IndraLab

Statements


| 409

sparser
"Based on these data, we wondered whether such up-regulation of USP2-AS1 had some association with the malignant behaviors of ovarian cancer cells."

sparser
"Therefore, loss-of-function assays were designed via transfecting sh-USP2-AS1#1 or sh-USP2-AS1#2 into A2780 and SKOV3 cells to effectively induce USP2-AS1 silence ( xref )."

sparser
"Then EdU assay, TUNEL assay, JC-1 assay, and transwell assay were conducted to observe the effect of USP2-AS1 knockdown on cell biological behaviors."

sparser
"The results of EdU assay demonstrated that USP2-AS1 depletion obviously suppressed ovarian cancer cell proliferation ( xref )."

sparser
"However, the outcomes of TUNEL assay and JC-1 assay indicated that cell apoptosis was remarkably strengthened due to USP2-AS1 knockdown ( xref and xref )."

sparser
"Moreover, transwell assay results revealed that USP2-AS1 deficiency seriously hampered the migration of ovarian cancer cells ( xref )."

sparser
"Collectively, USP2-AS1 is up-regulated in ovarian cancer tissues and cells and its down-regulation is effective to depress the malignant phenotypes of ovarian cancer cells."

sparser
"USP2-AS1 is Capable of Binding with miR-520d-3p."

sparser
"Therefore, we aimed to study the in-depth mechanism of USP2-AS1 in ovarian cancer."

sparser
"First, FISH assay and subcellular fractionation assay assisted us to analyze the location of USP2-AS1, and the results indicated the existence of most USP2-AS1 in the cytoplasm of A2780 and SKOV3 cells ( xref and xref )."
USP2 affects Ubiquitin
| 22 23
USP2 binds Ubiquitin.
| 9 23
| 9 23

reach
"Consistent with this hypothesis, the structures of USP2, USP8, and OTUB2 with Ub bound in the distal binding site reveal that the relevant serine residues (Ser57 for USP2, XREF_FIG K; Ser20 and 57 for USP8, XREF_FIG J and 6M; and Ser65 for OTUB2, XREF_FIG N) are solvent exposed in the distal binding sites (XREF_FIG H)."

sparser
"For USP16 we chose the model based on the USP2Ub complex (PDB ID: ;), which has a sequence identity of 34%."

sparser
"In addition, a layer of ordered water molecules mediates key interactions between ubiquitin and USP2."

sparser
"The kinetic constants are similar to those determined for native monoubiquitin, suggesting that the conjugation of an AMC group to the C-terminus of ubiquitin does not affect the kinetic behavior of ubiquitin binding to USP2."

sparser
"Kinetic studies with peptidic substrates and truncated ubiquitin mutants show that neither the C-terminal peptide nor ubiquitin deletion mutants lacking the C-terminal residues bind to USP2 with detectable affinity."

sparser
"Notably, both native ubiquitin and Ub-AMC bind to USP2 with monophasic kinetics."

sparser
"This suggests that ubiquitin binds to USP2 in a single step despite that USP2 catalytic core interacts with ubiquitin at multiple sites as revealed by the cocrystal structure of USP2-ubiquitin complex ( xref )."

sparser
"These values are close to those determined for ubiquitin binding to the wild-type USP2 ( k on = 0.40 ± 0.01 μM −1 s −1 and k off = 2.64 ± 0.56 s −1 )."

sparser
"Specific Interactions of USP2 with Ubiquitin."

sparser
"However, for a significant fraction of the total contact area the interactions are indirect, as numerous solvent molecules mediate H bond interactions between ubiquitin backbone and USP2 side chain/backbone atoms ( xref D and 2E)."
USP2 activates Ubiquitin.
| 7
| 7

reach
"USP2 promotes stabilization of cyclinD1 via direct interaction with cyclin D1 and antagonizes ubiquitin dependent degradation (Shan et al., 2009)."

reach
"At the molecular level, the simplest explanation for this result is that USP2 drives the Ub K48 -Imd molecules to the proteasome and/or ensures the removal of Ub K48 from Imd at the level of the proteasome, then allowing for Imd degradation by the proteasome machinery."

reach
"Moreover, P5091 inhibits USP7-, but not USP2- or USP8 mediated cleavage of poly K48 linked ubiquitin chains (visualized by the presence or absence of mono-ubiquitin) (XREF_SUPPLEMENTARY)."

reach
"The specificity of SJB was confirmed by various experiments : first, we show that SJB potently and selectively block USP1 activity without inhibiting other DUBs (USP2/USP5/USP7/USP14/UCH37); second, SJB inhibited binding of USP1 with HA-Ub-VS probe, but it did not affect labeling of other DUBs with probe; and third, SJB inhibited USP1, but not USP2 or USP7, triggered cleavage of ubiquitin tetramer chains."

reach
"As shown in XREF_FIG, USP2, a DUB that non specifically targets all ubiquitin linkages, completely reduced polyubiquitinated TLR2."

reach
"Since overexpression of Usp2 promotes the deubiquitination of IDOL, USP2 was considered to suppress LDL uptake via ubiquitin-dependent degradation of LDL [212]."

reach
"In agreement with this interpretation, no cleavage was detected if USP2 was allowed to remove ubiquitin chains prior to DDI2-RAD23 incubation, showing that DDI2 requires the ubiquitin chain yet cleaves in the body of the NRF1 protein (Figure 4A, compare lanes 17–20 with 11–14)."
USP2 inhibits Ubiquitin.
| 5
| 5

reach
"The inhibition of USP2 antagonizes ubiquitin-dependent degradation of cyclin D1 resulting in its stabilization."

reach
"USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin dependent degradation."

reach
"Moreover, USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin dependent degradation."

reach
"The ubiquitin dependent proteolysis of cyclin D1 is antagonized by the deubiquitinating enzyme USP2, thus increasing cyclin D1 half-life."

reach
"USP2 reverses ubiquitin‑mediated degradation of proteins and is involved in aberrant proliferation, migration, invasion, apoptosis and drug resistance of tumors."
USP2 decreases the amount of Ubiquitin.
| 1
USP2 decreases the amount of Ubiquitin. 1 / 2
| 1

reach
"Immunoblot analyses showed that USP2 treatment reduced the amount of associated ubiquitin by ~ 44% (XREF_FIG)."
Ubiquitin affects USP2
| 2 14 27
Ubiquitin binds USP2.
| 9 23
| 9 23

reach
"Consistent with this hypothesis, the structures of USP2, USP8, and OTUB2 with Ub bound in the distal binding site reveal that the relevant serine residues (Ser57 for USP2, XREF_FIG K; Ser20 and 57 for USP8, XREF_FIG J and 6M; and Ser65 for OTUB2, XREF_FIG N) are solvent exposed in the distal binding sites (XREF_FIG H)."

sparser
"For USP16 we chose the model based on the USP2Ub complex (PDB ID: ;), which has a sequence identity of 34%."

sparser
"In addition, a layer of ordered water molecules mediates key interactions between ubiquitin and USP2."

sparser
"The kinetic constants are similar to those determined for native monoubiquitin, suggesting that the conjugation of an AMC group to the C-terminus of ubiquitin does not affect the kinetic behavior of ubiquitin binding to USP2."

sparser
"Kinetic studies with peptidic substrates and truncated ubiquitin mutants show that neither the C-terminal peptide nor ubiquitin deletion mutants lacking the C-terminal residues bind to USP2 with detectable affinity."

sparser
"Notably, both native ubiquitin and Ub-AMC bind to USP2 with monophasic kinetics."

sparser
"This suggests that ubiquitin binds to USP2 in a single step despite that USP2 catalytic core interacts with ubiquitin at multiple sites as revealed by the cocrystal structure of USP2-ubiquitin complex ( xref )."

sparser
"These values are close to those determined for ubiquitin binding to the wild-type USP2 ( k on = 0.40 ± 0.01 μM −1 s −1 and k off = 2.64 ± 0.56 s −1 )."

sparser
"Specific Interactions of USP2 with Ubiquitin."

sparser
"However, for a significant fraction of the total contact area the interactions are indirect, as numerous solvent molecules mediate H bond interactions between ubiquitin backbone and USP2 side chain/backbone atoms ( xref D and 2E)."
Ubiquitin inhibits USP2.
| 2 5 4
| 2 5 4

reach
"Ubv2.3 is an engineered Ub variant, known to selectively inhibit USP2, which has C-terminal in close proximity to the catalytic active site."

reach
"While ubiquitin inhibits USP2 with a K of 2.8 μM, mutants lacking residues Arg74-Gly75-Gly76 or Leu73-Arg74-Gly75-Gly76 affect the proteolytic activity of USP2 only marginally even at the highest inhibitor concentration."

sparser
"While ubiquitin inhibits USP2 with a K i of 2.8 μM, mutants lacking residues Arg74-Gly75-Gly76 or Leu73-Arg74-Gly75-Gly76 affect the proteolytic activity of USP2 only marginally even at the highest inhibitor concentration."

eidos
"Figure 4 : USP2 Inhibition by Full-Length Ubiquitin and the C-Terminally Truncated Mutants , Ubiquitin 1-74 and 1-73 Residual enzymatic activity was determined at 22degreesC with ubiquitin-AMC as substrate ."

reach
"We therefore equipped a synthetic Ub variant, which is known to inhibit USP2 (Ubv2.3), with an aldehyde warhead at the C-terminus to improve its potency (Figure 4A)."

reach
"Inhibition of USP2 by Ubiquitin and Ubiquitin Truncation Mutants."

eidos
"While ubiquitin inhibits USP2 with a Ki of 2.8 muM , mutants lacking residues Arg74-Gly75-Gly76 or Leu73-Arg74-Gly75-Gly76 affect the proteolytic activity of USP2 only marginally even at the highest inhibitor concentration ."

sparser
"Inhibition of USP2 by Ubiquitin and Ubiquitin Truncation Mutants."

sparser
"USP2 is inhibited by ubiquitin with a K i of 2.8 μM, which is in the same range as the K i of 3 μM for the related enzyme USP5, also known as isopeptidase T ( xref )."

reach
"USP2 is inhibited by ubiquitin with a K of 2.8 μM, which is in the same range as the K of 3 μM for the related enzyme USP5, also known as isopeptidase T (Dang et al., 1998)."
USP2 affects SKP2
2 | 20 23
USP2 binds SKP2.
2 | 11 23
2 | 11 23

reach
"However, USP2 can interact with NBS1 protein when both proteins are overexpressed (Fig. 2C), and direct interaction between USP2 and SKP2 was also observed in a previous study (42)."

sparser
"In this study, we verified that the natural product Tan IIA markedly reduced the phosphorylation and total protein level of Skp2 by disrupting the interaction between Skp2 and USP2, resulting in increased Skp2 K48-linked ubiquitination and degradation."

reach
"It has been reported that E2F4 binds directly with USP2 (ubiquitin specific peptidase 2) in gastric cancer cells [29], while USP2 could bind with SKP2 and USP2-stabilized SKP2 did not destabilize its substrate P21 [30]."

reach
"We first screened a panel of DUBs and found that both USP2 and USP21 bound to endogenous SKP2, but only USP2 deubiquitylated and stabilized SKP2 protein."

reach
"Mechanistically, USP2 bound to SKP2 via the leucine-rich repeat substrate-binding domain on SKP2 to disrupt the SKP2-substrate binding, leading to stabilization of both SKP2 and these substrates."

reach
"In this study, we investigated a panel of DUBs for their potential binding of endogenous SKP2 by a conventional pull-down assay and found that both USP2 and USP21 bind to SKP2, but only USP2 stabilized SKP2."

reach
"USP2 bound to and stabilized SKP2."

reach
"Among the 9 DUBs, USP2 and USP21 selectively bound to SKP2 (Fig. 1A)."

reach
"Collectively, these results showed that USP2 interacts and stabilizes SKP2."

reach
"In this study, we showed that USP2 is a new bona fide deubiquitylase of SKP2, which is supported by the following lines of evidence: (1) SKP2 binding with USP2 under physiological conditions; (2) the levels of endogenous SKP2 are directly regulated by USP2 manipulation with USP2 knockdown to decrease and USP2 overexpression to increase SKP2; (3) USP2 stabilized SKP2 by cleaving SKP2 polyubiquitylation chains to extend SKP2 half-life; (4) pharmacologic inhibition of USP2 significantly impaired SKP2 stability; and (5) USP2 catalytic-inactive mutant USP2–C276A had no effect on SKP2."
USP2 increases the amount of SKP2.
| 3
USP2 increases the amount of SKP2. 3 / 3
| 3

reach
"Given that USP2 knockdown reduced the levels of both SKP2 and SKP2 substrates, p21 and p27 (Fig. 6B), this growth inhibitory effect appears to be mediated by other substrates of USP2 in a manner largely independent of SKP2."

reach
"Consistently, knockdown of USP2, but not USP21, reduced endogenous levels of SKP2 in three lung cancer cell lines (Fig. 1C), suggesting USP2, but not USP21, is likely an SKP2 deubiquitylase."

reach
"We found that while both USP2-WT and USP2–C276A rescued the levels of SKP2 substrates, only USP2-WT restored SKP2 protein levels (Fig. 5H)."
USP2 deubiquitinates SKP2.
| 3
USP2 deubiquitinates SKP2. 3 / 3
| 3

reach
"Likewise, USP2 knockdown shortened the protein half-life of SKP2 (Fig. 2C) and significantly increased the polyubiquitylation of SKP2 (Fig. 2D)."

reach
"We first screened a panel of DUBs and found that both USP2 and USP21 bound to endogenous SKP2, but only USP2 deubiquitylated and stabilized SKP2 protein."

reach
"The results demonstrated that SKP2 polyubiquitylation is triggered by CDH1 and FBXW2, and deubiquitylated by USP2, which is inhibited by ML364."
USP2 activates SKP2.
| 3
USP2 activates SKP2. 3 / 3
| 3

reach
"ML364, a small-molecule inhibitor of USP2, accelerates SKP2 degradation."

reach
"Surprisingly, ectopic USP2 expression caused the dose-dependent accumulation of both SKP2 and SKP2 substrates, p21 (11, 12) and p27 (13, 14, 15) in H1299 and H2170 cells (Fig. 4A)."

reach
"Interestingly and unexpectedly, USP2-induced SKP2 stabilization did not cause degradation of SKP2 substrates, rather USP2 disrupts SKP2 binding with its substrates via occupying the same LRR domain of SKP2 substrate binding."
USP2 affects KMT2A
| 45
| 45

sparser
"This issue commonly arises in KMT2A-USP2 fusion, involving a short inversion within the 11q23 region."

sparser
"Fusion-driven primary subtypes of AML identified (n=3) were: RUNX1::RUNX1T1 , KMT2Ar ( KMT2A::MLLT1, KMT2A::MLLT10; KMT2A:USP2 ), and NUP98::NSD1 (Table 1; xref )."

sparser
"In a previously published study of a cohort of 11 pediatric B-ALL patients with the KMT2A-USP2 fusion, three patients presented with central nervous system disease and 8 patients had positive-minimal residual disease at day 33, suggesting a poor prognosis of patients carrying this fusion ( xref )."

sparser
"LAMP5 expression effectively predicted the infrequent gene fusion KMT2A-USP2, frequently overlooked by routine methods, and the resultant gene product may serve as a viable therapeutic target in KMT2A-r leukemia."

sparser
"Our results identifying GATA2 as a potentially novel target of the KMT2A-USP2 fusion might provide novel clues for understanding the function of this fusion."

sparser
"In other cases, clinical break-apart FISH assays only identified one component of the likely fusion ( NUP98 in sample 0158 and KMT2A in sample 0172), while the partner gene with prognostic significance was only identified after nanopore sequencing, ( NUP98 :: NSD1 and KMT2A::USP2 )."

sparser
"Our data showed that in addition to the trans-activation of GATA2 by the KMT2A-USP2 fusion, a cis-activating mechanism is also involved in GATA2 dysregulation."

