IndraLab

Statements


OTUD5 affects TP53
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OTUD5 binds TP53.
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"Taken together, these results provide clinical evidence for p53OTUD5 axis‐mediated regulation of GPX4 protein stability, suggesting that the expression level of OTUD5 is important for ferroptosis of gastric cancer cells."

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"A recent study showed that in case of stress, programmed cell death 5 (PDCD5) is required to promote the interaction between OTUD5 and p53."

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"Since OTUD5 was shown to interact with p53, we also performed mapping analysis of the interaction between OTUD5 and p53."

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"However, upon prolonged genotoxic stress, the interaction between OTUD5 and PDCD5 was reduced, while the association between OTUD5 and p53 gradually increased, leading to p53 induction ( Fig. 4A )."

sparser
"OTUD5 can bind to TP53 and regulate its ubiquitination levels [ xref ]."

sparser
"A recent study showed that in case of stress, programmed cell death 5 (PDCD5) is required to promote the interaction between OTUD5 and p53."

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"OTUD5 can bind to TP53 and regulate its ubiquitination levels [56]."

sparser
"Since OTUD5 was shown to interact with p53, we also performed mapping analysis of the interaction between OTUD5 and p53."

sparser
"OTUD5 forms a direct complex with p53 and controls level of ubiquitination."

sparser
"P53 interacts with OTUD5."
OTUD5 activates TP53.
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OTUD5 activates TP53. 10 / 18
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"Accordingly, OTUD5 overexpression has been shown to effectively enhance the activation of PDCD5 and TP53."

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"This sequential interplay between OTUD5 and either PDCD5 or p53 was abolished by knockdown of either OTUD5 or PDCD5 ( Fig. 4B , second and third panel), suggesting that PDCD5 stabilization by OTUD5 pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"OTU deubiquitinase (OTUD5) binds to PDCD5inresponseto etoposide treatment and effectively mediates the sequential activation of both PDCD5 and P53."

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"The function of OTUD5 is required to allow the rapid activation of p53 dependent transcription and a p53 dependent apoptosis in response to DNA damage stress."

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"These data demonstrated that OTUD5 mediated sequential activation of PDCD5 and p53 in response to genotoxic stress.We further validated our findings using PDCD5- depleted mouse embryonic fibroblast (M[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"OTUD5 can increase the stability of TP53 and PDCD5 through deubiquitination [22] (Figure 3D)."

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"Conversely, reintroduction of PDCD5 in PDCD5 −/− MEF cells dramatically restored OTUD5 WT function in promoting p53 activation."

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"To this end, 1mug of p53 and increasing amounts of Flag-OTUD5 were cotransfected into H1299 cells."

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"Importantly, we found that OTUD5 promotes the p53 pathway in a PDCD5-dependent manner."

sparser
"As expected, overexpression of the PDCD5 E94D mutant failed to reactivate p53 even with OTUD5 WT present, suggesting that OTUD5 activated p53 via PDCD5 stabilization during response to genotoxic stres[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 activates mutated TP53. 1 / 1
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"Conversely, in gastric cancer cells harbouring p53 mutations, OTUD5 transcription is upregulated due to the inactivation of p53, allowing OTUD5 to stabilise GPX4 protein, inhibit ferroptosis and thereby promoting gastric cancer progression (Figure 6J, right panel)."
OTUD5 deubiquitinates TP53.
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OTUD5 deubiquitinates TP53. 10 / 12
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"In this study, we provide new evidences that OTUD5 can also function to deubiquitinate and stabilize p53."

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"To verify that the deubiquitination of p53 by OTUD5 is specific, MDM2 was also tested in the in vitro deubiquitination assay and found thatMDM2 can not be deubiquitinated by OTUD5 (XREF_FIG)."

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"These data indicate that both PDCD5 and OTUD5 are required for p53-mediated apoptosis in response to genotoxic stress.Previous work has shown that OTUD5 stabilizes p53 via deubiquitination of p53 [16][MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Other DUBs, OTUD1 and OTUD5, are also involved in the deubiquitination and stabilization of TP53 in response to DNA damage stress in U2OS cells [51,52]."

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"Besides, OTUB1 and OTUD5 are also discovered to positively regulate p53 stability by deubiquitinating p53 [8] [9] ."

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"OTU deubiquitinase 5 (OTUD5) can also deubiquitinate p53 protein and stabilize it, thus affecting apoptosis that is induced by DNA damage [38]."

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"OTUD1, OTUD5 and USP11 directly deubiquitinating p53 and functional proteins were required for p53 stabilization [XREF_BIBR - XREF_BIBR]."

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"We show that (1) OTUD5 is a novel p53 interacting protein; (2) OTUD5 deubiquitinates p53, leading to the stabilization of p53; (3) OTUD5 can help in the stabilization and activation of p53 in response to DNA damage; (4) OTUD5 is required for p53 induced apoptosis in response to DNA damage."

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"Another DUB, OTUD5, interacts and deubiquitylates p53 in response to DNA damage stress."

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"To verify that the deubiquitination of p53 by OTUD5 is specific, MDM2 was also tested in the in vitro deubiquitination assay."
OTUD5 inhibits TP53.
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OTUD5 inhibits TP53. 3 / 5
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sparser
"The knockdown of OTUD5 inactivated p53 and PDCD5, promoting the proliferation and metastasis of NSCLC cells while inhibiting their apoptosis."
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"In line with this, overexpression of OTUD5 can block p53 activation‐induced GPX4 downregulation (Figure 4C)."

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"Moreover, ectopic expression of phosphomimic KAP1 also rescued the telomere MMEJ defect in OTUD5-depleted Trf1 ;Trf2 ;Ku70 ;p53 ;Cre-ER MEFs (Fig. 5g, Supplementary Fig. 7d, e)."
OTUD5 increases the amount of TP53.
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OTUD5 increases the amount of TP53. 3 / 3
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"Collectively, these results demonstrated that OTUD5 stabilized PDCD5 via dequibiquitination of PDCD5 at Lys-97 and 98.Because overexpression of OTUD5 enhanced the levels of PDCD5, we next examined whe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Overexpression of OTUD5 increased the levels of PDCD5 and p53 in a time-dependent manner; whereas knockdown of OTUD5 substantially impaired the positive effect of ET on PDCD5 and p53 activation ( Fig.[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Intriguingly, overexpression of OTUD5 failed to increase the levels of p53 activity when PDCD5 was knocked down."
OTUD5 decreases the amount of TP53.
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OTUD5 decreases the amount of TP53. 1 / 2
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"Using shRNA mediated depletion of OTUD5 in U2OS cells caused marked decrease of p53 protein levels with no effect on p53 mRNA levels (XREF_FIG)."
OTUD5 affects PDCD5
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OTUD5 binds PDCD5.
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5 | 8 15

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"Therefore, dysregulation of p53-interacting deubiquitinating enzymes may lead to aberrant regulation of genotoxic responses in cancer cells.To understand how the tumor suppressor PDCD5 is regulated du[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"However, knocking-down of OTUD5 markedly diminished the association of PDCD5 with p53, indicating that OTUD5 binds to PDCD5 and mediates PDCD5-p53 network during genotoxic stress responses ( Fig. 1B )[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Thus, we next explored the possibility that OTUD5 initially binds to and stabilizes PDCD5 and subsequently stabilizes p53 in response to genotoxic stress."

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"However, upon prolonged genotoxic stress, the interaction between OTUD5 and PDCD5 was reduced, while the association between OTUD5 and p53 gradually increased, leading to p53 induction ( Fig. 4A )."

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"We corroborated these findings by a reconstitution experiment where a point mutant defective for interaction with OTUD5 (PDCD5 E94D ) showed that OTUD5 interaction with PDCD5 is a prerequisite for p53[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Park et al ( xref ) identified the programmed cell death 5-OTUD5 network as a central hub for regulating p53-mediated apoptosis."

sparser
"Under genotoxic stress, OTUD5 binds to PDCD5 and increases PDCD5 stability by mediating the deubiquitination of PDCD5 [ xref ]."

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"Park et al. xref further revealed that deubiquitinase OTUD5 directly binds to PDCD5 and promotes its accumulation by mediating PDCD5 deubiquitination at Lys‐97/98 upon genotoxic stress‐induced apoptosis."

sparser
"To understand how the tumor suppressor PDCD5 is regulated during apoptosis, we employed yeast two-hybrid assays to identify interacting partners of PDCD5 and found that OTUD5 bound to PDCD5 in respons[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Park and colleagues [17] found that OTUD5 binds to PDCD5 in response to etoposide treatment via its ubiquitin interacting motif (UIM)."
OTUD5 activates PDCD5.
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OTUD5 activates PDCD5. 10 / 17
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"Importantly, we found that OTUD5 promotes the p53 pathway in a PDCD5-dependent manner."

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"Accordingly, OTUD5 overexpression has been shown to effectively enhance the activation of PDCD5 and TP53."

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"This sequential interplay between OTUD5 and either PDCD5 or p53 was abolished by knockdown of either OTUD5 or PDCD5 ( Fig. 4B , second and third panel), suggesting that PDCD5 stabilization by OTUD5 pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Therefore, OTUD5 and YAF2 promote the TP53 signal pathway in a PDCD5-dependent manner, highlighting the critical role of the PDCD5 network, which acts as a central hub in the regulation of genotoxic s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The knockdown of OTUD5 inactivated p53 and PDCD5, promoting the proliferation and metastasis of NSCLC cells while inhibiting their apoptosis."

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"Moreover, we showed that OTUD5 mediated the sequential activation of PDCD5 and p53 in response to genotoxic stress."

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"Because overexpression of OTUD5 increased the PDCD5 stability via its UIM motif, we also investigated whether OTUD5 increased PDCD5 stability by deubiquitinating PDCD5."

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"By both in situ PLA and immunoprecipitation assays, we demonstrated that knockdown of OTUD5 eliminated the induction of PDCD5 at the early stage of the DNA damage response and failed to induce the p53[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Treatment with MG132 efficiently increased PDCD5 levels; whereas, knockdown of OTUD5 greatly diminished the effect of MG132 on the stabilization of PDCD5, indicating that OTUD5 played a critical role [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"A time-course experiment with cycloheximide (CHX) treatment showed that OTUD5 WT enhanced PDCD5 stability; however, inactive mutant OTUD5 C224S failed to increase the stability of PDCD5 ( Fig. 2B )."
OTUD5 deubiquitinates PDCD5.
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OTUD5 deubiquitinates PDCD5. 3 / 4
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"Because overexpression of OTUD5 increased the PDCD5 stability via its UIM motif, we also investigated whether OTUD5 increased PDCD5 stability by deubiquitinating PDCD5."

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"Strikingly, wild-type OTUD5 efficiently reduced the ubiquitination of PDCD5 when compared with the inactive mutant OTUD5 C224S ( Fig. 2D ), while knockdown of OTUD5 increased the ubiquitination of PDC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"We also observed that OTUD5 robustly reduced the ubiquitination of PDCD5 when compared to overexpression of other deubiquitinating enzymes, indicating the selective action of OTUD5 on PDCD5 ubiquitina[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 deubiquitinates PDCD5 on K97. 3 / 3
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"OTUD5 thus deubiquitinates PDCD5 at Lys97 and Lys98 subsequently regulating p53 levels."

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"Furthermore, OTUD5 increased PDCD5 levels by mediating deubiquitination of PDCD5 at Lys-97/98."

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"OTUD5 expression increased the stability of PDCD5 by mediating deubiquitination of PDCD5 at Lys-97/98, efficiently enhancing the activation of both PDCD5 and TP53, followed by elevated BAX and P21 pro[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 deubiquitinates PDCD5 on K98. 3 / 3
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"Furthermore, OTUD5 increased PDCD5 levels by mediating deubiquitination of PDCD5 at Lys-97/98."

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"OTUD5 expression increased the stability of PDCD5 by mediating deubiquitination of PDCD5 at Lys-97/98, efficiently enhancing the activation of both PDCD5 and TP53, followed by elevated BAX and P21 pro[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"OTUD5 thus deubiquitinates PDCD5 at Lys97 and Lys98 subsequently regulating p53 levels."
OTUD5 increases the amount of PDCD5.
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OTUD5 increases the amount of PDCD5. 6 / 6
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"Importantly, overexpression of wild-type OTUD5 robustly increased the levels of endogenous PDCD5, as well as p53."

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"Collectively, these results demonstrated that OTUD5 stabilized PDCD5 via dequibiquitination of PDCD5 at Lys-97 and 98.Because overexpression of OTUD5 enhanced the levels of PDCD5, we next examined whe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Overexpression of OTUD5 increased the levels of PDCD5 and p53 in a time-dependent manner; whereas knockdown of OTUD5 substantially impaired the positive effect of ET on PDCD5 and p53 activation ( Fig.[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Taken together, we concluded that OTUD5 increased PDCD5 levels via interaction with its UIM motif.Because overexpression of OTUD5 enhanced PDCD5 expression, we next examined if OTUD5 increased the sta[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Importantly, knockdown of OTUD5 reversed the effect of PDCD5 overexpression on p53 activation, indicating that OTUD5 is required for PDCD5-mediated p53 activation ( Fig. 3C )."

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"Furthermore, OTUD5 increased PDCD5 levels by mediating deubiquitination of PDCD5 at Lys-97/98."
OTUD5 inhibits PDCD5.
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OTUD5 inhibits PDCD5. 1 / 1
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sparser
"The knockdown of OTUD5 inactivated p53 and PDCD5, promoting the proliferation and metastasis of NSCLC cells while inhibiting their apoptosis."
| PMC
OTUD5 affects GPX4
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OTUD5 binds GPX4.
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"Next, we determined the interaction of OTUD5 and GPX4 by co‐immunoprecipitation and mass spectrometry (Figure  4A; Table S2, Supporting Information)."

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"The direct interaction of OTUD5 and GPX4 was further confirmed by GST‐pull down, surface plasmon resonance (SPR) and microscale thermophoresis (MST) (Figure 4B,C; Figure S5E, Supporting Information), and the colocalization of OTUD5 and GPX4 proteins in the NRCMs was verified by immunofluorescence (Figure 4D)."

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"As OTUD5 belongs to the DUBs family and plays a critical role in controlling the Ub‐dependent degradation of various proteins, 4‐HNE may reduce the interaction between OTUD5 and GPX4 to promote GPX4 ubiquitination and subsequent degradation.To test whether OTUD5 affects the protein stability of GPX4, we co‐transfected GPX4 with Flag‐OTUD5 or its enzyme‐inactive mutant C224S into HEK293T cells."

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"Taken together, OTUD5 specifically deubiquitinated and stabilized the protein level of GPX4.Furthermore, we confirmed that GPX4 and OTUD5 could form a complex through reciprocal coimmunoprecipitation, immunofluorescence, duolink proximity ligation in situ assay (PLA), and GST‐pull down assay, and that 4‐HNE could weak this binding (Figure  5A–D; Figure S6E, Supporting Information)."

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"Taken together, these results indicate that OTUD5 depletion results in increased ferroptosis in cardiomyocytes.2.6 4-HNE Affects the Interaction between OTUD5 and GPX4 by Addition and Carbonylation."

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"We used the database to predict potential ubiquitination‐related enzymes targeting GPX4 (Figure S5A, Supporting Information) and found that E3 ligases MIB2, NEDD4L, STUB1, and deubiquitinase OTUD5 could bind to GPX4 in co‐immunoprecipitation experiments (Figure S5B, Supporting Information)."

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"In addition, our results demonstrated that only the binding between OTUD5 and GPX4 was affected under 4‐HNE treatment, while the interactions between E3 ligases (MIB2, NEDD4L, and STUB1) and GPX4 were not changed (Figure S5C, Supporting Information)."

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"In ALDH2 cKO mice, co‐immunoprecipitation assays showed that the levels of 4‐HNE adducted to GPX4 and OTUD5 were increased and the binding between GPX4 and OTUD5 was reduced (Figure S8E,F, Supporting Information)."

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"In mutant‐transfected HEK293T cells, the carbonylation of GPX4 and OTUD5 were significantly abolished (Figure 6G), the ability of 4‐HNE to weaken the interaction of GPX4 and OTUD5 and promote GPX4 ubiquitin–proteasomal degradation was eliminated (Figure 6H)."

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"N‐acetyl cysteine (NAC), an analog of cysteine that competitively binds to 4‐HNE, and amino acid substitution rescued 4‐HNE‐induced cysteine carbonylation of GPX4 and OTUD5, recovered the interaction between GPX4 and OTUD5, alleviated ubiquitination‐dependent degradation of GPX4, and inhibited myocyte ferroptosis."
OTUD5 deubiquitinates GPX4.
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OTUD5 deubiquitinates GPX4. 10 / 12
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"OTUD5 mediates GPX4 deubiquitination to regulate its stability.Deletion of OTUD5 promotes ferroptosis and inhibits tumor growth.Wild type p53 inhibits OTUD5 transcription, thereby promoting GPX4 degradation and inhibiting the development of gastric cancer.OTUD5, GPX4 expression and p53 activity are highly correlated and correlates with clinical progression in STAD."

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"Co‐IP showed that OTUD5 interacts with GPX4 in cells (Figure 1E–H), which was further confirmed by their spatial proximity through immunofluorescence (Figure 1I).2.2 OTUD5 deubiquitinates and stabilises GPX4."

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"Therefore, identifying proteins that regulate GPX4 degradation is imperative for promoting ferroptosis without leading to the total elimination of GPX4.Next, we examined whether OTUD5 mediates the deubiquitination of GPX4 and found that overexpression of OTUD5 led to a significant reduction of GPX4 polyubiquitination (Figure 2J,K)."

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"Notably, the expression of wild‐type OTUD5 (OTUD5‐WT), but not the catalytically inactive mutant (OTUD5‐C224S) reduced GPX4 ubiquitination (Figure 2L)."

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"These data suggested that OTUD5 deubiquitinates GPX4, thereby enhancing its protein stability."

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"Additionally, we found that knockdown or depletion of OTUD5 markedly increased GPX4 ubiquitination (Figure 2N,O)."

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"In conclusion, these results demonstrated that OTUD5 mediates the K48‐linked deubiquitination of GPX4 and stabilises GPX4.2.3 OTUD5 suppresses ferroptosis via GPX4."

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"When experiencing I/R-induced AKI in mice or H/R-induced AKI in HK-2 cells, the deubiquitinase OTUD5 reportedly decreases the ubiquitination of GPX4, thus protecting GPX4 from TAX1BP1-dependent autophagy [131]."

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"Silencing OTUD5 increased GPX4 ubiquitination in HBMECs co-cultured with siTHBS1-AMφ-exo (Fig. 8G)."

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"OTUD5 increased GPX4 protein levels and decreased GPX4 ubiquitination in a dose‐dependent manner, whereas ectopic expression of OTUD5 did not affect GPX4 (Figure 4E,F; Figure S5E, Supporting Information), indicating that OTUD5 enzyme activity is indispensable for GPX4 protein stabilization."
OTUD5 increases the amount of GPX4.
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OTUD5 increases the amount of GPX4. 7 / 7
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"OTU deubiquitinase 5 (OTUD5), a protein that interacts with GPX4, can promote ferroptosis resistance during ischemia/reperfusion injury by stabilizing GPX4 expression; in turn, hypoxia/ischemia-induced OTUD5 autophagy can destabilize GPX4, leading to ferroptosis-dependent kidney injury (96)."

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"OTUD5 increased GPX4 protein levels and decreased GPX4 ubiquitination in a dose‐dependent manner, whereas ectopic expression of OTUD5 did not affect GPX4 (Figure 4E,F; Figure S5E, Supporting Information), indicating that OTUD5 enzyme activity is indispensable for GPX4 protein stabilization."

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"Notably, OTUD5 overexpression, knockdown or knockout did not impact the mRNA levels of GPX4 (Figure S1A–C), indicating that OTUD5 upregulates GPX4 expression independently of transcriptional level in gastric cancer cells.Subsequently, we determined the regulation of OTUD5 on the half‐life of GPX4."

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"Next, we employed CRISPR‐Cas9 genome editing technology to delete OTUD5 in AGS and mouse forestomach cancer (MFC) cells, and found that depletion of OTUD5 significantly suppressed the expression of GPX4 in both of the cells (Figure 5A,B)."

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"Results revealed that overexpression of wild‐type OTUD5 led to elevated GPX4 protein levels, whereas the catalytic mutant C224S‐OTUD5 did not influence GPX4 expression (Figure 2A), indicating that OTUD5 upregulated GPX4 protein levels depending on its DUB activity."

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"Therefore, we explored whether OTUD5 mediates the regulatory effect of THBS1 on GPX4 expression."

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"Conversely, knockdown or knockout of OTUD5 decreased the endogenous GPX4 protein levels in HEK293T, AGS and HGC‐27 cells (Figure 2C–E)."
OTUD5 activates GPX4.
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"Collectively, our data illustrate that 4‐HNE triggers ubiquitin–proteasomal degradation of GPX4 in cardiomyocytes.2.5 OTUD5 Mediates 4-HNE-Induced GPX4 Degradation and Ferroptosis."

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"p53 inhibits OTUD5 transcription to promote GPX4 degradation and induce ferroptosis in gastric cancer."

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"We then asked if OTUD5 could prevent GPX4 degradation."

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"We further investigated whether OTUD5 protected renal tubular cells from H/R-induced ferroptosis by stabilizing GPX4."

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"Conversely, in gastric cancer cells harbouring p53 mutations, OTUD5 transcription is upregulated due to the inactivation of p53, allowing OTUD5 to stabilise GPX4 protein, inhibit ferroptosis and thereby promoting gastric cancer progression (Figure 6J, right panel)."
OTUD5 bound to GPX4 activates GPX4. 1 / 1
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"As OTUD5 belongs to the DUBs family and plays a critical role in controlling the Ub‐dependent degradation of various proteins, 4‐HNE may reduce the interaction between OTUD5 and GPX4 to promote GPX4 ubiquitination and subsequent degradation.To test whether OTUD5 affects the protein stability of GPX4, we co‐transfected GPX4 with Flag‐OTUD5 or its enzyme‐inactive mutant C224S into HEK293T cells."
OTUD5 ubiquitinates GPX4.
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OTUD5 leads to the ubiquitination of GPX4. 4 / 4
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"Silencing OTUD5 increased GPX4 ubiquitination in HBMECs co-cultured with siTHBS1-AMφ-exo (Fig.  xref G)."

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"Additionally, we found that knockdown or depletion of OTUD5 markedly increased GPX4 ubiquitination (Figure  xref )."

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"In addition, OTUD5 rescued the GPX4 de‐ubiquitination phenotype in HEK293T cells, while E3 ligases MIB2, NEDD4L, and STUB1 didn't affect the ubiquitination of GPX4 caused by 4‐HNE (Figure S6A–D, Supporting Information)."

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"Silencing OTUD5 increased GPX4 ubiquitination in HPMECs co-cultured with siGBP2-SMφ-EVs ( xref H)."
OTUD5 inhibits GPX4.
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OTUD5 inhibits GPX4. 2 / 2
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"In conclusion, these results demonstrated that OTUD5 mediates the K48‐linked deubiquitination of GPX4 and stabilises GPX4.2.3 OTUD5 suppresses ferroptosis via GPX4."

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"Overall, this study highlights a new autophagy-dependent ferroptosis module: hypoxia/ischemia-induced OTUD5 autophagy triggers GPX4 degradation, offering a potential therapeutic avenue for I/R-related kidney diseases."
OTUD5 decreases the amount of GPX4.
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"As shown in ubiquitination assays, OTUD5 significantly downregulated the ubiquitination level of GPX4 in the presence of WT ubiquitin and K48 ubiquitin."

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"Importantly, OTUD5 did not decrease the ubiquitination level of GPX4 in K48R‐transfected cells, but the levels decreased in cells transfected with the ubiquitin mutant K63R, indicating that OTUD5 cleaves the K48‐linked ubiquitination of GPX4 rather than K63‐linked chains (Figure 4G)."
OTUD5 is modified
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OTUD5 is phosphorylated.
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OTUD5 is phosphorylated. 10 / 29
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"xref shows the overlaid 1 H- 15 N TROSY spectra of phosphorylated and non-phosphorylated DUBA, which display only minor differences."

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"It is conceivable that also in the DDR, an upstream kinase phosphorylates OTUD5 to amplify DDR signaling through a similar positive feedback mechanism."

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"Unlike other deubiquitinating enzymes, the phosphorylation of OTUD5 at a single residue (Ser177) is required for the activation of the enzyme [ xref ]."

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"For instance, phosphorylation of OTUD5 catalytic domain induces conformational changes thereby promoting substrate binding and facilitating deubiquitination ( xref )."

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"By comparing NMR and enzyme kinetics data among different forms of DUBA with low and high activities, we concluded that a two-state equilibrium that was present only in phosphorylated DUBA is important for DUBA activity."

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"NMR analysis and crystal structures of apo-DUBA in both phosphorylated and unphosphorylated forms and of phospho-DUBA in complex with ubiquitin-aldehyde reveal the role of the phosphate in enzyme acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Consistent with previous NMR studies on the isoform 1 of DUBA ( xref ), the chemical shifts of only a few residues near S177, including G174, Y175 and E178, are perturbed in the phosphorylated DUBA."

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"The phosphorylated DUBA displays essentially the same predicted secondary structure, which is in good agreement with the crystal structures of both non-phosphorylated DUBA in free form (PDB ID: 3PFY) and phosphorylated DUBA covalently linked to ubiquitin aldehyde (PDB ID: 3TMP)."

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"Moreover, phosphorylated OTUD5 was more efficient than wild-type OTUD5 in removing the ubiquitin chain of TRIM25 (Fig.  xref , lane 2 versus lane 3)."

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"Phosphorylation of the human deubiquitinase DUBA (OTUD5) at a single residue, Ser177, is both necessary and sufficient to activate the enzyme [23] ."
OTUD5 is phosphorylated on S177. 7 / 13
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"Phosphorylation of OTUD5 (also known as DUBA) at Ser177 is both necessary and sufficient to activate the enzyme which is a negative regulator of type I interferon [ xref ]."

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"Phosphorylation of OTUD5 at S177 is an absolute requirement for DUB activity because pS177 acts as a lynchpin that locks together two helices that are disordered in the unphosphorylated state."

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"The phosphorylation of OTUD5 at Ser177 is required for the activation of the enzyme."

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"In the OTU family, phosphorylation of DUBA (OTUD5) on S177 results in activation of its enzymatic activity ( xref )."

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"A nuclear magnetic resonance (NMR) analysis revealed that the phosphorylation of Ser177 alone does not induce significant structural changes, and only after binding ubiquitin does Ser177-phosphorylated OTUD5 show significant structural changes that activate its enzymatic activity."

sparser
"Phosphorylation of OTUD5 at Ser177 is critical to the enzyme activity of OTUD526.in our results, the most important of kinase-target networks related primarily to the kinases PLK1, ATM, PIK3CA, CDK5 and CHEK1."

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"In response to TCR stimulation, the protein levels of the Ser177-phosphorylated OTUD5 were found to be upregulated."
OTUD5 is ubiquitinated.
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OTUD5 is ubiquitinated. 7 / 7
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"UBR5 expression promoted both K29- and K48-linked ubiquitylation of OTUD5, which was almost completely diminished by TRIP12 knockdown (Fig.  xref , lanes 2, 4, 6, and 8)."

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"However, the TRIP12-dependent K29-linked ubiquitylation of OTUD5 was partially decreased by UBR5 knockdown, whereas TRIP12-dependent increase of K48-linked chains was massively diminished by UBR5 knockdown (Fig.  xref , lanes 2, 4, 6, and 8)."

sparser
"TRIP12 promotes OTUD5 ubiquitylation and degradation."

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"Next, we investigated whether OTUD5 DUB activity counteracted OTUD5 ubiquitylation by TRIP12 or UBR5."

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"We also confirmed that TRIP12 knockdown decreased ubiquitylation of endogenous OTUD5 under proteasomal inhibition (Fig.  xref )."

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"We subsequently investigated whether TRIP12 promoted ubiquitylation of OTUD5 in cell and in vitro."

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"Using TUBE2 (tandem ubiquitin-binding entity 2), which serves as a pan-ubiquitin chain binder when four repeats of the UBA1 domains from HR23A are tandemly fused xref , we found that OTUD5 pan-ubiquitylation was enhanced by the expression of wild-type, but not catalytically inactive (C1959A), TRIP12 (Fig.  xref )."
OTUD5 is produced.
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OTUD5 is produced. 3 / 3
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"Taken together, we concluded that OTUD5 increased PDCD5 levels via interaction with its UIM motif.Because overexpression of OTUD5 enhanced PDCD5 expression, we next examined if OTUD5 increased the sta[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

trips
"Here, we found that OTU deubiquitinase 5 (OTUD5) was bound to PDCD5 in response to etoposide treatment and increased the stability of PDCD5 by mediating deubiquitination of PDCD5 at Lys-97/98."

reach
"The MDP-NOD2 axis downregulates TLR9-induced ISG by upregulating DUBA expression (67, 68)."
UBR5 affects OTUD5
5 | 21 25
UBR5 binds OTUD5.
5 | 15 20
5 | 15 20

reach
"UBR5-OTUD5 complex mitigates DNA replication fork stress."

reach
"UBR5-OTUD5 complex mitigates transcription-replication conflicts."

reach
"We hypothesized that the OTUD5-UBR5 complex may mitigate TRCs, based on our previous finding that the OTUD5-UBR5 complex regulates DNA double strand break (DSB)-induced transcription arrest, and that OTUD5-UBR5 proteins are present at replication forks (Figure 1G)."

sparser
"UBR5 co-precipitates with OTUD5 (Figure xref ), and the purified recombinant OTUD5 and UBR5 proteins interact in vitro (Figure xref ; xref for western blot), suggesting that they interact directly."

sparser
"PLA detected the interaction between UBR5 and OTUD5 at damaged chromatin in situ , and that the interaction spots partially coincide with the DNA DSB marker 53BP1 (Figure xref ), suggesting that at least a fraction of the interaction indeed occurs at damage (DSB) sites."

sparser
"Previous studies report that OTUD5 interacts with UBR5, a HECT-type K48-linkage specific E3, in the regulation of DNA damage response and NF-κB signaling xref , xref ."

sparser
"Mass spectrometry coupled to short interfering RNA (siRNA) screening of ubiquitinated proteins in Duba −/− Th17 cells revealed that DUBA directly bound and deubiquitinated UBR5 (Ubiquitin protein ligase E3 component n-recognin 5)."

