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RPS6KB1 phosphorylates RPS6. 546 / 633
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sparser
"Rapamycin strongly inhibits S6K1 phosphorylation of S6 but also has a minor effect on Akt phosphorylation."

sparser
"S6K1 directly phosphorylates the 40S ribosomal protein S6, and then promotes ribosome biogenesis."

reach
"Similarly, an inhibitor of p70 S6 kinase (PF-4708671) partially inhibited phosphorylation of S6 and preserved RP, while only marginally prevented cyclin D1 induction."

reach
"Phosphorylated 4EBP1 allows active eIF4E.eIF4G complex formation and activated S6K1 phosphorylates ribosomal protein S6."

sparser
"This positive effect on nucleotide synthesis partners with the strong mTORC1 effects on protein translation through phosphorylation of 4EBP and ribosomal protein S6 by S6K1 and S6K2, and promotes overall ribosomal biogenesis."

reach
"S6 is phosphorylated by ribosomal protein p70 S6 kinase (S6K), which is activated by mTORC1."

reach
"mTORC1 activates p70S6K, which in turn phosphorylates the ribosomal protein S6 and 4E-BP1, both involved in protein translation."

sparser
"For this procedure to be effective, S6 must be phosphorylated by S6K1 ( xref )."

reach
"In the present study, we have examined whether Ang II activates p70 S6 kinase (p70S6K), which has been reported to phosphorylate S6 protein."

sparser
"Upon activation by receptor tyrosine kinases or G-protein coupled receptors, AKT activates mTOR, which in turn stimulates protein translation through inhibitory phosphorylation of eukaryotic translational initiation factor eIF4E-binding protein 1 (4E-BP1) that inhibits eIF4E, and activating phosphorylation of p70S6 kinase (S6K) that phosphorylates ribosomal S6 protein."

sparser
"Another important locus for regulation of mRNA binding to 40S ribosomal subunits occurs through phosphorylation of ribosomal protein S6 by S6K1."

sparser
"mTOR activates S6K1, which phosphorylates eIF4B and S6, so promoting translation initiation, and this may be enhanced by ASF/SF2 xref ."

sparser
"RAME, not RA, inhibits the phosphorylation of S6 by S6K1."

sparser
"Adiponectin enhances insulin signaling by reducing p70 S6 kinase-mediated serine phosphorylation of IRS-1 [ xref ] but systemic adiponectin levels are not influenced by carbohydrate uptake."

sparser
"In turn, p70S6K1 phosphorylates a S6 protein of the 40S ribosomal subunit (S6rp) and 4E-BP1, with the latter negatively regulating eukaryotic initiation factor 4E (eIF-4E)."

sparser
"After activation, S6K1 phosphorylates the ribosomal subunit protein S6, leading to an increase in protein translation."
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reach
"mTORC1 activated p70S6K phosphorylates the 40S-ribosomal protein S6."

sparser
"Activated p70S6K phosphorylates the ribosomal protein S6, leading to initiation of protein synthesis ( xref )."

sparser
"For example, S6Ks phosphorylate ribosomal protein S6, which promotes translation initiation and elongation."

sparser
"Downstream of mTORC1, both S6K1 and S6K2 can phosphorylate S6, xref , xref although S6K1 has been shown to regulate muscle growth and insulin sensitivity independent of S6K2 activity. xref , xref In the present study, all genetic mouse groups showed increased phosphorylation and thus activity of S6K2 ( Figure   xref )."

reach
"The inhibitor of RPS6KB1, LY2584702, significantly reduced the phosphorylation of RPS6KB1 and rpS6 in NSCLC cell lines."

sparser
"P70s6k in turn phosphorylates rpS6 which is a ribosomal protein specific to anabolic protein synthesis in myotubes."

reach
"RPS6 is phosphorylated by p70S6K, which in turn is activated by mTOR (Hay and Sonenberg, 2004)."

sparser
"mTOR complex 1 (mTORC1) promotes phosphorylation (activation) of p70 S6 kinases (p70S6Ks) that phosphorylate the small ribosomal subunit protein S6 (Rps6), and this was proposed to promote translation of TOP mRNAs."

reach
"S6K1, a key regulator of cell growth, phosphorylates ribosomal protein S6 and enhances the translation of 5 '-terminal oligopyrimidine tract mRNAs in some models."

reach
"While p70S6K phosphorylates ribosomal S6 protein potentially modulating its activity, 4EBP1 binds and inactivates a factor of initiation of Cap dependent transcription eIF4E."

reach
"Upon activation, S6K1 phosphorylates ribosomal protein S6 (RPS6) to stimulate ribosomal biogenesis [XREF_BIBR], eukaryotic elongation factor 2 kinase (eEF2K) to de-repress translational elongation [XREF_BIBR], as well as eIF4B to heighten translational initiation [XREF_BIBR, XREF_BIBR]."

sparser
"Regulation of protein synthesis via mTOR also occurs through its phosphorylation of p70S6K, which then phosphorylates the S6 40S ribosomal subunit."

reach
"S6K1 phosphorylates the S6 ribosome, thus stimulating protein synthesis."

reach
"S6, a component of the 40S ribosomal subunit, is phosphorylated by S6K1, which under growth promoting conditions causes increased rates of protein synthesis."

sparser
"mTOR is activated by AKT and it stimulates protein translation by phosphorylating eIF4E binding protein (4E-BP1) as well as p70S6K which phosphorylates S6."

sparser
"Phosphorylation of ribosomal protein S6 by p70 S6 kinase, which is believed to be involved in regulating 5′-TOP mRNA translation [ xref ], is one of the earliest events detected following mitogenic stimuli, as part of liver regeneration of following fasting and refeeding."

sparser
"Additionally, p70S6K can further phosphorylate the ribosomal protein S6 (RPS6) (S236, +81% CFC), and reduction of the levels of p70S6K has been shown to improve the AD-like phenotype of 3xTg-AD mice [ xref ]."

reach
"It is a serine/threonine kinase, which phosphorylates at least three downstream substrates : the translation repressor eIF4E binding protein (4E-BP) (von Manteuffel et al, 1997; Gingras et al, 1999; Nojima et al, 2003); p70S6K, which phosphorylates ribosome protein S6 (Nojima et al, 2003); and signal transducer and activator of transcription 3 (Stat3) (Yokogami et al, 2000; Kusaba et al, 2004)."

reach
"The activated S6K1 phosphorylates ribosomal protein S6 (RpS6), and promotes protein translation."

reach
"While p70S6K can phosphorylate RPS6 at all sites, S235/236 is a specific target for RSK [XREF_BIBR]."

sparser
"In response to various stimuli, mTOR can be activated through phosphorylation, and the activated p-mTOR is involved in the regulation of protein synthesis by inducing the phosphorylation of p70S6K, which can phosphorylate the ribosomal protein S6 that is involved in protein translation ( xref , xref )."

reach
"The mTORC1 controls the rate of protein synthesis through phosphorylation and activation of its substrates, p70S6 ribosomal kinase 1 (p70S6K) and eukaryotic translation initiation factor 4E (eIF4E) binding protein-1 (4E-BP1) and once phosphorylated, p70S6K phosphorylates ribosomal protein S6 and 4E-BP1 becomes dissociated from eIF4 and promote mRNA translation and protein synthesis [XREF_BIBR]."

sparser
"Activation of mTOR leads to phosphorylation of several proteins, including p70 ribosomal S6 (phospho-S6) kinase which phosphorylates ribosomal protein S6 thereby resulting in increased protein translation and cell growth."

sparser
"Activated mTOR phosphorylates p70 ribosomal protein S6 kinase (p70S6K) followed by p70S6K-induced phosphorylation of ribosomal protein S6 and eukaryotic initiation factor 4B (eIF-4B) which promotes the initiation of protein translation."

sparser
"2 Phosphorylation of S6 by p70S6K is involved in translational control of 50 oligopyrimidine tract mRNAs and ultimately causes upregulation of cellular protein synthesis."

reach
"Phosphorylation of S6 by S6K1 mediates unwinding and initiates translation of 5 terminal oligopyrimidine tract (TOP) mRNAs."

sparser
"When activated, p70 S6 kinase phosphorylates the S6 protein of the 40S ribosome, and this has been suggested to allow preferential translation of a family of mRNAs with 5′ polypyrimidine tracts [86, [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Phosphorylated S6K1 in turn phosphorylates S6 (40S ribosomal protein S6), enhancing the translation of mRNAs, while phosphorylation of 4EBP1 suppresses its ability to act on downstream effectors."

sparser
"Active mTORC1 phosphorylates and activates p70S6K which then phosphorylates ribosomal protein S6 and inhibits the binding of 4E-BP1 to eIF4E ( xref ; xref )."

sparser
"Although S6K1 was first identified as a kinase of ribosomal protein S6 (rpS6), the functional purpose of rpS6 phosphorylation by S6K1 is still unresolved."

sparser
"Using the SA model and in vitro models of muscle hypertrophy, it was discovered that the activation of mTORC1 signaling and subsequent ribosomal p70 S6 kinase phosphorylation is a driving factor in ribosome biogenesis ( xref , xref )."

sparser
"FRAP may activate p70 S6 kinase which phosphorylates the S6 subunit of the 40S ribosomal subunit, an event that correlates with growth factor-regulated increases of translation of defined, growth regu[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"It is known that mTOR phosphorylates p70S6K followed by p70S6K-induced phosphorylation of eIF4B and S6 ( xref ) ( xref )."

reach
"In response to various anabolic stimuli, mTOR phosphorylates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (rpS6) to positively affect muscle growth ."

reach
"In response to various anabolic stimuli, mTOR phosphorylates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (rpS6) to positively affect muscle growth ."

reach
"AA signaling is integrated by the mammalian target of rapamycin (mTOR), and mTOR is involved in the sensing of nutrient availability and modulation of insulin action in vivo via its further effects to activate S6 kinase 1 (S6K1), which phosphorylates the S6 ribosomal protein to enhance protein synthesis."

sparser
"The kinase p70S6K phosphorylates the S6 protein on the 40S ribosomal subunit allowing mRNA translation xref ."

sparser
"However, S6 phosphorylation by S6K1 is only minimally influenced by removal of the CTD, or by phospho-defective or -mimetic mutations ( xref ; xref ; xref )."

reach
"During the activation of mTORC1, its substrate p70S6K phosphorylates ribosomal protein S6 [XREF_BIBR]."

reach
"Activation of p70S6K by mTORC1 via phosphorylation at threonine 389 promotes phosphorylation and activation of ribosomal protein S6 (rpS6) and eukaryotic initiation factor 4B (eIF4B)."

reach
"XREF_FIG, S6K1 siRNA reduced S6K1 expression and S6 phosphorylation but increased AKT phosphorylation and LC3-II lipidation."

reach
"PF-4708671, a specific inhibitor of S6K1, modestly inhibited S6 phosphorylation but induced the feedback activation of the PI-3 kinase pathway, evidenced by increased AKT and S6K1 phosphorylation."

reach
"mTORC1 kinase activation causes the phosphorylation of the kinase p70S6K, which then phosphorylates the ribosomal protein S6."

sparser
"Once activated by mTORC1-mediated phosphorylation, S6K1 phosphorylates ribosomal protein S6, a component of the 40S ribosomal subunit, to upregulate rRNA transcription and promote the translation elongation of spliced transcripts."

reach
"Activation of p70S6K by mTORC1 via phosphorylation at threonine 389 promotes phosphorylation and activation of ribosomal protein S6 (rpS6) and eukaryotic initiation factor 4B (eIF4B)."

reach
"S6K1 phosphorylates ribosomal protein S6 (RPS6) (a component of the 40S ribosomal subunit), leading to active translation of mRNAs involved in ribosome biogenesis [8] ."

sparser
"mTORC1 promotes translation by increasing the activity of eukaryotic translation initiation factor 4E (eIF4E) binding proteins (4E-BP1, 2, and 3) and p70 ribosomal S6 kinase (S6K), which phosphorylates the ribosomal S6 subunit, promoting eukaryotic translation ( xref )."

reach
"Furthermore mTORC1 phosphorylates and activates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (RPS6), a component of the 40S ribosomal protein which regulates the initiation of protein translation."

reach
"Furthermore mTORC1 phosphorylates and activates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (RPS6), a component of the 40S ribosomal protein which regulates the initiation of protein translation."

sparser
"It is suggested that the phosphorylation of rpS6 by S6Ks downstream of the TOR pathway is needed to promote protein synthesis required for growth processes ( xref )."

sparser
"Both RSK and S6K1 phosphorylate and activate the ribosomal subunit S6, which is essential for translation."

sparser
"It has been demonstrated that the activation of the PI3K/Akt/mTOR signaling pathway increases the phosphorylation of p70S6K and 4EBP1, and that p-p70S6K subsequently induces the phosphorylation of the ribosomal protein S6."

sparser
"Once activated, S6K1 phosphorylates S6 ( xref )."

reach
"Growth factors and mitogens influence signaling pathways and often induce the activity of p70S6 kinase (p70S6K), which in turn phosphorylates the ribosomal S6 protein (S6)."

reach
"Growth factors and mitogens influence signaling pathways and often induce the activity of p70S6 kinase (p70S6K), which in turn phosphorylates the ribosomal S6 protein (S6)."