sparser
"MPAL with KMT2A usually has a B/myeloid immunophenotype (an example case of MPAL with KMT2A::USP2 fusion, B/myeloid (bilineal, pattern 1b, in xref )), although other immunophenotypic variants, including rare cases of AUL and T/myeloid MPAL, have been reported [ xref , xref , xref ]."

sparser
"Of importance, high expression levels of LAMP5 estimated the occurrence of all KMT2A-USP2 fusions."

sparser
"As expected, the two samples with out-of-frame did not present a similar expression profile of the other KMT2A fusions, as well as one sample with KMT2A-USP2 , which had higher expression of NEDD4 and lower expression of SKIDA1 , HOXA9 and MEIS1 ( xref )."
USP2 affects TP53
| 2 28 8
USP2 inhibits TP53.
| 1 10
USP2 inhibits TP53. 10 / 11
| 1 10

reach
"Since Mdm2 is responsible for p53 degradation, USP2 could negatively regulate the p53 pathway activity through up-regulation of Mdm2."

reach
"Overexpression of USP2 was also shown to reduce p53 stability [XREF_BIBR], and another de-ubiquitylase USP8 (also called UBPY) promotes epithelial growth factor receptor degradation [XREF_BIBR, XREF_BIBR, XREF_BIBR]."

reach
"USP2 additionally prevents p53 signaling."

eidos
"USP2 additionally prevents p53 signaling ."

reach
"Targeting USP2 regulation of VPRBP-mediated degradation of p53 and PD-L1 for cancer therapy."

reach
"Inhibition of USP2 activates p53 but Mdm2-mediated regulation of p53 is largely unaffected."

reach
"Here we show that VPRBP levels are tightly controlled by USP2, a deubiquitinase that binds and stabilizes VPRBP, and that loss of USP2 function can activate p53 responses, including its tumor suppressive effects."

reach
"We postulate that upregulation of USP2 antagonizes p53 effect via deubiquitination of Mdm2."

reach
"Our data demonstrate that USP2 inhibition activates p53 function by destabilizing VPRBP and that it does so without compromising p53-Mdm2 feedback loop."

reach
"Overexpression of USP2 was shown to reduce p53 stability, conversely suppression led to destabilisation of MDM2 and increased p53 level and activation (9)."
USP2 binds TP53.
| 2 8
| 2 8

sparser
"In vivo experiments in mice explored the specific role of the USP2-p53 axis in a xenograft tumor model."

sparser
"Although this study provides a preliminary exploration of the role of the USP2-p53 axis-mediated ferroptosis in reversing cisplatin resistance in NSCLC, several limitations persist."

sparser
"Subsequently, COIP was utilized to identify the binding of USP2 and p53, which showed the direct binding between USP2 and p53 (Fig.  xref B)."

sparser
"In our study, USP2 overexpression inhibited p53 ubiquitination and directly interacted with p53."

sparser
"USP2, which is usually suppressed in cisplatin-resistant NSCLC, can directly interact with p53 to promote its stability and nuclear translocation, eventually facilitating ferroptosis xref ."

reach
"Moreover, USP2 directly bound to p53 and USP2 overexpression stabilized p53 protein by suppressing its ubiquitination."

reach
"Subsequently, COIP was utilized to identify the binding of USP2 and p53, which showed the direct binding between USP2 and p53 (Fig. 5B)."

sparser
"Our study preliminarily unveils a novel mechanism of DDP resistance in NSCLC, highlighting the critical role of the USP2-p53 axis in regulating cisplatin sensitivity in NSCLC cells."

sparser
"Moreover, USP2 directly bound to p53 and USP2 overexpression stabilized p53 protein by suppressing its ubiquitination."

sparser
"This study revealed that USP2 and p53 were associated with NSCLC cisplatin resistance."
USP2 activates TP53.
| 1 5
USP2 activates TP53. 6 / 6
| 1 5

reach
"Therefore, we speculate that USP2 overexpression may lower the ubiquitination levels of p53 by acting on the K305R site of p53 protein and promote the translocation of p53 from the cytoplasm to the nucleus."

reach
"These data demonstrate that CD8 + T cells, but not CD4 + T cells, are the primary effector cells underlying the combination treatment in these tumor models.Finally, to ascertain whether USP2 inhibition potentiates the efficacy of PD-1/PD-L1 blockade by enhancing p53 function in a tumor cell autonomous manner, we also examined the effects of combination therapy in Balb/c mice bearing p53-null EMT6 tumors."

reach
"We speculate that the overexpression of USP2 promoted the nuclear translocation of p53."

reach
"Although p53 can be activated robustly under both conditions, in contrast to Mdm2 inhibition, USP2 inhibition neither affects Mdm2 induction by p53 nor suppresses Mdm2-mediated repression of p53."

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

eidos
"On the other hand , USP2 has also been shown to increase p53 in a hepatoma cell line ( HepG2 ) and a breast cancer cell line ( MCF7 ) after leptin stimulation [ 36 ] ."
USP2 decreases the amount of TP53.
| 5
USP2 decreases the amount of TP53. 5 / 5
| 5

reach
"Therefore, we speculate that USP2 overexpression may lower the ubiquitination levels of p53 by acting on the K305R site of p53 protein and promote the translocation of p53 from the cytoplasm to the nucleus."

reach
"The WB results revealed that USP2 overexpression inhibited GPX4, SLC7A11 and cytoplasm p53 protein expression, while promoted the nucleus p53 protein expression."

reach
"Notably, USP2 overexpression promoted p53 expression in the nucleus, while suppressing p53 expression in the cytoplasm (Fig. 4D)."

reach
"USP2 shows a tendency to bind and deubiquitinate Mdm2 and thus suppress the expression of p53 [152] ."

reach
"The results demonstrated that USP2 overexpression significantly inhibited p53 ubiquitination levels (Fig. 5A)."
USP2 increases the amount of TP53.
| 3
USP2 increases the amount of TP53. 3 / 3
| 3

reach
"Indeed, inactivation of USP2 increases the levels of both p53 and PD-L1 in tumors, suggesting that the tumor suppression activity of USP2 inhibition may be potentiated by the PD-1/PD-L1 immune checkpoint blockade."

reach
"Notably, USP2 overexpression promoted p53 expression in the nucleus, while suppressing p53 expression in the cytoplasm (Fig. 4D)."

reach
"Ubiquitination assays evaluated the ubiquitination levels of p53 following USP2 overexpression."
USP2 deubiquitinates TP53.
| 3
USP2 deubiquitinates TP53. 3 / 3
| 3

reach
"The results demonstrated that USP2 overexpression significantly inhibited p53 ubiquitination levels (Fig. 5A)."

reach
"In our study, USP2 overexpression inhibited p53 ubiquitination and directly interacted with p53."

reach
"Unlike USP7, USP2 does not deubiquitinate p53 (9)."
Gallic acid affects USP2
| 20 16
Gallic acid binds USP2.
| 14 14
| 14 14

sparser
"These findings collectively suggested the specific interaction between GA and USP2."

sparser
"Next, we performed a Cellular Thermal Shift Assay (CETSA) to determine whether GA directly binds to USP2 in cells."

sparser
"Considering that USP2 functions as a ubiquitin-specific protease, we hypothesized that the binding of GA to USP2 may inhibit its enzymatic activity."

sparser
"Cysteine 284 is critical for the covalent binding of GA to USP2."

sparser
"We then investigated the mode of interaction between GA and USP2."

sparser
"Interestingly, preincubation of iodoacetic acid (IAA), a cysteine alkylating agent, with USP2 completely inhibited the binding of biotin-GA to USP2 ( xref ), suggesting that GA may covalently bind to the cysteine(s) of USP2, which is consistent with previous reports that GA can covalently bind to its targets through cysteine residues xref ."

sparser
"It can be observed from xref C that the binding of GA to USP2 was dependent on the incubation time, following the pattern of an irreversible binding mechanism."

sparser
"These data suggest that GA covalently binds to USP2 in a dose- and time-dependent manner."

sparser
"GA binds to USP2 through an allosteric pocket."

sparser
"As shown in xref F, molecular docking indicated that P565 and R289 are critical for forming the pocket for GA to bind to USP2."
Gallic acid inhibits USP2.
| 3 2
| 3 2

reach
"In contrast, our results indicate that GA could inhibit USP2 activity by covalently binding to the Cys284 residue, instead of the Cys276 residue within the catalytic site, of USP2."

reach
"In addition to the CETSA results, the drug affinity-responsive target stability assay showed that GA could prevent the pronase-mediated digestion of USP2 in a dose-dependent manner (Supporting Information Fig. S4)."

sparser
"Specifically, we employed GA as a chemical probe and demonstrated that GA could covalently bind to and inhibit USP2 through an allosteric pocket."

reach
"As shown in Fig. 2I and J, GA inhibited the hydrolytic activity of USP2 against Ub-AMC in a concentration-dependent manner (Fig. 2I)."

sparser
"We further revealed that the USP2 stabilizes KRAS by deubiquitination; consequently, USP2 inactivation induced by GA could lead to the destabilization of KRAS, resulting in the suppression of MM cell proliferation in vitro and in vivo ."
Gallic acid activates USP2.
| 3
| 3

reach
"As shown in Fig. 2E and F, compared with DMSO treatment, GA increased the thermal stability of USP2 in MM.1S cells; meanwhile, no significant change in the thermal stability of vinculin (negative control) could be observed."

reach
"We further revealed that the USP2 stabilizes KRAS by deubiquitination; consequently, USP2 inactivation induced by GA could lead to the destabilization of KRAS, resulting in the suppression of MM cell proliferation in vitro and in vivo."

reach
"In addition to the CETSA results, the drug affinity-responsive target stability assay showed that GA could prevent the pronase-mediated digestion of USP2 in a dose-dependent manner (Supporting Information Fig. S4)."
SKP2 affects USP2
2 | 11 23
2 | 11 23

reach
"However, USP2 can interact with NBS1 protein when both proteins are overexpressed (Fig. 2C), and direct interaction between USP2 and SKP2 was also observed in a previous study (42)."

sparser
"In this study, we verified that the natural product Tan IIA markedly reduced the phosphorylation and total protein level of Skp2 by disrupting the interaction between Skp2 and USP2, resulting in increased Skp2 K48-linked ubiquitination and degradation."

reach
"It has been reported that E2F4 binds directly with USP2 (ubiquitin specific peptidase 2) in gastric cancer cells [29], while USP2 could bind with SKP2 and USP2-stabilized SKP2 did not destabilize its substrate P21 [30]."

reach
"We first screened a panel of DUBs and found that both USP2 and USP21 bound to endogenous SKP2, but only USP2 deubiquitylated and stabilized SKP2 protein."

reach
"Mechanistically, USP2 bound to SKP2 via the leucine-rich repeat substrate-binding domain on SKP2 to disrupt the SKP2-substrate binding, leading to stabilization of both SKP2 and these substrates."

reach
"In this study, we investigated a panel of DUBs for their potential binding of endogenous SKP2 by a conventional pull-down assay and found that both USP2 and USP21 bind to SKP2, but only USP2 stabilized SKP2."

reach
"USP2 bound to and stabilized SKP2."

reach
"Among the 9 DUBs, USP2 and USP21 selectively bound to SKP2 (Fig. 1A)."

reach
"Collectively, these results showed that USP2 interacts and stabilizes SKP2."

reach
"In this study, we showed that USP2 is a new bona fide deubiquitylase of SKP2, which is supported by the following lines of evidence: (1) SKP2 binding with USP2 under physiological conditions; (2) the levels of endogenous SKP2 are directly regulated by USP2 manipulation with USP2 knockdown to decrease and USP2 overexpression to increase SKP2; (3) USP2 stabilized SKP2 by cleaving SKP2 polyubiquitylation chains to extend SKP2 half-life; (4) pharmacologic inhibition of USP2 significantly impaired SKP2 stability; and (5) USP2 catalytic-inactive mutant USP2–C276A had no effect on SKP2."
USP2 affects UBD
| 8 12
| 8 12

sparser
"We carried out stopped-flow fluorescence analyses of the binding of mono- and diubiquitin to an inactive USP2 mutant and unveiled interesting differences in the binding kinetics between the two substrates."

sparser
"Binding of K48-linked IQF diubiquitin to USP2 under an equilibrium condition."

sparser
"Binding was monitored by following the enhancement of TAMRA fluorescence upon the binding of K48-linked IQF diubiquitin substrate (25 nM) to USP2 (0-8 μM)."

sparser
"Transient kinetics of K48-linked diubiquitin binding to USP2."

sparser
"Stopped-flow fluorescence analysis was used to follow K48-linked IQF diubiquitin binding to USP2 under a pre-steady-state condition."

sparser
"Based on the regression analysis of the plots of k obs,1 and k obs,2 , using xref - xref we were able to calculate each microscopic binding rate constant for a two-step process describing the binding of diubiquitin to USP2."

sparser
"The binding interaction between USP2 and native K48-linked diubiquitin was also characterized using stopped-flow fluorescence analysis under the pre-steady-state condition."

reach
"This apparent dissociation constant was later used in calculating the individual microscopic binding rate constants in a proposed two-step process of diubiquitin binding to USP2 (vide infra)."

reach
"Transient kinetics of K48 linked diubiquitin binding to USP2."

reach
"Stopped-flow fluorescence analysis was used to follow K48 linked IQF diubiquitin binding to USP2 under a pre-steady-state condition."
USP2 affects gallic acid
| 14 14
| 14 14

sparser
"These findings collectively suggested the specific interaction between GA and USP2."

sparser
"Next, we performed a Cellular Thermal Shift Assay (CETSA) to determine whether GA directly binds to USP2 in cells."

sparser
"Considering that USP2 functions as a ubiquitin-specific protease, we hypothesized that the binding of GA to USP2 may inhibit its enzymatic activity."

sparser
"Cysteine 284 is critical for the covalent binding of GA to USP2."

sparser
"We then investigated the mode of interaction between GA and USP2."

sparser
"Interestingly, preincubation of iodoacetic acid (IAA), a cysteine alkylating agent, with USP2 completely inhibited the binding of biotin-GA to USP2 ( xref ), suggesting that GA may covalently bind to the cysteine(s) of USP2, which is consistent with previous reports that GA can covalently bind to its targets through cysteine residues xref ."

sparser
"It can be observed from xref C that the binding of GA to USP2 was dependent on the incubation time, following the pattern of an irreversible binding mechanism."

sparser
"These data suggest that GA covalently binds to USP2 in a dose- and time-dependent manner."

sparser
"GA binds to USP2 through an allosteric pocket."

sparser
"As shown in xref F, molecular docking indicated that P565 and R289 are critical for forming the pocket for GA to bind to USP2."

reach
"As shown in Fig. 5A and B, USP2 knockout significantly inhibited the proliferation of MM.1S and RPMI 8226 cell lines."

reach
"These results suggest that USP-2 promotes cell proliferation pertinent to the features of human hepatic and breast cancers."

reach
"For example, USP2, USP22, USP28, and USP37 stabilize c-Myc expression and, thus, stimulate the proliferation of many tumour cells [58–61] ."

reach
"The results of the Cell Counting Kit-8 and clone formation assays showed that ubiquitin-specific peptidase 2 knockdown enhanced the proliferation rate of 769-P, 786-O, and Caki-1 cells (Figures 2(g,h))."

reach
"This study highlighted that USP2 inhibition or knockdown could inhibit the proliferation of MM cells and induce MM cell apoptosis in vitro and in vivo."

reach
"USP2 overexpression promoted proliferation and metastasis, and induced the EMT in CM cells in vitro, while specific inhibition of USP2 by ML364 produced the opposite effects."

reach
"Mutations in the CYLD deubiquitinating enzyme leads to familial cylindromatosis XREF_BIBR and overexpression of USP2 has been reported to increase prostate cancer cell proliferation through the stabilization of fatty acid synthase XREF_BIBR."

reach
"In addition, USP2 inhibition suppressed CFs proliferation, collagen synthesis and cell cycle progression."

