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EGF activates ERK. 1000 / 2951
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sparser
"In fact, ERK activation by EGF appeared even to be stimulated upon PKC inhibition ( xref I)."

reach
"Here, PDGF, FGF2, and EGF treatment of PA-SMCs resulted in rapid ERK1/2 activation, and this effect was completely blocked by suramin without changes in total ERK1/2 levels."

reach
"Since 2-APB did not block EGF activation of the ERK pathway, the elevated level of intracellular calcium is a previously undescribed independent regulator of EGF induced gene expression that works together with ERK activation to regulate immediate early gene expression."

sparser
"Interestingly, this correlated with synergy in dephosphorylation of the adaptor protein, KSR, suggesting that GH potentiates EGF-induced KSR-mediated MEK-ERK activation."

reach
"In addition, NGF stimulation of ERK is known to involve an early transient phase of activation followed by a late sustained phase of activation; in contrast, epidermal growth factor (EGF) elicits only early transient ERK activation."

sparser
"To test the hypothesis that MAPK activation could be dependent upon EGF-R transactivation, we also tested the effect of AG1478 on LTD 4 -induced ERK1/2 phosphorylation and found that 1 hour pretreatment with 250 nM of the EGF-R phosphorylation inhibitor totally prevented ERK1/2 activation by EGF and LTD 4 alone or by their combination (Fig. xref )."

sparser
"We found that antiestrogens failed to inhibit ERK activation induced by EGF (data not shown), suggesting that high concentrations of antiestrogens may fail to activate the MAPK/ERK."

sparser
"As EGF-dependent ERK activation ( xref – xref ) was suggested as a relevant pathway in CRC, the related signaling pathways on E-cadherin, EGFR, ERK, or β-catenin should be further investigated."

sparser
"Although it is known that EGF-activated Erks mediate hyper-phosphorylation of Smad2/3 that can affect the nuclear translocation of Smad2/3 ( xref ), MEK1 inhibitor PD98059 does not reverse the EGF-mediated attenuation of TGF-β signaling in our system."

reach
"In PC-12 pheochromocytoma cells, epidermal growth factor (EGF) and nerve growth factor (NGF) activate ERK in a different temporal schedule."

reach
"Epidermal growth factor receptors (EGFRs) have been shown to be polarized at the leading edge of migrating epithelial cells in the presence of an electric field and asymmetrically activated downstream MAP kinase ERK1/2 XREF_BIBR."

reach
"Thus, EGF induced a rapid and transient ERK activation that resulted in cell proliferation, whereas NGF induced sustained MAP kinases activation associated with cell differentiation."

sparser
"Similarly, there is evidence that EGF activation of ERK1/2 in non-cardiac muscle is PKC-independent and Ras-dependent ( Robin et al., 2004 )."

sparser
"In contrast, c-Src is dominantly involved in EGF-induced activation of ERK."

trips
"MDA-MB-231 cells, which express ERbeta, but not ERalpha, and low levels of GPR30 protein, are unable to stimulate adenylyl cyclase or promote estrogen-mediated blockade of EGF-induced activation of Erk-1/-2."

sparser
"Similar results were obtained by evaluating ERK1/2 phosphorylation: PP1 prevents both SDF-1α- and EGF-induced ERK1/2 activation (data not shown)."

reach
"The role of endocytosis in ERK1/2 activation by EGF has been previously demonstrated using general inhibitors of CME, although other studies using the same inhibitors produced conflicting results with regard to the role of endocytosis in ERK activity."

sparser
"Concomitant with a decline in the steady-state levels of the receptor, we observed a decrease in EGF-induced activation of Erk1/2 in Tsg101-deficient cells compared to their isogenic controls that express endogenous Tsg101."

reach
"On the other hand, the EGF treatment adopted by Albeck and Brugge stimulates not only ERK, but also other signaling molecules, such as PI3K and Akt."

reach
"UVB generated hydrogen peroxide stimulates membrane epidermal growth factor receptors (EGFRs) that activate ERK proteins which contribute to cell proliferation and differentiation that may be involved in tumor promotion [XREF_BIBR]."

sparser
"Accordingly, we did not observe significant differences in EGF-induced ERK or AKT activation between pBabe, WTATG3, and KR cells ( xref )."

reach
"We also demonstrated that while overexpression of the dominant negative form of Ras, N17-Ras, could inhibit ERK activation induced by EGF as reported [27,28]; it could not inhibit its activation induc[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"However, when Ras was targeted to the plasma membrane in a methylation-independent way, AFC could still block Ras-dependent EGF-stimulated ERK (extracellularly regulated kinase) activation [ 37 • ]."

sparser
"Therefore, we investigated whether the p38 MAPK and/or PI3K/Akt pathways affect EGF-induced ERK or JNK activations in human skin fibroblasts."

sparser
"Inhibition of the P2Y1 also reduced EGF-induced AKT and ERK activation ( xref ), indicating that both P2Y1 and P2Y12 contributed to EGFR signalling."

reach
"EGF or IGF-1-induced activation of the mitogenic ERK1/2 was blocked by pretreating the cells with sorafenib."

reach
"The result of Western blot analysis demonstrated that EGF (10 ng/ml) induced a sharp and transient activation of the ERK signals, which was rapidly detected at 15 min following the addition of EGF and[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Wild type BRCA1 reportedly inhibits estradiol- as well as EGF-induced activation of ERK and cell proliferation, but not the activation of AKT ( xref )."

reach
"The ERK activation induced by the combination of EGF and TGF-β1 was abrogated by PD98059 (50 μM), confirming the efficacy of this inhibitor ( Fig. 8 A )."

sparser
"Thus, SPRY2 inhibits fibroblast growth factor signaling but potentiates the activation of RAS-ERK by epidermal growth factor (EGF)."

reach
"However, HIP1R knockdown suppressed the enhancement of ERK phosphorylation and the p-ERK/ERK ratio induced by EGF, with or without BIBX (Figures 6A,I,J)."

sparser
"Interestingly, a previous study showed that knockdown of hnRNPA2 inhibits the Ras-MEK-ERK pathway and prevents ERK1/2 activation by epidermal growth factor [ xref ]."

reach
"Immunoblot analyses of ERK phosphorylation showed that EGF and PMA activated ERK in an n -butanol independent way in Cl8 cells suggesting that for these mitogens, PLD is not important in the ERK activ[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGF, another critical agonist in liver fibrogenesis, leads to a rapid and transient activation of p38, ERK1/2 and the substrate ATF-2 (XREF_FIG, II)."

reach
"Remarkably, down-regulation of AGK drastically reduced ERK1/2 activation induced by EGF, which, as expected, was bypassed by addition of the AGK product, LPA."

reach
"Our data suggest that AGK plays an important role in EGF induced mitogenic ERK signaling."

reach
"The EGF-stimulated ERK/MAPK pathway is an important regulator of cellular proliferation and differentiation."

reach
"As a functional consequence of diminished PP1beta activity, we find that somatostatin- and substance P induced but not epidermal growth factor induced ERK activation was aberrantly enhanced and prolonged."

reach
"The prediction that ErbB2 overexpression will sustain EGF induced ERK signaling is corroborated by studies of ErbB2 overexpression in HMECs, which showed that at 10 and 30 min after EGF stimulation, ERK activation is between 1.15- and 2-fold higher for cells overexpressing ErbB2."

reach
"Although EGF induced NF-kappaB activation was defected in CARMA3 or Bcl10 knockdown cells, EGF induced ERK activation was not affected in these cells (XREF_FIG)."

reach
"In addition, Fujiwara et al. (1997) reported that 3,4-dephostatin, a protein-tyrosine phosphatase inhibitor, sustained for 1-3 h the tyrosine phosphrylation and activation of ERK1/2 induced by NGF and[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For example, Figure 2 suggests that EGF induces most notably ERK and JNK, with perhaps a little p38 activation."

reach
"The extracellular signal regulated kinase (ERK) pathway is activated by EGF stimulation [1]."

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"This result suggests that mutant p53 may promote EGF induced sustained ERK1/2 activation via the miR-27a and EGFR axis."

reach
"In support of this, in p53-273H-expressing H1299 cells, induction of miR-27a by its mimics substantially attenuated EGF initiated sustained ERK1/2 activation, which was correlated with the decreased levels of EGFR (XREF_FIG)."

reach
"In monolayer culture, expression of Gab2 at levels comparable with those detected in human breast cancer cells accelerated epidermal growth factor (EGF)-induced cell cycle progression and was associated with increased basal Stat5 tyrosine phosphorylation and enhanced and/or more sustained EGF induced Erk and Akt activation."

reach
"We note that targeted depletion of caspase-8 (> 90%) using shRNAmirs is insufficient to inhibit EGF induced Erk signaling, implying a signaling role that can be effected by relatively low levels of the protein (Supp Fig."

sparser
"These results demonstrate that HRAS is competent for prolonging EGF-induced ERK activation, and subsequently MCF-7 differentiation, only if its signals emanate from plasma membrane DM microdomains."

sparser
"Widely expressed with highest levels in bone marrow, spleen, brain, testes, and embryonic brain, Abi1 may negatively regulate cell growth and transformation by interacting with the nonreceptor tyrosine kinases ABL1 and/or ABL2, thus regulating EGF-induced Erk pathway activation and EGFR signaling."

sparser
"AG1478, an inhibitor of EGFR, also attenuated EGF-induced activation of EGFR, JNK2, ERK1/2, and Akt."

sparser
"We found that Cdc42(F28Lss) fails to induce EGFR accumulation, consequently, EGF-induced activation of ERK, Akt, and JNK signals by Cdc42(F28L) were not sustained in Cdc42(F28Lss)-expressing cells."

sparser
"In both High Na + and Low Na + buffers, EGF activated EGFR, Erk1/2 and Akt to a similar extend suggesting that activation of canonical EGFR signaling is independent of the EGF-induced sodium influx (Fig.  xref )."

reach
"We also checked whether the EGF signaling-mediated activation of ERK was required for the binding of the two proteins."

reach
"Additionally, we further validated whether the EGF signaling-mediated activation of ERK affected the conformational change of HSP70."

reach
"The EGF induced activation of Erk1/2 was also apparent in MDCK and control cells but not in MDCK and CIIA-Flag cells (XREF_FIG)."

reach
"Our findings supported these data; the ERK1/2 was activated by the EGF stimulation and the active ERK1/2 was immediately localized within the nucleus (Figs. 4 and 5 b)."

reach
"In addition, EGF treatment combined with HA activated the MAPK and ERK signaling pathway in Caov-3 cells when compared with the group stimulated with EGF alone (XREF_FIG)."

reach
"It binds to Cat1 and Git1 through the Spa2-homology domain,24 and Cool-Cat1 and Git1 complexes are localized throughout the cytosol, along the cell membrane, and especially to focal adhesions, distinct structures that physically connect a cell to its extracellular environment (focal adhesions willbe discussed in more detail later).31 Because both Cooland Cat1 and Git1 can form homodimers, it is thought thatCool-Cat1 and Git1 complexes assemble into large oligo-meric complexes in cells as a mechanism to promote signal amplification.27,32,33 The localization of Cool-Cat1 and Git1 complexes to thecell surface has been shown to be important for mediat-ing G protein coupled receptor- and growth factorreceptor induced activation of Cdc42 and Rac, as well as for the regulation of Arf GTPase activation.27,31 Activa-tion of these signaling pathways lead to cytoskeletal rear-rangements which are required for promoting cancer cellmigration and invasion, and enhancement of cell growth.27,31 The ectopic expression of a mutant form ofCat1 and Git1 that is unable to associate with Cool abolishesthese effects, highlighting the importance of this proteinprotein interaction.34,35 The MEK1, ERK1/2, Cat1, and Git1 complex- Cat1 and Git1 hasalso been shown to promote the activation of the mitogen activated protein kinase ERK1/2.36 This is an out-come of a complex that can form at focal adhesions between Cat1 and Git1, MEK1.36 (i.e., the kinase that phosphorylates ERK1/2), and ERK1/2.37 Similar to Cool, MEK1 binds to the Spa-2 homology domain ofCat1 and Git1, an interaction that is enhanced when Cat1 and Git1 is phosphorylated by the non receptor tyrosine kinase c-Src.36,37 While it is not entirely clear how Cat1 and Git1 binds to ERK1/2, this interaction is augmentedfollowing the treatment of cells with epidermal growth factor (EGF).36 Interestingly, knocking-down Cat1 and Git1expression levels in vascular smooth-muscle cells, orhuman embryonic kidney 293 cells, using siRNAs was sufficient to block EGF- and c-Src-mediated ERK activation and cell migration.36,37 On the other hand, theectopic expression of Cat1 and Git1 in these same cells was found to extend EGF induced ERK activation.36 These findings suggested for the first time that Cat1 and Git1, atleast in certain contexts, could function as a signalingscaffold that connects growth factor receptor signaling tothe ERK pathway."

sparser
"By contrast, wild-type p53 failed to promote EGF-induced sustained ERK1/2 activation ( xref )."

reach
"PMA enhanced, EGF induced ERK activation amplifies the c-Fos expression response."

sparser
"Silencing β-arrestin2 gene expression attenuated ≥10-fold EGF activation of ERK by 40%, thereby implicating β-arrestin2 in the growth factor phase."

sparser
"The presented hMEC model was interrogated to ensure that it is able to reproduce some of the known global biological phenomena as previously observed experimentally ( xref ), including EGF-induced activation of Akt and Erk, EGF-independent regulation of Erk via activated Ras, integrin-dependent stimulation of Erk, Rac, and Cdc42, G-protein Coupled Receptor activation of adenylyl cyclase, as well as ErbB receptor dimerization hierarchy."

reach
"It enhances ERK activation in response to serum, lysophosphatidic acid (LPA), and phorbol ester (PMA), but has no effect on EGF-stimulation of ERK."

sparser
"It is well-established that Ras/ERK and PI3K/Akt are two major signaling pathways downstream of EGFR. xref Indeed, EGF treatment of MDA-MB-468 cells activated ERK and Akt ( xref ), while the EGFR inhibitor AG1478 abolished the effect of EGF ( xref )."

reach
"Moreover, RNAi-mediated silencing of MORG-1 inhibited serum and LPA stimulation of ERK, but did not alter EGF stimulation of ERK ( Vomastek et al., 2004 )."

reach
"However, in medulloblastoma cells, ouabain did not transactivate EGFR as has been reported in other cell lines, and ouabain inhibited EGF induced Erk1/2 and Akt activation."

reach
"Celecoxib inhibited basal and EGF stimulated proliferation, hypoxia related HIF-1alpha recruitment and stabilization as well as hypoxia- and EGF dependent activation of ERK and PI3K."

reach
"Epidermal growth factor stimulates Rac1 activation through Src and PI3K to promote colonic epithelial cell migration XREF_BIBR and induces the ERK and Rac1 signaling pathway to promote cell migration in human hepatoma HepG2 cells XREF_BIBR."

sparser
"As noted earlier, EGF transiently activates ERKs but causes neither their nuclear accumulation nor neurite extension."

sparser
"Both cell lines exhibit weak sensitivity towards gefitinib since 0.1 μM gefitinib did not affect and 1 μM gefitinib only slightly reduced the EGF-induced activation of ERK and Akt."

reach
"Under normal conditions, EGF stimulates a rapid but transient activation of ERK as the signal is rapidly shutdown, whereas under cancerous conditions, the ERK signal can not be shutdown and is sustained."

sparser
"Inhibition of ERK activity by U0126 treatment significantly prevented EGF-induced cell invasion, suggesting that EGF-induced ERK activation is responsible for the invasion of these cells."

reach
"We found EGF stimulated the activation of PI3K/Akt/mTOR and ERK1/2, and their phosphorylation extent is much higher in HCC827 cell (EGFR mutant) than H292 cells (EGFR wide type)."

sparser
"EGF Induced ERK Activation in CHO-k1 Cells Expressing EGFP-EGFR wt The activity was observed with a FRET sensor EKAREV."

sparser
"Therefore, it is reasonable to speculate that EGF-induced ERK activation is indispensible for GEP100/Arf6 mediated breast cancer cell invasion."

sparser
"This mutation (replacement of the linker by GGSGGS) breaks the differential Erk activation by EGF and TGF-α ( xref , xref )."

reach
"Both EGF and CCK octapeptide rapidly increased the activity of the MAPKs [extracellular signal regulated kinase (ERK) 1 and ERK2], reaching a maximum within 2.5 min when 3.9- and 8.5-fold increases, respectively, were observed."

reach
"IGF-1, EGF, and estradiol induce cancer cell adhesion via IGF1R and activation of ERK1/2 [61]."

sparser
"Furthermore, siRNA-mediated down-regulation of uPAR in MDA-MB-231 cells did not alter EGF-induced ERK activation."

sparser
"Most of them induced a reduction in ERK and AKT activation by EGF, a marker of RAS activity."

sparser
"Several stimuli and pathways cooperate in the regulation of SNAIL1 and SNAIL2 in different cell contexts, including the canonical TGF-β ( Zhang, Tian, & Xing, 2016 ) or TGF-β-induced MAPK, HGF and EGF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Furthermore, GPx3 could attenuate EGF (Epidermal Growth Factor) and HGF (Hepatocyte Growth Factor) induced Erk activation through the restoration of MKP3 expression."

sparser
"The insertion of this flexible segment results in a similar extent of Erk activation by EGF and TGF-α ( xref , xref ), which is consistent with the importance of this linker in coupling the conformations of the extracellular module and the transmembrane helix."

reach
"For control, we demonstrated the dose-dependent activation of ERK by EGF in PE/CA-PJ15 cells."

sparser
"Re-expression of wild-type PTEN or inhibition of PI 3-kinase caused a recovery of the sensitivity of these cells to IGF1 and insulin, without altering the activation of ERK by EGF and PDGF ( xref )."

reach
"By contrast, EGF induced high amplitude ERK activity in both mutant and wild‐type RAS cells, without evidence of H/K/NRAS‐independent activity in the BRAF cell line."

reach
"Across all cell lines, EGF stimulation initiated a rapid ERK activity peak ~ 15 min after stimulation, followed by attenuation over 1.5–2 h to reach a steady‐state level (Fig 3B and C), with HRAS and NRAS cells exhibiting slower attenuation than any of the KRAS isoforms."

sparser
"NT and EGF activate EGFR, ERK and AKT, stimulating DNA synthesis in PC3 cells."

reach
"Although prior studies have profiled cellular responses to PDGF and EGF, which respectively drive sustained and transient Ras and Erk module dynamics, these growth factors activate multiple pathways and differ from one another in many more ways than just their dynamic responses."

reach
"EGF induced ERK activation is mediated by EGF receptor type 1 while opioid receptor activation seems to brings about stimulation via EGF receptor type."

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"It was originally reported that transfection of HEK293 or COS-7 cells with either wild-type PI3KC2beta or a kinase inactive mutant PI3KC2betaDN construct did not affect EGF induced ERK activation XREF_BIBR."

reach
"On the other hand both basal and EGF mediated ERK1/2 activation appeared to be inhibited in A-431 cells overexpressing either wild type or kinase dead PI3KC2beta D1213A-17 and D1213A-32 XREF_BIBR."

sparser
"Catechol inhibits activation of EGFR, AKT, and ERK by EGF."

sparser
"These results are substantiated using β1-integrin mutants with enhanced transmembrane associations, resulting in larger integrin adhesions and increased EGF-induced ERK activation on soft substrates."

reach
"In quiescent cells, EGF pretreatment potentiated both EGFR and ERK activation in response to radiation, whereas erlotinib abrogated this activation (XREF_FIG)."

sparser
"To investigate whether catechol antagonizes against activations of EGFR, AKT, and ERK by EGF, HCC cells were pretreated with catechol 5 and 10 μM for 2 h in prior to stimulated with 100 ng/mL of EGF for 1 h, and p-EGFR, p-AKT, and p-ERK were detected by Western blotting and densitometric analysis."

reach
"Alone, EGF activated ERK1/2, increased and sustained levels of c-jun mRNA, but had almost no effect on JNK1 activation."

reach
"Interestingly trastuzumab strongly inhibits EGF-induced Akt phosphorylation and slightly inhibits EGF-induced Erk activation, in both BT474 and SKBR3 cells."

sparser
"These experiments, comparing the sensitivity of ERK activation by IGF1, insulin, PDGF and EGF, showed that when PTEN expression was reduced by RNA interference, both cell types displayed normal activation of ERK by PDGF and EGF, but were significantly suppressed in their ability to activate ERK in response to IGF1 or insulin ( xref )."

reach
"Abeta-CM almost completely abolished EGF induced activation of ERK1/2 (XREF_FIG)."

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"Further validation for DEL-22379 as an ERK dimerization inhibitor in vivo was obtained using SDS-PAGE electrophoresis (Philipova and Whitaker, 2005) for lysates from HEK293 cells, in which ERK dimeriz[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Interestingly, S1P and EGF promote nuclear translocation of activated ERK-1/2 in MCF-7 cells lacking HER2, while in cells over-expressing HER2 these growth factors activate a pool of ERK-1/2 that is r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"EGF clearly activated Akt and Erk in adipocytes whereas it activated Erk in preadipocytes, but had only a slight effect on Akt and IκB ( xref )."

reach
"EGF also triggered strong activation of Akt and ERK1/2 signaling pathways (Figures 1A, S1B, and S1C) in both parental and transfected HCT116 and HEK293T cells, except that Akt was found to be constitutively activated in HEK293T cells."

sparser
"In contrast, EGF-induced ERK activation is independent of CNK2 expression."

sparser
"Accordingly, EGF-induced Erk activation was reduced upon plaques ablation in β5-integrin-depleted cells (Fig.  xref c, d)."

reach
"13,20–23 Perhaps ERKs are activated by over-expression of EGF and ras mutation resulting in the accumulation of protein."

reach
"ERK is activated by the binding of growth factors (EGF, FGF) to their receptors (EGFR, FGFR)."

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"Stable knockdown of Sur8 in NIH3T3 cells (shSur8-GFP) decreased epidermal growth factor (EGF)-induced activation of ERKs and Elk-1 reporter compared to control (shCon-GFP) cells (XREF_SUPPLEMENTARY and XREF_SUPPLEMENTARY)."

reach
"Epiregulin (EREG), an epidermal growth factor family member, stimulated the MEK/ERK and JNK signaling cascades, resulting in increased SCAPs proliferation."

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"In turn, the EGF- and human serum-mediated rescue of drug-treated A431 cells restores ERK1/2 activity in functional tests."

sparser
"In contrast, ERK1/2 activation by epidermal growth factor (EGF) was not affected by C1orf27 knockdown ( xref )."

reach
"Previous studies indicated that activation of ERK1/2 by EGF could enhance ER-mediated transactivation [49–53] ."

sparser
"We also determined that the increased CXCR7 expression in prostate cancer cells during ADT enhances EGF-induced EGFR and ERK1/2 activation (Fig.  xref )."