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC

sparser
"These results unequivocally suggest that both the UBR5OTUD5 interaction and OTUD5–SPT16 interaction are necessary for transcriptional repression at the damaged chromatin."

reach
"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
UBR5 ubiquitinates OTUD5.
| 3 5
UBR5 ubiquitinates OTUD5. 8 / 8
| 3 5

reach
"UBR5-dependent ubiquitylation of OTUD5 was almost completely diminished with TRIP12 knockdown (Fig. 2g, lanes 2 and 4), whereas TRIP12-dependent ubiquitylation was partially decreased by UBR5 knockdown (Fig. 2h, lanes 2 and 4, and supplementary Fig. 1d)."

sparser
"UBR5-dependent ubiquitylation of OTUD5 was almost completely diminished with TRIP12 knockdown (Fig.  xref , lanes 2 and 4), whereas TRIP12-dependent ubiquitylation was partially decreased by UBR5 knockdown (Fig.  xref , lanes 2 and 4, and supplementary Fig.  xref )."

reach
"Notably, UBR5 ubiquitylation of wild-type OTUD5 was less effective than that of C224R OTUD5 (Fig. 5b (i))."

sparser
"The data on a catalytically-inactive OTUD5 (Fig.  xref ) suggest that the ubiquitylation of OTUD5 by TRIP12 and UBR5 appears as if they act sequentially because OTUD5 DUB activity negates UBR5 activity, which is different from typical chain initiation (priming)-elongation model xref , xref ."

sparser
"Since UBR5 can stably associate with OTUD5 xref , UBR5 may transiently ubiquitylate OTUD5 with K48-linked chains."

reach
"The data on a catalytically-inactive OTUD5 (Fig. 5) suggest that the ubiquitylation of OTUD5 by TRIP12 and UBR5 appears as if they act sequentially because OTUD5 DUB activity negates UBR5 activity, which is different from typical chain initiation (priming)-elongation model ."

sparser
"Next, we investigated whether OTUD5 DUB activity counteracted OTUD5 ubiquitylation by TRIP12 or UBR5."

sparser
"Notably, UBR5 ubiquitylation of wild-type OTUD5 was less effective than that of C224R OTUD5 (Fig.  xref (i))."
UBR5 activates OTUD5.
| 2
UBR5 activates OTUD5. 2 / 2
| 2

reach
"UBR5 destabilizes DUBA through ubiquitylation, whereas DUBA stabilizes UBR5 in activated T cells by attenuating degradative auto-ubiquitylation, triggering UBR5 mediated ubiquitylation of the transcription factor RORyt in response to TGF-beta [XREF_BIBR]."

reach
"Considering other potential post-transcriptional mechanisms, we investigated the ubiquitin ligase UBR5, which has been reported to mediate OTUD5 abundance post-transcriptionally ."
UBR5 inhibits OTUD5.
| 1
UBR5 bound to OTUD5 inhibits OTUD5. 1 / 1
| 1

reach
"DUBA interacted with the ubiquitin ligase UBR5, which suppressed DUBA abundance in naive T cells."
OTUD5 affects TRAF3
2 2 1 1 | 28 14
OTUD5 binds TRAF3.
2 1 | 8 12
2 1 | 7 12

sparser
"TRAF3-DUBA interaction."

sparser
"xref shows the TRAF3-DUBA interaction model that we obtained by PRISM xref , xref , xref , in which the concave crevice on TRAF-C of TRAF3 is in contact with DUBA."

reach
"DUBA interacts with and de-ubiquitinates TRAF3, thereby attenuating TLR dependent and TLR independent antiviral responses XREF_BIBR."

sparser
"DUBA physically binds with TRAF3 that is an E3 ubiquitin ligase and could assemble lysine-63–linked polyubiquitin chains [ 8 ]."

reach
"Interestingly, DUBA physically interacts with TRAF3 and inhibits the self ubiquitylation of TRAF3 (XREF_FIG)."

sparser
"Interaction of endogenous TRAF3 and DUBA takes place upon TLR3 stimulation ( 70 )."

sparser
"There are no clinically observed oncogenic mutations on DUBA that fall onto DUBA-TRAF3 interface residues or nonsense/frameshift mutations."

sparser
"In fact, DUBA physically interacts with TRAF3 and inhibits the self-ubiquitination of TRAF3."

sparser
"The interaction between DUBA and TRAF3 occurs during TLR3 activation [197] ."

sparser
"The interaction between DUBA and TRAF3 occurs during TLR3 activation [197] ."
| DOI
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

reach
"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
OTUD5 deubiquitinates TRAF3.
1 1 | 12
OTUD5 deubiquitinates TRAF3. 10 / 14
1 1 | 12

reach
"However, it is uncertain whether DUBA also deubiquitinates TRAF3 in rainbow trout through a similar mechanism."

reach
"DUBA interacts with and de-ubiquitinates TRAF3, thereby attenuating TLR dependent and TLR independent antiviral responses XREF_BIBR."

reach
"For example, the deubiquitinase DUBA reverses TRAF3 ubiquitination, disconnecting TRAF3 from its substrates TBK1 and IKKepsilon XREF_BIBR."

reach
"Furthermore, OTUD5 deubiquitylates TRAF3 resulting in diminished type I interferon and interleukin-10 responses [XREF_BIBR]."

reach
"Indeed, the blockade of DUBA-mediated deubiquitination of TRAF3 by DUBA-specific siRNA cancelled the negative regulatory effects of NOD2 on DSS-induced colitis in mice treated with CpG, and mice treated with DUBA-siRNA displayed severe DSS-induced colitis even after repeated MDP injections (67, 68)."

reach
"Two close relatives of DUBA, OTUB1 and OTUB2, were shown to reduce TRAF3 ubiquitination coupled with decreased anti-viral signaling responses in Sendai virus infected HEK 293T cells."

reach
"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."

reach
"TRIAD3A dependent TRAF3 proteasomal destruction, like TRAF3 de-ubiquitination by DUBA, halts the activation of TBK1 and IKKepsilon and subsequent IFN production."

reach
"DUBA deubiquitinates TRAF3 to down-regulate its interaction with TBK1 and consequent activation of IRF-3."

reach
"In fact, DUBA physically interacts with TRAF3 and inhibits the self-ubiquitination of TRAF3."
OTUD5 inhibits TRAF3.
1 | 5 2
OTUD5 inhibits TRAF3. 7 / 8
1 | 5 2

sparser
"231 DUBA also targets and inhibits TRAF3 by cleaving its K63-linked polyubiquitin chains."

reach
"In the study showing that OTUD5 suppresses the type I interferon (IFN-1) production, the proposed mechanism is that OTUD5 antagonizes the TRAF3 signaling (21)."

reach
"TRAF3 activation is negatively regulated by SOCS3 and DUBA."

reach
"TRAF3 activation is inhibited by SOCS3 and DUBA and TRAF 6 is targeted by numerous inhibitors including A20, USP4, CYLD, and others [73,74]."
| PMC

sparser
"TRAF3 are inhibited by DUBA, OTUB1 and UCHL1 that specilally remove TRAF3 K63‐linked ubiquitination."

reach
"Upregulated miR-210 could inhibit the expression of its target gene DUBA that is as the inhibitor of TRAF3 ( Fig. 5 )."

reach
"TRAF3 are inhibited by DUBA, OTUB1 and UCHL1 that specilally remove TRAF3 K63‐linked ubiquitination."
OTUD5 ubiquitinates TRAF3.
| 2
OTUD5 leads to the ubiquitination of TRAF3. 2 / 2
| 2

reach
"Other DUBs that target the RIG-I and MDA5 pathway include CYLD and DUBA which target RIG-I and TRAF3 ubiquitination respectively (XREF_FIG)."

reach
"In addition to USP21, A20, and CYLD, DUBA has been shown to down-regulate antiviral signaling by deubiquitinating TRAF3."
OTUD5 activates TRAF3.
| 1
OTUD5 activates TRAF3. 1 / 1
| 1

reach
"Moreover, the adaptor proteins TRAF3 and TRAF6 are targeted by the cellular proteins DUBA and A20 XREF_BIBR, XREF_BIBR, and TBK1 is sequestered by SIKE XREF_BIBR."
OTUD5 affects UBR5
5 1 | 17 20
OTUD5 binds UBR5.
5 | 15 20
5 | 15 20

reach
"UBR5-OTUD5 complex mitigates DNA replication fork stress."

reach
"UBR5-OTUD5 complex mitigates transcription-replication conflicts."

reach
"We hypothesized that the OTUD5-UBR5 complex may mitigate TRCs, based on our previous finding that the OTUD5-UBR5 complex regulates DNA double strand break (DSB)-induced transcription arrest, and that OTUD5-UBR5 proteins are present at replication forks (Figure 1G)."

sparser
"UBR5 co-precipitates with OTUD5 (Figure xref ), and the purified recombinant OTUD5 and UBR5 proteins interact in vitro (Figure xref ; xref for western blot), suggesting that they interact directly."

sparser
"PLA detected the interaction between UBR5 and OTUD5 at damaged chromatin in situ , and that the interaction spots partially coincide with the DNA DSB marker 53BP1 (Figure xref ), suggesting that at least a fraction of the interaction indeed occurs at damage (DSB) sites."

sparser
"Previous studies report that OTUD5 interacts with UBR5, a HECT-type K48-linkage specific E3, in the regulation of DNA damage response and NF-κB signaling xref , xref ."

sparser
"Mass spectrometry coupled to short interfering RNA (siRNA) screening of ubiquitinated proteins in Duba −/− Th17 cells revealed that DUBA directly bound and deubiquitinated UBR5 (Ubiquitin protein ligase E3 component n-recognin 5)."

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC

sparser
"These results unequivocally suggest that both the UBR5OTUD5 interaction and OTUD5–SPT16 interaction are necessary for transcriptional repression at the damaged chromatin."

reach
"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
OTUD5 deubiquitinates UBR5.
1 | 1
OTUD5 deubiquitinates UBR5. 1 / 2
1 | 1

reach
"When DUBA deubiquitylates UBR5 and rescues it from degradation, UBR5 is free to polyubiquitylate RORgammaT, driving its degradation and consequently limiting Th17 differentiation."
OTUD5 inhibits UBR5.
| 1
OTUD5 inhibits UBR5. 1 / 1
| 1

reach
"These properties of the three enzymes constitute a complex “rock–paper–scissors”-like interplay.OTUD5 protects UBR5 from self-ubiquitylation , suggesting that OTUD5 antagonizes UBR5-mediated K48 ubiquitylation."
TP53 affects OTUD5
9 | 14 13
TP53 binds OTUD5.
9 | 9 13
9 | 9 13

sparser
"Taken together, these results provide clinical evidence for p53OTUD5 axis‐mediated regulation of GPX4 protein stability, suggesting that the expression level of OTUD5 is important for ferroptosis of gastric cancer cells."

reach
"A recent study showed that in case of stress, programmed cell death 5 (PDCD5) is required to promote the interaction between OTUD5 and p53."

reach
"Since OTUD5 was shown to interact with p53, we also performed mapping analysis of the interaction between OTUD5 and p53."

reach
"However, upon prolonged genotoxic stress, the interaction between OTUD5 and PDCD5 was reduced, while the association between OTUD5 and p53 gradually increased, leading to p53 induction ( Fig. 4A )."

sparser
"OTUD5 can bind to TP53 and regulate its ubiquitination levels [ xref ]."

sparser
"A recent study showed that in case of stress, programmed cell death 5 (PDCD5) is required to promote the interaction between OTUD5 and p53."

reach
"OTUD5 can bind to TP53 and regulate its ubiquitination levels [56]."

sparser
"Since OTUD5 was shown to interact with p53, we also performed mapping analysis of the interaction between OTUD5 and p53."

sparser
"OTUD5 forms a direct complex with p53 and controls level of ubiquitination."

sparser
"P53 interacts with OTUD5."
TP53 increases the amount of OTUD5.
| 2
TP53 increases the amount of OTUD5. 2 / 2
| 2

reach
"P53 inhibits OTUD5 transcription to promote GPX4 degradation and induce ferroptosis in gastric cancer."

reach
"Collectively, these findings elucidate a mechanism whereby p53 inactivation upregulates OTUD5 transcription to deubiquitylate and stablize GPX4, resulting in ferroptosis inhibition and gastric cancer progression."
TP53 inhibits OTUD5.
| 1
TP53 inhibits OTUD5. 1 / 1
| 1

reach
"Furthermore, we investigated the specific mechanism by which p53 regulates OTUD5 and GPX4 in gastric cancer and found that p53 transcriptionally suppresses OTUD5, thereby regulating ferroptosis in gastric cancer cells."
TP53 decreases the amount of OTUD5.
| 1
TP53 decreases the amount of OTUD5. 1 / 1
| 1

reach
"P53 inhibits OTUD5 transcription to promote GPX4 degradation and induce ferroptosis in gastric cancer."
TP53 activates OTUD5.
| 1
TP53 activates OTUD5. 1 / 1
| 1

reach
"To determine whether p53 directly regulates OTUD5, ChIP‐qPCR experiment was performed and results showed that p53 could bind to the OTUD5 promoter region (Figure S3I)."
PDCD5 affects OTUD5
5 | 1 13 15
PDCD5 binds OTUD5.
5 | 8 15
5 | 8 15

reach
"Therefore, dysregulation of p53-interacting deubiquitinating enzymes may lead to aberrant regulation of genotoxic responses in cancer cells.To understand how the tumor suppressor PDCD5 is regulated du[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, knocking-down of OTUD5 markedly diminished the association of PDCD5 with p53, indicating that OTUD5 binds to PDCD5 and mediates PDCD5-p53 network during genotoxic stress responses ( Fig. 1B )[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Thus, we next explored the possibility that OTUD5 initially binds to and stabilizes PDCD5 and subsequently stabilizes p53 in response to genotoxic stress."

reach
"However, upon prolonged genotoxic stress, the interaction between OTUD5 and PDCD5 was reduced, while the association between OTUD5 and p53 gradually increased, leading to p53 induction ( Fig. 4A )."

reach
"We corroborated these findings by a reconstitution experiment where a point mutant defective for interaction with OTUD5 (PDCD5 E94D ) showed that OTUD5 interaction with PDCD5 is a prerequisite for p53[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Park et al ( xref ) identified the programmed cell death 5-OTUD5 network as a central hub for regulating p53-mediated apoptosis."

sparser
"Under genotoxic stress, OTUD5 binds to PDCD5 and increases PDCD5 stability by mediating the deubiquitination of PDCD5 [ xref ]."

sparser
"Park et al. xref further revealed that deubiquitinase OTUD5 directly binds to PDCD5 and promotes its accumulation by mediating PDCD5 deubiquitination at Lys‐97/98 upon genotoxic stress‐induced apoptosis."

sparser
"To understand how the tumor suppressor PDCD5 is regulated during apoptosis, we employed yeast two-hybrid assays to identify interacting partners of PDCD5 and found that OTUD5 bound to PDCD5 in respons[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Park and colleagues [17] found that OTUD5 binds to PDCD5 in response to etoposide treatment via its ubiquitin interacting motif (UIM)."
PDCD5 activates OTUD5.
| 1 4
PDCD5 activates OTUD5. 4 / 4
| 1 3

reach
"Conversely, PDCD5 knockdown greatly attenuated the effect of OTUD5 on TP53 activation."

eidos
"Therefore , it was concluded that OTUD5 could impede the proliferation , migration , and invasion of HCC827 cells while facilitating their apoptosis , and the role of OTUD5 was partly mediated by PDCD5 ."
| PMC

reach
"Although OTUD5 enhanced ET-induced PDCD5 stabilization and p53 activation, genetic depletion of PDCD5 reversed the positive action of OTUD5 on PDCD5 stabilization ( Fig. 5A )."

reach
"Conversely, reintroduction of PDCD5 in PDCD5 −/− MEF cells dramatically restored OTUD5 WT function in promoting p53 activation."
PDCD5-E16D activates OTUD5. 1 / 1
| 1

reach
"Importantly, ectopic expression of PDCD5 WT in PDCD5 −/− MEF cells led to substantial genotoxic stress-induced DNA damage and apoptosis; however, expression of neither PDCD5 E16D nor PDCD5 E94D induce[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
PDCD5 decreases the amount of OTUD5.
| 1
PDCD5 decreases the amount of OTUD5. 1 / 1
| 1

reach
"Consequently, knocking-down PDCD5 abrogates the effect of OTUD5 overexpression on p53 stabilization, highlighting the critical role of PDCD5 in OTUD5-mediated p53 activation ( Fig. 5E )."
TRIP12 affects OTUD5
| 14 16
TRIP12 binds OTUD5.
| 2 12
| 2 12

sparser
"First, whether TRIP12 constitutively associates with OTUD5 is unknown."

sparser
"TRIP12OTUD5 antagonism regulates TNF-α–induced NF-κB signaling."

sparser
"We next verified TRIP12OTUD5 antagonism in TNF-α–induced NF-κB signaling."

sparser
"The data presented thus far indicate that OTUD5 is cooperatively modified by K29-specific TRIP12 and K48-specific UBR5, resulting in the conjugation of K29/K48-linked branched ubiquitin chains, and the biological importance of TRIP12OTUD5 antagonism is observed in TNF-α–induced NF-κB signaling."

sparser
"Collectively, these results indicate that TRIP12 interacts with OTUD5 and promotes its ubiquitylation and degradation."

sparser
"The biological importance of the TRIP12OTUD5 interplay was demonstrated in the regulation of NF-κB signaling."

sparser
"Because the simultaneous knockdown of OTUD5 partially restores NF-κB signaling under TRIP12 knockdown, the TRIP12OTUD5 antagonism may account for the regulatory function of TRIP12 in NF-κB signaling."

sparser
"This equilibrium favors ubiquitin chain assembly and subsequent OTUD5 degradation when TRIP12 interacts with OTUD5; UBR5 assembles K48-linked ubiquitin chains onto previously formed K29-linked chains more efficiently than de novo ubiquitylation (Fig.  xref ), presumably through ubiquitin interaction of the UBA domain (Fig.  xref and ref. xref )."

sparser
"TRIP12 interacted with OTUD5 during co-immunoprecipitation (Fig.  xref )."

sparser
"Whether the TRIP12OTUD5 axis also regulates other pathways should be studied further."
TRIP12 ubiquitinates OTUD5.
| 8 4
TRIP12 ubiquitinates OTUD5. 10 / 11
| 7 4

reach
"TRIP12 promotes OTUD5 ubiquitylation and degradation."

reach
"We subsequently investigated whether TRIP12 promoted ubiquitylation of OTUD5 in cell and in vitro."

reach
"Using TUBE2 (tandem ubiquitin-binding entity 2), which serves as a pan-ubiquitin chain binder when four repeats of the UBA1 domains from HR23A are tandemly fused , we found that OTUD5 pan-ubiquitylation was enhanced by the expression of wild-type, but not catalytically inactive (C1959A), TRIP12 (Fig. 1f)."

reach
"We also confirmed that TRIP12 knockdown decreased ubiquitylation of endogenous OTUD5 under proteasomal inhibition (Fig. 1g)."

reach
"In an in vitro ubiquitylation assay using purified OTUD5 and TRIP12 proteins (supplementary Fig. 1c), OTUD5 was ubiquitylated by TRIP12 (Fig. 1h)."

sparser
"However, wild-type and C224R OTUD5 were ubiquitylated by TRIP12 at similar levels (Fig.  xref (ii))."

sparser
"The data on a catalytically-inactive OTUD5 (Fig.  xref ) suggest that the ubiquitylation of OTUD5 by TRIP12 and UBR5 appears as if they act sequentially because OTUD5 DUB activity negates UBR5 activity, which is different from typical chain initiation (priming)-elongation model xref , xref ."

reach
"UBR5 expression promoted both K29- and K48-linked ubiquitylation of OTUD5, which was almost completely diminished by TRIP12 knockdown (Fig. 2i, lanes 2, 4, 6, and 8)."

reach
"The data on a catalytically-inactive OTUD5 (Fig. 5) suggest that the ubiquitylation of OTUD5 by TRIP12 and UBR5 appears as if they act sequentially because OTUD5 DUB activity negates UBR5 activity, which is different from typical chain initiation (priming)-elongation model ."

sparser
"Next, we investigated whether OTUD5 DUB activity counteracted OTUD5 ubiquitylation by TRIP12 or UBR5."
TRIP12 ubiquitinates OTUD5-C224R. 1 / 1
| 1

reach
"However, wild-type and C224R OTUD5 were ubiquitylated by TRIP12 at similar levels (Fig. 5b (ii))."
TRIP12 inhibits OTUD5.
| 2
TRIP12 inhibits OTUD5. 1 / 1
| 1

reach
"We investigated whether TRIP12 antagonizes OTUD5 cellular function."
TRIP12 bound to OTUD5 inhibits OTUD5. 1 / 1
| 1

reach
"Because UBR5 appears to stably associate with OTUD5 , and because TRIP12 possesses phosphorylation sites (https://www.phosphosite.org), we presume that TRIP12 may associate with OTUD5 under certain cellular conditions, triggering rapid OTUD5 degradation to fulfill cellular demand.Second, our model suggests complex ubiquitin architectures consisting of K29 and K48 linkages."
TRIP12 activates OTUD5.
| 2
TRIP12 activates OTUD5. 2 / 2
| 2

reach
"TRIP12 knockdown massively slowed OTUD5 turnover (Fig. 1d)."

reach
"As shown in Fig. 2c, OTUD5 modification with K29-linked chains was massively increased by wild-type TRIP12 but not by catalytically inactive TRIP12 (Fig. 2c)."
TRAF3 affects OTUD5
2 1 | 8 12
2 1 | 7 12

sparser
"TRAF3-DUBA interaction."

sparser
"xref shows the TRAF3-DUBA interaction model that we obtained by PRISM xref , xref , xref , in which the concave crevice on TRAF-C of TRAF3 is in contact with DUBA."

reach
"DUBA interacts with and de-ubiquitinates TRAF3, thereby attenuating TLR dependent and TLR independent antiviral responses XREF_BIBR."

sparser
"DUBA physically binds with TRAF3 that is an E3 ubiquitin ligase and could assemble lysine-63–linked polyubiquitin chains [ 8 ]."

reach
"Interestingly, DUBA physically interacts with TRAF3 and inhibits the self ubiquitylation of TRAF3 (XREF_FIG)."

sparser
"Interaction of endogenous TRAF3 and DUBA takes place upon TLR3 stimulation ( 70 )."

sparser
"There are no clinically observed oncogenic mutations on DUBA that fall onto DUBA-TRAF3 interface residues or nonsense/frameshift mutations."

sparser
"In fact, DUBA physically interacts with TRAF3 and inhibits the self-ubiquitination of TRAF3."

sparser
"The interaction between DUBA and TRAF3 occurs during TLR3 activation [197] ."

sparser
"The interaction between DUBA and TRAF3 occurs during TLR3 activation [197] ."
| DOI
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

reach
"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
OTUD5 affects SYNM
| 24
OTUD5 activates SYNM.
| 11
OTUD5 activates SYNM. 10 / 11
| 11

reach
"Furthermore, confocal microscopic analysis visualized that the co‐localization of α‐Syn and LAMP1, a marker of lysosomes, was significantly enhanced by OTUD5 overexpression (Figure 2B)."

reach
"In contrast, a pronounced reduction in TH levels induced by α‐Syn PFF was rescued by OTUD5 overexpression in SH‐SY5Y cells (Figure S13B Supporting Information)."

reach
"35 Interestingly, the degradation of endogenous α‐Syn induced by OTUD5 was evidently blocked by CQ, but not MG132 or 3‐MA (Figure  2A)."

reach
"We then defined the type of polyubiquitin chains attached to α‐Syn mediated by OTUD5."

reach
"19 , 41 Therefore, we investigated whether the degradation of α‐Syn mediated by OTUD5 relied on NEDD4."

reach
"Firstly, we transfected siRNA specific to human NEDD4 into SH‐SY5Y cells and found that NEDD4 knockdown mitigated the degradation of α‐Syn induced by OTUD5 overexpression (Figure  4A)."

reach
"In contrast, OTUD5 knockout in HEK293T cells impaired the binding of NEDD4 and α‐Syn (Figure 4H)."

reach
"42 To validate our hypothesis that OTUD5 may recruit NEDD4 to degrade α‐Syn, we first demonstrated that OTUD5 failed to enhance the degradation of α‐Syn when NEDD4 was knockdown, while NEDD4 overexpression promoted the degradation of α‐Syn induced by OTUD5, which indicated that OTUD5 mediated α‐Syn degradation relying on NEDD4."

reach
"Our results showed that the degradation of α‐Syn induced by OTUD5 changed rarely in ATG5 knockout cells (Figure 2C)."

reach
"These data indicated that the degradation of α‐Syn induced by OTUD5 may not occur through autophagy‐lysosomal pathway."
OTUD5 ubiquitinates SYNM.
| 5
OTUD5 leads to the ubiquitination of SYNM. 5 / 5
| 5

reach
"Meanwhile, OTUD5 knockout greatly decreased polyubiquitination of α‐Syn (Figure 3H)."

reach
"Based on our finding that OTUD5 promoted α‐Syn polyubiquitination independent of its enzymatic activity, we hypothesized that OTUD5 may serve as an adaptor protein to recruit an E3 ligase for K63‐linked polyubiquitination and degradation of α‐Syn."

reach
"Furthermore, OTUD5 deficiency further attenuated the endogenous K63‐linked polyubiquitination of α‐Syn induced by α‐Syn PFF (Figure 7D)."

reach
"Particularly, OTUD5 selectively promoted K63‐linked polyubiquitination of α‐Syn and mediated its endolysosomal degradation by recruiting the E3 ligase NEDD4."

reach
"Mechanically, we found that OTUD5 deficiency further reduced endogenous K63‐linked polyubiquitination of α‐Syn and upregulated its protein level, then promoted the formation of pathologic p‐α‐Syn and aggravated the pathogenic consequences induced by α‐Syn PFF.In summary, we demonstrated that OTUD5 negatively regulated α‐Syn protein levels by promoting endolysosomal degradation independent of its deubiquitinate activity."
OTUD5 inhibits SYNM.
| 4
OTUD5 inhibits SYNM. 4 / 4
| 4

reach
"As there are physiological functions for α‐Syn, we investigated whether OTUD5 overexpression and knockout can perturb the cellular steady state by regulating α‐Syn."

reach
"Besides, double‐immunofluorescence staining showed that OTUD5 in DA neurons of SN was significantly downregulated after α‐Syn PFF injection (Figure 5E)."

reach
"Immunofluorescence also showed that OTUD5 deficiency upregulated the formation of pathologic p‐α‐Syn (Ser129) in TH‐positive neurons in SN (Figure 7B) and STR (Figure 7C)."

reach
"Notably, OTUD5 conditional deletion further increased pathological p‐α‐Syn induced by α‐Syn PFF (Figure  7A)."
OTUD5 decreases the amount of SYNM.
| 3
OTUD5 decreases the amount of SYNM. 3 / 3
| 3

reach
"Mechanically, we found that OTUD5 deficiency further reduced endogenous K63‐linked polyubiquitination of α‐Syn and upregulated its protein level, then promoted the formation of pathologic p‐α‐Syn and aggravated the pathogenic consequences induced by α‐Syn PFF.In summary, we demonstrated that OTUD5 negatively regulated α‐Syn protein levels by promoting endolysosomal degradation independent of its deubiquitinate activity."

reach
"Consistently, knockdown of OTUD5 significantly enhanced α‐Syn protein expression in SH‐SY5Y cells (Figure 1C), while the mRNA levels remained unaffected (Figure 1D)."

reach
"Additionally, OTUD5 deficiency significantly enhanced α‐Syn protein levels in primary cultured midbrain neurons (Figure 1E)."
OTUD5 binds SYNM.
| 1
| 1

reach
"Besides, an association between endogenous OTUD5 and α‐Syn was evident in SH‐SY5Y cells (Figure 3D,E)."
OTUD5 affects SUPT16H
5 | 9 10
OTUD5 binds SUPT16H.
5 | 4 10
5 | 4 10

reach
"To study the cellular consequence of specifically blocking the interaction between OTUD5 and FACT (OTUD5 overexpression or complete knockout causes aberration in cell cycle and apoptosis), we generated a HeLa knock-in (KI) cell line in which the D537A mutation is introduced into the genomic loci of OTUD5 (OTUD5 ) using the CRISPR-Cas9 method (Supplementary Figure S2; sequencing information of 4 clones)."

reach
"Thus, blocking the interaction between FACT and OTUD5 enhances FACT nucleosome remodeling function.The OTUD5 cells display a noticeable increase in micronuclei and cytoplasmic bridges (Figure 3E), which can form as a consequence of replication stress."

sparser
"In addition, OTUD5 can interact with the SPT16 protein in the FACT complex and antagonize the deposition of histone H2A upon DSB damage by acting as a scaffold [ xref , xref ] ( xref D)."

sparser
"OTUD5 interacts with SPT16 and regulates the SPT16-dependent Pol II elongation at damaged chromatin."