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

reach
"USP2 promotes cell proliferation and metastasis in choroidal melanoma via stabilizing Snail."

reach
"Ubiquitin-specific peptidase 2 overexpression inhibited clear cell renal cell carcinoma cell proliferation, migration, and invasion in vitro and in vivo."

reach
"In contrast, we found that USP2 inhibits the TGF-β signaling and the proliferation of myeloid cells."

reach
"Inactivation or knockdown of USP2 leads to the degradation of KRAS, resulting in the suppression of MM cell proliferation in vitro and in vivo."

reach
"USP2a overexpression in HepG2 and Huh 7 cells significantly increased cell proliferation while inhibition of USP2a activity by chemical inhibitor or stable knockout of USP2 by CRISPR markedly reduced cell proliferation."

reach
"We further found that USP2 inhibits the proliferation of myeloid cells to suppress the activation and expansion of CD4 T cells and CD8 T cells and the production of IL-22 and IFNγ."

reach
"Taken together, these data suggest that USP2 restricts the proliferation of myeloid cells and thereby inhibits the activation and expansion of IL-22- and IFNγ-producing T cells in the lamina propria during DSS-induced colitis.2.6 USP2 in myeloid cells inhibits immune responses to bacterial infections in the gut."

reach
"Ubiquitin-specific peptidase 2 inhibited the proliferation, migration, and invasion of clear cell renal cell carcinoma cells."

reach
"We further revealed that the USP2 stabilizes KRAS by deubiquitination; consequently, USP2 inactivation induced by GA could lead to the destabilization of KRAS, resulting in the suppression of MM cell proliferation in vitro and in vivo."

reach
"The proliferative abilities shown by the Cell Counting Kit-8 and clone formation assays indicated that cell proliferation was inhibited by ubiquitin-specific peptidase 2 overexpression in all three cell lines (Figures 3(b,c))."

reach
"These results demonstrate that ubiquitin-specific peptidase 2 inhibited Caki-1 cell proliferation both in vivo and in vitro."
USP2 affects MDM2
5 1 1 | 14 2
USP2 binds MDM2.
5 1 | 3 2
5 1 | 3 2

reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

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reach
"The C-terminal region of USP2 forms part of the catalytic domain and binds to the C-terminal Zinc and Ring finger domain of mdm2, the same region of mdm2 that we demonstrated to bind to USP7.Crystal s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Specifically, protein-protein interaction assays using the bacterial two-hybrid system showed that USP2 can deubiquitinate MDM2 and promotes p53 degradation, showing the association between USP2 and MDM2 (Stevenson et al., 2007)."

sparser
"Subsequent analysis demonstrated that USP2 can bind and deubiquitinate Mdm2."
USP2 deubiquitinates MDM2.
1 | 5
USP2 deubiquitinates MDM2. 6 / 6
1 | 5

reach
"USP2 deubiquitinates both MDM2 and MDMX [XREF_BIBR, XREF_BIBR] whereas USP4 deubiquitinates ARF-BP1 [XREF_BIBR], another ubiquitin ligase for p53, thus indirectly destabilizing p53 and inhibiting its function."

reach
"MDM2 can also be deubiquitinated by USP2, and this may contribute to osteosarcoma tumorigenesis [85]."

reach
"Specifically, protein-protein interaction assays using the bacterial two-hybrid system showed that USP2 can deubiquitinate MDM2 and promotes p53 degradation, showing the association between USP2 and MDM2 (Stevenson et al., 2007)."

reach
"USP2 deubiquitinates and stabilizes MDM2 thus inhibiting the proapoptotic activity of p53."

reach
"USP2 deubiquitinates the E3 ligase MDM2 and prevents its degradation [23] , and thus, we explored the effects of UbVs on MDM2 ubiquitination."
USP2 decreases the amount of MDM2.
| 2
USP2 decreases the amount of MDM2. 2 / 4
| 2

reach
"Conversely, the PD-L1 levels were induced upon USP2 inhibition in all cell types regardless of their p53 status.Since Mdm2 levels were induced in p53-positive cells by USP2 inhibition (Fig. 6a, c), we examined whether USP2 inhibition affects Mdm2-mediated degradation of p53."

reach
"Knockdown of USP2 decreases Mdm2 protein level and increases p53 transcriptional activity."
USP2 activates MDM2.
| 3
USP2 activates MDM2. 3 / 4
| 3

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

reach
"USP2 has been shown to enhance cancer cell proliferative and invasive capabilities by stabilizing and elevating murine double minute 2 (MDM2) protein, which subsequently promotes p53 protein elimination [17, 18]."

reach
"Although p53 can be activated robustly under both conditions, in contrast to Mdm2 inhibition, USP2 inhibition neither affects Mdm2 induction by p53 nor suppresses Mdm2-mediated repression of p53."
USP2 inhibits MDM2.
| 1
USP2 inhibits MDM2. 1 / 2
| 1

reach
"However, our Western blot assays demonstrated that depleting USP2 expression in multiple TNBC cells neither downregulated MDM2 nor upregulated p53 expression, suggesting that the MDM2 and p53 axis is not involved in USP2 's functions in CSCs."
1 | 1 19 1
1 | 11 1
1 | 11 1

reach
"USP2 knockout also induced apoptosis in these two MM cell lines (Fig. 5C and D)."

reach
"Moreover, USP2 silencing enhanced apoptosis after PUVA treatment in MyLa2000 suggesting that USP2 enhanced therapeutic resistance."

reach
"In addition, we observed that the ectopic of USP2 WT diminishes the apoptosis, whereas overexpression of catalytically mutated USP2 (C276A) causes apoptosis in ML362-plus-TRAIL-treated cells (Figure 6B)."

reach
"Here, we found that USP2 showed decreased expression in advanced CTCL compared with the early stage disease and blocked apoptosis in malignant p53 wt T-lymphocytes."

reach
"Finally, the knockdown of USP2 in Hut-78 cells did not increase apoptosis after PUVA, in contrast to what was observed in MyLa2000 with intact p53."

reach
"Overexpression of survivin or USP2 significantly prevented apoptosis through cotreatment with ML364 and TRAIL, whereas a knockdown of USP2 increased sensitivity to TRAIL."

reach
"USP2 knockdown via siRNA inhibits actinomycin and TNF-α-induced apoptosis via c-FLIP upregulation in hepatocytes [27]."

reach
"USP2 reduces cell apoptosis via p53 signaling."

reach
"Stevenson, et al. showed that USP2 prevents cell apoptosis induced by stabilizing MDM2, in turn leading to p53 reduction XREF_BIBR, XREF_BIBR."

reach
"Pharmacological USP2 targeting suppresses ovarian cancer growth by potentiating apoptosis and ferroptosis."
USP2 activates apoptotic process.
| 1 8

reach
"We have previously shown that the 41 kDa USP2 isoform induces apoptosis [3] and that the 69 kDa USP2a isoform mediates the apoptosis signal of the TNF receptor [5]."

reach
"USP2 Plays an Important Role in TRAIL-Mediated Apoptosis ."

eidos
"Significantly , siRNA knockdown of USP2 resulted in apoptosis , which could be reversed by overexpression of fatty acid synthase ( FAS ) ( 8) ."

reach
"These data collectively suggest that the USP2 plays a critical role in the proliferation and survival of MM cells in vitro and in vivo.3.8 GA induces the apoptosis of MM cells by targeting USP2 and destabilizing KRAS."

reach
"Caspase dependent apoptosis caused by overexpression of USP2 was first described for USP2-2 [XREF_BIBR] but also for isoform 1 [XREF_BIBR]."

reach
"Knockdown of USP2 did not increase cell apoptosis in Hut-78 cells compared to cells with control siRNA, further suggesting the pivotal role of p53 in USP2 function."

reach
"Our studies indicate that apoptosis can be modulated by peroxisomal import of USP2, resulting in its spatial separation from its target proteins in the cytosol and nucleus."

reach
"In vitro an artificial knockdown of USP2 inhibited actinomycin D/TNFalpha induced hepatocyte apoptosis, which was associated with elevated levels of the anti-apoptotic protein c-Flip (L/S) and a concomitant decrease of cellular levels of the ubiquitinligase Itch, a negative regulator of c-Flip."

reach
"Hence, the USP2 variants seem to cause the activation of apoptosis pathways by the specific stabilising of their RIP1 substrate.Based on the pro apoptotic effect when USP2c was targeted by RNAi, we fo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects UBC
24 | 1
24 | 1

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USP2 affects Cyclin
| 23
USP2 activates Cyclin.
| 9
USP2 activates Cyclin. 9 / 9
| 9

reach
"Although, USP-2, as a deubiquitinating enzyme, is well known to prevent degradation of cyclins, including cyclin A1 and cyclin D1 (Kim et al., 2012; Shan et al., 2009), no studies have been reported r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Pharmacological inhibition of USP2 accelerates cyclin D1 degradation and leads to cell cycle arrest in several cancer cell lines among which the HCT116 colon cancer cell line and MCF7 breast cancer cell line."

reach
"USP2 directly interacts with cyclin D1 to perform deubiquitylation and thereby antagonize proteasome-dependent degradation of cyclin D1."

reach
"USP2 overexpression also increased the levels of two additional SKP2 substrates, cyclin D1 (12) and cyclin E2 (32) in H2170 cells with the low basal levels, but not in H1299 cells with high basal levels (Fig. 4A)."

reach
"The ubiquitin dependent proteolysis of cyclin D1 is antagonized by the deubiquitinating enzyme USP2, thus increasing cyclin D1 half-life."

reach
"APN can inhibit the expression of USP2 in tumor cells and promote the degradation of cyclin D1 [XREF_BIBR]."

reach
"Again, knockdown of USP2 in MCF-7 and PC3 cells significantly reduced the steady state levels of cyclin D1 protein (XREF_FIG, also in XREF_SUPPLEMENTARY) and also induced cell growth suppression (XREF_FIG)."

reach
"As expected, the half-life of cyclin D1 protein in MCF-7 cells was longer than that of other cancer lines such as HCT116; however, USP2 knockdown drastically reduced the cyclin D1 half-life from 60 to 20 minutes (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"Our results reveal that USP2 specifically modulates cyclin D1 function in vivo and demonstrate the importance of this regulation in human cancer cell growth."
USP2 deubiquitinates Cyclin.
| 6
USP2 deubiquitinates Cyclin on D1. 6 / 6
| 6

reach
"ML364 is the inhibitor of the deubiquitinase USP2, which mediates the ubiquitination of cyclin D1."

reach
"In particular, it has been recently shown that USP-2 deubiquitinates cyclin D1, prevents degradation, accumulates cyclin D1 and finally leads to cell cycle progression from G1 to S phase (Lee et al., [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"reported that USP2 specifically deubiquitinates and stabilizes cyclin D1, whereas knockdown of this protease causes cyclin D1 degradation and growth arrest in cancer cells dependent on this cell cycle regulator."

reach
"Cyclin D1 ubiquitination is reversed by the deubiquitinases USP2 and USP22."

reach
"Several compounds have been shown to suppress the activity of USP2, which can promote the deubiquitylation and stability of cyclin D1 [137]."

reach
"In a screen of potential deubiquitination enzymes, Shan et al. identified USP2 to specifically deubiquitinate cyclin D1, preventing the degradation of cyclin D1, and leading to the accumulation of cyclin D1."
USP2 inhibits Cyclin.
| 4
USP2 inhibits Cyclin. 4 / 4
| 4

reach
"These data demonstrate that inactivation of USP2 downregulates the levels of endogenous cyclin D1 and suppresses cell growth by delaying G1/S progression."

reach
"The ubiquitin dependent proteolysis of cyclin D1 is antagonized by the deubiquitinating enzyme USP2, thus increasing cyclin D1 half-life."

reach
"The inhibition of USP2 antagonizes ubiquitin-dependent degradation of cyclin D1 resulting in its stabilization."

reach
"Based on our results, it is highly likely that USP-2 suppression by adiponectin enhanced proteasomal activity, thereby increasing cyclin D1 degradation, whereas enhancement in USP-2 expression by lept[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 increases the amount of Cyclin.
| 2
USP2 increases the amount of Cyclin. 2 / 2
| 2

reach
"We found that USP2 knockdown consistently downregulated cyclin D1 expression in multiple TNBC cells."

reach
"Furthermore, the role of USP-2 induction in cyclin D1 expression was examined by gene silencing of USP-2."
USP2 binds Cyclin.
| 2
USP2 binds Cyclin and D1. 2 / 2
| 2

reach
"USP2 binds to Cyclin D1 and Mdm2 and induces their deubiquitination and stabilization, and its silencing causes downregulation of Cyclin D1 and Mdm2, leading to inhibiting cell proliferation."

reach
"By using co-immunoprecipitation approaches, we tried to determine whether elevation of O GlcNAcylation favors the binding of USP2 to cyclin D1, as described for BAP1 and PGC1alpha."
USP2 affects PER1
1 | 12 8
USP2 binds PER1.
| 6 8
| 6 8

sparser
"These findings show that USP2 binds PER1 directly and indicate that the interaction of USP2 with the other clock proteins is indirect and possibly occurring within a complex that contains components known to participate in the main negative feedback loop of the circadian clock, including PER1."

sparser
"Earlier studies from our lab showed that USP2 directly interacts with ubiquitinated PER1 (Yang et al.  xref ) and facilitates its localization to the nucleus (Yang et al.  xref )."

reach
"USP2 can be coimmunoprecipitated with several clock proteins but directly interacts specifically with PER1 and deubiquitinates it."

sparser
"In our hands, whereas it forms a complex with several clock proteins, USP2 directly binds only to PER1 ( xref )."

reach
"These findings show that USP2 binds PER1 directly and indicate that the interaction of USP2 with the other clock proteins is indirect and possibly occurring within a complex that contains components known to participate in the main negative feedback loop of the circadian clock, including PER1."

reach
"For example, USP2 binds directly to PER1 in order to regulate PER1 intracellular localization [XREF_BIBR]."

sparser
"Previous studies have shown that USP2 directly interacts with PER1 and affects its localization (Yang et al.  xref )."

sparser
"However, in vitro assays indicated that USP2 could only interact directly with PER1, suggesting that USP2 interacts with the other clock proteins by virtue of their being in a complex with PER1 ( xref )."

sparser
"The USP2 protein binds and deubiquitinates PER1, regulating its nuclear localization [ xref , xref ]."

sparser
"For example, USP2 binds directly to PER1 in order to regulate PER1 intracellular localization [ xref ]."
USP2 deubiquitinates PER1.
1 | 6
USP2 deubiquitinates PER1. 7 / 7
1 | 6

reach
"USP2 is essential to deubiquitinating PER1, CRY1 and CRY2 in vivo [54–57]."

reach
"Deubiquitination of PER1 by USP2 does not stabilize it."

reach
"In light of the shorter half-life of USP2 (compared to that of PER1) (XREF_FIG) and of its rhythmic expression peaking almost simultaneously with that of PER1, the deubiquitination of PER1 by USP2 could be restricted to a specific circadian time and therefore be altering the function and/or localization of PER1 to fine tune the molecular clock in physiological conditions."

reach
"The USP2 protein binds and deubiquitinates PER1, regulating its nuclear localization [31,78]."