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"Knockdown of Hck or Blk reduced EGF-induced ERK1/2, p38, or AKT phosphorylation (Figure 5GA, 5HA) in HaCaT cells, as well as inhibited the phosphorylation of ERK1/2, p38, and AKT in A431 cells (Figure 5GB, 5HB)."

reach
"However, previous single cell studies in different mammalian cell lines have reported both graded [XREF_BIBR, XREF_BIBR] and " all-or-nothing " [XREF_BIBR] EGF induced ERK activation responses."

reach
"Thus, in a dose dependent manner, EGF increases the probability that a cell will have ERK turned on."

reach
"Thus, the EGF induced ERK response on the population level is complex consisting of both analog and digital elements."

sparser
"Consistent with previously published results [10,11] , we find that overexpression of GRK2 in HEK-293 cells results in a ~ 2-fold increase in EGF-induced ERK activation, as detected by Western blottin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"As showed inFigure xref , EGF activated ERK in a concentration dependent manner, and it exerted the best effect at a concentration of 100 ng for 1 h (Figures xref ; n = 4, P < 0.05)."

reach
"We also show that necdin suppresses astrocyte differentiation induced by EGF-dependent activation of the EGFR/ERK signaling pathway in CPCs."

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"For example, NGF induces sustained activation of extracellular signal regulated kinase (ERK) and promotes differentiation of PC12 cells, whereas epidermal growth factor (EGF) induces transient ERK activation and provokes cell proliferation."

reach
"Interestingly, it did not affect EGF stimulated ERK1/2 activation."

sparser
"The activation of ERK1/2 by EGF was, however, reduced to a lesser extent."

sparser
"In addition, the G i2α ribozyme resulted in a partial decrease of the EGF-induced ERK1/2 activation, in accordance with a role of G i ."

sparser
"Notably, 11a-1 efficaciously attenuates cell viability and blocks EGF-induced ERK1/2 activation in H1975 lung cancer cells; abrogates ERK1/2 and AKT activation in SKBR3 breast cancer cells and suppresses cell growth in a 3D Matrigel environment; and inhibits oncogenic KITD814V-induced constitutive cell growth."

sparser
"Taken together, these results suggest that Rho-activated ERK scaffolding by GRK2 can function to increase EGF-induced ERK activation in HEK-293 cells."

reach
"Pretreatment with the EGF receptor kinase inhibitor AG1478 had, remarkably, no significant effect on Erk1/2 activation induced by any of these adrenergic agonists (although it fully abolished EGF induced Erk1/2 activation), demonstrating absence of EGF receptor mediated transactivation."

reach
"Intriguingly, several studies have also demonstrated that mammalian Sprouty proteins can increase EGF mediated Erk and MAPK signaling in a cell type dependent manner."

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"LH-mediated production of EGF-like peptides triggers ERK activation through EGFR/Ras/Raf/MEK cascade in cumulus cells, which is essential for cumulus expansion and meiotic maturation of follicle-enclosed murine oocytes (Park et al., 2004; Reizel et al., 2010)."

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"Apparently, activation of ERK1/2 by EGF stimulation was independent of increased RAS-GTP levels."

sparser
"The results showed that EGF activated STAT3, Akt and ERK1/2 in a dose- and time-dependent manner and increased the expression of p-EGFR, p-STAT3, p-Akt and p-ERK1/2 to a peak after 45 min of incubatio[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The binding of EGF and dimerization of EGFR, a transmembrane glycoprotein, activates EGFR and the downstream PI3K-AKT and ERK signaling pathway, which can regulate cell survival, proliferation, anti-apoptosis, and metastasis (Fig. 4A)."

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"The final ERK * was enhanced by HRG or EGF."

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"PAK5 expression does not potentiate EGF stimulated ERK and MAPK activation."

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"Tyr1068 is a major autophosphorylation site that, in its phosphorylated state, recruits the adaptor protein Grb2, the main step in EGF dependent induction of the Ras-MAPkinase and ERK and Akt and PKB [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To confirm that the transfected dominant negative Ras construct was functional we verified that EGF induced ERK activation is significantly attenuated in cells transfected with dominant negative Ras ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"These results suggest that EGF activates the ERK and Akt and NF-kappaB signaling pathways in HNSCC."

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"Taken as a whole, these results indicate that PAK5 is an efficient Raf-1 kinase, but that its expression is not limiting for EGF stimulated Raf-1 and ERK and MAPK pathway activation in these cells."

sparser
"Phosphoimmunoblotting of PCMO indicated that both EGF and HB-EGF activated MEK-1/2 and ERK1/2 in a concentration-dependent fashion with the effect of EGF being more prominent [ xref ]."

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"EGF stimulation increased ERK *, but did not induce a high final ERK * (> 0.9) without HRG stimulation."

reach
"In addition, EGF activated Akt and Erk whereas TNF solely activated IkappaB in OVCAR-3 cells."

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"Positive rate of ER-alpha36 was increased in high-stage (P =.03) and high-grade (P =.224) endometrial cancer; expression of ER-alpha36 and EGFR exhibited a significant positive correlation (r = 0.334, P =.025) and they showed substantial colocalization on the plasma membrane of glandular cells; phospho-extracellular signal regulated kinase positive rate in ER-alpha36 positive group and EGFR positive group was higher than that of ER-alpha36 negative group (P =.014) and EGFR negative group (P =.016); finally, ER-alpha36 mediated epidermal growth factor stimulated extracellular signal regulated kinase activation in Hec1A cells."

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"The ERK pathway inhibitor suppressed HRG- or EGF induced ERK * [XREF_BIBR]."

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"However EGF activated Erk without IkappaB phosphorylation whereas TNF highly activated IkappaB and had a small effect on the activation of Erk."

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"However, overexpression of EGF overly activated the EGFR and ERK1/2 proteins and malignant growth of gastric epithelial cells."

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"EGF activated IkappaB, Akt and Erk whereas TNF activated only IkappaB and reduced EGF activated Akt and Erk in these cells."

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"EGF activated IkappaB and Erk whereas TNF activated only IkappaB."

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"At concentrations over 10-50 fold higher than their IC 50, these inhibitors failed to prevent EGF induced ERK1/2 activation (XREF_FIG)."

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"For example, photolabile molecular glues or polymers can be modified to photo-protect EGF/NGF and TNFα, allowing controlled activation of the ERK and NF-κB pathways, respectively."

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"MyD88 independent activation of ERK by epidermal growth factor increased p-ERK and c-Myc and restored the multiple intestinal neoplasia (Min) phenotype in Apc (min/+)/Myd88 (-/-) mice."

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"The behavior of the ERK members of the MAPK family -with respect to the duration of their activation -shows differ-ent patterns in PC12 cells treated with differentiating agents (such as NGF) or with [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The pharmacological activation of ERK by epidermal growth factor (EGF) in tanycytes can restore leptin transport and rescue the normal phenotype in obese mice xref ."

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"Identification of a dominant negative mutant of Sprouty that potentiates fibroblast growth factor- but not epidermal growth factor induced ERK activation."

sparser
"Unlike platelet-derived growth factor, EGF does not stimulate Tyr-phosphorylation of PLCγ or PtdIns(4,5)P 2 hydrolysis in cardiac myocytes, nor does inhibition of PKC reduce EGF-induced ERK1/2 activat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Though EGF induced ERK activation in the SHPS-1 mutant cells is similar to that in wild-type cells, IGF-1-induced ERK activation in the mutant cells is enhanced [62]."

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"Although epidermal growth factor stimulation of the cells induces the activation and nuclear accumulation of ERK1/2, it does not induce the prolonged nuclear retention of the activated ERK1/2, and fails to induce cell motility."

sparser
"Overexpression of Abi-1 inhibits epidermal growth factor (EGF)-induced activation of extracellular signal-regulated kinases (Erks) but does not affect EGF-induced activation of c-Jun N-terminal kinase or Akt."

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"32 confirmed the existence of a dynamic MEK independent compensatory circuit for ERK activation in T47D cells that is induced by epidermal growth factor and other ligands of ErbB family receptors."

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"The role of protein kinase C (PKC) in the EGF stimulated ERK signaling pathway was further examined by inhibition of PKC with the staurosporine analog, CGP41251, and by down-regulation of PKC via chronic treatment with PMA."

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"Epidermal growth factor (EGF) also stimulated the activity of ERK, in both WKY and SHR cells, but nor of JNK or p38, with stimulation of ERK activity by EGF occurring more rapidly in SHR cells than in those from WKY rats."

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"By contrast, EGF mediated ERK activation was unaffected."

reach
"As shown in Figure 2 (c)), heterodimerization of NF1catR1276P did not only fail to attenuate Ras activation in response to EGF but instead prolonged Ras-GTP accumulation and lowered Erk activation induced by EGF."

sparser
"Notably, in both in vivo and in vitro breast cancer models, matrix stiffening induces integrin clustering to enhance EGF-dependent ERK activation and Rho-generated force, thereby promoting tumor growth and progression [ xref , xref , xref ]."

sparser
"In the parietal cell, extracellular signal-regulated kinases are activated by epidermal growth factor, gastrin, and acetylcholine."

reach
"Pre‐stimulation with 100 ng/ml EGF led to the characteristic adaptive ERK transient."

reach
"In contrast, EGF induced ERK activation is independent of CNK2 expression."

sparser
"Interestingly, neither ELMO1 nor ELMO2 is required for either CXCL12- or EGF-induced activation of PDK1/AKT or ERK1/2 ( xref and xref ), or for EGF-induced migration of the breast cancer cells in vitro ( xref )."

reach
"We found that in this case the system can distinguish between sequences ' 10 ' and ' 11 ', because the second EGF pulse increases the ERK activity tail."

reach
"However, rapid activation of Erk by EGF did result in significant smad2 linker phosphorylation (XREF_FIG)."

reach
"FGF2 and EGF also promote metastasis by activating the ERK/MAPK signaling pathway."

reach
"These waves, although not yet fully characterized, are dependent on EGF receptor ligands because inhibition of EGF receptors completely blocks ERK activity propagation [XREF_BIBR], which in turn suppresses collective cell migration [XREF_BIBR]."

reach
"Similar to our observations of CNK2 function in PC12 cells, CNK1 did not appear to contribute to EGF mediated activation of ERK in HeLa cells."

sparser
"EGF-induced activation of ERK has been extensively studied by both experimental and theoretical approaches."

sparser
"Treatment with 10 nM GA-AuNPs or 1000 nM gallic acid attenuated the EGF-induced ERK activation ( Fig. 3 B)."

sparser
"However, in medulloblastoma cells, ouabain did not transactivate EGFR as has been reported in other cell lines, and ouabain inhibited EGF-induced Erk1/2 and Akt activation (Wolle et al., xref )."

sparser
"Treatment with U0126 (MEK1 and MEK2 inhibitor) suppressed the EGF-induced ERK activation, c-Jun activation, and MMP-9 up-regulation ( Fig. 3 D)."

sparser
"One hypothesis that explains these disparate actions of Erk in PC12 cells proposes that the time course of Erk activity determines the downstream effect of pathway stimulation: NGF produces prolonged [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Nonetheless, over-expression of CA-Akt did not abrogate the inhibitory effect of GA-AuNPs on EGF-induced ERK activation."

reach
"While the EGF initiated signaling usually results in a transient extracellular-signal-regulated kinase (ERK) activity in normal fibroblast and epithelial cells, a more sustained ERK signal of sufficient duration is required to pass the R-point [XREF_BIBR]."

reach
"In the cytosol, EGF induces transient and HRG induces sustained ERK activation."

reach
"EGF was also under these conditions able to activate ERK1/2."

reach
"The 5 h EGF exposure rendered the cells resistant to further ERK1/2 activation by EGF, as well as EGF-induced alterations in the Cx43 band pattern, all in agreement with the GJIC experiments.Pre-treat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The two tyrosine residues (Tyr 627 and Tyr 659) in the carboxyl-terminal region of Gab1 are required for SHP-2 binding to Gab1 and for EGF stimulated ERK activation, in which another study reported Gab1 and SHP -2 interaction was independent of ERK activation XREF_BIBR."

reach
"Overexpression of EGFR1 or HER2 as well as mutations that undermine the negative feedback regulation of the EGFR-ERK signaling pathway may transform the transient mode of the EGF induced ERK signaling to the sustained mode, and uncontrollably drive cells past the R-point [XREF_BIBR]."

sparser
"Expression of a constitutively active Rap1 (RapV12) inhibited the ERK activation induced by LPA and EGF in Rat-1 fibroblasts [7] ."

sparser
"For instance, it has been shown that EGF-induced ERK activation diminishes after 15 min despite the continuous presence of EGF due to rapid degradation of EGF receptors and other feedback mechanisms xref ."

sparser
"Furthermore, activation of ERK1/2 through EpEX-Fc and EGF in FaDu, Cal27, and Kyse30 cells was entirely blocked by an inhibitor of the upstream kinase MEK1 (AZD6244), whereas tyrosine kinase inhibitor (TKI) AG1478 entirely (Cal27) or partially (FaDu, Kyse30) blocked ERK1/2 activation by EpEX-Fc and EGF ( xref )."

reach
"EGF stimulation of Erk was inhibited by AFC (N-acetyl-S-farnesyl-L-cysteine), an inhibitor of methylation, but not AGC (N-acetyl-S-geranyl-L-cysteine), an inactive analog of AFC."

reach
"Transient transfection of human prenylcysteine directed carboxyl methyltransferase increased EGF stimulated Erk activation."

reach
"It was shown that although EGF and NRG can both stimulate ERK1/2, EGF acts primarily through EGFR and ERBB2 heterodimers to preferentially stimulate PKC, whereas NRG acts primarily through ERBB2 and ERBB3 heterodimers to preferentially stimulate AKT [XREF_BIBR]."

reach
"37 However, our results suggest it may also act as noise filter, filtering out small and slow changes in growth factor such that EGFR is internalized before the EGF reaches a high enough concentration in the local microenvironment to trigger ERK activation."

reach
"Of course, the precise interpretation of ERK signaling patterns may depend on the target gene.39 To our knowledge, no study has addressed how the length of EGF-induced ERK activation decodes to cellular phenotypic decisions."

reach
"EGF production by tumor-infiltrating M2 TAMs within the TME can stimulate the NF-κB, STAT3, EGFR, and extracellular signal-regulated kinase signaling axes in tumor cells, promoting their invasive traits (140, 141)."

reach
"Soluble EGF induced rapid activation of both EGFR and the downstream target ERK 1/2 with peaks in phosphorylation after 30 and 60 minutes, respectively (XREF_FIG) and a return to near-baseline levels by 4 hr."

reach
"Both low and high EGF activated ERK and p38 in preadipocytes."

reach
"EGF secreted by TAMs activates the EGFR/extracellular signal-regulated kinase 1/2 signal pathway in some types of cancer cells, which results in the promotion of EMT (144)."

reach
"Selective inhibition of the EGF receptor (EGFR) with AG1478 blocked ERK and p38 activation at both concentrations; however, selective inhibition of the ErbB2 receptor (EB2R) with AG825 or small interfering RNA (siRNA) blocked low but not high EGF activation of ERK and p38."

sparser
"Silencing of hSPRY2 decreased serum- or epidermal growth factor (EGF)-elicited activation of AKT and ERK1/2 and reduced the levels of EGF receptor."

reach
"As expected, EGF and PDBu both caused rapid and transient ERK activation, increasing ppERK to the maxima at 5-15 min (XREF_FIG C)."

reach
"Interestingly, in this study silencing of clathrin heavy chain augmented EGF stimulation of Erk, a result inconsistent with studies using dominant negative dynamin as a way of blocking endocytosis [XREF_BIBR]."

sparser
"We also demonstrate that p53-273H-mediated suppression of miR-27a expression increases EGFR levels and enhances EGF-induced sustained extracellular signal–regulated kinase 1/2 (ERK1/2) activation, thus facilitating cell proliferation and tumor growth."

reach
"In PC12 cells, transient ERK activation induced by EGF resulted in proliferation, whereas sustained activation by NGF resulted in differentiation XREF_BIBR XREF_BIBR."

sparser
"Our data suggest basal AKT phosphorylation and the degree of EGF-induced activation of AKT and ERK as molecular determinants of erlotinib efficiency in PC cells."

reach
"We found that hNatB knock-down markedly reduced ERK1/2 phosphorylation at 72 h and 96 h after siRNA transfection and that EGF induced ERK1/2 activation was severely blunted in hNatB deficient cells."

reach
"Introduction of a SAICAR-insensitive or a non phosphorylatable PKM2 mutant inhibited the EGF stimulated ERK activation and consequent cancer cell proliferation, suggesting SAICAR induced PKM2 protein kinase activity is required for EGF mediated cancer cell proliferation [XREF_BIBR]."

reach
"For example, when monitoring EGF-stimulated Erk activation, it was found that, in contrast to control fibroblasts, where EGF-stimulated Erk activity typically peaked within 5–15 min, in Cdc42(F28L)-ex[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For instance, epidermal growth factor induces a transient extracellular signal-regulated kinase (ERK) activation in MCF-7 breast cancer cells, eliciting proliferation, whereas heregulin induces sustained ERK activation causing differentiation [139]."
| PMC

trips
"EBP50 knockdown abolished its inhibition on EGF-induced ERK activation, suggesting EBP50 regulated EGFR signaling."

sparser
"Thus, it was demonstrated that the ubiquitination-dependent downregulation of the EGFR is the major mechanism for EGF-induced transient ERK activation."

sparser
"Given the important role of EGFR in mediating EGF-initiated cellular signaling cascade, we next determined whether mutant p53-273H regulates EGF-induced AKT or ERK activation."

reach
"Unexpectedly, beta-arrestin-2 siRNA diminished both EGF induced ERK activation and primary astrocyte proliferation suggesting that this adaptor protein plays a novel role in EGF signaling as well as in the opioid receptor phase of this pathway."

reach
"For example, epidermal growth factor (EGF) induces transient ERK activation in PC12 cells, resulting in cell proliferation."

sparser
"As shown in xref , EGF-induced sustained ERK1/2 activation in p53-273H-expressing H1299 cells was largely abolished by EGFR knockdown, indicating that mutant p53-273H promotes sustained EGF-induced ERK1/2 activation through EGFR."

sparser
"In concordance, knockdown of EpCAM in Kyse30 cells resulted in enhanced ERK1/2 activation by EGF in our in vitro experiments."

sparser
"This differential activation of ERK1/2 by NGF or EGF translates into distinct biological responses: NGF promotes PC12 cell differentiation, whereas EGF promotes their proliferation ( xref )."

reach
"Notably, EGF-mediated activation of ERK 1/2 was associated with a downregulation of endogenous CD protein level.The present work demonstrates, for the first time, a novel antiproliferative role of cathepsin D that may be exploited to improve neuroblastoma management and treatment."

reach
"The ERK pathway is activated mainly by growth factors (epidermal growth factor, EGF, and platelet derived growth factor, PDGF) and cytokines (IL-1beta and TNF-alpha), and its activation is related to the stimulation of tyrosine kinase receptors [XREF_BIBR, XREF_BIBR]."

sparser
"Hence, in addition to an inhibitory role of EpEX on the strong activation of ERK1/2, Snail, Zeb1, and Slug by EGF, EpEX has itself an intermediately activating effect on ERK activation in serum-starved HNSCC lines that results in proliferation."

sparser
"Zhang et al. (2014) showed that EGF induces Snail through the PI3K-Akt pathway; EGF and endothelial cell-secreted factors activate STAT3, Akt, and ERK, whereas the inhibition of PI3K/Akt signalling prevents EGF- and endothelial cell-induced Snail expression [ xref ]."
| PMC

reach
"The functionality of this interaction was demonstrated by the inhibition of the epidermal growth factor dependent activation of the Erk pathway when PMCA4b and RASSF1 were co-expressed."

sparser
"To further examine whether the promoting effect of mutant p53 on EGF-induced sustained ERK1/2 activation is due to enhanced phosphorylation of ERK1/2 or decreased dephosphorylation of phosphorylated-ERK1/2 (p-ERK1/2), p53-273H-expressing or control H1299 cells were pre-treated with EGF before the addition of MEK-1 (MAP ERK kinase) inhibitor U0126 at the different time points."

sparser
"We determined if EGF activates ERK 1/2 to stimulate secretion in cultured goblet cells."

sparser
"This result suggests that mutant p53 may promote EGF-induced sustained ERK1/2 activation via the miR-27a/EGFR axis."

reach
"Interestingly, recent progress in understanding the key downstream effectors that mediate endocannabinoid signaling has implicated mammalian target of rapamycin complex 1 (mTORC1) rather than PI3K or ERK 103 as downstream signal effector; with biochemical and transcriptional experiments suggesting that PI3K activity is largely regulated by the EGF receptor, while the EGF and FGF receptors synergistically regulate the ERK pathway 168."

reach
"EGF induced ERK activation is not affected by pretreatment with BiPS (data not shown)."

sparser
"We next investigated whether miR-27a regulates EGF-induced ERK1/2 activation."

reach
"As morphine had not been previously tested for its long-term effect on EGF activation of ERK, it was examined and found to mimic the DAMGO action (XREF_FIG)."

sparser
"Mechanistically, mutant p53-mediated miR-27a suppression increases cellular levels of EGFR and promotes EGF-induced ERK1/2 sustained activation."

reach
"Altogether, these results indicate that MKP-3 plays a role in a negative feedback pathway that curtails EGF stimulated ERK1/2 activity in DLD-1 cells."

reach
"This effect was selective since Barbadin did not block EGF stimulated ERK1/2 (XREF_FIG)."

sparser
"We therefore investigated the effect of CIIA on the EGF-induced activation of Ras and Erk1/2."

sparser
"The EGF-induced activation of Erk1/2 was also apparent in MDCK/control cells but not in MDCK/CIIA-Flag cells ( xref )."

reach
"Furthermore, reducing levels of WNK2 resulted in an enhancement of EGF activation of ERK1/2."

sparser
"RNAi-mediated depletion of CIIA potentiated the EGF-induced activation of both Ras and Erk1/2 compared with that apparent in cells expressing GFP siRNA ( xref )."

reach
"This was revealed when we found that knockdown of MKP-3 by siRNA prolonged the EGF- and TNF-alpha-stimulated elevation of phospho-ERK1/2 in DLD-1 cells."

reach
"Further research is required to determine why the regulation of EGF stimulated ERK1/2 differs so dramatically between H-ras MCF10A cells and DLD-1 cells, however."

reach
"These experiments showed that GDF15 application did not lead to cilia lengthening while MEK1/2 was inhibited, whereas activation of ERK signalling by EGF was not sufficient to rescue cilia morphology."

reach
"We found that SHIP2 enhances SHC1 interaction with the mutant receptor in an EGF-dependent manner, which leads to sustained ERK and AKT signaling, ultimately increasing cell migration and invasiveness[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The specificity of the inhibitors was confirmed by EGF stimulation: none of the inhibitors of ATP signaling – that is, apyrase, suramin, or PPADS – suppressed the EGF‐induced ERK activation, whereas trametinib and AG1478 did completely suppress the ERK activation (Fig.  xref F)."

reach
"The results of this and our previous [14] studies show that both wild-type and Golgi-targeted RIG1 efficiently downregulate the ERK/ELK1-mediated transactivation induced by EGF."