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"Further, the interaction between OTUD5 and SPT16 can be detected in situ by the PLA (Figure xref )."

sparser
"These results unequivocally suggest that both the UBR5–OTUD5 interaction and OTUD5SPT16 interaction are necessary for transcriptional repression at the damaged chromatin."

sparser
"Independently of UBR5, OTUD5 also interacts with SPT16, through the UIM domain at the C-terminal tail."

reach
"These results confirm that the interaction between OTUD5 and FACT is important for the transcriptional regulation, and provide a new possible link between the FACT-dependent transcriptional process and tumor suppression."

reach
"Further investigation into the nature of the interaction between OTUD5 and FACT will provide more insight into the regulation."
OTUD5 inhibits SUPT16H.
| 5
| 5

reach
"Based on this and our results, we hypothesized that UBR5 and OTUD5 may temporarily antagonize the FACT activity to regulate the timely stalling and subsequent resumption of Pol II at the damage sites."

reach
"The FACT-limiting activity of OTUD5 is, at least in part, due to the recruitment and stabilization of the HDAC1 and HDAC2 proteins."

reach
"We have previously shown that the FACT histone chaperone, necessary for remodeling nucleosomes to allow for transcription, is negatively regulated by OTUD5 upon nuclease induction of DSBs (37)."

reach
"Thus, our work propose that FACT is an important factor in this in cis regulation, and that FACT is subject to regulation by OTUD5–UBR5 complex for the non-lesion stalling of Pol II.Our work leaves several open-ended questions; our data collectively suggest that OTUD5 and UBR5 repress FACT histone exchange activity, but how does this complex exactly achieve this?"

reach
"It is possible that OTUD5 prevents the FACT engagement on the histones or DNA, by directly binding to FACT."
OTUD5 affects GBP2
| 9 14
| 9 14

sparser
"However, the interaction between GBP2 and OTUD5 in HPMECs was partially inhibited by PD ( xref H)."

sparser
"This reduces GBP2's interaction with OTUD5 and its inhibitory effect on OTUD5."

sparser
"Furthermore, the precise mechanism by which PD affects the GBP2-OTUD5 interaction remains unclear and requires further investigation."

sparser
"Second, although this study confirmed the regulatory interaction between GBP2 and OTUD5 in endothelial cells, the possibility that GBP2 modulates ferroptosis through interactions with endothelial cell receptors cannot be entirely excluded."

sparser
"The mechanism by which GBP2 binds directly to OTUD5 and promotes GPX4 ubiquitination was elucidated using RNA interference, adeno-associated virus transfection, and endothelial-specific Gpx4 knockout mice."

sparser
"PD treatment inhibited the interaction between GBP2 and OTUD5, leading to a reduction in GPX4 ubiquitination."

sparser
"We further found that PD treatment inhibits the interaction between GBP2 and OTUD5 in HPMECs, reducing GPX4 ubiquitination."

reach
"Protein-protein docking analysis was performed to predicted the interaction between GBP2 and OTUD5, which revealed a binding energy of −67.42 kcal/mol, indicating a significant probability of interaction (Fig. 8D)."

sparser
"Mechanistically, GBP2 interacts with OTU deubiquitinase 5 (OTUD5) in endothelial cells, promoting the ubiquitination of GPX4, thereby increasing sensitivity to ferroptosis and compromising the endothelial barrier."

sparser
"GBP2 and OTUD5 interaction promotes the ubiquitination and degradation of GPX4 in HPMECs."
OTUD5 affects SLC38A1
| 17 5
OTUD5 binds SLC38A1.
| 4 5
| 4 5

sparser
"Our further experiments verified the interaction between OTUD5 and SLC38A1."

reach
"Coimmunoprecipitation analysis confirmed the interaction between OTUD5 and SLC38A1."

reach
"OTUD5 binds to SLC38A1."

reach
"Co-IP assays confirmed that both exogenous and endogenous OTUD5 bound to SLC38A1 (Fig. 5B and C)."

reach
"Our further experiments verified the interaction between OTUD5 and SLC38A1."

sparser
"Coimmunoprecipitation analysis confirmed the interaction between OTUD5 and SLC38A1."

sparser
"Finally, the interaction between OTUD5 and solute carrier family 38 member 1 (SLC38A1) and the underlying regulatory mechanism were investigated."

sparser
"OTUD5 binds to SLC38A1."

sparser
"Co-IP assays confirmed that both exogenous and endogenous OTUD5 bound to SLC38A1 (Fig.  xref B and C)."
OTUD5 deubiquitinates SLC38A1.
| 5
OTUD5 deubiquitinates SLC38A1. 5 / 5
| 5

reach
"OTUD5 promotes the growth of hepatocellular carcinoma by deubiquitinating and stabilizing SLC38A1."

reach
"Our results suggest that OTUD5 is an oncogene in HCC that controls the growth of cancer cells by deubiquitinating and stabilizing SLC38A1."

reach
"Next, we determined whether OTUD5 prevents the ubiquitination of SLC38A1."

reach
"Therefore, these results support that OTUD5 deubiquitinates and stabilizes SLC38A1 in HCC."

reach
"SLC38A1 is deubiquitinated and stabilized by OTUD5 and mediates the tumour-promoting role of OTUD5 in HCC."
OTUD5 activates SLC38A1.
| 4
OTUD5 activates SLC38A1. 4 / 4
| 4

reach
"Therefore, our results suggest that OTUD5 promotes HCC cell proliferation by increasing the stability of SLC38A1."

reach
"We found that OTUD5 knockdown led to faster SLC38A1 degradation and that OTUD5 overexpression prolonged the half-life of SLC38A1 in HCC cells (P < 0.05, Fig. 6B and C and Supplementary Fig. 4B and 4 C)."

reach
"OTUD5 knockdown significantly reduced the IHC score of SLC38A1 in xenograft tumour tissues (P < 0.05, Supplementary Fig. 2)."

reach
"The SLC38A1 protein was the most strongly downregulated by OTUD5 knockdown (Fig. 5A and Supplementary Table 2)."
OTUD5 increases the amount of SLC38A1.
| 3
OTUD5 increases the amount of SLC38A1. 3 / 3
| 3

reach
"The results showed that SLC38A1 protein expression was markedly decreased by OTUD5 knockdown (P < 0.05, Fig. 5D)."

reach
"OTUD5 knockdown reduced and OTUD5 overexpression increased SLC38A1 protein levels in HCC cells."

reach
"We subsequently found that the protein level of SLC38A1 was increased by OTUD5 overexpression (Fig. 5E)."
OTUD5 decreases the amount of SLC38A1.
| 1
OTUD5 decreases the amount of SLC38A1. 1 / 1
| 1

reach
"To determine whether the reduction in SLC38A1 expression induced by OTUD5 knockdown is dependent on the proteasomal degradation pathway, MG132 was used to block proteasome activity."
OTUD5 affects DBC1
| 21
OTUD5 binds DBC1. 10 / 21
| 21

sparser
"Lots of interaction assays suggested that the potential DUB OTUD5 might interact with DBC1 ( xref ) and regulate its ubiquitination ( xref )."

sparser
"Co-IP analysis demonstrated the exogenous interaction between OTUD5 and DBC1 ( xref )."

sparser
"Furthermore, we confirmed that endogenous DBC1 could directly interact with OTUD5 ( xref , xref ), which was further verified by an in vitro pull-down assay using purified His-tagged DBC1 to pull down OTUD5 from cell lysates ( xref , xref )."

sparser
"Interestingly, we found that either hypoxia treatment or overexpression of SIAH2 promoted the interaction between DBC1 and SIAH2, but inhibited the interaction of DBC1 with OTUD5 ( xref , xref )."

sparser
"Moreover, we still need to further investigate the details of how the OTUD5-DBC1 complex dissociates under hypoxia to modulate DBC1 stability."

sparser
"Figure 4: the authors should study OTUD5 binding to DBC1 under hypoxia compared with normoxia to substantiate the claim for preferred interaction under normoxia."

sparser
"The authors propose that in normoxia, OTUD5 is bound to DBC1 which is displaced by SIAH2 upon activation by hypoxia, however despite the comprehensive study of the interaction between DBC1 this is not fully supported by the data."

sparser
"1) The authors propose a hypothesis that OTUD5 is bound to DBC1 in normoxic conditions which is displaced upon activation of SIAH2."

sparser
"Does OTUD5 bind to non-ubiquitinated DBC1?"

sparser
"Direct binding of recombinant DBC1 and OTUD5 could confirm this."
GPX4 affects OTUD5
1 | 13 7
GPX4 binds OTUD5.
1 | 12 7
1 | 12 7

reach
"Next, we determined the interaction of OTUD5 and GPX4 by co‐immunoprecipitation and mass spectrometry (Figure  4A; Table S2, Supporting Information)."

reach
"The direct interaction of OTUD5 and GPX4 was further confirmed by GST‐pull down, surface plasmon resonance (SPR) and microscale thermophoresis (MST) (Figure 4B,C; Figure S5E, Supporting Information), and the colocalization of OTUD5 and GPX4 proteins in the NRCMs was verified by immunofluorescence (Figure 4D)."

reach
"As OTUD5 belongs to the DUBs family and plays a critical role in controlling the Ub‐dependent degradation of various proteins, 4‐HNE may reduce the interaction between OTUD5 and GPX4 to promote GPX4 ubiquitination and subsequent degradation.To test whether OTUD5 affects the protein stability of GPX4, we co‐transfected GPX4 with Flag‐OTUD5 or its enzyme‐inactive mutant C224S into HEK293T cells."

reach
"Taken together, OTUD5 specifically deubiquitinated and stabilized the protein level of GPX4.Furthermore, we confirmed that GPX4 and OTUD5 could form a complex through reciprocal coimmunoprecipitation, immunofluorescence, duolink proximity ligation in situ assay (PLA), and GST‐pull down assay, and that 4‐HNE could weak this binding (Figure  5A–D; Figure S6E, Supporting Information)."

reach
"Taken together, these results indicate that OTUD5 depletion results in increased ferroptosis in cardiomyocytes.2.6 4-HNE Affects the Interaction between OTUD5 and GPX4 by Addition and Carbonylation."

reach
"We used the database to predict potential ubiquitination‐related enzymes targeting GPX4 (Figure S5A, Supporting Information) and found that E3 ligases MIB2, NEDD4L, STUB1, and deubiquitinase OTUD5 could bind to GPX4 in co‐immunoprecipitation experiments (Figure S5B, Supporting Information)."

reach
"In addition, our results demonstrated that only the binding between OTUD5 and GPX4 was affected under 4‐HNE treatment, while the interactions between E3 ligases (MIB2, NEDD4L, and STUB1) and GPX4 were not changed (Figure S5C, Supporting Information)."

reach
"In ALDH2 cKO mice, co‐immunoprecipitation assays showed that the levels of 4‐HNE adducted to GPX4 and OTUD5 were increased and the binding between GPX4 and OTUD5 was reduced (Figure S8E,F, Supporting Information)."

reach
"In mutant‐transfected HEK293T cells, the carbonylation of GPX4 and OTUD5 were significantly abolished (Figure 6G), the ability of 4‐HNE to weaken the interaction of GPX4 and OTUD5 and promote GPX4 ubiquitin–proteasomal degradation was eliminated (Figure 6H)."

reach
"N‐acetyl cysteine (NAC), an analog of cysteine that competitively binds to 4‐HNE, and amino acid substitution rescued 4‐HNE‐induced cysteine carbonylation of GPX4 and OTUD5, recovered the interaction between GPX4 and OTUD5, alleviated ubiquitination‐dependent degradation of GPX4, and inhibited myocyte ferroptosis."
GPX4 activates OTUD5.
| 1
GPX4 activates OTUD5. 1 / 1
| 1

reach
"In contrast, overexpression of GPX4 can reverse OTUD5 knockdown‐induced cellular lipid peroxidation (Figure S2B)."
DBC1 affects OTUD5
| 21
OTUD5 binds DBC1. 10 / 21
| 21

sparser
"Lots of interaction assays suggested that the potential DUB OTUD5 might interact with DBC1 ( xref ) and regulate its ubiquitination ( xref )."

sparser
"Co-IP analysis demonstrated the exogenous interaction between OTUD5 and DBC1 ( xref )."

sparser
"Furthermore, we confirmed that endogenous DBC1 could directly interact with OTUD5 ( xref , xref ), which was further verified by an in vitro pull-down assay using purified His-tagged DBC1 to pull down OTUD5 from cell lysates ( xref , xref )."

sparser
"Interestingly, we found that either hypoxia treatment or overexpression of SIAH2 promoted the interaction between DBC1 and SIAH2, but inhibited the interaction of DBC1 with OTUD5 ( xref , xref )."

sparser
"Moreover, we still need to further investigate the details of how the OTUD5-DBC1 complex dissociates under hypoxia to modulate DBC1 stability."

sparser
"Figure 4: the authors should study OTUD5 binding to DBC1 under hypoxia compared with normoxia to substantiate the claim for preferred interaction under normoxia."

sparser
"The authors propose that in normoxia, OTUD5 is bound to DBC1 which is displaced by SIAH2 upon activation by hypoxia, however despite the comprehensive study of the interaction between DBC1 this is not fully supported by the data."

sparser
"1) The authors propose a hypothesis that OTUD5 is bound to DBC1 in normoxic conditions which is displaced upon activation of SIAH2."

sparser
"Does OTUD5 bind to non-ubiquitinated DBC1?"

sparser
"Direct binding of recombinant DBC1 and OTUD5 could confirm this."

reach
"According to the results of the EdU assays, cell proliferation was strongly inhibited by OTUD5 knockdown in HCC cells (P < 0.05, Fig. 2C)."

reach
"Collectively, these results imply that OTUD5 orchestrates ferroptosis in gastric cancer cells.Moreover, we evaluated the impact of OTUD5 knockout on the cell cycle of gastric cancer cells by an EdU incorporation assay, and the results revealed that OTUD5 depletion decreased the proportion of cells in S phase, indicating impaired cell proliferation (Figure 3M,N)."

reach
"Consistently, overexpression of OTUD5 promoted the proliferation, while OTUD5 depletion suppressed the proliferation of gastric cancer cells (Figure 3O,P)."

reach
"These findings demonstrate that both OTUD5 knockdown and nutlin‐3a treatment can induce ferroptosis and suppress cell proliferation."

reach
"We treated control and OTUD5‐KO cells with nutlin‐3a and observed that knockout of OTUD5, in combination with nutlin‐3a treatment, led to a reduction in protein levels of OTUD5 and GPX4 (Figure 5A,B), thereby triggering ferroptosis in cancer cells and consequently inhibiting cell proliferation (Figure 5C,D)."

reach
"Taken together, these findings indicate that OTUD5 promotes HCC cell proliferation via SLC38A1."

reach
"Thus, the above evidence seems to support the high expression of OTUD5 in HCC, especially in HBV-related HCC.In our study, overexpression of OTUD5 promoted the proliferation of HCC cells, while OTUD5 knockdown inhibited cell proliferation, which was also confirmed in vivo."

reach
"SLC38A1 silencing prominently attenuated the OTUD5-induced increase in HCC cell proliferation."

reach
"OTUD5 promotes the proliferation of HCC cells."

reach
"Importantly, SLC38A1 knockdown significantly attenuated the OTUD5-induced increase in proliferation of HCC cells."

reach
"Consistently, overexpression of OTUD5 promoted the proliferation, while OTUD5 depletion suppressed the proliferation of gastric cancer cells (Figure 3O,P)."

reach
"These data collectively support the notion that OTUD5 inhibits the proliferation of gastric cancer cells.2.4 Wild-type p53 transcriptionally suppresses OTUD5 to promote ferroptosis in gastric cancer cells."

reach
"On the other hand, over-expression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."

reach
"In addition, OTUD5 knockdown has been shown to inhibit the proliferation of cancer cell lines with mutations that activate the mTOR pathway; these include HT29 colon cancer cells and MCF7 breast cancer cells [10]."

reach
"Overexpression of OTUD5 suppresses NSCLC cell proliferation, invasion, and migration."

reach
"Overall, these findings suggested that overexpression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."

reach
"We treated control and OTUD5‐KO cells with nutlin‐3a and observed that knockout of OTUD5, in combination with nutlin‐3a treatment, led to a reduction in protein levels of OTUD5 and GPX4 (Figure 5A,B), thereby triggering ferroptosis in cancer cells and consequently inhibiting cell proliferation (Figure 5C,D)."

reach
"Li et al. found that OTUD5 stabilizes PTEN through deubiquitination and that the overexpression of OTUD5 inhibits NSCLC cell proliferation, invasion and migration [9] (Figure 3E)."

eidos
"Two different lung tumor cell lines , H1299 and A549 , were used to investigate whether siRNA-mediated depletion of OTUD5 also promoted cell proliferation ."
OTUD5 affects TRIM25
1 1 | 6 12
OTUD5 binds TRIM25.
1 | 12
1 | 12

sparser
"The enriched biological processes were dominated by cell cycle and cell division processes, implying that the OTUD5-TRIM25 axis may affect cell proliferation by regulating the cell cycle (Fig.  xref )."

sparser
"The OTUD5-TRIM25 axis regulates liver cancer development."

sparser
"We tested whether the OTUD5-TRIM25 axis would affect tumor growth in a mouse model."

sparser
"In addition, the OTUD5/TRIM25 double knockdown cells formed smaller tumor sphere than did the OTUD5-depleted cells, demonstrating that the OTUD5-TRIM25 axis plays an important role in the regulation of tumor growth."

sparser
"OTUD5-TRIM25 transcriptionally regulates the expression of the putative tumor suppressor PML, which is dependent on the intact TRIM25 RING domain."

sparser
"Further investigation is needed to understand the precise regulatory mechanism underlying the OTUD5-TRIM25 function."

sparser
"Taken together, our study sheds light into regulation and function of the OTUD5-TRIM25 axis, suggesting it as a potential target for tumor therapy."

sparser
"However, some TRIM25 coprecipitated with the Flag-OTUD5-ΔUIM, indicating that the UIM may not be the sole interaction site and that TRIM25 may bind to the other parts of the OTUD5 protein (Fig.  xref )."

sparser
"We confirmed the interaction between endogenous OTUD5 and TRIM25 by the immunoprecipitation (IP) of Hep3B cells lysates."

sparser
"Our studies reveal a critical role of OTUD5 in tumorigenesis, as indicated through its interaction with TRIM25, suggesting that the OTUD5-TRIM25 axis acts as an important transcriptional regulator in tumor progression."
OTUD5 deubiquitinates TRIM25.
1 | 2
OTUD5 deubiquitinates TRIM25. 2 / 3
1 | 2

reach
"In tumor cells, OTUD5 promotes tumor progression by deubiquitinating TRIM25 [9]."

reach
"Besides, OTUD5 regulates gene transcription and inhibits tumorigenesis by deubiquitinating TRIM25, providing forceful evidence for the synergistic effect of deubiquitinating enzyme and ubiquitin ligase [34]."
OTUD5 increases the amount of TRIM25.
| 2
OTUD5 increases the amount of TRIM25. 2 / 2
| 2

reach
"By reducing the ubiquitination level of TRIM25, OTUD5 suppresses promyelocytic leukemia protein (PML) expression [7] (Figure 3C)."

reach
"Further investigation reveals that OTUD5 depletion leads to the enhanced transcriptional activity of TRIM25 and the inhibited expression of PML by altering the ubiquitination level of TRIM25."
OTUD5 ubiquitinates TRIM25.
| 1
Modified OTUD5 leads to the ubiquitination of TRIM25. 1 / 1
| 1

reach
"Our findings reveal a mechanism whereby OTUD5 regulates gene transcription and suppresses tumorigenesis by deubiquitinating TRIM25, providing a potential target for oncotherapy."
OTUD5 decreases the amount of TRIM25.
| 1
OTUD5 decreases the amount of TRIM25. 1 / 1
| 1

reach
"OTUD5 has been shown to reduce the ubiquitination level of TRIM25, resulting in its decreased transcriptional activity."
OTUD5 affects MYD88
| 12 8
OTUD5 deubiquitinates MYD88.
| 8
OTUD5 leads to the deubiquitination of MYD88 on K250. 2 / 2
| 2

reach
"Notably, OTUD5 abrogates K11-linked polyubiquitination of MyD88 at Lys95, Lys231 and Lys250, which subsequently promotes MyD88 oligomerization and the assembly of Myddosome."

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."
OTUD5 leads to the deubiquitination of MYD88 on K231. 2 / 2
| 2

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."

reach
"Notably, OTUD5 abrogates K11-linked polyubiquitination of MyD88 at Lys95, Lys231 and Lys250, which subsequently promotes MyD88 oligomerization and the assembly of Myddosome."
OTUD5 leads to the deubiquitination of MYD88 on K95. 2 / 2
| 2

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."

reach
"Notably, OTUD5 abrogates K11-linked polyubiquitination of MyD88 at Lys95, Lys231 and Lys250, which subsequently promotes MyD88 oligomerization and the assembly of Myddosome."
OTUD5 leads to the deubiquitination of MYD88. 2 / 2
| 2

reach
"To further investigate whether OTUD5 abrogated MyD88 polyubiquitination in physiological conditions, we detected the levels of endogenous MyD88 polyubiquitination."

reach
"In this study, we hypothesized that OTUD5 may abrogate MyD88 polyubiquitination."
OTUD5 binds MYD88.
| 8
| 8

sparser
"OTUD5 can directly interact with MyD88 and removed its K11-linked polyubiquitination."

sparser
"OTUD5 interacts with MyD88."

sparser
"The coimmunoprecipitation (Co-IP) analysis revealed that OTUD5 interacts with MyD88 but not IRAK4, IRAK2, or TRAF6 (Fig.  xref )."

sparser
"Taken together, these findings suggested that OTUD5 interacts with MyD88 but not the other adapter molecules involved in TLR/IL-1R signaling."

sparser
"To further confirm the interaction of OTUD5 with MyD88, we obtained PMs from WT mice."

sparser
"These collective findings indicated that OTUD5 interacts with MyD88 through the amino acids 360-571 on OTUD5 and the death domain of MyD88."

sparser
"OTUD5 directly interacts with MyD88 and cleaves its K11-linked polyubiquitin chains at Lys95, Lys231 and Lys250."

sparser
"A previous study revealed that OTUD5 interacts with MyD88, therefore promoting TLR/IL-1R mediated signal transduction."
OTUD5 ubiquitinates MYD88.
| 4
OTUD5 leads to the ubiquitination of MYD88. 1 / 1
| 1

reach
"The Co-IP analysis showed that OTUD5 could abrogate polyubiquitination of MyD88 (WT), while it does not possess the ability to eliminate the ubiquitination of MyD88 (K95R, K231R, K250R) mutant forms (Fig. 3H)."
OTUD5 leads to the ubiquitination of mutated MYD88 on K250. 1 / 1
| 1

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."
OTUD5 leads to the ubiquitination of MYD88 on K231. 1 / 1
| 1

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."
OTUD5 leads to the ubiquitination of MYD88 on K95. 1 / 1
| 1

reach
"The Co-IP analysis showed that OTUD5 inhibited polyubiquitination of MyD88 (WT) and its mutants MyD88 (K95), MyD88 (K231), and MyD88 (K250) in HEK293T cells (Fig. 3G)."
SUPT16H affects OTUD5
5 | 4 10
5 | 4 10

reach
"To study the cellular consequence of specifically blocking the interaction between OTUD5 and FACT (OTUD5 overexpression or complete knockout causes aberration in cell cycle and apoptosis), we generated a HeLa knock-in (KI) cell line in which the D537A mutation is introduced into the genomic loci of OTUD5 (OTUD5 ) using the CRISPR-Cas9 method (Supplementary Figure S2; sequencing information of 4 clones)."

reach
"Thus, blocking the interaction between FACT and OTUD5 enhances FACT nucleosome remodeling function.The OTUD5 cells display a noticeable increase in micronuclei and cytoplasmic bridges (Figure 3E), which can form as a consequence of replication stress."

sparser
"In addition, OTUD5 can interact with the SPT16 protein in the FACT complex and antagonize the deposition of histone H2A upon DSB damage by acting as a scaffold [ xref , xref ] ( xref D)."

sparser
"OTUD5 interacts with SPT16 and regulates the SPT16-dependent Pol II elongation at damaged chromatin."

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"Further, the interaction between OTUD5 and SPT16 can be detected in situ by the PLA (Figure xref )."

sparser
"These results unequivocally suggest that both the UBR5–OTUD5 interaction and OTUD5SPT16 interaction are necessary for transcriptional repression at the damaged chromatin."

sparser
"Independently of UBR5, OTUD5 also interacts with SPT16, through the UIM domain at the C-terminal tail."

reach
"These results confirm that the interaction between OTUD5 and FACT is important for the transcriptional regulation, and provide a new possible link between the FACT-dependent transcriptional process and tumor suppression."

reach
"Further investigation into the nature of the interaction between OTUD5 and FACT will provide more insight into the regulation."
OTUD5 affects Interferon
| 16
OTUD5 inhibits Interferon.
| 12
| 12

reach
"Indeed, DUBA and OTUD5, a DUB which selectively disassembles Lys63 linked ubiquitin chains on TRAF3, has been shown to suppress the production of type I interferon by reducing the level of TRAF3-TBK1 interaction XREF_BIBR."

reach
"Since DUBA is known to negatively regulate type I IFN and IFN is associated with IBD, it is important to understand association of DUBA and IBD as well."

reach
"OTUD5 (DUBA) is activated by phosphorylation ( Huang et al., 2012 ) and suppresses the type I interferon response by cleaving the polyubiquitin chain from tumor necrosis factor receptor-associated fac[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In the study showing that OTUD5 suppresses the type I interferon (IFN-1) production, the proposed mechanism is that OTUD5 antagonizes the TRAF3 signaling (21)."

reach
"The activation of TLR4 and TLR3 stimulates the K 63 -linked ubiquitination of TRAF3, which is required for the activation of IRF3 and the production of type I IFNs, and DUBA removes K 63 -linked ubiquitin chains from TRAF3, thereby inhibiting type I IFN responses."

reach
"For example, DUBA, a member of the Otubain (OTUB) family, has been shown to deubiquitinate TRAF3 and negatively regulate TLR3- and RIG-I and MDA5-mediated IFN induction."

reach
"Thus, DUBA mediated inhibition of TRF3 inhibition blocks TLR-dependent type 1 IFN production, but does not inhibit NF-κB activation [86] ."

reach
"OTUD5 has been shown to suppress the type I interferon dependent innate immune response by cleaving the polyubiquitin chain from an essential type I interferon adaptor protein."

reach
"It has been demonstrated that the deubiquitinating enzymes A20 and CYLD inhibit NF-kappaB signaling by targeting TRAF6 upstream of IKK, while the deubiquitinating protein DUBA inhibits type I interferon activity by targeting TRAF3."

reach
"For example, deubiquitinating enzyme A (DUBA) is able to negatively regulate the interferon response induced by PRRs to maintain homeostasis (Kayagaki et al., 2007)."
OTUD5 activates Interferon.
| 2
| 2

reach
"In the presence of CD-associated NOD2 mutations, impaired recognition of MDP results in inefficient induction of DUBA, which leads to excessive and colitogenic type I IFN and Th1 responses upon TLR9 activation ( Figure 4B )."

reach
"Among identified candidates were known regulators of IFN signaling such as Usp25 ( Lin et al., 2015 ), Usp15 ( Pauli et al., 2014 ), and DUBA, which displayed reduced expression in H1N1-infected cells[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 deubiquitinates Interferon.
| 2
OTUD5 deubiquitinates Interferon. 2 / 2
| 2

reach
"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."

reach
"TRIAD3A dependent TRAF3 proteasomal destruction, like TRAF3 de-ubiquitination by DUBA, halts the activation of TBK1 and IKKepsilon and subsequent IFN production."
OTUD5 affects NP
| 17
OTUD5 binds NP. 10 / 17
| 17

sparser
"Motions in the apo state of enzymes can facilitate substrate binding by enabling transitions into to a binding competent state xref , xref and can also regulate the catalytic cycle by allowing access to conformational states conducive for catalytic turnover. xref – xref The low activity of np-DUBA likely results from its inability to form a highly productive enzyme-substrate complex rather than low binding affinity to substrates because only 3-fold enhancement in the affinity to the fluorogenic ubiquitin-7-amino-4-methylcoumarin (Ub-AMC) substrate was observed upon phosphorylation, whereas the enhancement in k cat is 300-fold. xref We can speculate that an important feature of the productive enzyme-substrate complex is that the α1 helix can adopt a closed conformation, as defined by the crystal structure of p-DUBA in the ubiquitin-bound state. xref The only conformer in np-DUBA lies between the a and b conformers in terms of structural characteristics according to the relative positions of cross-peaks. xref This observation indicates that at least in the free state, the α1 helix in np-DUBA adopts a more open conformation than the a conformer in p-DUBA."

sparser
"In the enzyme-substrate complex formed from np-DUBA, the α1 helix either cannot adopt a closed conformation or fluctuates rapidly between open and closed conformations."

sparser
"Broadening of NMR spectral lines and the resulting low signal intensities are signatures of conformational exchanges on the μs-ms scales. xref Among ~ 120 cross-peaks detectable in the 15 N TROSY spectra of p-DUBA, many display broad 1 H lines, whereas the 15 N transverse relaxation rates of majority of the resonances are consistent with the molecular weight. xref , xref TROSY spectra of nonphosphorylated DUBA (np-DUBA) display similar features."

sparser
"The backbone assignments of p-DUBA and np-DUBA are available in the Biological Magnetic Resonance Bank (BMRB) under accession codes 27495 and 27494, respectively."

sparser
"Based on this observation, we performed constant-time amide 1 H Carr-Purcell-Meiboom-Gill (CPMG) experiments xref to characterize the conformational dynamics of both p-DUBA and np-DUBA."

sparser
"The data were subsequently group fit with a common exchange rate for residues that satisfy χ 2 group /χ 2 invididual < 2.0. xref Group fitting yielded k ex = 4954 ± 82 s −1 for p-DUBA ( xref ). np-DUBA undergoes a similar conformational exchange process with the exchange rate of 5750 ± 110 s −1 ( xref )."

sparser
"A network of dynamic residues underlies the synchronized motions observed in both p-DUBA and np-DUBA and explains the similarity in the global conformational dynamics in these two forms."

sparser
"Differences in the dynamics of p-DUBA and np-DUBA."

sparser
"Despite the overall similarity in the global conformational process between p-DUBA and np-DUBA, the α1 helix and neighboring regions display detectable differences."

sparser
"Most notably, S177 in np-DUBA shows a flat dispersion profile but elevated R 2,eff compared to the b conformer in p-DUBA ( xref )."
THBS1 affects OTUD5
| 7 8
THBS1 binds OTUD5.
| 6 8
| 6 8

reach
"Salvianolic acid B inhibits GPX4 ubiquitination by disrupting the interaction between THBS1 and OTUD5."

sparser
"These findings support the hypothesis that the interaction between THBS1 and OTUD5 mediates increased GPX4 ubiquitination; however, the precise mechanisms require further experimental validation."

reach
"Finally, the precise details of the interaction between THBS1 and OTUD5 remain undefined and will be the focus of our future research."

sparser
"Our study revealed that THBS1 in AMφ-derived exosomes interacted with OTUD5 in endothelial cells, leading to increased GPX4 ubiquitination."

sparser
"Salvianolic acid B inhibits GPX4 ubiquitination by disrupting the interaction between THBS1 and OTUD5."

sparser
"Finally, the precise details of the interaction between THBS1 and OTUD5 remain undefined and will be the focus of our future research."

sparser
"SAB treatment inhibited the interaction between THBS1 and OTUD5, leading to reduced GPX4 ubiquitination."

sparser
"Mechanistically, THBS1 interacted with OTUD5 to promote GPX4 ubiquitination, thereby increasing the sensitivity of endothelial cells to ferroptosis and contributing to endothelial barrier disruption."

sparser
"These findings suggest that SAB inhibits GPX4 ubiquitination by interfering with the interaction between THBS1 and OTUD5."

reach
"These findings support the hypothesis that the interaction between THBS1 and OTUD5 mediates increased GPX4 ubiquitination; however, the precise mechanisms require further experimental validation."
THBS1 activates OTUD5.
| 1
THBS1 activates OTUD5. 1 / 1
| 1

reach
"These findings indicated that in CIRI, THBS1 in macrophage-derived exosomes modulates OTUD5 deubiquitinase activity, thereby affecting GPX4 levels in endothelial cells."
Phosphatase affects OTUD5
| 1 14
Phosphatase dephosphorylates OTUD5. 10 / 15
| 1 14

reach
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S177 is phosphorylated by casein kinase II (CK2), a constitutively active kinase, making it likely that changes in the activity of the protein phosphatase that dephosphorylates OTUD5 controls its acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects ferroptosis
| 15
OTUD5 inhibits ferroptosis.
| 10
| 10

reach
"Collectively, our data illustrate that 4‐HNE triggers ubiquitin–proteasomal degradation of GPX4 in cardiomyocytes.2.5 OTUD5 Mediates 4-HNE-Induced GPX4 Degradation and Ferroptosis."