sparser
"Phosphorylation of S6 by S6K1 mediates unwinding and initiates translation of 5 terminal oligopyrimidine tract (TOP) mRNAs."

sparser
"Activation of mTOR leads to the phosphorylation of eukaryotic translation initiation factor eIF4E-binding protein 1 (4E-BP1) and p70S6K which phosphorylates S6. xref , xref xref show that pp70S6K-Thr-389/p70SK protein ratios were increased 1.8- and 3.1-fold in 1- and 10-day-old mutant hearts compared to wild-type, respectively."

sparser
"For example, p70 S6 kinase 1 (p70S6K) phosphorylates the S6 protein, a component of the 40S ribosomal subunit, and is an intermediary between mTOR and pS6."

sparser
"P70S6 Kinase phosphorylates the 40S ribosomal subunit protein S6 and thereby allows translation of proteins involved in cell growth and hypertrophy."

sparser
"P70S6K is activated by mTORC1 and phosphorylates S6."

reach
"mTOR allows eukaryotic cells to adjust their protein biosynthetic capacity XREF_BIBR, XREF_BIBR, XREF_BIBR through downstream effectors : (1) S6K1, a kinase that phosphorylates Rps6 and promotes protein synthesis and cell proliferation, and (2) eukaryotic translation initiation factor 4E binding protein 1 (4EBP1), a protein that prevents translation when its unphosphorylated form interacts with eIF4E XREF_BIBR, XREF_BIBR, XREF_BIBR."

reach
"S6K1, in turn, phosphorylates its target - ribosomal protein S6 - on serines 240 and 244 (Ser240 and Ser244) and serines 235 and 236 (Ser235 and Ser236) [XREF_BIBR - XREF_BIBR]."

reach
"For example, mTOR activates ribosomal S6 kinase-1 (S6K1), which phosphorylates the ribosomal protein S6, promoting ribosomal biogenesis and protein translation."

reach
"P70 S6 kinase 1 (S6K1), another downstream effector of mTORC1, phosphorylates ribosomal protein S6, which also is associated with increased translation."

sparser
"Another important target of mTOR is the S6Rs, p70 S6 kinases phosphorylates the 40S ribosomal subunit protein S6 and stimulates the translation of 5′ oligopyrimidine tract containing mRNAs ( Martin an[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"p70 S6 kinase 1 (S6K1), another downstream effector of mTORC1, phosphorylates ribosomal protein S6, which also is associated with increased translation."

sparser
"P70S6K subsequently phosphorylates the ribosomal S6 subunit (S6) [ xref , xref – xref ]."

sparser
"As mentioned, S6K1 phosphorylates S6, eIF4B, as well as eEF2 kinase, suggesting that S6K1 is involved with both the initiation and elongation phases of protein synthesis xref ."

sparser
"It was originally thought that the phosphorylation of S6 by S6K1 was responsible for the translation of TOP mRNAs."

sparser
"In contrast, S6K1 phosphorylation of GST-S6 was detectable at both Ser235/Ser236 and Ser240/Ser244."

sparser
"S6K1 phosphorylates and activates S6, a ribosomal subunit involved in initiating protein synthesis machinery."

reach
"Upon activation by mTOR, S6K1 phosphorylates S6 ribosomal protein, leading to increased translation of mRNA with oligopyrimidine tract at the 5 ' terminal (5 ' TOP) [XREF_BIBR]."

sparser
"Activated S6K1 phosphorylates ribosomal protein S6, which promotes production of ribosomal proteins and elongation factors to increase translation [ xref , xref , xref ]."

sparser
"Activated S6K1 phosphorylates S6 to promote ribosome biogenesis."

reach
"Key downstream targets that modulate protein translation include eukaryotic initiation factor 4E and p70S6 kinase, the latter of which phosphorylates ribosomal S6 protein."

sparser
"After phosphorylation, p70S6K becomes activated and phosphorylates its substrate S6 protein of the 40S ribosomal subunit which is involved in translational regulation of mRNAs [ xref ]."

sparser
"mTOR-activated p70S6K phosphorylates ribosomal protein S6 and directly phosphorylates mTOR at Serine 2448 ( xref , xref ), although the biological significance of mTOR S2448 phosphorylation is not fully understood."

reach
"Both leucine and isoleucine have been shown to enhance the fractional protein synthesis rates in bovine mammary cells with the phosphorylation of mTOR and rpS6, or mTOR, S6K1, and rpS6 respectively [XREF_BIBR], while decreasing the abundance of proteasome protein, ubiquitinated protein, and the rate of protein degradation [XREF_BIBR]."

sparser
"On the other hand, activated S6K1 causes phosphorylation of S6 [33] and eIF4B [34] , which then produce an increase in eIF4A RNA helicase activity [35] ."

sparser
"P70S6K phosphorylates ribosomal protein S6, in vivo, which has been implicated in stimulating the translation of mRNA transcripts that contain a polypyrimidine tract at their 5′ transcriptional start [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Similarly, an inhibitor of p70 S6 kinase (PF-4708671) partially inhibited phosphorylation of S6 and preserved RP, while only marginally prevented cyclin D1 induction."

reach
"In mammalian cells, mTORC1 phosphorylates proteins involved in protein synthesis and cell growth, including p70S6K1, which phosphorylates the ribosomal protein S6."

reach
"Activated mTOR and ERK phosphorylate p70S6 kinase (p70S6K), which, in turn, phosphorylates the ribosomal protein S6 (RP-S6) and the eukaryotic translation initiation factor 4E (eIF4E) binding protein [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Small interfering RNA knockdown of the mTOR substrate p70 S6 kinase abrogated PAK1 phosphorylation and enhanced HCV RNA abundance, whereas overexpression of a constitutively active alternate substrate, eukaryotic translation initiation factor 4E-binding protein 1, increased cap-independent viral translation and viral RNA abundance without influencing PAK1 phosphorylation."

reach
"Activated mTOR and ERK phosphorylate p70S6 kinase (p70S6K), which, in turn, phosphorylates the ribosomal protein S6 (RP-S6) and the eukaryotic translation initiation factor 4E (eIF4E) binding protein [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"[16] The resulting activated S6K1 can phosphorylate ribosomal protein S6."

sparser
"Furthermore, S6Ks phosphorylate ribosomal p70 S6 (rpS6) stimulating translation initiation [52–55] ( Fig. 1 )."

reach
"S6K1 knockdown led to decreased S6 phosphorylation but increased the levels of LC3-II."

reach
"Similar to A77 1726, PF-4708671, a specific inhibitor of S6K1, inhibited S6 phosphorylation but induced the feedback activation of the PI-3 kinase pathway, as shown by increased AKT and S6K1 phosphorylation."

sparser
"S6K1 phosphorylates the S6 protein of the 40s ribosomal subunit at several sites, including Ser 235/236 and Ser 240/244, leading to the initiation of protein synthesis [ xref ]."

reach
"As expected, inhibition of mTORC1 greatly inhibited both p70S6K and S6 phosphorylation, whereas inhibition of p70S6K diminished S6 phosphorylation without impairing mTORC1 activity (as judged by p70S6K phosphorylation)."

sparser
"Since activated S6K1 can phosphorylate the ribosomal protein S6, making it easier to translate mRNAs with a 5-terminal oligopolypyrimidine, this shows that activation of S6K1 speeds up the creation of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"One target of mTOR is the ribosomal S6 kinase (S6K1) that phosphorylates the ribosomal protein S6 XREF_BIBR."

sparser
"Treatment of mammalian cells with rapamycin abolishes in vivo S6 phosphorylation by p70s6k; however, ectopic expression of AtS6k2 rescues the rapamycin block."

sparser
"P70 S6K kinase directly phosphorylates the 40S ribosomal protein S6, which results in enhanced translation of proteins that contain a polypyrimidine tract in the 5′-untranslated region [28] ."

sparser
"S6 is phosphorylated by both S6K1 and the closely related S6K2 on Ser 240 /Ser 244 as well as Ser 235 /Ser 236 ."

sparser
"mTOR is known to regulate the p70 S6K kinase, which phosphorylates ribosomal protein S6."

reach
"The most well characterized rapamycin sensitive mTORC1 target is S6K1, a protein kinase that phosphorylates the ribosomal subunit S6 to control translation initiation."

sparser
"For translational regulation, a clue as to how Rac acts may lie in its reported ability to interact with p70S6K , which phosphorylates the ribosomal protein S6 and thereby promotes the translation of [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"S6K1 then phosphorylates ribosomal protein S6, a component of the 40S ribosomal subunit and regulates mRNA translation."

sparser
"mTORC1 controls mRNA translation, in part, by phosphorylating and activating S6K1 which phosphorylates the S6 protein of the 40S ribosomal protein ( xref )."

reach
"In addition to the eIF4E binding proteins, p70 S6k1 also phosphorylates rpS6 [35] and eIF4B [36] ."

reach
"A caveat to this idea is that rpS6 phosphorylation is also enhanced by mitogens in S6k1 −/− /S6k2 −/− cells, probably through activation of p90 rsk , implying that although p70 S6k1 is not required fo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Whereas mTORC1-mediated phosphorylation of 4E-BP1 induces its dissociation from eIF4E, the rate-limiting factor for initiation of Cap-dependent translation ( xref ), phosphorylated S6K1 phosphorylates ribosomal protein S6, an essential regulator of protein translation and cell growth ( xref )."

reach
"Unstimulated COS cells contain active p70S6K, which phosphorylates ribosomal S6 protein as well as CREM in vitro."

reach
"mTOR phosphorylates S6K1 at T389 to enhance S6K1 activity, which amongst other things phosphorylates the S6 subunit of the ribosome to promote translation."

sparser
"P70S6K phosphorylates RPS6 and leads to an increase in the rate of translation of 5′TOP mRNA transcripts which encode components of the cellular translational apparatus, thus facilitating an increase [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"While p70S6K can phosphorylate RPS6 at all sites, S235/236 is a specific target for RSK [ xref ]."

sparser
"The activated p70s6k then phosphorylates 40S ribosomal protein S6, leading to an increased rate of translation of the class of 5′TOP mRNA transcripts."

reach
"Likewise, activated S6K1 phosphorylates ribosomal protein S6 that also participates in translation initiation."

reach
"mTOR signaling leads to the activation of p70S6 kinase, which phosphorylates S6 ribosomal protein."

reach
"For instance, in an mTOR independent manner, phospholipase D-2-derived phosphatidic acid directly binds and activates p70S6K, 55 and p90 ribosomal S6 kinases directly phosphorylate RPS6."

sparser
"Activated mTORC1 can also phosphorylate and activate S6K1 at T389 to further facilitate translation initiation and elongation via S6K1-induced subsequent phosphorylation of the ribosomal protein S6 (RPS6), eIF4B and elongation factor eEF2K, respectively [ xref ]."

reach
"Active mTORC1 in turn phosphorylates p70S6K, which then phosphorylates eukaryotic initiation factor 4B (eIF4B) and ribosomal protein S6 (RPS6), which are directly involved in the formation of the mRNA translation pre and initiation complex (XREF_FIG) XREF_BIBR."

sparser
"Although S6Ks phosphorylate ribosomal protein S6, it is not clear whether or how this affects mRNA translation."

reach
"Active mTORC1 phosphorylates p70S6K and, in turn, activated p70S6K phosphorylates RPS6 XREF_BIBR."

reach
"Stimulation of mammalian cells with growth factors or amino acids leads to phosphorylation and activation of p70 S6 kinase, which in turn phosphorylates the ribosomal protein S6 (Chung et al., 1992; J[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"mTORC1 is sensitive to rapamycin and phosphorylates its downstream effectors, including p70 S6 kinase, which in turn phosphorylates ribosomal S6 protein, thereby promoting protein biosynthesis."

sparser
"In response to mitogen stimulation, mTOR phosphorylates and activates S6K1, which in turn phosphorylates the 40S ribosomal protein S6, leading to the enhancement of translation of mRNAs with a 5′-terminal oligopyrimidine, including mRNAs that encode for ribosomal proteins and elongation factor-1."

reach
"There are several documented downstream targets for Akt, including p70S6 kinase, which phosphorylates the S6 ribosomal protein (reviewed in [43])."

sparser
"We have recently identified S6K2, a homolog of S6K1, which phosphorylates S6 in vitro and is regulated by the phosphatidylinositide 3-kinase (PI3-K) and mammalian target of rapamycin pathways in vivo."

reach
"However, rpS6 can also be phosphorylated by S6K1 on both its Ser235/236 and Ser240/244 regulatory sites [XREF_BIBR]."

sparser
"S6K1 phosphorylates ribosomal protein S6 (RPS6) (a component of the 40S ribosomal subunit), leading to active translation of mRNAs involved in ribosome biogenesis [8] ."

sparser
"In addition to the eIF4E binding proteins, p70 S6k1 also phosphorylates rpS6 [35] and eIF4B [36] ."

reach
"Phosphorylation of the ribosomal S6 protein by S6K1 regulates the translation of both ribosomal proteins and initiation factors, leading to an increase in protein synthesis."

reach
"P70S6 kinase phosphorylates ribosomal protein S6, and possibly the initiation factors eIF4B and eIF4G."

