic expression of the myristoylated form of AKT1 (myr-AKT1) significantly repressed USP12 expression in both human and mouse tumour cells (Fig. 1k); knockdown of endogenous AKT1 or AKT2, the two major isoforms of AKT, induced higher levels of USP12 than did the control treatment (Fig. 1l)."
Β-propeller proteins affects USP12
| 1
Β-propeller proteins activates USP12. 1 / 1
| 1

reach
"Without increasing substrate affinity of USP12, the two β-propeller proteins potentiate the enzyme through different allosteric mechanisms."
Vorinostat affects USP12
1 |
Vorinostat decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Vincristine affects USP12
1 |
Vincristine increases the amount of USP12. 1 / 1
1 |

No evidence text available
Urethane affects USP12
1 |
Urethane increases the amount of USP12. 1 / 1
1 |

No evidence text available

No evidence text available
Transmitting long-range allosteric signal BL1 affects USP12
| 1
Transmitting long-range allosteric signal BL1 activates USP12. 1 / 1
| 1

eidos
"UAF1 activates USP12 by binding to the tip of its Fingers domain and transmitting a long-range allosteric signal to BL1 ."
1 |
Titanium dioxide increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Testosterone decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Sunitinib affects USP12
1 |
Sunitinib increases the amount of USP12. 1 / 1
1 |

No evidence text available
Substrate affinity affects USP12
| 1
Substrate affinity activates USP12. 1 / 1
| 1

eidos
"First , our results rule out the possibility that UAF1 and WDR20 activate USP12 by enhancing its substrate affinity ."
Sirolimus affects USP12
| 1
Sirolimus decreases the amount of USP12. 1 / 1
| 1

reach
"Treatment with inhibitors targeting MEK (U0126), ERK (ERK inhibitor), JNK (JNK inhibitor II), JAK-STAT3 (WP1066), c-Myc (c-Myc inhibitor), p38 MAPK (SB203580), and TGF-β-SMAD (SB431542 and SIS3) failed to produce a substantial change in USP12 expression (Supplementary Fig. 1e); however, blocking AKT-mTOR signalling using the selective AKT inhibitor API-2 or the mTOR inhibitor rapamycin significantly increased USP12 expression (Fig. 1i, j)."
1 |
Silicon dioxide decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Rotenone affects USP12
1 |
Rotenone decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Potassium chromate decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Pirimiphos-methyl decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |

No evidence text available
PcDNA3.1 affects USP12
| 1
USP12 binds pcDNA3.1. 1 / 1
| 1

reach
"Human USP1, USP12, and UAF1 cDNAs were ligated to the pcDNA3.1 vector (Clontech Laboratories) modified with N-terminal Myc, hemagglutinin (HA), and FLAG peptides, respectively."
Pan affects USP12
| 1
Pan acetylates USP12. 1 / 1
| 1

sparser
"Intriguingly, we found that USP12 significantly inhibited CBP-mediated pan-acetylation effects in cells."
Oxycodone affects USP12
1 |
Oxycodone decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Optimizing catalytic cleft affects USP12
| 1
Optimizing catalytic cleft activates USP12. 1 / 1
| 1

eidos
"These results strongly suggest that WDR20 can activate USP12 by allosterically optimizing its catalytic cleft , whereas the effect of UAF1 binding cannot be directly transmitted to the catalytic cysteine through USP12 ."
Mutation Ub globular domain-binding residue BL1 affects USP12
| 1
Mutation Ub globular domain-binding residue BL1 activates USP12. 1 / 1
| 1

eidos
"Finally , consistent with the expected substrate-induced interactions between Ub and BL1 of USP12 in all forms , mutation of a Ub globular domain-binding residue on BL1 , Y264 , compromised the ability of both UAF1 and WDR20 to activate USP12 for Ub-AMC hydrolysis ( Figures 6D , 6E , and S5F ) ."
Medroxyprogesterone acetate increases the amount of USP12. 1 / 1
1 |

No evidence text available
| 1
MTOR inhibitor decreases the amount of USP12. 1 / 1
| 1

reach
"Treatment with inhibitors targeting MEK (U0126), ERK (ERK inhibitor), JNK (JNK inhibitor II), JAK-STAT3 (WP1066), c-Myc (c-Myc inhibitor), p38 MAPK (SB203580), and TGF-β-SMAD (SB431542 and SIS3) failed to produce a substantial change in USP12 expression (Supplementary Fig. 1e); however, blocking AKT-mTOR signalling using the selective AKT inhibitor API-2 or the mTOR inhibitor rapamycin significantly increased USP12 expression (Fig. 1i, j)."
MHTT affects USP12
| 1
USP12 binds mHTT. 1 / 1
| 1

reach
"Using the available association data for Usp12 and mHTT interactions, we mapped a protein protein interaction network between Usp12 and mHTT, and identified the autophagy receptor Optineurin (Optn) as a potential link."
Lipopolysaccharide decreases the amount of USP12. 1 / 1
| 1

reach
"Further analysis showed that LPS reduced the levels of Sp1 which enhanced the transcriptional levels of USP12."
Leflunomide affects USP12
1 |
Leflunomide increases the amount of USP12. 1 / 1
1 |

No evidence text available
Lead atom affects USP12
1 |
Lead atom increases the amount of USP12. 1 / 1
1 |