reach
"p53 inhibits OTUD5 transcription to promote GPX4 degradation and induce ferroptosis in gastric cancer."

reach
"These findings demonstrate that both OTUD5 knockdown and nutlin‐3a treatment can induce ferroptosis and suppress cell proliferation."

reach
"To evaluate whether OTUD5 inhibited ferroptosis, we assessed the levels of iron, MDA, and GSH (Figure 7E,F; Figure S9B, Supporting Information)."

reach
"Additionally, OTUD5 silencing and nutlin-3a-induced GPX4 degradation enhances the sensitivity of gastric cancer cells to ferroptosis in vivo."

reach
"We treated control and OTUD5‐KO cells with nutlin‐3a and observed that knockout of OTUD5, in combination with nutlin‐3a treatment, led to a reduction in protein levels of OTUD5 and GPX4 (Figure 5A,B), thereby triggering ferroptosis in cancer cells and consequently inhibiting cell proliferation (Figure 5C,D)."

reach
"Upregulation of OTUD5 deubiquitinates and stabilizes GPX4, thus reversing 4-HNE-induced ferroptosis and alleviating myocardial I/R injury [96]."

reach
"Conversely, in gastric cancer cells harbouring p53 mutations, OTUD5 transcription is upregulated due to the inactivation of p53, allowing OTUD5 to stabilise GPX4 protein, inhibit ferroptosis and thereby promoting gastric cancer progression (Figure 6J, right panel)."

reach
"In conclusion, these results demonstrated that OTUD5 mediates the K48‐linked deubiquitination of GPX4 and stabilises GPX4.2.3 OTUD5 suppresses ferroptosis via GPX4."

reach
"OTUD5 inhibited ferroptosis and alleviated myocardial injury by directly binding to glutathione peroxidase 4 (GPX4), the key protein of ferroptosis, and removing the K48-linked ubiquitin chains of GPX4 (Ref."
OTUD5 activates ferroptosis.
| 5
| 5

reach
"These findings suggest that OTUD5, through GPX4 stabilization, protects renal tubular cells from I/R-induced ferroptosis."

reach
"We then identified ovarian tumor domain-containing 5 (OTUD5) as a GPX4-interacting protein that promotes resistance to ferroptosis during I/R by stabilizing GPX4 expression."

reach
"These data implied that the protective effect of OTUD5 against I/R‐induced myocardial injury may be related to GPX4 dysfunction‐mediated ferroptosis."

reach
"We further investigated whether OTUD5 protected renal tubular cells from H/R-induced ferroptosis by stabilizing GPX4."

reach
"Specific targeting of OTUD5 may mitigate I/R‐induced ferroptosis and cardiac injury."
OTUD5 affects TRIP12
| 3 12
OTUD5 binds TRIP12.
| 2 12
| 2 12

sparser
"First, whether TRIP12 constitutively associates with OTUD5 is unknown."

sparser
"TRIP12OTUD5 antagonism regulates TNF-α–induced NF-κB signaling."

sparser
"We next verified TRIP12OTUD5 antagonism in TNF-α–induced NF-κB signaling."

sparser
"The data presented thus far indicate that OTUD5 is cooperatively modified by K29-specific TRIP12 and K48-specific UBR5, resulting in the conjugation of K29/K48-linked branched ubiquitin chains, and the biological importance of TRIP12OTUD5 antagonism is observed in TNF-α–induced NF-κB signaling."

sparser
"Collectively, these results indicate that TRIP12 interacts with OTUD5 and promotes its ubiquitylation and degradation."

sparser
"The biological importance of the TRIP12OTUD5 interplay was demonstrated in the regulation of NF-κB signaling."

sparser
"Because the simultaneous knockdown of OTUD5 partially restores NF-κB signaling under TRIP12 knockdown, the TRIP12OTUD5 antagonism may account for the regulatory function of TRIP12 in NF-κB signaling."

sparser
"This equilibrium favors ubiquitin chain assembly and subsequent OTUD5 degradation when TRIP12 interacts with OTUD5; UBR5 assembles K48-linked ubiquitin chains onto previously formed K29-linked chains more efficiently than de novo ubiquitylation (Fig.  xref ), presumably through ubiquitin interaction of the UBA domain (Fig.  xref and ref. xref )."

sparser
"TRIP12 interacted with OTUD5 during co-immunoprecipitation (Fig.  xref )."

sparser
"Whether the TRIP12OTUD5 axis also regulates other pathways should be studied further."
OTUD5 inhibits TRIP12.
| 1
OTUD5 inhibits TRIP12. 1 / 1
| 1

reach
"We hypothesized that changes in gene expression caused by OTUD5 accumulation under TRIP12 knockdown, may be canceled by simultaneous knockdown of OTUD5."
OTUD5 affects THBS1
| 6 8
| 6 8

reach
"Salvianolic acid B inhibits GPX4 ubiquitination by disrupting the interaction between THBS1 and OTUD5."

sparser
"These findings support the hypothesis that the interaction between THBS1 and OTUD5 mediates increased GPX4 ubiquitination; however, the precise mechanisms require further experimental validation."

reach
"Finally, the precise details of the interaction between THBS1 and OTUD5 remain undefined and will be the focus of our future research."

sparser
"Our study revealed that THBS1 in AMφ-derived exosomes interacted with OTUD5 in endothelial cells, leading to increased GPX4 ubiquitination."

sparser
"Salvianolic acid B inhibits GPX4 ubiquitination by disrupting the interaction between THBS1 and OTUD5."

sparser
"Finally, the precise details of the interaction between THBS1 and OTUD5 remain undefined and will be the focus of our future research."

sparser
"SAB treatment inhibited the interaction between THBS1 and OTUD5, leading to reduced GPX4 ubiquitination."

sparser
"Mechanistically, THBS1 interacted with OTUD5 to promote GPX4 ubiquitination, thereby increasing the sensitivity of endothelial cells to ferroptosis and contributing to endothelial barrier disruption."

sparser
"These findings suggest that SAB inhibits GPX4 ubiquitination by interfering with the interaction between THBS1 and OTUD5."

reach
"These findings support the hypothesis that the interaction between THBS1 and OTUD5 mediates increased GPX4 ubiquitination; however, the precise mechanisms require further experimental validation."
TRIM25 affects OTUD5
1 | 12
1 | 12

sparser
"The enriched biological processes were dominated by cell cycle and cell division processes, implying that the OTUD5-TRIM25 axis may affect cell proliferation by regulating the cell cycle (Fig.  xref )."

sparser
"The OTUD5-TRIM25 axis regulates liver cancer development."

sparser
"We tested whether the OTUD5-TRIM25 axis would affect tumor growth in a mouse model."

sparser
"In addition, the OTUD5/TRIM25 double knockdown cells formed smaller tumor sphere than did the OTUD5-depleted cells, demonstrating that the OTUD5-TRIM25 axis plays an important role in the regulation of tumor growth."

sparser
"OTUD5-TRIM25 transcriptionally regulates the expression of the putative tumor suppressor PML, which is dependent on the intact TRIM25 RING domain."

sparser
"Further investigation is needed to understand the precise regulatory mechanism underlying the OTUD5-TRIM25 function."

sparser
"Taken together, our study sheds light into regulation and function of the OTUD5-TRIM25 axis, suggesting it as a potential target for tumor therapy."

sparser
"However, some TRIM25 coprecipitated with the Flag-OTUD5-ΔUIM, indicating that the UIM may not be the sole interaction site and that TRIM25 may bind to the other parts of the OTUD5 protein (Fig.  xref )."

sparser
"We confirmed the interaction between endogenous OTUD5 and TRIM25 by the immunoprecipitation (IP) of Hep3B cells lysates."

sparser
"Our studies reveal a critical role of OTUD5 in tumorigenesis, as indicated through its interaction with TRIM25, suggesting that the OTUD5-TRIM25 axis acts as an important transcriptional regulator in tumor progression."
OTUD5 affects VDAC2
| 7 6
OTUD5 binds VDAC2.
| 2 6
| 2 6

sparser
"Because VDAC2 mainly interacts with OTUD5 via the N-terminal stretch, we hypothesized that OTUD5 likely modulates the ubiquitination of lysine residues (s) in the N-terminal stretch of VDAC2."

sparser
"OTUD5 directly interacts with VDAC2 in an OTU domain-dependent manner and sustains VDAC2 stability by cleaving K48-linked polyubiquitin chains from VDAC2 K23 and K31."

reach
"Immunofluorescence staining assays using confocal microscopy were then performed in HEK-293T cells, and a distinct interaction between OTUD5 and VDAC2 was detected (Figure 10A)."

reach
"Moreover, the direct interaction between OTUD5 and VDAC2 was confirmed using glutathione S-transferase precipitation assays (Figure 10C)."

sparser
"OTUD5 Interacts With VDAC2 and Cleaves the K48-linked Polyubiquitin Chains From VDAC2."

sparser
"Immunofluorescence staining assays using confocal microscopy were then performed in HEK-293T cells, and a distinct interaction between OTUD5 and VDAC2 was detected ( xref A )."

sparser
"Moreover, the direct interaction between OTUD5 and VDAC2 was confirmed using glutathione S-transferase precipitation assays ( xref C )."

sparser
"Collectively, these data indicate that OTUD5 directly interacts with VDAC2 in an OTU domain-dependent manner."
OTUD5 deubiquitinates VDAC2.
| 2
OTUD5 leads to the deubiquitination of VDAC2. 1 / 1
| 1

reach
"Consistently, we found that OTUD5 suppressed ubiquitination of both endogenous and exogenous VDAC2 in PAOA-treated HepG2 cells (Figure 12C and D)."
OTUD5 deubiquitinates VDAC2 on K23. 1 / 1
| 1

reach
"Consistently, OTUD5 overexpression did not affect the half-life of VDAC2-2KR, suggesting that OTUD5 mainly deubiquitinated VDAC2 at K23 and K31 (Figure 12J)."
OTUD5 decreases the amount of VDAC2.
| 2
OTUD5 decreases the amount of VDAC2. 2 / 2
| 2

reach
"41 We found that the OTUD5 knockdown-mediated decrease in VDAC2 expression level was abolished by MG132, supporting that OTUD5 restrained the proteasomal degradation of VDAC2 (Figure 12B)."

reach
"The data revealed that OTUD5 decreased the ubiquitination level of VDAC2 in cells transfected with K48O (Ub with the intact Lys48 residue alone), rather than K6O, K11O, K27O, K29O, K33O, or K63O (Figure 12G)."
OTUD5 inhibits VDAC2.
| 1
OTUD5 inhibits VDAC2. 1 / 1
| 1

reach
"In particular, OTUD5 knockdown-induced mitochondrial dysfunction was abolished in the presence of VDAC2 in PAOA-stimulated primary hepatocytes."
OTUD5 affects YAP1
| 12
OTUD5 deubiquitinates YAP1.
| 5
OTUD5 deubiquitinates YAP1. 5 / 5
| 5

reach
"Furthermore, we report for the first time that OTUD5 deubiquitinates and stabilizes YAP in macrophage."

reach
"In breast cancer, OTUD5 deubiquitinates and stabilizes YAP in macrophages, and YAP promotes M2 macrophage polarization, which is beneficial for enhancing the metastatic ability of breast cancer cells [12]."

reach
"In addition, OTUD5 can also mediate YAP deubiquitination in macrophages and subsequently promote M2 phenotype polarization, which facilitates the progression of triple-negative breast cancer [10]."

reach
"We next examined the possibility that OTUD5 might deubiquitinate YAP through its deubiquitinating enzyme activity."

reach
"We proved that dapagliflozin decreased the expression level of OTU deubiquitinase 5 (OTUD5), which further increased the ubiquitination and degradation of YAP1."
OTUD5 increases the amount of YAP1.
| 2
OTUD5 increases the amount of YAP1. 2 / 2
| 2

reach
"OTUD5 is able to promote macrophage differentiation toward the M2 type by stabilizing the protein levels of YAP [12]."

reach
"Depletion of OTUD5 by siRNA or sgRNA significantly decreased YAP levels, and ectopic expression of OTUD5 increased YAP ( Fig. 3 C–E)."
OTUD5 activates YAP1.
| 2
OTUD5 activates YAP1. 2 / 2
| 2

reach
"Depletion of OTUD5 by siRNA or sgRNA significantly decreased YAP levels, and ectopic expression of OTUD5 increased YAP ( Fig. 3 C–E)."

reach
"The CHX assay indicated that OTUD5 overexpression prolonged the half-life of YAP ( Fig. 3 F), while OTUD5 depletion reduced the protein stability of YAP ( Figs."
OTUD5 ubiquitinates YAP1.
| 1
OTUD5 leads to the ubiquitination of YAP1. 1 / 1
| 1

reach
"We proved that dapagliflozin decreased the expression level of OTU deubiquitinase 5 (OTUD5), which further increased the ubiquitination and degradation of YAP1."
OTUD5 inhibits YAP1.
| 1
OTUD5 inhibits YAP1. 1 / 1
| 1

reach
"The CHX assay indicated that OTUD5 overexpression prolonged the half-life of YAP ( Fig. 3 F), while OTUD5 depletion reduced the protein stability of YAP ( Figs."
OTUD5 binds YAP1.
| 1
OTUD5 binds YAP1 and 3 L. 1 / 1
| 1

reach
"Indeed, TNBC cells enhanced the binding between OTUD5 and YAP ( Fig. 3 L), and the ubiquitination assays showed the ubiquitin chain of YAP in co-culture group was dramatically removed ( Fig. 3 M)."
VDAC2 affects OTUD5
| 5 6
VDAC2 binds OTUD5.
| 2 6
| 2 6

sparser
"Because VDAC2 mainly interacts with OTUD5 via the N-terminal stretch, we hypothesized that OTUD5 likely modulates the ubiquitination of lysine residues (s) in the N-terminal stretch of VDAC2."

sparser
"OTUD5 directly interacts with VDAC2 in an OTU domain-dependent manner and sustains VDAC2 stability by cleaving K48-linked polyubiquitin chains from VDAC2 K23 and K31."

reach
"Immunofluorescence staining assays using confocal microscopy were then performed in HEK-293T cells, and a distinct interaction between OTUD5 and VDAC2 was detected (Figure 10A)."

reach
"Moreover, the direct interaction between OTUD5 and VDAC2 was confirmed using glutathione S-transferase precipitation assays (Figure 10C)."

sparser
"OTUD5 Interacts With VDAC2 and Cleaves the K48-linked Polyubiquitin Chains From VDAC2."

sparser
"Immunofluorescence staining assays using confocal microscopy were then performed in HEK-293T cells, and a distinct interaction between OTUD5 and VDAC2 was detected ( xref A )."

sparser
"Moreover, the direct interaction between OTUD5 and VDAC2 was confirmed using glutathione S-transferase precipitation assays ( xref C )."

sparser
"Collectively, these data indicate that OTUD5 directly interacts with VDAC2 in an OTU domain-dependent manner."
VDAC2 inhibits OTUD5.
| 2
VDAC2 inhibits OTUD5. 2 / 2
| 2

reach
"Mechanistically, VDAC2 mediates the inhibitory effects of OTUD5 on hepatocytes under metabolic stress."

reach
"VDAC2 overexpression restored OTUD5 knockdown induced transformation of mitochondrial morphology (Figure 14F)."
VDAC2 ubiquitinates OTUD5.
| 1
Mutated VDAC2 leads to the ubiquitination of OTUD5. 1 / 1
| 1

reach
"The results showed that both individual and combination mutants of VDAC2 resisted OTUD5-mediated deubiquitination (Figure 12I)."
SLC38A1 affects OTUD5
| 6 5
SLC38A1 binds OTUD5.
| 4 5
| 4 5

sparser
"Our further experiments verified the interaction between OTUD5 and SLC38A1."

reach
"Coimmunoprecipitation analysis confirmed the interaction between OTUD5 and SLC38A1."

reach
"OTUD5 binds to SLC38A1."

reach
"Co-IP assays confirmed that both exogenous and endogenous OTUD5 bound to SLC38A1 (Fig. 5B and C)."

reach
"Our further experiments verified the interaction between OTUD5 and SLC38A1."

sparser
"Coimmunoprecipitation analysis confirmed the interaction between OTUD5 and SLC38A1."

sparser
"Finally, the interaction between OTUD5 and solute carrier family 38 member 1 (SLC38A1) and the underlying regulatory mechanism were investigated."

sparser
"OTUD5 binds to SLC38A1."

sparser
"Co-IP assays confirmed that both exogenous and endogenous OTUD5 bound to SLC38A1 (Fig.  xref B and C)."
SLC38A1 inhibits OTUD5.
| 1
Ubiquitinated SLC38A1 inhibits OTUD5. 1 / 1
| 1

reach
"The results showed that the knockdown of OTUD5 increased the level of ubiquitinated SLC38A1 in HCC cells (Fig. 6D and Supplementary Fig. 4D), while the level of ubiquitinated SLC38A1 was significantly reduced in HCC cells overexpressing OTUD5 (Fig. 6E and Supplementary Fig. 4E)."
SLC38A1 activates OTUD5.
| 1
Ubiquitinated SLC38A1 activates OTUD5. 1 / 1
| 1

reach
"The results showed that the knockdown of OTUD5 increased the level of ubiquitinated SLC38A1 in HCC cells (Fig. 6D and Supplementary Fig. 4D), while the level of ubiquitinated SLC38A1 was significantly reduced in HCC cells overexpressing OTUD5 (Fig. 6E and Supplementary Fig. 4E)."
OTUD5 affects MAP3K7
| 7 4
OTUD5 binds MAP3K7.
| 3 4
| 3 4

sparser
"We confirmed the endogenous interaction between OTUD5 and TAK1 in podocytes using a Co-immunoprecipitation (co-IP) assay (Fig.  xref )."

sparser
"Similarly, endogenous OTUD5 and TAK1 could form a complex in mouse kidney tissues (Fig.  xref )."

sparser
"Moreover, Flag-OTUD5 and His-TAK1 plasmids were co-transfected into NIH/3T3 cells, confirming the exogenous interaction between OTUD5 and TAK1 (Fig.  xref )."

reach
"Therefore, we hypothesize that OTUD5 negatively regulates inflammation and injury through deubiquitinating TAK1 in podocytes.We confirmed the endogenous interaction between OTUD5 and TAK1 in podocytes using a Co-immunoprecipitation (co-IP) assay (Fig. 4c)."

reach
"Similarly, endogenous OTUD5 and TAK1 could form a complex in mouse kidney tissues (Fig. 4d)."

reach
"Moreover, Flag-OTUD5 and His-TAK1 plasmids were co-transfected into NIH/3T3 cells, confirming the exogenous interaction between OTUD5 and TAK1 (Fig. 4e)."

sparser
"Mechanistically, we demonstrated that OTUD5 directly interacts with TAK1 and reduces K63-linked polyubiquitination of TAK1 at residue K158 through its active site C224, subsequently inhibiting the TAK1-mediated inflammatory response and injury in podocytes."
OTUD5 deubiquitinates MAP3K7.
| 2
OTUD5 deubiquitinates MAP3K7. 2 / 2
| 2

reach
"Podocyte OTUD5 alleviates diabetic kidney disease through deubiquitinating TAK1 and reducing podocyte inflammation and injury."

reach
"As shown in Fig. 4f, OTUD5 overexpression decreased the polyubiquitination of TAK1 in NIH/3T3 cells."
OTUD5 dephosphorylates MAP3K7.
| 2
OTUD5 leads to the dephosphorylation of MAP3K7. 2 / 2
| 2

reach
"Together, OTUD5 negatively regulates TAK1-MAPK activation and inflammation in podocytes under HG/PA treatment.Then, we tried to explore how OTUD5-mediated TAK1 deubiquitination prevents TAK1 phosphorylation."

reach
"Western blot assay confirmed that OTUD5 overexpression significantly inhibited HG/PA-induced TAK1 phosphorylation in podocytes but did not change the total TAK1 protein level (Fig. 5a, Supplementary Fig. 5a)."
OTUD5 affects STING1
2 | 5 3
OTUD5 binds STING1.
2 | 3
2 | 3

sparser
"USP11 exacerbates radiation induced pneumonitis by activating endothelial cell inflammatory response via OTUD5-STING signaling."

sparser
"Here, we report that the deubiquitinase OTUD5 interacts with STING, cleaves its K48-linked polyubiquitin chains, and promotes its stability."

sparser
"OTUD5 interacts with STING, cleaves its K48-linked polyubiquitin chain and promotes its stability ( xref B) [ xref ]."
OTUD5 activates STING1.
| 5
OTUD5 activates STING1. 5 / 5
| 5

reach
"25 OTUD5 enhanced anti‐viral infection and anti‐tumor immunity by stabilizing STING."

reach
"In the innate immune response, OTUD5 can promote the stability of STING [37]."

reach
"Together, these results suggest that USP11/OTUD5 axis induces RILI through stabilizing STING in endothelial cells [74]."

reach
"Similarly, OTUD5 promotes the protein stability of STING via cleaving the K48-linked polyubiquitin chains."

reach
"On the other hand, OTUD5 and USP20 catalyze the K48-linked deubiquitination of STING and inhibit STING degradation, thus maintain the stability of STING and promote activation of STING pathway (87–89)."
OTUD5 affects Flag
| 10
| 10

sparser
"In ubiquitination experiments, we transfected 3 µg His‐α‐Syn, 2 µg FlagOTUD5 or its mutant and 1 µg HA‐ubiquitin."

sparser
"To validate which type of MyD88 polyubiquitin chain was abrogated by OTUD5, HA-ubiquitin mutants K0, K6, K11, K27, K29, K33, K48 and K63 (containing only one lysine residue with other lysine residues replaced by arginine), together with Myc-MyD88 and Flag-OTUD5 (WT) were co-transfected into HEK293T cells."

sparser
"Each myc-tagged PDCD5 point mutant was co-transfected with Flag-p53 or Flag-OTUD5, and then each PDCD5 mutant was screened by immunoprecipitation analysis."

sparser
"Moreover, Flag-OTUD5 and His-TAK1 plasmids were co-transfected into NIH/3T3 cells, confirming the exogenous interaction between OTUD5 and TAK1 (Fig.  xref )."

sparser
"To investigate whether ubiquitination was involved in MyD88 oligomerization, we transfected Flag-OTUD5 (WT) or its mutant C224S and GFP-MyD88 into HEK293T cells."

sparser
"The results showed that transfection of Flag-OTUD5 (WT) enhanced MyD88 oligomerization (Fig.  xref )."

sparser
"However, transfection of the Flag-OTUD5 mutant C224S did not promote MyD88 oligomerization (Fig.  xref )."

sparser
"HA-YAP and Flag-OTUD5 were constructed by cloning cDNA into pcDNA3-HA or pcDNA3-Flag vector, respectively."

sparser
"HCT116 cells were co-transfected with plasmids encoding HA-Ub and Myc-PDCD5 or Flag-OTUD5."

sparser
"To test this hypothesis, we transfected Flag-OTUD5 (WT) or C224S (mutation of cysteine to serine at position 224) [ xref , xref ], Myc-MyD88 and HA-Ubiquitin into HEK293T cells."
OTUD5 affects HNF4A
| 9
OTUD5 increases the amount of HNF4A.
| 4
OTUD5 increases the amount of HNF4A. 4 / 4
| 4

reach
"Meanwhile, overexpression of OTUD5 could inhibit the MAPK pathway and then increase the expression of HNF4a, and ERK1/2 signaling was required for OTUD5-mediated activation of HNF4alpha, promoting HBV replication."

reach
"Together, these data indicate that OTUD5 could deubiquitinate HBV core protein degradation by its deubiquitinase function and promote HBV activity by up-regulating HNF4alpha expression via inhibition of the ERK1/2 pathway."

reach
"Another result showed that by activating the ERK1/2 signaling pathway, OTUD5 promoted the expression of HNF4α, a well-known activator of HBV core promoter which can promote the replication of HBV."

reach
"In this study, we proved OTUD5 could downregulate the expression of phosphorylation ERK1/2 in hepG2.2.15 cells, and the results illustrated that OTUD5 could promote the expression of HNF4α via inhibiting ERK1/2 of MAPK pathway (Fig. 7A–D)."
OTUD5 activates HNF4A.
| 4
OTUD5 activates HNF4A. 4 / 4
| 4

reach
"However, the inhibition of ERK1/2 but not JNK and p38 MAPK pathways enhanced OTUD5-mediated promotion of HNF4α (Fig. 7D)."

reach
"The data of PCR showed that knockdown of OTUD5 significantly downregulated the mRNA of HNF1ɑ and HNF4ɑ (Fig. 6B,C)."

reach
"Conversely, the overexpression of OTUD5 could upregulate the mRNA of HNF1ɑ and HNF4ɑ (Fig. 6E–F)."

reach
"Meanwhile, overexpression of OTUD5 could inhibit the MAPK pathway and then increase the expression of HNF4a, and ERK1/2 signaling was required for OTUD5-mediated activation of HNF4alpha, promoting HBV replication."
OTUD5 decreases the amount of HNF4A.
| 1
OTUD5 decreases the amount of HNF4A. 1 / 1
| 1

reach
"Together, these data indicate that OTUD5 could deubiquitinate HBV core protein degradation by its deubiquitinase function and promote HBV activity by up-regulating HNF4alpha expression via inhibition of the ERK1/2 pathway."
Ubiquitin affects OTUD5
| 3 5
Ubiquitin binds OTUD5.
| 1 5
| 1 5

reach
"Huang et al. reported the crystal structure of the pSer177 OTUD5Ub complex."

sparser
"In the second method, after phosphorylation modification, the conformation of DUBs changes only after binding to the substrate, such as the phosphorylated DUBA bound to Ub, which alters its conformation instead of the phosphorylated DUBA [ xref ]."

sparser
"Huang et al. reported the crystal structure of the pSer177 OTUD5Ub complex."

sparser
"The crystal structure of the Ub-aldehyde complex with phosphorylated OTUD5 reveals a remarkable interaction between the phosphorylated Ser177 of OTUD5 and the guanidinium moiety of Arg74 of the bound ubiquitin showing that the phosphorylation is essential for the DUB activity [ xref ]."

sparser
"In the crystal structure of OTUD5 bound with ubiquitin ( xref ), Asp256, Arg274, and Asn293 are located on the surface of the protein, and form no direct interactions with ubiquitin ( xref )."

sparser
"In the crystal structure of the Ser177-phosphorylated form of OTUD5 bound with ubiquitin ( xref ), the phosphate group has intramolecular interactions with other OTUD5 polar residues (including Arg272), and also has intermolecular interactions with the C-terminal tail of the bound ubiquitin, precisely orienting its C-terminus for catalysis."
Ubiquitin activates OTUD5.
| 2
| 2

reach
"Considering other potential post-transcriptional mechanisms, we investigated the ubiquitin ligase UBR5, which has been reported to mediate OTUD5 abundance post-transcriptionally ."

reach
"An intricate web of interactions involving the phosphate and the C-terminal tail of ubiquitin cause DUBA to fold around its substrate, revealing why phosphorylation is essential for deubiquitinase activity."
OTUD5 affects Ubiquitin
| 3 5
OTUD5 binds Ubiquitin.
| 1 5
| 1 5

reach
"Huang et al. reported the crystal structure of the pSer177 OTUD5Ub complex."

sparser
"In the second method, after phosphorylation modification, the conformation of DUBs changes only after binding to the substrate, such as the phosphorylated DUBA bound to Ub, which alters its conformation instead of the phosphorylated DUBA [ xref ]."

sparser
"Huang et al. reported the crystal structure of the pSer177 OTUD5Ub complex."

sparser
"The crystal structure of the Ub-aldehyde complex with phosphorylated OTUD5 reveals a remarkable interaction between the phosphorylated Ser177 of OTUD5 and the guanidinium moiety of Arg74 of the bound ubiquitin showing that the phosphorylation is essential for the DUB activity [ xref ]."

sparser
"In the crystal structure of OTUD5 bound with ubiquitin ( xref ), Asp256, Arg274, and Asn293 are located on the surface of the protein, and form no direct interactions with ubiquitin ( xref )."

sparser
"In the crystal structure of the Ser177-phosphorylated form of OTUD5 bound with ubiquitin ( xref ), the phosphate group has intramolecular interactions with other OTUD5 polar residues (including Arg272), and also has intermolecular interactions with the C-terminal tail of the bound ubiquitin, precisely orienting its C-terminus for catalysis."
OTUD5 activates Ubiquitin.
| 2
| 2

reach
"OTUD5 overexpression significantly increased α‐Syn polyubiquitination in WT and K63 ubiquitin transfected cells but not in K48 ubiquitin transfected cells (Figure 3I)."

reach
"An intricate web of interactions involving the phosphate and the C-terminal tail of ubiquitin cause DUBA to fold around its substrate, revealing why phosphorylation is essential for deubiquitinase activity."
OTUD5 affects FlaG
| 8
| 8

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"At last, the PLA assay showed that FLAG-OTUD5 WT interacted with endogenous UBR5, measured by increased PLA spots, but the FLAG-OTUD5 ΔN17 mutant did not (Figure xref ), further supporting the conclusions."

sparser
"First, we found that expressing the FLAG-OTUD5 WT reversed (repressed) the YFP-MS2 reporter expression at the Fok1-induced DSB sites in OTUD5 knockdown cells (Figure xref )."

sparser
"To identify OTUD5 target(s) during NF-κB activation, we analyzed FLAGOTUD5 associated protein identified by LC-MS."

sparser
"However, neither ΔN17 nor ΔUIM mutant were able to rescue the phenotype (Figure xref ; presence of FLAG-OTUD5 mutants at the spots indicate the normal expression and localization)."

sparser
"Below panel shows the expression confirmation of the FLAG-OTUD5 truncates."

sparser
"The depleted UBR5 proteins were effectively rescued by expressing FLAG-UBR5 wild-type (WT), but not by a catalytically inactive FLAG-OTUD5 mutant (C224S) ( xref ) (Figure xref ), suggesting that the deubiquitinating activity of OTUD5 is necessary for preserving the UBR5 stability."

sparser
"Indeed, we have observed this in several ways; foci of endogenous OTUD5 and GFP-OTUD5 were detected at UVC-induced γH2AX spots (Figure xref and  xref , respectively; FLAG-OTUD5 in xref )."

reach
"The function of OTUD5 is required to allow the rapid activation of p53 dependent transcription and a p53 dependent apoptosis in response to DNA damage stress."

