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PKA phosphorylates USP20 on S333. 2 / 3
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sparser
"Subsequently, they found that protein kinase A phosphorylates USP20 on serine 333, inhibits the trafficking of the substrate β2AR (β2 adrenergic receptors) and decreases degradation via autophagosomes [ xref ]."

reach
"Subsequently, they found that protein kinase A phosphorylates USP20 on serine 333, inhibits the trafficking of the substrate β2AR (β2 adrenergic receptors) and decreases degradation via autophagosomes [41]."