sparser
"These results are consistent with the conclusion that phosphorylation of S6(S240) by S6K1 is an essential prerequisite for subsequent phosphorylation of S6(S235) by RSK."

reach
"The best known rapamycin sensitive mTORC1 target is S6K1, a protein kinase that phosphorylates the ribosomal subunit S6 to control translation initiation."

reach
"S6K1 phosphorylates ribosomal protein S6 (S6) and enhances the translation of mRNAs [XREF_BIBR, XREF_BIBR]."

reach
"Furthermore, mTORC1 phosphorylates p70 ribosomal protein S6 (rpS6) kinase beta-1 (S6K1), which stimulates protein synthesis through phosphorylating rpS6 protein, a component of the 40S ribosomal subunit (16)."

sparser
"The mTORC1 controls the rate of protein synthesis through phosphorylation and activation of its substrates, p70S6 ribosomal kinase 1 (p70S6K) and eukaryotic translation initiation factor 4E (eIF4E) binding protein-1 (4E-BP1) and once phosphorylated, p70S6K phosphorylates ribosomal protein S6 and 4E-BP1 becomes dissociated from eIF4 and promote mRNA translation and protein synthesis [ xref ]."

sparser
"Furthermore, S6K1 phosphorylates its substrate ribosome S6 protein, which promotes the initiation and processes of protein translation [ xref ]."

reach
"The S6 ribosomal protein belongs to the PI3K/AKT/PTEN/mTOR signaling pathway and is phosphorylated by p70 S6 kinase when this pathway is activated."

reach
"FRAP may activate p70 S6 kinase which phosphorylates the S6 subunit of the 40S ribosomal subunit, an event that correlates with growth factor regulated increases of translation of defined, growth regu[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The most well characterized rapamycin sensitive mTORC1 target is S6K1, a protein kinase that phosphorylates the ribosomal subunit S6 to control translation initiation."

sparser
"P70S6K kinase directly phosphorylates the 40S ribosomal protein S6, which results in enhanced translation of proteins that contain a polypyrimidine tract in the 5′-untranslated region [ xref ]."

sparser
"S6 is phosphorylated by the p70 S6 kinase directly downstream of the mammalian target of rapamycin (mTOR) complex 1 (TORC1) [ xref ]."

sparser
"The activated mTORC1 phosphorylates and activates S6K1 which subsequently phosphorylates the 40s protein S6 of ribosome resulting to initiation of mRNA translation with a 5′-terminal oligopyrimidine ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In conclusion, our data show that mTORC1 and S6K1 influence T cell activation and differentiation independently of their capacity to mediate rpS6 phosphorylation."

reach
"In turn, phosphorylated (P-) p70S6K phosphorylates S6 ribosomal protein (S6RP), which also stimulates translation."

sparser
"Finally, the p70 S6 kinases also phosphorylate RPS6 and the eukaryotic elongation factor 2α (eEF2α) kinase (eEF2K) [ xref , xref ]."

sparser
"RPS6KB1 phosphorylates the 40 S ribosomal protein S6 in response to mitogen allowing the upregulation of mRNA translation."

sparser
"Subsequently, p70S6K phosphorylates ribosomal protein S6 to selectively translate 5’-terminal polypyrimidine mRNA encoding ribosomal proteins and translation factors [ xref ], while phosphorylating 4E-BP1 would enable dissociation from eukaryotic initiation factor 4E (eIF4E)."

reach
"The p70-S6K1 protein is activated by mTOR and subsequently phosphorylates the ribosomal S6 protein (RPS6), eukaryotic transcription initiation factor 4B (EIF4B), and eukaryotic elongation factor 2-kinase (eEF2k) and is a positive regulator of protein translation."

reach
"Increased p70S6K activity by mTOR activity induces the phosphorylation of the ribosomal protein S6, which induces protein synthesis, 4E-BP1 inhibits eIF4E activity."

reach
"Although S6K1 was first identified as a kinase of ribosomal protein S6 (rpS6), the functional purpose of rpS6 phosphorylation by S6K1 is still unresolved."

sparser
"S6K1 phosphorylates the S6 protein of the 40s ribosomal subunit at several sites, including Ser 235/236 and Ser 240/244, leading to initiation of protein synthesis ( xref ; xref )."

sparser
"AKT phosphorylation leads to activation of mammalian target of rapamycin (mTOR), a protein kinase that controls cell proliferation via activation of the eukaryotic initiation factor 4E-binding protein-1 complex and S6K1, which phosphorylates the ribosomal S6 protein."
| PMC

sparser
"Since LKB1 is a regulator of mTOR activity through the phosphorylation of the energy sensor AMPK, in the absence of this regulation, mTOR is active whereby it phosphorylates S6K1, that in turn phosphorylates pS6, ultimately leading to protein translation xref ."

sparser
"In mammalian cells, mTORC1 phosphorylates proteins involved in protein synthesis and cell growth, including p70S6K1, which phosphorylates the ribosomal protein S6."

sparser
"Activated mTORC1 phosphorylates S6K1, which phosphorylates S6 (40S ribosomal protein S6), enhancing the translation of mRNAs with a 5′-terminal oligopolypyrimidine (5′-TOP)."

reach
"In addition, we examined two phosphorylation sites on rpS6, Ser240/244, that are uniquely phosphorylated by S6K1 and S6K2."

sparser
"During the activation of mTORC1, its substrate p70S6K phosphorylates ribosomal protein S6 [ xref ]."

sparser
"Additionally, mTOR phosphorylates and activates p70S6K and 4E-BP1, which in turn phosphorylates S6 and triggers the ribosomal 40S subunit to participate in protein translation [ xref ]."

sparser
"In particular, phosphorylated p70S6K enhances phosphorylation and activity of ribosomal protein S6 and initiates various translation activities required for cell growth."

sparser
"P70S6K phosphorylates ribosomal protein S6 (S6) to promote protein synthesis (Taniguchi et al ., xref )."

reach
"mTOR triggers activity of ribosomal S6 kinase-1 (S6K1), which phosphorylates the ribosomal protein S6, promoting ribosomal biogenesis and protein translation."

reach
"Once phosphorylated, 4E-BP1 dissociates from eIF4E whereas p70S6K phosphorylates ribosomal protein S6, thus promoting mRNA translation [XREF_BIBR]."

sparser
"P70S6 kinase phosphorylates the ribosomal protein S6 with subsequent enhancement of translation of mRNAs bearing a 5′-terminal oligopyrimidine tract."

reach
"In addition, the p70S6K immunoprecipitated from nuclear extracts of unlabeled COS cells phosphorylates S6 protein in an in vitro immunokinase assay."

sparser
"mTORC1 promotes protein synthesis by phosphorylation of p70S6 kinases and eukaryotic initiation factor 4E (eIF4E)-binding proteins (4E-BPs), and subsequently, the p70S6 kinase phosphorylates the S6 protein."
| PMC

sparser
"Therefore, we might hypothesize that the phosphorylation of rpS6 by p70S6K is under the control of a CK2-mediated regulatory mechanism."

sparser
"P70S6K not only phosphorylates the small ribosomal subunit RPS6 to increase translation but also enhances recruitment of mRNAs with a 5′ terminal oligopyrimidine (TOP) sequence to polyribosomes for efficient translation."

sparser
"However, rpS6 can also be phosphorylated by S6K1 on both its Ser235/236 and Ser240/244 regulatory sites [ xref ]."

sparser
"Furthermore, p70S6K phosphorylates ribosomal protein S6 (S6RP)."

sparser
"Accordingly, mTOR regulates protein synthesis by hyperphosphorylating 4EBP-1, which decreases its affinity for the translation initiation factor eIF4E [ xref ] and phosphorylating and activating p70S6K, which in turn phosphorylates the 40S ribosomal subunit S6 protein [ xref ]."

sparser
"These results show that haloperidol promotes mTORC1- and S6K1-dependent phosphorylation of rpS6 at Ser240/244, in a subpopulation of striatal MSNs expressing D2Rs."

sparser
"Phosphorylated p70S6K further phosphorylates the 40S ribosomal protein S6, leading to enhancement of the translation of mRNAs."

sparser
"Increased mTORC1 activity ultimately results in the phosphorylation of multiple protein substrates, including p70S6K; when activated, p70S6K phosphorylates its substrate RPS6."

reach
"S6K1 deficient mice phosphorylated S6 but had a small body phenotype."

reach
"Phosphorylation (denoted by P in circle) activates S6K1 which in turn phosphorylates S6, a ribosomal protein."

reach
"mTOR is activated by AKT and it stimulates protein translation by phosphorylating eIF4E binding protein (4E-BP1) as well as p70S6K which phosphorylates S6."

sparser
"Consequently, p70S6K phosphorylates ribosomal protein S6, resulting in increased translation of mRNA messages containing 5′ terminal oligopolypyrimidine (5′TOP) tracts."

reach
"S6K is represented by two homologous cellular proteins, S6K1 and S6K2, both of which act downstream of mTOR and phosphorylate S6."

reach
"AKT1S1 (PRAS40) contains a consensus phosphorylation motif and is a direct substrate for AKT, a mediator of PI3K signaling [XREF_BIBR], while RPS6 is phosphorylated at Ser235/236 by p70S6 kinase."

reach
"Both mTOR and Akt are upstream activators of translational regulators such as p70 S6-kinase, an enzyme that phosphorylates rpS6."

reach
"Whereas mTORC1 mediates cell growth and organ size through S6K1-dependent phosphorylation of ribosomal protein S6 (rpS6), mTORC1-dependent 4E-BP1 phosphorylation mediates cell proliferation."

sparser
"Wang C, Mao X, Wang L, Liu M, Wetzel MD, Guan KL et al. Adiponectin sensitizes insulin signaling by reducing p70 S6 kinase-mediated serine phosphorylation of IRS-1."

sparser
"In addition, NAD(P)H oxidase p22phox, p47phox, gp91phox expression, and mitogen-activated protein kinase and its downstream effector p70 S6 kinase phosphorylation in DS rats was also inhibited by telmisartan."

sparser
"In turn S6K1 phosphorylates and activates S6, a ribosomal protein involved in translation xref , xref , xref ."

reach
"However, recent data suggest that ribosomal S6 phosphorylation by S6K1 is not acting alone in this growth pathway as growth factors are able to stimulate translation of 5 ' TOP containing mRNAs even i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"S6 is phosphorylated at Ser235/236 by p70S6 kinase."

reach
"Silencing of S6K1 or S6K2 effectively depleted the homolog and attenuated phosphorylation of the substrate S6."

sparser
"S6 is phosphorylated by S6K1, which itself is a target of mTOR."

reach
"S6K1 phosphorylates rpS6, whereas protein phosphatase 1 negatively regulates rpS6 phosphorylation of all 5 sites in intact cells."

sparser
"Activated RPS6KB1 phosphorylates RPS6 (ribosomal protein S6) and other components involved in the translation machinery, enhancing the translation of mRNAs that encode ribosomal proteins and translation factors [ xref ]."

reach
"Pharmacologic inhibition of S6K1 decreased rpS6 phosphorylation, leading to blunted podocyte hypertrophy and FSGS lesions."

reach
"We proposed that, regardless of etiology, the initial podocyte injury or podocyte loss activates mTORC1 signaling of S6K1-dependent phosphorylation of rpS6, resulting in hypertrophy of the remaining podocytes."

sparser
"mTORC1 phosphorylates and activates p70S6K which in turn phosphorylates ribosomal protein S6 to enhance translation efficiency xref ."

sparser
"Upon activation, S6K1 phosphorylates its downstream target ribosomal protein S6 (rpS6) on a cluster of five serine residues at the carboxy terminus [ xref , xref ]."

sparser
"RPS6 is phosphorylated by p70S6K, which in turn is activated by mTOR ( xref )."

reach
"Moreover, S6K1 is a major kinase that phosphorylates rpS6, and the S6K1 inhibitor PF4708671 has been demonstrated to selectively inhibit rpS6 phosphorylation."

sparser
"S6K1 phosphorylates ribosomal protein S6 (S6) and enhances the translation of mRNAs [ xref , xref ]."

reach
"These include the ribosome kinase S6K1 (p70S6K1), which phosphorylates the ribosomal protein rpS6 to facilitate protein synthesis."

reach
"One such substrate is p70S6 kinase, which phosphorylates ribosomal protein S6 as well as several other proteins XREF_BIBR."

sparser
"P70S6K phosphorylates ribosomal protein S6 and regulates protein translation [11] ."

sparser
"mTOR phosphorylates downstream targets such as p70S6 kinase (p70S6K) which then phosphorylates ribosomal protein S6 (S6) [ xref ]."

sparser
"Anabolic pathways that promote fiber hypertrophy result in activation via phosphorylation of protein kinase B (Akt) and mechanistic target of rapamycin (mTOR), which leads to phosphorylation and activation of p70 S6 kinase (p70S6K) that in turn phosphorylates the ribosomal protein S6 [ xref ]."

reach
"mTORC1 kinase modulates translation by phosphorylating two substrates, the 4E-BP1 protein and S6K1 kinase, which in turn phosphorylates the ribosomal protein S6 (rpS6)."

sparser
"The p70 S6 kinase-mediated phosphorylation of rictor (Thr1135) has been reported to be ( xref ) or not to be ( xref ) a negative regulator of mTORC2-mediated phosphorylation of Akt(Ser473)."