No evidence text available
Jinfukang affects USP12
1 |
Jinfukang decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Ionomycin affects USP12
1 |
Ionomycin increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hypochlorous acid increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-9500 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-92a-2-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-8083 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-7515 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-744-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-7109-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6887-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6885-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6876-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6795-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6780b-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-625-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-5196-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4747-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4725-3p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4533 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4476 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4472 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4447 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4271 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-373-3p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-328-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3202 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-215-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-192-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-1260b decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-1252-5p decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Geldanamycin increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Formaldehyde increases the amount of USP12. 1 / 1
1 |

No evidence text available
Doxorubicin affects USP12
1 |
Doxorubicin decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Dicrotophos affects USP12
1 |
Dicrotophos decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Cre affects USP12
| 1
USP12 binds cre. 1 / 1
| 1

sparser
"However, after exclusion of samples in which CNVs overlay the gene itself, amplification of the CRE was not associated with differential expression of USP12 or any other gene in the proximity of the CNV segments ( xref , xref )."
Clofibrate affects USP12
1 |
Clofibrate increases the amount of USP12. 1 / 1
1 |

No evidence text available
Cisplatin affects USP12
1 |
Cisplatin decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Bun107 affects USP12
| 1
Bun107 activates USP12. 1 / 1
| 1

reach
"Moreover, the human ortholog of Bun107 activates USP12 and USP46 XREF_BIBR."
Bucladesine affects USP12
1 |
Bucladesine increases the amount of USP12. 1 / 1
1 |

No evidence text available
Binding tip Fingers domain affects USP12
| 1
Binding tip Fingers domain activates USP12. 1 / 1
| 1

eidos
"UAF1 activates USP12 by binding to the tip of its Fingers domain and transmitting a long-range allosteric signal to BL1 ."
Avobenzone affects USP12
1 |
Avobenzone increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Aluminium atom increases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Aflatoxin M1 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
1 |
Aflatoxin B1 demethylates USP12. 1 / 1
1 |

No evidence text available
Abrine affects USP12
1 |
Abrine increases the amount of USP12. 1 / 1
1 |

No evidence text available
ZAP70 affects USP12
1 |
1 |

No evidence text available
WDR77 affects USP12
1 |
1 |

No evidence text available
WDR48 affects Ubiquitin
| 1
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
WDR26 affects USP12
1 |
1 |

No evidence text available
WDR20 affects Ubiquitin
| 1
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
WDR20 affects PHLPP2
| 1
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
WDR20 affects DMWD
| 1
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
WD40-repeat-containing protein affects USP12
| 1
WD40-repeat-containing protein activates USP12. 1 / 1
| 1

eidos
"USP12 and USP46 , but not USP1 , can be independently activated by another WD40-repeat-containing protein , WDR20 ( Burska et al ., 2013 ; Dahlberg and Juo , 2014 ; Kee et al ., 2010 ; McClurg et al ., 2014 ) ."
WD40 affects USP12
| 1
WD40 activates USP12. 1 / 1
| 1

reach
"For example, GMP synthetase (GMPS) interacts and activates USP7 (Faesen et al., 2011; Sarkari et al., 2009; van der Knaap et al., 2005), whereas the WD40 repeat containing UAF1 (WDR48) activates USP1,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
VSX1 affects USP12
1 |
1 |

No evidence text available
Ubiquitin affects WDR48
| 1
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."

eidos
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases ( USPs ) constitute the largest family of deubiquitinating enzymes , whose catalytic competency is often modulated by their binding partners through unknown mechanisms ."
USPs affects USP46
| 1
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USPs affects PHLPP1
| 1
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
USP46 affects USPs
| 1
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USP46 affects TBK1
| 1
TBK1 binds USP46 and USP12. 1 / 1
| 1

sparser
"In addition, USP12 and USP46 were not associated with TBK1 ( xref )."
USP46 affects DMWD
| 1
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
USP39 affects USP12
1 |
1 |

No evidence text available
USP12 affects tumour-promoting secretome resulting insufficient PPM1B deubiquitination
| 1
USP12 inhibits tumour-promoting secretome resulting insufficient PPM1B deubiquitination. 1 / 1
| 1

eidos
"Mechanistically , USP12 downregulation creates a tumour-promoting secretome resulting from insufficient PPM1B deubiquitination that causes NF-kappaB hyperactivation in tumour cells ."
USP12 affects tumor growth
| 1
USP12 activates tumor growth. 1 / 1
| 1

eidos
"Downregulation of USP12 inhibits tumor growth via the p38 / MAPK pathway in hepatocellular carcinoma ."
| 1
| 1

reach
"Thus, USP12 and OTUD7B counterbalance inhibitory E3 ligases to stabilize TCR signaling.Aside from acting at the proximal signalosome, E3 ligases also inhibit T-cell activation by acting further downstream, controlling the key transduction events that lead to the nuclear translocation of the T-cell activating transcription factors."
| PMC
USP12 affects transcription downstream METTL3
| 1
USP12 activates transcription downstream METTL3. 1 / 1
| 1

eidos
"Mechanistically , USP12 binds and stabilizes p300 , thereby activating the transcription of its downstream gene METTL3 ."
USP12 affects ribH2
| 1
USP12 inhibits ribH2. 1 / 1
| 1

reach
"The largely constant substrate affinity of USP12 in different forms negates a role of the BLs in controlling the accessibility of the Ub-binding surface of the enzyme."
USP12 affects protein stability cells
| 1
USP12 activates protein stability cells. 1 / 1
| 1

eidos
"USP12 could directly interact with MDK , decrease its polyubiquitination and increase its protein stability in cells ."
USP12 affects polyubiquitination
| 1
USP12 inhibits polyubiquitination. 1 / 1
| 1

eidos
"USP12 could directly interact with MDK , decrease its polyubiquitination and increase its protein stability in cells ."
USP12 affects pcDNA3.1
| 1
USP12 binds pcDNA3.1. 1 / 1
| 1

reach
"Human USP1, USP12, and UAF1 cDNAs were ligated to the pcDNA3.1 vector (Clontech Laboratories) modified with N-terminal Myc, hemagglutinin (HA), and FLAG peptides, respectively."

