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"Moreover, hypoxia/reoxygenation (H/R) induces FOXO4 upregulation in rat H9C2 cardiomyocytes, and FOXO4 overexpression can reverse the protective effects of ubiquitin-specific peptidase 10 (USP10) overexpression on H/R-induced H9C2 cells by regulating the Hippo/yes-associated protein 1 signaling pathway (10)."