"Recently, we showed that the deubiquitinating enzyme ubiquitin-specific peptidase 2 (<span class="match term0">USP2</span>) associates with clock proteins and deubiquitinates PERIOD1 (<span class="match term1">PER1</span>) but does not affect its overall stability."

reach
"Further, USP2 is able to deubiquitinate the ubiquitinated forms of the clock proteins BMAL1 (Scoma et al. 2011), PER1 (Yang et al. 2012) and CRY1 (Tong et al. 2012)."

reach
"Our previous studies have shown that PER1 is deubiquitinated by USP2 (Yang et al. 2012)."
USP2 affects PPARG
| 11 8
USP2 binds PPARG.
| 8
| 8

sparser
"To further investigate whether USP2 directly interacts with PPAR-γ, we performed endogenous and exogenous co-immunoprecipitation (Co-IP) analysis."

sparser
"USP2 interacted with PPAR-γ in C2C12 cells and the Gas muscle ( xref )."

sparser
"Further immuno-precipitation analysis and pulldown assay also joined to confirm the interaction between USP2 and PPARγ ( xref )."

sparser
"Targeting the USP2PPAR-γ axis may offer promising therapeutic strategies for metabolic disorders and sarcopenia."

sparser
"In addition, the computational modeling was used to determine whether USP2 binds to PPARγ and which residues are required for their interaction."

sparser
"Data also showed the USP2-PPAR-γ interaction was key in mediating the effects of USP2 on muscle atrophy and insulin signaling ( xref and xref )."

sparser
"By identifying USP2 as a novel PPAR-γ deubiquitinase and protective factor, we have broadened the PPAR-γ regulatory network and revealed the potential of targeting the USP2PPAR-γ axis for skeletal muscle and insulin signaling disorders."

sparser
"These findings highlight the potential of targeting the USP2PPAR-γ axis for therapeutic interventions aimed at combating muscle atrophy and enhancing insulin sensitivity, thereby presenting novel avenues for addressing metabolic disorders and sarcopenia."
USP2 increases the amount of PPARG.
| 7
USP2 increases the amount of PPARG. 7 / 7
| 7

reach
"In this study, USP2 deficiency reduced PPAR-γ expression and increased its protein ubiquitination (Fig. 4)."

reach
"WT USP2 overexpression induced an increase in the protein level of PPAR-γ and decreased PPAR-γ ubiquitination, but not USP2 C276A (an enzyme-activity dead mutation) overexpression (Fig. 4C)."

reach
"Usp2 overexpression increased PPAR-γ protein levels in the Gas muscles of diabetic mice (Fig. 5C)."

reach
"Knockdown of PPAR-γ reduced and Usp2 overexpression increased the levels of PPAR-γ, IRS1, and GLUT4 in C2C12 cells (Fig. 6A)."

reach
"USP2 Upregulates PPAR-gamma Expression and Improves Insulin Signaling."

reach
"Usp2 knockdown reduced the expression of PPAR-γ, GLUT4, and IRS1, and increased atrogene expression under DEX stimulation (Supplementary Fig. 4A)."

reach
"In contrast, Usp2 overexpression induced the expression of PPAR-γ, GLUT4, and IRS1, and reduced the expression of atrogenes under DEX stimulation (Supplementary Fig. 4B)."
USP2 inhibits PPARG.
| 2
USP2 inhibits PPARG. 2 / 2
| 2

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"Taken together, these results revealed a previously unrecognized yet critical role of USP2 in the regulation of PPAR-γ signaling and in the pathogenesis of muscle atrophy.Our mechanistic investigations revealed that USP2 prevents muscle atrophy by regulating the stability of PPAR-γ, a key TF implicated in muscle metabolism and insulin sensitivity (31–33)."

reach
"Therefore, we hypothesized that Usp2 depletion in the diabetes- and DEX-induced muscle atrophy models might destabilize PPAR-γ protein."
USP2 activates PPARG.
| 2
USP2 activates PPARG. 2 / 2
| 2

reach
"In contrast, USP2 deficiency also decreased peroxisome proliferator-activated receptor gamma coactivator 1alpha (PGC1alpha) protein in C2C12 cells, while it upregulated Ppargc1a mRNA."

reach
"We also observed that adenovirus-mediated Usp2 (AdUsp2) overexpression increased PPAR-γ, GLUT4, and IRS1 expression under TNF-α stimulation (Fig. 5F)."
USP2 affects ARID2
3 | 6 9
USP2 binds ARID2.
3 | 1 9
3 | 1 9

sparser
"USP2 Is Significantly Suppressed and Interacts with ARID2 in Lung Cancer Cells."

sparser
"Based on the above findings, we then utilized HEK-293T cell line as a model to explore and confirm the potential interaction of USP2-ARID2."

reach
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure 1 B) and interactive Flag-ARID2 (Supplementary Figure 1 C) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

No evidence text available

sparser
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure xref ) and interactive Flag-ARID2 (Supplementary Figure xref ) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

No evidence text available

sparser
"The results indicated that in both A549 and H1299 cells, ARID2 protein can be detected in anti-USP2 antibody precipitated protein samples, and vice versa ( xref ), suggesting endogenously USP2-ARID2 protein-protein interaction (PPI) in lung cancer cells."

No evidence text available

sparser
"The quantitative analysis of the USP2-ARID2 colocalization ratio also provided consistent results (Figures xref and xref )."

sparser
"To explore the potential role of proteasome-dependent protein degradation in USP2-ARID2 regulation, MG132 was then applied in A549 cells transfected with USP2 overexpression vectors."
USP2 increases the amount of ARID2.
| 4
USP2 increases the amount of ARID2. 4 / 4
| 4

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"USP2 Modulated ARID2 Protein Level via Ubiquitination Suppression and Subsequent ARID2 Protein Stabilization."

reach
"The USP2 overexpression vector transfection enhanced ARID2 protein expression."

reach
"The result of the anti-ARID2 antibody precipitated protein indicated that USP2 silencing significantly enhanced the ubiquitination level of ARID2 in H1299 cells, while USP2 overexpression significantly reduced the ubiquitination level of ARID2 in A549 cells (Figures 3(e) and 3(f))."

reach
"In summary, the above evidence supported our theory that USP2 enhances cellular ARID2 protein level through its inhibitory function on ARID2 protein degradation via the ubiquitination pathway."
USP2 ubiquitinates ARID2.
| 1
USP2 ubiquitinates ARID2. 1 / 1
| 1

reach
"Additionally, USP2 inhibits lung cancer through ubiquitinating ARID2 protein [20]."
USP2 affects DCAF1
4 | 5 7
USP2 binds DCAF1.
4 | 3 7
4 | 3 7

reach
"This interaction appears to be direct since purified Flag-VPRBP binds a purified GST-USP2 fusion protein but not GST alone (Fig. 5d)."

reach
"Collectively, these results indicate that USP2 binds, deubiquitinates, and thereby stabilizes VPRBP in vivo (Supplementary Fig. 1)."

sparser
"Interestingly, we found that the USP2-VPRBP axis modulates PD-L1 expression in a p53-independent manner through two distinct mechanisms (Supplementary Fig.  xref )."

reach
"Here we show that VPRBP levels are tightly controlled by USP2, a deubiquitinase that binds and stabilizes VPRBP, and that loss of USP2 function can activate p53 responses, including its tumor suppressive effects."

No evidence text available

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sparser
"Interestingly, the USP2-VPRBP axis also regulates PD-L1 (programmed death-ligand 1) expression."

sparser
"As both mechanisms serve to reduce PD-L1 levels (Supplementary Fig.  xref ), the USP2-VPRBP axis should suppress PD-L1-mediated immune evasion and thereby enhance tumor immunosurveillance."

No evidence text available

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USP2 increases the amount of DCAF1.
| 2
USP2 increases the amount of DCAF1. 2 / 2
| 2

reach
"Moreover, RNAi-mediated depletion of endogenous USP2 reduced the levels of endogenous VPRBP protein (Fig. 5g), but not VPRBP mRNA (Fig. 5h), in both H1299 and Cal-33 cells."

reach
"Endogenous VPRBP protein levels were also significantly decreased in H1299 cells by three independent USP2-specific siRNAs (Fig. 5i)."
USP2 affects CCND1
5 1 | 8
USP2 binds CCND1.
5 | 8
5 | 8

No evidence text available

sparser
"USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin-dependent degradation."

sparser
"We next investigated whether USP2 can directly interact with cyclin D1 upon co-expression of both proteins in 293 cells."

sparser
"Finally, the interaction of USP2 and cyclin D1 was tested in vitro using GST pulldown assays."

sparser
"These data demonstrate a direct interaction between USP2 and cyclin D1."

sparser
"Moreover, USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin-dependent degradation."

sparser
"USP2 directly interacts with cyclin D1 to perform deubiquitylation and thereby antagonize proteasome-dependent degradation of cyclin D1."

sparser
"USP2 directly interacted with cyclin D1 and decreased polyubiquitination-dependent degradation [ xref ]."

sparser
"By using co-immunoprecipitation approaches, we tried to determine whether elevation of O -GlcNAcylation favors the binding of USP2 to cyclin D1, as described for BAP1 and PGC1α ( xref )."

No evidence text available
USP2 deubiquitinates CCND1.
1 |
USP2 deubiquitinates CCND1. 1 / 1
1 |

"<span class="match term0">Usp2</span> is a deubiquitination enzyme targeting various factors including <span class="match term1"> CyclinD1</span> in cancer cells and PTH receptor 1 in osteoblasts"
USP2 affects FASN
6 1 | 3 3
USP2 binds FASN.
6 | 2 3
6 | 2 3

reach
"For example, USP-2 interacts and stabilizes fatty acid synthase which is overexpressed in prostate cancer cells (Graner et al., 2004) or causes murine double minute 2 (MDM2) accumulation and enhance M[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Among these, USP2 interacts with FASN and promotes its protein stability by reducing the ubiquitination level of FASN in prostate cancer [ xref ]."

No evidence text available

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reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

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sparser
"USP2 interacts with FASN in the mouse liver, human hepatic cancer, and HepG2 cells, and subsequently prevents the ubiquitination-dependent digestion of FASN [ xref , xref ]."
USP2 activates FASN.
| 1
USP2 activates FASN. 1 / 3
| 1

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"Usp2 enhances the stability of fatty acid synthase (FASN) by impeding proteasome-dependent degradation in human HCC (Calvisi et al., 2011; Kitamura and Hashimoto, 2021)."
USP2 deubiquitinates FASN.
1 |
USP2 deubiquitinates FASN. 1 / 1
1 |

"Here we show that the isopeptidase <span class="match term0">USP2a</span> (ubiquitin-specific protease-2a) interacts with and stabilizes fatty acid synthase (<span class="match term1">FAS</span>), "
CCND1 affects USP2
5 | 8
5 | 8

No evidence text available

sparser
"USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin-dependent degradation."

sparser
"We next investigated whether USP2 can directly interact with cyclin D1 upon co-expression of both proteins in 293 cells."

sparser
"Finally, the interaction of USP2 and cyclin D1 was tested in vitro using GST pulldown assays."

sparser
"These data demonstrate a direct interaction between USP2 and cyclin D1."

sparser
"Moreover, USP2 directly interacts with cyclin D1 and promotes its stabilization by antagonizing ubiquitin-dependent degradation."

sparser
"USP2 directly interacts with cyclin D1 to perform deubiquitylation and thereby antagonize proteasome-dependent degradation of cyclin D1."

sparser
"USP2 directly interacted with cyclin D1 and decreased polyubiquitination-dependent degradation [ xref ]."

sparser
"By using co-immunoprecipitation approaches, we tried to determine whether elevation of O -GlcNAcylation favors the binding of USP2 to cyclin D1, as described for BAP1 and PGC1α ( xref )."

No evidence text available
USP2 affects E2F4
6 | 2 6
6 | 2 6

sparser
"Emetine, an autophagy inhibitor, can block the interaction between USP2 and E2F4 and promote E2F4 degradation for an oncogenic effect, which can be reversed upon USP2 overexpression ( xref )."

sparser
"In clinical gastric cancer specimens, both USP2 and E2F4 were upregulated and associated with poor outcome of patients."

No evidence text available

sparser
"These findings indicate that therapeutic targeting of the USP2-E2F4 axis inhibits autophagic machinery essential for zinc homeostasis in cancer progression."

No evidence text available

sparser
"By blocking the USP2-E2F4 interaction, emetine inhibited autophagy and GC aggressiveness, suggesting the therapeutic potential of targeting the USP2-E2F4 axis [ xref ]."

No evidence text available

No evidence text available

sparser
"It has been reported that E2F4 binds directly with USP2 (ubiquitin specific peptidase 2) in gastric cancer cells [ xref ], while USP2 could bind with SKP2 and USP2-stabilized SKP2 did not destabilize its substrate P21 [ xref ]."

No evidence text available
USP2 affects CTNNB1
6 | 6 2
USP2 binds CTNNB1.
6 | 3 2
6 | 3 2

No evidence text available

No evidence text available

sparser
"The deubiquitinating enzyme USP2 interacts with β-catenin to deubiquitinate β-catenin without degradation, which in turn activates Wnt/β-catenin signaling xref ."

No evidence text available

reach
"GST pull-down assay and Co-IP assay verified the interaction between USP2 and β-catenin (Figure 6B,C)."

reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

reach
"Furthermore, USP2 interacted with beta-catenin, thereby regulating its deubiquitination and stabilization in CFs."

No evidence text available

No evidence text available
USP2 deubiquitinates CTNNB1.
| 3
USP2 deubiquitinates CTNNB1. 3 / 3
| 3

reach
"Xu et al. found 29 that the expression of the deubiquitinating enzyme USP2 was elevated in angiotensin II-induced cardiac fibroblasts, and that its interaction with β-catenin prevented β-catenin deubiquitination from being degraded, thereby promoting the activation, proliferation, and collagen synthesis of cardiac fibroblasts."

reach
"Additionally, USP2-induced Wnt/beta-catenin signalling pathway activation via deubiquitination of beta-catenin protein."

reach
"The deubiquitinating enzyme USP2 interacts with β-catenin to deubiquitinate β-catenin without degradation, which in turn activates Wnt/β-catenin signaling 29."
USP2 affects BIRC5
1 | 8 5
USP2 binds BIRC5.
1 | 3 5
1 | 3 5

reach
"In addition, USP2 bound to survivin, and ML364 increased ubiquitination of survivin."