reach
"Using RNAi, we found that certain deubiquitinases, BAP1, USP14 and OTUD5 each promote proviral transcription."

reach
"We found that OTUD5, and its binding partner UBR5, are modulators of transcription-induced replication fork stress."

reach
"21 In addition, OTUD5 promotes DNA damage repair by inhibiting Ku80 degradation or regulating FACT‐dependent transcription at damaged chromatin."

reach
"Using the ROC curve and Kaplan–Meier method, OTUD5 was demonstrated as an interesting biomarker for predicting HBeAg seroconversion.In summary, we revealed that deubiquitinase OTUD5 could positively regulate HBV transcription and replication in HepG2.2.15 cells by deubiquitinating the K48-mediated ubiquitination of hepatitis B precore/core components to promote HBV infection."

reach
"OTUD5 promoted HBV transcription and replication."

reach
"These results suggest that both OTUD5 and UBR5 repress transcription nearby DSB lesions."

reach
"OTUD5 downregulates the stability of the transcription factor RORγt, which induces IL-17 production in T cells, and interestingly, it does so indirectly by stabilizing UBR5 (43); the model suggested that UBR5 ubiquitinates and degrades RORγt. Therefore, the role of OTUD5 in cytokine production has a similarity with the role described herein, in that it negatively impacts transcription, at least in part through UBR5 stabilization."
MYD88 affects OTUD5
| 8
| 8

sparser
"OTUD5 can directly interact with MyD88 and removed its K11-linked polyubiquitination."

sparser
"OTUD5 interacts with MyD88."

sparser
"The coimmunoprecipitation (Co-IP) analysis revealed that OTUD5 interacts with MyD88 but not IRAK4, IRAK2, or TRAF6 (Fig.  xref )."

sparser
"Taken together, these findings suggested that OTUD5 interacts with MyD88 but not the other adapter molecules involved in TLR/IL-1R signaling."

sparser
"To further confirm the interaction of OTUD5 with MyD88, we obtained PMs from WT mice."

sparser
"These collective findings indicated that OTUD5 interacts with MyD88 through the amino acids 360-571 on OTUD5 and the death domain of MyD88."

sparser
"OTUD5 directly interacts with MyD88 and cleaves its K11-linked polyubiquitin chains at Lys95, Lys231 and Lys250."

sparser
"A previous study revealed that OTUD5 interacts with MyD88, therefore promoting TLR/IL-1R mediated signal transduction."
MAP3K7 affects OTUD5
| 4 4
MAP3K7 binds OTUD5.
| 3 4
| 3 4

sparser
"We confirmed the endogenous interaction between OTUD5 and TAK1 in podocytes using a Co-immunoprecipitation (co-IP) assay (Fig.  xref )."

sparser
"Similarly, endogenous OTUD5 and TAK1 could form a complex in mouse kidney tissues (Fig.  xref )."

sparser
"Moreover, Flag-OTUD5 and His-TAK1 plasmids were co-transfected into NIH/3T3 cells, confirming the exogenous interaction between OTUD5 and TAK1 (Fig.  xref )."

reach
"Therefore, we hypothesize that OTUD5 negatively regulates inflammation and injury through deubiquitinating TAK1 in podocytes.We confirmed the endogenous interaction between OTUD5 and TAK1 in podocytes using a Co-immunoprecipitation (co-IP) assay (Fig. 4c)."

reach
"Similarly, endogenous OTUD5 and TAK1 could form a complex in mouse kidney tissues (Fig. 4d)."

reach
"Moreover, Flag-OTUD5 and His-TAK1 plasmids were co-transfected into NIH/3T3 cells, confirming the exogenous interaction between OTUD5 and TAK1 (Fig. 4e)."

sparser
"Mechanistically, we demonstrated that OTUD5 directly interacts with TAK1 and reduces K63-linked polyubiquitination of TAK1 at residue K158 through its active site C224, subsequently inhibiting the TAK1-mediated inflammatory response and injury in podocytes."
MAP3K7 activates OTUD5.
| 1
MAP3K7 activates OTUD5. 1 / 1
| 1

reach
"In contrast, OTUD5 knockdown by siRNA aggravated TAK1 activation in HG/PA-challenged podocytes (Fig. 5b)."
FlaG affects OTUD5
| 8
| 8

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"At last, the PLA assay showed that FLAG-OTUD5 WT interacted with endogenous UBR5, measured by increased PLA spots, but the FLAG-OTUD5 ΔN17 mutant did not (Figure xref ), further supporting the conclusions."

sparser
"First, we found that expressing the FLAG-OTUD5 WT reversed (repressed) the YFP-MS2 reporter expression at the Fok1-induced DSB sites in OTUD5 knockdown cells (Figure xref )."

sparser
"To identify OTUD5 target(s) during NF-κB activation, we analyzed FLAGOTUD5 associated protein identified by LC-MS."

sparser
"However, neither ΔN17 nor ΔUIM mutant were able to rescue the phenotype (Figure xref ; presence of FLAG-OTUD5 mutants at the spots indicate the normal expression and localization)."

sparser
"Below panel shows the expression confirmation of the FLAG-OTUD5 truncates."

sparser
"The depleted UBR5 proteins were effectively rescued by expressing FLAG-UBR5 wild-type (WT), but not by a catalytically inactive FLAG-OTUD5 mutant (C224S) ( xref ) (Figure xref ), suggesting that the deubiquitinating activity of OTUD5 is necessary for preserving the UBR5 stability."

sparser
"Indeed, we have observed this in several ways; foci of endogenous OTUD5 and GFP-OTUD5 were detected at UVC-induced γH2AX spots (Figure xref and  xref , respectively; FLAG-OTUD5 in xref )."
OTUD5 affects cell growth
| 4 3
OTUD5 inhibits cell growth.
| 4 2
| 4 2

reach
"Through an unbiased RNAi screen, knockdown of OTUD5 is shown to significantly accelerate cell growth."

eidos
"OTUD5 knockdown accelerated cell growth ."

eidos
"Through an unbiased RNAi screen , knockdown of OTUD5 is shown to significantly accelerate cell growth ."

eidos
"OTUD5 knockdown promoted cell growth in a TRIM25-dependent manner ."

reach
"In a study conducted by Li et al. (2020), an impartial RNAi screen was employed to demonstrate that the knockdown of OTUD5 substantially promotes cell growth."

eidos
"In the current study , we find that OTUD5 knockdown accelerates cell growth in a TRIM25-dependent manner ."
OTUD5 activates cell growth.
| 1
| 1

reach
"To further confirm whether OTUD5-induced cell growth involves SLC38A1, we knocked down SLC38A1 expression in HCC cells overexpressing OTUD5 (P < 0.05, Fig. 7A)."
OTUD5 affects XRCC5
| 6 1
OTUD5 inhibits XRCC5.
| 3
OTUD5 inhibits XRCC5. 3 / 3
| 3

reach
"21 In addition, OTUD5 promotes DNA damage repair by inhibiting Ku80 degradation or regulating FACT‐dependent transcription at damaged chromatin."

reach
"This suggests the possibility that OTUD5 is directly phosphorylated by ATM to function as an amplifier of ATM-dependent DDR signaling in a potential positive-feedback circuit, resembling its role in mTOR signaling.In the context of DNA repair, OTUD5 has been suggested to promote NHEJ by counteracting the degradation of Ku80, a core NHEJ factor, in certain cell lines ."

reach
"OTUD5 also stabilises Ku80, loss of OTUD5 reduces NHEJ repair efficiency [11] (Figure 2)."
OTUD5 binds XRCC5.
| 2 1
| 2 1

reach
"In addition, the new autoantigen reported here, the deubiquitinase OTUD5, associates with Ku80 to regulate its function ."

sparser
"OTUD5 interacts with Ku80 to positively regulate non-homologous end joining NHEJ-mediated repair ( xref )."

reach
"OTUD5 interacts with Ku80 to positively regulate non homologous end joining NHEJ mediated repair."
OTUD5 activates XRCC5.
| 1
OTUD5 activates XRCC5. 1 / 1
| 1

reach
"OTUD5 promotes NHEJ repair by increasing the stability of Ku80 [52] (Figure 2E)."
OTUD5 affects TRIM28
| 4 3
OTUD5 phosphorylates TRIM28.
| 3 3
OTUD5 leads to the phosphorylation of TRIM28 on S824. 5 / 5
| 2 3

sparser
"We find that OTUD5 promotes ATM-mediated phosphorylation of KAP1 S824 in response to telomere deprotection, as well as in response to genome-wide, ionizing radiation (IR)-induced DNA damage."

sparser
"We identified that the OTU family deubiquitinase (DUB) OTUD5 promotes KAP1 S824 phosphorylation by facilitating ATM activation, through stabilization of the ubiquitin ligase UBR5 that is required for DNA damage-induced ATM activity."

reach
"We identified that the OTU family deubiquitinase (DUB) OTUD5 promotes KAP1 S824 phosphorylation by facilitating ATM activation, through stabilization of the ubiquitin ligase UBR5 that is required for DNA damage-induced ATM activity."

sparser
"Next, we assessed the potential functional involvement of UBR5 in OTUD5-promoted KAP1 S824 phosphorylation, by depleting OTUD5, UBR5, or both in U2OS cells."

reach
"Loss of OTUD5 impairs KAP1 S824 phosphorylation, which suppresses end-joining mediated DNA repair at deprotected telomeres and at DNA breaks in heterochromatin."
OTUD5 leads to the phosphorylation of TRIM28. 1 / 1
| 1

reach
"Next, we assessed the potential functional involvement of UBR5 in OTUD5-promoted KAP1 phosphorylation, by depleting OTUD5, UBR5, or both in U2OS cells."
OTUD5 dephosphorylates TRIM28.
| 1
OTUD5 leads to the dephosphorylation of TRIM28. 1 / 1
| 1

reach
"This shows that the residual ATM activity in OTUD5-deficient cells is inadequate to support sufficient phosphorylation of KAP1 , a key step in DNA repair in KAP1-rich genomic environments such as the telomeres and heterochromatin, thereby translating into a repair defect specifically in these regions."
OTUD5 affects IFN-I
| 7
OTUD5 inhibits IFN-I.
| 5
OTUD5 inhibits IFN-I. 5 / 5
| 5

reach
"Notably, UCHL1, OTUB1, OTUB2, OTUD5, USP25, MYSM1, and MCPIP1 (Figure 3A) negatively regulate virus-induced IFN-I production and antiviral activities by cleaving K63-linked or other types of polyubiquitin chains from TRAF3 or TRAF6."

reach
"Studies have shown that OTUD5 is able to attenuate the IFN-I response induced by TLR and RLR signaling [5]."

reach
"The knockout of OTUD5 leads to faster turnover of STING and impairs IFN-I signaling following cytosolic DNA stimulation, whereas Lyz2-Cre Otud5fl/Y mice and CD11-Cre Otud5fl/Y mice show higher susceptibility to HSV-1 infection than their corresponding control littermates (69)."

reach
"Reduction of DUBA could augment IFN-I response, whereas ectopic expression of DUBA has the opposite effect."

reach
"As a DUB, DUBA can selectively weaken the K63-linked ubiquitination of TRAF3, assist in the dissociation of TRAF3 from the signal complex containing TBK1, and negatively regulate the production of IFN-I mediated by TRAF3 (75)."
OTUD5 activates IFN-I.
| 2
OTUD5 activates IFN-I. 2 / 2
| 2

reach
"Among the DUBs that interact with STING, five members, namely CYLD, OTUD5, USP18 (also termed UBP43), USP20, and USP44, have been demonstrated to promote IFN-I production and antiviral responses."

reach
"Notably, UCHL1, OTUB1, OTUB2, OTUD5, USP25, MYSM1, and MCPIP1 (Figure 3A) negatively regulate virus-induced IFN-I production and antiviral activities by cleaving K63-linked or other types of polyubiquitin chains from TRAF3 or TRAF6."
OTUD5 affects HYCC1
| 7
OTUD5 activates HYCC1.
| 6
OTUD5 activates HYCC1. 6 / 6
| 6

reach
"Therefore, our results suggest that OTUD5 promotes HCC cell proliferation by increasing the stability of SLC38A1."

reach
"Taken together, these findings indicate that OTUD5 promotes HCC cell proliferation via SLC38A1."

reach
"Thus, the above evidence seems to support the high expression of OTUD5 in HCC, especially in HBV-related HCC.In our study, overexpression of OTUD5 promoted the proliferation of HCC cells, while OTUD5 knockdown inhibited cell proliferation, which was also confirmed in vivo."

reach
"Finally, OTUD5 knockdown markedly suppressed the growth of HCC cells in vivo."

reach
"OTUD5 promotes the proliferation of HCC cells."

reach
"CCK-8 assays verified that OTUD5 silencing decreased the viability of HCC cells (P < 0.05, Fig. 2B)."
OTUD5 deubiquitinates HYCC1.
| 1
OTUD5 deubiquitinates HYCC1. 1 / 1
| 1

reach
"Therefore, these results support that OTUD5 deubiquitinates and stabilizes SLC38A1 in HCC."
OTUD5 affects ATM
| 7
OTUD5 activates ATM.
| 6
OTUD5 activates ATM. 6 / 6
| 6

reach
"OTUD5 promotes ATM-dependent DDR signaling."

reach
"As independently, UBR5 was found to inhibit the interaction between ATM and ATMIN, a negative regulator of canonical DSB-induced ATM activation , we hypothesized that OTUD5 may promote ATM-dependent DDR signaling via stabilization of UBR5."

reach
"OTUD5 promotes ATM-dependent DDR activation by regulating the UBR5-ATMIN axis."

reach
"Indeed, we found OTUD5 to promote ATM-dependent DDR signaling in a manner that is epistatic with UBR5, dependent on its catalytic activity and involving ATMIN, supporting that OTUD5 acts via UBR5.Early biochemical studies performed on OTUD5 suggested that phosphorylation at residue Ser-177 is vital for its DUB activity in vitro ."

reach
"These previously established links prompted us to hypothesize that OTUD5 promotes ATM-dependent DDR and telomeric DNA repair in a UBR5 and ATMIN-dependent manner."

reach
"If OTUD5 indeed promotes ATM-dependent DDR and telomeric DNA repair through the UBR5-ATMIN axis, ablation of ATMIN is expected to reverse the DDR defects exhibited by OTUD5-depleted cells."
OTUD5 binds ATM.
| 1
| 1

reach
"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
IFNG affects OTUD5
| 1 6
IFNG increases the amount of OTUD5.
| 5
IFNG increases the amount of OTUD5. 5 / 5
| 5

reach
"IFN-γ has been detected to upregulate the expression of OTUD5 through p38-dependent mechanism to amplify the aberrant inflammatory cytokine response in IBD (Dinallo et al., 2022)."

reach
"It has been found that IFN-γ stimulation increases OTUD5 expression in LPMC from patients with inflammatory bowel disease."

reach
"IFN-γ and IL-1β have been shown to upregulate and downregulate expression of DUBA, respectively."

reach
"Additionally, they examined the molecular mechanisms of DUBA expression and found that IFN-γ promotes the DUBA expression through p38/mitogen-activated protein kinase pathways."

reach
"OTUD5 expression was enhanced by IFN-gamma through a p38 and MAPK dependent mechanism and the AS induced knockdown of OTUD5 in LPMCs of IBD patients and colitic mice reduced TNF-alpha."
IFNG activates OTUD5.
| 1 1
IFNG activates OTUD5. 2 / 2
| 1 1

eidos
"OTUD5 expression was enhanced by IFN-gamma through a p38 / MAPK-dependent mechanism and the AS-induced knockdown of OTUD5 in LPMCs of IBD patients and colitic mice reduced TNF-alpha ."

reach
"Recently, OTUD5 was found to be up-regulated in the intestinal inflammatory tissues of IBD patients and TNBS-induced colitis mice, and IFN-γ was found to up-regulate OTUD5’s expression through a p38/MAPK-dependent mechanism."

reach
"Also known as DUBA (deubiquitinating enzyme A), the OTUD5 protein was shown to suppress type-I interferon (IFN-I) dependent innate immune response, by cleaving the poly-ubiquitin chain from the IFN-I adaptor protein, thus causing the disassociation of the adaptor from the downstream signaling complex, and ultimately the interruption of the IFN-I signaling cascade."

reach
"Three human DUBs, CYLD, A20 and DUBA, have been shown to negatively regulate the innate immune response by removing K63 based chains XREF_BIBR, XREF_BIBR, and USP15 inhibits the NFkappaB pathway by removing K48-Ub from IkappaBalpha, preventing its degradation XREF_BIBR."

reach
"OTUD5 has been shown to suppress the type I interferon dependent innate immune response by cleaving the polyubiquitin chain from an essential type I interferon adaptor protein."

reach
"DUBA, a member of the ovarian tumor (OTU) domain containing cysteine protease superfamily, was shown to suppress the type I IFN dependent innate immune response by cleaving the K63 polyubiquitin chain on TRAF3."

reach
"It has been reported that OTUD5 could suppress the type I interferon dependent innate immune response by cleaving the polyubiquitin chain from an essential type I interferon adaptor protein TRAF3."
| 1

reach
"In the innate immune response, OTUD5 can promote the stability of STING [37]."
| 2 4
| 2 2

reach
"The function of OTUD5 is required to allow the rapid activation of p53 dependent transcription and a p53 dependent apoptosis in response to DNA damage stress."

eidos
"It was proved that OTUD5 inhibited the proliferation and metastasis of NSCLC cells but promoted their apoptosis via regulating p53 and PDCD5 ."
| PMC

reach
"To study the cellular consequence of specifically blocking the interaction between OTUD5 and FACT (OTUD5 overexpression or complete knockout causes aberration in cell cycle and apoptosis), we generated a HeLa knock-in (KI) cell line in which the D537A mutation is introduced into the genomic loci of OTUD5 (OTUD5 ) using the CRISPR-Cas9 method (Supplementary Figure S2; sequencing information of 4 clones)."

eidos
"Knockdown of OTUD5 accelerated the proliferation , migration , and invasion of A549 cells , and inhibited their apoptosis Next , A549 cells were used for the following experiment ."
| PMC
| 2

reach
"Knockdown of OTUD5 clearly impeded the apoptosis, consistent with a weaker activation of p53 (XREF_FIG)."

reach
"Moreover, overexpression of OTUD5 attenuated podocyte apoptosis (Fig. 1m) and recovered the expression levels of Nephrin in podocytes (Fig. 1n) with HG/PA treatment."
OTUD5 affects PTEN
| 6
OTUD5 increases the amount of PTEN.
| 2
OTUD5 increases the amount of PTEN. 2 / 2
| 2

reach
"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We further detected the protein levels of PTEN in A549 and HCI-H460 cells with another Western blot assay and uncovered that overexpression of OTUD5 promoted the protein level of PTEN, while PTEN protein levels were further increased on treatment with MG132 (a proteasome inhibitor) (p < 0.05, Figure 3C), indicating that OTUD5 may regulate PTEN expression through the proteasome pathway."
OTUD5 deubiquitinates PTEN.
| 2
OTUD5 deubiquitinates PTEN. 2 / 2
| 2

reach
"Nevertheless, whether miR-652-3p regulates that the OTUD5/PTEN axis has not been discussed before and warrants further exploration.In light of the abovementioned evidence, we put forth a hypothesis that OTUD5 deubiquitinates and stabilizes PTEN to suppress NSCLC cell proliferation, invasion, and migration, whereas this mechanism could be counteracted by miR-652-3p overexpression."

reach
"The obtained findings revealed that OTUD5 deubiquitinated and stabilized PTEN to suppress NSCLC proliferation, invasion, and migration, and miR-652-3p serving as an upstream target of OTUD5 abolished the tumor-suppressing role of OTUD5/PTEN in NSCLC.There is a plethora of evidence to highlight the correlation between decreased levels of OTUD5 and poor prognoses in NSCLC patients, such that OTUD5 knockdown exerts an enhancing effect on NSCLC cell proliferation, migration, and chemoresistance [13]."
OTUD5 decreases the amount of PTEN.
| 2
OTUD5 decreases the amount of PTEN. 2 / 2
| 2

reach
"Furthermore, we quantified the ubiquitination levels of PTEN in A549 and HCI-H460 cells and documented that OTUD5 overexpression reduced the ubiquitination level of PTEN (Figure 3D)."

reach
"Subsequent experimentation revealed that miR-652-3p inhibition brought about an increase in OTUD5 expression levels (p < 0.05, Figure 6B and C) and PTEN protein levels (p < 0.05, Figure 6D and E), while simultaneously reducing the ubiquitination level of PTEN, cell viability (p < 0.05, Figure 6F), and the number of migrated and invaded cells (p < 0.05, Figure 6G and H)."
OTUD5 affects OTUD5
| 6
OTUD5 increases the amount of OTUD5.
| 2
OTUD5 increases the amount of OTUD5. 2 / 2
| 2

reach
"DUBA inhibited the development of severe DSS colitis through downregulation of type I IFN responses, as knockdown of DUBA expression by administration of DUBA-specific siRNA cancelled the therapeutic effects of MDP injection via upregulation of type I IFN responses."

reach
"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 decreases the amount of OTUD5.
| 2
OTUD5 decreases the amount of OTUD5. 2 / 2
| 2

reach
"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Altogether, the abovementioned findings and valuable evidence highlighted that OTUD5 exerts an anti-tumor function by means of deubiquitinating and stabilizing PTEN, while enhancing the ubiquitination of PTEN reverses the inhibition of OTUD5 overexpression on NSCLC progression.Furthermore, we focused our efforts on the upstream targets of OTUD5."
OTUD5 deubiquitinates OTUD5.
| 1
OTUD5 deubiquitinates OTUD5. 1 / 1
| 1

reach
"These DUBs have been reported to be involved in regulating various biological processes in fish, including regulating the innate immune system.The deubiquitinating enzyme A (DUBA), also known as ovari[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 activates OTUD5.
| 1
OTUD5 activates OTUD5. 1 / 1
| 1

reach
"Conversely, in gastric cancer cells harbouring p53 mutations, OTUD5 transcription is upregulated due to the inactivation of p53, allowing OTUD5 to stabilise GPX4 protein, inhibit ferroptosis and thereby promoting gastric cancer progression (Figure 6J, right panel)."
STING1 affects OTUD5
2 | 3
2 | 3

sparser
"USP11 exacerbates radiation induced pneumonitis by activating endothelial cell inflammatory response via OTUD5-STING signaling."

sparser
"Here, we report that the deubiquitinase OTUD5 interacts with STING, cleaves its K48-linked polyubiquitin chains, and promotes its stability."

sparser
"OTUD5 interacts with STING, cleaves its K48-linked polyubiquitin chain and promotes its stability ( xref B) [ xref ]."

reach
"Specifically, podocyte-specific overexpression of OTUD5 potentially serves as a feasible strategy to inhibit podocyte inflammation by down-regulating TAK1 activity, ultimately controlling the progression of DKD."

reach
"Lipid accumulation and inflammation were exacerbated by Otud5 knockdown but attenuated by Otud5 overexpression under PAOA treatment."

reach
"OTUD5 overexpression in primary hepatocytes inhibited PAOA-induced hepatocyte lipid accumulation and inflammation (Figure 5A–D)."

reach
"Therefore, we hypothesize that OTUD5 negatively regulates inflammation and injury through deubiquitinating TAK1 in podocytes.We confirmed the endogenous interaction between OTUD5 and TAK1 in podocytes using a Co-immunoprecipitation (co-IP) assay (Fig. 4c)."

reach
"Macrophage-specific ablation of DUBA attenuated MDP-induced systematic inflammation and liver injury in mice."
| 5

reach
"OTUD5 promotes innate antiviral and antitumor immunity through deubiquitinating and stabilizing STING."

reach
"This is highlighted in antigen-stimulated macrophages, in which pSer177 DUBA levels are increased to regulate the immune response."

reach
"25 OTUD5 enhanced anti‐viral infection and anti‐tumor immunity by stabilizing STING."

reach
"Our previous study indicated that OTUD5 promotes innate antiviral and antitumor immunity by removing K48-linked ubiquitin chains of STING and strengthening its stability [33]."

reach
"The experimental data obtained from knockout mouse models clearly indicate that the lack of OTUD5 leads to heightened vulnerability to viral infections and accelerated melanoma progression, highlighting the crucial function of OTUD5 in STING-mediated immunity [21]."
| 3

reach
"Intriguingly, this separation-of-function mutant (OTUD5 DeltaCterm) failed to support neural crest differentiation and showed aberrant CNS precursor formation (XREF_FIG and fig."
| PMC

reach
"Previous research has demonstrated that the ability of OTUD5 to cleave K48‐ubiquitin chains is required to support differentiation and that, during embryonic development, OTUD5 regulates cell fate decisions by cleaving the degrading K48‐ubiquitin junctions of a substrate (Beck et al., 2021; Park et al., 2015; Yin et al., 2019)."

reach
"Thus, OTUD5 controls neural cell fate commitment by regulating chromatin accessibility at neural- and neural crest specific enhancers to enable activation of transcriptional networks that drive the differentiation program."
| PMC
| 2

reach
"Indeed, knockout of Otud5 is embryonic lethal in mice and OTUD5 depleted hESCs are defective in neuroectodermal differentiation, which can be rescued by re-expression of wild-type OTUD5 [XREF_BIBR]."

reach
"DUBA inhibits Th17 differentiation by stabilizing UBR5, which mediates ubiquitination and degradation of RORγt."
OTUD5 affects TRAF2
1 | 3 1
OTUD5 deubiquitinates TRAF2.
1 | 2
OTUD5 deubiquitinates TRAF2. 2 / 3
1 | 2

reach
"We found that K63-ubiquitylation of TRAF2 was increased by OTUD5 knockdown and decreased by TRIP12 knockdown (Fig. 3h)."

reach
"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."
OTUD5 binds TRAF2.
| 1 1
| 1 1

reach
"Indeed, we found that TRAF2 associated with OTUD5 in cells (Fig. 3g)."

sparser
"Indeed, we found that TRAF2 associated with OTUD5 in cells (Fig.  xref )."
OTUD5 affects RIGI
| 5
OTUD5 inhibits RIGI.
| 3
OTUD5 inhibits RIGI. 3 / 3
| 3

reach
"In addition, A20, NLRX1, PIN1, DUBA and Triad3A deregulate RIG-I mediated signaling by inhibiting MAVS, IRF3 and TRAF3."

reach
"For example, DUBA, a member of the Otubain (OTUB) family, has been shown to deubiquitinate TRAF3 and negatively regulate TLR3- and RIG-I and MDA5-mediated IFN induction."

reach
"The deubiquitinating enzyme A (DUBA), a novel OTU-domain DUB has been shown to negatively regulate IFN signaling following RIG-I, MDA-5 or TLR3 stimulation [60] ."
OTUD5 deubiquitinates RIGI.
| 2
OTUD5 deubiquitinates RIGI. 2 / 2
| 2

reach
"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."

reach
"Other DUBs that target the RIG-I and MDA5 pathway include CYLD and DUBA which target RIG-I and TRAF3 ubiquitination respectively (XREF_FIG)."
OTUD5 affects MTOR
| 5
OTUD5 activates MTOR.
| 4
OTUD5 activates MTOR. 4 / 4
| 4

reach
"Interestingly, OTUD5 positively regulates the mTOR pathway by removing ubiquitin molecules from the ubiquitin ligases β-TrCP1 and RNF186, thereby stabilizing both proteins."

reach
"OTUD5 knockdown inhibited the cell proliferation, and OTUD5 positively regulated the mTOR signaling pathway to promote cell proliferation."

reach
"In breast cancer and colon cancer, OTUD5 promotes carcinogenesis by regulating the mammalian target of rapamycin (mTOR) and Hippo signaling pathways [10,11,12]."

reach
"In bladder cancer, OTUD5 activates mechanistic target of rapamycin complex 1 (mTORC1) signalling by deubiquitinating and stabilizing ring finger protein 186 (RNF186) and promoting sestrin2 degradation [10]."
OTUD5 inhibits MTOR.
| 1
OTUD5 inhibits MTOR. 1 / 1
| 1

reach
"The knockdown of OTUD5 has caused several mTOR-related phenotypes, such as a decrease in cell size and an increase in autophagy in mammalian cells."
OTUD5 affects HDAC2
| 4 1
| 2 1

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC

reach
"S5), we show that OTUD5 interacts with and targets ARID1A/B and HDAC2, proteins that are mutated in Coffin-Siris and Cornelia de Lange syndromes, respectively."
| PMC

sparser
"HDAC2 was a notable interactor, as OTUD5 also interacted with HDAC2 in a recent report ( xref )."
| 2

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them."