reach
"To clarify whether changes in Cx43 levels influence the activation of Akt and ERK induced by EGF, we examined U87 GBM tumorspheres with nearly absent Cx43 expression that were cultured in stem cell medium."

sparser
"EGF also activated Erk-1/2 in these cells; however the EGF-receptor inhibitor PD153035 did not affect the LTD(4)-induced activation of Erk-1/2."

reach
"However, EGF-dependent activation of ERKs was not affected by GM6001 at the same concentration."

reach
"As a positive control, we showed that ERK1/2 were strongly activated by EGF in the interphase ( Fig. 6 A–C)."

reach
"Together our results indicate that during the M phase EGF was unable to stimulate activation of ERK1/2 and the ERK substrates RSK and Elk1.It was suggested that the decoupling of EGFR activation from [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Inhibition of Cdc2 activity by roscovitine in mitotic cells restored ERK activation by EGF [40] ."

reach
"We showed that while MEK was well expressed in mitosis at a level similar to that in the interphase, MEK was not phosphorylated in response to EGF in mitotic Hela H2B–GFP cells ( Fig. 8 E & F), which [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In addition, LTD(4) did not induce phosphorylation of the EGF receptor, nor did pertussis toxin (PTX) block EGF-induced activation of Erk-1/2, thus refuting a possible crosstalk between the receptors."

sparser
"In MG-63 and Saos-2 osteosarcoma cells, it has been reported to induce apoptosis and inhibit both the constitutive as well as EGF-induced activation of Raf, MEK and ERK in a dose-dependent manner."

sparser
"In contrast, only epidermal growth factor (EGF) can activated ERK1/2 in HEK293 cells transfected with empty plasmid ( Fig. 1 D)."

reach
"In particular, aspirin decreased phosphorylated Akt and Erk activated by EGF."

sparser
"The EGF-induced ERK activation was inhibited to the basal level in cells pretreated with 10 nM MEK inhibitor."

reach
"SKCOX-1 cells were more responsive to aspirin when compared to cells transfected with empty vector, and decreased EGF activated Akt and Erk as well as cell viability."

reach
"We also showed that inhibition of Cdc2 activity with roscovitine did not restore EGF-induced ERK activation in Hela H2B–GFP cells ( Fig. 7 )."

sparser
"Control experiments showed that the activation of ERK and STAT1 by EGF and IFN-γ, respectively, was not compromised in U2OS-shUb-Ub(K63R) cells ( xref )."

sparser
"In a study by Impagnatiello et al. [ xref ] indicating an anti-proliferative effects of the overexpressed Spry1 and Spry2 on endothelial cells in the presence of FGF, VEGF, and EGF, while FGF- and VEGF-induced activation of ERK were repressed by Sprouty, EGF-activated ERK was left unaffected."

reach
"XIA W, MULLIN RJ, KEITH BR et al. : Anti-tumor activity of GW572016 : a dual tyrosine kinase inhibitor blocks EGF activation of EGFR and erbB2 and downstream Erk1/2 and AKT pathways."

reach
"To test this possibility, EGF activation of ERK was measured using beta-arrestin2 siRNA transfected immortalized astrocytes (XREF_FIG)."

reach
"We confirmed that EGF stimulated ERK1/2 activation in hTERT-C3 cells, and this response was blocked by treatment with AG-1478 (1 muM), a specific inhibitor of the EGFR."

sparser
"It was revealed in a study by Sasaki et al. [ xref ] that the expression of Spry2 and Spry4 in HEK293 cells inhibited FGF-induced ERK signaling but did not affect EGF or PDBu activation of ERK."

reach
"EGF-stimulated ERK and AKT signaling is comparable in KRAS G12X/+ oncogenic mutants and KRAS G12G(WT)/+ cells."

reach
"The inhibitory activity of DSF-102 was furtherly investigated evaluating the p44/42 ERK and p38 MAPK activation induced by EGF exogenously administered."

reach
"Both neural growth factor (NGF) and epidermal growth factor (EGF) activate ERK in PC12 cells (a cell line derived from rat pheochromocytoma)."

reach
"Therefore, the present study was conducted to determine the molecular mechanisms by which EGF activates the PI3 K-AKT1 or ERK1/2 MAPK cell signal transduction pathway for proliferation, migration and [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The Src homology domain-2 containing tyrosine phosphatase-1 (SHP-1), highly expressed in hematopoietic cells, plays a positive role in epidermal growth factor-induced ERK1/2 activation and negative role in anti-apoptotic signaling pathway.23 Inhibition of the SHP-1 gene expression by miRNA has shown reduced cardiomyocyte apoptosis and increased cardio-protection through Akt activation in a rat model of cardiac ischemia-reperfusion injury.24 Combining gene therapies involves utilizing a hypoxia-inducible plasmid expressing both HO-1 and miSHP-1, demonstrating higher levels of HO-1 expression and miSHP-1 production in hypoxic cells compared to normoxic cells."

reach
"Western blot results showed that EGF could activate MAPK/ERK pathway proteins."

sparser
"SPRY2-OE Enhances EGF-Induced Activation of PLCγ1, Whereas shSPRY2 Reduces EGF-Induced Activation of PLCγ1 and ERK."

reach
"In contrast, EGF induces transient ERK activation, and eventually increases cell proliferation (Murphy et al., 2002)."

sparser
"EGF-induced activation of PLCγ1 and ERK was inhibited by shSPRY2 in SF126 cells ( xref B)."

reach
"Quantification of EGF-induced ERK activation in individual cells and in a population of cells confirmed the suitability of electric stimulation for ERK activation dynamics interrogation (Figures S1B–S1E)."

reach
"ERK activation induced by EGF was consistent with that reported before (Albeck et al., 2013)."

reach
"We examined this issue by quantifying the dynamics of Erk signaling induced by epidermal growth factor, basic fibroblast growth factor (bFGF), and platelet derived growth factor (PDGF) in NIH-3T3 fibroblasts."

reach
"Therefore, ERK activation induced by EF and EGF, including the dynamics of Fc/Fn ratio are comparable to each other (Figure 2 and Video S1, compared to Figure S1 and Video S2)."

reach
"Adhesion to fibronectin coated surfaces enhances Erk signaling elicited by epidermal growth factor but not by bFGF or PDGF."

reach
"Co-transfection of cells with dominant negative mutant N17 Ras or with a betagamma scavenger, CD8-betaARK-C, suppressed opioid stimulation of ERK1 and ERK2.When EGF was used to activate ERK, chronic m[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In addition, adenoviral overexpression of dominant negative ras (Ad.HA-Ras (N17)) partially inhibited the ATP- and PDBu induced activation of ERK1/2 and blocked ionomycin- and EGF stimulated ERK1/2, and inhibition of tyrosine kinases with AG-1478, an EGFR inhibitor, or the src family kinase inhibitor PP2 attenuated ATP stimulated ERK1/2 activation."

reach
"This is in consistent with ERK activation induced by EGF that in MCF10A cells, EGF leads to multiple stochastic pulses of ERK, whereas in PC12 cells, EGF induces one ERK activity pulse (Vaudry et al., 2002; Worster et al., 2012)."

reach
"Furthermore, EGF activated Erk within 5-15 minutes and pre-addition of aspirin inhibited the EGFR activated Erk in OVCAR-3 cells."

reach
"We confirmed that the silkworm carboxypeptidase inhibitor inhibited the expression of c-Myc through the MAPK/ERK pathway initiated by EGF/EGFR, thereby affecting the related cyclins and inhibiting the proliferation of gastric cancer cells.Currently, many scholars are focused on the discovery of lead compounds from active ingredients of natural products."

reach
"EGF activated ERK1/2 has also been found to both stabilize and induce the expression of FOXC1, a transcription factor with implications in triple negative breast cancer [XREF_BIBR, XREF_BIBR]."

reach
"In contrast to the previous reports, our study show that silencing of Erbin expression completely blocked catecholamine-induced ERK activation, whereas overexpression of Erbin PDZ domain efficiently r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We also found that EGF mediated ERK itself is sensitive to PP2 being reduced by 45% (data not shown)."

reach
"As expected, EGF stimulated a 5-fold increase in phospho-Erk1/2 (Figure 1c) that was unaffected by UBO-QIC and completely inhibited by AG1478."

reach
"As a biologically active ingredient, it can inhibit the expression of the proto-oncogene c-Myc by affecting the MAPK/ERK pathway initiated by EGF/EGFR, thereby inhibiting the proliferation of gastric cancer cells."

reach
"Both LPA and EGF stimulated ERK activation in HASM cells, assessed by immunoblotting with antibodies specific for phospho-ERK [34]."

reach
"To determine whether G z could act as the transducer of the chronic opioid agonist actions on EGF stimulated ERK in the presence or absence of PTX, COS-7 cells were cotransfected with mu or kappa OR +[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"ALA also inhibited activation of Erk mediated by 5-hydroxytryptamine (5-HT) [19] and epidermal growth factor (EGF), basic fibroblast growth factor (bFGF), and platelet-derived growth factor (PDGF) [20]."

reach
"Spatio-temporal regulation of expression of these genes by EGF requires further investigation to better understand specific functions and regulatory mechanisms associated with development of the pig c[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, EGF activated EGFR, Erk and Akt in both SKpcDNA and SKCOX-1 cells, but aspirin inhibited EGFR, Erk and Akt activation in SKCOX-1 cells but not in SKpcDNA cells."

reach
"Stimulation by LPA alone and by EGF alone was completely blocked by the MAPK-ERK kinase (MEK) inhibitor U0126; however, modest stimulation by the combination of LPA+EGF was still observed in the prese[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, detailed comparisons of the time courses for ERK activation by LPA, EGF and LPA+EGF indicated that ERK is not the locus of synergism [34]."

reach
"On the other hand, H89 had no effect on the activation of ERK by EGF, although the PKA inhibitor diminished P-ERK expression when EGF and 8-Br-cAMP were added conjointly Figure 9 c and d."

reach
"In PTX treated cells, mu agonist inhibited EGF activation of ERK by ~ 40%, in the presence or absence of Galpha z transfection."

reach
"LPA induced strong ERK activation within 2 min, which peaked at 10 min and then gradually declined, with no significant stimulation after 6 h. EGF also induced activation within 2 min, which peaked at[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"By contrast, in HKC cells the activation of ERK by EGF was not accentuated by 8-Br-cAMP; rather, the nucleotide decreased the expression of P-ERK to a small extent Figure 9 e and f."

reach
"Transfected Galpha z did not replace ADP ribosylated G proteins and couple kappa OR mediated inhibition of EGF activation of ERK."

reach
"In Csk (-/-) fibroblasts, suppression of SFK results in prolonged EGF induced activation of Erk1/2, with concomitant suppression of EGFR degradation."

sparser
"In SF126 cells, shSPRY2 increased FGF2-induced PLCγ1 but not ERK signaling, whereas it reduced EGF-induced ERK and PLCγ1 activation."

reach
"Because the combination of the PI-3-kinase and Mek and Erk inhibitors synergistically increased beta-casein promotor luciferase activity and because neither LY294002 nor Rapamycin affects EGF induced Erk activation, we conclude that the PI-3-kinase and Mek and Erk signaling pathways are independent and synergistic in their ability to block lactogenic differentiation in HC11 cells."

reach
"Together, these data indicate that ERK activation by both LPA and EGF is required for synergism; however, signaling pathways in addition to ERK are also required since ERK activation does not exhibit [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Chronic mu OR inhibition of EGF stimulation of ERK activity occurred when palmitoylation deficient mutant alpha z C3A, which as described above mediated opioid stimulation of adenylate cyclase II, was[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"SPRY2 inhibits degradation of FGFR3, thereby enhancing FGF2-induced ERK activation, and it inhibits EGFR degradation and enhances EGF-induced ERK activation."

reach
"The more extensive model captures Akt and Erk signaling induced by EGF and HRG, 21 whereas the second, smaller model focuses specifically on activation of Akt by HRG and BTC."

reach
"These results are consistent with a role for Galpha z as transducer of chronic mu opioid inhibition of EGF activation of ERK."

sparser
"In contrast to IGF-1, AICAR did not block epidermal growth factor (EGF)-dependent Raf-1 and Erk activation, but our results demonstrated that multiple Raf-1 upstream pathways induced by EGF were differentially affected by AICAR: inhibition of Ras activation and simultaneous induction of Ras-independent Raf activation."

trips
"Inhibition of Src tyrosine kinase activity abrogated EGF-stimulated Erk activation and cell migration."

reach
"Thus, our studies indicate that EGF activates ERK through the sequential stimulation of Ras, Raf-1 and MEK, whereas PKA activates ERK through its capacity to activate B-Raf and MEK Figure 10."

reach
"Whereas seribantumab was effective at inhibiting HRG induced p-ErbB3 and p-Akt and erlotinib was effective at inhibiting EGF induced p-Erk, only the combination of both drugs shut down both pathways in the presence of HRG and EGF, which is qualitatively in agreement with the simulation results."

reach
"21 EGF transiently activates Erk, a member of the MAPK family."

sparser
"Our findings provide a possible resolution for this apparent discrepancy by demonstrating that the localization of EGF-induced ERK activation rather than the absolute level of activation is the critic[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"This Rap dependent mechanism may also be operating in the case of both Galpha z and Galpha 12, but because Galpha 12 uses predominantly its alpha subunit to act on the effector, this G protein may be [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"NT stimulated ERK activation via a protein kinase C (PKC)-dependent (but EGF receptor independent) pathway in PANC-1 and MIA PaCa-2 cells, whereas EGF promoted ERK activation through a PKC independent pathway in these cells."

reach
"The overexpressed, constitutively active mutant of Galpha z (Q205L) inhibited EGF activation of ERK to a comparable extent in the absence and presence of DAMGO."

reach
"In senescent human diploid fibroblasts, the phosphorylation of ERK1/2 was not induced by epidermal growth factor (EGF), suggesting that the down-regulation of EGF-induced ERK activation might be due t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Although further studies are required to clarify the relationship between EGFR signaling pathways and COX-1 in ovarian cancer, here we show that aspirin blocked EGF stimulated Akt and Erk activation in COX-1 stable transfected cells."

reach
"ERK activation by both LPA and EGF presumably involves the Ras regulated kinase cascade in these as in other cells [30], although the involvement of Ras and each of the downstream kinases has not been[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGFR tyrosine kinase activity is required for both EGF stimulation of ERK and for mitogenic synergism."

sparser
"Moreover, some studies have confirmed that ERK 1 and epidermal growth factor can activate ERK and calpain-2, but the activation of calpain-2 depends on ERK [35,36] ."

sparser
"As a positive control, we showed that ERK1/2 were strongly activated by EGF in the interphase ( Fig. 6 A–C)."

reach
"After differential reduction of p14 protein levels by p14 siRNA1 and p14 siRNA2, EGF induced activation of endogenous ERK1/2 decreased in a p14 concentration dependent manner."

reach
"While blocking the increase in intracellular Ca 2+ by locking the IP 3 R calcium channel with 2-APB had no effect on EGF activation of the ERK or AKT signaling pathways, it abolished the rapid EGF mediated induction and repression of gene expression."

sparser
"In a study by Edwin et al. [ xref ], where Spry2 unexpectedly inhibited EGF activation of AKT and exhibited no significant effect on EGF activation of EGFR and ERK, they observed that Spry2 increases the amount and activity of PTEN that was found necessary for Sprouty to attenuate EGF-activated AKT and to inhibit cell proliferation."

sparser
"Interestingly, in human intestinal epithelial cells cortical active ERK1/2 has been recently associated with a stronger EGF-induced ERK activation as compared to cytoplasmic ERK in under-differentiate[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The present results suggest that Src kinases may be involved in ERK1/2 activation induced by PGF 2alpha and norepinephrine, while the EGF induced ERK1/2 response appears to be independent of these Src kinases."

sparser
"Inhibition of Cdc2 activity by roscovitine in mitotic cells restored ERK activation by EGF [40] ."

sparser
"Since intracellular calcium levels fail to reach the near-millimolar concentrations required for calpain activation, epidermal growth factor (EGF) triggers the MAPK cascade and eventually activates ERK, which phosphorylates calpain at Ser50 and leads to calpain activation xref , xref ."

reach
"Furthermore, we show that similar to the light intensity the dose of epidermal growth factor induces a change in CNK1 complex composition and thereby allows RAF and ERK signalling or exertion of an AKT and RAF crosstalk which suppresses RAF and ERK signalling."

reach
"In cells where p14 was reduced by p14 siRNA2, EGF induced ERK1/2 activation was only moderately affected after 5 min of EGF stimulation."

sparser
"In ShSrx cells, a similar pattern of time-dependent activation of ERK is observed, however, both the basal as well as EGF-induced activation of ERK are significantly reduced ( xref )."

reach
"Pharmacological inhibition of EGFR and ERK1/2 abrogated expression of GLI/EGF target genes, suggesting that EGF can signal via MEK/ERK to cooperate with GLI proteins in regulation of target gene expression [82]."

reach
"EGF treatment (1mg/kg, 15 min) markedly activated ERK in ME tanycytes (XREF_FIG)."

sparser
"Sprouty evidently inhibits c-Cbl-induced ubiquitination and degradation of EGFR, thereby sustaining EGF-activated ERK [ xref , xref , xref , xref , xref ]."

reach
"While several studies have suggested that Erk activation can be regulated through the PI-3-kinase pathway [XREF_BIBR, XREF_BIBR] our data demonstrated that EGF stimulation of Erk activation in HC11 mammary epithelial cells was not altered by blocking PI-3-kinase signaling with LY294002."

reach
"In the EGFR signalling pathway, in coordination with EGF induced EGFR phosphorylation, methylation of EGFR by PRMT5 determines EGF mediated ERK activation and cell proliferation XREF_BIBR."

reach
"EGF activated predominantly ERK pathway."

sparser
"We showed that while MEK was well expressed in mitosis at a level similar to that in the interphase, MEK was not phosphorylated in response to EGF in mitotic Hela H2B–GFP cells ( Fig. 8 E & F), which [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The activation of ERK1/2 by EGF was, however, reduced to a lesser extent."

sparser
"Sasaki A, Taketomi T, Wakioka T, Kato R, Yoshimura A. Identification of a dominant negative mutant of Sprouty that potentiates fibroblast growth factor- but not epidermal growth factor-induced ERK activation."

reach
"These results were consistent with the findings in XREF_FIG and XREF_FIG that suggested that beta-arrestin2 reduced EGF induced ERK activation by blocking the growth factor pathway."

reach
"EGF stimulated ERK and PI3K and AKT activity and increased the viability of MCF-7 cells."

reach
"Paulsen et al. showed that EGF-mediated signaling results in H O production and oxidation of the EGFR active site (Cys797) to enhance kinase activity and downstream AKT and ERK activation [38]."

sparser
"It is noteworthy that NSC-87877 inhibited epidermal growth factor (EGF)-induced activation of Shp2 PTP, Ras, and Erk1/2 in cell cultures but did not block EGF-induced Gab1 tyrosine phosphorylation or Gab1-Shp2 association."

reach
"Transfection studies in COS-1 cells demonstrated that in vivo phosphorylation of epidermal growth factor stimulated ERK depended on VHR protein levels."

reach
"In our 3D experiments, it is likely that the activation of AKT signaling by the HIF pathway counteracted its induction by collagen I.The kinetics of EGF-mediated ERK activation were different in monolayer and spheroid cultures."

reach
"In contrast, EGF strongly stimulated ERK without reducing its activity upon longer treatment in CNK1 knockdown cells."

reach
"Moreover, prolonged epidermal growth factor stimulation, which activates the ERK pathway, induces apoptosis in normal gastric epithelial cells [24]."

reach
"We further found that MKP3 activity was not altered by the addition of GST-VRK3 protein ( Fig. 5 D), suggesting that VRK3 can indirectly modulate ERK activity by activating VHR in embryos.Stimulation [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In addition, EGF activated Akt and Erk whereas TNF solely activated IκB in OVCAR-3 cells (Figure  xref C)."

reach
"Western blot analysis of kinetics of EGFR and MAPK proteins activation treated with As 2 O 3 ( Fig. 1 A,B ) or EGF ( Fig. 1 C,D) found that both agents could induce activation of ERK1/2 and JNK1."

sparser
"However EGF activated Erk without IκB phosphorylation whereas TNF highly activated IκB and had a small effect on the activation of Erk (Figure  xref C)."

reach
"Similarly, neither DAMGO nor morphine displayed proliferative activities and proved to inhibit EGF stimulated ERK activation and cell proliferation."

reach
"A novel finding of this study is the implication of beta-arrestin2 in exogenous EGF stimulation of ERK mediated astrocyte proliferation (XREF_FIG)."

reach
"Some signaling components or adaptor proteins involved in EGF stimulated ERK activation undergo endocytosis and are recruited along with EGFR to endosomes."

sparser
"EGF activated IκB, Akt and Erk whereas TNF activated only IκB and reduced EGF-activated Akt and Erk in these cells (Figure  xref C)."

sparser
"EGF activated IκB and Erk whereas TNF activated only IκB (Figure  xref C)."

sparser
"In vascular smooth muscle cells, angiotensin II-induced tyrosine phosphorylation of two major proteins (p120, p70) and ERK activation are markedly reduced by α-tocopherol, whereas ERK activation by ep[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Addition of TPA or EGF did produce the expected increase in ERK activation within 5 min, but this was lost and reduced to below control levels within 15 min."

reach
"We have previously demonstrated that siRNA depletion of MEK1 resulted in sustained EGF stimulated ERK activation in G2 phase HeLa cells [21], suggesting that tMEK is involved in the transient ERK acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We found that the activation of Erk by EGF stimulation was attenuated in the cells under overexpression of GFP-Rab21, suggesting that Rab21 reduces the duration of EGFR activation indicated by Erk pho[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"ERK MAPK could be activated by diverse growth factors (e.g., EGF, HGF, IGF, PDGF, and TGFβ) and regulated at its origin by Ras GTPases, which lead to the activation of MAPKKK, constituted by Raf family kinases."

reach
"Similar observations have been reported in a computational model of EGF induced ERK activation in HeLa cells."

sparser
"While our finding that ERK was not activated by EGF during the M phase is consistent with early reports, the previous reports suggest that the ERK inactivation is due to the inactivation of EGFR durin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In particular, SHP-2 is involved in activation of ERK1/2 by epidermal growth factor (EGF) [10] ."

reach
"For example, ERK1/2 and PKB and Akt, components activated upon insulin stimulation, can be activated by hepatic growth factor and epidermal growth factor treatments in rat primary hepatocytes."

sparser
"We also showed that inhibition of Cdc2 activity with roscovitine did not restore EGF-induced ERK activation in Hela H2B–GFP cells ( Fig. 7 )."

reach
"EGF induced ERK signaling in HeLa cells is increased 1 h after stimulation when translation is blocked by cycloheximide."

sparser
"EGF-induced ERK activation is also slower and more persistent in cysts than in plastic-grown cells (see Supplemental Figure S1 at http://www.developmentalcell.com/cgi/content/full/7/1/21/DC1 and data [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Similarly, in SHP2 D61G-MB231 cells, the EGF induced ERK and PI3K and Akt pathway activities were markedly enhanced."