No evidence text available

sparser
"Mechanistically, USP2 interacted with survivin, and ML364 degraded survivin protein expression by increasing the ubiquitination of survivin."

sparser
"To investigate the interaction between USP2 and survivin, we performed immunoprecipitation (IP) assay using a USP2 antibody."

sparser
"USP2 bound to survivin ( xref A)."

sparser
"In addition, USP2 interacted with survivin and inhibition of USP2 by ML364-induced ubiquitination of survivin ( xref A,B)."

sparser
"In addition, USP2 bound to survivin, and ML364 increased ubiquitination of survivin."

reach
"USP2 Interacts with and Deubiquitinates Survivin."

reach
"To investigate the interaction between USP2 and survivin, we performed immunoprecipitation (IP) assay using a USP2 antibody."
USP2 increases the amount of BIRC5.
| 3
USP2 increases the amount of BIRC5. 3 / 3
| 3

reach
"The pharmacological inhibitor (ML364) and siRNA of USP2 downregulated survivin expression at the post-translational level."

reach
"ML364 or USP2 siRNA in the presence of CHX further decreased survivin protein levels compared to CHX alone (Figure 2C,E)."

reach
"Therefore, these data showed that pharmacological and genetic inhibition of USP2 downregulate survivin expression in many cancer cells."
USP2 activates BIRC5.
| 2
USP2 activates BIRC5. 2 / 2
| 2

reach
"Therefore, these results indicated that survivin protein stability is modulated by the expression level of USP2."

reach
"We observed that USP2 WT rescues survivin downregulation through USP2 siRNA (Figure 1D)."
TP53 affects USP2
| 6 8
TP53 binds USP2.
| 2 8
| 2 8

sparser
"In vivo experiments in mice explored the specific role of the USP2-p53 axis in a xenograft tumor model."

sparser
"Although this study provides a preliminary exploration of the role of the USP2-p53 axis-mediated ferroptosis in reversing cisplatin resistance in NSCLC, several limitations persist."

sparser
"Subsequently, COIP was utilized to identify the binding of USP2 and p53, which showed the direct binding between USP2 and p53 (Fig.  xref B)."

sparser
"In our study, USP2 overexpression inhibited p53 ubiquitination and directly interacted with p53."

sparser
"USP2, which is usually suppressed in cisplatin-resistant NSCLC, can directly interact with p53 to promote its stability and nuclear translocation, eventually facilitating ferroptosis xref ."

reach
"Moreover, USP2 directly bound to p53 and USP2 overexpression stabilized p53 protein by suppressing its ubiquitination."

reach
"Subsequently, COIP was utilized to identify the binding of USP2 and p53, which showed the direct binding between USP2 and p53 (Fig. 5B)."

sparser
"Our study preliminarily unveils a novel mechanism of DDP resistance in NSCLC, highlighting the critical role of the USP2-p53 axis in regulating cisplatin sensitivity in NSCLC cells."

sparser
"Moreover, USP2 directly bound to p53 and USP2 overexpression stabilized p53 protein by suppressing its ubiquitination."

sparser
"This study revealed that USP2 and p53 were associated with NSCLC cisplatin resistance."
TP53 activates USP2.
| 3
TP53 activates USP2. 3 / 3
| 3

reach
"These results suggested that activated p53 increased USP2, which in turn counteracted the pro apoptotic activity of p53, forming a negative feedback loop."

reach
"USP2 is upregulated by PUVA and a p53 activator, nutlin3a."

reach
"It would be interesting to investigate whether USP2 aggravates the resistance to other therapeutic modalities, such as ionizing radiation or ultraviolet B. Mechanistically we were able to demonstrate that the activation of p53 upregulated USP2."
TP53 inhibits USP2.
| 1
TP53 inhibits USP2. 1 / 1
| 1

reach
"Interestingly, leptin induced increase in p53 expression was significantly suppressed by gene silencing of USP2 in both HepG2 and MCF-7 cells (XREF_FIG), suggesting the crucial role of USP2 and probably modulation of ubiquitination in leptin induced p53 expression."
PER1 affects USP2
| 6 8
| 6 8

sparser
"These findings show that USP2 binds PER1 directly and indicate that the interaction of USP2 with the other clock proteins is indirect and possibly occurring within a complex that contains components known to participate in the main negative feedback loop of the circadian clock, including PER1."

sparser
"Earlier studies from our lab showed that USP2 directly interacts with ubiquitinated PER1 (Yang et al.  xref ) and facilitates its localization to the nucleus (Yang et al.  xref )."

reach
"USP2 can be coimmunoprecipitated with several clock proteins but directly interacts specifically with PER1 and deubiquitinates it."

sparser
"In our hands, whereas it forms a complex with several clock proteins, USP2 directly binds only to PER1 ( xref )."

reach
"These findings show that USP2 binds PER1 directly and indicate that the interaction of USP2 with the other clock proteins is indirect and possibly occurring within a complex that contains components known to participate in the main negative feedback loop of the circadian clock, including PER1."

reach
"For example, USP2 binds directly to PER1 in order to regulate PER1 intracellular localization [XREF_BIBR]."

sparser
"Previous studies have shown that USP2 directly interacts with PER1 and affects its localization (Yang et al.  xref )."

sparser
"However, in vitro assays indicated that USP2 could only interact directly with PER1, suggesting that USP2 interacts with the other clock proteins by virtue of their being in a complex with PER1 ( xref )."

sparser
"The USP2 protein binds and deubiquitinates PER1, regulating its nuclear localization [ xref , xref ]."

sparser
"For example, USP2 binds directly to PER1 in order to regulate PER1 intracellular localization [ xref ]."
USP2 is modified
| 9 5
USP2 is phosphorylated.
| 6 5
USP2 is phosphorylated. 7 / 7
| 6 1

sparser
"Phosphorylation of two critical residues in USP2 N terminus is required for its recruitment to DSB sites and interaction with RECQL4."

sparser
"As the N-terminal domain of USP2 showed ATM-dependent recruitment to DSB sites, we further investigated whether phosphorylation of USP2 N terminus is important for its recruitment to DSB sites."

sparser
"We then examined whether the phosphorylation of USP2 influences its interaction with RECQL4."

sparser
"Western blotting showed that supernatant from USP2 KD cells repressed insulin-elicited phosphorylation of Akt in 3T3-L1 cells significantly more than media from control HL-60 cells ( xref A)."

sparser
"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."

rlimsp
"However, USP12 was still able to exert an indirect effect on AR Vs by controlling its phosphorylation through AKT activation (Figure 7F), confirming that AR Vs are still phosphorylated at S213 while this phosphorylation is most likely to be of no consequence for AR V activity as phosphorylation at this site reportedly decreases AR-androgen binding and targets AR for ubiquitination by MDM2 [44], both processes that seem irrelevant to AR V biology."

sparser
"Live-cell imaging of cells expressing EGFP-fused USP2 after laser microirradiation, mapping of domains responsible for recruitment to DSB sites, and analyses of phospho-deficient and phospho-mimetic mutants of USP2 showed that USP2 is recruited to DSB sites in a ATM-dependent manner, and phosphorylation of two critical residues in the N terminus of USP2 is essential for its recruitment to DSBs and interaction with RECQL4."
USP2 is phosphorylated on S57. 2 / 2
| 2

rlimsp
"Ser57 phosphorylation increased USP2-mediated cleavage of K29-linked Ub, but it decreased USP2-mediated cleavage of K6 and K48 linkages (Figure 6K)."

rlimsp
"The cleavage of K29-linked dimers by USP2 was activated by phosphorylation at Ser57, but the cleavage of K29-linked dimers of Ub by USP2 was deactivated by Ser65 phosphorylation (Figures 5D and 6E)."
USP2 is phosphorylated. 2 / 2
| 2

rlimsp
"(G) Phosphorylation on the distal Ub in USP2 may inhibit cleavage of all Ub linkages."

rlimsp
"Ser65 of the distal Ub is buried at the interface with USP2, and we suggest that phosphorylation of USP2 may control the cleavage of all Ub linkages by inhibiting binding of the distal Ub (Figure 7G)."
USP2 is ubiquitinated.
| 3
USP2 is ubiquitinated. 3 / 3
| 3

sparser
"In addition, cancer-associated SPOP mutants showed reduced activity for promoting USP2 ubiquitination in cells ( xref )."

sparser
"We further confirmed that SPOP could promote the ubiquitination of wild-type USP2, but not the degron 2 mutant form of USP2 ( xref )."

sparser
"Although we have identified USP2 as a pivotal DUB enzyme that deubiquitinates and stabilizes the CD47 protein, the physiological ubiquitin E3 ligase responsible for promoting USP2 ubiquitination and subsequent degradation remains largely unknown."
ML364 affects USP2
| 14
ML364 inhibits USP2.
| 11
ML364 inhibits USP2. 10 / 11
| 11

reach
"Given that ML364 inhibits USP2, we next investigated the role of USP2 in antithrombin turnover."

reach
"We demonstrated that pharmacological inhibition of USP2 by ML364 prominently reduced the formation of TNBC tumorspheres in a dose dependent fashion."

reach
"Moreover, USP2 inhibition by ML364 considerably enhanced the cytotoxicity of doxorubicin or paclitaxel in multiple TNBC cell models."

reach
"We reasoned that while ML364 inactivated USP2 catalytic activity, this small molecule is unlikely to disrupt the USP2–SKP2 binding, leading to less “free” SKP2 to degrade its substrates."

reach
"USP2 overexpression promoted proliferation and metastasis, and induced the EMT in CM cells in vitro, while specific inhibition of USP2 by ML364 produced the opposite effects."

reach
"In support of this notion, USP2 inhibition by ML364 led to a dose dependent reduction in Twist and Bmi1 expression."

reach
"ML364 is an inhibitor of ubiquitin specific peptidase 2 (USP2) in human breast cancer (Davis et al., 2016)."

reach
"The small-molecular inhibitor of USP2, ML364, was purchased from TargetMol."

reach
"Together, these observations provide compelling evidence that ML364, by inhibiting USP2 and subsequently reducing ACE2 protein, effectively counteracts SARS-CoV-2 infection both in vitro and in vivo."

reach
"However, the commercially available inhibitors for these DUBs had no obvious influence on the SOX2 expression, including b-AP15 (the inhibitor of USP14 and UCHL5), P5091 (the inhibitor of USP7 and USP[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
ML364 binds USP2.
| 2
ML364 binds USP2. 2 / 2
| 2

reach
"Direct binding of ML364 to USP2 was demonstrated using microscale thermophoresis."

reach
"ML364 directly binds to USP2 and selectively inhibits its enzymatic activity 31."
ML364 activates USP2.
| 1
ML364 activates USP2. 1 / 1
| 1

reach
"These effects of ML364 support the possibility of USP2 as a therapeutic target in cancer therapy."
E2F4 affects USP2
6 | 2 6
6 | 2 6

sparser
"Emetine, an autophagy inhibitor, can block the interaction between USP2 and E2F4 and promote E2F4 degradation for an oncogenic effect, which can be reversed upon USP2 overexpression ( xref )."

sparser
"In clinical gastric cancer specimens, both USP2 and E2F4 were upregulated and associated with poor outcome of patients."

No evidence text available

sparser
"These findings indicate that therapeutic targeting of the USP2-E2F4 axis inhibits autophagic machinery essential for zinc homeostasis in cancer progression."

No evidence text available

sparser
"By blocking the USP2-E2F4 interaction, emetine inhibited autophagy and GC aggressiveness, suggesting the therapeutic potential of targeting the USP2-E2F4 axis [ xref ]."

No evidence text available

No evidence text available

sparser
"It has been reported that E2F4 binds directly with USP2 (ubiquitin specific peptidase 2) in gastric cancer cells [ xref ], while USP2 could bind with SKP2 and USP2-stabilized SKP2 did not destabilize its substrate P21 [ xref ]."

No evidence text available
DCAF1 affects USP2
4 | 3 7
4 | 3 7

reach
"This interaction appears to be direct since purified Flag-VPRBP binds a purified GST-USP2 fusion protein but not GST alone (Fig. 5d)."

reach
"Collectively, these results indicate that USP2 binds, deubiquitinates, and thereby stabilizes VPRBP in vivo (Supplementary Fig. 1)."

sparser
"Interestingly, we found that the USP2-VPRBP axis modulates PD-L1 expression in a p53-independent manner through two distinct mechanisms (Supplementary Fig.  xref )."

reach
"Here we show that VPRBP levels are tightly controlled by USP2, a deubiquitinase that binds and stabilizes VPRBP, and that loss of USP2 function can activate p53 responses, including its tumor suppressive effects."

No evidence text available

No evidence text available

sparser
"Interestingly, the USP2-VPRBP axis also regulates PD-L1 (programmed death-ligand 1) expression."

sparser
"As both mechanisms serve to reduce PD-L1 levels (Supplementary Fig.  xref ), the USP2-VPRBP axis should suppress PD-L1-mediated immune evasion and thereby enhance tumor immunosurveillance."

No evidence text available

No evidence text available
USP2 affects TWIST1
| 10 3
USP2 binds TWIST1.
| 3 3
| 3 3

reach
"We carried out a co-immunoprecipitation experiment and showed a strong interaction between endogenous USP2 and Twist in BT549 cells."

sparser
"We carried out a co-immunoprecipitation experiment and showed a strong interaction between endogenous USP2 and Twist in BT549 cells (Fig.  xref )."

reach
"Further characterization of the interaction between USP2 and Twist uncovered that the N-terminal domain of USP2 mediates its binding to Twist."

reach
"We demonstrate that TJP1 functions as a scaffold for the interaction between USP2 and TWIST1 and this may provide potential therapeutic targets in bladder cancer."

sparser
"We demonstrate that TJP1 functions as a scaffold for the interaction between USP2 and TWIST1 and this may provide potential therapeutic targets in bladder cancer."

sparser
"Further characterization of the interaction between USP2 and Twist uncovered that the N-terminal domain of USP2 mediates its binding to Twist (Supplementary Figure  xref )."
USP2 activates TWIST1.
| 3
USP2 activates TWIST1. 3 / 3
| 3

reach
"Indeed, MG132 treatment reverted Twist downregulation mediated by USP2 depletion."

reach
"In addition, USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats containing proteins (beta-TrCP), a cellular E3 ubiquitin ligase."

reach
"Our work reveals that USP2 mediated Twist and Bmi1 pathway represents a cell-intrinsic mechanism crucial for CSC regulation besides EMT and provides pre-clinical evidence that targeting USP2 is indeed a promising approach for anti-CSC therapy."
USP2 deubiquitinates TWIST1.
| 2
USP2 deubiquitinates TWIST1. 2 / 2
| 2

reach
"Furthermore, TJP1 recruited USP2, which deubiquitinated TWIST1, thereby protecting TWIST1 from proteasome-mediated protein degradation."

reach
"USP2 regulates Bmi1 and EMT by Twist stabilization and prevents Twist ubiquitination driven by beta-TrCP and subsequent proteasome mediated protein degradation."
USP2 increases the amount of TWIST1.
| 1
Modified USP2 increases the amount of TWIST1. 1 / 1
| 1

reach
"Altogether, our study illustrates that USP2 overexpression in TNBC promotes Twist and Bmi1 expression, CSC expansion, cancer cell migration and tumor development."
USP2 decreases the amount of TWIST1.
| 1
Modified USP2 decreases the amount of TWIST1. 1 / 1
| 1

reach
"Western blot assays illustrated that ablating USP2 expression in BT549 and MDA-MB-157 cells considerably reduced protein expression of Twist as expected."
USP2 affects LDLR
1 | 1 9
USP2 activates LDLR.
| 3
USP2 activates LDLR. 3 / 5
| 3

reach
"Conversely, loss of USP2 reduces LDLR protein in an IDOL dependent manner and limits LDL uptake."

reach
"USP2 promoting LDLR activity was not simply an artefact of overexpression, as convincingly demonstrated by the Usp2 siRNA experiments, which showed a 70% reduction in surface LDLR levels and a halving of LDL uptake."

reach
"USP2 not only interacts with and stabilises the sterol regulated E3-ligase IDOL but also rescues LDLR from lysosomal degradation, recalling the example of USP7 described in detail below 53."
USP2 deubiquitinates LDLR.
| 4
USP2 deubiquitinates LDLR. 4 / 4
| 4

reach
"We identify a tri and partite complex encompassing IDOL, USP2, and LDLR and demonstrate that in this context USP2 promotes deubiquitylation of the LDLR and prevents its degradation."

reach
"The authors attempt to answer this question by introducing a tripartite complex model, in which USP2 interacts with LDLR at the plasma membrane in an IDOL dependent manner to deubiquitylate and stabilize both LDLR and IDOL."

reach
"Such a temporal expression pattern would leave little opportunity to alleviate the USP2 mediated antagonism of IDOL, and the tripartite complex model would imply that IDOL is constantly engaged in a futile reaction, in which its ubiquitylation of LDLR is immediately reversed by USP2."

reach
"In summary, USP2 activity leads to the deubiquitylation and stabilization of cell surface LDLR in and IDOL dependent manner, but it remains uncertain exactly why such a phenomenon is accompanied by an extended IDOL protein half-life."
USP2 inhibits LDLR.
| 1 1
USP2 inhibits LDLR. 2 / 2
| 1 1

eidos
"Both USP2-1 and USP2-4 deubiquitinate and stabilize IDOL , suggesting that USP2 promotes LDLR degradation [ 110 ] ."

reach
"Using a genetic screening approach, we identify the ubiquitin specific protease 2 (USP2) as a post-transcriptional regulator of IDOL mediated LDLR degradation."
USP2 binds LDLR.
1 | 1
1 | 1

No evidence text available

reach
"The authors attempt to answer this question by introducing a tripartite complex model, in which USP2 interacts with LDLR at the plasma membrane in an IDOL dependent manner to deubiquitylate and stabilize both LDLR and IDOL."
USP2 affects Imd
| 13
USP2 binds Imd.
| 5
| 5

reach
"The control of Imd homeostasis by USP2 is associated with the hydrolysis of Imd linked K48-ubiquitin chains and the synergistic binding of USP2 and Imd to the proteasome, as evidenced by both mass-spectrometry analysis of USP2 partners and by co-immunoprecipitation experiments."

reach
"Focusing on USP2 biochemical function, we show that USP2 binds to Imd and promotes the cleavage of Ub K48 chains from the protein in both cultured cells and flies."