sparser
"Phosphorylated S6K1 further phosphorylates ribosomal protein S6 (S6) to promote ribosome biogenesis."

reach
"mTOR also activates S6K-1, which phosphorylates ribosomal protein S6, resulting in increased translation."

reach
"S6K-1, which is phosphorylated and activated by mTOR, in turn, phosphorylates ribosomal protein S6, which is a commonly used readout of mTOR activity."

reach
"mTOR is the key component of mTORC1 complex that promotes protein synthesis largely through the phosphorylation of two effectors, p70 S6 kinase (S6K1, which further phosphorylates S6) and 4E-BP1 [XREF_BIBR]."

reach
"Genetic reduction of S6K1 in Fmr1-KO mice prevents the increased phosphorylation of S6 and eIF4B and the increased levels of eEF2, and this correlates with the resetting of protein synthesis to levels comparable to those in wild-type mice 15."

sparser
"P70S6K directly phosphorylates the 40S ribosomal protein S6, which correlated with enhanced translation of transcripts with 5’-terminal oligopyrimidine (5’-TOP) sequences that encode components of tra[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The serine/threoinine kinase mTOR phosphorylates the 70 kDa ribosomal protein (rp) S6 kinase (p70 S6k1 ), which in turn phosphorylates and activates the eIF4E binding proteins, rpS6, and eIF4B."

reach
"S6K1 phosphorylates multiple sites of the 40 S ribosomal protein S6 resulting in a specific increase in the translation of a subset of mRNAs containing a polypyrimidine tract in their 5 '-untranslated region (5 ' TOP mRNAs) XREF_BIBR."

sparser
"Furthermore mTORC1 phosphorylates and activates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (RPS6), a component of the 40S ribosomal protein which regulates the initiation of protein translation (see review [ xref ])."

reach
"Although the deregulation of the upstream regulatory mechanisms differs between the E198K and E420K variant cell lines, inhibition of mTORC1 with rapamycin or of p70S6K with LY2584702 reduces RPS6 phosphorylation in both variant cell lines."

reach
"mTORC1 phosphorylates proteins involved in protein synthesis and cell growth, including 4E-BP1 and p70S6 kinase 1 (S6K1), which phosphorylates the ribosomal protein S6."

sparser
"The activated S6K1 phosphorylates ribosomal protein S6 (RpS6), and promotes protein translation."

reach
"Furthermore, inhibition of p70S6K with LY2584702 reduced RPS6 phosphorylation at S235:236 and S240:244 in the E420K and E198K variant cell lines (Figure 6D–F)."

reach
"The serine/threonine kinase p70 S6 kinase (p70S6K) phosphorylates the 40 S ribosomal protein S6, modulating the translation of an mRNA subset that encodes ribosomal proteins and translation elongation factors."

sparser
"Importantly, inhibition of PI3K pathway activation by GDC-0941 was clearly manifested by decreases in ribosomal p70 S6 kinase (S6 kinase) phosphorylation (p-S6 kinase, Ser235/236 and Ser240/244), particularly in the 5-FU-tolerant cells."

reach
"The serine/threonine kinase p70 S6 kinase (p70S6K) phosphorylates the 40 S ribosomal protein S6, modulating the translation of an mRNA subset that encodes ribosomal proteins and translation elongation factors."

sparser
"Detection of LPS by TLR4 activates PI3K through the recruitment of the MyD88 and TRIF adaptors, leading to AKT and mTOR activation and finally ending by S6 protein phosphorylation by S6K1."

reach
"For translational regulation, a clue as to how Rac acts may lie in its reported ability to interact with p70S6K (Bishop and Hall, 2000), which phosphorylates the ribosomal protein S6 and thereby promo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"RPS6KB1 phosphorylates specifically ribosomal protein S6."

reach
"A major downstream molecule of the mTOR axis is p70 S6 kinase, which, activated by mTOR, phosphorylates the S6 ribosomal protein to initiate protein synthesis."

reach
"P70S6K phosphorylates ribosomal protein S6 (S6) to promote protein synthesis."

reach
"mTORC1 phosphorylates and activates p70S6K which in turn phosphorylates ribosomal protein S6 to enhance translation efficiency XREF_BIBR."

reach
"S6K1 phosphorylates ribosomal protein S6 (RPS6) which is an essential component of the 40S ribosomal subunit."

sparser
"It is well established that S6 can be phosphorylated at S 235 and S 236 sites by p70S6K. Taking S6 as the substrate, the consensus motif (R/KXR/KXXS*/T*) of p70S6K was determined using the NetPhos2.0p[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"S6K1 (p70S6 kinase) phosphorylates S6 which controls the translation of mRNA that possesses an unusual oligopyrimidine tract at its transcription starting site termed 5 ' TOP."

reach
"S6K1 (p70S6 kinase) phosphorylates S6 which controls the translation of mRNA that possesses an unusual oligopyrimidine tract at its transcription starting site termed 5 ' TOP."

reach
"mTORC1 phosphorylates the p70S6 kinase (p70S6K), which in turn phosphorylates the S6 ribosomal protein and 4E-BP1, leading to protein translation."

reach
"mTORC1 phosphorylates the p70S6 kinase (p70S6K), which in turn phosphorylates the S6 ribosomal protein and 4E-BP1, leading to protein translation."

reach
"S6 phosphorylation is mediated by S6K1, which requires phosphorylation at its ' T loop ' site by PDK1."

sparser
"However an alternate event, p70 S6 kinase phosphorylation of IRS-1 on S636/639, could also occur resulting in decreased IGF-1R signaling."

sparser
"Since phosphorylation of S6 ribosome protein (S6RP) by p70S6K can induce protein synthesis at the ribosome, the dramatic reduction of p70S6K after ATA treatment led to great reductions of new protein synthesis on several cell cycle-related proteins including cyclin D3, aurora kinase A, polo-like kinase, cyclin B1, survivin; and reduced the levels of EGFR and MET."

reach
"This complex directly phosphorylates p70 ribosomal protein S6 kinase 1 (p70S6K1, phosphorylates ribosomal protein S6) and eukaryote translation initiation factor 4E binding protein 1 (4E-BP1) to regulate cell growth and survival [XREF_BIBR]."

sparser
"Treatment of cells with the phosphatidylinositol 3-kinase (PI3K) inhibitors wortmannin and LY294002 [2-(4-morpholinyl)-8-phenyl-4H-1-benzopyran-4-one] or rapamycin, an inhibitor of mammalian target of rapamycin and ribosomal p70 S6 kinase (p70S6K) phosphorylation, or with an adenovirus containing green fluorescent protein and a dominant-negative and kinase-dead Akt, effectively inhibited the insulin-mediated increase in alpha-class GST expression and GST activity toward NBD."

reach
"Similarly, changes in muscular mTOR pathway signaling were probed by measuring insulin induced phosphorylation of S6 kinase 1 (S6K1)."

reach
"S6K1 phosphorylates ribosomal protein S6 (rpS6) leading to increased translation of ribosomal and transcription factor mRNA resulting in protein translation initiation."

reach
"P70S6 kinase phosphorylates the ribosomal protein S6 with subsequent enhancement of translation of mRNAs bearing a 5 '-terminal oligopyrimidine tract."

sparser
"Once activated, S6K1 phosphorylates rpS6 and eIF4B (reviewed in Fingar and Blenis, 2004 )."

reach
"Alterations in cell size are associated with changes in activity of mTORC1 acting through its downstream targets S6Kinase1 (S6K1), which phosphorylates ribosomal S6 protein (RPS6), and 4EBP1."

reach
"Although S6 ribosomal protein (S6) is phosphorylated by p70S6 kinase and therefore lies downstream of mTOR, previous studies have not found a correlation between S6 phosphorylation and either PTEN levels or PKB and Akt phosphorylation in primary GBM specimens [XREF_BIBR], likely reflecting the importance of other pathways in controlling mTOR activity in these tumors."

sparser
"To determine if mTORC1 activity is enhanced in GSK3-deficient B cells, we measured the abundance of mTORC1-phosphorylated p70S6K (T389), as well as p70S6K-phosphorylated ribosomal protein S6 (S235, S236, S240 and S244) in anti-CD40+IL-4–stimulated B cells ( xref )."

reach
"While S6 is predominantly phosphorylated by p70S6K in most mammalian adult tissues, phosphorylation of S6 by p90rsk is important during meiosis in mammalian and amphibian oocytes [XREF_BIBR, XREF_BIBR], and to a lesser extent in growth factor stimulated hepatocytes [XREF_BIBR]."

sparser
"The antagonist effect of dominant kinase deficient PKC shows PKC can activate p70 and acts to integrate signals through PI3K and the Rapamycin sensitive pathway detailed previou[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Phosphorylation of ribosomal protein S6, which is mediated by an mTORC1 target p70 S6 kinase (S6K), is dramatically increased in tumors isolated from Sesn2 -/- mice (XREF_FIG and XREF_FIG)."

sparser
"S6K1 phosphorylates the S6 protein on the 40S ribosomal subunit and leads to cell proliferation."

sparser
"S6K1 phosphorylates ribosomal protein S6, in some models enhancing the translation of mRNAs possessing a 5′ terminal oligopyrimidine tract ( xref )."

sparser
"Downstream substrates become phosphorylated including glycogen synthase kinase 3α and β (GSK3α/β, which are inhibited), and p70 ribosomal S6 kinases (p70S6K) which phosphorylate the small ribosomal subunit Rps6 to regulate translation of specific mRNAs, particularly increasing the protein synthetic machinery."

reach
"More downstream, the ribosomal protein S6 is phosphorylated by p-p70S6K1."

sparser
"After phosphorylation by activated mTORC1, S6K1 phosphorylates S6 (40S ribosomal protein S6), which enhances mRNA translation through 5 ′- terminal oligopyrimidine (5′-top)."

sparser
"S6K1 phosphorylates ribosomal protein S6 (RPS6), which can increase translation rates by stimulation of ribosome biogenesis facilitated by increased translation of mRNAs containing 5′ (TOP) sequences."

reach
"In LN229 and GAMG glioblastoma cells treated with either non targeting or PTEN targeting siRNA (siNT and siPTEN, respectively), both AD80 and LY-2779964 (LY64) were effective in reducing the S6K1 dependent phosphorylation of the ribosomal protein S6 (rpS6) at 3 hours (XREF_FIG, XREF_SUPPLEMENTARY)."

reach
"In LN229 and GAMG glioblastoma cells treated with either non targeting or PTEN targeting siRNA (siNT and siPTEN, respectively), both AD80 and LY-2779964 (LY64) were effective in reducing the S6K1 dependent phosphorylation of the ribosomal protein S6 (rpS6) at 3 hours (XREF_FIG, XREF_SUPPLEMENTARY)."

sparser
"MTOR phosphorylates a series of growth- and metabolism-associated substrates, among which the best-characterized target is ribosomal protein S6 (RPS6) kinase 1 (S6K1). xref MTOR phosphorylates S6K1 at Thr389, and activated S6K1 further phosphorylates RPS6 to regulate protein translation, synthesis, and metabolism. xref To investigate the role of MET in MTOR activation, we used WT and Met -KO HepG2 cells and observed that the amino acid-enhanced phosphorylation of S6K1 at Thr389 was repressed in MET-KO cells (Fig. xref )."

reach
"The S6K1 gene can promote protein translation by stimulating the phosphorylation of downstream eIF-4B, RPS6, eIF-2, and PAPB [40], and the SLC3A2, IRS1, PDK, P13K, TSC1, TSC2, mTORC1, eIF4EBP1, S6K1, and eIF4B genes are related to nitrogen metabolism in the liver; moreover, these genes would become overexpressed when blood ammonia increased to increase urea synthesis and balance the blood ammonia [41]."

sparser
"S6K1 phosphorylates rpS6, whereas protein phosphatase 1 negatively regulates rpS6 phosphorylation of all 5 sites in intact cells."

sparser
"RPS6KB1 is ribosomal protein S6 kinase B1, and its upstream is the key protein mTOR, which activates RPS6KB1 through phosphorylated mTOR and then activated RPS6KB1 phosphorylates its downstream molecules RPS6 and EIF4B, thereby controlling the downstream process of protein synthesis during cell growth ( xref ; xref )."

reach
"Phosphorylation activated p70S6K then phosphorylates S6 protein (small ribosomal S6) and eIF4B (eukaryotic translation initiation factor 4B)."

sparser
"P70 S6 kinase phosphorylates ribosomal S6 protein, resulting in an increase of the protein synthesis complex."

reach
"P70 S6 phosphorylates the 40S ribosomal subunit protein S6 and stimulates the translation of the 5 ' oligopyrimidine tract containing mRNAs [XREF_BIBR]."

sparser
"Active p70S6K phosphorylates the ribosomal protein S6, enhancing translation of mRNA transcripts involved in cell cycle progression ( xref )."

sparser
"For example, Kim et al. (2004) reported that the PI3K inhibitors (wortmannin and LY294002), dominant-negative Akt, and rapamycin, an inhibitor of mTOR (mammalian target of rapamycin) and ribosomal p70[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"It was originally thought that the phosphorylation of S6 by S6K1 was responsible for the translation of TOP mRNAs."