reach
"Deubiquitylase USP12 induces pro-survival autophagy and bortezomib resistance in multiple myeloma by stabilizing HMGB1."

reach
"We also investigated whether USP12 expression in tumour cells might modulate macrophage polarization."
USP12 affects lung tumour resistance PD-1
| 1
USP12 activates lung tumour resistance PD-1. 1 / 1
| 1

eidos
"USP12 downregulation orchestrates a protumourigenic microenvironment and enhances lung tumour resistance to PD-1 blockade ."
USP12 affects lung metastasis 4T1 cells
| 1
USP12 activates lung metastasis 4T1 cells. 1 / 1
| 1

eidos
"Knockdown of USP12 decreased the lung metastasis ability of 4T1 cells , while USP12 overexpression increased the lung metastasis ability of these cells in vivo ."
USP12 affects function monocytic MDSCs
| 1
USP12 activates function monocytic MDSCs. 1 / 1
| 1

eidos
"In the present study , we reported that USP12 deficiency decreased infiltration and impaired the suppressor function of monocytic ( M ) - MDSCs , resulting in increased CD8 + T cell response and decelerated tumor growth ."
USP12 affects function M-MDSCs
| 1
USP12 activates function M-MDSCs. 1 / 1
| 1

eidos
"Furthermore , USP12 deficiency inhibited the suppressor function of M-MDSCs by downregulating the negative regulatory molecules iNOS and PD-L1 , through deubiquitinating and stabilizing p65 ."
USP12 affects dephosphorylation TCPTP p-STAT1
| 1
USP12 inhibits dephosphorylation TCPTP p-STAT1. 1 / 1
| 1

eidos
"Upon IFN treatment , USP12 accumulates in the nucleus , blocks the CREB-binding protein-induced acetylation of p-STAT1 , and thus inhibits the dephosphorylation effects of TCPTP on p-STAT1 , which ultimately maintains the nuclear p-STAT1 levels and IFN antiviral efficacy ( 114 ) ."

reach
"Gene ontology (GO) analysis revealed that the pathways significantly enriched in USP12-downregulated genes were mostly related to cytokine-mediated signalling pathway and immune effector process (Fig. 3a); further in-depth examination showed that USP12 expression decreased levels of a number of chemokine, such as CXCL8, CXCL1, CXCL2, CCL2 and CCL5, most of which were associated with immune cell recruitment (Fig. 3b and Supplementary Fig. 2a)."
USP12 affects cre
| 1
USP12 binds cre. 1 / 1
| 1

sparser
"However, after exclusion of samples in which CNVs overlay the gene itself, amplification of the CRE was not associated with differential expression of USP12 or any other gene in the proximity of the CNV segments ( xref , xref )."
USP12 affects clonogenicity
| 1
USP12 activates clonogenicity. 1 / 1
| 1

eidos
"Depletion of USP12 suppressed cell growth and clonogenicity and inhibited autophagy ."

reach
"USP12 negatively regulates protumourigenic chemokine production."

reach
"Transient depletion of USP12, USP46, and USP53 did not induce a senescence arrest."
USP12 affects breast cancer angiogenesis
| 1
USP12 activates breast cancer angiogenesis. 1 / 1
| 1

eidos
"USP12 promotes breast cancer angiogenesis by maintaining midkine stability ."
USP12 affects bortezomib
| 1
| 1

reach
"Deubiquitylase USP12 induces pro-survival autophagy and bortezomib resistance in multiple myeloma by stabilizing HMGB1."
USP12 affects ZAP70
1 |
1 |

No evidence text available
USP12 affects WDR77
1 |
1 |

No evidence text available
USP12 affects WDR48.USP12 complex
| 1
USP12 activates WDR48.USP12 complex. 1 / 1
| 1

reach
"In fact, overexpression of WDR48 and USP12 together with PHLPP1 resulted in reduction of Akt phosphorylation, supporting the synergistic effect of the WDR48.USP12 complex along with PHLPP1 in suppressing Akt signaling (XREF_FIG D)."
USP12 affects WDR26
1 |
1 |

No evidence text available
USP12 affects Val-Ser
| 1
USP12 activates Val-Ser. 1 / 1
| 1

reach
"Our previous observations for the LNCaP cell line that express AR FL but not AR Vs have demonstrated that USP12 silencing causes a decrease in transcript expression of all the AR target genes [XREF_BIBR] supporting our data that AR Vs are not targeted by USP12 in the AR V expressing CWR22Rv1 cell line."
USP12 affects VSX1
1 |
1 |

No evidence text available
USP12 affects Ubiquitin, WDR20, and WDR48
| 1
| 1

sparser
"This model is inferred from a combination of structures of USP46 bound to WDR48 and ubiquitin (Yin et al., 2015) and USP12 bound to WDR48 and WDR20 (Li et al., 2016)."
USP12 affects USP46, USPs, and WDR48
| 1
| 1

sparser
"As stated earlier, UAF1 interacts with two other USPs, USP12 and USP46."
USP12 affects USP39
1 |
1 |

