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USP4 leads to the deubiquitination of SMAD4. 2 / 2
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reach
"USP4 inhibits SMAD4 monoubiquitination and promotes activin and BMP signaling."

"We demonstrated that ubiquitin-specific protease (USP) 4 strongly induces activin/BMP signaling by removing the inhibitory monoubiquitination from SMAD4."