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USP4 activates TUT1. 2 / 2
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"As a result of SART3 co-expression with USP4, the efficiency of TUT1 translocation was increased ( Fig. 4 E and F), demonstrating that USP4 induces TUT1 subnuclear redistribution in a similar manner o[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In addition, co-expression of USP4 augmented TUT1-dependent U6-snRNA upregulation ( Fig. 4 G)."