No evidence text available
USP12 affects USP12
| 1
USP12 bound to WDR48 activates USP12. 1 / 1
| 1

reach
"In C. elegans, USP46 and USP12 both bind to USP1 associated factor 1 (UAF1 and WDR48) and the binding dramatically enhances the activity of USP12 and USP46."
USP12 affects UL138
1 |
1 |

No evidence text available
USP12 affects UCHL5
1 |
1 |

No evidence text available
USP12 affects UBC
1 |
1 |

No evidence text available
USP12 affects UAF1DeltaSLD
| 1
USP12 binds UAF1DeltaSLD. 1 / 1
| 1

reach
"The UAF1DeltaSLD, WDR20, and USP12 complex was prepared by mixing the UAF1DeltaSLD and USP12 complex with proteolytically treated WDR20."
USP12 affects UAF1DSLD
| 1
USP12 binds UAF1DSLD. 1 / 1
| 1

reach
"The UAF1DSLD-WDR20-USP12 complex was prepared by mixing the UAF1DSLD-USP12 complex with proteolytically treated WDR20."
USP12 affects TSPYL6
1 |
1 |

No evidence text available
USP12 affects TRAT1
1 |
1 |

No evidence text available
USP12 affects TBK1
| 1
TBK1 binds USP46 and USP12. 1 / 1
| 1

sparser
"In addition, USP12 and USP46 were not associated with TBK1 ( xref )."
| 1

reach
"Thus, USP12 and OTUD7B counterbalance inhibitory E3 ligases to stabilize TCR signaling.Aside from acting at the proximal signalosome, E3 ligases also inhibit T-cell activation by acting further downstream, controlling the key transduction events that lead to the nuclear translocation of the T-cell activating transcription factors."
| PMC
USP12 affects Snail1
| 1
USP12 increases the amount of Snail1. 1 / 1
| 1

reach
"When these DUBs were co-expressed with Snail1 in HEK293 cells, we found that USP12, Dub3 and USP28 significantly increased Snail1 levels, similar to results obtained when cells were treated with the proteasome inhibitor MG132 (XREF_SUPPLEMENTARY)."
USP12 affects Sf9
| 1
USP12 increases the amount of Sf9. 1 / 1
| 1

reach
"USP1, USP12, and UAF1 were produced using Sf9 and Sf21 insect cell expression."
USP12 affects SDF4
1 |
1 |

No evidence text available
USP12 affects RRM2
| 1
USP12 deubiquitinates RRM2. 1 / 1
| 1

reach
"Here we showed that ubiquitin-specific peptidase 12 (USP12) directly interacts with and deubiquitinates RRM2 in non-small cell lung cancer (NSCLC) cells."
USP12 affects RNF19B
1 |
1 |

No evidence text available
USP12 affects RELA
| 1
USP12 deubiquitinates RELA. 1 / 1
| 1

reach
"Furthermore, USP12 deficiency inhibited the suppressor function of M-MDSCs by downregulating the negative regulatory molecules iNOS and PD-L1, through deubiquitinating and stabilizing p65."
USP12 affects RAD51AP1
1 |

No evidence text available
USP12 affects PRRC2A
| 1
USP12 inhibits PRRC2A. 1 / 1
| 1

eidos
"In vivo experiments showed that USP12-knockdown could suppress tumor growth in mice , and immuno-blotting revealed that USP12 could induce G2 / M arrest through the cyclin dependent kinase 1 / cyclinB1 axis , and trigger apoptosis via the p38 / mitogen-activated protein kinase pathway ."
USP12 affects PHLPP1.WDR48
| 1
USP12 binds PHLPP1.WDR48. 1 / 1
| 1

reach
"We further confirmed the existence of a specific PHLPP1.WDR48 and USP12 complex in vivo by demonstrating that WDR48 and USP12 but not other WD repeat protein WDR5 co-immunoprecipitated with exogenously expressed PHLPP1 in 293T cells (XREF_FIG D)."
USP12 affects PARD6B
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
USP12 affects PAIP1
1 |
1 |

No evidence text available
USP12 affects NUP160
1 |
1 |

No evidence text available
USP12 affects NOTCH1
1 |
1 |

No evidence text available
USP12 affects NLRP3
| 1
USP12 increases the amount of NLRP3. 1 / 1
| 1

reach
"Additionally, the study also found out that the UAF1/USP12 and UAF1/USP46 complexes increase p65 expression, which promotes NF-κB activation and increases NLRP3 and IL-1β expression levels."
| PMC
| 1

reach
"Deubiquitylase USP12 induces pro-survival autophagy and bortezomib resistance in multiple myeloma by stabilizing HMGB1."
USP12 affects Mice
| 1
USP12 activates Mice. 1 / 1
| 1

reach
"Exogenous USP12 expression markedly increased the survival of mice and decreased overall tumour number or burden in the entire lung compared with that of control mice (Fig. 2a–c)."
USP12 affects MON2
| 1
USP12 increases the amount of MON2. 1 / 1
| 1

reach
"USP1, USP12, and UAF1 were produced using Sf9 and Sf21 insect cell expression."
USP12 affects MMP2
1 |
1 |