reach
"USP2 interacts with Imd and promotes cleavage of Imd linked Ub K48."

reach
"By using GST pull-down assays with GST tagged full length or truncated forms of USP2 (USP2-N-ter [AA :1-531] and USP2-C-ter [AA :475-856]), we finally showed that USP2 preferentially interacts with Imd through its non catalytic N-ter domain."

reach
"Thus, the Imd and USP2 complex apparently binds more efficiently to the proteasome than do Imd or USP2 alone."
USP2 inhibits Imd.
| 4
USP2 inhibits Imd. 4 / 4
| 4

reach
"However, while USP2 is also required to prevent excessive activation of the Imd pathway in infected flies, USP34 displays differential requirement depending on the antimicrobial peptide gene analysed : silencing Usp34 enhanced the activation of AttA, Drs and IM1."

reach
"However, while USP2 also prevents excessive Imd dependent signalling in vivo, USP34 shows differential requirement depending on NF-kappaB target genes, in response to fly infection by either Gram positive or Gram negative bacteria."

reach
"USP2 promotes Imd degradation by the proteasome."

reach
"We report here that USP2 and USP34 and Puf, in addition to the previously described USP36 and Scny, prevent inappropriate activation of Imd dependent immune signal in unchallenged conditions."
USP2 deubiquitinates Imd.
| 2
USP2 deubiquitinates Imd. 2 / 2
| 2

reach
"Identifying USPs regulating immune signals in Drosophila : USP2 deubiquitinates Imd and promotes its degradation by interacting with the proteasome."

reach
"USP2, for instance, deubiquitinates Imd, promoting its degradation."
USP2 activates Imd.
| 2
USP2 activates Imd. 2 / 2
| 2

reach
"We further show that USP2 prevents the constitutive activation of signalling by promoting Imd proteasomal degradation."

reach
"Surprisingly, USP2 also targets Imd for degradation, a function which, from our proteomic analysis of USP2 partners and subsequent co-immunoprecipitation experiments, likely occurs at the level of the proteasome."
USP2 affects CD47
| 13
| 13

sparser
"Mechanistically, USP2 interacts with CD47, stabilizing it through deubiquitination."

sparser
"Mechanistically, USP2 interacts with CD47 and removes ubiquitination on CD47, leading to the stabilization of CD47 (Fig.  xref )."

sparser
"Meanwhile, we observed a strong endogenous interaction between USP2 and CD47 in both PC9 and H1975 lung cancer cells (Fig.  xref )."

sparser
"In this study, our findings reveal that USP2 interacts with CD47, facilitating the deubiquitination of CD47 and leading to the stabilization of the CD47 protein."

sparser
"Collectively, these results demonstrate that USP2 interacts with CD47 in cells."

sparser
"Considering USP2 as a deubiquitinating enzyme, we hypothesized that USP2 might interact with CD47 to erase the ubiquitination on CD47."

sparser
"To further dissect the mechanism of how USP2 regulates the expression of CD47, we performed co-immunoprecipitation assays and found that USP2 interacted with CD47 ( xref , xref )."

sparser
"CD47 mainly interacted with the N-terminal domain of USP2, but not with the C-terminal USP domain ( xref )."

sparser
"The results demonstrated that the N-terminal domain, but not the C-terminal domain of USP2, interacted with CD47 (Fig.  xref )."

sparser
"These findings identify the USP2-CD47 axis as a critical driver of tumor immune evasion and anti-PD-1 therapy resistance."
ARID2 affects USP2
3 | 1 9
3 | 1 9

sparser
"USP2 Is Significantly Suppressed and Interacts with ARID2 in Lung Cancer Cells."

sparser
"Based on the above findings, we then utilized HEK-293T cell line as a model to explore and confirm the potential interaction of USP2-ARID2."

reach
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure 1 B) and interactive Flag-ARID2 (Supplementary Figure 1 C) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

No evidence text available

sparser
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure xref ) and interactive Flag-ARID2 (Supplementary Figure xref ) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

No evidence text available

sparser
"The results indicated that in both A549 and H1299 cells, ARID2 protein can be detected in anti-USP2 antibody precipitated protein samples, and vice versa ( xref ), suggesting endogenously USP2-ARID2 protein-protein interaction (PPI) in lung cancer cells."

No evidence text available

sparser
"The quantitative analysis of the USP2-ARID2 colocalization ratio also provided consistent results (Figures xref and xref )."

sparser
"To explore the potential role of proteasome-dependent protein degradation in USP2-ARID2 regulation, MG132 was then applied in A549 cells transfected with USP2 overexpression vectors."

reach
"Ubiquitin-specific peptidase 2 inhibited epithelial-mesenchymal transition in clear cell renal cell carcinoma cells."

reach
"Ubiquitin-specific peptidase 2-mediated inhibition of epithelial-mesenchymal transition in clear cell renal cell carcinoma cells is dependent on the nuclear factor-kappaB pathway."

reach
"The results from our in vitro and in vivo research demonstrated that ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition in clear cell renal cell carcinoma metastasis by downregulating the NF-κB pathway."

reach
"Hence, ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition of clear cell renal cell carcinoma by downregulating the NF-κB pathway."

reach
"Additionally, ubiquitin-specific peptidase 2 was found to negatively regulate the epithelial-mesenchymal transition pathway by RNA sequencing results."

reach
"The results from both in vitro and in vivo showed that ubiquitin-specific peptidase 2 inhibited epithelial-mesenchymal transition by downregulating the NF-κB pathway."

reach
"Ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition in clear cell renal cell carcinoma metastasis by downregulating the NF-kappaB pathway."

reach
"It has been demonstrated that the TGF-β pathway has an effective role in metastasis; for instance, USP2 promotes EMT after interacting with TGFBR1 and TGFBR2 after TGF-β stimulation[76]."

reach
"Likewise, ubiquitin-specific peptidase 2 (USP2) downregulates the NF-κB pathway, inhibiting EMT in clear cell renal cell carcinoma metastasis (Duan et al., 2022)."

reach
"USP2 overexpression promoted proliferation and metastasis, and induced the EMT in CM cells in vitro, while specific inhibition of USP2 by ML364 produced the opposite effects."

eidos
"In addition , USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats-containing proteins ( beta-TrCP ) , a cellular E3 ubiquitin ligase ( 104 ) ."

reach
"In addition, USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats containing proteins (beta-TrCP), a cellular E3 ubiquitin ligase."
USP2 affects cell cycle
| 3 9
USP2 activates cell cycle.
| 3 7
| 3 7

eidos
"In addition , USP2 inhibition suppressed CFs proliferation , collagen synthesis and cell cycle progression ."

reach
"Taken together, these results demonstrate that USP2 mediated cell cycle control occurs primarily through its effect on cyclin D1, regardless of p53 status."

reach
"USP2 modulates cell cycle progression, and therefore carcinogenesis, via the deubiquitination of cyclins and Aurora-A."

reach
"USP2 stabilizes cyclin D1 and promotes cell cycle progression [31]."

eidos
"USP2 modulates cell cycle progression , and therefore carcinogenesis , via the deubiquitination of cyclins and Aurora-A ."

reach
"In addition, USP2 inhibition suppressed CFs proliferation, collagen synthesis and cell cycle progression."

reach
"To test this hypothesis, we first examined cell cycle distribution in cells treated with control versus USP2 siRNAs."

eidos
"USP2 inhibition causes cell cycle arrest in activated cardiac fibroblasts ."

reach
"USP22 and USP2 promote cell cycle progression by stabilizing cyclins ( Shan et al., 2009; Gennaro et al., 2018 )."

reach
"USP2 can promote cell cycle progression through its effect on cyclin D1."
USP2 inhibits cell cycle.
| 2
| 2

reach
"Results from gene expression profiling assays suggested that knockout of USP2 resulted in upregulation of cell cycle and DNA replication pathways in lamina propria myeloid cells after DSS treatment."

reach
"Strikingly, ectopic overexpression of cyclin D1 in USP2-knockdown cells effectively reverses the changes in cell cycle distribution caused by USP2 knockdown (XREF_FIG)."
| 7

reach
"To summarize, we confirmed that USP2 suppressed cancer cell migration and invasion capabilities through ARID2."

reach
"Ubiquitin-specific peptidase 2 inhibited the proliferation, migration, and invasion of clear cell renal cell carcinoma cells."

reach
"Functional experiments suggested USP2 inhibited lung cancer cell invasion and migration by reducing ARID2 protein degradation."

reach
"Ubiquitin-specific peptidase 2 overexpression prevented the migration and invasion ability of clear cell renal cell carcinoma cells (Figures 3(d,e))."

reach
"Through our experiments, the results showed that ubiquitin-specific peptidase 2 negatively regulated the migration and invasion abilities in clear cell renal cell carcinoma cell lines."

reach
"Ubiquitin-specific peptidase 2 inhibition promoted clear cell renal cell carcinoma cell proliferation, migration, and invasion."

reach
"Ubiquitin-specific peptidase 2 overexpression inhibited clear cell renal cell carcinoma cell proliferation, migration, and invasion in vitro and in vivo."
| 5

reach
"USP2 promotes cell migration and invasion in triple negative breast cancer cell lines."

reach
"As shown in Figures 4(a) and (b), USP2 shRNA transfection uniformly enhanced H1299 and A549 cell migration and invasion, while USP2 overexpression vector transfection significantly suppressed the migrative and invasive capability of cancer cells.Subsequent wound healing assay suggested that H1299 cells with USP2 shRNAs transfection elevated cancer cell migration and wound healing after 48 h of incubation (Supplementary Figure 2 A-B)."

reach
"Previously, USP2 was revealed to enhance tumor migration and invasion by increasing MMP-2 activity in metastatic triple-negative breast cancer (101)."

reach
"In breast cancer, overexpression of USP2 in TNBC cell lines MDA-MB-468 and MDA-MB-231 enhances migration and invasion with the elevated MMP2 protein level."

reach
"It is upregulated in various malignancies, such as triple negative breast cancer [17] and choroidal melanoma [18] and USP2 overexpression stimulates cell migration and invasion."
USP2 affects MDM4
5 1 | 3 2
USP2 binds MDM4.
5 | 1 2
5 | 1 1

No evidence text available

No evidence text available

reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

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sparser
"Ubiquitin-specific protease 2a (USP2a) is a deubiquitinating enzyme that protects MDM4 against degradation, so USP2-MDM4 interaction may be a key determinant of the malignant potential of human cancers."
EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
| 1

sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
USP2 deubiquitinates MDM4.
1 | 2
USP2 deubiquitinates MDM4. 3 / 3
1 | 2

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"USP2 deubiquitinates both MDM2 and MDMX [XREF_BIBR, XREF_BIBR] whereas USP4 deubiquitinates ARF-BP1 [XREF_BIBR], another ubiquitin ligase for p53, thus indirectly destabilizing p53 and inhibiting its function."

reach
"MdmX, another target of Mdm2, is also deubiquitinated by USP2."

"<span class="match term0">USP2</span> deubiquitinates both Mdm2 and <span class="match term1">MdmX</span>"
PPARG affects USP2
| 4 8
PPARG binds USP2.
| 8
| 8

sparser
"To further investigate whether USP2 directly interacts with PPAR-γ, we performed endogenous and exogenous co-immunoprecipitation (Co-IP) analysis."

sparser
"USP2 interacted with PPAR-γ in C2C12 cells and the Gas muscle ( xref )."

sparser
"Further immuno-precipitation analysis and pulldown assay also joined to confirm the interaction between USP2 and PPARγ ( xref )."

sparser
"Targeting the USP2PPAR-γ axis may offer promising therapeutic strategies for metabolic disorders and sarcopenia."

sparser
"In addition, the computational modeling was used to determine whether USP2 binds to PPARγ and which residues are required for their interaction."

sparser
"Data also showed the USP2-PPAR-γ interaction was key in mediating the effects of USP2 on muscle atrophy and insulin signaling ( xref and xref )."

sparser
"By identifying USP2 as a novel PPAR-γ deubiquitinase and protective factor, we have broadened the PPAR-γ regulatory network and revealed the potential of targeting the USP2PPAR-γ axis for skeletal muscle and insulin signaling disorders."

sparser
"These findings highlight the potential of targeting the USP2PPAR-γ axis for therapeutic interventions aimed at combating muscle atrophy and enhancing insulin sensitivity, thereby presenting novel avenues for addressing metabolic disorders and sarcopenia."
PPARG activates USP2.
| 4
PPARG activates USP2. 4 / 4
| 4

reach
"To directly demonstrate that PPAR-γ is necessary for USP2 to regulate muscle atrophy and insulin signaling, we infected C2C12 cells with AdUsp2 and/or siPPAR-γ."

reach
"Loss of PPAR-gamma abolishes the regulatory effects of USP2 on insulin sensitivity and muscle atrophy."

reach
"Loss of PPAR-gamma Blocks the Effects of USP2 on Insulin Sensitivity and Muscle Atrophy in Diabetic Mice."

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"PPAR-gamma Deficiency Eliminates the Effects of USP2 on Insulin Signaling and Muscle Atrophy In Vitro."
MDM2 affects USP2
5 1 | 3 2
5 1 | 3 2

reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

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reach
"The C-terminal region of USP2 forms part of the catalytic domain and binds to the C-terminal Zinc and Ring finger domain of mdm2, the same region of mdm2 that we demonstrated to bind to USP7.Crystal s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Specifically, protein-protein interaction assays using the bacterial two-hybrid system showed that USP2 can deubiquitinate MDM2 and promotes p53 degradation, showing the association between USP2 and MDM2 (Stevenson et al., 2007)."

sparser
"Subsequent analysis demonstrated that USP2 can bind and deubiquitinate Mdm2."
ETS1 affects USP2
| 8 4
ETS1 activates USP2.
| 5 2
ETS1 activates USP2. 7 / 7
| 5 2

sparser
"ETS1 transcriptionally activates USP2."

sparser
"To sum up, ETS1 transcriptionally activates USP2."

reach
"Moreover, ETS1 promoted the osteogenic differentiation of HBMSCs by up-regulating USP2."

reach
"ETS1 transcriptionally activates USP2."

reach
"To sum up, ETS1 transcriptionally activates USP2."

reach
"ETS1 up-regulates USP2 to induce the osteogenic differentiation of HBMSCs."

reach
"Therefore, it is confirmed that ETS1 can promote the osteogenic differentiation of HBMSCs by up-regulating USP2."
ETS1 binds USP2.
| 1 2
| 1 2

sparser
"In our study, we carried out assays and validated that ETS1 could bind to USP2 promoter region and activate USP2 transcription."

reach
"Therefore, we further conducted mechanism experiment to prove the interaction between ETS1 and USP2."

sparser
"Therefore, we further conducted mechanism experiment to prove the interaction between ETS1 and USP2."
ETS1 increases the amount of USP2.
| 2
ETS1 increases the amount of USP2. 2 / 2
| 2

reach
"Western blot analysis consistently indicated ETS1 knockdown could decrease USP2 protein level (Figure 3C)."

reach
"In our study, we carried out assays and validated that ETS1 could bind to USP2 promoter region and activate USP2 transcription."
USP2 affects USP2
| 9
USP2 inhibits USP2.
| 4
USP2 inhibits USP2. 4 / 4
| 4

reach
"Specifically, USP2 overexpression could markedly reduce elevated Nox2, Nox4, and p22phox at the mRNA level in pressure-overloaded hearts, indicating an unprecedented regulatory role of USP2 in mediating oxidative stress and Nox protein expression during cardiac pressure overload."

reach
"The above data suggested that USP2 overexpression attenuated calcium overload and mitochondrial dysfunction in cardiomyocytes of mice with cardiac hypertrophy.We further validated the role of USP2 in [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In summary, our data show that the ubiquitin core and the ubiquitin C terminus alone bind USP2 very weakly with affinities at best in the high micromolar range, while full-length ubiquitin inhibits USP2 with a K of 2.8 μM. Discussion."

reach
"Collectively, these results confirmed the ACE2 dependency in the inhibition of USP2 by ML364 or MS102, thereby resulting in their inhibitory effects on viral infections."
USP2 activates USP2.
| 3
USP2 activates USP2. 3 / 4
| 3

reach
"On the other hand, a defect of peroxisomal protein import or even release of USP2 from defective peroxisomes because of cell injury or dysfunction would increase the cytosolic pool of USP2, which might contribute to the initiation of apoptosis."

reach
"Altogether, we initially demonstrated that USP2 overexpression suppressed hypertrophy, calcium overload, and mitochondrial dysfunction and increased protein levels of MFN2 via deubiquitination in Ang [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"To further investigate the potential involvement of USP2 in cisplatin resistance in NSCLC through ferroptosis, USP2 expression was overexpressed and the impact on cellular ferroptosis was assessed."
USP2 decreases the amount of USP2.
| 2
USP2 decreases the amount of USP2. 1 / 2
| 1

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"Conversely, transfection with sh-USP2 effectively decreased the expression of USP2 (Fig. 2G)."
USP2 decreases the amount of USP2-C276A. 1 / 1
| 1

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"WT USP2 overexpression induced an increase in the protein level of PPAR-γ and decreased PPAR-γ ubiquitination, but not USP2 C276A (an enzyme-activity dead mutation) overexpression (Fig. 4C)."
USP2 affects TRAF2
8 1 | 2
8 1 | 2

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sparser
"USP2 interacts with TRAF2 and promotes deubiquitination of TRAF2, enhancing its stability."