sparser
"S6K1 in turn phosphorylates the ribosomal protein S6 ( xref )."

reach
"Active S6K1 can stimulate the protein synthesis by phosphorylating S6 ribosomal proteins."

reach
"Indeed, the exogenous leucine was able to restore TCR and CD28 stimulated phosphorylation of S6 and its kinase S6K1 (XREF_SUPPLEMENTARY)."

sparser
"Well-characterized effector targets of mTORC1 includes 4E-BP1 (4E-binding protein), ribosomal protein S6 kinase and S6K1 (p70S6 kinase) which, in turn, phosphorylates the ribosomal protein S6 (S6/RPS6) ( xref )."

sparser
"Activated RPS6KB1 phosphorylates ribosomal protein S6 leading to an increase in protein synthesis, cell growth/size, and cell survival xref ."

reach
"S6K1 phosphorylates the ribosomal protein S6 (RPS6) thereby connecting the TOR pathway with ribosomes."

sparser
"S6K1 may phosphorylate and activate 40S ribosomal protein S6."

sparser
"mTORC1 triggers phospho-S6K1 which in turn phosphorylates S6, a critical protein in ribosomal constitution and thus in protein synthesis."

reach
"S6K1 phosphorylates ribosomal protein S6, in some models enhancing the translation of mRNAs possessing a 5 ' terminal oligopyrimidine tract."

sparser
"P70S6K phosphorylates ribosomal protein S6 (rpS6), which is a crucial component of the ribosome itself and initiates translation [ xref ] ( xref )."

sparser
"Upon activation, p70S6K directly phosphorylates the 40S ribosomal protein S6, which correlates with enhanced translation of transcripts with a 5'-terminal oligopyrimidine tract, such as elongation factor-1 alpha and ribosomal proteins [ xref ]."

reach
"P70S6K is downstream of the mTOR pathway and directly phosphorylates RPS6, a marker of the activation of both pathways ."

reach
"Activated mTOR phosphorylates the eukaryotic initiation factor 4E binding protein 1 (4E-BP1) and the p70 ribosomal S6 kinase 1 (S6K1), which phosphorylates the ribosomal protein S6 and consequently regulates protein synthesis."

reach
"The PI3K and mTOR signaling pathway controls phosphorylation of 4E-BP1 and S6K, and S6K1 also phosphorylates S6, and has been proposed to phosphorylate EF2K and pdcd4, controlling the activity of S6 and EF2K, and protein stability of pdcd4 [XREF_BIBR, XREF_BIBR - XREF_BIBR]."

reach
"This subsequently leads to increased phosphorylation of eIF4E binding protein 1 (4EBP1) and activation of p70 ribosomal S6 protein kinase (p70 S6K), which phosphorylates S6 ribosomal protein XREF_BIBR."

reach
"mTOR also allows translation initiation by phosphorylating and activating the kinase p70S6K which in turn phosphorylates the 40S ribosomal subunit S6 protein 30."

reach
"Once phosphorylated, S6K1 regulates protein translation in part by phosphorylating and activating ribosomal protein S6 (rpS6)."

sparser
"P70 S6 kinase 1 (S6K1), another downstream effector of mTORC1, phosphorylates ribosomal protein S6, which also is associated with increased translation ( xref , xref )."

reach
"P70S6K phosphorylates and activates S6, a ribosomal subunit involved in initiating translation of 5 ' terminal oligopyrimidine tract containing mRNA encoding components of the protein synthesis machinery."

reach
"This result is consistent with the RxRxxS motif on S6 that is known to be phosphorylated by S6K1 (Flotow and Thomas, 1992)."

reach
"Among other things, mTOR activates the ribosomal protein S6 kinase 1 (S6K1), which phosphorylates the ribosomal S6 protein, leading to increased cell growth."

sparser
"In addition, it phosphorylates S6K1, which in turn phosphorylates small ribosomal subunit protein RPS6."

reach
"S6 phosphorylation is mediated by the 70-kD ribosomal S6 kinase 1 (S6K1) which requires phosphorylation at its " T loop " site by PDK1."

reach
"Subsequently, S6K1 phosphorylates ribosomal protein rpS6, eIF4B, and eukaryotic elongation factor (eEF2) kinase."

sparser
"Once phosphorylated, 4E-BP1 dissociates from eIF4E whereas p70S6K phosphorylates ribosomal protein S6, thus promoting mRNA translation [ xref ]."

reach
"Upon activation, p70S6K phosphorylates ribosomal protein S6 (RPS6) and other downstream targets involved in protein synthesis and cell growth."

reach
"Rapamycin strongly inhibits S6K1 phosphorylation of S6 but also has a minor effect on Akt phosphorylation."

reach
"The p70 S6 kinase (p70S6K), which is activated by mTOR and phosphorylates RPS6, is also inhibited in SU-DHL-1 cells treated with U0126 [XREF_BIBR], but surprisingly not in the SR-786 ALK+ ALCL cell line [XREF_BIBR]."

reach
"mTORC1 controls mRNA translation via at least two well-established downstream targets: p70S6 kinase (p70S6K), which phosphorylates several translation factors including the ribosomal protein S6 (pS6), and repressor proteins of eukaryotic initiation factor 4E (eIF4E) termed 4E-BPs [5]."

reach
"mTORC1 controls mRNA translation via at least two well-established downstream targets: p70S6 kinase (p70S6K), which phosphorylates several translation factors including the ribosomal protein S6 (pS6), and repressor proteins of eukaryotic initiation factor 4E (eIF4E) termed 4E-BPs [5]."

sparser
"Upon activation, p70S6K phosphorylates the S6 protein of ribosomal subunit 40S leading to the selective translation of protein synthesis-related mRNA family [ xref – xref ]."

sparser
"mTORC1 activates p70S6K, which in turn phosphorylates the ribosomal protein S6 and 4E-BP1, both involved in protein translation."

reach
"However, our data indicated that inhibition of IKK had no impact on palmitate induced phosphorylation of S6 protein, a downstream of S6K1."

reach
"This confirmed that miR-455-5p inhibited the phosphorylation of RPS6 by targeting RPS6KB1, which in turn decreased the translation process of the protein."

sparser
"The p70S6K protein phosphorylates the 40S ribosomal protein S6 and the translation initiation factor eIF4B. The pivotal role of mTOR pathway in the generation of cardiac hypertrophy is underscored by the studies which show that rapamycin, an inhibitor for mTOR, attenuates the development of cardiac hypertrophy induced by aortic constriction. xref Our data show that the mTOR pathway is upregulated in the hypertrophic heart, which is attenuated in the cathepsin K knockout mice."

reach
"The p70 S6 kinase (p70S6K), which is activated by mTOR and phosphorylates RPS6, is also inhibited in SU-DHL-1 cells treated with U0126 [XREF_BIBR], but surprisingly not in the SR-786 ALK+ ALCL cell line [XREF_BIBR]."

reach
"Upon activation by mTORC1, S6K1 phosphorylates S6 ribosomal protein to stimulate mRNA translation."

sparser
"Interestingly, BDNF-induced mTORC1 activation (as measured by S6 phosphorylation at specific p70 S6K phosphorylation sites) occurred normally in Lamtor1 -deficient neurons, despite the reduction of LAMTOR1 protein levels by >95% ( xref )."

reach
"One of these enzymes was MAP k inase- a ctivated p rotein k inase- 1 (MAPKAP-K1, also known as p90 rsk), a component of the classical MAP kinase cascade, while the other was p70 S6 kinase, the enzyme [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The activated S6K1 can phosphorylate downstream molecule ribosomal protein S6 (RpS6) to promote protein translation."

sparser
"mTORC1 activates S6K1, which in turn phosphorylates the S6 protein of the 40S ribosomal subunit to promote biosynthetic pathways and protein synthesis important for cell growth ( Ruvinsky and Meyuhas,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Akt activation can also induce stimulation of mTOR that will lead to phosphorylation of the 40S ribosomal S6 protein by the p70S6 kinase, thereby enabling efficient translation of 5′ terminal oligopyrimidine tract (5′TOP) mRNA [ xref ]."

sparser
"When active, mTORC1 promotes cell growth by stimulating the protein synthesis through the phosphorylation of the eIF4E-binding protein 1 (4E-BP1) and of p70S6 kinase 1 that in turn phosphorylates the ribosomal protein S6 (S6)."

reach
"Active mTORC1 also phosphorylates and activates p70S6K which in turn phosphorylates ribosomal protein S6 to enhance translation efficiency 6."

sparser
"S6K1 phosphorylates the ribosomal protein S6 (RPS6) thereby connecting the TOR pathway with ribosomes."

reach
"RAD001 specifically blocks S6K1 activation and subsequent phosphorylation of Rps6."

reach
"4,31 In addition, inhibition of mTORC1, an upstream activator of S6K1/S6K2, inhibited the phosphorylation of rpS6 in vivo ."

reach
"In response to numerous intracellular and extracellular stimuli, mTORC1 phosphorylates eukaryotic initiation factor 4E binding protein-1 (4E-BP1) and S6 kinase 1 (S6K1), which exerts an essential role in increasing translation of a subset of mRNAs and accelerating growth and proliferation XREF_BIBR XREF_BIBR."

reach
"P70S6K, activated by mTOR, phosphorylates S6 ribosomal protein to regulate translation of 5 '-TOP mRNA and ribosome biogenesis [XREF_BIBR]."

reach
"Although the dysregulation of the upstream regulatory mechanisms differs between the E198K and E420K variant cell lines, inhibition of mTORC1 with rapamycin or of p70S6K with LY2584702 reduces RPS6 phosphorylation in both variant cell lines."

reach
"Phospho-S6K1 phosphorylates S6, a key component of ribosome, whose phosphorylation is critical for the reconstitution of ribosome and for protein synthesis."

sparser
"mTORC1 phosphorylates the p70 ribosomal S6 Kinase, which in turn phosphorylates the ribosomal protein S6, as well as the initiation factor 4E binding protein ( xref )."

reach
"Consequently, p70S6K phosphorylates ribosomal protein S6, resulting in increased translation of mRNA messages containing 5 ' terminal oligopolypyrimidine (5 ' TOP) tracts."

sparser
"Since p70S6K can also phosphorylate rpS6, the actual effect of the RSK inhibitors on rpS6 phosphorylation could be masked."

sparser
"These data suggest that SIRT1 deacetylates S6K1 in ISCs during CR to facilitate Thr389 phosphorylation by mTORC1 and S6 phosphorylation by S6K1 resulting in ISC expansion."

reach
"Furthermore, inhibition of p70S6K with LY2584702 reduced RPS6 phosphorylation at S235/236 and S240/244 in both the E198K and E420K variant cell lines (Fig. 7, D–F)."

sparser
"Upon activation, p70S6K phosphorylates ribosomal protein S6 (RPS6) and other downstream targets involved in protein synthesis and cell growth."

sparser
"The authors hypothesized that the mechanism behind the requirement of mTOR activation for HIF-1α translation is likely through the phosphorylation of the 40S ribosomal protein S6 by the mTOR target S6[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The activated S6K1 can phosphorylate downstream molecule ribosomal protein S6 (RpS6) to promote protein translation."

sparser
"LN-18 cells were transfected and serum-starved before pretreatment with rapamycin to prevent p70S6K-dependent phosphorylation of rpS6."

reach
"The S6 ribosomal protein is phosphorylated by p70 S6 kinase (p70S6k)."

sparser
"mTORC1 phosphorylates proteins involved in protein synthesis and cell growth, including 4E-BP1 and p70S6 kinase 1 (S6K1), which phosphorylates the ribosomal protein S6."

sparser
"Likewise, activated S6K1 phosphorylates ribosomal protein S6 that also participates in translation initiation ( xref )."

sparser
"P70S6, a serine/threonine kinase, is a downstream signal of the PI3K/Akt/mammalian target of rapamycin and mitogen-activated protein kinase cascades. xref On activation, p70S6 subsequently phosphorylates ribosomal protein S6, which, in turn, promotes protein translation and cell growth. xref In addition, we also found that boehmenan remarkably inhibited the expression of p-Akt, p-70S6, and p-S6, which is critical for cell survival and resistant to apoptosis. xref Thus, suppression of STAT3 and PI3K/Akt pathway contributed to cell apoptosis and inhibition of cell survival triggered by boehmenan."

reach
"A number of studies in which either or both S6K genes have been deleted in mice suggest that the control of cell growth is a non redundant function specific to S6K1, while phosphorylation of the canon[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Cell proliferation requires the co-ordinate triggering of several protein kinases of Ser/Thr specificity such as p70 S6 kinase (S6K), which phosphorylates the ribosomal S6 protein and thus increases translation of mRNAs with polypyrimidine tracts."

reach
"Once activated, S6K1 phosphorylates rpS6 and eIF4B (reviewed in Fingar and Blenis, 2004)."

sparser
"Rapamycin effectively inhibited rpS6 phosphorylation by WT S6K1 or WT S6K2 but did not affect rpS6 phosphorylation in cells expressing the activated, rapamycin-resistant forms of S6K1 or S6K2."

