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USP4 activates BMP. 3 / 3
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eidos
"( B ) BMP signaling : USP4 can deubiquitinate TI3RI and Smad4 , and enhance BMP signaling ."

eidos
"Moreover , USP4 stabilizes Smad4 and activates BMP signaling by inhibiting Smad4 monoubiquitination ( Zhou et al ., 2017 ) ."

reach
"Furthermore, USP4 stabilizes Smad4 through inhibiting its monoubiquitination and enhances activin as well as BMP signaling [86]."