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them (Figure 7)."
OTUD5 affects DNA repair
| 5
| 5

reach
"In addition to its role in promoting the signaling axis of the DDR, we found that OTUD5 promotes the efficiency of DNA repair exclusively at telomeres and heterochromatin, but not in general, non-specific chromatin contexts."

reach
"Taken together, we conclude that OTUD5 is required both for efficient NHEJ and for efficient MMEJ at deprotected telomeres.Having established that OTUD5 promotes DNA repair by NHEJ and MMEJ at telomeres, we assessed if the role of OTUD5 in DNA repair extends beyond a telomeric context, especially since we found OTUD5 to promote both telomere-specific and genome-wide DDR activation in mouse and human cells."

reach
"Moreover, we found that OTUD5 promotes DNA repair at deprotected telomeres and at heterochromatin."

reach
"If OTUD5 indeed promotes ATM-dependent DDR and telomeric DNA repair through the UBR5-ATMIN axis, ablation of ATMIN is expected to reverse the DDR defects exhibited by OTUD5-depleted cells."

reach
"Hence, we next considered the possibility that KAP1, the central factor of our screening approach, is itself of key importance for how OTUD5 promotes efficient DNA repair activity at telomeres."
OTUD5 affects DDR
| 5
OTUD5 activates DDR. 5 / 5
| 5

reach
"Indeed, we found OTUD5 to promote ATM-dependent DDR signaling in a manner that is epistatic with UBR5, dependent on its catalytic activity and involving ATMIN, supporting that OTUD5 acts via UBR5.Early biochemical studies performed on OTUD5 suggested that phosphorylation at residue Ser-177 is vital for its DUB activity in vitro ."

reach
"If OTUD5 indeed promotes ATM-dependent DDR and telomeric DNA repair through the UBR5-ATMIN axis, ablation of ATMIN is expected to reverse the DDR defects exhibited by OTUD5-depleted cells."

reach
"OTUD5 promotes ATM-dependent DDR signaling."

reach
"As independently, UBR5 was found to inhibit the interaction between ATM and ATMIN, a negative regulator of canonical DSB-induced ATM activation , we hypothesized that OTUD5 may promote ATM-dependent DDR signaling via stabilization of UBR5."

reach
"OTUD5 promotes ATM-dependent DDR activation by regulating the UBR5-ATMIN axis."
OTUD5 affects Colitis
| 5
OTUD5 inhibits Colitis.
| 3
| 3

reach
"DUBA inhibited the development of severe DSS colitis through downregulation of type I IFN responses, as knockdown of DUBA expression by administration of DUBA-specific siRNA cancelled the therapeutic effects of MDP injection via upregulation of type I IFN responses."

reach
"In addition, DUBA deficiency in macrophages rendered mice hypersensitive to DSS-induced colitis and eliminated the protective effect of MDP treatment in colitis."

reach
"Furthermore, administration of DUBA antisense oligonucleotides (ASOs) attenuates the development of trinitrobenzene sulfonic acid (TNBS) colitis via downregulation of TNF-α expression."
OTUD5 activates Colitis.
| 2
OTUD5 activates Colitis. 2 / 2
| 2

reach
"Alternatively, knockdown of DUBA expression by ASO may inhibit the development of TNBS colitis through downregulation of pathogenic IL-17 production, since DUBA has been shown to function as a posttranslational brake against IL-17 production by T cells."

reach
"As such, increased OTUD5 aggravates DSS-induced colitis [ 100 ]."
OTUD5 affects BRINP1
| 5
OTUD5 binds BRINP1.
| 3
| 3

reach
"Interestingly, we further confirmed that SIAH2 and OTUD5 competitively bind to DBC1 at the same N-terminal region, and hypoxia promotes the interaction of DBC1 with SIAH2 rather than OTUD5, resulting in ubiquitination and degradation of DBC1 to promote tumor progression."

reach
"Moreover, we still need to further investigate the details of how the OTUD5-DBC1 complex dissociates under hypoxia to modulate DBC1 stability."

reach
"Co-IP analysis demonstrated the exogenous interaction between OTUD5 and DBC1 (Figure 4—figure supplement 1D and E)."
OTUD5 inhibits BRINP1.
| 1
OTUD5 inhibits BRINP1. 1 / 1
| 1

reach
"Consistently, hypoxia-induced endogenous DBC1 degradation was specifically inhibited by expression of wildtype OTUD5 rather than the inactive mutant (Figure 4E)."
OTUD5 activates BRINP1.
| 1
OTUD5 bound to BRINP1 activates BRINP1. 1 / 1
| 1

reach
"Moreover, we still need to further investigate the details of how the OTUD5-DBC1 complex dissociates under hypoxia to modulate DBC1 stability."
HDAC2 affects OTUD5
| 4 1
| 2 1

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC

reach
"S5), we show that OTUD5 interacts with and targets ARID1A/B and HDAC2, proteins that are mutated in Coffin-Siris and Cornelia de Lange syndromes, respectively."
| PMC

sparser
"HDAC2 was a notable interactor, as OTUD5 also interacted with HDAC2 in a recent report ( xref )."
| 2

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them."

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them (Figure 7)."
TRIM28 affects OTUD5
| 4
TRIM28 activates OTUD5.
| 3
TRIM28 activates OTUD5. 3 / 3
| 3

reach
"Depletion of OTUD5 in TRF2ts MEFs with 2 independent sgRNAs (g1, g1B) recapitulated the pKAP1 defect observed with shRNA-mediated knockdown of OTUD5 (Fig. 1i, j)."

reach
"Moreover, ectopic expression of phosphomimic KAP1 also rescued the telomere MMEJ defect in OTUD5-depleted Trf1 ;Trf2 ;Ku70 ;p53 ;Cre-ER MEFs (Fig. 5g, Supplementary Fig. 7d, e)."

reach
"Indeed, overexpression of KAP1 restored telomere fusion efficiency in OTUD5-depleted cells, while ectopic expression of wild-type (KAP1 ) failed to do so (Fig. 5e, f, Supplementary Fig. 7b, c)."
TRIM28 inhibits OTUD5.
| 1
TRIM28 inhibits OTUD5. 1 / 1
| 1

reach
"Next, we assessed the potential functional involvement of UBR5 in OTUD5-promoted KAP1 phosphorylation, by depleting OTUD5, UBR5, or both in U2OS cells."
OTUD5 affects TNF-α production
| 4
OTUD5 activates TNF-α production. 4 / 4
| 4

reach
"Given that DUBA deubiquitinates K63-liked polyubiquitination of TRAF3 and thereby inhibits type I IFN responses, the administration of DUBA ASOs is likely to augment the NF-κB dependent TNF-α production."

reach
"In contrast to this hypothesis, Dinnalo’s data suggest that DUBA is an inducer of TNF-α production."

reach
"Up-regulation of OTUD5 further increased TNF-α production in LPMCs of IBD patients (160)."

reach
"Knockdown of DUBA expression by its antisense oligonucleotides reduces TNF-α production by colonic lamina propria mononuclear cells isolated from mice challenged with intrarectal TNBS administration (74)."
OTUD5 affects TLR
| 4
OTUD5 activates TLR. 4 / 4
| 4

reach
"Some others are deubiquitinases (DUBs), such as A20 (also known as TNFAIP3), CYLD (cylindromatosis D), and DUBA (deubiquitinating enzyme A), which negatively modulate TLR signaling by removing the K63[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These results suggested that OTUD5 enhances TLR/IL-1R-mediated NF-κB and MAPK signaling."

reach
"Taken together, these findings revealed that OTUD5 stimulates TLR/IL-1R-mediated production of proinflammatory cytokines.Since TLR/IL-1R-mediated activation of NF-κB and MAPKs lead to the production of inflammatory cytokines [45, 46], we next explored the role of OTUD5 in the NF-κB and MAPK signaling pathways by transfecting Otud5-siRNA into PMs."

reach
"OTUD5 promotes TLR/IL1-R-mediated NF-kappaB and MAPK signaling."
OTUD5 affects Neoplasms
| 2 2
OTUD5 inhibits Neoplasms.
| 1 1
| 1 1

reach
"Thus, OTUD5 knockdown was able to accelerate tumor growth in a nude mouse model."

eidos
"Briefly , OTUD5 potentially inhibits tumor progression by regulating PML transcription by deubiquitinating TRIM25 ( Fig. 8g ) ."
OTUD5 activates Neoplasms.
| 1 1
| 1 1

reach
"25 OTUD5 enhanced anti‐viral infection and anti‐tumor immunity by stabilizing STING."

eidos
"Here , we report that the deubiquitinating enzyme , ovarian tumor domain-containing protein 5 ( OTUD5 ) , can activate the mTOR signaling pathway , promote cancer progression , and show its oncogenic potential in bladder cancer ."

reach
"Overexpression of OTUD5 suppresses NSCLC cell proliferation, invasion, and migration."

reach
"Overall, these findings suggested that overexpression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."

reach
"On the other hand, over-expression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."
| 1

reach
"To elaborate our understanding of OTUD5 functionality in NSCLC cells, we further overexpressed OTUD5 in A549 and HCI-H460 cells and learnt that OTUD5 overexpression led to a reduction in cell viability, migration, and invasion."
OTUD5 affects NSCLC
| 4
OTUD5 inhibits NSCLC. 4 / 4
| 4

reach
"Overall, these findings suggested that overexpression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."

reach
"Overexpression of OTUD5 suppresses NSCLC cell proliferation, invasion, and migration."

reach
"Li et al. found that OTUD5 stabilizes PTEN through deubiquitination and that the overexpression of OTUD5 inhibits NSCLC cell proliferation, invasion and migration [9] (Figure 3E)."

reach
"On the other hand, over-expression of OTUD5 suppressed NSCLC cell proliferation, invasion, and migration."
OTUD5 affects NEDD4
| 2 2
OTUD5 binds NEDD4.
| 1 2
| 1 2

sparser
"Second, an interaction between OTUD5 and NEDD4 was indeed detected."

sparser
"Next, Co‐IP assays revealed that OTUD5 interacted with NEDD4 exogenously (Figure  xref ), and 1–220 residues of OTUD5 were required for the interaction with NEDD4 (Figure xref , Supporting Information)."

reach
"Second, an interaction between OTUD5 and NEDD4 was indeed detected."
OTUD5 activates NEDD4.
| 1
OTUD5 activates NEDD4. 1 / 1
| 1

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"In contrast, OTUD5 knockout in HEK293T cells impaired the binding of NEDD4 and α‐Syn (Figure 4H)."
OTUD5 affects AKT
| 4
OTUD5 decreases the amount of AKT.
| 2
OTUD5 decreases the amount of AKT. 2 / 2
| 2

reach
"OTUD5 overexpression has been shown to significantly downregulate the phosphorylation level of Akt in cervical cancer cells."

reach
"Interestingly, OTUD5 overexpression also significantly downregulated the ubiquitination level of Akt in cervical cancer cells, suggesting that OTUD5 may be a deubiquitinating enzyme for Akt [73] (Table 1)."
OTUD5 ubiquitinates AKT.
| 1
OTUD5 leads to the ubiquitination of AKT. 1 / 1
| 1

reach
"Accordingly, the enhanced Akt ubiquitination and activation caused by the downregulation of OTUD5 gives rise to radioresistance in cervical cancer [119]."
OTUD5 inhibits AKT.
| 1
OTUD5 inhibits AKT. 1 / 1
| 1

reach
"Interestingly, OTUD5 overexpression also significantly downregulated the ubiquitination level of Akt in cervical cancer cells, suggesting that OTUD5 may be a deubiquitinating enzyme for Akt [73] (Table 1)."
MTOR affects OTUD5
| 2 2
MTOR phosphorylates OTUD5.
| 1 2
MTOR phosphorylates OTUD5. 3 / 3
| 1 2

sparser
"A recent study revealed that OTUD5 can be phosphorylated by mTOR and activate its catalytic activity and is positively correlated with the mTOR signalling pathway [ xref ]."

sparser
"We also showed that mTOR directly phosphorylated OTUD5 and activated its DUB activity."

reach
"A recent study revealed that OTUD5 can be phosphorylated by mTOR and activate its catalytic activity and is positively correlated with the mTOR signalling pathway [13]."
MTOR activates OTUD5.
| 1
MTOR activates OTUD5. 1 / 1
| 1

reach
"To date, little is known about the kinases that phosphorylate OTUD5, apart from recent in vivo evidence that mTOR kinase promotes OTUD5 stability and activity via direct phosphorylation of multiple sites on OTUD5."
XRCC5 affects OTUD5
| 2 1
| 2 1

reach
"In addition, the new autoantigen reported here, the deubiquitinase OTUD5, associates with Ku80 to regulate its function ."

sparser
"OTUD5 interacts with Ku80 to positively regulate non-homologous end joining NHEJ-mediated repair ( xref )."

reach
"OTUD5 interacts with Ku80 to positively regulate non homologous end joining NHEJ mediated repair."
TLR9 affects OTUD5
| 3
TLR9 increases the amount of OTUD5.
| 2
TLR9 increases the amount of OTUD5. 2 / 2
| 2

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"The MDP-NOD2 axis downregulates TLR9-induced ISG by upregulating DUBA expression (67, 68)."

reach
"Intact NOD2 plays a negative regulatory function in TLR9-induced type I IFN responses by upregulating DUBA expression (67, 68)."
TLR9 decreases the amount of OTUD5.
| 1
TLR9 decreases the amount of OTUD5. 1 / 1
| 1

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"Intact NOD2 plays a negative regulatory function in TLR9-induced type I IFN responses by upregulating DUBA expression (67, 68)."
| 2

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"Using an inhibitor of ERK1/2, we demonstrated OTUD5 inhibited ERK1/2 MAPK signaling pathway to accumulate HNF4α expression to benefit HBV replication.In this study, the healthy cohort, HBV carriers and chronic hepatitis B patients were enrolled, the expression OTUD5 in different groups was analyzed, and significant differences of OTUD5 were observed between the HBV-negative cohort and HBV positive carriers, meaning the expression of OTUD5 in human serum was promoted by infecting hepatitis B virus."

reach
"As the RLRs signaling pathway were negatively regulated by DUBA and miR-210 could target and regulate DUBA, therefore, miR-210 could indirectly positively modulate the immune responses in miiuy croake[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 1

reach
"OTUD5 knockdown inhibited the cell proliferation, and OTUD5 positively regulated the mTOR signaling pathway to promote cell proliferation."
OTUD5 affects polyubiquitin chain
| 3
OTUD5 inhibits polyubiquitin chain. 3 / 3
| 3

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"It has been reported that OTUD5 could suppress the type I interferon dependent innate immune response by cleaving the polyubiquitin chain from an essential type I interferon adaptor protein TRAF3."

reach
"OTUD5 (DUBA) is activated by phosphorylation ( Huang et al., 2012 ) and suppresses the type I interferon response by cleaving the polyubiquitin chain from tumor necrosis factor receptor-associated fac[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"DUBA, a member of the ovarian tumor (OTU) domain containing cysteine protease superfamily, was shown to suppress the type I IFN dependent innate immune response by cleaving the K63 polyubiquitin chain on TRAF3."
OTUD5 affects polB
| 3
OTUD5 inhibits polB.
| 2
OTUD5 inhibits polB. 2 / 2
| 2

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"By inhibiting the FACT-mediated nucleosomal rearrangement and subsequent dimer exchange, which may be possibly reflected in the H2A deposition assay (Figure 5D), OTUD5 may prevent the efficient Pol II progression nearby the damaged chromatin."

reach
"OTUD5 localizes to DNA double strand breaks (DSBs), interacts with UBR5 and represses the RNA Pol II elongation and RNA synthesis."
OTUD5 activates polB.
| 1
OTUD5 activates polB. 1 / 1
| 1

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"These results altogether suggest that UBR5 and OTUD5 induce transcriptional silencing at DSB lesions by controlling the PolII access."

reach
"OTUD5 promotes NHEJ repair by increasing the stability of Ku80 [52] (Figure 2E)."

reach
"This suggests the possibility that OTUD5 is directly phosphorylated by ATM to function as an amplifier of ATM-dependent DDR signaling in a potential positive-feedback circuit, resembling its role in mTOR signaling.In the context of DNA repair, OTUD5 has been suggested to promote NHEJ by counteracting the degradation of Ku80, a core NHEJ factor, in certain cell lines ."

reach
"OTUD5 also stabilises Ku80, loss of OTUD5 reduces NHEJ repair efficiency [11] (Figure 2)."
OTUD5 affects TRAF6
1 | 2
OTUD5 deubiquitinates TRAF6.
1 | 1
OTUD5 deubiquitinates TRAF6. 1 / 2
1 | 1

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"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."
OTUD5 activates TRAF6.
| 1
OTUD5 activates TRAF6. 1 / 1
| 1

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"Moreover, the adaptor proteins TRAF3 and TRAF6 are targeted by the cellular proteins DUBA and A20 XREF_BIBR, XREF_BIBR, and TBK1 is sequestered by SIKE XREF_BIBR."
OTUD5 affects TNF
| 3
OTUD5 decreases the amount of TNF.
| 2
OTUD5 decreases the amount of TNF. 2 / 2
| 2

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"Podocyte deficiency of OTUD5 also increased the mRNA levels of Il6 and Tnfa in renal tissues of T2DM mice (Fig. 2m, n)."

reach
"Similarly, deficiency of OTUD5 in podocytes enhanced inflammatory Il6 and Tnfa gene expression in T1DM mouse kidneys (Fig. 3k, l)."
OTUD5 binds TNF.
| 1
| 1

reach
"DUBA bound tumor necrosis factor receptor associated factor 3 (TRAF3), an adaptor protein essential for the IFN-I response."
| 3
| 3

reach
"These results suggested that OTUD5 enhances TLR/IL-1R-mediated NF-κB and MAPK signaling."

reach
"OTUD5 promotes TLR/IL1-R-mediated NF-kappaB and MAPK signaling."

reach
"These findings demonstrated that knockdown of OTUD5 suppresses LPS-induced NF-κB and MAPK signaling in human and mouse macrophages."
OTUD5 affects IRF3
| 2 1
OTUD5 inhibits IRF3. 3 / 3
| 2 1

reach
"These results suggest that DUBA inhibits IRF3 by functioning as a K63 specific DUB that antagonizes the function of TRAF3."

reach
"Removal of Lys 63 -linked polyubiquitin chains from RIG-I and TRAF3 by cylindromatosis (CYLD) and deubiquitinating enzyme A (DUBA), respectively, inhibits the RIG-I-dependent activation of IRF3."

sparser
"These results suggest that DUBA inhibits IRF3 by functioning as a K63-specific DUB that antagonizes the function of TRAF3."
OTUD5 affects IL17A
| 1 2
OTUD5 inhibits IL17A.
| 1 1
OTUD5 inhibits IL17A. 2 / 2
| 1 1

eidos
"OTUD5 inhibits the production of IL-17A by blocking the UBR5-mediated proteasomal degradation of RORgammat17 ."

reach
"T-cell specific loss of DUBA therefore results in reduced levels of UBR5, stabilization of RORgammaT, and increased IL-17A production in response to TCR stimulation 76."
OTUD5 decreases the amount of IL17A.
| 1
OTUD5 decreases the amount of IL17A. 1 / 1
| 1

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"Accumulation of the deubiquitinating enzyme DUBA (OTUD5) negatively regulates RORgammat stability and IL-17 expression in T cells by stabilizing the ubiquitin ligase UBR5, which ubiquitinates RORgammat in response to TGF-beta, leading to RORgammat degradation and decreased IL-17 expression."
OTUD5 affects HNF1A
| 3
OTUD5 activates HNF1A. 3 / 3
| 3

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"In the present study, we discovered that overexpression of OTUD5 promoted HNF1α/HNF4α protein and mRNA expression while down-expression of OTUD5 inhibited HNF1α/HNF4α protein and mRNA expression in hepG2 2.15 cells (Fig. 6A–E)."

reach
"The data of PCR showed that knockdown of OTUD5 significantly downregulated the mRNA of HNF1ɑ and HNF4ɑ (Fig. 6B,C)."

reach
"Conversely, the overexpression of OTUD5 could upregulate the mRNA of HNF1ɑ and HNF4ɑ (Fig. 6E–F)."
OTUD5 affects GST
| 3
| 3

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"Recombinant GST-OTUD5 WT, but not GST-OTUD5 ΔUIM interacted with SPT16, further confirming that the UIM is necessary for the interaction (Figure xref )."

sparser
"We thus extended this finding with bacterially purified GST-OTUD5 proteins; both full-length OTUD5 and a C-terminal fragment of OTUD5 (residues 501–571) similarly co-purified SPT16 (Figure xref ), suggesting that the C-terminus is sufficient to bind FACT."
OTUD5 affects FYN
| 3
| 3

sparser
"To further explore how OTUD5 promoted α‐Syn degradation, we examined whether OTUD5 interacted with α‐Syn."

sparser
"Co‐immunoprecipitation (Co‐IP) assay clearly showed the interaction between OTUD5 and α‐Syn ( Figure   xref )."

sparser
"OTUD5 Interacted with α‐Syn and Promoted Its K63‐Linked Polyubiquitination."
OTUD5 affects ERK
| 3
OTUD5 phosphorylates ERK.
| 1
OTUD5 phosphorylates ERK. 1 / 1
| 1

reach
"In addition, Otud5 knockdown was associated with decreased phosphorylation of ERK, JNK, and P38 in the MAPK signaling pathway (Fig. S1D)."
OTUD5 inhibits ERK.
| 1
OTUD5 inhibits ERK. 1 / 1
| 1

reach
"Using an inhibitor of ERK1/2, we demonstrated OTUD5 inhibited ERK1/2 MAPK signaling pathway to accumulate HNF4α expression to benefit HBV replication.In this study, the healthy cohort, HBV carriers and chronic hepatitis B patients were enrolled, the expression OTUD5 in different groups was analyzed, and significant differences of OTUD5 were observed between the HBV-negative cohort and HBV positive carriers, meaning the expression of OTUD5 in human serum was promoted by infecting hepatitis B virus."
OTUD5 decreases the amount of ERK.
| 1
OTUD5 decreases the amount of ERK. 1 / 1
| 1

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"In this study, we proved OTUD5 could downregulate the expression of phosphorylation ERK1/2 in hepG2.2.15 cells, and the results illustrated that OTUD5 could promote the expression of HNF4α via inhibiting ERK1/2 of MAPK pathway (Fig. 7A–D)."
OTUD5 affects Dnmt1
| 3
| 3

sparser
"The OTUD5 UIM domain can bind both Lys 48 - and Lys 63 -linked polyubiquitin chains xref , xref ."

sparser
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."

sparser
"The stimulator of interferon genes (STING), suppressor of ty 16 homolog (SPT16), TRIM25, TP53 and PDCD5 have all been reported to interact with the UIM structural domain of OTUD5 [ xref , xref , xref ]."
| 3
| 3

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"To discern whether OTUD5 could mitigate GPX4-mediated cell ferroptosis and enhance cell survival, we knocked down OTUD5 in HK2 cells using siRNA."

reach
"Specifically, the role of OTUD5 in stabilizing GPX4 suggests that therapeutic modulation of its activity might be a feasible strategy for controlling GPX4 levels, thus manipulating the course of ferroptosis and ultimately, the progression of AKI.Through a series of experiments, we demonstrate that OTUD5 plays a protective role against ferroptosis and contributes to cell survival."

reach
"To elaborate our understanding of OTUD5 functionality in NSCLC cells, we further overexpressed OTUD5 in A549 and HCI-H460 cells and learnt that OTUD5 overexpression led to a reduction in cell viability, migration, and invasion."
| 1 2
OTUD5 activates Carcinogenesis.
| 2

reach
"In breast cancer and colon cancer, OTUD5 promotes carcinogenesis by regulating the mammalian target of rapamycin (mTOR) and Hippo signaling pathways [10,11,12]."

reach
"OTUD5 acts as a tumor inhibitor in non-small cell lung cancer and cervical cancer.36 37 However, OTUD5 promotes carcinogenesis in triple-negative breast cancer and bladder cancer.38 39 Hence, the specific function of OTUD5 in different cancers needs to be analyzed on a case-to-case basis."
OTUD5 inhibits Carcinogenesis.
| 1

eidos
"Our findings reveal a mechanism whereby OTUD5 regulates gene transcription and suppresses tumorigenesis by deubiquitinating TRIM25 , providing a potential target for oncotherapy ."
NEDD4 affects OTUD5
| 1 2
| 1 2

sparser
"Second, an interaction between OTUD5 and NEDD4 was indeed detected."

sparser
"Next, Co‐IP assays revealed that OTUD5 interacted with NEDD4 exogenously (Figure  xref ), and 1–220 residues of OTUD5 were required for the interaction with NEDD4 (Figure xref , Supporting Information)."

reach
"Second, an interaction between OTUD5 and NEDD4 was indeed detected."
Hypoxia affects OTUD5
| 3
Hypoxia inhibits OTUD5.
| 1
| 1

reach
"Hypoxia triggers autophagic degradation of OTUD5."
Hypoxia decreases the amount of OTUD5.
| 1
Hypoxia decreases the amount of OTUD5. 1 / 1
| 1

reach
"Our findings indicated that hypoxia inhibited mTORC1 activity as well as the protein levels of OTUD5 and GPX4 (Fig. 6d), suggesting that H/R-induced OTUD5 reduction occurs at the early stage of hypoxia."
Hypoxia activates OTUD5.
| 1
Hypoxia activates OTUD5. 1 / 1
| 1

reach
"Interestingly, we further confirmed that SIAH2 and OTUD5 competitively bind to DBC1 at the same N-terminal region, and hypoxia promotes the interaction of DBC1 with SIAH2 rather than OTUD5, resulting in ubiquitination and degradation of DBC1 to promote tumor progression."
GST affects OTUD5
| 3
| 3

sparser
"To test whether OTUD5 also interacts with SPT16 in an unbiased method, we performed FLAG-OTUD5 IP and GST-OTUD5 pulldown experiments from HeLa S3 cells and subjected the eluate to mass spec analysis (Figure xref )."

sparser
"Recombinant GST-OTUD5 WT, but not GST-OTUD5 ΔUIM interacted with SPT16, further confirming that the UIM is necessary for the interaction (Figure xref )."

sparser
"We thus extended this finding with bacterially purified GST-OTUD5 proteins; both full-length OTUD5 and a C-terminal fragment of OTUD5 (residues 501–571) similarly co-purified SPT16 (Figure xref ), suggesting that the C-terminus is sufficient to bind FACT."
FYN affects OTUD5
| 3
| 3

sparser
"To further explore how OTUD5 promoted α‐Syn degradation, we examined whether OTUD5 interacted with α‐Syn."

sparser
"Co‐immunoprecipitation (Co‐IP) assay clearly showed the interaction between OTUD5 and α‐Syn ( Figure   xref )."

sparser
"OTUD5 Interacted with α‐Syn and Promoted Its K63‐Linked Polyubiquitination."
Dnmt1 affects OTUD5
| 3
| 3

sparser
"The OTUD5 UIM domain can bind both Lys 48 - and Lys 63 -linked polyubiquitin chains xref , xref ."

sparser
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."

sparser
"The stimulator of interferon genes (STING), suppressor of ty 16 homolog (SPT16), TRIM25, TP53 and PDCD5 have all been reported to interact with the UIM structural domain of OTUD5 [ xref , xref , xref ]."
CACYBP affects OTUD5
1 | 1 1
CACYBP binds OTUD5.
1 | 1
1 | 1

sparser
"Further mechanistic studies revealed that CacyBP interacts with the tumour suppressor OTUD5, enhances the ubiquitination and proteasomal degradation of OTUD5, and regulates tumorigenesis via OTUD5."
CACYBP activates OTUD5.
| 1
CACYBP activates OTUD5. 1 / 1
| 1

reach
"CacyBP promotes the development of lung adenocarcinoma by regulating OTUD5."
BRINP1 affects OTUD5
| 3
| 3

reach
"Interestingly, we further confirmed that SIAH2 and OTUD5 competitively bind to DBC1 at the same N-terminal region, and hypoxia promotes the interaction of DBC1 with SIAH2 rather than OTUD5, resulting in ubiquitination and degradation of DBC1 to promote tumor progression."

reach
"Moreover, we still need to further investigate the details of how the OTUD5-DBC1 complex dissociates under hypoxia to modulate DBC1 stability."

reach
"Co-IP analysis demonstrated the exogenous interaction between OTUD5 and DBC1 (Figure 4—figure supplement 1D and E)."
ATM affects OTUD5
| 2 1
ATM phosphorylates OTUD5.
| 1 1
ATM phosphorylates OTUD5. 2 / 2
| 1 1

sparser
"This suggests the possibility that OTUD5 is directly phosphorylated by ATM to function as an amplifier of ATM-dependent DDR signaling in a potential positive-feedback circuit, resembling its role in mTOR signaling."

reach
"This suggests the possibility that OTUD5 is directly phosphorylated by ATM to function as an amplifier of ATM-dependent DDR signaling in a potential positive-feedback circuit, resembling its role in mTOR signaling.In the context of DNA repair, OTUD5 has been suggested to promote NHEJ by counteracting the degradation of Ku80, a core NHEJ factor, in certain cell lines ."
ATM binds OTUD5.
| 1
| 1

reach
"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
Vasotocin affects OTUD5
| 1 1
| 1 1

reach
"XREF_FIG d, AVT also decreases Otud4, Otud5 and Otud7b in addition to Otub1 and c-Maf."

eidos
"As shown in Fig. 5d , AVT also decreases Otud4 , Otud5 and Otud7b in addition to Otub1 / c-Maf ."
Type p53 affects OTUD5
| 2
Type p53 increases the amount of OTUD5.
| 1
Type p53 increases the amount of OTUD5. 1 / 1
| 1

reach
"OTUD5 mediates GPX4 deubiquitination to regulate its stability.Deletion of OTUD5 promotes ferroptosis and inhibits tumor growth.Wild type p53 inhibits OTUD5 transcription, thereby promoting GPX4 degradation and inhibiting the development of gastric cancer.OTUD5, GPX4 expression and p53 activity are highly correlated and correlates with clinical progression in STAD."
Type p53 decreases the amount of OTUD5.
| 1
Type p53 decreases the amount of OTUD5. 1 / 1
| 1

reach
"OTUD5 mediates GPX4 deubiquitination to regulate its stability.Deletion of OTUD5 promotes ferroptosis and inhibits tumor growth.Wild type p53 inhibits OTUD5 transcription, thereby promoting GPX4 degradation and inhibiting the development of gastric cancer.OTUD5, GPX4 expression and p53 activity are highly correlated and correlates with clinical progression in STAD."
Region affects OTUD5
| 2
Region bound to 1 and SIAH2 activates OTUD5. 1 / 1
| 1

reach
"Domain mapping analysis further revealed that the N-terminal region (amino acid residues 1–230) of DBC1, which binds with SIAH2, was also necessary for its interaction with OTUD5 (Figure 4H)."
Region bound to SIAH2 activates OTUD5. 1 / 1
| 1

reach
"Domain mapping analysis further revealed that the N-terminal region (amino acid residues 1–230) of DBC1, which binds with SIAH2, was also necessary for its interaction with OTUD5 (Figure 4H)."
Polyubiquitin chains affects OTUD5
| 2
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 binds polyubiquitin chains and K48. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
Pan affects OTUD5
| 2
Pan ubiquitinates OTUD5. 2 / 2
| 2

sparser
"Therefore, we verified cooperative actions of TRIP12 and UBR5 in OTUD5 pan-ubiquitylation using TUBE2."

sparser
"Using TUBE2 (tandem ubiquitin-binding entity 2), which serves as a pan-ubiquitin chain binder when four repeats of the UBA1 domains from HR23A are tandemly fused xref , we found that OTUD5 pan-ubiquitylation was enhanced by the expression of wild-type, but not catalytically inactive (C1959A), TRIP12 (Fig.  xref )."
MiR-652-3p affects OTUD5
| 2
MiR-652-3p inhibits OTUD5.
| 1
MiR-652-3p inhibits OTUD5. 1 / 1
| 1

reach
"Above all, our findings demonstrated that miR-652-3p may abolish the tumor-suppressing role of OTUD5/PTEN, thereby facilitating NSCLC cell proliferation, invasion, and migration."
MiR-652-3p decreases the amount of OTUD5.
| 1
MiR-652-3p decreases the amount of OTUD5. 1 / 1
| 1

reach
"MiR-652-3p inhibits OTUD5 expression."
| 1 1

eidos
"As OTUD5 is induced by LPS stimulation , it might act as a negative feedback regulator of IFN signaling by targeting TRAF3 [ 119 ] ."

reach
"As DUBA protein is also increased in macrophages by LPS stimulation, it may also function as part of a negative feedback circuit."
HgpA affects OTUD5
| 2
HgpA decreases the amount of OTUD5. 2 / 2
| 2

reach
"This finding identifies OTUD5 as an inhibitor of TAK1 activation, which may serve as an alternative therapeutic strategy to regulate TAK1 hyperactivation.A limitation of this study may be the unclear mechanism by which diabetes or HG/PA downregulates OTUD5 expression in podocytes."