No evidence text available
USP12 affects MDM2-P53
| 1
USP12 leads to the ubiquitination of MDM2-P53. 1 / 1
| 1

reach
"Prior studies suggested a role of USP12 in prostate carcinomas, wherein USP12 promoted AR signalling and tumour cell proliferation, as well as therapy resistance, by regulating AR ubiquitination and MDM2-P53 signalling ."
USP12 affects MAPK
| 1
USP12 activates MAPK. 1 / 1
| 1

reach
"Invivo experiments showed that USP12-knockdown could suppress tumor growth in mice, and immuno blotting revealed that USP12 could induce G2/M arrest through the cyclin dependent kinase1 and cyclinB1 axis, and trigger apoptosis via the p38 and mitogen activated protein kinase pathway."
USP12 affects M-MDSC
| 1
USP12 activates M-MDSC. 1 / 1
| 1

reach
"USP12 positively regulates M-MDSC function to inhibit anti-tumor immunity through deubiquitinating and stabilizing p65."

reach
"Thus, in this study we probed the involvement of USP12 in macrophage mediated inflammatory responses using bacterial endotoxin, LPS, as the model system."
USP12 affects KLK3
| 1
USP12 increases the amount of KLK3. 1 / 1
| 1

reach
"For example, USP26 physically interacts with AR and influences AR ubiquitination and transcriptional activation; 17 USP12 stabilizes AR, enhances its cellular function, and thereby triggers the gene expression of PSA."
USP12 affects KLF5
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
USP12 affects ISG65
| 1
USP12 increases the amount of ISG65. 1 / 1
| 1

reach
"UbH1 knockdown reduced ISG75 but not ISG65 expression (XREF_FIG), suggesting that deubiquitination by UbH1 specifically affects ISG75 copy number; clearly this mimics the direct effect of RNAi against ISG75."
USP12 affects IL6
| 1
Modified USP12 decreases the amount of IL6. 1 / 1
| 1

reach
"We observed that siRNA mediated ablation of USP12 expression in mouse macrophages suppressed the induction of LPS induced iNOS and IL-6 expression but failed to alter IFN-beta synthesis, oxidative stress and phagocytic ability of macrophages."
USP12 affects IL1B
| 1
USP12 increases the amount of IL1B. 1 / 1
| 1

reach
"Additionally, the study also found out that the UAF1/USP12 and UAF1/USP46 complexes increase p65 expression, which promotes NF-κB activation and increases NLRP3 and IL-1β expression levels."
| PMC
USP12 affects Histone
| 1
USP12 deubiquitinates Histone-H2A. 1 / 1
| 1

reach
"Joo et al. demonstrated that USP46 and USP12 containing fractions from HeLa cells can deubiquitinate histones H2A and H2B to control cell fate and gastrulation during Xenopus development [XREF_BIBR]."
USP12 affects HTT
| 1
USP12 inhibits HTT. 1 / 1
| 1

reach
"We propose a mechanism by which Usp12 suppresses HD related neurodegeneration via induction of autophagy."
USP12 affects HSPB9
1 |
1 |

No evidence text available
USP12 affects HMGB1-mediated autophagy
| 1
USP12 activates HMGB1-mediated autophagy. 1 / 1
| 1

eidos
"Knockdown of USP12 decreased the level of HMGB1 and suppressed HMGB1-mediated autophagy in MM ."
USP12 affects H2BK120ub
| 1
USP12 deubiquitinates H2BK120ub. 1 / 1
| 1

reach
"Other major DUBs with specificity for histone H2AK119ub over H2BK120ub are BAP1 and USP16, while USP3, USP12, USP22, and USP44 deubiquitinate both H2AK119ub and H2BK120ub, as well as different non histone substrates [XREF_BIBR]."
USP12 affects H2BC10
1 |
USP12 deubiquitinates H2BC10. 1 / 1
1 |

"It was shown that USP12 deubiquitinates histones H2A and H2B both in vitro and in vivo"
USP12 affects H2AK119ub
| 1
USP12 deubiquitinates H2AK119ub. 1 / 1
| 1

reach
"Other major DUBs with specificity for histone H2AK119ub over H2BK120ub are BAP1 and USP16, while USP3, USP12, USP22, and USP44 deubiquitinate both H2AK119ub and H2BK120ub, as well as different non histone substrates [XREF_BIBR]."
USP12 affects H2AC17
1 |
USP12 deubiquitinates H2AC17. 1 / 1
1 |

"It was shown that USP12 deubiquitinates histones H2A and H2B both in vitro and in vivo"
USP12 affects GRIA
| 1
USP12 deubiquitinates GRIA. 1 / 1
| 1

reach
"Knock-down of USP46, but not USP12, results in increased levels of ubiquitinated GluA1, decreased surface and total levels of GluA1, and reduced mEPSC amplitudes, consistent with a role for USP46 in deubiquitinating mammalian AMPARs (Huo et al., 2015)."
USP12 affects GRB2
1 |
1 |

No evidence text available
USP12 affects GPR161
1 |
1 |

No evidence text available
USP12 affects GPR12
| 1
| 1

sparser
"Three SNPs (SBNO2 rs2024092, CARD9 rs10781499, and rs17085007 between GPR12-USP12) were significantly associated with increased UC risk while NKX2-3 rs4409764 was a significant protective factor."
USP12 affects GORASP1
1 |
1 |

No evidence text available
USP12 affects FOXH1
| 1
| 1

sparser
"On comparing the binding characteristics of these mutants with USP12 WT , we noted that the UAF1 3X mutant only binds at very high concentrations and does not release easily, while the USP12 E190K mutant binds with a fast release ( xref a)."
USP12 affects E1
1 |
1 |