No evidence text available

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USP2 affects RECQL4
2 | 2 7
2 | 2 7

sparser
"To support this notion, USP2 interacted with RECQL4 in coimmunoprecipitation experiments and NCS treatment significantly increased this interaction ( xref E )."

sparser
"The expression of helicase-defective RECQL4 mutant proteins (D605A and E606A in the Walker B motif) as well as WT RECQL4 proteins restored USP2 binding to laser-induced DSB sites in RECQL4-depleted cells."

sparser
"As shown in xref D , interaction between phospho-deficient USP2 (AA) and RECQL4 was not observed in the coimmunoprecipitation analysis."

sparser
"In contrast, phosphomimetic USP2 (EE) proteins stably interacted with RECQL4, both in the absence and presence of DNA damage ( xref D ), and inhibition of ATM kinase activity did not disrupt their interaction, although the interaction appeared to be slightly weak ( xref E )."

sparser
"Taken together, these results suggest that interaction of USP2 with RECQL4 through ATM-dependent phosphorylation of these two sites is not sufficient for the recruitment of USP2 to DSB sites."

sparser
"As shown in xref , xref , xref , USP2 stably interacted with RECQL4 in an ATM-dependent manner and recruitment of USP2 to DSB sites also requires RECQL4."

reach
"As shown in Figure 5D, interaction between phospho-deficient USP2 (AA) and RECQL4 was not observed in the coimmunoprecipitation analysis."

sparser
"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."

reach
"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."

No evidence text available
USP2 affects Neoplasms
| 11
| 11

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"Previously, USP2 was revealed to enhance tumor migration and invasion by increasing MMP-2 activity in metastatic triple-negative breast cancer (101)."

reach
"We found that USP2 knockout significantly suppressed tumor growth, as indicated by the smaller tumor volumes and weights in the knockout groups compared with the control group (Fig. 5E–G)."

reach
"These data indicate that a combination of USP2 inhibition and PD-1/PD-L1 blockade potently suppresses tumor growth in vivo, and that the contribution of USP2 inhibition to this effect is dependent on the tumor cell autonomous activity of p53."

reach
"As illustrated in Fig. 6A-C, the combination of USP2 overexpression and DDP treatment promoted tumor regression, evidenced by a significant reduction in tumor volume and notable decrease in weight."

reach
"USP2 promotes tumor immune evasion via deubiquitination and stabilization of PD-L1."

reach
"Pharmacological USP2 inhibition inhibits tumor progression in breast cancer and sensitizes tumor cells to chemotherapy.As well as the canonical EMT-associated markers discussed previously, matrix meta[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The combination of USP2 inhibition and the PD-1/PD-L1 immune checkpoint blockade completely suppresses tumor growth with no obvious toxicity."

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

reach
"Indeed, USP2 inhibition alone partially induces tumor growth suppression but more strikingly, the combination of USP2 inhibition and PD-1/PD-L1 blockade promote vigorous tumor regression and long-term survival of all tumor-bearing mice."

reach
"Moreover, although a small-molecule inhibitor of USP2 alone can partially suppress the in vivo growth of p53-wild-type mammary tumor xenografts, more strikingly, USP2 inhibition and PD-1/PD-L1 blockade in combination promote vigorous tumor regression and long-term survival of all tumor-bearing mice."
TRAF2 affects USP2
8 1 | 2
8 1 | 2

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sparser
"USP2 interacts with TRAF2 and promotes deubiquitination of TRAF2, enhancing its stability."

No evidence text available

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RECQL4 affects USP2
2 | 2 7
2 | 2 7

sparser
"To support this notion, USP2 interacted with RECQL4 in coimmunoprecipitation experiments and NCS treatment significantly increased this interaction ( xref E )."

sparser
"The expression of helicase-defective RECQL4 mutant proteins (D605A and E606A in the Walker B motif) as well as WT RECQL4 proteins restored USP2 binding to laser-induced DSB sites in RECQL4-depleted cells."

sparser
"As shown in xref D , interaction between phospho-deficient USP2 (AA) and RECQL4 was not observed in the coimmunoprecipitation analysis."

sparser
"In contrast, phosphomimetic USP2 (EE) proteins stably interacted with RECQL4, both in the absence and presence of DNA damage ( xref D ), and inhibition of ATM kinase activity did not disrupt their interaction, although the interaction appeared to be slightly weak ( xref E )."

sparser
"Taken together, these results suggest that interaction of USP2 with RECQL4 through ATM-dependent phosphorylation of these two sites is not sufficient for the recruitment of USP2 to DSB sites."

sparser
"As shown in xref , xref , xref , USP2 stably interacted with RECQL4 in an ATM-dependent manner and recruitment of USP2 to DSB sites also requires RECQL4."

reach
"As shown in Figure 5D, interaction between phospho-deficient USP2 (AA) and RECQL4 was not observed in the coimmunoprecipitation analysis."

sparser
"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."

reach
"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."

No evidence text available
LEP affects USP2
| 10
LEP increases the amount of USP2.
| 8
LEP increases the amount of USP2. 8 / 9
| 8

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"Herein, we present the evidence that changes in USP-2 expression by adiponectin and leptin contribute to the regulation of cyclin D1 degradation by proteasomes that impacts cell cycle modulation in ca[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"While it has been shown that USP2 is involved in the regulation of cancer growth and leptin induces USP2 expression, the functional role of USP2 in leptin induced p53 expression has not been explored."

reach
"Similarly, leptin also increased USP-2 protein expression in a dose- and time dependent manner in HepG2 cells and MCF-7 cells."

reach
"As shown in Fig. 4C, transfection with siRNA targeting USP-2 significantly suppressed leptin induced cyclin D1 protein expression in HepG2 cells, suggesting that increase in USP-2 expression by leptin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Leptin increases expression of p53 and USP2 in HepG2 tumor Xenograft model."

reach
"Although USP-2 seems to play a critical role in USP-2 expression modulated by adiponectin and leptin, the only little information regarding molecular mechanisms underlying USP-2 expression is availabl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In the present study, we clearly demonstrated that adiponectin and leptin modulates expression of USP-2 expression in the opposite manner, which in turn directly impacts cyclin D1 degradation (Figs. 3[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Based on our results, it is highly likely that USP-2 suppression by adiponectin enhanced proteasomal activity, thereby increasing cyclin D1 degradation, whereas enhancement in USP-2 expression by lept[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
LEP activates USP2.
| 2
LEP activates USP2. 2 / 2
| 2

reach
"We next examined whether USP-2 modulation affects proteasomal activity in HepG2 cells treated with globular adiponectin and leptin."

reach
"We therefore next inquired the role of USP-2 in cyclin D1 expression in cancer cells treated with leptin."
FASN affects USP2
6 | 2 3
6 | 2 3

reach
"For example, USP-2 interacts and stabilizes fatty acid synthase which is overexpressed in prostate cancer cells (Graner et al., 2004) or causes murine double minute 2 (MDM2) accumulation and enhance M[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Among these, USP2 interacts with FASN and promotes its protein stability by reducing the ubiquitination level of FASN in prostate cancer [ xref ]."

No evidence text available

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reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

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sparser
"USP2 interacts with FASN in the mouse liver, human hepatic cancer, and HepG2 cells, and subsequently prevents the ubiquitination-dependent digestion of FASN [ xref , xref ]."
CTNNB1 affects USP2
6 | 3 2
6 | 3 2

No evidence text available

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sparser
"The deubiquitinating enzyme USP2 interacts with β-catenin to deubiquitinate β-catenin without degradation, which in turn activates Wnt/β-catenin signaling xref ."

No evidence text available

reach
"GST pull-down assay and Co-IP assay verified the interaction between USP2 and β-catenin (Figure 6B,C)."

reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

reach
"Furthermore, USP2 interacted with beta-catenin, thereby regulating its deubiquitination and stabilization in CFs."

No evidence text available

No evidence text available
| 5

reach
"In addition, results from Masson trichromatic staining showed that collagens (major components of ECM in the gut) (Rieder et al., 2013) accumulated more intensively in the colon tissues from Usp2 mice than Lyz2-Cre;Usp2 mice after colitis induction (Fig. 5D), indicating that knockout of USP2 impairs the ECM remodeling in the colon inflammation."

reach
"Meanwhile, Usp2-overexpressed hepatocytes displayed aggressive inflammation, as evidenced by increased production of inflammatory cytokines IL-1β, IL-6, and TNF-α (Fig. 5F)."

reach
"Collectively, these data suggest that USP2 in myeloid cells promotes colon inflammation after DSS treatment.2.3 Knockout of USP2 alters gene expression profile in myeloid cells after DSS treatment."

reach
"USP2 may also mediate the response of the clock to inflammation, as the expression of the gene is increased in response to TNFalpha treatment, and CRY1 protein induction in response to this cytokine is abrogated when Usp2 expression is knocked down."

reach
"In macrophages, USP2 promotes inflammation by activating the nuclear factor kappa B (NF-κB) pathway17, 18."
| 4

reach
"Moreover, macrophage USP2 represses adipose tissue inflammation and subsequent insulin resistance [79]."

reach
"Macrophage USP2 may therefore selectively prevent age- and/or adiposity related chronic inflammation and subsequent insulin resistance."

reach
"Substrates of USP2 remaining unidentified, its presence in adipose tissue macrophages inhibits chronic inflammation in adipose tissue [20]."

reach
"Overexpression of USP2 via tail injection of recombinant adeno-associated virus serotype 9 (AAV9) in TAC-treated male mice has been reported to attenuate pressure-overload-induced ventricular dysfunction, cardiac fibrosis, inflammation, and overall oxidative stress compared with AAV9 control-treated counterparts [31]."
USP2 affects CD274
4 | 3 3
USP2 binds CD274.
4 | 3
4 | 3

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sparser
"USP2’s interaction with PD-L1 is a testament to its significance."

sparser
"A recent study by Kuang et al ( xref ) reported that Thr288, Arg292 and Asp293 of USP2 bind to PD-L1 through the resolution of K48 link polyubiquitination at the PD-L1 lysine 270 site."

sparser
"In colorectal cancer and prostate cancer cells, USP2 can directly interact with PD-L1 and promote the K48 deubiquitination of PD-L1."
USP2 activates CD274.
| 3
USP2 activates CD274. 3 / 3
| 3

reach
"USP2 promotes tumor immune evasion via deubiquitination and stabilization of PD-L1."

reach
"USP2 directly interacts with and increases PD-L1 abundance in colorectal and prostate cancer cells."

reach
"These data demonstrate that CD8 + T cells, but not CD4 + T cells, are the primary effector cells underlying the combination treatment in these tumor models.Finally, to ascertain whether USP2 inhibition potentiates the efficacy of PD-1/PD-L1 blockade by enhancing p53 function in a tumor cell autonomous manner, we also examined the effects of combination therapy in Balb/c mice bearing p53-null EMT6 tumors."
MYOD1 affects USP2
2 | 5 3
MYOD1 increases the amount of USP2.
| 4
MYOD1 increases the amount of USP2. 4 / 4
| 4

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"Taken together, these findings demonstrate that MYOD1 transcriptionally activates USP2 expression by binding to its promoter region."

reach
"MYOD1 activates USP2 transcription by binding to its promoter region."

reach
"Hence, we first determined whether MYOD1 regulates USP2 expression in C2C12 cells and found that Myod1 overexpression increased, whereas Myod1 knockdown inhibited Usp2 protein levels in C2C12 cells (Fig. 8C)."

reach
"Additionally, MYOD1 induces the transcription of USP2, which is downregulated during muscle atrophy."
MYOD1 decreases the amount of USP2.
2 | 1
MYOD1 decreases the amount of USP2. 3 / 3
2 | 1

No evidence text available

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"Hence, we first determined whether MYOD1 regulates USP2 expression in C2C12 cells and found that Myod1 overexpression increased, whereas Myod1 knockdown inhibited Usp2 protein levels in C2C12 cells (Fig. 8C)."
MYOD1 binds USP2.
| 3
| 3

sparser
"Furthermore, TNF-α and DEX treatment inhibited the binding of MYOD1 to the promoter of USP2 in the C2C12 cells ( xref )."

sparser
"Two putative MYOD1 binding motif sequences (MBE) were predicted in the USP2 promoter region (JASPAR analysis), and ChIP-qPCR assays further demonstrated the binding of MYOD1 to the promoter of the USP2 ( xref )."

sparser
"Our study further showed that TNF-α–induced inflammation significantly reduced MYOD1 binding to the USP2 promoter ( xref )."
USP2 affects glucose
| 9
USP2 activates glucose.
| 5
USP2 activates glucose. 5 / 5
| 5

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"Consistently, we found that mice transduced with siUSP2 adenovirus have impaired ability to convert pyruvate into glucose during PTT (XREF_FIG), suggesting that hepatic gluconeogenesis and glucose output are impaired by RNAi knockdown of USP2 in the liver."

reach
"Several USPs, including USP2, USP10, USP14, USP15, USP18, and USP22, have been associated with obesity and related metabolic disorders.In diet-induced obese mice, it is reported that USP2 can increase the production of hepatic glucose and promote glucose intolerance (143)."

reach
"Nfil3 is a known negative regulator of gluconeogenesis (Kang et al. 2017), while Usp2 stimulates glucose production (Molusky et al. 2012)."

reach
"Correspondingly, infection with an adenovirus encoding Usp2 shRNA significantly lowered blood glucose, blood insulin, hepatic glycogen content, and insulin sensitivity in high fat diet–fed mice, suggesting that USP2 plays a role in aggravating T2DM pathogenesis [102]."

reach
"USP2 modulates glucose and lipid metabolism, insulin sensitivity, and inflammatory processes."
USP2 inhibits glucose.
| 2
USP2 inhibits glucose. 2 / 2
| 2

reach
"USP2 deficiency in the liver also perturbs normal diurnal glucose rhythms under restricted feeding conditions."

reach
"For example, USP2 in ventromedial hypothalamic neurons attenuates aberrant increases in blood glucose by mitigating glycogenolysis in the liver, whereas hepatic USP2 regulates gluconeogenesis and diurnal glucose metabolism in the liver [28,31]."
USP2 decreases the amount of glucose.
| 2
USP2 decreases the amount of glucose. 2 / 2
| 2

reach
"Administration of a USP2-selective inhibitor, ML364 (20 ng/head), into the VMH elicited a rapid increase in the circulating glucose level in male mice, suggesting USP2 has a suppressive role on glucose mobilization."