sparser
"In contrast to HGF, EGF-induced ERK activation attenuates after 3 hr (see Supplemental Figure S1 at http://www.developmentalcell.com/cgi/content/full/7/1/21/DC1 )."

reach
"When EGF and heregulin were contacted with a breast cancer cell line, EGF resulted in a short-term (~ 30 min) spike in Erk activation, whereas heregulin elicited a longer duration of Erk activation."

sparser
"The overexpression or reduction of IQGAP1 expression by siRNA both caused the activation of Erk2 by EGF or IGF-1 in MCF-7 cells to be reduced, probably due to an optimal level of intracellular IQGAP1 being necessary for maximal level of activation of Erk by EGF or IGF-1 [ xref ]."

sparser
"However, these modeling analyses also revealed non-obvious modes of potential regulation: (i) ERK activation is robust to parameter perturbation at high ligand doses, whereas Akt activation is not and (ii) PI-3K is a dominant factor for long-term but not short-term EGF-induced ERK activation."

sparser
"Knockdown of either SOS1 or Ras isoforms decreased the EGF-induced activation of pCRAF, pMEK and pERK after 24 hours of vemurafenib treatment in A375 cells, suggesting that reactivation of ERK signaling requires these proteins ( xref , respectively; lanes 8 vs 4)."

sparser
"Celecoxib inhibited basal and EGF-stimulated proliferation, hypoxia-related HIF-1α recruitment/stabilization as well as hypoxia- and EGF-dependent activation of ERK and PI3K. Interestingly, celecoxib prevented EMT-related changes, as shown by modifications of β-catenin intracellular localization or vimentin and E-cadherin levels, as well as HT-29 invasiveness induced by hypoxia, EGF, or hypoxia plus EGF."

sparser
"In fact, increased cellular CTCF activity resulted from EGF-induced Erk activation plays functional roles in inhibition of Pax6 since suppression of Pax6 activity is required for EGF to stimulate corn[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"2) EGF stimulation resulted in increased serine and threonine phosphorylation of nuclear symplekin, which was abrogated by the MEK and ERK inhibitor U0126, indicating that ERK1/2 regulated symplekin phosphorylation can be induced by EGF treatment."

sparser
"EGFR inhibitors (gefitinib and AEE788) reduced clonogenicity, proliferation and survival of ependymoma-SC lines dose-dependently, and blocked EGF-induced activation of EGFR, Akt and extracellular signal-regulated kinase 1/2."

reach
"In addition, EGF may also regulate the expression functions of claudins in ovarian and colon cancer cells during cancer development via the EGF activated ERK1/2 and PI3K-Akt pathways [XREF_BIBR, XREF_BIBR]."

reach
"In pancreatic acinar cells both cholecystokinin (CCK) and epidermal growth factor (EGF) are known to stimulate ERK."

sparser
"Previous reports have shown that p150 Glued of dynactin, as well as the dynein motor complex, are involved in signal-induced phosphorylation of MKK3/6 and p38 MAPKs, but not activation of ERKs by EGF ( xref )."

reach
"To test whether elevated intracellular calcium was stimulating ERK activation, and activated ERK then induced immediate early gene expression, we examined the effect of 2-APB on rapid EGF activation of the ERK and AKT signaling pathways."

sparser
"It was reported that EGF-induced activation of extracellular signal-regulated kinase (ERK) results in phosphorylation and activation of CK2α ( Ji et al., 2009 )."

reach
"Under the same conditions used in the gene expression experiment, 2-APB did not inhibit rapid and robust EGF activation of the ERK and AKT (protein kinase B) signaling pathways (XREF_FIG)."

reach
"Takahashi-Niki K, Kato-Ose I, Murata H, Maita H, IguchiAriga SM, Ariga H. Epidermal growth factor dependent activation of the extracellular signal regulated kinase pathway by DJ-1 protein through its direct binding to c-Raf protein."

reach
"As controls, we showed that EGF stimulated ERK activation and this ERK activation is abolished in the presence of U0126 (XREF_FIG)."

reach
"Furthermore, Cu-B inhibits MEK 1/2 as well as the constitutive and EGF-induced ERK 1/2 activation, indicating a definitive involvement of MAPK signal transducers in regulating Cu-B-mediated anti-melanoma activity."

sparser
"This synergistic increase of EGF-induced ERK activation afforded by GH costimulation was also detected in 3T3-L1 preadipocytes and in HEK-293 cells that transfected to express GHR and JAK2 (data not shown)."

sparser
"However, EGF-induced ERK activation was more sustained and easily detected."

sparser
"3.2 GH augmentation of EGF-induced ERK activation in 3T3-F442A cells resides at the Raf-MEK level."

sparser
"Like other growth factors, EGF activates ERKs mainly by engaging the Grb-2-SOS-Ras-Raf-MEK-ERK pathway [ xref ]."

sparser
"Moreover, ERK and JNK, which can be activated by TNFα and EGF [80] , phosphorylate S84 of the A/B domain of PPARγ1 in vitro and in turn inhibit both ligand-independent and ligand-dependent transactiva[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"As shown in Figure xref , activation of MEK1/2 and ERK1/2 by EGF was severely impaired in IEC-6/KRAS and DLD-1 cells knocked-down for SHP-2."

sparser
"As our data suggest that GH synergistically augments EGF-induced ERK activation in 3T3-F442A cells, we sought to localize the level(s) in EGF-induced downstream ERK signaling at which GH exerts this effect."

sparser
"Importantly, both of these siRNAs also markedly reduced EGF-induced ERK1/2 activation but did not decrease LPA-induced ERK activation ( xref E), suggesting that LPA can bypass the effects of down-regulation of AGK."

sparser
"These data strongly suggest that GH's ability to synergistically augment EGF-induced ERK activation resides at the level of Raf's ability to activate MEK."

reach
"EGF induced ERK activation downstream of FAK requires rac1-NADPH oxidase."

reach
"Our studies revealed that Beclin 1 regulates both EGF and insulin like growth factor-1 (IGF-1) stimulated AKT and extracellular signal regulated kinase (ERK) activation in breast carcinoma cells."

sparser
"In contrast, EGF induced ERK1/2, p38 MAPK and Akt activation in MPSC1 cells."

sparser
"Remarkably, down-regulation of AGK drastically reduced ERK1/2 activation induced by EGF, which, as expected, was bypassed by addition of the AGK product, LPA."

sparser
"These compelling results lead us to conclude that GH facilitates the synergistic lessening of KSR serine-392 phosphorylation, suggesting that the mobilization of this important EGFR scaffold is an important part of GH's sensitizing effect on EGF-induced ERK activation."

reach
"Next, we examined the effect of SF1126 on EGF mediated activation of ERK (phospho-ERK) in these cells."

sparser
"We sought to determine whether selective GH augmentation of EGF-induced ERK activation was mirrored by interaction of another cytokine receptor-engaging hormone (PRL) with another RTK-engaging growth factor (PDGF)."

reach
"Since 2-APB blocks EGF induced immediate early gene expression without affecting EGF activation of the ERK and AKT signaling pathways, 2-APB represents a useful new probe for dissecting the roles of immediate early gene expression and ERK and AKT activation in downstream actions of EGF."

reach
"The duration of Epidermal Growth Factor (EGF) stimulation modulates ERK dynamics and controls differentiation."

sparser
"Forskolin, a direct activator of adenylyl cyclase, inhibited CCK- and EGF-induced activation of c-Raf-1 and ERK1/2 in a manner similar to that of PTx."

sparser
"In PTx-treated cells, the cAMP content was increased and forskolin did not further inhibit CCK- and EGF-induced activation of c-Raf-1 or ERK1/2."

sparser
"Because SHP2 phosphatase activity is required for the full activation of the Ras-Erk1/2 pathway, xref , xref we assessed the cellular effect of 11a - 1 on EGF-induced Erk1/2 activation in H1975 cells."

reach
"EGF infusion for 6 days induced continued CREB and ERK activation in nestin+ cells paralleled by intense periventricular cell proliferation."

reach
"Overexpression of EGFR in MCF-7 cells confers estrogen independent growth, and studies using an MCF-7 cell line model of hormone resistance showed that EGF dependent activation of heterodimers of EGFR and HER1 and HER2 resulted in activation of the ERK pathway and increased cell proliferation."

sparser
"EGF-induced ERK activation in MC3T3-E1 OBs ultimately leads to proliferation and differentiation xref ."

sparser
"Short-term EGCG exposure substantially decreased EGF-induced EGFR, AKT and ERK1/2 activation."

reach
"Since the above data indicate that USP10 is involved in the EGF response, the next question was what molecular pathways might underlie this phenomenon.As EGF is a potent stimulator of the PI3K/AKT/mTOR and MAPK/ERK pathways and dysregulation of these pathways is common in CRC, we next tested whether USP10 knockout affects EGF-dependent activation of the PI3K/AKT/mTOR and MAPK/ERK signaling pathways.First, we examined EGF-induced phosphorylation of EGFR tyrosine 1068 (p-EGFR)."

sparser
"In MCF-7 and MDA-MB-231, breast cancer cells E2 was reported to not alter Erk1/2 phosphorylation, while EGF induced robust Erk1/2 activation [6,17–21] ."

reach
"These findings sharply contrasted with the transient phosphorylation patterns of EGF receptor, Erk1/2 and Akt that were induced by soluble EGF supply ( Fig. 7 )."

sparser
"Niacin showed a selected additive effect on chemoattractant-induced activation of ERK1/2, JNK and PI3K pathways, but only the MEK inhibitor UO126 reduced niacin-mediated inhibition of macrophage chemotaxis, while activation of ERK1/2 by EGF alone did not inhibit fMLF-mediated migration of HEK293T cells co-expressing HCA2 and fMLF receptor FPR1."

sparser
"Our results show that (i) both high [Ca 2+ ] o and EGF activated ERK1/2 inducing TRPC1 over-expression and MCF-7 cells proliferation, and (ii) the inhibition of the MMPs, known to be involved in the p[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Blockade of the epidermal growth factor (EGF) receptor by AG1478 significantly reduces activation of ERK, suggesting that EGF receptor transactivation could act upstream of the ERK pathway in PC12 cells."

reach
"For example, in rat PC12 cells, sustained epidermal growth factor (EGF) stimulation of ERK signaling was found to induce proliferation, while sustained nerve growth factor (NGF) stimulation resulted in differentiation."

reach
"XREF_BIBR, XREF_BIBR In MCF-10A cells, sustained EGF stimulation was observed to cause oscillations of ERK activity in single cells."

sparser
"We found that 1 μM of gefitinib was sufficient to inhibit phosphorylation of the EGFR at Tyr1068, as well as EGF-induced activation of ERK, in each of these MPNST cell lines ( xref )."

reach
"The MEK/ERK pathway that can be activated, for example, by EGF (Epidermal growth factor) also controls RSK2 activity which is responsible for activation of ELK3, a gene positively involved in proliferation and metastatic formation."

reach
"In addition, EGF is known to stimulate the ERK1/2 signaling pathway."

sparser
"The 5 h EGF exposure rendered the cells resistant to further ERK1/2 activation by EGF, as well as EGF-induced alterations in the Cx43 band pattern, all in agreement with the GJIC experiments."

reach
"In PC12 cells, epidermal growth factor induces transient ERK activation while nerve growth factor leads to a sustained ERK response, a response that may be essential for cell differentiation XREF_BIBR - XREF_BIBR."

reach
"On the other hand, EGF activated ERK at the same time, which could target MMP9 directly to promote the migration ( Figure 6 )."

reach
"Because the Erk signaling pathway is activated by EGF and is a major contributor to cellular growth, we have studied the phosphorylation of this kinase after exposure to anti-EGFR antibody or rapamyci[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, we have demonstrated that CCL17 transcription is upregulated by TNF-α-activated p38 and downregulated by EGF-activated ERK1/2."

reach
"EGF family members contribute to periderm migration by activating the ERK signaling pathway."

reach
"Fumos inhibits EGF induced ERK1/2 activation."

reach
"Shp2 is required for full activation of EGF induced ERK1/2 XREF_BIBR."

sparser
"Furthermore, we investigated how the minimal EGF-induced ERK activation affects the downstream transcription in HeLa cells by comprehensive transcriptional analysis."

sparser
"For example, EGF activated ERK and stimulated proliferation not only in liver but also in cell lines of colonic, bronchial, pancreatic and gastric origin [8] ."

reach
"In these cells EGF causes a transient activation of ERK, whereas NGF causes a sustained activation of ERK, thus the duration of ERK activation specifies signal identity [14]."

sparser
"Expression of wild-type (wt) Cbp remarkably suppressed EGF-induced activation of Src, ERK1/2, and Akt-1 enzymes, and NIH3T3 cell transformation, as well as colony formation of a breast cancer cell line (MDA-MB-468) in soft agar."

reach
"The results suggest that Fumos inhibits EGF stimulated ERK1/2 activation in a dose dependent manner in HeLa and HEK293T cell lines that contain wild type Ras."

reach
"Trastuzumab- and pertuzumab mediated effects on EGF induced ERK1/2 signaling were also minor in HCC1954 cells (XREF_SUPPLEMENTARY)."

reach
"For this reason, it is useful to have a better understanding of the key mutations and their significance and the synergistic effect of multiple mutations which are responsible for neoplastic transform[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In PC3 and LNCaP cells EGF stimulates Erk to various extents, however in MDA-PCa-2b cells EGF had little effect on Erk and strongly increased p-RPS6 along with IGF1 which was not seen to have an effect LNCaP or PC3 cells."

reach
"In the ApcMin/+ model, initiation of intestinal tumorigenesis by aberrant Wnt signaling is promoted by IL-17 XREF_BIBR, EGF activated ERK signaling XREF_BIBR, or TLR activation XREF_BIBR."

reach
"Our data indicate that Fumos does not inhibit EGF stimulated ERK1/2 activation in MDA-MB-231 cell line (XREF_FIG)."

reach
"To test, we determined whether Fumos inhibits EGF stimulated ERK1/2 activation in other cell lines expressing mutant Ras."

reach
"Robust phosphorylation (activation) of ERK was found in EGF treated cells, as predicted (XREF_FIG), whereas the EGF conjugated dendrimer induced minimal ERK activation."

reach
"These findings indicate that Fumos inhibits EGF induced ERK1/2 activation."

reach
"Moreover, treatment of cells with epidermal growth factor (EGF) that activates MEK and ERK, as well as genetic ERK activation by a constitutively active MEK (MEK CA), increased T562 phosphorylation levels; pharmacological or genetic MEK inhibition using the selective U0126 inhibitor or byexpressing a dominant negative MEK mutant (MEK DN); and reduced T562 phosphorylation; finally, in Mfn1 -/- cells re-expressing the MFN1 T562A mutant, phosphorylation was undetectable even in the presence of EGF (XREF_FIG G)."

reach
"Shimada et al. [60] identified the epidermal growth factor (EGF)-like factor that affects EGF receptor and then induces the ERK1/2 and Ca 2+ -PLC pathways in cumulus cells."

sparser
"Importantly, the chemical antagonism of CaM prevents KSR1 translocation to the plasma membrane and reduces activation of ERK by epidermal growth factor (EGF) (Parvathaneni et al.,  xref )."

reach
"Santos et al. used the SSPRs and pseudo-SSPRs to calculated the LRCs for NGF and EGF stimulated ERK pathway 15 respectively."

reach
"Therefore, Fumos inhibits EGF induced activation of ERK1/2 downstream of Ras in cells expressing wild type Ras."

reach
"BAK oligomerization in response to recombinant caspases-8-cleaved BID (cBID) was increased in mitochondria from cells where ERK was activated by EGF treatment (XREF_FIG D-3F)."

sparser
"We first examined effects of AG1478 and PP2 on EGF-induced ERK1/2 activation to compare the IR-induced EGFR activation with the ligand-dependent one."

sparser
"AG1478 suppressed EGF-induced ERK1/2 activation at 2–5 min after the addition of EGF into the culture medium but it failed to suppress the ERK1/2 activation later ( Fig. 2 A)."

sparser
"PP2 did not suppress EGF-induced ERK1/2 activation ( Fig. 2 A)."

reach
"In conclusion, our results show a novel role for claudin-3 overexpression in promoting the malignant potential of colorectal cancer cells, which is potentially regulated by the EGF activated ERK1/2 and PI3K-Akt pathways."

sparser
"These results suggest that EGF-induced ERK1/2 activation is independent of Src."

sparser
"In MCF7 and PC12 cells, the transient activation of ERK by the epidermal growth factor (EGF) leads to cell proliferation, while the sustained activation of ERK by heregulin (HRG) or nerve growth factor (NGF) leads to cell differentiation ( xref ; xref ; xref )."

reach
"We confirmed here that EGF treatment activates PI3K/AKT and MAPK/ERK signaling pathways and activates YAP1 in GCs."

reach
"Thus, the different duration of ERK signalling is biologically sensed as differences in c-Fos protein stability as result of at least two feedback loops and a time delay: a negative feedback that inac[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"This feedback may underlie the differential activation of ERK by EGF and NGF in PC12 cells."

sparser
"Therefore, we investigated whether impairment of Ca 2+ /CaM binding to KSR1 influences ERK activation by EGF."

reach
"For example, EGF and another member of the EGF family, heregulin (HRG), induce transient and sustained ERK activity in cell lines such as MCF-7 that express low and high levels of ErbB1 and ErbB3 receptors, respectively [XREF_BIBR, XREF_BIBR]."

sparser
"We validated by Western blot that EGF activates ERK in cells expressing GFP only, with maximal ERK activation seen 1 min after adding EGF (Figure  xref )."

reach
"For example, EGF and another member of the EGF family, heregulin (HRG), induce transient and sustained ERK activity in cell lines such as MCF-7 that express low and high levels of ErbB1 and ErbB3 receptors, respectively [XREF_BIBR, XREF_BIBR]."

reach
"Conversely, when the ERK pathway was stimulated by addition of EGF (25 ng/ml), TGF-β (10 ng/ml), VEGF (5 ng/ml), and basic fibroblast growth factor (bFGF, 40 ng/ml) to the culture medium, VEGF express[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"While GHR knockout is associated with diminished expression of the EGFR and a concomitant decrease in EGF signaling, GH overexpression results in EGFR overexpression with different effects depending on the signaling pathway analyzed : AKT and ERK1/2 pathways are induced by EGF, while STAT3 and STAT5 activation is heterologously desensitized."

reach
"In PC12 cells, EGF stimulation results in transient activation of Erk to induce cell proliferation, whereas NGF stimulation results in sustained activation of Erk to cause growth arrest and outgrowth of neurites."

sparser
"In this report we show that external Qi of YXQ has opposite effects on the Akt and ERK pathways in BxPC3 pancreatic cancer cells versus fibroblasts; while it inhibits basal Akt and ERK1/2 activity and[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Stimulation with EGF caused ERK activation that was also inhibited by both drugs."

reach
"By adding modifications to the established EGFR kinase inhibitor gefitinib, an approach to small molecule conjugates with a unique nuclear targeting peptoid sequence was tested in both human and murine breast tumor cell models for their capacity to inhibit EGF stimulated activation of ERK1/2 and STAT3."

reach
"It was found that EGF activates ERK signaling transiently, while NGF produces sustained ERK activation, suggesting that it is the difference in the dynamics of ERK activity that determines the cellula[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"However, rapid activation of Erk by EGF did result in significant smad2 linker phosphorylation ( xref )."

reach
"These data also indicated HLP46 has some selective cytotoxic effects on different human cell lines.Shp2 is essential for full activation of ERK1/2 induced by EGF (Epidermal Growth Factor)."

reach
"We examined whether HLP46 could inhibit the activation of ERK1/2 induced by EGF."

reach
"We found that in HaCaT cells ADAM17 depletion reduces ERK1/2 phosphorylation, independent from the treatment with PsVs.To solidify the hypothesis that ADAM17 acts via the ERK signaling pathway in HPV16 infection and entry platform formation, we restored ERK activation by the addition of recombinant soluble form of the epidermal growth factor (EGF) which is one of the EGFR-activating ligands."

reach
"Anti-tumor activity of GW572016 : a dual tyrosine kinase inhibitor blocks EGF activation of EGFR and erbB2 and downstream Erk1/2 and AKT pathways."

reach
"HLP46 also inhibits EGF-induced ERK1/2 activation in 293 T cells ( Fig. 4 A)."

reach
"All results suggested that HLP46 inhibits EGF-stimulated ERK1/2 activation dependent of Shp2.The PTP activity of Shp2 is required for full activation of Ras/ERK, and thereby Shp2 functions upstream of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"While activation of ERKs by EGF treatment was significantly increased, total ERKs did not change with any cell treatments.Western blot analysis demonstrated rapid time-dependent pMAP kinase activation[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Importantly, the kinetics of ERK1/2 activation by EGF (4 ng/ml) in gene-edited cells was essentially similar to that in parental cells ( xref and xref ), indicating that increased expression of endogenous mNG-KRAS did not affect signal transduction along the EGFR-ERK1/2 pathway."

reach
"Our previous studies have revealed that neuronal leucine-rich repeat 1 (NLRR1), a type І transmembrane protein, is associated with tumorigenesis by promoting cell proliferation through the activation of ERK mediated by EGF and IGF-1 (10) and negatively regulating anaplastic lymphoma kinase (ALK) (11) in NB, although the contribution of NLRR1 to other types of cancers is not understood."

sparser
"The ERK activations by WNT3a and EGF were totally abolished in the EGFR−/− fibroblasts ( Fig. 2 C)."

reach
"While ERK1/2 expression and basal phosphorylation did not significantly differ between GHR-KO and normal mice, EGF activation of ERK1/2 was significantly diminished in GHR-KO mice (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"The ERK signaling by IL-6 and EGF included several cases that lie outside of the 95% confidence interval."

sparser
"Axin inhibits cellular proliferation and ERK pathway activation induced by either epidermal growth factor or Ras, indicating a role of Axin in the regulation of growth induced by ERK pathway activation."

sparser
"We showed that EGF-induced Raf-1 and ERK activation was reduced (≈60%) in luteal cells rendered PKC-deficient by chronic PMA treatment."

sparser
"Partial inhibition of EGF-induced Raf-1 and ERK activation in the absence of PKC indicates that mechanisms other than PKC also account for Raf-1 and ERK activation in response to EGF in bovine luteal [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Gab1-SHP2 interaction has been shown to mediate ERK (extracellular-signal-regulated kinase) activation by EGF (epidermal growth factor)."

sparser
"Using T47D and MCF-7 human breast carcinoma cells that express endogenous Gab1 and Gab2, we examined the role of these docking proteins in EGF-induced ERK activation."

reach
"To further investigate the bifurcation in EGF mediated activation of ERK1/2 versus STAT3 signaling upon subcellular compartmentalization of gefitinib, MDA-MB-468 cells were exposed to Gef-NArg, Gef-SV40-NArg, or gefitinib in a two-fold series of concentrations up to 1 muM for gefitinib or 5 muM for each peptoid conjugate (XREF_FIG)."