No evidence text available
USP12 affects Death
| 1
USP12 inhibits Death. 1 / 1
| 1

reach
"Usp12 overexpression significantly decreased the risk of death of HDQ109 neurons, but did not affect the survival of control neurons."
| 1

reach
"Knockdown of USP12 causes DNA replication stress and retards tumor growth in vivo and in vitro."
USP12 affects DHX8
1 |
1 |

No evidence text available
USP12 affects Cyclin
| 1
USP12 increases the amount of Cyclin. 1 / 1
| 1

reach
"In this study, we showed that knockdown of USP12 effectively induced cell cycle arrest in HeLa cells and decreased BMI-1, c-Myc and cyclin D2 transcription levels."
| 1

eidos
"In addition , USP12 promotes cell survival , proliferation , tumorigenesis , and cell cycle progression by upregulating BMI-1 , c-Myc , and cyclin D2 [ 24,25,26,27 ] ."
USP12 affects CTNNB1
| 1
USP12 increases the amount of CTNNB1. 1 / 1
| 1

reach
"Likewise, depletion of USP12, USP7, or FBXO36 increased total cellular levels of beta-catenin."
USP12 affects CREB-binding protein-induced acetylation p-STAT1
| 1
USP12 inhibits CREB-binding protein-induced acetylation p-STAT1. 1 / 1
| 1

eidos
"Upon IFN treatment , USP12 accumulates in the nucleus , blocks the CREB-binding protein-induced acetylation of p-STAT1 , and thus inhibits the dephosphorylation effects of TCPTP on p-STAT1 , which ultimately maintains the nuclear p-STAT1 levels and IFN antiviral efficacy ( 114 ) ."
USP12 affects CD4+
| 1
USP12 activates CD4+. 1 / 1
| 1

reach
"USP12 promotes CD4+ T cell responses through deubiquitinating and stabilizing BCL10."
USP12 affects CD4 T cell responses
| 1
USP12 activates CD4 T cell responses. 1 / 1
| 1

eidos
"USP12 promotes CD4 + T cell responses through deubiquitinating and stabilizing BCL10 ."
USP12 affects CD274
| 1
USP12 decreases the amount of CD274. 1 / 1
| 1

reach
"Accordingly, we found that silencing USP12 enhanced PD-L1 expression in tumour cells."
USP12 affects CBLB
1 |
1 |

No evidence text available
USP12 affects CASC3
| 1
USP12 activates CASC3. 1 / 1
| 1

reach
"USP12 depletion, concomitant with a reduced expression of HMGB1, suppressed autophagy and increased the sensitivity of resistant cells to BTZ."
USP12 affects BMI1
| 1
USP12 increases the amount of BMI1. 1 / 1
| 1

reach
"In this study, we showed that knockdown of USP12 effectively induced cell cycle arrest in HeLa cells and decreased BMI-1, c-Myc and cyclin D2 transcription levels."
| 1

reach
"USP12 deficiency also significantly attenuate the phenotype of Ang II induced cardiac hypertrophy in vivo."
USP12 affects Ang II-induced myocardial hypertrophy
| 1
USP12 inhibits Ang II-induced myocardial hypertrophy. 1 / 1
| 1

eidos
"Inhibition of USP12 ameliorate Ang II-induced myocardial hypertrophy , while overexpression of USP12 have the opposite effect ."
USP12 affects APP
1 |
1 |

No evidence text available
USP12 affects ANO7
| 1
| 1

sparser
"To establish if this was the case in PC cell lines, we firstly investigated potential Usp12-PHLPP/L interactions in androgen-dependent LNCaP cells by immunoprecipitation."
USP12 affects AKT-mTOR
| 1
USP12 inhibits AKT-mTOR. 1 / 1
| 1

reach
"Deletion analysis of USP12 promoter activity showed that the region spanning −2928 to −1337 bases relative to the transcriptional start site was responsible for the downregulation of USP12 by AKT-mTOR signalling (Supplementary Fig. 1h)."
USP affects USP12
| 1
USP activates USP12. 1 / 1
| 1

sparser
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases (USPs) constitute the largest family of deubiquitinating enzymes, whose catalytic competency is often modulated by their binding partners through unknown mechanisms."
UL138 affects USP12
1 |
1 |

No evidence text available
UCHL5 affects USP12
1 |
1 |

No evidence text available
UBC affects USP12
1 |
1 |

No evidence text available
UAF1DeltaSLD affects USP12
| 1
USP12 binds UAF1DeltaSLD. 1 / 1
| 1

reach
"The UAF1DeltaSLD, WDR20, and USP12 complex was prepared by mixing the UAF1DeltaSLD and USP12 complex with proteolytically treated WDR20."
UAF1DSLD affects USP12
| 1
USP12 binds UAF1DSLD. 1 / 1
| 1

reach
"The UAF1DSLD-WDR20-USP12 complex was prepared by mixing the UAF1DSLD-USP12 complex with proteolytically treated WDR20."
TSPYL6 affects USP12
1 |
1 |

No evidence text available
TRAT1 affects USP12
1 |
1 |

No evidence text available
TP53 affects USP12
1 |
TP53 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
TBK1 affects USP46
| 1
TBK1 binds USP46 and USP12. 1 / 1
| 1

sparser
"In addition, USP12 and USP46 were not associated with TBK1 ( xref )."
T-2 Toxin affects USP12
1 |
T-2 Toxin decreases the amount of USP12. 1 / 1
1 |