reach
"The protein expression of OTUD5 was also decreased by HG/PA stimulation in a time-dependent manner in podocytes (Fig. 1d, Supplementary Fig. 1b)."
Doxorubicin affects OTUD5
| 2
Doxorubicin inhibits OTUD5.
| 1
| 1

reach
"As shown in XREF_FIG after Doxorubicin treatment depletion of OTUD5 clearly prevented the efficient deubiquitination of p53 that is associated with protein accumulation (XREF_FIG)."
Doxorubicin activates OTUD5.
| 1
| 1

reach
"U2OS cells transfected with scramble shRNA or OTUD5 shRNA were treated with Doxorubicin."
ZGPAT affects OTUD5
| 1 1
| 1 1

sparser
"However, their DUB activity toward K29-linked diubiquitin was much weaker (Fig.  xref ), suggesting that K29-linked ubiquitin chains were relatively resistant to DUB activity of ARMC7–OTUD5 or ZGPATOTUD5 complexes."

reach
"Indeed, we found that ARMC7 and ZGPAT, whose protein levels are co-regulated by TRIP12 and UBR5 (Fig. 6b), stably associated with OTUD5 in cells (Fig. 6c)."
UBD affects OTUD5
| 1 1
| 1 1

sparser
"So far, no crystal structures of diubiquitin bound to DUBA are available and therefore the structural basis for ubiquitin chain linkage specificity is not known."

reach
"So far, no crystal structures of diubiquitin bound to DUBA are available and therefore the structural basis for ubiquitin chain linkage specificity is not known."
TRAF2 affects OTUD5
| 1 1
| 1 1

reach
"Indeed, we found that TRAF2 associated with OTUD5 in cells (Fig. 3g)."

sparser
"Indeed, we found that TRAF2 associated with OTUD5 in cells (Fig.  xref )."
SYNM affects OTUD5
| 2
SYNM inhibits OTUD5.
| 1
SYNM inhibits OTUD5. 1 / 1
| 1

reach
"The data provided direct evidence that OTUD5 reduced the protein levels of α‐Syn by promoting its degradation.2.2 OTUD5 Promoted Endolysosomal Degradation of alpha-Syn."
SYNM binds OTUD5.
| 1
| 1

reach
"Besides, an association between endogenous OTUD5 and α‐Syn was evident in SH‐SY5Y cells (Figure 3D,E)."
SMARCA4 affects OTUD5
| 1 1
| 1 1

reach
"Further supporting the idea that OTUD5 regulates cell fate commitment through stabilizing chromatin remodelers, we observed that OTUD5 regulated neural enhancers were often bound by SMARCA4, a component of the BAF complex recruited by ARID1A/B in neural progenitor cells (XREF_FIG)."
| PMC

sparser
"Further supporting the idea that OTUD5 regulates cell fate commitment through stabilizing chromatin remodelers, we observed that OTUD5-regulated neural enhancers were often bound by SMARCA4 ( xref ), a component of the BAF complex recruited by ARID1A/B ( xref ) in neural progenitor cells ( xref )."
| PMC
SLC7A11 affects OTUD5
| 2
| 2

sparser
"Mechanistically, OTUD5 interacted with SLC7A11 and cleaved K48-linked polyubiquitin chains from SLC7A11 to enhance the stability of SLC7A11."

sparser
"88 Furthermore, differentiated CSCs can counter ferroptosis by increasing the levels of the deubiquitinase DUBA, which interacts with SLC7A11 and stabilizes SLC7A11 protein levels through deubiquitina[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
RNF146 affects OTUD5
| 1 1
RNF146 ubiquitinates OTUD5. 2 / 2
| 1 1

sparser
"Nie et al. found that RNF146 can ubiquitinate OTUD5 and lead to its degradation by the proteasome [ xref ]."

reach
"Nie et al. found that RNF146 can ubiquitinate OTUD5 and lead to its degradation by the proteasome [63]."
RING domain affects OTUD5
| 2
OTUD5 binds TRAF-C region and RING domain. 1 / 1
| 1

reach
"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 binds TRAF-C and RING domain. 1 / 1
| 1

reach
"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
Protease affects OTUD5
| 2
Protease increases the amount of OTUD5.
| 1
Protease increases the amount of OTUD5. 1 / 1
| 1

reach
"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Protease decreases the amount of OTUD5.
| 1
Protease decreases the amount of OTUD5. 1 / 1
| 1

reach
"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2
OTUD5 translocates to the cytoplasm. 1 / 1
| 1

reach
"After binding to HER-2 and being internalized, the cleavable linker binding both components of the ADC is cleaved in the lysosome, and the active drug DUBA is released in the cytoplasm [94]."
OTUD5 translocates to the replication fork. 1 / 1
| 1

reach
"We found that OTUD5 is recruited to replication forks, and its depletion causes replication fork stress."
OTUD5 affects transformation malignant A549 cells
| 2
OTUD5 inhibits transformation malignant A549 cells. 2 / 2
| 2

eidos
"Therefore , it was concluded that OTUD5 impeded the transformation of malignant phenotypes of A549 cells via positively regulating p53 expression ."
| PMC

eidos
"This study demonstrated that OTUD5 could inhibit the transformation of malignant phenotypes of A549 cells by inducing the expression of p53 , suggesting that OTUD5 may be used as a potential therapeutic target for NSCLC treatment ."
| PMC
OTUD5 affects polyubiquitin chains
| 2
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 binds polyubiquitin chains and K48. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 affects migration invasion HCC827 cells
| 2
OTUD5 inhibits migration invasion HCC827 cells. 2 / 2
| 2

eidos
"Therefore , it was concluded that OTUD5 could impede the proliferation , migration , and invasion of HCC827 cells while facilitating their apoptosis , and the role of OTUD5 was partly mediated by PDCD5 ."
| PMC

eidos
"Knockdown of OTUD5 promoted the proliferation , migration , and invasion of HCC827 cells while inhibiting their apoptosis via regulating PDCD5 ."
| PMC
OTUD5 affects migration invasion A549 cells
| 2
OTUD5 inhibits migration invasion A549 cells. 2 / 2
| 2

eidos
"Knockdown of OTUD5 accelerated the proliferation , migration , and invasion of A549 cells , and inhibited their apoptosis Next , A549 cells were used for the following experiment ."
| PMC

eidos
"Knockdown of OTUD5 accelerated the proliferation , migration , and invasion of A549 cells while inhibiting their apoptosis ."
| PMC
OTUD5 affects mTORC1
| 2
OTUD5 activates mTORC1. 2 / 2
| 2

reach
"Prior research has demonstrated that OTUD5 modulates mTORC1 activity by destabilizing its inhibitory core proteins ."

reach
"In bladder cancer, OTUD5 activates mechanistic target of rapamycin complex 1 (mTORC1) signalling by deubiquitinating and stabilizing ring finger protein 186 (RNF186) and promoting sestrin2 degradation [10]."
OTUD5 affects doxorubicin
| 2
| 2

reach
"In addition, OTUD5 knockdown also enhances the resistance of NSCLC cells to doxorubicin and cisplatin [8]."

reach
"OTUD5 knockdown also enhanced the resistance of NSCLC cells to doxorubicin and cisplatin."
OTUD5 affects cisplatin
| 2
| 2

reach
"OTUD5 knockdown also enhanced the resistance of NSCLC cells to doxorubicin and cisplatin."

reach
"In addition, OTUD5 knockdown also enhances the resistance of NSCLC cells to doxorubicin and cisplatin [8]."
OTUD5 affects ZGPAT
| 1 1
| 1 1

sparser
"However, their DUB activity toward K29-linked diubiquitin was much weaker (Fig.  xref ), suggesting that K29-linked ubiquitin chains were relatively resistant to DUB activity of ARMC7–OTUD5 or ZGPATOTUD5 complexes."

reach
"Indeed, we found that ARMC7 and ZGPAT, whose protein levels are co-regulated by TRIP12 and UBR5 (Fig. 6b), stably associated with OTUD5 in cells (Fig. 6c)."
OTUD5 affects UBD
| 1 1
| 1 1

sparser
"So far, no crystal structures of diubiquitin bound to DUBA are available and therefore the structural basis for ubiquitin chain linkage specificity is not known."

reach
"So far, no crystal structures of diubiquitin bound to DUBA are available and therefore the structural basis for ubiquitin chain linkage specificity is not known."
OTUD5 affects TLR3
| 2
OTUD5 inhibits TLR3. 2 / 2
| 2

reach
"The deubiquitinating enzyme A (DUBA), a novel OTU-domain DUB has been shown to negatively regulate IFN signaling following RIG-I, MDA-5 or TLR3 stimulation [60] ."

reach
"For example, DUBA, a member of the Otubain (OTUB) family, has been shown to deubiquitinate TRAF3 and negatively regulate TLR3- and RIG-I and MDA5-mediated IFN induction."
OTUD5 affects SPT16
| 2
OTUD5 inhibits SPT16.
| 1
OTUD5 inhibits SPT16. 1 / 1
| 1

reach
"Consistent with the roles of Fanconi Anemia proteins in CFS stability (50) and limiting aberrant R-loop formations (21,22), depletion of FANCD2 was synthetically lethal with loss of FACT regulation in the OTUD5 cells.A previous study showed that depleting SPT16 increases DNA–RNA hybrids and genomic instability in yeast and human cells (35)."
OTUD5 binds SPT16.
| 1
| 1

reach
"Further, the interaction between OTUD5 and SPT16 can be detected in situ by the PLA (Figure 4E)."
OTUD5 affects SMARCA4
| 1 1
| 1 1

reach
"Further supporting the idea that OTUD5 regulates cell fate commitment through stabilizing chromatin remodelers, we observed that OTUD5 regulated neural enhancers were often bound by SMARCA4, a component of the BAF complex recruited by ARID1A/B in neural progenitor cells (XREF_FIG)."
| PMC

sparser
"Further supporting the idea that OTUD5 regulates cell fate commitment through stabilizing chromatin remodelers, we observed that OTUD5-regulated neural enhancers were often bound by SMARCA4 ( xref ), a component of the BAF complex recruited by ARID1A/B ( xref ) in neural progenitor cells ( xref )."
| PMC
OTUD5 affects SLC7A11
| 2
| 2

sparser
"Mechanistically, OTUD5 interacted with SLC7A11 and cleaved K48-linked polyubiquitin chains from SLC7A11 to enhance the stability of SLC7A11."

sparser
"88 Furthermore, differentiated CSCs can counter ferroptosis by increasing the levels of the deubiquitinase DUBA, which interacts with SLC7A11 and stabilizes SLC7A11 protein levels through deubiquitina[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects RNF186
| 2
OTUD5 deubiquitinates RNF186. 2 / 2
| 2

reach
"OTUD5 can activate the mTOR signalling pathway by deubiquitinating ring finger protein 186 (RNF186), thereby promoting the progression of bladder cancer [10]."

reach
"The specific mechanism is that OTUD5 deubiquitinates RNF186 and stabilizes its protein levels, which, in turn, leads to the increased degradation of Sestrin2 and the subsequent activation of the mTOR pathway [11]."
OTUD5 affects RLH
| 2
OTUD5 inhibits RLH. 2 / 2
| 2

reach
"Consistent with these findings, a close family member of DUBA, A20, and known inhibitor of NF-κB signaling, also inhibits RLH signaling."

reach
"Recently, a deubiquitinase named DUBA was shown to deubiquitinate TRAF3 and suppress RLH signaling [ 31 • ]."
OTUD5 affects RING domain
| 2
OTUD5 binds TRAF-C region and RING domain. 1 / 1
| 1

reach
"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 binds TRAF-C and RING domain. 1 / 1
| 1

reach
"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
OTUD5 affects PML
| 2
OTUD5 decreases the amount of PML. 2 / 2
| 2

reach
"By reducing the ubiquitination level of TRIM25, OTUD5 suppresses promyelocytic leukemia protein (PML) expression [7] (Figure 3C)."

reach
"Further investigation reveals that OTUD5 depletion leads to the enhanced transcriptional activity of TRIM25 and the inhibited expression of PML by altering the ubiquitination level of TRIM25."
OTUD5 affects NSCLC cells
| 2
OTUD5 inhibits NSCLC cells. 2 / 2
| 2

eidos
"* p < .05 , * * p < .01 , and * * * p < .001 [ Colour figure can be viewed at wileyonlinelibrary.com ] OTUD5 impeded the progression of NSCLC cells via regulating p53 In order to study whether OTUD5 regulates p53 expression in NSCLC , OTUD5 siRNA and NC siRNA were transfected into A549 cells , respectively ."
| PMC

eidos
"OTUD5 impeded the progression of NSCLC cells via regulating p53 ."
| PMC
OTUD5 affects MAP1LC3
| 1 1
| 1 1

reach
"An immunoprecipitation assay showed an increased interaction between LC3 and OTUD5 under H/R conditions (Fig. 5j)."

sparser
"An immunoprecipitation assay showed an increased interaction between LC3 and OTUD5 under H/R conditions (Fig.  xref )."
OTUD5 affects K63
| 2
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

reach
"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
OTUD5 affects IHC
| 2
OTUD5 activates IHC. 2 / 2
| 2

reach
"OTUD5 knockdown significantly reduced the IHC score of Ki-67 in xenograft tumour tissues (P < 0.05, Fig. 4C)."

reach
"OTUD5 knockdown significantly reduced the IHC score of SLC38A1 in xenograft tumour tissues (P < 0.05, Supplementary Fig. 2)."
OTUD5 affects IFNB1
| 2
OTUD5 activates IFNB1. 2 / 2
| 2

reach
"DUBA knockdown by siRNA transfection in Il1r1 -/- BMDCs (XREF_FIG) resulted in significantly increased IL-10 and IFN-beta production and decreased secretion of IL-6, but not TNF, in CpG stimulated Il1r1 -/- BMDCs (XREF_FIG)."

reach
"The removal of endogenous DUBA by siRNA produced an elevated IFN-β and IFN-α response to poly:IC or Sendai virus, whereas NF-κB activation was predominantly unaffected ( Kayagaki et al., 2007 )."
OTUD5 affects IFN-1
| 2
OTUD5 inhibits IFN-1.
| 1
OTUD5 inhibits IFN-1. 1 / 1
| 1

reach
"In the study showing that OTUD5 suppresses the type I interferon (IFN-1) production, the proposed mechanism is that OTUD5 antagonizes the TRAF3 signaling (21)."
OTUD5 decreases the amount of IFN-1.
| 1
OTUD5 decreases the amount of IFN-1. 1 / 1
| 1

reach
"The first report of function of OTUD5 is to negatively regulate IFN-1 expression by cleaving the polyubiquitin chains on TRAF3 (137)."
OTUD5 affects HDAC1
| 2
| 2

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them."

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them (Figure 7)."
OTUD5 affects Figs
| 2
OTUD5 inhibits Figs.
| 1
OTUD5 inhibits Figs. 1 / 1
| 1

reach
"The CHX assay indicated that OTUD5 overexpression prolonged the half-life of YAP ( Fig. 3 F), while OTUD5 depletion reduced the protein stability of YAP ( Figs."
OTUD5 activates Figs.
| 1
OTUD5 activates Figs. 1 / 1
| 1

reach
"The CHX assay indicated that OTUD5 overexpression prolonged the half-life of YAP ( Fig. 3 F), while OTUD5 depletion reduced the protein stability of YAP ( Figs."
| 2
| 2

reach
"Given that TRAF3 may be dispensable for RIG-I signalling XREF_BIBR, it can not be excluded, however, that DUBA also targets other E3 ligases like TRAF2, 5 and 6."

reach
"Mechanistically, OTUD5 promotes K63-linked polyubiquitination of alpha-Syn independent of its deubiquitinating enzyme activity and mediates its endolysosomal degradation by recruiting the E3 ligase neural precursor cell expressed developmentally downregulated 4 (NEDD4)."

reach
"These results suggest that the aberrant Pol II elongation near DSB sites caused by UBR5 or OTUD5 deficiency depend on the FACT activity."

reach
"These results suggest that both OTUD5 and UBR5 repress transcription nearby DSB lesions."
OTUD5 affects CACYBP
1 | 1
1 | 1

sparser
"Further mechanistic studies revealed that CacyBP interacts with the tumour suppressor OTUD5, enhances the ubiquitination and proteasomal degradation of OTUD5, and regulates tumorigenesis via OTUD5."
OTUD5 affects ARID1A/B
| 2
OTUD5 binds ARID1A/B. 2 / 2
| 2

reach
"S5), we show that OTUD5 interacts with and targets ARID1A/B and HDAC2, proteins that are mutated in Coffin-Siris and Cornelia de Lange syndromes, respectively."
| PMC

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC

reach
"Furthermore, administration of DUBA antisense oligonucleotides (ASOs) attenuates the development of trinitrobenzene sulfonic acid (TNBS) colitis via downregulation of TNF-α expression."

reach
"Alternatively, knockdown of DUBA expression by ASO may inhibit the development of TNBS colitis through downregulation of pathogenic IL-17 production, since DUBA has been shown to function as a posttranslational brake against IL-17 production by T cells."
MAP1LC3 affects OTUD5
| 1 1
| 1 1

reach
"An immunoprecipitation assay showed an increased interaction between LC3 and OTUD5 under H/R conditions (Fig. 5j)."

sparser
"An immunoprecipitation assay showed an increased interaction between LC3 and OTUD5 under H/R conditions (Fig.  xref )."
K63 affects OTUD5
| 2
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

reach
"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
Ischemia affects OTUD5
| 2
| 2

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"OTU deubiquitinase 5 (OTUD5), a protein that interacts with GPX4, can promote ferroptosis resistance during ischemia/reperfusion injury by stabilizing GPX4 expression; in turn, hypoxia/ischemia-induced OTUD5 autophagy can destabilize GPX4, leading to ferroptosis-dependent kidney injury (96)."

reach
"Overall, this study highlights a new autophagy-dependent ferroptosis module: hypoxia/ischemia-induced OTUD5 autophagy triggers GPX4 degradation, offering a potential therapeutic avenue for I/R-related kidney diseases."
IL1B affects OTUD5
| 2
IL1B increases the amount of OTUD5. 2 / 2
| 2

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"IFN-γ and IL-1β have been shown to upregulate and downregulate expression of DUBA, respectively."

reach
"Given the negative regulation of DUBA expression by IL-1β, impaired IL-1 receptor signaling pathways may be attributed to increased DUBA expression in the remission phases of CD."
IL-1RI affects OTUD5
| 2
IL-1RI decreases the amount of OTUD5. 2 / 2
| 2

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"Consistent with the protein data, Duba mRNA levels were increased in freshly isolated splenocytes and mesenteric LN derived cells from Il1r1 -/- mice (XREF_FIG), suggesting that IL-1RI signaling negatively regulates DUBA transcription."

reach
"We show that IL-1RI signaling down-regulates the expression of deubiquitinating enzyme A (DUBA) and consequently enhances the Lys63 linked ubiquitination of TNF receptor associated factor 3 (TRAF3), which is necessary for the transcription of antiinflammatory cytokines."
HDAC2 affects HDAC1
| 2
| 2

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"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them."

reach
"Based on the data, we propose a model where the OTUD5-HDAC1-HDAC2 complex modulates FACT recruitment to nucleosomes by deacetylating them (Figure 7)."
BACH2 affects OTUD5
| 2
BACH2 increases the amount of OTUD5. 2 / 2
| 2

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"Whether the transcription factor Bach2 directly regulates the expression of OTUD5 needs to be determined."

reach
"In CD4 T lymphocytes from chronic pancreatitis tissues, the knockdown of the transcription factor Bach2 downregulates OTUD5 expression [81]."
ATG5 affects OTUD5
| 2
ATG5 inhibits OTUD5.
| 1
ATG5 inhibits OTUD5. 1 / 1
| 1

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"Further validating these findings, we revealed that genetic knockdown of autophagy regulator ATG5 prevented H/R-induced OTUD5 and GPX4 loss (Fig. 5h)."
ATG5 activates OTUD5.
| 1
ATG5 activates OTUD5. 1 / 1
| 1

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"Further validating these findings, we revealed that genetic knockdown of autophagy regulator ATG5 prevented H/R-induced OTUD5 and GPX4 loss (Fig. 5h)."
ARID1A/B affects OTUD5
| 2
OTUD5 binds ARID1A/B. 2 / 2
| 2

reach
"S5), we show that OTUD5 interacts with and targets ARID1A/B and HDAC2, proteins that are mutated in Coffin-Siris and Cornelia de Lange syndromes, respectively."
| PMC

reach
"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC
Proteasome inhibitor decreases the amount of OTUD5. 1 / 1
| 1

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"The proteasome inhibitor MG-132 was also found to increase the protein level of OTUD5 in T cells, suggesting that OTUD5 is degraded by the proteasome in the absence of TCR stimulation [6]."
Polyubiquitin affects OTUD5
| 1
OTUD5 binds polyubiquitin. 1 / 1
| 1

sparser
"DUBA physically binds with TRAF3 that is an E3 ubiquitin ligase and could assemble lysine-63–linked polyubiquitin chains [ 8 ]."
Polyubiquitin chains affects K63
| 1
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

reach
"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
Polyubiquitin chains affects K48
| 1
OTUD5 binds polyubiquitin chains and K48. 1 / 1
| 1

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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
| 1
P53 activator decreases the amount of OTUD5. 1 / 1
| 1

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"Moreover, the p53 activator nutlin-3a suppresses OTUD5 transcription, leading to GPX4 degradation and ferroptosis of gastric cancer cells."
P38 affects OTUD5
| 1
P38 activates OTUD5. 1 / 1
| 1

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"To further investigate the role of ERK1/2 and p38 signaling in OTUD5-mediated promotion of HNF4ɑ or HNF1ɑ, we treated the OTUD5-sh HepG2.2.15 cells with ERK1/2-specific pharmacological inhibitor (PD98059), JNK-specific pharmacological inhibitor (SP600125) and p38-specific pharmacological inhibitor (SB203580), and then tested the effect of OTUD5 on the protein level of HNF1α and HNF4α."
MiR-210 affects OTUD5
| 1
MiR-210 activates OTUD5. 1 / 1
| 1

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"As the RLRs signaling pathway were negatively regulated by DUBA and miR-210 could target and regulate DUBA, therefore, miR-210 could indirectly positively modulate the immune responses in miiuy croake[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
MTORC1 affects OTUD5
| 1
MTORC1 increases the amount of OTUD5. 1 / 1
| 1

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"Instead of the mTORC1 inhibitor rapamycin, the mTORC1 activator MHY1485 significantly restored the levels of OTUD5 and GPX4 reduced by H/R (Fig. 6c), suggesting that mTORC1 plays a beneficial role in maintaining OTUD5 homeostasis."
Ligand affects OTUD5
| 1
Ligand increases the amount of OTUD5. 1 / 1
| 1

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"The stimulation of bone marrow-derived dendritic cells (BMDCs) from WT and Il1r1 mice with CpG, a ligand for TLRs, showed that CpG induced higher levels of the OTUD5 protein in Il1r1 compared to WT BMDC."
Hsa-mir-3149 affects OTUD5
| 1
Hsa-mir-3149 decreases the amount of OTUD5. 1 / 1
| 1

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"Moreover, hsa-mir-137, hsa-mir-1913, hsa-mir-937, hsa-mir-607, hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5."
Hsa-mir-144 affects OTUD5
| 1
Hsa-mir-144 decreases the amount of OTUD5. 1 / 1
| 1

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"Moreover, hsa-mir-137, hsa-mir-1913, hsa-mir-937, hsa-mir-607, hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5."
Hsa-mir-137 affects OTUD5
| 1
Hsa-mir-137 decreases the amount of OTUD5. 1 / 1
| 1

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"Moreover, hsa-mir-137, hsa-mir-1913, hsa-mir-937, hsa-mir-607, hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5."
| 1
Dapagliflozin decreases the amount of OTUD5. 1 / 1
| 1

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"We proved that dapagliflozin decreased the expression level of OTU deubiquitinase 5 (OTUD5), which further increased the ubiquitination and degradation of YAP1."
Autophagy affects OTUD5
| 1
| 1

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"During I/R, ferroptosis is induced by mTORC1-mediated autophagy, causing OTUD5 degradation and subsequent GPX4 decay."
ZUP1 affects OTUD5
| 1
ZUP1 activates OTUD5. 1 / 1
| 1

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"In this study, we identified an elegant mechanism to counteract deubiquitylation that allows a DUB, OTUD5, to be efficiently targeted for degradation."
YAP1 affects OTUD5
| 1
OTUD5 binds YAP1 and 3 L. 1 / 1
| 1

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"Indeed, TNBC cells enhanced the binding between OTUD5 and YAP ( Fig. 3 L), and the ubiquitination assays showed the ubiquitin chain of YAP in co-culture group was dramatically removed ( Fig. 3 M)."
TRAF3 affects K63
| 1
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

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"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
TRAF-C affects RING domain
| 1
OTUD5 binds TRAF-C and RING domain. 1 / 1
| 1

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"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
TRAF-C region affects RING domain
| 1
OTUD5 binds TRAF-C region and RING domain. 1 / 1
| 1

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"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
TNF affects OTUD5
| 1
| 1

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"DUBA bound tumor necrosis factor receptor associated factor 3 (TRAF3), an adaptor protein essential for the IFN-I response."
TGFB affects OTUD5
| 1
TGFB activates OTUD5. 1 / 1
| 1

eidos
"In immunomodulation , TGF-beta stimulation leads to excessive OTUD5 production ."
TAK1-TAB2 affects OTUD5
| 1
TAK1-TAB2 binds OTUD5. 1 / 1
| 1

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"These findings elucidate that OTUD5 inhibits TAK1 activation by blocking TAK1-TAB2 interaction via OTUD5-mediated TAK1 deubiquitination at the K158 site."

reach
"TRAF3 activation is negatively regulated by Suppressor of Cytokine Signalling 3 (SOCS3) and Deubiquitinating Enzyme A (DUBA) [XREF_BIBR]."
SPT16 affects OTUD5
| 1
| 1

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"Further, the interaction between OTUD5 and SPT16 can be detected in situ by the PLA (Figure 4E)."

reach
"Additionally, we demonstrated that I/R downregulates OTUD5 through the activation of lysosomal degradation controlled by the mammalian target of rapamycin complex 1 (mTORC1)."
RSL3 affects OTUD5
| 1
RSL3 activates OTUD5. 1 / 1
| 1

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"The use of the GPX4 inhibitor RSL3 triggered physiological cell ferroptosis and eliminated the protective effects of OTUD5 under H/R conditions (Fig. 4f), thus confirming the essential nature of the OTUD5/GPX4 axis in mediating renal tubular cell ferroptosis."
RING domain affects TRAF-C
| 1
OTUD5 binds TRAF-C and RING domain. 1 / 1
| 1

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"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
RING domain affects TRAF-C region
| 1
OTUD5 binds TRAF-C region and RING domain. 1 / 1
| 1

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"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
PTEN affects OTUD5
| 1
PTEN increases the amount of OTUD5. 1 / 1
| 1

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"Increasing ubiquitination level of PTEN reverses the inhibition of OTUD5 overexpression on NSCLC cell proliferation, invasion, and migration."

reach
"We hypothesized that OTUD5 could promote HBV viral transcription and replication through interactions with individual viral proteins."
| 1
| 1

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"25 OTUD5 enhanced anti‐viral infection and anti‐tumor immunity by stabilizing STING."