reach
"In fact, in addition to blocking DHT induced Erk activation, paxillin knockdown similarly blocks EGF induced Erk activation, indicating that paxillin functions downstream of the EGFR to regulate Erk activation in response to both direct (EGF) and indirect (DHT) EGFR activation [XREF_BIBR]."

sparser
"Expression of either SHP2-binding defective Gab1 or Gab2 mutant blocked EGF-induced ERK activation."

sparser
"Furthermore, Gab1 and Gab2 are critical signalling threshold proteins for ERK activation by EGF."

reach
"However, EGF stimulation of ERK activation was suppressed in PTEN expressing cells."

sparser
"Neither CHC depletion nor Dyngo-4a significantly altered the kinetics of EGF-dependent ERK1/2 activation, although there was a trend of increased magnitude of ERK1/2 activation at early time points after EGF stimulation in CHC-depleted cells ( xref )."

sparser
"Furthermore, treatment of primary hepatocytes with the proteasome inhibitor MG-132 did not affect EGF-induced Erk activation ( Figure S5 C)."

sparser
"Previous studies showed that cells without all three ShcA isoforms are defective in H 2 O 2 or EGF induced ERK activation, indicating that ShcA proteins are required for optimal ERK activation under t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Interestingly, CMTM7 knockdown selectively enhances AKT but not ERK pathway activation, which is stimulated by EGF."

sparser
"In contrast, Gγ9 and p110γ knockout did not affect ERK1/2 activation by epidermal growth factor (EGF) and insulin-like growth factor 1 ( xref G and xref )."

reach
"For instance, EGFR endocytosis to early endosomes was found to be required for the full stimulation of EGF induced ERK1/2 and PI3K and AKT signaling."

sparser
"The activations of MEK1/2, ERK1/2 and p70S6K by EGF are key elements in the classical MAPK signalling cascade EGF → ERBB1/1 → GRB2 → SOS1 → RAS → RAF1 → MEK1/2 → ERK1/2 → p70S6K."

reach
"As shown previously [2], in HEK293 cells that do not express Klotho, ERK1/2 was activated by EGF but not by FGF23, as indicated by elevated levels of phosphorylated ERK1/2 (Figure 5A)."

reach
"Since EGF, a ligand of receptor tyrosine kinase, is widely reported to promote tumor growth, cell proliferation and apoptosis deregulation through activation of its downstream signaling, including MEK/ERK [25], we clarified whether 22-(4′py)-JA was able to suppress MEK/ERK signaling mediated by EGF."

reach
"Indeed, EGF induction of STAT5 and ERK1/2 was more pronounced than the activation of STAT3 and AKT in normal mice (XREF_FIG)."

sparser
"This is reinforced by the inhibition of clathrin-mediated endocytosis using two independent approaches, which did not significantly affect the time course of EGF-induced ERK1/2 activity ( xref ) and which confirmed that EGFR endocytosis is not an essential process for EGF-induced ERK1/2 activation in HeLa cells."

reach
"In conclusion, our data reveal the functional relationship between YB-1 and CXCL14 in EGF mediated ERK signaling, and YB-1 expression is a significant prognostic marker to predict prostate cancer."

sparser
"Nonetheless, compound 9 was capable of blocking EGF-induced activation of SHP2, Ras, and ERK1/2 in cell cultures but did not inhibit EGF-induced Gab1 tyrosine phosphorylation or Gab1-SHP2 association."

reach
"This phenomenon is similar to transient ERK activation induced by mitogens, such as EGF in PC-3 cells."

reach
"In addition, EGF and FBS rapidly activated ERK signalling in prostate cells (XREF_FIG)."

reach
"In our studies we found that adiponectin attenuated ERK signalling (the MEK-ERK-p90RSK axis), which is activated by EGF or FBS."

reach
"(3) Both p38MAPK and ERK signalling pathways were activated by EGF in JEG-3 cells."

reach
"Interestingly, even though the cell cycle is profoundly altered by inhibiting CME, inhibition of CME by knocking out dynamin in MEFs does not significantly affect Akt and ERK mitogenic signaling stimulated by EGF."

eidos
"Differences in the cell lines were generated by stimulation with either epidermal growth factor ( EGF ) ( E ) , which induces ERK and other kinase activity ,30 or anisomycin ( A ) , which has been reported to activate both the JNK and p38 MAPK cascades.31 Addition of U0126 , a MEK inhibitor ( U ) ,32 achieved inhibition of ERK ."

sparser
"These same cells also downregulated elastin mRNA in response to heparin-binding epidermal growth factor induced activation of extracellular signal-regulated kinase (ERK) [9] ."

sparser
"Our failure to confirm downstream network-specific explanations for sustained versus transient ERK activation by NGF and EGF, together with other reports that receptor-level mechanisms encode ERK dynamics ( xref – xref ), prompted us to compare RTK and ERK signaling kinetics."

reach
"In the present study we show that EGF activates both EGFR- and ERK1/2-signaling but not the Smad2/3-signaling pathway."

reach
"None of the other treatments had any significant effects on the uPAR induced response, although pertussis toxin did inhibit lysophosphatidic acid induced ERK activation, bisindolylmaleimide I and Go6983 inhibited PMA induced ERK activation, and PD98059 and U0126 inhibited platelet derived growth factor- and EGF induced ERK activation (data not shown)."

reach
"In conclusion, we have shown that claudin-3 overexpression is associated with the increased malignant potential of HT-29 cells and that the ERK1/2 and PI3K-Akt pathways, activated by EGF, are important regulators of this event."

sparser
"EGF-induced ERK1/2 and Akt activation in vehicle-treated cells was 14–15-fold at the 10 min time point and was sustained for up to 4 h with minimal decline."

sparser
"In agreement with our results, a more recent study conducted in HeLa cells showed that knockdown of clathrin heavy chain increased EGF-induced ERK activation [40] , suggesting that CME might have nega[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We propose that EGF may enhance TGFβ-induced EMT in LECs by potentiating both EGFR- and ERK1/2- signaling.Although EGF can activate EGFR- and ERK1/2-signaling, it alone cannot induce EMT in our lens e[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In contrast, knockdown of hnRNP A2 inhibited the Ras-MAPK-ERK pathway and prevented ERK1/2 activation by EGF."

sparser
"EGF-induced ERK activation in PC12 cells peaked at 5 min and fell back to near-background levels within 15 min ( xref ), whereas NGF-induced ERK activation persisted for >60 mins ( xref )."

sparser
"Based on our knowledge concerning the intimate links between signaling pathways involved in regulating proliferation and oxidative stress responsiveness, and prior findings indicating that EGF-induced[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, the strong early activation of ERK by EGF and its prolonged tail of activation, together with weaker activation of the other MAPKs (JNK and p38), might play a role in shaping the long-t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Together, these data argue against the idea that distinct network topologies in the Raf-MEK-ERK cascade alone (or in the identity of the GTPase that activates the cascade) define the difference in dynamics of ERK activation by EGF and NGF in PC12 cells."

reach
"There may be a difference in the EGFR- and/or ERK1/2-signaling pathways induced by EGF compared to that induced by TGFβ, and further experiments are required to clarify the precise role of these indiv[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGF-induced activation of the ERK cascade induces the expression of IFN-dependent genes, but HCV infection may result in their down-regulation (Tan et al. 1999)."

reach
"Treatment of cultured fibroblasts with MCD caused decreased levels of Erk2 in cholesterol-depleted caveolae fraction, but endothelial growth factor (EGF) caused hyperactivation of the remaining Erk is[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Thus, it is possible that NMDA receptor mediated Ca 2+ influx induces transactivation of EGF, which in turn activates ERK1/2."

reach
"Further studies demonstrated that epidermal growth factor activated robust amplification of ERK and moderate upregulation of AKT in Eps8 overexpressing cells."

reach
"Induction of PKC and ERK Signal cascade by HB-EGF, EGF and HGF."

reach
"EGF mediated ERK activation has also been shown to phosphorylate caspase-9 and inactivate its caspase activity that is also similar to AKT function (Allan et al."

sparser
"Furthermore, these effects appear to be Src-specific because CD2 expression had no effect on EGF-induced activation of EGF receptor and ERK."

reach
"Chronic ethanol treatment results in suppression of p42/44 MAPK activation induced by EGF."

reach
"Gonias and co-authors demonstrated that in MB-231 breast cancer cells and in MEFs, EGF administration resulted in EGFR tyrosine phosphorylation and ERK/MAP kinase activation mediated by STAT5 and acco[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"For example, WNK1 is required for activation of extracellular signal-regulated kinases 5 (ERK5) by epidermal growth factor (EGF) and Mitogen-activated Protein/ERK Kinase Kinases (MEKK2/3) pathway [ xref ], and Wnt/β-catenin signaling [ xref ]."

reach
"These data suggest that coactivation of both receptors enhances the release of sIL-6R from microglial cells.In DRG neurons, both hyper-IL-6 and EGF stimulation activated Stat3 and Erk1/2, but we were unable to show that IL-6-induced Stat3 activation was affected by blocking EGFR activation."

reach
"We next determined whether the stimulatory effects of EGF on activation of P70RSK and RPS6 proteins are mediated via the EGF induced ERK1/2 MAPK pathway."

reach
"In the presence of E2, EGF also induced activation of ERK1/2, and the amplitude of the ERK1/2 activation was similar (P> 0.05) to that observed in the absence of E2."

reach
"SP, PGE 2, and EGF activated Erk1/2 and stimulated proliferation of Caco-2 cells."

reach
"In humans, migration and invasion of trophoblast cells are mediated by the EGF induced ERK1/2 MAPK cell signaling cascade that stimulates downstream targets (Qiu et al., 2004a, 2004b)."

reach
"In pTr cells, EGF stimulates both the PI3K and AKT and ERK1/2 MAPK signaling cascades, and these EGF induced pathways appear to function independently in pTr cells (Jeong et al., 2013)."

sparser
"Moreover, BAPTA-AM did not suppress EGF-induced ERK1/2 activation ( xref )."

reach
"Thus, our findings indicate a central role for EGFR dependent ERK1/2 MAPK cell signaling on migration inducing effects of EGF on pLE cells.Collectively, our results provide novel insight into the mech[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Widely expressed with highest levels in bone marrow, spleen, brain, testes, and embryonic brain, Abi1 may negatively regulate cell growth and transformation by interacting with the nonreceptor tyrosine kinases ABL1 and/or ABL2, thus regulating EGF induced Erk pathway activation and EGFR signaling."

reach
"However, ethanol did not potentiate EGF stimulated p42/44 MAPK activation in embryonic liver cells (Reddy and Shukla, 1996) ."

reach
"In addition, TGFalpha and EGF have been reported to transiently activate ERK1/2, but this transient activation is not sufficient to induce beta-cell proliferation."

sparser
"RNAi studies showed that MLK3 can promote microtubule instability, is required for cell proliferation in epithelial and fibroblast cells, and is required for TNF and EGF activation of extracellular signal-regulated kinase (ERK) and JNK [ xref , xref ]."

reach
"According to this model, Rabex5 and Rabaptin5 are tightly bound, and the complex does not dissociate after Rab5 activation, whereas binding of SPIN90 and Gapex5 is dependent on the phosphorylation status of SPIN90, as determined by EGF-induced ERK signaling."

reach
"Moreover, Rabex5 and Rabaptin5 are tightly bound, and the complex does not dissociate, even after Rab5 activation , whereas binding of SPIN90 and Gapex5 is signal-dependently regulated by the phosphorylation status of SPIN90, which is determined by EGF-induced ERK signaling."

sparser
"The modified biosensor, EKAREN5-gl showed an increased sensitivity to EGF-induced ERK activation responding to a very low dose (20 pg/ml) of EGF stimulation."

reach
"Further, EGF receptors activate the extracellular signal regulated kinase (ERK) signaling pathway, which results in increased NF-kappaB and Sp1 transcription factors, followed by MUC5AC upregulation [XREF_BIBR]."

reach
"For example, hSef overexpression attenuates EGFR degradation and promotes activity in the EGF triggered Erk pathway XREF_BIBR."

reach
"However, ethanol induced prolonged activation of EGF stimulated p42/44 MAPK activation in hepatocytes has been shown to be insensitive to PKC inhibition (Chen et al., 1998) , thus also suggesting PKC [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Huber and colleagues identified a late endosomal protein p14, which was indispensable for efficient EGF signalling in the ERK cascade [59 •• ] ."

sparser
"Both insulin and EGF activated extracellular regulated kinase 2 (ERK-2), p70rsk and protein kinase B (PKB) and inactivated glycogen synthase kinase-3 (GSK-3)."

sparser
"The ALIX-supported MVB sorting has been demonstrated to be a critical determinant in the highly transient nature (10 min) of EGF-induced ERK1/2 activation [ xref ], which reflects rapid silencing of activated EGFR before its degradation."

reach
"All of these results showed that ER-alpha36 mediated the EGF stimulated ERK activation in Hec1A cells."

reach
"Thereby, Ruk (l)/CIN85 led to a more rapid and prolonged epidermal growth factor dependent activation of Src, Akt and ERK1/2 and treatment with the Src inhibitor PP2 and the PI3K inhibitor LY294002 abolished the Ruk (l)/CIN85-dependent changes in cell motility."

reach
"Tensin4 is up-regulated by EGF induced ERK1/2 activity and promotes cell proliferation and migration in hepatocellular carcinoma."

reach
"Up-regulation of ERK signaling by EGF in the HCC cells resulted in a change to a mesenchymal cell like morphology through modulation of the actin cytoskeleton."

sparser
"Here we show that treatment of MCF-7 breast cancer cells with EGF activated Akt and ERK, induced morphological changes, and increased cell motility."

sparser
"EGF induced intense activation of pEGFR (tyr1173, tyr 1045) and pERK."

sparser
"EGCG had no effect on activation of EGFR (tyr 1173 and tyr1045), pERK and pAKT by EGF, but specifically augmented ligand-induced downregulation of total EGFR protein, whereas erlotinib substantially abrogated the activation of pEGFR, pERK and pAKT by EGF without altering the total levels of EGFR protein."

reach
"However, while epidermal growth factor, which increases proliferation of PC12 cells, activates ERK1/2 only transiently, neural growth factor, which promotes their differentiation, induces a prolonged ERK1/2 activation [89], indicating that different patterns or duration of ERK1/2 activation are specifically and differently decoded by the cells."

reach
"By contrast, the allosteric SHP2 inhibitor SHP099 effectively suppressed PDGF- or EGF stimulated ERK MAPK pathway in Swiss 3T3 cells or HEK293T cells, respectively."

sparser
"EBP50 knockdown abolished its inhibition on EGF-induced ERK activation, suggesting EBP50 regulated EGFR signaling."

reach
"Since agonist occupancy of the EGFR also activates Rho [32] and we have found that EGF treatment promotes the interaction between GRK2 and RhoA-V14 ( Fig. 4 C and 4 D), we investigated whether Rho-act[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Consistent with previously published results [10,11] , we find that overexpression of GRK2 in HEK-293 cells results in a ~ 2-fold increase in EGF-induced ERK activation, as detected by Western blottin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Taken together, these results suggest that Rho-activated ERK scaffolding by GRK2 can function to increase EGF-induced ERK activation in HEK-293 cells.Several studies have linked increased levels of GR[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, the protein expression level of c-FOS is post-transcriptionally regulated by ERK activation kinetics, where transient ERK signals induced by EGF resulted in little induction of c-FOS, whereas sustained ERK activation by PDGF induced markedly higher amounts of the same protein [XREF_BIBR, XREF_BIBR]."

reach
"Consistent with this hypothesis and in agreement with previously published data [10] , we find that overexpression of GRK2 results in increased EGF-induced ERK activation."

reach
"Previously, EGF-mediated ERK activation was shown to be potentiated in HEK-293 cells by over-expression of phosphodiesterase-γ (PDEγ), a GRK2 substrate [36] ."

sparser
"Pretreatment with AG markedly blocked ERK1/2 activation induced by EGF but did not have any effect on AII-induced ERK1/2 activation (Figure 6C)."

reach
"In light of the kinase-independent ERK scaffolding function for GRK2 reported herein, we can conclude that GRK2 is able to potentiate EGF-mediated ERK activation by both kinase-dependent and independe[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Unlike AG1478, PP2 had no effect on EGF-induced ERK1/2 activation."

reach
"Since the results presented here indicate that ERK mediates positive regulation of PDX-1 expression by EGF and metformin has recently been shown to inhibit ERK activation in PDAC cells [XREF_BIBR, XREF_BIBR, XREF_BIBR], we hypothesized that the metformin inhibits PDX-1 expression by abrogating EGF induced ERK activation."

reach
"Metformin pretreatment also inhibited EGF stimulated ERK kinase activation, as evidenced by decreased phosphorylation of ERK at Thr201 and Tyr204 (XREF_FIG), consistent with the notion that metformin inhibits PDX-1 expression through a mechanism involving inhibition of EGF- stimulated ERK signaling (XREF_FIG)."

reach
"GnRH-, PMA- and EGF induced a robust ERK1/2 activation within 5 min of stimulation in both alphaT3-1 (XREF_FIG) and LbetaT2 cells (XREF_FIG)."

reach
"We demonstrated an EGF induced ERK nuclear accumulation in SMMC-7721 and LO2 cells, which may probably be associated with Tensin4 expression."

reach
"In order to understand how EGF levels modulated ERK activity, Albeck and coworkers monitored the dynamics of an ERK phosphorylation sensor at the single-cell level using timelapse microscopy."

sparser
"13,62 Blocking of phosphatidylinositol-3-kinase activity with InsP 6 profoundly impaired JB6 cell transformation and ERK activation induced by EGF or TPA, as well as AP1 ac[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Given that metformin inhibited EGF stimulated ERK kinase activation, it is, thus, reasonable to speculate that metformin inhibits tumor cell proliferation through inhibiting EGF stimulated ERK kinase activation, leading to down-regulation of PDX-1 expression at post-translational level since ERK has kinase activity towards PDX-1 (XREF_FIG)."

reach
"For example, while over-expression of GRK2 in HEK-293 cells results in increased EGF-induced ERK activation, as previously described, ERK activation in response to ANG II , which also activates Rho, i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In gastrointestinal epithelia EGF induces MEK and ERK activation that, in turn, induces Rac1 activation."

sparser
"Moreover, we found that the EGF-induced activation of the downstream signalling proteins Erk and Akt was also inhibited by 17-AAG ( xref )."

reach
"EGF exposure resulted in a transient, significant induction of PKC (81kDa) and ERK (43kDa) activation, with maximal ~ 2-fold (p < = 0.001) and ~ 1.6-fold (p < = 0.001) increases seen at 15 and 30min, respectively (XREF_FIG); activation of the other PKC and ERK proteins remained unaffected (XREF_FIG)."

sparser
"E-cadherin, however, might promote proliferation via interaction with EGFR, resulting in EGF-dependent ERK activation in CRC ( xref – xref )."

reach
"Importantly, when used at physiologically relevant concentrations, EGF, insulin and IGF-I activated the 81kDa PKC and ERK, 116kDa PKC and ERK, and 116kDa PKC, respectively and at different times."

reach
"For example, EGF activation of ERK combines digital and analog signaling mechanisms."

reach
"Stimulation of ERK signalling by EGF, which was particularly observed at the tegument, might be important during early host invasion during which the parasite will not only sense human EGF for the first time but must rapidly adapt to the host; thus, host mediated ERK activation might drive tegument remodelling ensuring parasite survival in addition to promoting cell growth/differentiation."

reach
"Rapid and sustained nuclear cytoplasmic ERK oscillations induced by epidermal growth factor."

sparser
"Inhibition of endocytosis by expression of dominant-negative mutants of dynamin (K44A) abolished activation of MEK1/2 and ERK1/2 by EGF ( xref , xref , xref )."

reach
"However, in breast cancer cells, abolishing PRMT5 mediated EGFR Arg1175 methylation increased EGF induced extracellular signal regulated kinase (ERK) activation by preventing the recruitment of Src homology 2 domain containing protein tyrosine phosphatase (SHP) 1 to EGFR, leading to enhanced cell proliferation, migration, and invasion of EGFR overexpressing tumor cells [XREF_BIBR]."

reach
"In ARPE-19 cells, EGF stimulation of phospho-ERK1/2 was predominantly unperturbed by LCLAT1 suppression with two oligonucleotides (Figure 5, G and H)."

reach
"The treatment of the cells with the ERK activator epidermal growth factor (EGF, 200 ng/mL) decreased the protection effect of ATF6 on IR."

reach
"Results from this study show that triggering of GPR120 with EPA, DHA or AA in Caco-2 cells activated three independent intracellular signalling events; accumulation of cytosolic Ca 2+, EGF receptor- and Raf-1-dependent activation of MAP kinase ERK1/2, and EGF receptor- and Raf-1-independent inhibition of IL-1beta induced NF-kappaB activation."

sparser
"EGF-dependent ERK activation ( xref – xref ) might be a relevant pathway in CRC."

reach
"Compounds 1 and 4 displayed strong cytotoxicity against three human tumor cell lines with GI50 values in the submicromolar range, whereas 2 showed subnanomolar activity as an inhibitor of EGFR-MAPK-AP1-mediated mitogenic signaling, causing inhibition of EGF mediated AP1 trans-activation and EGF mediated ERK activation and slight inhibition of EGF mediated JNK activation."

sparser
"Braconi Quintaje et al. (1998) showed both PMA and EGF activate cardiomyocyte ERK1/2, however PKC effects were insensitive to RTK inhibitors and the EGF response appeared PKC-independent."

sparser
"For example, knockdown of IQGAP1 by siRNA abrogates the activation of MEK [ xref ] and ERK [ xref ] by epidermal growth factor."

sparser
"Activation of Erk by EGF, HGF or activated Ras can induce the phosphorylation of S/TP or PXS/TP motifs in the linker region ( Calonge and Massagué, 1999 ; Kretzschmar et al., 1997a,b, 1999 )."

sparser
"Cook S J, Rubinfeld B, Albert I, McCormick F: RapV12 antagonizes ras-dependent activation of ERK-1 and ERK- 2 by LPA and EGF in Rat-1 fibroblasts."

sparser
"To determine whether high concentrations of antiestrogens failed to activate the MAPK/ERK or inhibited the ERK activation, we examined the effects of high concentrations of ICI and 4-OHT on the ERK activation induced by EGF."

trips
"Kinase-inactive MEKK1, but not MEKK2, 3 or 4, strongly inhibited EGF-stimulated ERK activity."

sparser
"This makes sense biologically because both EGF and HRG activate ERK through Ras and Raf."

sparser
"Indeed, a concentration-dependent inhibition of the EGF-induced ERK1/2 activation by 9 was observed ( xref )."

sparser
"In this context, overexpression of E-cadherin ( xref ) or soluble E-cadherin ( xref ) can stimulate EGFR and EGF-dependent ERK activation, resulting in cancer stem cell properties such as spheroid formation."

reach
"Silencing of SPRY2 increased epidermal growth factor-induced extracellular signal-regulated kinase (ERK) and Akt activation causing premature onset of DNA replication, increased DNA damage, and impaired proliferation, suggesting that SPRY2 suppresses DNA replication stress."