No evidence text available
Small hairpin affects USP12
| 1
Small hairpin inhibits USP12. 1 / 1
| 1

reach
"Small hairpin RNA-mediated silencing of USP12 confirmed the findings in both human and mouse lung cancer cells, as evidenced by ELISA quantification (Supplementary Fig. 2b, c)."
STUB1 affects USP12
| 1
STUB1 activates USP12. 1 / 1
| 1

reach
"XREF_BIBR, XREF_BIBR In addition, other E3 ubiquitin ligases are probably involved in the AR regulation process, such as CHIP XREF_BIBR and SKP2 XREF_BIBR which also promote AR degradation, and Siah2, XREF_BIBR RNF6, XREF_BIBR and USP12 XREF_BIBR, XREF_BIBR which promote deubiquitination and AR activation."
SQSTM1 affects OPTN
| 1
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
SP1 affects USP12
| 1
SP1 activates USP12. 1 / 1
| 1

reach
"Over-expression of Sp1 and p65 upregulates the promoter activity of USP12."
SKP2 affects USP12
| 1
SKP2 activates USP12. 1 / 1
| 1

reach
"XREF_BIBR, XREF_BIBR In addition, other E3 ubiquitin ligases are probably involved in the AR regulation process, such as CHIP XREF_BIBR and SKP2 XREF_BIBR which also promote AR degradation, and Siah2, XREF_BIBR RNF6, XREF_BIBR and USP12 XREF_BIBR, XREF_BIBR which promote deubiquitination and AR activation."
SDF4 affects USP12
1 |
1 |

No evidence text available
RNF19B affects USP12
1 |
1 |

No evidence text available
RNAi affects USP12
| 1
RNAi activates USP12. 1 / 1
| 1

reach
"qPCR data suggest that RNAi mediated inactivation of USP49, FBXO36, and USP12 depleted levels of the corresponding transcripts."
RELA affects USP12
| 1
RELA activates USP12. 1 / 1
| 1

reach
"Over-expression of Sp1 and p65 upregulates the promoter activity of USP12."
RAD51AP1 affects USP12
1 |

No evidence text available
Protease affects USP12
| 1
| 1

eidos
"Highlights d Free USP12 deubiquitinase is inactive and has several flexible structural elements d UAF1 and WDR20 can both activate USP12 without increasing its substrate affinity d UAF1 and WDR20 bind USP12 at two distinct sites away from its catalytic center d Two distinct allosteric mechanisms underlie USP12 activation by UAF1 and WDR20 Ubiquitin-specific proteases ( USPs ) constitute the largest family of deubiquitinating enzymes , whose catalytic competency is often modulated by their binding partners through unknown mechanisms ."
PPM1B affects USP12
| 1
| 1

reach
"The interaction between PPM1B and USP12 was then validated (Fig. 4c), raising the question of whether USP12 deubiquitinates PPM1B."
PPARA affects USP12
1 |
PPARA decreases the amount of USP12. 1 / 1
1 |

No evidence text available
PHLPP2 affects WDR20
| 1
| 1

sparser
"Both Usp12 and WDR20 alone were able to interact with both PHLPP (Figure xref ) and PHLPPL (Figure xref )."
PHLPP2 affects ANO7
| 1
| 1

sparser
"To establish if this was the case in PC cell lines, we firstly investigated potential Usp12-PHLPP/L interactions in androgen-dependent LNCaP cells by immunoprecipitation."
PHLPP1.WDR48 affects USP12
| 1
USP12 binds PHLPP1.WDR48. 1 / 1
| 1

reach
"We further confirmed the existence of a specific PHLPP1.WDR48 and USP12 complex in vivo by demonstrating that WDR48 and USP12 but not other WD repeat protein WDR5 co-immunoprecipitated with exogenously expressed PHLPP1 in 293T cells (XREF_FIG D)."
PHLPP1 affects USPs
| 1
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
PHLPP1 affects USP12, and WDR48
| 1
| 1

sparser
"We confirmed the association of endogenous PHLPP1 with WDR48 and USP12 by immunoprecipitation with PHLPP1 antibody ( xref B )."
PHLPP1 affects USP12, USPs, and WDR48
| 1
| 1

sparser
"Interestingly, we observed that PHLPP1 specifically associated with WDR48 and USP12 but not other USPs ( xref C )."
PAX4 affects USP12
1 |
PAX4 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
PARD6B affects USP12
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
PAIP1 affects USP12
1 |
1 |

No evidence text available
OPTN affects SQSTM1, and USP12
| 1
| 1

sparser
"We have found an intriguing interaction of Usp12 with p62 and Optn, two key receptors necessary for the selective-autophagic clearance of misfolded proteins, mitochondria, and protein aggregates."
NUP160 affects USP12
1 |
1 |

No evidence text available
NR2F2 affects USP12
1 |
NR2F2 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
NOTCH1 affects USP12
1 |
1 |

No evidence text available
MYC affects USP12
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
MMP2 affects USP12
1 |
1 |

No evidence text available
LAT affects USP12
1 |
1 |

No evidence text available
KLF5 affects USP12
| 1
PARD6B binds USP12, KLF5, and MYC. 1 / 1
| 1

sparser
"These super-enhancer amplifications are associated with over-expression of the MYC oncogene as well as the KLF5 , USP12 and PARD6B genes."
JUN affects USP12
1 |
JUN decreases the amount of USP12. 1 / 1
1 |