sparser
"S6K1 and S6K2 phosphorylate and activate the 40S ribosomal protein S6, which promotes protein synthesis through an increased rate of mRNA transcription. xref S6K1 also regulates cell size and progression through the cell cycle, xref − xref in addition to promoting cell survival by inactivating the proapoptotic protein BAD. xref "

sparser
"S6K1 (p70S6 kinase) phosphorylates S6 which controls the translation of mRNA that possesses an unusual oligopyrimidine tract at its transcription starting site termed 5'TOP."

sparser
"Activation of mTOR results in the phosphorylation and activation of S6K1, which in turn phosphorylates ribosomal protein S6, and finally stimulates translation initiation and protein synthesis."

sparser
"In mammals, the TORC1 controlled S6K1 kinase phosphorylates the ribosomal protein S6 thereby co-ordinating cell size and nutritional status."

reach
"In the present study, we have examined whether Ang II activates p70 S6 kinase (p70S6K), which has been reported to phosphorylate S6 protein."

sparser
"S6K1 phosphorylates S6, which leads to enhanced translation initiation of the 5′ terminal oligopyrimidine tract-containing mRNAs encoding components of the protein synthesis machinery (ribosomal prote[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"32 MTOR phosphorylates S6K1 at Thr389, and activated S6K1 further phosphorylates RPS6 to regulate protein translation, synthesis, and metabolism."

sparser
"Phosphorylation by mTORC1 at Thr389 activates p70S6K, which in turn phosphorylates ribosomal protein S6, and may influence translation of TOP RNAs ( xref ; xref ; xref ; xref ) ."

sparser
"mTORC1 activation leads to phosphorylation of p70 S6-kinase, which phosphorylates its downstream protein S6 [ xref , xref ]."

sparser
"Activated S6K1 phosphorylates S6 (40S ribosomal protein S6), enhancing the translation of mRNA ( Jastrzebski et al., 2007 )."

sparser
"Its major effects are mediated by phosphorylation of proteins involved in the regulation of protein synthesis such as 4EBP1 and p70S6K, with subsequent phosphorylation of ribosomal S6 protein by p70S6K, and the components of ULK1 kinase that play a critical role in the regulation of autophagy [ xref ]."

reach
"However, S6 phosphorylation by S6K1 is only minimally influenced by removal of the CTD, or by phospho defective or -mimetic mutations."

reach
"Phosphorylation by mTORC1 at Thr389 activates p70S6K, which in turn phosphorylates ribosomal protein S6, and may influence translation of TOP RNAs."

sparser
"We conclude that Tyr-phosphorylated insulin receptor substrate-1 is the main upstream regulator of insulin-induced S6 phosphorylation by p70s6k, whereas MAPK signaling seems to be activated in these cells primarily through the adaptor molecule Shc."

reach
"mTOR activates S6K1, which phosphorylates eIF4B and S6, so promoting translation initiation, and this may be enhanced by ASF and SF2 XREF_BIBR."

reach
"mTORC1 consists of mTOR, mLST8, Raptor, Deptor, and PRAS40 and phosphorylates the elF4E binding protein (4F-BP) and p70 S6 kinase 1 (S6K1), a serine/threonine kinase that phosphorylates the ribosomal protein S6."

sparser
"While S6K1 phosphorylates rpS6 on all phosphorylation sites (S235/236 and S240/244), RSK phosphorylates rpS6 exclusively on S235/236 in an mTOR-independent mechanism, suggesting that ERK/RSK pathway contributes to rpS6 phosphorylation upon mitogen stimulation [ xref ]."

reach
"mTORC1 consists of mTOR, mLST8, Raptor, Deptor, and PRAS40 and phosphorylates the elF4E binding protein (4F-BP) and p70 S6 kinase 1 (S6K1), a serine/threonine kinase that phosphorylates the ribosomal protein S6."

sparser
"To increase its own protein synthesis, HSV-1 augments phosphorylation of p70 S6 kinase, which in turn phosphorylates S6 protein ( xref )."

reach
"In order to evaluate the possible targeting of the mTOR pathway by both MTH (through raptor) and RAPA (through mTOR binding by FKBP12), we compared the effects of these two inducers on the phosphorylation of p70S6Kinase (p70S6K), which specifically phosphorylates the ribosomal protein S6, strongly involved in sustaining protein synthesis and cell growth [XREF_BIBR, XREF_BIBR]."

sparser
"Also, downstream of mTOR and PI3K are S6K1 and S6K2, which phosphorylate the ribosomal protein S6 [3] ."

reach
"P70S6K phosphorylates ribosomal protein S6 and regulates protein translation [11]."

reach
"P70S6K subsequently phosphorylates the ribosomal S6 subunit (S6) [20, 23-26]."

sparser
"We also studied the effects of RSK on the phosphorylation of rpS6 (S235/236) without the inhibition of p70S6K, which also phosphorylates the same residues of rpS6."

reach
"β Cells from S6K1–/– embryos exhibit reduced cell size and insulin content, however, re-expression of S6K1 in β cells of S6K1–/– mice restored embryonic β cell size, insulin levels, glucose tolerance, and RPS6 phosphorylation (183)."

sparser
"As mentioned above, S6K1 phosphorylates S6 at multiple sites to drive protein translation; therefore, it is intriguing to ask whether those phosphorylation events play roles in T cells."

sparser
"While S6 is predominantly phosphorylated by p70S6K in most mammalian adult tissues, phosphorylation of S6 by p90rsk is important during meiosis in mammalian and amphibian oocytes [ xref , xref ], and to a lesser extent in growth factor stimulated hepatocytes [ xref ]."

reach
"Activated mTOR phosphorylates p70 S6 kinase, followed by activated p70 S6 kinase that phosphorylates ribosomal protein S6, and as a result, increases the protein synthesis complex."

sparser
"In response to mitogenic stimuli, p70S6K phosphorylates S6 on multiple sites, and it is believed that these modifications favor the recruitment of the 40S subunit into actively translating polysomes [[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"S6K1 in turn phosphorylates the ribosomal protein S6."

reach
"Since a key downstream target of PDK1 is p70S6K, which phosphorylates S6 [XREF_BIBR, XREF_BIBR], we hypothesized that if UCN-01 inhibited PDK1, we would observe reduced pS6."

sparser
"mTOR phosphorylates p70S6K, which in turn phosphorylates the S6 protein in the 40S small ribosomal subunit."

sparser
"In response to various anabolic stimuli, mTOR phosphorylates p70S6 kinase (p70S6K) which in turn phosphorylates the ribosomal protein S6 (rpS6) to positively affect muscle growth xref ."

sparser
"P70S6K phosphorylates and activates ribosomal protein S6 (S6), a component of the 40S ribosomal subunit , and phosphorylates the eukaryotic initiation factor 4B (eIF4B), a regulator of protein synthes[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, siRNA mediated knockdown of S6K1 reduced phosphorylation of its substrate RPS6 without altering phosphorylation of 4E-BP1, and suppressed both basal rDNA transcription and 47/45S rRNA synthesis in the presence of amino acids (Lane 2 vs. Lane 4)."

sparser
"Phosphorylation (denoted by P in circle) activates S6K1 which in turn phosphorylates S6, a ribosomal protein."

sparser
"The latter activates pp70 S6-kinase, which phosphorylates the S6 subunit of the ribosome to enhance translation of selected mRNAs by promoting their interaction with the 40S subunit [42–44] ."

reach
"Together, these results suggest either that the FL772 inhibitor has poor cell membrane permeability or that inhibition of S6K1 in cells does not significantly reduce S6 phosphorylation."

reach
"To inhibit p70S6 kinase, which phosphorylates the ribosomal RPS6 protein, we used LY2584702 (Selleckchem, #S7698)."

sparser
"This is done through activation of ribosomal S6 kinase 1 (S6K1) and inhibition of eukaryotic initiation factor 4E binding protein 1 (4E-BP1) by phosphorylation.[16] The resulting activated S6K1 can phosphorylate ribosomal protein S6.[17] Generally, phosphorylation of S6 is a good indicator of S6K1 activity."

sparser
"S6K1 also phosphorylates ribosomal protein S6 (rpS6; Ser235/236 and Ser240/244) [ xref , xref , xref ], a protein within the small (40S) ribosomal subunit."

sparser
"P70S6K phosphorylates ribosomal protein S6 (S6RP), whose hyperphosphorylation status correlates with translation activity ( van Gorp et al., 2009 )."

sparser
"The phosphorylation of ribosomal S6 protein by S6K1 regulates the initiation of 5′ terminal oligopyrimidine tract mRNA translation protein (5′ TOP)."

reach
"256 Mitogenic stimulation causes S6 C-terminal phosphorylation by p70 S6 kinases (S6Ks) and p90 ribosomal S6 kinases (RSKs) on Ser 235, Ser 236, Ser 240, and Ser 244."

sparser
"S6K1 phosphorylates the S6 ribosome, thus stimulating protein synthesis."

sparser
"S6K1 phosphorylates ribosomal protein S6 (rpS6) leading to increased translation of ribosomal and transcription factor mRNA resulting in protein translation initiation."

sparser
"On the other hand, activated p70S6K1 phosphorylates the ribosomal protein S6 and eukaryotic elongation factor 2 kinase (eEF2K) leading to increased translational efficiency."

sparser
"One clue, involving translational control, is the activation of p70 S6 kinase, which phosphorylates the ribosomal S6 protein ()."

sparser
"mTORC1-activated p70S6K phosphorylates the 40S-ribosomal protein S6 ( xref )."

sparser
"A significant decrease in phosphorylation of 40S ribosomal subunit S6 by p70S6K has been observed in insular, cingulated, and prefrontal cortex of rat brain ( xref )."

reach
"Once activated, S6K1 phosphorylates S6."

sparser
"mTOR-phosphorylated S6K1 in turn phosphorylates the ribosomal protein S6, the initiation factor eIF4B, and the translation elongation factor eEF2 kinase, inhibiting protein translation [30,31] ."

reach
"One clue, involving translational control, is the activation of p70 S6 kinase, which phosphorylates the ribosomal S6 protein (Shioi et al., 2002)."

sparser
"S6K1 and S6K2 both phosphorylate the 40S ribosomal subunit protein S6 [2,3] , a process that was thought to increase translation of mRNAs with a 5′-terminus oligopyrimidine tract (5′TOP mRNA)."

reach
"Recent conflicting studies, however, showed that over-expression of a dominant-negative S6K1 variant, which was sufficient to repress rpS6 phosphorylation, had no effect on the translation of 5′TOP mR[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"However, recent studies showed that cells from S6K1 and S6K2 double knockout mice have impaired S6 phosphorylation but maintain mTOR-dependent 5′TOP mRNA translation, putting into question the functio[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Activated mTOR induces phosphorylation of p70S6K followed by p70S6K-induced phosphorylation of S6 and the eukaryotic translation initiation factor 4B (eIF-4B), which promotes protein translation ( Wan[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"P70S6K phosphorylates and activates ribosomal protein S6 (RPS6) ( xref )."

sparser
"Additionally, AKT via mTOR pathway indirectly regulates cell growth and proliferation and protein synthesis by phosphorylating p70S6K which subsequently phosphorylates the ribosomal protein S6 leading to enhanced translation [ xref , xref ]."

reach
"Among the best characterized mTOR substrates are S6K1, which phosphorylates the ribosomal protein S6 and regulates the translation process [XREF_BIBR], and eIF4E-BP1, which is involved in the formation of an active translation initiation complex [XREF_BIBR, XREF_BIBR, XREF_BIBR]."

reach
"Cells were pretreated or not with an inhibitor of p70S6K (PF-4708671), which caused RPS6 de-phosphorylation (Figure 4)."

reach
"Additionally, the mGluR1/5 stimulated ERK1/2 activation may impinge on S6K1 (p70-S6 kinase 1), which phosphorylates ribosomal protein S6 and contributes to the enhancement of 5 ' TOP (terminal 5 ' oligopyrimidine tract) mRNA translation."

sparser
"Evidence points to some common functions shared by the two; activities of both are regulated by the same upstream activating pathways such as mTOR, PI3K, and MEK pathways, and both S6K1 and S6K2 phosp[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Additionally, the mGluR1/5-stimulated ERK1/2 activation may impinge on S6K1 (p70-S6 kinase 1), which phosphorylates ribosomal protein S6 and contributes to the enhancement of 5′TOP (terminal 5′ oligopyrimidine tract) mRNA translation ( xref )."

reach
"Inhibited p70S6K, in turn, fails to phosphorylate rpS6 at Ser235/236, compromising its involvement in ribosome biogenesis (43, 44)."

reach
"S6 phosphorylation in mammalian cells is catalyzed by the protein kinase RPS6KB1, usually called p70 S6 [XREF_BIBR, XREF_BIBR], which in turn is activated by sequential phosphorylation by mTOR complex1 (most importantly at S6K1 (Thr389) and PDPK1 [XREF_BIBR, XREF_BIBR]."

sparser
"Activated mTORC1 phosphorylates downstream effector p70S6K1, which subsequently phosphorylates 40s ribosomal protein S6 (pS6), promoting protein translation and cell growth."

sparser
"Later work showed that the p70S6 kinase, which phosphorylates the ribosomal protein S6, and an ERK were also activated by PTTH ( Song and Gilbert, 1994; Rybczynski et al., 2001 )."