reach
"HRP-3, a family member of HDGF-related proteins (HRPs), markedly prolonged EGF-mediated ERK1/2 activation and knockdown of HRP-3 was shown to enhance apoptosis of HCC cells induced by multiple chemoth[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGF treatment significantly induced ERK activity (peaked at 30 min) and significantly increased COX-2 protein (peaked at 6 hr), production of prostaglandin E2 (PGE2), and cell proliferation."

reach
"EGF and TNFalpha ligands stimulate ERK, JNK and p38 MAPK cascades, the PI3K and AKT pathways, and the NFkB cascade."

reach
"ERK1/2 MAPK signalling in cardiac fibroblasts is activated by TGF-β, PDGF, and EGF and has been found to be responsible for fibrosis [133]."

reach
"Growth factors like fibroblast growth factor and epidermal growth factor activates ERK and Akt signaling pathways, which in turn induce keratinocytes motility and causes re-epithelialization in wound (Seeger and Paller, 2015, Perrone et al., 2018)."

sparser
"These findings are consistent with our data that EGF stimulation causes PI3K-independent activation of ERK in BT-474."

sparser
"We confirmed their observation in RH7777 cells as well ( Fig. 2 A ), in which EGF-induced ERK activation was lower in GPR4-transfected cells than vector-transfected cells at a physiological pH of 7.4."

sparser
"Chrysophanol was observed to suppress EGF-induced activation of ERK1/2 in SNU-C5 cells ( xref )."

sparser
"Thus, EGF-induced ERK activation is dependent on extracellular pH."

sparser
"As expected, EGF induced activation of both ERK1/2 and Akt as previously reported [ xref ]."

sparser
"Actually, cAMP-increasing agents, forskolin and isoproterenol, inhibited the basal and EGF-induced ERK activation in vector cells ( Fig. 2 B)."

sparser
"EGF‐induced activation of ERK and AKT is significantly increased in AEG‐1‐L24K/L25H compared with AEG‐1‐WT (Fig. xref )."

reach
"It was reported that EGF activates ERKs mainly through the Grb-2-SOS-Ras-Raf-MEK-ERK pathway [XREF_BIBR]."

reach
"It has been reported that EGF stimulation of ERK is enhanced in Rat-1 cells treated with MbetaCD XREF_BIBR, but in contrast MbetaCD treatment alone has been shown to suppress ERK activation in human breast cancer cells XREF_BIBR."

reach
"Given that human EGF, insulin and IGF-I stimulated ERK and PKC activities in somules in vitro we wished to establish the importance of ERK and PKC activity to somule phenotype."

sparser
"EGF and HB-EGF activate EGFR, ERBB4, AKT and extracellular signal-regulated kinase 1/2 in EGFR(+) primary trophoblasts; however, they do not show an effect on HLA-G(+) EVTs."

sparser
"Moreover, neither pre-incubation of cultures with the selective EGFR antagonist, AG1478, nor down-regulation of the EGFR to a point where EGF could no longer activate ERKs had an inhibitory effect on ERK activation by DSLET."

sparser
"The stimulation of cAMP accumulation and inhibition of EGF-induced ERK activation in GPR4-expressing cells were attenuated by decreasing the extracellular proton concentrations."

sparser
"The pH-dependent cAMP accumulation may mediate the ligand-independent inhibition of ERK activation, as evidenced by the inhibition of EGF-induced ERK activation by cAMP-increasing agents ( Fig. 2 B)."

reach
"Taken together, these data indicate that nucleobindin-2 regulates EGF stimulated MAPK kinase and Erk signaling, cell proliferation, and adipocyte differentiation."

reach
"Overexpression of the SH3 domains of ITSN1 inhibits EGF activation of Ras and extracellular signal regulated kinase (ERK), suggesting that ITSN1 might activate the ERK pathway."

sparser
"Knockdown of p14 partially reduced the extent of EGF-induced ERK1/2 activation as revealed by western blotting and immunofluorescence staining with the phospho-ERK1/2 (pERK1/2) antibody ( xref )."

reach
"It was reported that EGF-induced ERK1/2 activation occurs mainly through EGFR activation in porcine renal cell (Dong et al., 2004)."

sparser
"Silencing QKI in the U343 glioblastoma cell line, results in miR-7 expression and cell cycle arrest, through a mechanism involving miR-7 negative regulation of epidermal growth factor (EGF) receptor (EGFR) protein levels, thus blunting the EGF-dependent ERK activation (Wang et al., xref )."

sparser
"Reminiscent of the distinct phenotypic outcomes observed in PC12 cells as a result of acute or sustained Erk activation by EGF or nerve growth factor (NGF) respectively xref , the authors showed that osteogenic differentiation of hMSCs was governed exclusively by differential activation of the PI3K pathway."

reach
"For measuring EGF-mediated ERK reporter activities, the EGFR-transfected cells were treated with (1) the vehicle only as the control; (2) EGF only at 20 ng/mL; and (3) each of the five compounds at 10 μM or erlotinib at 1 μM in the presence of EGF at 20 ng/mL."

sparser
"As one of the nuclear targets of Erk is Elk-1, a transcription factor of the Ets family, FGF- or EGF-induced activation of Erk can be monitored by measuring the rate of Elk-1-dependent transcription."

reach
"Therefore, the effects of the five compounds on FGF23- and EGF-mediated ERK reporter activities were measured."

sparser
"These data suggested that the SPR domain of Spred-1 is responsible for the effective suppression of FGF- and EGF-induced Erk activation."

reach
"None of the five compounds exhibits EGFR tyrosine kinase inhibition and disrupts EGF/EGFR interaction and EGF-mediated ERK activation (Figure 5c)."

reach
"In contrast, erlotinib, a small-molecule EGFR tyrosine kinase inhibitor, completely abolishes EGF-mediated ERK reporter activities (Figure 5c)."

reach
"Both GAS6 and EGF stimulated downstream PI3K/AKT and MAPK/ERK signaling in parental cells, but only GAS6 activated these pathways in OSI-resistant (OSIR) derivative cell lines."

sparser
"We reported that mammalian Sproutys selectively inhibit FGF-induced Erk activation but do not inhibit EGF-induced Erk activation [20] ."

sparser
"However, transient ERK1/2 activation induced by epidermal growth factor could not mimic the gene profile of the senescent hHSCs."

sparser
"c-Jun N-terminal kinase (JNK), extracellular signal-regulated kinase (ERK) and P38 were activated by EGF in BEAS-2B cells ( xref )."

sparser
"Primary monolayer cultures of noncystic and cystic murine collecting duct principal cells were used to identify aberrant EGF-dependent ERK1/2 activation and regulation of Na+ transport associated with autosomal recessive PKD."

reach
"As shown in XREF_FIG, a 3.5-fold increase of ERK1/2 phosphorylated Exo70 was detected in cells treated with EGF (see XREF_SUPPLEMENTARY for all the statistical analyses of the binding results shown in XREF_FIG)."

reach
"Use of suboptimal EGF concentrations revealed the potentiating action of PTHRP on EGF induced ERK activation more clearly (XREF_FIG)."

reach
"Conversely, the SHOC2 (M173I)-tRFP mutant expressed to a similar level as WT SHOC2-tRFP was only able to restore ~ 10% above the basal level of EGF induced ERK1/2 activity (XREF_FIG, lanes 7-9 and XREF_FIG)."

reach
"Conversely, potentiation of EGF induced activation of ERKs was also dependent on the concentration of PTHRP administered (XREF_FIG)."

reach
"In comparison, EGF induced motility of the same undifferentiated keratinocytes requires the previously described extracellular signal regulated kinase to the M-calpain pathway."

reach
"Thus, although PTHRP induced mitogenesis may occur through an ERK independent mechanism, the ability of PTHRP to potentiate the activation of ERK by EGF and IGF-1, together with the ability of cells to respond to autocrine PTHRP, could provide one explanation for the increased mitogenic responsiveness and reduced doubling times of MCF-7 cells overexpressing the PTHRP-R."

reach
"While EGF/Spitz activates the RAS/extracellular signal-regulated kinase (ERK) pathway through the EGF receptor (EGFR) in an autocrine manner, the RAS/ERK pathway promotes cell migration by repressing [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGF, an EGFR ligand, efficiently activated ERK1/2 while binding of the anti-EGFR antibody to the extracellular domain of EGFR inhibited EGFR/ERK1/2 activation by occluding ligand binding [XREF_BIBR]."

sparser
"Epidermal growth factor (EGF) activated ERK and p38MAPK but not JNK."

reach
"In this study, we used Luminex multiplexing assays and high-content single-cell imaging to demonstrate that EGF triggers temporal activation of extracellular-signal-regulated kinases 1/2 (ERK1/2) in Hs578T cells."

reach
"In contrast, treatment of PC-12 cells with NGF promotes a very strong and sustained ERK1/2 activation that leads to cell cycle arrest and terminal differentiation; in this case epidermal growth factor[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The effects of ZM323881 on LH responses were attributable to specific inhibition of VEGFR2 because although ZM323881 simultaneously and efficiently prevented the activation of ERK1/2 by VEGF it had no[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Moreover, EGF, which is known to activate RAS, caused the dissociation of AK2 from BRAF and triggered concomitant activation of ERK at the same time points (Fig. 6D)."

sparser
"Both TNF and exogeneous C6-ceramide interfered with the mitogenic activation of Raf-1 and ERK by epidermal growth factor and down-regulated v-Src-induced Raf-1 kinase activity."

reach
"We found that HER2 mediated effects on EGFR dimerization and trafficking were sufficient to explain the observed HER2 mediated amplification of epidermal growth factor induced ERK signaling."

reach
"EGF dependent activation of Erk1/2 and Akt pathways may regulate cell cycle progression through control of p27 protein levels or by causing p27 to be sequestered away from its nuclear site of action [[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Indeed, such a scenario is supported by experiments in which the expression of the isolated PH domain of Gab1 suppressed EGF induced ERK and AKT activation in breast cancer cell lines [XREF_BIBR]."

reach
"XREF_BIBR These observations are in agreement with previous reports conducted both invivo and invitro that have indicated that BMP-4 inhibits EGF stimulated ERK activation in isolated parietal cells XREF_BIBR and that prolonged overexpression of growth factors in the stomach of mice alters the normal architecture of the gastric mucosa, leading to increased ERK activation, loss of parietal cells, and foveolar hyperplasia."

reach
"Our data showed that the initial EGF mediated activation of the ERK and MAPK pathway is similar in both cell lines."

reach
"Mitogenesis in response to EGF can not be studied in COS-7 cells as they are partially transformed, but it is known that picomolar EGF doses are able to activate the Ras and extracellular signal regulated kinase (ERK) signaling cascade, the central driver of cell proliferation in a PI3K dependent mode, in this cell type XREF_BIBR."

reach
"Afatinib had little effect on the phosphorylation of EGFR receptors tested or on downstream kinase activity caused by EGF stimulation on either cell line with the exception of ERK1/2 in the MCF-7 cell[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In monolayer culture, overexpression of Gab2 accelerated EGF induced cell cycle progression and was associated with enhanced and/or more sustained EGF induced ERK and AKT activation."

reach
"The EGF dependent activation of Erk1/2 was unaffected by LY294002 (data not shown)."

reach
"Furthermore, picomolar concentrations of EGF activate selectively ERK and PI3K and Akt pathways while PLCgamma, which produces IP3 and triggers Ca 2+ store release, is activated only by nanomolar EGF concentrations XREF_BIBR."

reach
"Since our data show that EGF activates Erk1/2 in high-density cells but they do not divide, Erk1/2 activation by itself is not sufficient to lower p27 below the level critical to allow proliferation."

reach
"EGF‐like factors and EGF receptors are upstream activators of ERK1/2."

sparser
"EGF Activates ERK and p38MAPK but Not JNK."

reach
"ERK can be activated by EGF concentrations as low as 2 pM and 40 pM, resulting in proliferation of 8% and 55% of the cells respectively XREF_BIBR."

reach
"We controlled that gefitinib was able to inhibit EGF-induced Erk activation (fig."

reach
"It was shown that EGF stimulation causes a strong but transient activation of the ERK cascade, while NGF causes an equally strong but sustained activation; 18 and both effects were shown to be dependent on the activity of ERKs."

sparser
"Phosphorylated ERK1/2, which was activated by EGF, promoted LF proliferation by accelerating cell-cycle progression from the G1 to S phase."

reach
"These results indicated that CK2α-mediated PGK1 phosphorylation promotes the binding of PGK1 to CDC7.It was reported that EGF-induced activation of extracellular signal-regulated kinase (ERK) results [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In this effort, we were able to show that EGF stimulation of EGF-free DU145 sphere cells resulted in ERK activation, but not STAT3 activation, in a time dependent manner (XREF_SUPPLEMENTARY)."

reach
"Ectopic overexpression of Sprouty2 inhibits fibroblast growth factor (FGF) and vascular endothelial growth factor (VEGF), but not epidermal growth factor- (EGF) induced ERK activation XREF_BIBR."

sparser
"EGF, which activates ERK and p38MAPK, but not JNK, ameliorates CyA-induced nuclear translocation of R-Smad and apoptosis through activation of p38MAPK in RPTCs."

reach
"Studies in T47D breast cancer cells have shown that EGF treatment induces Erk dependent PRB phosphorylation on serine residues [XREF_BIBR]."

reach
"Although NS5A inhibits MAPK and ERK activation induced by exogenous EGF, ORF3 was described as an activator of MAPK and ERK pathway like hPIV3-C."

reach
"Insulin regulated activation of AKT and EGF regulated activation of ERK1/2 is not affected by L-glutamine pretreatment (XREF_FIG)."

reach
"As expected, EGFRi but not ErbB2i abrogated EGF induced ERK activation, validating the specificity of the inhibitors."

reach
"In all the three cell lines, EGF stimulation increased P-ERK together with P-FAKSer732 and P-CDK5Tyr15, the active kinase isoform (XREF_FIG)."

reach
"Down-regulation of GEP100 or Arf6 by specific siRNAs largely inhibited EGF induced ERK activity."

reach
"Taken together, these experiments demonstrate that uPAR is a downstream target of GEP100 and Arf6 whose expression is required for EGF induced MDA-MB-231 cell invasion.Previous reports have shown that[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"It has been proposed that the transient activation of Ras and ERK by EGF in PC12 cells may be due to attenuation of the signal at or near the level of the EGFR [22,28] ."

reach
"Effects of EGF are maximal at 5 minutes and wane by 1 hour; EGF activates ERK and AKT with similar kinetics."

reach
"Extracellular signal-controlled kinase (ERK) signals demonstrate ERK1/2 activation is strongly induced by EGF [60] ."

reach
"In agreement with these findings, our results showed that EGF induced a time dependent increase in ERK activity."

reach
"There is ample evidence for the essential role of Ras in the EGF mediated ERK activation."

reach
"This observation agrees with the previous report showing that Ras isoforms are redundant for EGF activated ERK activation and that HRas is a minor isoform in HeLa cells compared with KRas and NRas."

reach
"Because a PKC inhibitor Go6983 did not inhibit EGF induced ERK activation, we speculated that a PLC mediated increase in Ca 2+ influx enhanced the Ras mediated ERK activation in a PKC independent manner."

reach
"Furthermore, inhibition GEP100 or Arf6 activity by siRNA blocked EGF induced ERK activation."

reach
"Therefore, it is reasonable to speculate that EGF induced ERK activation is indispensible for GEP100 and Arf6 mediated breast cancer cell invasion.Signals mediated by uPAR have been shown to play crit[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, siRNA mediated down-regulation of uPAR in MDA-MB-231 cells did not alter EGF induced ERK activation."

reach
"TU-100 and ginseng and to a lesser extent ginger or 6-gingerol, but not Japanese pepper, ginsenoside Rb1 or compound K inhibited EGF activation of ERK1/2."

sparser
"Moreover, treatment of cells with epidermal growth factor (EGF) that activates MEK and ERK, as well as genetic ERK activation ( xref ) by a constitutively active MEK (MEK CA ) ( xref ), increased T562 phosphorylation levels; pharmacological or genetic MEK inhibition using the selective U0126 inhibitor or by expressing a dominant negative MEK mutant (MEK DN ); and reduced T562 phosphorylation; finally, in Mfn1 −/− cells re-expressing the MFN1 T562A mutant, phosphorylation was undetectable even in the presence of EGF ( xref G)."

sparser
"Interestingly, although wortmannin had no effect on leptin- or H 2 O 2 -induced ERK phosphorylation, it partially attenuated EGF-induced ERK phosphorylation ( Fig. 3 C), suggesting that PI3K is involv[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"On the GF-RTK side, treatment with AG1478, a small-molecule inhibitor of EGFR tyrosine kinase activity, significantly decreased ERK activation by EGF or HRG (to levels close to unstimulated cells) in all conditions tested."

reach
"After light irradiation, ERK activation was induced by bath application of 50 ng/mL EGF at the final concentration."

reach
"Therefore, activation of glucocorticoid receptor reduces the amplitude and markedly shortens the duration of EGF-induced ERK activation by intercepting the RTK-to-ERK signalling at multiple levels [47[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In contrast to androgen signaling, however, phosphoserine-paxillin is not required for EGF induced nuclear localization of Erk, and does not appear to bind to Erk dependent promoters [XREF_BIBR]."

reach
"EGF and TNFalpha ligands stimulate ERK, JNK and p38 MAPK cascades, the PI3K and AKT pathways, and the NFkappaB cascade."

reach
"We have shown that inhibition of PLC suppressed EGF induced ERK activation to ca. 50% of the full activation (XREF_FIG, A and B)."

reach
"This is independent of alterations in receptor autophosphorylation and correlates with impaired c-Cbl phosphorylation and association with the EGFR, reduced EGFR ubiquitylation, and sustained EGF induced extracellular signal regulated kinase activation."

reach
"Interestingly, EGF-induced ERK and AKT activation increase concomitantly resulting in signaling stress with premature S-phase entry."

sparser
"While ERK1/2 expression and basal phosphorylation did not significantly differ between GHR-KO and normal mice, EGF activation of ERK1/2 was significantly diminished in GHR-KO mice ( xref and xref )."

sparser
"S1P has been shown to activate ERK in Swiss 3T3 fibroblasts, and the SphK inhibitor DHS blocked PDGF- but not EGF-induced activation of ERK, which was reversed by exogenous S1P [80] ."

sparser
"Similarly EGF, which activates ERK, increases BAK oligomerization in a Mfn1 T562 -dependent manner, placing MFN1 at the crossroad between extracellular signals and activation of the mitochondrial pathway of apoptosis."

sparser
"However, Enserink et al. reported that the Epac pathway specific cAMP analogue 8CPT-2Me-cAMP does not activate ERK, nor does it inhibit growth-factor-induced ERK activation."

sparser
"Using the setup in Fig. 1 , we previously demonstrated that a peptide (PVPE-Pmp-INQS) blocking the Grb2–SH2 domain can inhibit Erk1/2 activation by EGF when electroporated into mouse NIH3T3 fibroblas[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"These results demonstrate that the possibility for upscaling using this apparatus makes the examination of Erk1/2 activation by EGF in subconfluent NIH3T3 fibroblasts possible."

reach
"This was initially demonstrated in rat pheochromocytoma PC-12 cells, in which transient ERK activation, stimulated by epidermal growth factor (EGF), resulted in proliferation, whereas sustained ERK activation, evoked by nerve growth factor (NGF), induced cell differentiation."

reach
"Thus, it was shown that EGF stimulation causes a strong but transient activation of the ERK cascade, while NGF causes an equally strong but sustained activation [91] ; both effects were shown to be de[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For instance, mutant p53 (p53-273H) suppresses miR-27a expression by specifically binding to the miR-27a promoter region (nts -2899 to -2675) and subsequently enhances EGF mediated ERK1/2 activation in breast and lung cancer cells [XREF_BIBR]."

reach
"In particular, we use a right combination of multiplexing via fluorescent cell barcoding (FCB) and intracellular antibody staining for simultaneous detection of phospho-ERK1/2 (pERK) stimulated by epidermal growth factor (EGF) in multiple samples."

reach
"CTTN expression maintained EGF induced ERK activation."

reach
"We show that overexpression of HRAS and other constituents of the RAS-ERK pathway leads to the prolongation of EGF induced ERK activation and that this is sufficient to transform the EGF signal into a differentiating stimulus."

sparser
"Further analysis revealed the requirement of CNKSR2 for NGF but not EGF-induced activation of ERK [ xref ]."

reach
"RWPE1 cells are grown in media containing recombinant epidermal growth factor (EGF), which activates RAS and ERK signaling, but because the RAS and ERK pathway is intact, negative feedback loops can keep ERK activity low [XREF_BIBR]."

sparser
"Immunofluorescence revealed that EGF-activated EGFR, MEK1/2 and ERK1/2 co-localize on endosomes."

sparser
"Further investigations demonstrated in the TRAF2-shRNA NPC cells found that EGF-induced activation of ERK1/2 significantly decreased."

sparser
"For instance, epidermal growth factor-induced transient ERK activation via the Raf-MEK-ERK axis induces proliferation in PC12 neuroendocrine cells, whereas nerve growth factor-induced prolonged ERK activation via the Raf-MEK-ERK axis leads to the differentiation of PC12 cells into sympathetic neuron-like cells [ xref , xref ]."

sparser
"Interestingly, prolonged ERK activation by epidermal growth factor in PC12 cells overexpressing the epidermal growth factor receptor switches cell fate from proliferation to differentiation [ xref ]."

reach
"In particular, ERK1/2 signaling induced by EGF in PC-12 cells resulted in proliferation [17], whereas ERK1/2 signaling induced by nerve growth factor (NGF) resulted in differentiation [18]."

reach
"The activations of MEK1/2, ERK1/2 and p70S6K by EGF are key elements in the classical MAPK signalling cascade EGF --> ERBB1/1 --> GRB2 --> SOS1 --> RAS --> RAF1 --> MEK1/2 --> ERK1/2 --> p70S6K."

reach
"EGF treatment induced ERK activation as well as the rapid assembly of SHP2, EGFR, and Gab1."

sparser
"Importantly, control experiments show that this mutant retains the ability to mediate EGF-dependent activation of Erk and induction of an Elk reporter."

reach
"EGF was shown to increase time-dependently the activation of extracellular signal regulated kinase (ERK) and c-Jun amino terminal kinase (JNK) in A431 cells [79]."

sparser
"EGF can activate the intracellular MAP kinase signaling molecules ERK1/2, which are themselves phosphorylated in response to injury [ xref ]."

reach
"34 SCCs exhibit enhanced secretion of several cytokines, such as EGF, which induce ERK activation via a ras dependent signaling mechanism."

sparser
"Both neural growth factor (NGF) and epidermal growth factor (EGF) activate ERK in PC12 cells (a cell line derived from rat pheochromocytoma)."

sparser
"Quantification of EGF-induced ERK activation in individual cells and in a population of cells confirmed the suitability of electric stimulation for ERK activation dynamics interrogation ( xref B–S1E)."

sparser
"ERK activation induced by EGF was consistent with that reported before ( xref )."