No evidence text available
HSPB9 affects USP12
1 |
1 |

No evidence text available
HNF4A affects USP12
1 |
HNF4A decreases the amount of USP12. 1 / 1
1 |

No evidence text available
H2BC21 affects USP12
1 |
1 |

No evidence text available
H2AC20 affects USP12
1 |
1 |

No evidence text available
GRB2 affects USP12
1 |
1 |

No evidence text available
GPR161 affects USP12
1 |
1 |

No evidence text available
GPR12 affects USP12
| 1
| 1

sparser
"Three SNPs (SBNO2 rs2024092, CARD9 rs10781499, and rs17085007 between GPR12-USP12) were significantly associated with increased UC risk while NKX2-3 rs4409764 was a significant protective factor."
GORASP1 affects USP12
1 |
1 |

No evidence text available
Figures affects USP12
| 1
Figures activates USP12. 1 / 1
| 1

reach
"In agreement with our enzyme kinetics data, our binding studies revealed negligible changes in the substrate binding affinity of USP12 induced by UAF1 or WDR20 (Figures 6A and S6) ."
Figure affects USP12
| 1
Figure activates USP12. 1 / 1
| 1

eidos
"Based on our results , we propose a tenable model for USP12 activation by UAF1 and WDR20 ( Figure 7 ) ."
FOXH1 affects USP12
| 1
| 1

sparser
"On comparing the binding characteristics of these mutants with USP12 WT , we noted that the UAF1 3X mutant only binds at very high concentrations and does not release easily, while the USP12 E190K mutant binds with a fast release ( xref a)."
ESR1 affects USP12
1 |
ESR1 decreases the amount of USP12. 1 / 1
1 |

No evidence text available
EPO affects USP12
| 1
EPO increases the amount of USP12. 1 / 1
| 1

reach
"In a category of ubiquitinylation (XREF_SUPPLEMENTARY), EPO induced the expression of Usp12 and Fbxw7."
EBNA3 affects USP12
| 1
EBNA3 activates USP12. 1 / 1
| 1

reach
"Because we did not find peptides corresponding to other EBNA proteins in these complexes, each EBNA3 protein appears to target RBPJ and the USP46 and USP12 DUB complexes independently."
E1 affects USP12
1 |
1 |

No evidence text available
DMWD affects WDR20
| 1
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
DMWD affects USP46
| 1
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
DMWD affects USP12, and WDR20
| 1
WDR20 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Altogether, our findings suggest a model whereby mutually exclusive binding of DMWD and WDR20 to USP12 may lead to formation of deubiquitinase complexes with distinct subcellular localization, potentially targeting different substrate repertoires."
DMWD affects USP12, and USP46
| 1
USP46 binds USP12 and DMWD. 1 / 1
| 1

sparser
"Experimentally, we first confirmed that DMWD binds both USP12 and USP46 through direct co-immunoprecipitation of epitope-tagged proteins."
DHX8 affects USP12
1 |
1 |

No evidence text available
DEHP A2780 cells affects USP12
| 1
DEHP A2780 cells activates USP12. 1 / 1
| 1

eidos
"Our multiplex RT-PCR results showed that USP12 expression was increased by DEHP in A2780 cells , and thus , the above-mentioned findings of its influence on cell cycle progression and proliferation suggest that it may induce POF by influencing oocyte maturation ."
Cuprizone affects USP12
1 |
Cuprizone decreases the amount of USP12. 1 / 1
1 |

No evidence text available
CBLB affects USP12
1 |
1 |

No evidence text available

No evidence text available
ATXN3 affects PHB2, TXN, UCHL5, USP1, USP10, USP12, and USP14
| 1
| 1

sparser
"Other examples of protein complexes retrieved include host cell factor-1, galectin 7, p97, ras GTPase binding proteins 1 and 2, WD-repeat containing protein 48 and WD-repeat containing protein 20, and thioredoxin like protein, which interact with BAP, USP1, ataxin-3, USP10, USP12, and USP14 or UCH-L5 [ xref , xref ], respectively."
APP affects USP12
1 |
1 |

No evidence text available
ANO7 affects USP12
| 1
| 1

sparser
"To establish if this was the case in PC cell lines, we firstly investigated potential Usp12-PHLPP/L interactions in androgen-dependent LNCaP cells by immunoprecipitation."
AKT2 affects USP12
| 1
AKT2 decreases the amount of USP12. 1 / 1
| 1

reach
"Consistently, ectopic expression of the myristoylated form of AKT1 (myr-AKT1) significantly repressed USP12 expression in both human and mouse tumour cells (Fig. 1k); knockdown of endogenous AKT1 or AKT2, the two major isoforms of AKT, induced higher levels of USP12 than did the control treatment (Fig. 1l)."
AKT1 affects USP12
| 1
AKT1 decreases the amount of USP12. 1 / 1
| 1

reach
"Consistently, ectopic expression of the myristoylated form of AKT1 (myr-AKT1) significantly repressed USP12 expression in both human and mouse tumour cells (Fig. 1k); knockdown of endogenous AKT1 or AKT2, the two major isoforms of AKT, induced higher levels of USP12 than did the control treatment (Fig. 1l)."
1 |

No evidence text available
1 |

No evidence text available
1 |
17beta-estradiol increases the amount of USP12. 1 / 1
1 |

No evidence text available
4,4'-diaminodiphenylmethane increases the amount of USP12. 1 / 1
1 |

No evidence text available