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"Downregulation of OTUD5 reduces TNF production in lamina propria mononuclear cells (LPMCs), indicating that OTUD5 may exacerbate IBD by inducing proinflammatory cytokine production [ 101 ]."
OTUD5 affects tumor growth
| 1
OTUD5 inhibits tumor growth. 1 / 1
| 1

eidos
"Importantly , OTUD5 knockdown accelerates tumor growth in a nude mouse model ."
| 1
| 1

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"In contrast, OTUD5 overexpression in primary hepatocytes inhibited PAOA-induced lipid accumulation and reduced TG concentration (Figure 4C–I)."
OTUD5 affects transcriptional
| 1
OTUD5 inhibits transcriptional. 1 / 1
| 1

eidos
"OTUD5 inhibits transcriptional activity of TRIM25 by downregulating the ubiquitination level of TRIM25 but the question remains : how exactly does ubiquitination of TRIM25 relate to its functions ?"
OTUD5 affects receptor-interacting protein kinase 2
| 1
OTUD5 increases the amount of receptor-interacting protein kinase 2. 1 / 1
| 1

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"DUBA potentiated NOD2-mediated signal transduction by maintaining the protein levels of NOD2 and receptor-interacting protein kinase 2 (RIPK2)."
OTUD5 affects radiosensitivity cervical cancer cells
| 1
OTUD5 activates radiosensitivity cervical cancer cells. 1 / 1
| 1

eidos
"A recent study demonstrated that OTUD5 could increase the radiosensitivity of cervical cancer cells ( Yin et al ., 2019 ) , which was consistent with the results of this study ."
| PMC
OTUD5 affects proteolysis
| 1
| 1

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"38 We utilized ATG5 knockout cells to investigate whether OTUD5 mediated the protein degradation of α‐Syn through the autophagic pathway."
OTUD5 affects protein SFN
| 1
OTUD5 activates protein SFN. 1 / 1
| 1

eidos
"In addition , the ubiquitination level of MAVS was reduced following OTUD5 coexpression , and OTUD5 increased the protein level of SFN , demonstrating that OTUD5 was possibly involved in these signaling pathways to work with the TRIM25 E3 ubiquitin ligase in the protein complex ( Supplementary Fig. 5 ) ."

eidos
"Studies have shown that OTUD5 can activate the programmed cell death 5 ( PDCD5 , a tumor suppressor ; Park et al ., 2015 ) ."
| PMC
OTUD5 affects polyubiquitin
| 1
OTUD5 binds polyubiquitin. 1 / 1
| 1

sparser
"DUBA physically binds with TRAF3 that is an E3 ubiquitin ligase and could assemble lysine-63–linked polyubiquitin chains [ 8 ]."
OTUD5 affects poly I:C
| 1
| 1

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"In miiuy croaker, DUBA was up-regulated post poly I:C treatment, indicating DUBA was associated with the innate immune response [ 14 ]."

reach
"To directly evaluate the physiological role of OTUD5, myeloid-specific OTUD5 knockout mice were produced by breeding Otud5 mice with Lyz2-Cre mice."
OTUD5 affects paoA
| 1
OTUD5 inhibits paoA. 1 / 1
| 1

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"A qRT-PCR analysis revealed that OTUD5 knockdown in primary hepatocytes contributed to PAOA-induced hepatocyte lipid accumulation and inflammation."
OTUD5 affects p38
| 1
OTUD5 phosphorylates p38. 1 / 1
| 1

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"In addition, Otud5 knockdown was associated with decreased phosphorylation of ERK, JNK, and P38 in the MAPK signaling pathway (Fig. S1D)."

reach
"We have previously shown that the FACT histone chaperone, necessary for remodeling nucleosomes to allow for transcription, is negatively regulated by OTUD5 upon nuclease induction of DSBs (37)."
OTUD5 affects miR-210
| 1
OTUD5 activates miR-210. 1 / 1
| 1

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"In conclusion, these results indicated that miR-210 can target DUBA 3′UTR and regulate the luciferase activity in a dose- and time-dependent manner.To certify the prediction, we cloned miiuy croaker p[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects metastasis cells
| 1
OTUD5 inhibits metastasis cells. 1 / 1
| 1

eidos
"It was proved that OTUD5 inhibited the proliferation and metastasis of NSCLC cells but promoted their apoptosis via regulating p53 and PDCD5 ."
| PMC
OTUD5 affects metastasis NSCLC cells
| 1
OTUD5 inhibits metastasis NSCLC cells. 1 / 1
| 1

eidos
"The knockdown of OTUD5 inactivated p53 and PDCD5 , promoting the proliferation and metastasis of NSCLC cells while inhibiting their apoptosis ."
| PMC
| 1
OTUD5 increases the amount of malonaldehyde. 1 / 1
| 1

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"The results showed that OTUD5 overexpression restored the increase in iron and MDA levels and the decrease in GSH caused by MI/R."
OTUD5 affects maleate(2-)
| 1
| 1

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"Recently, a male specific multiple congenital anomaly disorder caused by pathogenic variants in OTUD5 was reported by two groups."
OTUD5 affects mTOR signaling pathway
| 1
OTUD5 activates mTOR signaling pathway. 1 / 1
| 1

eidos
"Here , we report that the deubiquitinating enzyme , ovarian tumor domain-containing protein 5 ( OTUD5 ) , can activate the mTOR signaling pathway , promote cancer progression , and show its oncogenic potential in bladder cancer ."
| 1
| 1

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"Furthermore, confocal microscopic analysis visualized that the co‐localization of α‐Syn and LAMP1, a marker of lysosomes, was significantly enhanced by OTUD5 overexpression (Figure 2B)."
OTUD5 affects iron atom
| 1
OTUD5 increases the amount of iron atom. 1 / 1
| 1

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"The results showed that OTUD5 overexpression restored the increase in iron and MDA levels and the decrease in GSH caused by MI/R."

reach
"OTUD5 can also reduce the IL-17 production by T cells [6]."
OTUD5 affects innate antiviral antitumor immunity
| 1
OTUD5 activates innate antiviral antitumor immunity. 1 / 1
| 1

eidos
"OTUD5 promotes innate antiviral and antitumor immunity through deubiquitinating and stabilizing STING ."
OTUD5 affects immune responses
| 1
OTUD5 activates immune responses. 1 / 1
| 1

eidos
"OTUD5 also modulates immune responses by deubiquitinating and stabilizing UBR5 to suppress the production of IL-17 and by selectively cleaving the polyubiquitin chains on TRAF3 to regulate IFN-I production17 ,18 ."
| 1

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"Thus, our work propose that FACT is an important factor in this in cis regulation, and that FACT is subject to regulation by OTUD5–UBR5 complex for the non-lesion stalling of Pol II.Our work leaves several open-ended questions; our data collectively suggest that OTUD5 and UBR5 repress FACT histone exchange activity, but how does this complex exactly achieve this?"

reach
"These results suggest that FACT and HDAC proteins are associated by tethering to the C-terminus of OTUD5.One plausible hypothesis as to how OTUD5 regulates FACT activity is through recruiting HDAC1/2 and inducing histone deacetylation."
| 1

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"In breast cancer and colon cancer, OTUD5 promotes carcinogenesis by regulating the mammalian target of rapamycin (mTOR) and Hippo signaling pathways [10,11,12]."
OTUD5 affects glutathione
| 1
| 1

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"The results showed that OTUD5 overexpression restored the increase in iron and MDA levels and the decrease in GSH caused by MI/R."
| 1

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"We hypothesized that changes in gene expression caused by OTUD5 accumulation under TRIP12 knockdown, may be canceled by simultaneous knockdown of OTUD5."
OTUD5 affects erastin
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OTUD5 inhibits erastin. 1 / 1
| 1

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"The results showed that OTUD5 overexpression inhibited RSL3 and erastin‐induced lipid peroxidation (Figure S2C), whereas OTUD5 knockdown enhanced this effect (Figure S2D)."
OTUD5 affects enrichment TRIM25 PML
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OTUD5 inhibits enrichment TRIM25 PML. 1 / 1
| 1

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"Indeed , OTUD5 depletion increased the enrichment of TRIM25 on the promoter of PML ( Fig. 6a ) ."
OTUD5 affects dopamine
| 1
| 1

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"In addition, genetic deletion of OTUD5 in DA neurons significantly aggravated the α‐Syn PFF‐induced neurodegeneration in vivo, while OTUD5 overexpression rescued the DA neuron damage induced by α‐Syn PFF in vitro.Recent studies emphasized the importance of DUBs in regulating the fate of α‐Syn."
| 1

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"Overexpression of OTUD5 in gastric cancer cells partly reversed the anti-tumor effect of dapagliflozin."
OTUD5 affects chemoradiotherapy resistance
| 1
OTUD5 inhibits chemoradiotherapy resistance. 1 / 1
| 1

eidos
"Since the FACT histone chaperone complex can promote the resistance of tumor cells to DNA damage factors ( Dejmek , Iglehart , & Lazaro , 2009 ; Koman et al ., 2012 ; Yarnell , Oh , Reinberg , & Lippard , 2001 ) , a follow-up research is necessary to determine whether OTUD5 can reduce chemoradiotherapy resistance by antagonizing the FACT histone chaperone complex ."
| PMC
OTUD5 affects autophagy
| 1
| 1

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"The knockdown of OTUD5 has caused several mTOR-related phenotypes, such as a decrease in cell size and an increase in autophagy in mammalian cells."
OTUD5 affects accessible chromatin neuroectodermal aberrant
| 1
OTUD5 activates accessible chromatin neuroectodermal aberrant. 1 / 1
| 1

eidos
"Loss of OTUD5 during differentiation leads to less accessible chromatin at neuroectodermal enhancers and aberrant gene expression ."
OTUD5 affects ZUP1
| 1
OTUD5 activates ZUP1. 1 / 1
| 1

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"Thus, TRIP12-generated K29-linkage besides protecting itself from OTUD5-mediated deubiquitylation, also 1) facilitates UBR5-dependent assembly of K48-linked chains and 2) modestly protects K48-linkages from disassembly as forms of mixed/branched linkages."
OTUD5 affects WT
| 1
OTUD5 leads to the ubiquitination of WT. 1 / 1
| 1

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"The Co-IP analysis showed that OTUD5 could abrogate polyubiquitination of MyD88 (WT), while it does not possess the ability to eliminate the ubiquitination of MyD88 (K95R, K231R, K250R) mutant forms (Fig. 3H)."
OTUD5 affects VSV-G pseudotype
| 1
OTUD5 activates VSV-G pseudotype. 1 / 1
| 1

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"In contrast, knockout of SBDS affected the VSV-G pseudotyped virus, but not HIV-1 envelope-mediated entry while OTUD5 knockout had an effect on both viruses but decreased the VSV-G pseudotype to a greater extent."
OTUD5 affects UBR5-ATMIN
| 1
OTUD5 activates UBR5-ATMIN. 1 / 1
| 1

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"OTUD5 promotes ATM-dependent DDR activation by regulating the UBR5-ATMIN axis."

reach
"OTUD5 acts as a tumor inhibitor in non-small cell lung cancer and cervical cancer.36 37 However, OTUD5 promotes carcinogenesis in triple-negative breast cancer and bladder cancer.38 39 Hence, the specific function of OTUD5 in different cancers needs to be analyzed on a case-to-case basis."
OTUD5 affects Th17
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OTUD5 inhibits Th17. 1 / 1
| 1

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"DUBA inhibits Th17 differentiation by stabilizing UBR5, which mediates ubiquitination and degradation of RORγt."
OTUD5 affects Th1 Cells
| 1
| 1

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"In the presence of CD-associated NOD2 mutations, impaired recognition of MDP results in inefficient induction of DUBA, which leads to excessive and colitogenic type I IFN and Th1 responses upon TLR9 activation ( Figure 4B )."
OTUD5 affects TRAF3-TBK1 interaction
| 1
OTUD5 activates TRAF3-TBK1 interaction. 1 / 1
| 1

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"Indeed, DUBA and OTUD5, a DUB which selectively disassembles Lys63 linked ubiquitin chains on TRAF3, has been shown to suppress the production of type I interferon by reducing the level of TRAF3-TBK1 interaction XREF_BIBR."
OTUD5 affects TRAF-C
| 1
OTUD5 binds TRAF-C and RING domain. 1 / 1
| 1

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"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
OTUD5 affects TRAF-C region
| 1
OTUD5 binds TRAF-C region and RING domain. 1 / 1
| 1

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"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects TLR9
| 1
OTUD5 inhibits TLR9. 1 / 1
| 1

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"Mice lacking IL-1RI signaling fail to produce a protective type I IFN response after administration of TLR9 ligand (CpG), suggesting that regulation of DUBA expression by IL-1 signaling attenuates TLR9-mediated proinflammatory responses.In virus-induced sepsis, the virus triggers a strong inflammatory response by activating the host’s immune system, where overactivation of the type I interferon pathway and the NF-κB signaling pathway often leads to SIRS and cytokine storm (61, 62)."
OTUD5 affects TH
| 1
OTUD5 increases the amount of TH. 1 / 1
| 1

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"Besides, overexpressing OTUD5 restored the protein levels of TH in α‐Syn PFF‐induced PD cell model."
OTUD5 affects TERF1
| 1
OTUD5 inhibits TERF1. 1 / 1
| 1

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"Moreover, ectopic expression of phosphomimic KAP1 also rescued the telomere MMEJ defect in OTUD5-depleted Trf1 ;Trf2 ;Ku70 ;p53 ;Cre-ER MEFs (Fig. 5g, Supplementary Fig. 7d, e)."
OTUD5 affects TAK1-TAB2
| 1
TAK1-TAB2 binds OTUD5. 1 / 1
| 1

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"These findings elucidate that OTUD5 inhibits TAK1 activation by blocking TAK1-TAB2 interaction via OTUD5-mediated TAK1 deubiquitination at the K158 site."
OTUD5 affects TAK1-MAPK
| 1
OTUD5 inhibits TAK1-MAPK. 1 / 1
| 1

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"In addition, OTUD5 inhibited the activation of the TAK1-MAPK pathway in HG/PA-treated podocytes but failed when its deubiquitinating activity was canceled in the C224S mutant (Fig. 5j)."
OTUD5 affects Sepsis
| 1
OTUD5 activates Sepsis. 1 / 1
| 1

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"Consistently, Otud5-deficient mice are less susceptible to LPS- and CLP-induced sepsis."
OTUD5 affects S phase
| 1
OTUD5 activates S phase. 1 / 1
| 1

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"Collectively, these results imply that OTUD5 orchestrates ferroptosis in gastric cancer cells.Moreover, we evaluated the impact of OTUD5 knockout on the cell cycle of gastric cancer cells by an EdU incorporation assay, and the results revealed that OTUD5 depletion decreased the proportion of cells in S phase, indicating impaired cell proliferation (Figure 3M,N)."
OTUD5 affects RORgammat
| 1
OTUD5 decreases the amount of RORgammat. 1 / 1
| 1

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"Therefore, DUBA or UBR5 knockout can enhance the level of RORgammat and promote T H 17 cell differentiation."
OTUD5 affects RORgammat protein
| 1
OTUD5 inhibits RORgammat protein. 1 / 1
| 1

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"For example, deubiquitinase DUBA can suppress RORgammat protein stability in TGF-beta stimulated Th17 cells [XREF_BIBR]."
| PMC
OTUD5 affects RNApo_II
| 1
| 1

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"Using a PLA-based readout for TRC (4), we found that OTUD5 or UBR5 depletion causes increased proximity of PCNA and an elongating form of RNA polymerase II (RNA Pol II (S2-P)) (Figure 2A)."
OTUD5 affects RNAPII
| 1
OTUD5 inhibits RNAPII. 1 / 1
| 1

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"Furthermore, loss of the deubiquitylase OTUD5, which inhibits RNAPII elongation in response to DNA double-strand breaks , alleviates the sensitivity of ELOF1-KO cells to Illudin S, while its loss has the opposite impact on CSB-KO cells (Fig. 7a, b)."

reach
"OTUD5 localizes to DNA double strand breaks (DSBs), interacts with UBR5 and represses the RNA Pol II elongation and RNA synthesis."
OTUD5 affects RNA Pol II
| 1
OTUD5 inhibits RNA Pol II. 1 / 1
| 1

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"Using a PLA-based readout for TRC (4), we found that OTUD5 or UBR5 depletion causes increased proximity of PCNA and an elongating form of RNA polymerase II (RNA Pol II (S2-P)) (Figure 2A)."
OTUD5 affects RIPK2
| 1
OTUD5 increases the amount of RIPK2. 1 / 1
| 1

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"DUBA potentiated NOD2-mediated signal transduction by maintaining the protein levels of NOD2 and receptor-interacting protein kinase 2 (RIPK2)."
OTUD5 affects RIPK1
| 1
OTUD5 deubiquitinates RIPK1. 1 / 1
| 1

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"For example, TNFAIP3, CYLD, USP3, USP4, USP15, USP25, and OTUD5 deubiquitinate key components of the IFN production pathway, including RIG-I, TRAF2, TRAF3, TRAF6, RIP1, and TRIF (reviewed in [93,94])."
OTUD5 affects RING domain, and TRAF-C
| 1
OTUD5 binds TRAF-C and RING domain. 1 / 1
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"If DUBA binds to both TRAF-C and the catalytic RING domain of TRAF3, it impedes TRAF3 signaling not only by removing the Ub chains, but also by preventing its interactions.4.5 TRAF3-OTUB1 interaction."
OTUD5 affects RING domain, and TRAF-C region
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OTUD5 binds TRAF-C region and RING domain. 1 / 1
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"If DUBA also binds to TRAF-C region in addition to the RING domain of TRAF3, it attenuates TLR-induced IRF activation not only through its catalytic (DUB) function, but also through interfering with t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
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"Overall, these results indicated that OTUD5 recruited NEDD4 to promote K63‐linked polyubiquitination of α‐Syn and subsequent degradation.2.5 OTUD5 Was Downregulated in alpha-Syn PFF-induced PD Model."
OTUD5 affects PML-NBs
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OTUD5 activates PML-NBs. 1 / 1
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"f Depletion of OTUD5 reduced the formation of PML-NBs in the Huh7 and Hep3B cells ."
OTUD5 affects NMRAL1
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"HSCARG neither interacted with DUBA nor promoted its recruitment (XREF_SUPPLEMENTARY, B), and depletion of DUBA had no effect on HSCARG in inhibiting TRAF3 ubiquitination (XREF_SUPPLEMENTARY)."
OTUD5 affects NFkappaB
| 1
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"OTUD5 promotes TLR/IL1-R-mediated NF-kappaB and MAPK signaling."

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"Treatment with MG132 efficiently increased PDCD5 levels; whereas, knockdown of OTUD5 greatly diminished the effect of MG132 on the stabilization of PDCD5, indicating that OTUD5 played a critical role [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects MFSD11
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OTUD5 activates MFSD11. 1 / 1
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"Overexpression of OTUD5 increased the levels of PDCD5 and p53 in a time-dependent manner; whereas knockdown of OTUD5 substantially impaired the positive effect of ET on PDCD5 and p53 activation ( Fig.[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects MDM2
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OTUD5 deubiquitinates MDM2. 1 / 1
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"As shown in XREF_FIG, MDM2 can not be deubiquitinated by OTUD5."
OTUD5 affects MASH
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OTUD5 inhibits MASH. 1 / 1
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"Conditional Knockout of OTUD5 in Hepatocytes Exacerbated HFF-induced MASH."
OTUD5 affects M3
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OTUD5 inhibits M3. 1 / 1
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"Metabolic tracking revealed that OTUD5 knockdown induced a significantly decreased abundance of tricarboxylic acid (TCA) cycle metabolites (such as M2 citrate, M2 ketoglutarate, M2 succinate, and M2 fumarate), with a compensatory increase in the abundance of glycolytic metabolites (such as M6 G-6-P, M6 F-6-P, M6 F-1, 6-P, M3 pyruvate, and M3 lactate)."
OTUD5 affects LAMP1
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OTUD5 activates LAMP1. 1 / 1
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"Furthermore, confocal microscopic analysis visualized that the co‐localization of α‐Syn and LAMP1, a marker of lysosomes, was significantly enhanced by OTUD5 overexpression (Figure 2B)."
OTUD5 affects K63-linked polyubiquitination AKT
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OTUD5 inhibits K63-linked polyubiquitination AKT. 1 / 1
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eidos
"DUB CYLD , OTUD5 and USP1 can reverse K63-linked polyubiquitination of AKT and inhibit its activation [ 85-87 ] ."
| PMC
OTUD5 affects K48
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OTUD5 binds polyubiquitin chains and K48. 1 / 1
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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
OTUD5 affects JNK
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OTUD5 leads to the dephosphorylation of JNK. 1 / 1
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"The results showed OTUD5 knockdown significantly enhanced the phosphorylation of ERK1/2, p38 and JNK (Fig. 7A), while after overexpression of OTUD5, the inverse results were only observed in ERK1/2 and p38 pathways (Fig. 7B)."

sparser
"Since DUBA is known to negatively regulate type I IFN and IFN is associated with IBD, it is important to understand association of DUBA and IBD as well."
OTUD5 affects IL6
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OTUD5 inhibits IL6. 1 / 1
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"DUBA knockdown by siRNA transfection in Il1r1 -/- BMDCs (XREF_FIG) resulted in significantly increased IL-10 and IFN-beta production and decreased secretion of IL-6, but not TNF, in CpG stimulated Il1r1 -/- BMDCs (XREF_FIG)."
OTUD5 affects IKBKE
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OTUD5 deubiquitinates IKBKE. 1 / 1
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"TRIAD3A dependent TRAF3 proteasomal destruction, like TRAF3 de-ubiquitination by DUBA, halts the activation of TBK1 and IKKepsilon and subsequent IFN production."
OTUD5 affects IFNA
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OTUD5 activates IFNA. 1 / 1
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"The removal of endogenous DUBA by siRNA produced an elevated IFN-β and IFN-α response to poly:IC or Sendai virus, whereas NF-κB activation was predominantly unaffected ( Kayagaki et al., 2007 )."
OTUD5 affects IFIH1
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OTUD5 inhibits IFIH1. 1 / 1
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"The deubiquitinating enzyme A (DUBA), a novel OTU-domain DUB has been shown to negatively regulate IFN signaling following RIG-I, MDA-5 or TLR3 stimulation [60] ."
OTUD5 affects Hypertrophy
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| 1

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"Meanwhile, OTUD5 overexpression reduced cardiac fibrotic areas (Figure 7K) and cardiac hypertrophy (Figure 7L)."
OTUD5 affects HES
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OTUD5 inhibits HES. 1 / 1
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"To corroborate these results and exclude any off-target effects originating from clonal selection during iPSC reprogramming, we next generated control or OTUD5 depleted hES H1 cells and subjected them to neural conversion."
| PMC
OTUD5 affects HDAC1/2
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OTUD5 binds HDAC1/2. 1 / 1
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"Altogether, these results suggest that the OTUD5-HDAC1/2 complex restricts aberrant FACT loading onto chromatin."
OTUD5 affects HDAC
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OTUD5 activates HDAC. 1 / 1
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"This result suggests that OTUD5 promotes stability of the HDAC proteins."
OTUD5 affects HCFC1
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| 1

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"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC
OTUD5 affects HBc
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OTUD5 activates HBc. 1 / 1
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"These results showed that OTUD5 strongly inhibited HBc degradation in vitro (Fig. S4G, H)."
OTUD5 affects GOT
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OTUD5 decreases the amount of GOT. 1 / 1
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"Moreover, Otud5 re-expression also reduced serum AST and ALT levels after 16 weeks of HFF feeding compared with AAV-GFP injection (Figure 15Q)."
OTUD5 affects Dronc
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OTUD5 activates Dronc. 1 / 1
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"However, this non-apoptotic role can be compromised by reducing the expression of the de-ubiquitylating enzyme DUBA, which physically interacts with and modulates the activity of Dronc in the sperm [ [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTUD5 affects DPEP1
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OTUD5 activates DPEP1. 1 / 1
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"In addition, DUBA deficiency in macrophages rendered mice hypersensitive to DSS-induced colitis and eliminated the protective effect of MDP treatment in colitis."

reach
"Moreover, we showed that OTUD5 mediated the sequential activation of PDCD5 and p53 in response to genotoxic stress."
OTUD5 affects DNA break DSB repair
| 1
OTUD5 activates DNA break DSB repair. 1 / 1
| 1

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"Our previous study indicated that OTUD5 promoted DNA double-strand break ( DSB ) repair by inhibiting Ku80 degradation14 ."
| 1

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"This novel payload, DUBA, induces DNA alkylation and damage in both dividing and non-dividing cells."
| 1
| 1

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"Intriguingly, introduction of WT OTUD5, as opposed to catalytically inactive C224S OTUD5, diminished H/R-induced cell ferroptosis (Fig. 4b), indicating a protective role of OTUD5 against renal tubular cell ferroptosis upon H/R."
OTUD5 affects Caspase
| 1
| 1

sparser
"DUBA physically interacts with the initiator caspase death regulator Nedd2-like caspase (Dronc) and de-ubiquitylates it, thereby contributing to efficient inhibitor of apoptosis-antagonist-induced apoptosis in the fly eye."
OTUD5 affects CD8
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OTUD5 inhibits CD8. 1 / 1
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"However, whether blockade of USP10 or OTUD5 can reactivate CD8 T-cell-mediated cytotoxicity against tumors in a MDSC-dependent manner remains unclear and requires further exploration."
OTUD5 affects CASP2
| 1
| 1

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"DUBA physically interacts with the initiator caspase death regulator Nedd2 like caspase (Dronc) and de-ubiquitylates it, thereby contributing to efficient inhibitor of apoptosis-antagonist-induced apoptosis in the fly eye."
OTUD5 affects ATMIN
| 1
| 1

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"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
OTUD5 affects ARMC7
| 1
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"Indeed, we found that ARMC7 and ZGPAT, whose protein levels are co-regulated by TRIP12 and UBR5 (Fig. 6b), stably associated with OTUD5 in cells (Fig. 6c)."
OTUD5 affects ACX4
| 1
OTUD5 inhibits ACX4. 1 / 1
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"Metabolic tracking revealed that OTUD5 knockdown induced a significantly decreased abundance of tricarboxylic acid (TCA) cycle metabolites (such as M2 citrate, M2 ketoglutarate, M2 succinate, and M2 fumarate), with a compensatory increase in the abundance of glycolytic metabolites (such as M6 G-6-P, M6 F-6-P, M6 F-1, 6-P, M3 pyruvate, and M3 lactate)."
OTUD5 affects 3 L
| 1
OTUD5 binds YAP1 and 3 L. 1 / 1
| 1

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"Indeed, TNBC cells enhanced the binding between OTUD5 and YAP ( Fig. 3 L), and the ubiquitination assays showed the ubiquitin chain of YAP in co-culture group was dramatically removed ( Fig. 3 M)."
Neoplasms affects OTUD5
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Neoplasms decreases the amount of OTUD5. 1 / 1
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"OTUD5 mediates GPX4 deubiquitination to regulate its stability.Deletion of OTUD5 promotes ferroptosis and inhibits tumor growth.Wild type p53 inhibits OTUD5 transcription, thereby promoting GPX4 degradation and inhibiting the development of gastric cancer.OTUD5, GPX4 expression and p53 activity are highly correlated and correlates with clinical progression in STAD."
NMRAL1 affects OTUD5
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| 1

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"HSCARG neither interacted with DUBA nor promoted its recruitment (XREF_SUPPLEMENTARY, B), and depletion of DUBA had no effect on HSCARG in inhibiting TRAF3 ubiquitination (XREF_SUPPLEMENTARY)."
Mice affects OTUD5
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Mice inhibits OTUD5. 1 / 1
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"To directly evaluate the physiological role of OTUD5, myeloid-specific OTUD5 knockout mice were produced by breeding Otud5 mice with Lyz2-Cre mice."
MIR937 affects OTUD5
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MIR937 inhibits OTUD5. 1 / 1
| 1

eidos
"Moreover , hsa-mir-137 , hsa-mir-1913 , hsa-mir-937 , hsa-mir-607 , hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5 ."
MIR607 affects OTUD5
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MIR607 inhibits OTUD5. 1 / 1
| 1

eidos
"Moreover , hsa-mir-137 , hsa-mir-1913 , hsa-mir-937 , hsa-mir-607 , hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5 ."
MIR1913 affects OTUD5
| 1
| 1

eidos
"Moreover , hsa-mir-137 , hsa-mir-1913 , hsa-mir-937 , hsa-mir-607 , hsa-mir-3149 and hsa-mir-144 may inhibit the expression of OTUD5 ."
MDP-NOD2 axis affects OTUD5
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MDP-NOD2 axis increases the amount of OTUD5. 1 / 1
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"The MDP-NOD2 axis downregulates TLR9-induced ISG by upregulating DUBA expression (67, 68)."
K63 affects polyubiquitin chains
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OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
K63 affects TRAF3
| 1
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

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"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
K63 affects OTUD5, and polyubiquitin chains
| 1
OTUD5 binds polyubiquitin chains and K63. 1 / 1
| 1

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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
K63 affects OTUD5, and TRAF3
| 1
OTUD5 binds TRAF3 and K63. 1 / 1
| 1

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"DUBA interacts with TRAF3 and removes K63 polyubiquitin chains from TRAF3 in a manner that depends on its ubiquitin-interaction motif (UIM)."
K48 affects polyubiquitin chains
| 1
OTUD5 binds polyubiquitin chains and K48. 1 / 1
| 1

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"Although the DUBA UIM domain binds both K48 and K63 polyubiquitin chains, DUBA specifically cleaves K63 polyubiquitin chains from TRAF3."
JNK affects OTUD5
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JNK inhibits OTUD5. 1 / 1
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"However, the inhibition of ERK1/2 but not JNK and p38 MAPK pathways enhanced OTUD5-mediated promotion of HNF4α (Fig. 7D)."
Interferon affects OTUD5
| 1
| 1

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"As TRAF3 has been demonstrated to be a significant gene for the production of type I interferon in RLRs signaling pathway [ 39 , 40 ], the overexpression of miR-210 could indirectly activate the RLRs [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Since DUBA is known to negatively regulate type I IFN and IFN is associated with IBD, it is important to understand association of DUBA and IBD as well."
IL1R1 affects OTUD5
| 1
IL1R1 inhibits OTUD5. 1 / 1
| 1

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"DUBA knockdown by siRNA transfection in Il1r1 -/- BMDCs (XREF_FIG) resulted in significantly increased IL-10 and IFN-beta production and decreased secretion of IL-6, but not TNF, in CpG stimulated Il1r1 -/- BMDCs (XREF_FIG)."
HDAC1/2 affects OTUD5
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OTUD5 binds HDAC1/2. 1 / 1
| 1

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"Altogether, these results suggest that the OTUD5-HDAC1/2 complex restricts aberrant FACT loading onto chromatin."
HCFC1 affects OTUD5
| 1
| 1

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"We could confirm the interactions between OTUD5 and ARID1A/B, HDAC2, HCF1, and UBR5 by immunoblotting at the endogenous level in hES H1 cells (XREF_FIG)."
| PMC
ERK affects OTUD5
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ERK activates OTUD5. 1 / 1
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"To further investigate the role of ERK1/2 and p38 signaling in OTUD5-mediated promotion of HNF4ɑ or HNF1ɑ, we treated the OTUD5-sh HepG2.2.15 cells with ERK1/2-specific pharmacological inhibitor (PD98059), JNK-specific pharmacological inhibitor (SP600125) and p38-specific pharmacological inhibitor (SB203580), and then tested the effect of OTUD5 on the protein level of HNF1α and HNF4α."
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E3_Ub_ligase ubiquitinates OTUD5. 1 / 1
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"In the case of OTUD5, restoring deubiquitylation activity, perhaps by destruction of the cognate E3 ubiquitin ligase(s) which ubiquitylate OTUD5 substrate proteins involved in chromatin organisation, could be employed to re‐establish neurodevelopmental gene expression."
| 1
Deubiquitinase decreases the amount of OTUD5. 1 / 1
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"In addition to inhibiting USP20, a recent study confirmed that 160 nM GSK2643943A also has an inhibitory effect on another DUB, ovarian tumor protease deubiquitinase 5 (OTUD5), which can increase the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Caspase affects OTUD5
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| 1

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"DUBA physically interacts with the initiator caspase death regulator Nedd2-like caspase (Dronc) and de-ubiquitylates it, thereby contributing to efficient inhibitor of apoptosis-antagonist-induced apoptosis in the fly eye."
CD4+ affects OTUD5
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CD4+ inhibits OTUD5. 1 / 1
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"Exposure of Bach2 silenced CD4+ T-lymphocytes to CP tissue extract showed increased Ahr, decreased OTUD5, and enhanced Th17 cell differentiation."
CASP2 affects OTUD5
| 1
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"DUBA physically interacts with the initiator caspase death regulator Nedd2 like caspase (Dronc) and de-ubiquitylates it, thereby contributing to efficient inhibitor of apoptosis-antagonist-induced apoptosis in the fly eye."
ATMIN affects OTUD5
| 1
| 1

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"To test our hypothesis, we first assessed whether there is any detectable interaction between OTUD5, UBR5, ATMIN and ATM."
ARMC7 affects OTUD5
| 1
| 1

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"Indeed, we found that ARMC7 and ZGPAT, whose protein levels are co-regulated by TRIP12 and UBR5 (Fig. 6b), stably associated with OTUD5 in cells (Fig. 6c)."

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"The results showed that nutlin‐3a significantly reduced the mRNA level of OTUD5, while it had no effect on the transcription levels of the E3 ubiquitin ligases (Figure S3C)."
3 L affects YAP1
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OTUD5 binds YAP1 and 3 L. 1 / 1
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"Indeed, TNBC cells enhanced the binding between OTUD5 and YAP ( Fig. 3 L), and the ubiquitination assays showed the ubiquitin chain of YAP in co-culture group was dramatically removed ( Fig. 3 M)."