sparser
"Otherwise, a recent study described a regulatory signalling pathway of OSCC metastasis consisting of miR-491-5p and its downstream target, GIT1, which functions as a scaffold for EGF-induced ERK activation. xref In both studies, it would be interesting to analyse whether the decrease in activity of these downstream signalling cascades could be related to EGFR inhibition by miR-491-5p."

sparser
"Therefore, ERK activation induced by EF and EGF, including the dynamics of Fc/Fn ratio are comparable to each other ( xref and xref , compared to xref and xref )."

reach
"Moreover, ERK1/2 present in the brush border are stimulated by EGF and feeding, suggesting a role in nutrient, ion transport or other brush border-associated functions."

reach
"In cooperation with integrins, the growth factors EGF, PDGF-BB, and basic FGF each produced a marked, transient activation of the ERK (extracellular signal regulated kinase) class of MAP kinase, but only if the integrins were both aggregated and occupied by ligand."

sparser
"Several mitogens, including epidermal growth factor, platelet-derived growth factor, and ROS can activate ERK1/2 [ xref ] The activation of ERK1/2 by oxidative stress is consistent with the idea that low, but adequate levels of ROS are mitogenic [ xref ]."

reach
"These findings demonstrate that activin A effectively hinders the migration of A549 cells induced by EGF through ERK1/2 signaling, suggesting that targeting activin A may hold promise in the treatment of EGF-dependent LUAD growth and metastasis."

reach
"Upon comparing the intensity of ERK activation at these sublocalizations, we found that both EGF and HRG stimulated stronger ERK activation at the cytoplasm than at the nucleus (XREF_FIG)."

sparser
"However, long-term application of U69,593 neither down-regulated EGFR nor inhibited EGF-induced ERK activation."

reach
"We defined the dynamic transcriptional effects elicited in MCF7, T-47D, and MDA-MB-436 breast cancer cell lines by nine regulators of growth and differentiation (17beta-estradiol, antiestrogens fulvestrant and tamoxifen, progestin R5020, antiprogestin RU486, all-trans-retinoic acid, epidermal growth factor, mitogen activated protein and extracellular signal regulated kinase 1/2 inhibitor U0126 and phorbol ester 12-O-tetradecanoylphorbol-13-acetate) and compared the patterns of gene regulation to published tumor expression profiles."

reach
"Pretreatment with the c-Src specific inhibitor PP2 attenuated the lysophosphatidic acid and serum mediated increase in Erk 1/2 phosphorylation but showed little effect on the EGF mediated Erk 1/2 acti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"EGF activates both ERK-MAPK and PI3K pathways, leading to the upregulation of Ugrp2 gene expression in mtCC cells."

sparser
"This is in consistent with ERK activation induced by EGF that in MCF10A cells, EGF leads to multiple stochastic pulses of ERK, whereas in PC12 cells, EGF induces one ERK activity pulse ( xref ; xref )."

reach
"Furthermore, the study revealed that EGF upregulated the ratio of p-ERK/ERK in A549 cells, which was weakened by activin A, and A549 cell migration regulated by activin A was not related to calcium signaling."

reach
"Addition of EGF dramatically increased ERK activities in all fractions, including the brush border-enriched fraction (P2)."

reach
"The p-EGFR and p-Erk1/2 were examined to see whether EGFR signaling was activated upon EGF addition and we found that addition of EGF activates EGFR and Erk1/2 signaling (XREF_FIG)."

sparser
"AACOCF3 did not influence EGF-induced ERK1/2 activation, indicating that ERK1/2 is upstream of PLA(2)."

reach
"ERK activity induced by EGF was not affected by SB203580, whereas HGF induced ERK activity was greatly increased."

reach
"In recent work XREF_BIBR, we observed similar pulses using a sensor based on phosphorylation regulated Forster resonance energy transfer (FRET) to monitor EGF stimulated ERK activity in single MCF10A cells."

sparser
"It is interesting to note that Spred-2•ΔN does not inhibit Erk activation induced by epidermal growth factor or nerve growth factor ( xref )."

reach
"We demonstrated that sustained EGF dependent activation of the MAPK and ERK kinase (MEK)/ERK pathway was involved in this blockage as specific transient inhibition of the cascade repotentiated hepatocytes to perform a new wave of replication and multiple cell cycles."

sparser
"This and previous data lead to the conclusion that EGF successively activates MEK, ERK1/2, and PLA(2), leading to an increased release of arachidonic acid."

sparser
"It profoundly inhibited the EGF-induced activation of ERK, as expected, but it also inhibited almost entirely the activation of ERK induced by Ang II ( A)."

reach
"This sequence illustrates how non uniform EGF stimulation, which is visualized in the top row of images, can trigger a traveling wave of activated RAS that spreads over the plasma membrane (XREF_SUPPLEMENTARY) and that produces a transient gradient of ERK activity within the cell, which is visualized in the bottom row of images."

sparser
"Since Ca has been implicated as a potential regulator of the ERK pathway in many cell types, two approaches were used to investigate the Ca sensitivity of ERK activation by Ang II and EGF in WB cells."

sparser
"Since the maximal ERK activation induced by EGF or Ang II could not be accounted for by PKC activation or Ca mobilization, another pathway(s) must be involved in mediating the effects of these hormone[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"EGF stimulated EGFR and Erk 1/2 activation reached a maximum at only 10 min after treatment, whereas TGFbeta 1 -stimulated activation reached a maximum at 30 min."

sparser
"In the case of EGF-induced activation of ERK, evidence points to the kinase domain of the receptor itself being responsible for the first committed tyrosine phosphorylation step."

reach
"EGF induced activation of the ERK signaling cascade has been implicated in a variety of cellular functions, including cell migration, cell cycle progression, and differentiation."

sparser
"Next, we pursued why IL-8 expression was triggered by the OA-activated KRAS/ERK axis but not the EGF-activated KRAS pathway or ERK pathway."

reach
"In contrast, for a relatively high EGF dose of 60pg/ml, even the shortest period of EGF stimulation, 30s, induces a pulse of high ERK activity (XREF_FIG)."

sparser
"To investigate the role of PKC in Ang II- and EGF-induced activation of ERK, WB cells were incubated with GF-109203X prior to the addition of EGF and Ang II."

sparser
"It is evident, therefore, that during the first hour of ERK activation by EGF and Ang II, PKC plays a minor role in the overall effects of the hormones on ERK activation."

sparser
"In contrast, the present study demonstrates that in WB cells, short-term activation of ERK by Ang II or EGF was essentially Ca independent."

sparser
"Since neither treatment affected the Ang II- or EGF-induced activation of ERK, we can conclude that the transient Ca increase was not obligatory for the early activation of ERK."

reach
"These results demonstrate that HRAS is competent for prolonging EGF induced ERK activation, and subsequently MCF-7 differentiation, only if its signals emanate from plasma membrane DM microdomains."

sparser
"The fact that neither PKC activation nor Ca mobilization could account for the observed ERK activation induced by Ang II and EGF in WB cells indicates that other pathways predominate."

sparser
"Control experiments showed that PTx had no effect on ERK activation by either VEGF or EGF (data not shown)."

sparser
"Tyrosine phosphorylation, through the intrinsic tyrosine kinase activity of the EGF receptor, is required for EGF to activate ERK, while tyrosine phosphorylation through cytosolic tyrosine kinases see[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Interestingly trastuzumab strongly inhibits EGF-induced Akt phosphorylation and slightly inhibits EGF-induced Erk activation, in both BT474 and SKBR3 cells."

sparser
"The augmented cellular endogenous forces induce cell-matrix adhesion kinase signaling, ROCK-mediated disruption of adherens junctions and growth-factor-dependent ERK activation, resulting in: (i) tumor cell proliferation, (ii) disruption of basal polarity, (iii) abrogation of glandular lumen formation, and (iv) remodeling of mammary tissue architecture [ xref , xref ]."

sparser
"21 EGF transiently activates Erk, a member of the MAPK family."

sparser
"In contrast to results with Abl, Erk activation by EGF was strongly inhibited only by Grb2 mutants; Crk and Nck mutants had little or no effect."

sparser
"This finding suggests that Grb2 is the only adapter involved in the activation of Erk by EGF."

reach
"First, we found that EGF activated ERK1/2 and Akt (data not shown) signal molecules in a time dependent manner in MCF-7 cells."

reach
"Whereas prolactin induced phosphorylation of Stat5, Erk and Akt in SKBR3 cells were suppressed at pH e of 6.8 and 6.5, Erk activation induced by EGF showed no evidence of inhibition by extracellular acidosis, compared to the response at normal pH e of 7.4."

reach
"To further confirm that EGFR was involved in Cd induced activation of ERK, the cells were transfected with siEGFR and then treated with Cd or EGF for 5 min prior to analyzing ERK phosphorylation."

reach
"The role played by Cdc42 in regulating the timing of EGF receptor-Cbl interactions is underscored by the fact that constitutively active Cdc42 (F28L), by persistently blocking the binding of Cbl to these receptors, leads to their aberrant accumulation and sustained EGF stimulated ERK activation, thus resulting in cellular transformation."

reach
"The inhibitor alone did not alter either basal or EGF mediated ERK activation (XREF_FIG, panel B)."

reach
"Anti-tumor activity of GW572016 : a dual tyrosine kinase inhibitor blocks EGF activation of EGFR and erbB2 and downstream Erk1/2 and AKT pathways."

sparser
"Since EGF activation of ERKs are independent of the PI 3-kinase pathway in NIH(M17) cells, our results indicate that the MEK-ERK- and the PI 3-kinase-dependent pathways constitute two independent sign[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"To explore the importance of the PY motif and putative ERK phosphorylation site in the acute downregulation of ENaC by EGF induced ERK1/2 activation, a stable mammalian expression system was developed."

reach
"It has been reported that the activation of ERK1/2 by EGF and EGFR is implicated in the transcription of E-cadherin repressors, such as Snail and Slug [XREF_BIBR, XREF_BIBR, XREF_BIBR]."

reach
"EGF dependent activation of ERK and serum response element (SRE) are both up-regulated in PC12 cells stably overexpressing PLC-gamma1, but knockdown of PLC-gamma1 by siRNA significantly reduces ERK activation."

reach
"Cdc42 activation inhibits the ubiquitination and degradation of EGFR; these actions result in cellular transformation due to EGFR accumulation and sustained EGF stimulated ERK activation XREF_BIBR."

reach
"Compared with the parental cells, the GNAS KO HEK293 cells had nearly complete impairment of ERK activation by Isoproterenol, while control EGF mediated activation of ERK was preserved (XREF_FIG)."

sparser
"Next, after activating Erk1/2 by EGF, we found that there was no significant change of LC3 puncta accumulation between TAIII- or TAIII plus EGF- treated cells ( Fig. 7 E, F and G), but the apoptosis r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Conversely, anastellin had no significant effect on EGF-stimulated Ras activation which was consistent with control experiments showing that anastellin did not inhibit the activation of ERK by EGF (data not shown)."

reach
"Stimulation of EGF receptor couples with the activation of PI 3-kinase in various cells [32], and EGF stimulates ERK in GH3 cells [33]."

reach
"Congruently, PD98059 increased the turnover rate of CSN6 in the presence of the de novo protein synthesis inhibitor cycloheximide (XREF_FIG), whereas EGF, which induced ERK activation, reduced CSN6 turnover (XREF_FIG)."

reach
"PKCzeta activity is required for EGF induced extracellular signal regulated kinase (ERK) activation in both normal human adult epidermal keratinocytes and five of seven SCCHN cell lines."

reach
"We have previously demonstrated by using monolayer cultures [13] that pGC contain immunodetectable ERK1 and ERK2 (p44 mapk and p42 mapk, respectively) and that EGF, in a time- and concentration depend[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"It was found that EGF stimulation actually results in a transient activation of ERK, while NGF elicits a sustained ERK activation, which leads to a neuronal differentiation program."

sparser
"Once this binding is disrupted, ERK activation induced by EGF is inhibited [ 39 ]."

reach
"Activation was maintained through the 10 ng/ml dose of the growth factor.In the last series of experiments, we investigated the time course for EGF activation of ERK in the freshly isolated pGC."

reach
"Activation was confirmed by demonstrating that EGF treatment in these cells increased the ability of ERKs to phosphorylate a substrate."

reach
"This EGF induced activation of ERKs occurred within 2.5 min of treatment, continued through 10 min of exposure, and was maximal with 10 ng/ml of EGF [13]."

reach
"Thus, as seen in primary monolayer cultures [13], the EGF induced activation of ERKs in freshly isolated pGC is sensitive and occurs rapidly."

reach
"For example, heterologous expression of ENaC and the EGF receptor in CHO cells reconstituted a response in which EGF induced inhibition of ENaC activity did not require ERK activation."

reach
"Preliminary results form our laboratory have also shown that Raf kinase activity in primary cultures of pGC is markedly increased by EGF treatment, suggesting that EGF stimulates ERKs in pGC through a[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Commensurate with the typical EGF induced ERK activation and BRAF/RAF1 heterodimerization kinetics [ xref ], Nluc activity was highest at 5 min and then gradually declined over the next 55 min (Fig.  xref G)."

sparser
"Both cause SHP2 catalytically inactive and significantly inhibit EGF induced ERK activation [ 78 ]."

reach
"In the studies presented herein that used freshly isolated pGCs, EGF induced activation of ERK occurred within 1 min of treatment, continued through 60 min, and began to decline thereafter."

reach
"EGF dependent activation of Ca 2+ influx as well as PLC-gamma, Shc, and ERK MAPK was normal in both CAML negative and CAML positive cells, however."

sparser
"In HEK293 and NIH3T3 cells, it significantly blocked SHP2 dependent ERK activation induced by EGF."

sparser
"Notably, it enhanced paxillin phosphorylation and attenuated ERK activation induced by EGF but did not affect phorbol 12-myristate 13-acetate (PMA) triggered ERK activation (SHP2 independent), suggest[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The chemotaxis assay was performed as previously described (Rabinovitz et al., 1999) with 0.1 and 10 ng/ml EGF.The assay for EGF induced ERK MAPK activity was performed as previously described (Karnit[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We also noticed both BAPTA/AM and thapsigargin partially blocked EGF induced ERK1/2 phosporylation in DU145 cells but not PC3 cells."

reach
"During C. elegans vulval development, an EGF signal from the anchor cell initiates the activation of Ras LET-60> Raf LIN-45> MEK MEK-2> ERK MPK-1 signaling cascade to induce the 1degrees cell."

reach
"Whilst even repetitive EGF stimulation cannot trigger a sustained ERK response, pulsing at the optimal frequency maximises the integral under the ERK response curve, thus suggesting that the cumulativ[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"NSCs from GD3S-KO mice showed decreased self-renewal ability compared with those from the wild-type animals, and that decreased ability was accompanied by reduced expression of EGF receptor (EGFR) and an increased degradation rate of EGFR and EGF induced ERK signaling."

reach
"To this end, we assessed the ability of wild-type and mutant SHP2 enzymes to dephosphorylate Paxillin and pY118, a physiological substrate of SHP2 required for EGF stimulated ERK1/2 activation."

reach
"As expected, knock-down of pp32 protein resulted in enhancement of MEK and ERK activation induced by EGF ( Fig. 4 )."

sparser
"A volume of 8 μM sorafenib was used to induce cellular senescence, and ERK was activated by 30 ng/ml ERK agonist EGF."

reach
"In PC12 cells, EGF stimulates a rapid but transient activation of ERK, peaking at 5 mins and returning to basal levels at 30 mins, leading to cellular proliferation [XREF_BIBR, XREF_BIBR]."

sparser
"Use of suboptimal EGF concentrations revealed the potentiating action of PTHRP on EGF-induced ERK activation more clearly ( xref )."

sparser
"However, both in cell lines in which cAMP inhibits growth-factor-induced ERK activation and in which cAMP activates ERK, 8CPT-2Me-cAMP did not affect ERK activity."

reach
"Interestingly, siRNA mediated knockdown of paxillin expression in androgen dependent LnCAP cells as well as in androgen independent PC3 cells abrogates DHT- and/or EGF induced Erk signaling."

reach
"Consistent with the previous results using RKIP-/- MEFs, EGF stimulated ERK activation remained severely compromised (XREF_FIG)."

reach
"EGF stimulates the rapid and transient activation of ERK that is required for proliferation."

reach
"EGF stimulation of ERK signaling in RKIP-/- MEFs is comparable to that of RKIP wild-type MEFs."

reach
"This suggestion is supported by results of Stefanovsky et al. (2001) showing that epidermal growth factor induces ERK dependent activation of ribosomal gene transcription."

sparser
"EGF-induced activation of ERK1/2 and Aurora A kinase was responsible for CPEB1 phosphorylation."

reach
"On the other hand, EGF triggered a brief ERK activation, insufficient to induce nuclear translocation and consequent neuronal differentiation [22, 41, 44, 114]."

sparser
"Conversely, potentiation of EGF-induced activation of ERKs was also dependent on the concentration of PTHRP administered ( xref )."

reach
"Growth factors such as epidermal growth factor (EGF) activate this pathway through receptor tyrosine kinases, enabling ERK to enter the nucleus and phosphorylate transcription factors [7]."

reach
"Furthermore, specific downregulation of Rac1 signaling in HepG2 cells did not alter EGF induced activation of ERK."

reach
"This interaction modulates EGF-dependent activation of MEK and ERK in MCF-7 human breast epithelial cells."

reach
"To investigate the role of protein kinase C (PKC) isozymes in epithelial growth factor (EGF)-induced activation of extracellular signal regulated kinase (ERK) and cell proliferation in cultured human corneal epithelial cells."

reach
"To determine the effect of TNQ on EGF-activated ERK1/2 and Akt signals, Western blot analysis with antibodies specific to phospho-ERK1/2 and phospho-Akt was performed."

sparser
"Thus, although PTHRP-induced mitogenesis may occur through an ERK-independent mechanism, the ability of PTHRP to potentiate the activation of ERK by EGF and IGF-1, together with the ability of cells to respond to autocrine PTHRP, could provide one explanation for the increased mitogenic responsiveness and reduced doubling times of MCF-7 cells overexpressing the PTHRP-R."

reach
"In addition, EGF, which activates ERK1/2, inhibited the downregulatory effect of shikonin on SIRT2 protein expression as well as the effects on proliferation and invasion of SW480 cells."

reach
"The study by Orton et al. was focused on investigating what effects different cancerous alterations exert on signaling through the EGF activated ERK pathway [68] ."

reach
"However, at 48 and 72 h, on RADA, RADA + RGD2 and RADA + RGD2 + HB peptide gels, tethered EGF induced statistically higher ERK activation than soluble EGF."

reach
"As activation of ERK increases its nuclear localization, we next investigated whether exposure to EGF enhanced P-ERK and LC3-II colocalization in the nucleus."

sparser
"We have shown that loss of Sprouty2 resulted in enhanced FGF- but not EGF-induced ERK activation in mouse embryonic fibroblasts (MEFs) [9] ."

sparser
"EGF induced weaker ERK activation in Sprouty4 KO than WT MEFs ( Fig. 3 B)."

reach
"Prior phosphorylation of ERK was not required for ERK to colocalize with LC3-II because U0126-pretreatment of NIH/3T3 cells did not decrease EGF induced ERK and LC3-II colocalization (XREF_SUPPLEMENTARY)."

reach
"XIA W, MULLIN RJ, KEITH BR et al. : Antitumor activity of GW572016 : a dual tyrosine kinase inhibitor blocks EGF activation of EGFR and erbB2 and downstream Erk1/2 and AKT pathways."

reach
"Tethered EGF induced higher levels of activation of EGFR and ERK1/2 on various peptide gel compositions at later time-points compared to soluble EGF."

reach
"In Drosophila, increased EGF signaling activates ERK to promote sleep."

reach
"Functionally we show that EGF mediated ERK1/2 translocation and activation in focal adhesions is required for cell spreading and migration."

reach
"17β-Estradiol stimulates the secretion of heparin-binding epidermal growth factor (HB-EGF) following activation of estrogen receptors, which in turn transactivates EGFR and leads to activation of extr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Ectopic expression GEP100 siRNA, GEP100-^ PH, or Arf6-T27N suppressed EGF induced ERK and Rac1 activity."

reach
"To test whether inhibition of endocytosis affects EGF-induced ERK1/2 activity, the effects of siRNA knockdown of clathrin heavy chain (CHC) and a small-molecule Dyngo-4a, treatments known to inhibit clathrin-mediated endocytosis of EGFR (Huang et al., 2004; Pinilla-Macua and Sorkin, 2015), on the time course of ERK1/2 activation were tested."

reach
"Despite the reduced amount of c-Raf and Erk2 in the cholesterol depleted caveolae fraction, EGF caused a hyperactivation of the remaining caveolae Erk isoenzymes."

sparser
"CXCL12–CXCR4 interaction elicits G protein signaling to activate Ras and PI3K and downstream ERK and Akt, which can also be activated by EGF-EGFR."

reach
"This is reinforced by the inhibition of clathrin-mediated endocytosis using two independent approaches, which did not significantly affect the time course of EGF-induced ERK1/2 activity (Fig. 8) and which confirmed that EGFR endocytosis is not an essential process for EGF-induced ERK1/2 activation in HeLa cells."

reach
"To validate the putative ERK-mediated phosphorylation site in PFAS, we performed tandem mass spectrometry (MS/MS) analyses of PFAS immunopurified from EGF-stimulated cells in the presence or absence of an ERK inhibitor to identify potential ERK-regulated phosphorylation sites."

reach
"Based on this information, we propose a model to explain the role of the ERK1/2 CC domain in EGF mediated ERK1/2 translocation and activation in focal adhesions."