reach
"Additionally, p70S6K can further phosphorylate the ribosomal protein S6 (RPS6) (S236, +81% CFC), and reduction of the levels of p70S6K has been shown to improve the AD-like phenotype of 3xTg-AD mice [62]."

reach
"The 70-kD ribosomal protein S6 kinase (p70S6K), which directly phosphorylates ribosomal protein S6, is a key effector of mTOR."

sparser
"P70S6K kinase directly phosphorylates the 40S ribosomal protein S6, which results in enhanced translation of proteins that contain a polypyrimidine tract in the 5’-untranslated region [ xref ]."

reach
"On activation, PI3-K phosphorylates inositol lipids that act as second messengers to activate different downstream effector kinases such as the PKB and AKT kinase, which is involved in cell survival, [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"After engagement of TCR and CD28, p70S6K becomes activated and phosphorylates S6, which leads to increased ribosome biogenesis."

reach
"On activation, PI3-K phosphorylates inositol lipids that act as second messengers to activate different downstream effector kinases such as the PKB and AKT kinase, which is involved in cell survival, [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Activation of mTOR results in phosphorylation and activation of the 70-kDa ribosomal protein S6 kinase 1 (S6K1) which, in turn, phosphorylates ribosomal protein S6 (S6)."

sparser
"P70S6K kinase directly phosphorylates the 40S ribosomal protein S6, which results in enhanced synthesis of proteins that contain a polypyrimidine tract in the 50-untranslated region ( xref )."

sparser
"However, phosphorylation of Akt at residue Ser473 by mTORC2 and phosphorylation of S6 by p70S6K, which is downstream of Akt, in residue Ser240 were reduced at all time points in the absence of RRAS2 compared with the WT."

sparser
"Although animal data presented by Nakajima et al. ( xref ) also observed no changes in metabolic stress after repeated bouts of REST + BFR versus REST (quantified as AMPK phosphorylation), they did observe greater p70S6K and ribosomal S6 phosphorylation in skeletal muscle tissue."

reach
"In mammals, upon stimulation by growth factors including insulin, the mammalian TOR (mTOR) cooperates with PI3K dependent effectors to activate p70 ribosomal protein S6 kinase 1 (S6K1), thereby phosphorylating the 40S-ribosomal protein S6, and subsequently enhances translation of the 5 '-terminal oligopyrimidine (5 '-TOP) sequences that encode components of the translational machinery."

sparser
"Activated mTOR and ERK phosphorylate p70 S6 kinase (p70S6k), which, in turn, phosphorylates the ribosomal protein S6 (RP-S6) and the eukaryotic translation initiation factor 4E (eIF4E) binding protein[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In turn, S6K1 phosphorylates and activates S6, a ribosomal protein involved in protein translation."

reach
"Interestingly, control of mRNA translation occurs via phosphorylation of another component of the 40S ribosomal subunit, RPS6, by the previously mentioned kinase RPS6KB1."

sparser
"The mTOR directly phosphorylates 4E-BP1 and p70S6K which then phosphorylate S6 to initiate protein translation step."

sparser
"Activated S6K1 then phosphorylates eIF4B and the 40S ribosomal subunit protein S6."

reach
"This indicates that p70S6K1 activated by Abeta phosphorylates the ribosomal S6 protein and 4E-BP1, which in turn prevents the interaction between 4E-BP1 and eIF-4E ultimately to enhance the cap dependent mRNA translation of HIF1alpha and HIF1alpha synthesis."

reach
"The p70 and p85 S6 kinase 1 (S6K1), which is able to catalyze the phosphorylation of rpS6 at all sites, was the first kinase identified."

sparser
"S6K1 is activated by phosphorylation and, in turn, phosphorylates the 40S ribosomal S6 protein."

reach
"The p70 and p85 S6 kinase 1 (S6K1), which is able to catalyze the phosphorylation of rpS6 at all sites, was the first kinase identified."

reach
"Although both RSK2 and S6K1 phosphorylate S6 in vivo, these kinases do not appear to be functionally redundant for a few reasons."

reach
"In parallel, FSH induced p70S6K activity triggers a robust phosphorylation of rpS6."

sparser
"The p70S6 kinase phosphorylates the ribosomal protein S6, resulting in the phosphorylation of transcription elongation factors, which facilitates translation and cell growth [ xref , xref , xref , xref ]."

sparser
"P70S6 kinase phosphorylates ribosomal protein S6, and possibly the initiation factors eIF4B and eIF4G."

reach
"P70 S6 kinases phosphorylate the 40S ribosomal subunit protein S6 and activate the translation of mRNA containing 5 ' ligopyrimidine tracts."

reach
"P70S6K is activated by mTORC1 and phosphorylates S6."

reach
"Activated mTORC1 increases protein synthesis by directly phosphorylating the ribosomal S6 kinase 1 (S6K1), which in turn directly phosphorylates ribosomal protein S6, resulting in increased ribosomal protein translation."

sparser
"Finally, S6Ks phosphorylate the ribosomal protein RPS6, the role of which is poorly described."

sparser
"Activated mTOR and ERK phosphorylate p70S6 kinase (p70S6K), which, in turn, phosphorylates the ribosomal protein S6 (RP-S6) and the eukaryotic translation initiation factor 4E (eIF4E) binding protein [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We also measured phosphorylation on ribosomal protein S6 (S6RP), which is phosphorylated by p70S6K in a mTORC1 dependent manner and thus indicates mTORC1 activity."

reach
"P70S6K phosphorylates RPS6 and leads to an increase in the rate of translation of 5′TOP mRNA transcripts which encode components of the cellular translational apparatus, thus facilitating an increase [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In this murine HF model, the downstream mTOR signaling proteins S6K1 and S6 were both increased in TAC-HF, although the abundance of Thr389 phosphorylated S6K1 and Ser235/236 phosphorylated S6 were not increased in HF ( xref )."

reach
"A preponderance of evidence suggests that translation of mRNAs containing a 5 '-TOP signal is strongly up-regulated in response to ribosome protein S6 phosphorylation triggered by mTORC1 stimulation and subsequent S6K1 activation."

sparser
"Subsequently, S6K1 phosphorylates the 40S ribosomal subunit protein rpS6, the eukaryotic translation factor eIF4B, as well as the eIF4A inhibitor PDCD4 (leading to PDCD4 degradation)."

reach
"Another well established target of the TORC1 complex is the S6 protein kinase S6K1, which phosphorylates ribosomal protein S6, an event that controls elongation factor 2 kinase, but probably not translation of mRNAs with a 5 '-oligopyrimidine tract as originally suggested [XREF_BIBR - XREF_BIBR]."

sparser
"mTOR also activates S6K-1, which phosphorylates ribosomal protein S6, resulting in increased translation."

sparser
"Activated S6K1 phosphorylates S6 and other translational regulators such as eIF2 kinase and eIF-4B to regulate the translation initiation [ xref , xref ] and ribosomal biogenesis."

reach
"LN-18 cells were transfected and serum starved before pretreatment with rapamycin to prevent p70S6K dependent phosphorylation of rpS6."

sparser
"As indexed to actin and compared to SHAM mice, abundance of total S6K1 and S6 but not Thr389 phosphorylated S6K1 or Ser235/236 phosphorylated S6 were increased in HF ( xref )."

reach
"Active p70S6K phosphorylates the ribosomal protein S6, enhancing translation of mRNA transcripts involved in cell cycle progression."

sparser
"Potential downstream targets of PI3-kinase activity include: p70 kinase phosphorylates the 40S ribosomal protein S6 in response to mitogenic stimuli and plays a key role in G1 to S phase transition th[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Since p70S6K can also phosphorylate rpS6, the actual effect of the RSK inhibitors on rpS6 phosphorylation could be masked."

reach
"XREF_BIBR, XREF_BIBR, XREF_BIBR S6K1 phosphorylates S6, which regulates the translation of 5 ' terminal oligopyrimidine mRNAs that encode ribosomal proteins and translation factors."

sparser
"6 In response to activation by a complex of mTOR with two proteins, Raptor and GβL (this complex is commonly referred to as TORC1), p70S6K phosphorylates the S6 protein of the 40S ribosomal subunit."

reach
"mTOR activation has been linked to cardiac hypertrophy by phosphorylating downstream targets including eukaryotic initiation factor 4E binding protein 1 (4E-BP1) and ribosomal S6 kinase (p70S6K), which leads to increased phosphorylation of ribosomal protein S6 (S6)."

sparser
"mTOR, structurally related to PI3K, is involved in the regulation of p70S6K, which in turn phosphorylates the S6 protein of the 40S ribosomal subunit."

reach
"mTOR activation has been linked to cardiac hypertrophy by phosphorylating downstream targets including eukaryotic initiation factor 4E binding protein 1 (4E-BP1) and ribosomal S6 kinase (p70S6K), which leads to increased phosphorylation of ribosomal protein S6 (S6)."

reach
"For example, p70 S6 kinase 1 (p70S6K) phosphorylates the S6 protein, a component of the 40S ribosomal subunit, and is an intermediary between mTOR and pS6."

reach
"For example, p70 S6 kinase 1 (p70S6K) phosphorylates the S6 protein, a component of the 40S ribosomal subunit, and is an intermediary between mTOR and pS6."

reach
"At a downstream level, mTORC1 activates p70 ribosomal protein S6 kinase1 (p70S6K), which further phosphorylates ribosomal protein S6 (RPS6) and carbamoyl phosphate synthetase 2-aspartate transcarbamoylase-dihydroorotase (CAD), responsible for inducing ribosome biogenesis and de novo pyrimidine biosynthesis, respectively [58]."

sparser
"Although the signaling pathways controlling translation of TOP mRNAs are still under intensive investigations, it seems that RPS6 phosphorylation by S6Ks, mTOR activity, and the PI3-kinase pathway activation in response to growth or mitotic signals is essential for the translational regulation of TOP mRNAs (for review [ xref , xref ])."

reach
"Among many other functions, mTORC1 is best known to control protein synthesis through phosphorylation of the translational regulators 4E-BP1 and S6K1, the latter of which in turn phosphorylates the ribosomal protein S6 and other substrates."

reach
"Active mTORC1- consisting besides the mTOR kinase itself, of Raptor and PRAS40 as well as other regulatory proteins [XREF_BIBR] -- phosphorylates proteins such as 4E-BP1 and S6 kinase 1 (S6K1), which in turn phosphorylates the ribosomal protein S6."

reach
"In response to mitogenic stimuli, p70S6K phosphorylates S6 on multiple sites, and it is believed that these modifications favor the recruitment of the 40S subunit into actively translating polysomes [[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Wang C, Mao X, Wang L, et al. Adiponectin sensitizes insulin signaling by reducing p70 S6 kinase-mediated serine phosphorylation of IRS-1."

reach
"S6K1 phosphorylates eIF-4B and S6 ribosomal protein (S6RP), initiating translation."

sparser
"Thus, it was proposed that the hyper-phosphorylation of rpS6 by S6Ks is implicated in the regulation of cell growth."

reach
"Thus, S6 phosphorylation does not occur through ERK regulated p90S6K but rather is mediated by p70S6K."

reach
"In addition, it phosphorylates S6K1, which in turn phosphorylates small ribosomal subunit protein RPS6."

reach
"Specifically, we found that our running protocol led to phosphorylation of S6 ribosomal protein, most likely by p70 S6 kinase."

sparser
"The C1 prototype is responsible for activating p70S6K, which in turn phosphorylates the ribosomal protein S6 and 4E-BP1 — which is inhibited by rapamycin (Corradetti et al., 2006;)."

sparser
"One such substrate is p70S6 kinase, which phosphorylates ribosomal protein S6 as well as several other proteins xref ."

sparser
"Within the cell, activation of mTORC1 leads to phosphorylation of 4E-BP1, which activates mRNA translation, and p70 S6 kinase, which in turn phosphorylates ribosomal protein S6, part of the 40S ribosomal subunit ( xref ; xref )."

reach
"Activation of mTOR by leucine leads to the phosphorylation of ribosomal protein p70 S6 kinase (p70S6K) which, in turn, phosphorylates the ribosomal protein S6 (rpS6) and up-regulates a subclass of mRN[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Activated mTORC1 phosphorylates S6Ks, which in turn phosphorylate the ribosomal protein S6 and other substrates involved in protein and lipid biosynthesis ( xref )."

reach
"mTORC1 phosphorylates the p70 ribosomal S6 Kinase, which in turn phosphorylates the ribosomal protein S6, as well as the initiation factor 4E binding protein."

sparser
"P70S6K phosphorylates and activates S6, a ribosomal subunit involved in initiating translation of 5′ terminal oligopyrimidine tract-containing mRNA encoding components of the protein synthesis machinery."

sparser
"Prime candidate targets include: eIF4E, the cap-binding initiation factor, in which the phosphorylation site is now recognized to be serine209 [57]; ribosomal protein S6, which is phosphorylated by bo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"mTOR activates the downstream p70S6 Ser/Thr kinase that phosphorylates ribosomal protein S6, required for biosynthesis of the cell's translational apparatus, a critical component of cell growth and proliferation xref ."

sparser
"Adiponectin sensitizes insulin signaling by reducing p70 S6 kinase-mediated serine phosphorylation of IRS-1."

