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"Abrogation of USP28 induces DNA damage and cell death."

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"Remarkably, the ablation of p53, 53bp1, or Usp28 in addition to Sas‐4 or Sass6 rescues the cell death and allows the double mutant embryos to develop 1 day further, when they resemble cilia mutants (Fig. 4) [51, 70, 100]."