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"GEP100 and Arf6 Is Required for Epidermal Growth Factor Induced ERK and Rac1 Signaling and Cell Migration in Human Hepatoma HepG2 Cells."

reach
"It would appear that, unlike cultures with soluble EGF on collagen I or peptide gels, DNA synthesis is stimulated via tethered EGF activating EGFR and ERK1/2 without altering levels of EGFR or ErbB2."

reach
"Importantly, the kinetics of ERK1/2 activation by EGF (4 ng/ml) in gene-edited cells was essentially similar to that in parental cells (Figs."

reach
"EGF also increased ERK and Rac1 activity."

reach
"DHT additionally upregulated the levels of phosphorylation of EGFR (pEGFR) and its downstream proteins AKT (pAKT) and ERK1/2 (pERK), induced by EGF treatment, in AR positive cells."

reach
"As shown in Fig. 2 A and D (upper panels), NTHi and EGF synergistically induced activation of p38 and ERK."

reach
"Sos1 and Sos2 have similar molecular structures, and both proteins mediate EGF induced Erk activation."

reach
"However, it has been shown that Sos1 participates in both short- and long-term Erk activation by EGF treatment, while Sos2 dependent signals are predominantly short-term [XREF_BIBR]."

reach
"In contrast, when EGF is used to elicit transient ERK activation, CHX has little or no effect (supplemental Fig. 2)."

sparser
"In this study, EGF activation of ERK1/2 and of expansion-related transcripts were not affected by inhibiting TrkA, indicated that TrkA is not involved in EGF receptor-mediated ERK activation."

sparser
"On the other hand, LL-Z1640-2, a natural compound isolated as an anti-protozoan (Ellestad et al., 1978) , has been reported to inhibit anisomycin-induced JNK and p38 activation but not epidermal grow[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In contrast, ERK MAPK is strongly activated by growth factors such as PDGF, hepatocyte growth factor (HGF), epidermal growth factor (EGF), transforming growth factor (TGF) α and other stimuli that med[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The results showed that both GEP100 siRNA and GEP100-^ PH largely inhibited EGF induced ERK activity (XREF_FIG)."

reach
"44 , 45 Furthermore, a recent study demonstrated that EGF stimulation in cells overexpressing EGFR resulted in more effective ERK activation in the presence of BRAFi."

reach
"As shown in Fig. 4 A and B, overexpression of dominant negative mutant form of PAK2 enhanced synergistic activation of p38 or ERK in HM3 cells treated with NTHi and EGF, whereas overexpression of domi[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In vascular smooth muscle cells, angiotensin II-induced tyrosine phosphorylation of two major proteins (p120, p70) and ERK activation are markedly reduced by α-tocopherol, whereas ERK activation by ep[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"On the other hand, EGF stimulates a transient ERK1/2 response in PC12 cells, leading to proliferation."

sparser
"Sucrose and PAO also inhibited EGFinduced internalization of the EGF-R in C9 cells, and the inability of these agents to impair EGF-induced ERK activation suggested that the latter is also independent of receptor endocytosis."

reach
"Most of them induced a reduction in ERK and AKT activation by EGF, a marker of RAS activity."

sparser
"It was shown that activation of ERK1/2 and Snail by EGF in SiHa cells was prevented by SkQ1 suggesting that EGFR signaling critically depends on mtROS."

sparser
"The present findings fit into this model of stimulus specificity since the NGF-induced sustained ERK activity and neurite outgrowth but not the EGF-induced ERK activation and mitogenic response were s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In Ca-Ski cells EGF-induced ERK1/2 activation was not affected by SkQ1 in contrast to SiHa cells."

reach
"EGF stimulated extracellular signal regulated kinase (ERK) and stress activated protein kinase and c-Jun N-terminal kinase (JNK)."

sparser
"In C3H10T1/2 cells, EGF activates Mek-Erk and stimulates AhR induction of CYP1B1 by TCDD ( Hanlon et al., 2005b )."

reach
"We therefore examined the relationship between SMYD3 and EGF stimulated ERK1/2 activation."

sparser
"Similarly, Kim et al. found that phosphorylation of the Nicastrin subunit by EGF-activation of ERK1/2 reduces γ-secretase activity ( xref )."

reach
"As a previous study reported, EGF induces the activation of transcription factor TAZ via EGFR and downstream factors STAT3 and ERK (Gao et al., 2021)."

reach
"Furthermore, transfection of GEP100-^ PH, inhibition GEP100 or Arf6 activity by GEP100 siRNA or Arf6 T27N failed to facilitate EGF induced ERK and Rac1 activation."

reach
"SMYD3 was also required when serum was used to activate ERK1/2 activation in LKR10 cells (XREF_FIG) and when EGF was used to activate ERK1/2 in human lung and pancreatic cancer cells (XREF_FIG)."

reach
"Therefore, it may be reasonable to speculate that EGF induced ERK and Rac1 activation and cell migration require the PH domain of GEP100 and are mediated through the GEP100 and Arf6 pathways."

reach
"We found that NLRR1 positively regulated cell proliferation through activation of extracellular signal regulated kinase mediated by EGF and IGF-1."

reach
"Knockdown and overexpression of IQGAP1 decreases EGF-stimulated activation of MEK and ERK, suggesting that IQGAP1 functions as a molecular scaffold by facilitating the coupling of kinases in the Ras–M[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Thus, a different mechanism of ERK activation by either EGF or oxidative stress was regulated via RSK1 activation/inhibition in cells ( xref )."

reach
"Thus, EGF strongly activated both the ERK1/2 MAP kinase pathway and Akt in IMCD cell suspensions."

reach
"Nevertheless, compound 6 is not specific to MEK5 since it also blocks ERK1/2 activation induced by EGF as well."

reach
"Although EGF is a well-known activator of ERK, EGF downregulated the transcriptional activity and protein levels of M-Runx2 in the present study."

reach
"Taken together, we hypothesized that EGF-induced ERK activation acutely enhances the transcriptional activity of Runx2 and subsequently Runx2-dependent transcription of Smurf1."

reach
"EGF induced activation of ERK1/2 has been shown to be highly transient in HEK293 cells [XREF_BIBR]."

reach
"Consistent with these previous results, immunoblotting of phosphorylated ERK1/2 in cells stimulated with EGF for 10, 20, 30 and 60 min showed that EGF stimulation induced a peak activation of ERK1/2 at 10 min, which quickly dropped to ~ 10% of its peak level at 20 min and remained at low levels afterwards (XREF_FIG)."

reach
"ALIX knockdown cells behaved similarly as the Y319F FLAG-ALIX Bro1 expressing or the HA-ALIX V expressing cells in sustaining the EGF induced ERK1/2 activation (XREF_FIG)."

sparser
"Once again, our results support that the inhibitory effect of gallic acid on the EGF-induced Akt and ERK activation, which leads to reduced MMP-9 expression levels in EGF-treated cells."

reach
"As BzATP and EGF stimulation triggered ERK activation and regulated the endogenous DUSP6 levels in cerebellar granule neurons [23], they might also modulate the levels of DUSP1 protein."

reach
"In both physiological and cell culture systems, EGF stimulated ERK activity occurs in discrete pulses within individual cells."

reach
"Agents that cause only transient ERK activation, such as EGF, do not cause neurite outgrowth."

reach
"The role of MEK in the AhR suppression mechanism has been definitively established by the complimentary experiment in which ERK activation is restored by either CA-MEK1 or EGF."

reach
"Most importantly, all three treatments greatly prolonged the EGF induced activation of ERK1/2 (XREF_FIG), also consistent with the results obtained under continuous EGF conditions."

reach
"EGF signaling enhances ligand independent AR transactivation and promotes activation of Src and the Ras-Raf-MAPK and ERK pathway."

sparser
"Here, EGF-induced activation of EGFR-p1068 ( xref – xref ), phosphorylated AKT (pAKT) and ERK1/2 (pERK1/2) ( xref – xref ) were evaluated in whole cell lysates via immunoblotting assays."

reach
"IL-1beta or EGF further increased the activities of the extracellular signal-regulated kinase and c-Jun-N-terminal kinase/stress-activated protein kinase MAP kinases."

reach
"As noted earlier, EGF transiently activates ERKs but causes neither their nuclear accumulation nor neurite extension."

reach
"In S. mansoni Vicogne and colleagues (2004) XREF_BIBR showed that the human epidermal growth factor (EGF) can activate the Ras and ERK pathway, which induces meiosis in oocytes."

reach
"Since either the ERK or the BMK1 pathway was activated by EGF treatment, further studies were performed to confirm which was involved in the proliferation of the prostate cancer cells."

reach
"The transfection of EGFR alone with EGF increased p-Erk modestly."

reach
"We first examined effects of AG1478 and PP2 on EGF induced ERK1/2 activation to compare the IR induced EGFR activation with the ligand dependent one."

reach
"In contrast to EGF, which induced transient EGFR signaling and ERK activation, Cd promoted sustained EGFR signaling to trigger Akt/mTOR cascade."

reach
"AG1478 suppressed EGF induced ERK1/2 activation at 2-5 min after the addition of EGF into the culture medium but it failed to suppress the ERK1/2 activation later."

sparser
"Similar to FBS, conditioning with various concentrations of EGF transiently activated Akt and ERK, but activities of these kinases returned to baseline within four hours."

reach
"PP2 did not suppress EGF induced ERK1/2 activation."

sparser
"PC-3 and Du145, but not LNCaP, proliferate in response to LPA and S1P. Epidermal growth factor (EGF), phorbol 12-myristate 13-acetate (PMA), LPA, and S1P induce activation of Erks in PC-3 and Du145; only EGF and PMA activate Erks in LNCaP."

reach
"These results suggest that EGF induced ERK1/2 activation is independent of Src."

sparser
"In contrast, EGF-induced ERK activation was inhibited by AG 1478, but not by cell exposure to galardin."

reach
"Our results show that under both continuous and pulse-chase EGF conditions, inhibition of the ALIX supported MVB sorting of activated EGFR by each of the three different approaches employed converts the sharp peak kinetics of the EGF induced activation of ERK1/2 into a slow slope kinetics, lengthening the EGF induced ERK1/2 activation from < 20 min to> 2 h."

sparser
"We previously found that EGF activates ERK1/2 to stimulate goblet cell proliferation. xref As PKC is known to activate ERK1/2, we investigated whether myr-PKC α activates ERK1/2."

reach
"The overexpression or reduction of IQGAP1 expression by siRNA both caused the activation of Erk2 by EGF or IGF-1 in MCF-7 cells to be reduced, probably due to an optimal level of intracellular IQGAP1 being necessary for maximal level of activation of Erk by EGF or IGF-1 [XREF_BIBR]."

sparser
"None of the other treatments had any significant effects on the uPAR-induced response, although pertussis toxin did inhibit lysophosphatidic acid–induced ERK activation, bisindolylmaleimide I and Gö6983 inhibited PMA-induced ERK activation, and PD98059 and U0126 inhibited platelet-derived growth factor– and EGF-induced ERK activation (data not shown)."

reach
"The selective EGF-R kinase antagonist, AG1478, completely abolished EGF-induced activation of ERK1/2 and its dependent protein p90 ribosomal S6 kinase (RSK-1), but only partially (40%) inhibited ET-1-[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Stable knock-down (KD) of CIP4 in lung adenocarcinoma H1299 cells, expressing wild-type EGFR, led to increased EGFR levels on the cell surface, and defects in sustained activation of Erk kinase in H1299 cells treated with EGF."

reach
"Kinase-inactive MEKK1, but not MEKK2, 3 or 4, strongly inhibited EGF stimulated ERK activity."

reach
"While the vehicle control induced a minimal response (Figure 2A), the canonical growth factor EGF induced a maximal ERK response, which lasted 60 to 120 minutes, with 99% of cells responding to the treatment (Figure 2B, Figure E1A)."

reach
"The JNK and ERK pathways were also activated in cells treated with EGF; however, activation of these pathways was not correlated to the induction of IL-1beta expression."

reach
"A N-terminal deletion of SHP-2 not only reduced ERK activity induced by IGF-I, EGF and PDGF, but also inhibited the expression of the beta subunit of platelet derived growth factor receptor in mouse embryonic fibroblast cells [XREF_BIBR, XREF_BIBR]."

reach
"Our finding that reconstitution of SHP2 Y542F in SHP2 depleted H1666 cells rescued ERK phosphorylation as efficiently as SHP2 WT suggests that SHP2 Y542 phosphorylation is dispensable for SHP2 mediated activation of ERK, consistent with previous findings regarding EGF mediated ERK activation in 3T3 fibroblasts."

reach
"Several studies demonstrated that stiff matrices increase the frequency of EGF-induced ERK activity pulses and foster a shift towards more sustained and higher ERK activity [25, 34, 51]."

reach
"DNPKCalpha blocked EGF stimulated activation of AKT and ERK1/2 while myrPKCalpha increased activation of these kinases."

reach
"Similarly, MEC cells exhibit a higher and more sustained EGF-induced ERK activity on stiffer matrices, associated with enhanced cell contractility [25]."

sparser
"Besides, 20 μM Go6983 abolished EGF-induced ERK activation (Fig. xref ), which further indicated that PKC was the upstream kinase of MAPK pathway involved in the PN-1 up-regulation."

reach
"Our results suggest that EBV promotes EGF-induced ERK1/2 signaling by activating STIM1-dependent Ca 2+ signaling, and that blocking such signaling may inhibit EBV-promoted angiogenesis in NPC."

reach
"Thus, loss of Sprouty4 may be compensated by other genes in this family.We have shown that loss of Sprouty2 resulted in enhanced FGF- but not EGF-induced ERK activation in mouse embryonic fibroblasts [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Inhibitors of EGF stimulated ERK1/2 activity did not inhibit AKT activation but blocked proliferation."

reach
"A stiff substrate promotes integrin clustering, FAK phosphorylation (FAKpY397), vinculin recruitment to β1 integrin adhesions, and EGF-induced ERK activity."

reach
"These results are substantiated using β1-integrin mutants with enhanced transmembrane associations, resulting in larger integrin adhesions and increased EGF-induced ERK activation on soft substrates."

reach
"EGF induced weaker ERK activation in Sprouty4 KO than WT MEFs ( Fig. 3 B)."

reach
"Our findings suggest a novel mechanism by which cytokine-mediated ERK activity can cause collective cell behavior in the airway epithelium that differs from canonical EGF-mediated ERK signaling."

reach
"This proposed mechanism is distinct compared to the barrier-protective effects of EGF-mediated ERK activity (46), which we demonstrate to be uniformly distributed in epithelial monolayers."

reach
"EGF also induced robust ERK activation in primary porcine mesenchymal stem cells."

reach
"Western blot analysis showed that Akt (also named protein kinase B), which is a downstream effector of PI3K, and ERK1/2 MAPK were activated by EGF."

reach
"In addition to strong activation of ERK by EGF, and AKT by serum, early transcription remarkably differed : while EGF induced early growth response-1 (EGR1), and this was required for migration, serum induced c-Fos and FosB to enhance proliferation."

reach
"For example, epidermal growth factor robustly activated ERK, but did not stimulate iNOS-derived NO production nor did it alter B1R-mediated iNOS activation (F. Kuhr et al., submitted)."

reach
"PR antagonism with mifepristone inhibited the suppression of EGFR by heterologously expressed PR and treatment with a PR agonist-suppressed EGF-induced ERK1/2 activity further."

reach
"Stable silencing of CIP4 resulted in defects in EGF induced Erk activation, and reduced motility and invasiveness of H1299 cells."

reach
"6 Serine 272, a residue localized in the inhibitory domain of the FOXC1 protein 7 (see XREF_FIG), can be phosphorylated by EGF activated ERK signaling, resulting in stabilization of the FOXC1 protein."

reach
"These results further supported our idea that flagellin, but not hydrogen peroxide, induced the internalization of EGFR.We next examined the EGF-induced activation of ERK after pretreatment of A549 ce[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In contrast, ERK was activated by EGF only 1.7±0.1-fold ( P <0.01) and 1.7±0.2-fold after pretreatment with flagellin and TNFα, respectively."

reach
"Pretreatment with hydrogen peroxide had no effect on the activation of ERK by EGF treatment, and ERK was activated 3.5±0.6-fold."

reach
"However, transient ERK1/2 activation induced by epidermal growth factor could not mimic the gene profile of the senescent hHSCs."

reach
"In addition, ERK activation by EGF was markedly weakened after flagellin pretreatment, suggesting the desensitization of EGFR."

sparser
"As presented in , EGF induced Erk-1/2 activation in renal interstitial fibroblast NRK-49F cells ()."

reach
"In contrast, EGF induced ERK1/2, p38 MAPK and Akt activation in MPSC1 cells."

sparser
"The inhibitors of MEK-1/2 activation, PD98058 (50 μM) and U0126 (10 μM), largely (PD98059) and completely (U0126) prevented EGF-induced ERK-1/2 activation in WB-F344 cells."

reach
"Recent work in both cell lines [XREF_BIBR] and xenograft models [XREF_BIBR] indicates that the paxilin protein is phosphorylated by either androgen or EGF induced ERK signaling, facilitating signal transduction and resulting in paxillin translocation to the nucleus."

reach
"The results of the study indicate that this is due to activation of ERK1/2 by EGF, which entails increased expression of tensin-4 and indicates the role of this protein in cancer cell proliferation and migration."

reach
"For example, enforced expression of IQGAP3 accelerated the migration and invasion of lung cancer cells by interacting with ERK1 and promoting EGF induced activation of ERK [XREF_BIBR]."

sparser
"We initially optimized the system with the more robust ERK 1/2 activation by EGF, and subsequently used this response as a positive control to establish other conditions for our method."

reach
"Even though EGFR was strongly inhibited in LAP-R cells, this was associated with a strong (>3-fold) basal activation of ERK1/2 (Figure 6E and G) and an equally strong (~3-fold) EGF-stimulated ERK1/2 activation."

reach
"Since polyQ-htt strongly inhibit the EGFR redistribution from early to late endosomes, it causes, as consequence, a decrease in the active ligand-receptor complexes present within the late endosomal c[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Inhibition of ERK activity by U0126 or suppression of Rac1 activity by ectopic expression of inactive mutant form of Rac1 (Rac1-T17N) significantly prevents EGF-induced cell migration, suggesting that EGF-induced ERK and Rac1 activation was responsible for the migration of these cancer cells."

reach
"On the other hand, withdrawal of lapatinib (LAP free) was associated with >5-fold EGF-stimulated activation of MAP kinase ERK1/2 (Figure 6E–G)."

sparser
"Furthermore, transfection of GEP100-△PH, inhibition GEP100 or Arf6 activity by GEP100 siRNA or Arf6 T27N failed to facilitate EGF-induced ERK and Rac1 activation."

reach
"This also restored the responsiveness of the cells to EGF and, consequently, activation of the MAP kinase ERK1/2."

sparser
"Therefore, it may be reasonable to speculate that EGF-induced ERK and Rac1 activation and cell migration require the PH domain of GEP100 and are mediated through the GEP100/Arf6 pathways."

reach
"Interestingly, in PC12 cells prolonged ERK activation, as induced by NGF (Neuronal Growth Factor), induces differentiation, while short term ERK activation, induced by EGF (Epidermal Growth Factor), causes growth."

reach
"Among the down-regulated not shared genes ( Tables 3 and S6 ) we can evidence: i) DHX58 (Asp-Glu-X-His box polypeptide 58) which participates in innate immune defense and induces the expression of cyt[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Furthermore, specific downregulation of Rac1 signaling in HepG2 cells did not alter EGF-induced activation of ERK."

sparser
"Down-regulation of Akt1 enhances migration and ERK activation induced by EGF."

reach
"EGF activates the MEK and ERK pathway to activate RSK2."

sparser
"Akt1 down-regulation enhanced EGF-induced ERK activation and migration."

sparser
"Addition of exogenous EGF rapidly activated MEK and ERK in HT-29 cells ( Figure 3 B)."

sparser
"In contrast, overexpression of wild-type Akt2 in MCF-10A did not affect ERK activation induced by EGF."

reach
"In this case, the addition of epidermal growth factor (EGF) induces transient activation of ERK whereas nerve growth factor (NGF) induces a prolonged activation."

reach
"Once this binding is disrupted, ERK activation induced by EGF is inhibited [ 39 ]."

sparser
"The delay between the peak of direct ERK activation by EGF at t = 5 min and indirect activation by TNF at t = 15 min is an estimate of the minimum time required to establish the TNF-induced TGF-α auto[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Our previous study demonstrated that PRB expression reduced EGF-induced NSCLC cell proliferation and the activation of ERK1/2 in the absence of progestin (46)."

sparser
"The results exhibited that EGF and IL-1β additively activated ERK1/2, increased migration and invasion than either EGF or IL-1β alone in AGS and MKN-45 cells."

reach
"Finally, Src and the EGF-stimulated ERK signaling are significantly activated ( Ren et al., 2004 ; Zhang et al., 2004 )."

sparser
"Initial insight into this specificity dilemma came from observations that PC-12 cells proliferated after a transient ERK activation by epidermal growth factor (EGF), but differentiated after a sustained ERK activation by nerve growth factor (NGF), showing that the duration of ERK signaling is critical for cell fate decisions ( xref )."

sparser
"For control, we demonstrated the dose-dependent activation of ERK by EGF in PE/CA-PJ15 cells."

reach
"A well-known example of a FRET based activity reporter is the extracellular signal regulated kinase activity reporter (EKAR) [XREF_BIBR], which has been used to study oscillatory EGF mediated ERK activation and relationships between ERK activity and nucleocytoplasmic shuttling [XREF_BIBR, XREF_BIBR]."

sparser
"In cells treated with AG1478, activation of ERK1/2 by both EGF and Ang II was eliminated."

reach
"More recently, epidermal growth factor (EGF) has been shown to induce ERK activation pulses, the frequency of which depends on the concentration of EGF and controls the velocity of cell proliferation [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Both cause SHP2 catalytically inactive and significantly inhibit EGF induced ERK activation [ 78 ]."

sparser
"EGF-induced ATMCs display activation of the Erk1/2, PI3/Akt and Smad-dependent (Smad2/3) pathways."

reach
"Interestingly, the unexpected results repeatedly show that EGF induced ERKs ' and RSKs ' phosphorylation was increased by fargesin treatment in a dose dependent manner (XREF_FIG b)."

reach
"Using T47D and MCF-7 human breast carcinoma cells that express endogenous Gab1 and Gab2, we examined the role of these docking proteins in EGF induced ERK activation."

reach
"Application of epidermal growth factor to PC12 cells produces transient ERK activation (though the EGF receptor) and cell proliferation while nerve growth factor binds to the TRK receptor and leads to sustained ERK activation and cell differentiation [3]."

reach
"Expression of either SHP2 binding defective Gab1 or Gab2 mutant blocked EGF induced ERK activation."

reach
"In support of this notion, it has been demonstrated that EGF induces sustained ERK activity near the plasma membrane in sharp contrast to the transient activity observed in the cytoplasm and nucleus [185]."

reach
"How do low and medium amplitude EGF pulse trains that differ by a factor of 25 produce similar extents of ERK activation?"

reach
"In HEK293 and NIH3T3 cells, it significantly blocked SHP2 dependent ERK activation induced by EGF."

reach
"In contrast, EGF stimulation in PC12 cells triggers cytoplasmic ERK activity only, resulting in cellular proliferation, whereas neural growth factor (NGF) causes nuclear localization, which is essential for neuronal differentiation."

reach
"Notably, it enhanced paxillin phosphorylation and attenuated ERK activation induced by EGF but did not affect phorbol 12-myristate 13-acetate (PMA) triggered ERK activation (SHP2 independent), suggest[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"From these considerations, we would predict that an EGF sinewave with average EGF concentration of between 0.02 and 0.04 Kd, and frequency of 2.5 × 10 –3 × 10 rad/s (i.e., a period of 0.5 h to 4 h) would produce a DC value of ERK* of 0.11–0.16 and an ERK* maximum peak value close to 1."

reach
"Interestingly, the transfer functions depended on average receptor occupancy, with low to moderate occupancy producing the highest EGF-triggered ERK activation fluctuations."

reach
"It would be interesting to change the parameters and compartments of the current model to see where fixed concentration EGF inputs can produce ERK activity oscillations and whether these could be entrained externally."

reach
"Down-regulation of either Gab1 or Gab2 by siRNAs (small interfering RNAs) effectively inhibited the EGF stimulated ERK activation pathway and cell migration."

sparser
"Taken together, the extract of C. sudachi peel may attenuate EGF-induced ERK activation through inhibition of not only EGFR but also its downstream molecule(s), such as Ras and Raf-1, or other signaling molecule(s)."