sparser
"More downstream, the ribosomal protein S6 is phosphorylated by p-p70S6K1."

reach
"S235/236 of rpS6 can be phosphorylated by p70S6K and/or RSK."

sparser
"S6Ks phosphorylates multiple residues on RPS6 (Ser 235 , Ser 236 , Ser 240 , Ser 244 , and Ser 247 ), all of which are proximally located at the C-terminal region of the protein [97] ."

sparser
"These include the ribosome kinase S6K1 (p70S6K1), which phosphorylates the ribosomal protein rpS6 to facilitate protein synthesis."

sparser
"S6Ks, which phosphorylates ribosomal S6 protein, are effectors of the mTOR pathway."
| PMC

reach
"These results suggest that the hyper-phosphorylation of rpS6 is not directly mediated by an increased activation of its kinases RSK and p70S6K."

reach
"These results demonstrate that, in imatinib resistant KCL22 cells, inhibition of either CK2 or p70S6K almost abolishes rpS6 phosphorylation, while the inhibition of one kinase does not influence the activity of the other."

reach
"CK2-inhibition strongly reduces rpS6 phosphorylation without affecting p70S6K, the main kinase active on this protein in R-KCL22 cells as indicated by cell treatment with rapamycin and PF-4708671."

reach
"Therefore, we might hypothesize that the phosphorylation of rpS6 by p70S6K is under the control of a CK2 mediated regulatory mechanism."

sparser
"P70S6k sequentially phosphorylates the S6 kinase that induces ribosomal proliferation."

sparser
"S6K1, a key regulator of cell growth, phosphorylates ribosomal protein S6 and enhances the translation of 5’-terminal oligopyrimidine tract mRNAs in some models ( xref )."

sparser
"S6K1 phosphorylates multiple sites of the 40 S ribosomal protein S6 resulting in a specific increase in the translation of a subset of mRNAs containing a polypyrimidine tract in their 5′-untranslated region (5′ TOP mRNAs) xref ."

sparser
"The p70-S6K1 protein is activated by mTOR and subsequently phosphorylates the ribosomal S6 protein (RPS6), eukaryotic transcription initiation factor 4B (EIF4B), and eukaryotic elongation factor 2-kinase (eEF2 k) and is a positive regulator of protein translation (reviewed in [ xref ])."

sparser
"In turn S6K1 phosphorylates and activates S6 ."

sparser
"The p70-S6K1 protein is activated by mTOR and subsequently phosphorylates the ribosomal S6 protein (RPS6), eukaryotic transcription initiation factor 4B (EIF4B), and eukaryotic elongation factor 2-kinase (eEF2 k) and is a positive regulator of protein translation (reviewed in [ xref ])."

sparser
"S6K1 phosphorylates ribosomal protein S6 and plays an important role in cell growth and proliferation [ 41 , 42 ]."

sparser
"S6 is phosphorylated by S6K1, a key regulator of the mTOR pathway, and regulates protein synthesis, cell differentiation, and immune function xref ."

reach
"Genetic knockout studies showed that both S6K1 and S6K2 can mediate rpS6 phosphorylation, suggesting that pharmacologic strategies to counteract PTEN loss in GBMs require suppression of both paralogs (15)."

reach
"Akt mediated phosphorylation of the mTOR phosphorylates the p70S6K which is known to phosphorylate the ribosomal protein S6 and consequently results in ribosome biogenesis and protein synthesis."

reach
"Single sgRNA inactivation of S6K1 or S6K2 modestly reduced rpS6 phosphorylation at Ser240/244 in both LN229 and U87MG GBMs (Fig. 4A–C)."

reach
"Based on our data pointing to critical involvement of mTOR signaling in TTL short / high-risk leukemia, we addressed mTOR pathway activity by investigating phosphorylation of key signaling molecules : ribosomal protein S6 (S6), a downstream molecule which is phosphorylated by the ribosomal protein S6 kinase (p70S6K1) upon mTOR activation; and AKT, an upstream signaling kinase that is activated by phosphatidylinositol 3-kinase (PI3K) also mediating mTOR activation."

reach
"These results show that haloperidol promotes mTORC1- and S6K1 dependent phosphorylation of rpS6 at Ser240/244, in a subpopulation of striatal MSNs expressing D2Rs."

reach
"Since LKB1 is a regulator of mTOR activity through the phosphorylation of the energy sensor AMPK, in the absence of this regulation, mTOR is active whereby it phosphorylates S6K1, that in turn phosphorylates pS6, ultimately leading to protein translation XREF_BIBR."

sparser
"While p70S6K phosphorylates ribosomal S6 protein potentially modulating its activity, 4EBP1 binds and inactivates a factor of initiation of Cap-dependent transcription eIF4E. In its hypophosphorylated forms, 4EBP1 has high avidity to eIF4E causing inhibition of translation of many mRNA with structured 5′UTR, encoding proteins critical for cell growth and metabolism."

sparser
"Two primary effectors of mTORC1 signaling are p70S6 kinase, which phosphorylates the ribosomal protein S6, and 4E-BP1 that controls formation of the translation initiation complex xref ."

sparser
"Binding of insulin to its receptor triggers activation of the phosphatidylinositol 3-kinase (PI3K)-Akt pathway and subsequent activation of mTOR; activation of mTOR results in the phosphorylation and activation of S6K1 and 4EBP1, which in turn phosphorylates ribosomal protein S6 and, finally, stimulates protein synthesis [ xref , xref ]."

reach
"It has been observed that there may be a relationship between the phosphorylation of ribosomal protein S6 (RPS6) by p70 ribosomal protein S6 kinase 1 and translational activation of TOP mRNAs XREF_BIBR."

reach
"Although single knockout or knockdown of S6K1 or S6K2 was insufficient to entirely reduce rpS6 phosphorylation, the combination of S6K1 and S6K2 inactivation cooperated to substantially reduce rpS6 phosphorylation, consistent with overlapping and compensatory functions in GBM (Fig. 4A–C)."

reach
"P70S6K phosphorylates ribosomal protein S6, in vivo, which has been implicated in stimulating the translation of mRNA transcripts that contain a polypyrimidine tract at their 5 ' transcriptional start[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Unstimulated COS cells contain active p70S6K, which phosphorylates ribosomal S6 protein as well as CREM in vitro (Figure 3A)."

sparser
"It is a serine/threonine kinase, which phosphorylates at least three downstream substrates: the translation repressor eIF4E-binding protein (4E-BP) ( xref ; xref ; xref ); p70S6K, which phosphorylates ribosome protein S6 ( xref ); and signal transducer and activator of transcription 3 (Stat3) ( xref ; xref )."

reach
"It has been observed that there may be a relationship between the phosphorylation of ribosomal protein S6 (RPS6) by p70 ribosomal protein S6 kinase 1 and translational activation of TOP mRNAs XREF_BIBR."

reach
"Moreover, activation of mTOR promotes phosphorylation of p70 ribosomal protein S6 kinase (p70S6K), which subsequently phosphorylates the ribosomal protein S6 (rpS6) and enhances HIF-1α translation [126]."

sparser
"S6K1 subsequently phosphorylates its namesake target, ribosomal protein S6, a component of the 40S subunit."

sparser
"S6K1, an evolutionarily conserved nutrient-sensing enzyme, phosphorylates the 40S ribosomal protein S6 to regulate the translation of 5′-terminal oligopyrimidine tract (5′TOP) -containing mRNA transcripts and is a downstream target of mTOR ( xref )."

reach
"To investigate mTOR pathway activity in acquired immunodeficiency syndrome (AIDS) related diffuse large B-cell lymphoma AR-DLBCL, we used immunohistochemistry to examine the presence of the phosphorylated 70 ribosomal S6 protein-kinase (p70S6K), an extensively studied effector of mTOR Complex 1 (mTORC1) and the phosphorylated phosphatase and tensin homolog (pPTEN), a negative regulator of mTORC1 pathway."

reach
"Based on our data pointing to critical involvement of mTOR signaling in TTL short / high-risk leukemia, we addressed mTOR pathway activity by investigating phosphorylation of key signaling molecules : ribosomal protein S6 (S6), a downstream molecule which is phosphorylated by the ribosomal protein S6 kinase (p70S6K1) upon mTOR activation; and AKT, an upstream signaling kinase that is activated by phosphatidylinositol 3-kinase (PI3K) also mediating mTOR activation."

reach
"The cellular effects of mTOR are largely mediated by its activation of the p70S6 kinase, which itself phosphorylates ribosomal protein S6 (S6)."

sparser
"Insulin further mediates translation initiation by increasing the activity of S6K1, which phosphorylates the ribosomal target S6 [ xref ]."

sparser
"The active form of S6Ks then phosphorylates ribosomal protein S6 and increases general protein synthesis."

reach
"In response to various stimuli, mTOR can be activated through phosphorylation, and the activated p-mTOR is involved in the regulation of protein synthesis by inducing the phosphorylation of p70S6K, which can phosphorylate the ribosomal protein S6 that is involved in protein translation."

reach
"Figure XREF_FIG shows the effects of AZD-8055 on the phosphorylation of Rps6, which is phosphorylated by the protein kinase Rps6kb1 (i.e., p70S6K), which is activated by mTOR."

reach
"mTOR promotes new protein synthesis by targeting the eukaryotic translation initiation factor 4E binding protein 1 (4EBP1) and S6 kinase 1 (S6K1), which in turn phosphorylates S6."

reach
"The S6 protein is a component of the 40S ribosomal subunit and, as phosphorylation of the S6 protein by p70S6K correlates with increased translation, this suggests one possible mechanism by which rapamycin could inhibit translation."

sparser
"Phosphorylation of S6 by S6K1 at Serine 235/Ser236 is associated with increased protein translation ( xref ; xref )."

reach
"The C1 prototype is responsible for activating p70S6K, which in turn phosphorylates the ribosomal protein S6 and 4E-BP1 -- which is inhibited by rapamycin (Corradetti et al., 2006; Gera and Lichtenste[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"S6Ks phosphorylate ribosomal protein S6 (rpS6) and additional proteins involved in the translational machinery, although the functional roles of these modifications remain elusive."

sparser
"mTOR also phosphorylates and activates p70S6K, which in turn phosphorylates the 40S ribosomal subunit S6 protein [ xref ]."

sparser
"mTORC1 kinase activation causes the phosphorylation of the kinase p70S6K, which then phosphorylates the ribosomal protein S6."

sparser
"Upon activation, S6K1 phosphorylates ribosomal protein S6 (RPS6) to stimulate ribosomal biogenesis [ xref ], eukaryotic elongation factor 2 kinase (eEF2K) to de-repress translational elongation [ xref ], as well as eIF4B to heighten translational initiation [ xref , xref ]."

reach
"Phosphorylation of ribosomal protein S6 by p70 S6 kinase, which is believed to be involved in regulating 5 '-TOP mRNA translation [XREF_BIBR], is one of the earliest events detected following mitogenic stimuli, as part of liver regeneration of following fasting and refeeding."

reach
"We demonstrate that while ribosomal S6 kinase 1 (S6K1) phosphorylates rpS6 at all sites, RSK exclusively phosphorylates rpS6 at Ser (235/236) in vitro and in vivo using an mTOR independent mechanism."

sparser
"S6 phosphorylation by S6K1 is involved in a plethora of biologic processes including cell proliferation, translation initiation, and metabolic regulation and is known to be activated downstream of the PI3K-mTOR axis ( xref – xref ); very little has been studied regarding the specific involvement of S6K1 in drug resistance."

reach
"S6 is phosphorylated by S6K1, which itself is a target of mTOR."

reach
"S6K1 phosphorylates S6, which leads to enhanced translation initiation of the 5 ' terminal oligopyrimidine tract containing mRNAs encoding components of the protein synthesis machinery (ribosomal prot[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"S6K1 phosphorylates and activates S6, a ribosomal subunit involved in initiating protein synthesis machinery."

reach
"Increased mTORC1 activity ultimately results in the phosphorylation of multiple protein substrates, including p70S6K; when activated, p70S6K phosphorylates its substrate RPS6."

reach
"Whether it is involved in the axis we have suggested remains to be confirmed.Furthermore, p70S6K, also called RPS6KB1, can phosphorylate and activate RPS6 to promote cell growth."

reach
"Well characterized effector targets of mTORC1 includes 4E-BP1 (4E binding protein), ribosomal protein S6 kinase and S6K1 (p70S6 kinase) which, in turn, phosphorylates the ribosomal protein S6 (S6/RPS6)."

sparser
"Although both RSK2 and S6K1 phosphorylate S6 in vivo , these kinases do not appear to be functionally redundant for a few reasons."

reach
"Well characterized effector targets of mTORC1 includes 4E-BP1 (4E binding protein), ribosomal protein S6 kinase and S6K1 (p70S6 kinase) which, in turn, phosphorylates the ribosomal protein S6 (S6/RPS6)."

sparser
"Conversely, TORC1 activation drives protein synthesis via phosphorylation of S6K1, which phosphorylates the ribosomal protein S6 and initiates the translation of messenger RNA."

reach
"Activation of mTOR results in the phosphorylation and activation of S6K1, which in turn phosphorylates ribosomal protein S6, and finally stimulates translation initiation and protein synthesis."