reach
"These results indicate hypothalamic USP2 attenuates perturbations in blood glucose levels by modifying the ROS-AMPK-sympathetic nerve axis."
USP2 affects SMAD7
4 | 3 2
USP2 binds SMAD7.
4 | 2
4 | 2

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sparser
"Specifically, USP2 interacted with SMAD7 and prevented SMAD7 ubiquitination."

sparser
"USP2 interacts with SMAD7 and prevents SMAD7 ubiquitination in Glioblastoma, while SMAD7 is a major inhibitor of TGFβ signaling xref ."
USP2 deubiquitinates SMAD7.
| 3
USP2 deubiquitinates SMAD7. 3 / 3
| 3

reach
"It has established that USP2 functions as a tumor suppressor in numerous malignancies through multiple mechanisms; for instance, USP2 deubiquitinates SMAD7 and suppresses TGF-β signaling pathway to inhibit glioblastoma progression [19]."

reach
"USP2 leads to de-ubiquitination and stabilization of SMAD7, and USP2 loss results in an increased SMAD7 ubiquitination and increased phospho-SMAD2 [61]."

reach
"Specifically, USP2 interacted with SMAD7 and prevented SMAD7 ubiquitination."
USP2 affects LEF1
| 7 2
USP2 binds LEF1.
| 2 2
| 2 2

reach
"LEF1 protein stability analysis and co-immunoprecipitation (Co-IP) assay were performed to test the USP2/LEF1 interaction."

reach
"Through Co-IP assays using an anti-USP2 or anti-LEF1 antibody, we demonstrated the interaction between USP2 and LEF1 in hBMSCs, as indicated by LEF1 enrichment in USP2-associating precipitates and USP2 enrichment in LEF1-related precipitates (Fig. 4D)."

sparser
"Through Co-IP assays using an anti-USP2 or anti-LEF1 antibody, we demonstrated the interaction between USP2 and LEF1 in hBMSCs, as indicated by LEF1 enrichment in USP2-associating precipitates and USP2 enrichment in LEF1-related precipitates (Fig.  xref D)."

sparser
"These findings reveal a novel mechanism by which USP2 and LEF1 interact to regulate hBMSC osteogenic differentiation."
USP2 activates LEF1.
| 4
USP2 activates LEF1. 4 / 4
| 4

reach
"Next, we identified whether USP2 can modulate LEF1 in a deubiquitination way."

reach
"Mechanistically, our findings indicate that USP2 promotes the stability of LEF1 through deubiquitination, thereby preventing its degradation."

reach
"USP2-induced upregulation of LEF1 through deubiquitination relieves osteoporosis development by promoting the osteogenic differentiation of bone marrow mesenchymal stem cells."

reach
"USP2 promotes LEF1 protein stability by deubiquitination."
USP2 increases the amount of LEF1.
| 1
USP2 increases the amount of LEF1. 1 / 1
| 1

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"To address this question, we elevated LEF1 expression using a LEF1 ORF construct (oe-LEF1) in USP2-silenced hBMSCs because USP2 depletion led to a reduction in LEF1 expression (Fig. 5A)."
USP2 affects CLOCK
| 9
USP2 activates CLOCK.
| 7
USP2 activates CLOCK. 7 / 7
| 7

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"Thus, the USP2 deubiquitinating enzyme modulates the two major features of clock function - its intrinsic circadian rhythm and its capacity to respond to external cues."

reach
"The ability of USP2 to modulate these central functions of the clock suggested that it acts on the molecular clock machinery."

reach
"By its action on the ubiquitination status of PER1, and possibly other clock proteins it interacts with, USP2 appears to exert a broad function, modulating both SCN central clock function (as seen by effects on both period length and response to photic stimuli) as well as peripheral clocks (as exemplified by our observations in embryonic fibroblasts)."

reach
"USP2 may also mediate the response of the clock to inflammation, as the expression of the gene is increased in response to TNFalpha treatment, and CRY1 protein induction in response to this cytokine is abrogated when Usp2 expression is knocked down."

reach
"The Deubiquitinase USP2 Modulates Photic Entrainment of the Circadian Clock at the Level of the Suprachiasmatic Nucleus."

reach
"Our group showed that USP2 modulates the response of the clock to light [XREF_BIBR]."

reach
"Apart from its functions within the clock, USP2 also mediates clock output."
USP2 deubiquitinates CLOCK.
| 2
USP2 deubiquitinates CLOCK. 2 / 2
| 2

reach
"For instance, in ST pol mice, there was a marked induction of the expression of Usp2 (encoding the Ubiquitin specific peptidase 2, which deubiquitinates core clock components) [30] and a strong down-r[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Further, USP2 is able to deubiquitinate the ubiquitinated forms of the clock proteins BMAL1 (Scoma et al. 2011), PER1 (Yang et al. 2012) and CRY1 (Tong et al. 2012)."
MDM4 affects USP2
5 | 1 2
5 | 1 1

No evidence text available

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reach
"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."

No evidence text available

No evidence text available

No evidence text available

sparser
"Ubiquitin-specific protease 2a (USP2a) is a deubiquitinating enzyme that protects MDM4 against degradation, so USP2-MDM4 interaction may be a key determinant of the malignant potential of human cancers."
EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
| 1

sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
BIRC5 affects USP2
1 | 3 5
1 | 3 5

reach
"In addition, USP2 bound to survivin, and ML364 increased ubiquitination of survivin."

No evidence text available

sparser
"Mechanistically, USP2 interacted with survivin, and ML364 degraded survivin protein expression by increasing the ubiquitination of survivin."

sparser
"To investigate the interaction between USP2 and survivin, we performed immunoprecipitation (IP) assay using a USP2 antibody."

sparser
"USP2 bound to survivin ( xref A)."

sparser
"In addition, USP2 interacted with survivin and inhibition of USP2 by ML364-induced ubiquitination of survivin ( xref A,B)."

sparser
"In addition, USP2 bound to survivin, and ML364 increased ubiquitination of survivin."

reach
"USP2 Interacts with and Deubiquitinates Survivin."

reach
"To investigate the interaction between USP2 and survivin, we performed immunoprecipitation (IP) assay using a USP2 antibody."
USP2 affects TGFB
| 8
USP2 inhibits TGFB.
| 6
USP2 inhibits TGFB. 6 / 6
| 6

reach
"USP2 depletion induced TGF-β signaling and progression of GBM."

reach
"Conversely, USP2 inhibits glioblastoma development by suppressing the TGF-B signaling pathway through the deubiquitination of SMAD7 [19]."

reach
"In contrast, we found that USP2 inhibits the TGF-β signaling and the proliferation of myeloid cells."

reach
"Whether USP2 down-regulates TGF-β pathway dependently on SMAD7 in myeloid cells in such a context as DSS treatment requires further investigation."

reach
"Collectively, these data suggest that USP2 negatively regulates TGF-β signaling to suppress myeloid cells proliferation in the lamina propria during DSS-induced colitis, although it promotes TGF-β signaling and exhibits cytostatic effects in multiple non-immune cells.One major function of myeloid cells is to activate T cells during infection and inflammation."

reach
"More recently, it was published that loss in USP2 expression due to DNMR3A-mediated aberrant DNA methylation enhances TGF-β signaling and GBM progression."
USP2 activates TGFB.
| 2
USP2 activates TGFB. 2 / 2
| 2

reach
"We have previously reported that USP2 positively regulates TGF-β signaling in tumor parenchymal cells by promoting deubiquitylation of TGFBR1 and participates in tumor genesis and metastasis (Zhao et al., 2018)."

reach
"To examine whether USP2 directly regulates TGF-β signaling in vitro, we isolated RFP  LPMCs from Lyz2-Cre;Usp2 Ai14 and Lyz2-Cre;Ai14 mice and stimulated them with TGF-β for 30 min or 4 h followed by flow cytometry with p-SMAD2 and Ki67."
| 1 7
| 1 5

reach
"Likewise, ubiquitin-specific peptidase 2 (USP2) downregulates the NF-κB pathway, inhibiting EMT in clear cell renal cell carcinoma metastasis (Duan et al., 2022)."

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

reach
"USP2 overexpression promoted proliferation and metastasis, and induced the EMT in CM cells in vitro, while specific inhibition of USP2 by ML364 produced the opposite effects."

eidos
"USP2 promotes cell proliferation and metastasis in choroidal melanoma via stabilizing Snail ."

reach
"Thus, USP2 appears to be an attractive anticancer target, and indeed, small-molecule inhibitors of USP2 have shown to inhibit growth and metastasis of cancer cells (22, 40)."

reach
"USP2 promotes cell proliferation and metastasis in choroidal melanoma via stabilizing Snail."
| 2

reach
"Overexpression of ubiquitin-specific peptidase 2 suppressed metastasis of clear cell renal cell carcinoma cells in vivo."

reach
"Ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition in clear cell renal cell carcinoma metastasis by downregulating the NF-kappaB pathway."
USP2 affects Mice
| 8
USP2 activates Mice. 8 / 8
| 8

reach
"Moreover, increased USP2 expression reduced bone loss and enhanced osteogenic differentiation in OVX mice."

reach
"Taken together, these findings suggest the loss of USP2 promotes muscle loss and weakness in diabetic or DEX-induced mice."

reach
"Meanwhile, USP2 ablation-induced PD-L1 clearance enhances antitumor immunity in mice via increasing CD8 + T cells infiltration and reducing immunosuppressive infiltration of myeloid-derived suppressor cells (MDSCs) and regulatory T cells (Tregs), whereas PD-L1 overexpression reverses the tumor growth suppression by USP2 silencing."

reach
"In a transcriptome analysis of renal cortex comparing diabetic mice with diabetic post-menopausal mice, Usp2 expression was slightly enhanced in the menopause model [130]."

reach
"Correspondingly, infection with an adenovirus encoding Usp2 shRNA significantly lowered blood glucose, blood insulin, hepatic glycogen content, and insulin sensitivity in high fat diet–fed mice, suggesting that USP2 plays a role in aggravating T2DM pathogenesis [102]."

reach
"Indeed, USP2 inhibition alone partially induces tumor growth suppression but more strikingly, the combination of USP2 inhibition and PD-1/PD-L1 blockade promote vigorous tumor regression and long-term survival of all tumor-bearing mice."

reach
"Moreover, although a small-molecule inhibitor of USP2 alone can partially suppress the in vivo growth of p53-wild-type mammary tumor xenografts, more strikingly, USP2 inhibition and PD-1/PD-L1 blockade in combination promote vigorous tumor regression and long-term survival of all tumor-bearing mice."

reach
"Upregulation of USP2 reduces bone loss in OVX mice."
USP2 affects EGFR
1 | 1 1
USP2 inhibits EGFR.
| 1
USP2 inhibits EGFR. 1 / 5
| 1

reach
"USP2 has also been shown to localise to endosomes and negatively regulate EGFR down-regulation 52."
USP2 binds EGFR.
1 | 1
1 | 1

sparser
"For example, USP2 interacts with early EGFR transporters and performs deubiquitination modifications, which prevent EGFR from being transported to lysosomes for degradation."

No evidence text available
USP2 affects EBF2
| 3 5
EBF2 binds USP2. 5 / 5
| 5

sparser
"Moreover, immunofluorescence staining confirmed the interaction of USP2 and EBF2 in primary brown adipocytes ( xref K)."

sparser
"Inhibition of the USP2EBF2 axis significantly reduced the half-life of EBF2 protein and suppressed adipocyte thermogenesis."

sparser
"Mechanistically, we demonstrated that USP2 directly interacts with EBF2 and reduces K63-linked polyubiquitination of EBF2 at residue K321, subsequently enhancing the EBF2-mediated adaptive thermogenesis."

sparser
"Overall, we revealed a USP2-EBF2 axis in BAT thermogenesis, expanding our understanding of how ubiquitination dynamics influence metabolic adaptations, and identified USP2 as a potential therapeutic target for metabolic diseases."

sparser
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."
| 3

reach
"Moreover, immunofluorescence staining confirmed the interaction of USP2 and EBF2 in primary brown adipocytes (Figure 6K)."

reach
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."

reach
"The association between Flag-USP2 and HA-EBF2 was detected in these cells (Figure 6I,J)."
USP2 affects CRY1
3 1 | 3 1
USP2 deubiquitinates CRY1.
1 | 3
USP2 deubiquitinates CRY1. 4 / 4
1 | 3

reach
"Further, USP2 is able to deubiquitinate the ubiquitinated forms of the clock proteins BMAL1 (Scoma et al. 2011), PER1 (Yang et al. 2012) and CRY1 (Tong et al. 2012)."

reach
"In addition to PER1 and BMAL1, USP2 deubiquitinates CRY1 in cultured cells in response to a serum shock, and in the mouse liver, Usp2 knockdown increases CRY ubiquitination and decreases CRY1 protein levels."

reach
"USP2 is essential to deubiquitinating PER1, CRY1 and CRY2 in vivo [54–57]."

"<span class="match term0">USP2a</span> protein deubiquitinates and stabilizes the circadian protein <span class="match term1">CRY1</span> in response to inflammatory signals"
USP2 binds CRY1.
3 | 1
3 | 1

sparser
"Targeting the CRY1-USP2 feedback loop ( xref ) offers a promising avenue for therapeutic intervention in Group 3 MB."

No evidence text available

No evidence text available

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TGFBR2 affects USP2
2 4 2 |
TGFBR2 phosphorylates USP2.
2 2 |
TGFBR2 phosphorylates USP2 on S225. 2 / 2
1 1 |

No evidence text available

"Here, we report the role of USP2a in promoting metastasis by facilitating TGF-β-triggered signaling. USP2a interacts with TGFBR1 and TGFBR2 upon TGF-β stimulation and removes K33-linked polyubiquitin chains from Lys502 of TGFBR1, promoting the recruitment of SMAD2/3. Simultaneously, TGFBR2 phosphorylates Ser207/Ser225 of USP2a, leading to the disassociation of SMAD2/3 from TGFBR1."
TGFBR2 phosphorylates USP2 on S207. 2 / 2
1 1 |

No evidence text available

"Here, we report the role of USP2a in promoting metastasis by facilitating TGF-β-triggered signaling. USP2a interacts with TGFBR1 and TGFBR2 upon TGF-β stimulation and removes K33-linked polyubiquitin chains from Lys502 of TGFBR1, promoting the recruitment of SMAD2/3. Simultaneously, TGFBR2 phosphorylates Ser207/Ser225 of USP2a, leading to the disassociation of SMAD2/3 from TGFBR1."
TGFBR2 binds USP2.
2 |
2 |

No evidence text available

No evidence text available
TGFBR2 activates USP2.
2 |
TGFBR2 activates USP2. 2 / 2
2 |

"Here, we report the role of USP2a in promoting metastasis by facilitating TGF-β-triggered signaling. USP2a interacts with TGFBR1 and TGFBR2 upon TGF-β stimulation and removes K33-linked polyubiquitin chains from Lys502 of TGFBR1, promoting the recruitment of SMAD2/3. Simultaneously, TGFBR2 phosphorylates Ser207/Ser225 of USP2a, leading to the disassociation of SMAD2/3 from TGFBR1."

"Here, we report the role of USP2a in promoting metastasis by facilitating TGF-β-triggered signaling. USP2a interacts with TGFBR1 and TGFBR2 upon TGF-β stimulation and removes K33-linked polyubiquitin chains from Lys502 of TGFBR1, promoting the recruitment of SMAD2/3. Simultaneously, TGFBR2 phosphorylates Ser207/Ser225 of USP2a, leading to the disassociation of SMAD2/3 from TGFBR1."
PTH affects USP2
| 1 7
PTH increases the amount of USP2.
| 4
PTH increases the amount of USP2. 4 / 4
| 4

reach
"Parathyroid hormone (hPTH 1-38) stimulates the expression of UBP41, an ubiquitin specific protease, in bone."

reach
"PTH (1-31), which stimulates intracellular cAMP, and PTHrP (1-34) both induced UBP41 mRNA expression; whereas PTH analogs (3-34) and (7-34), that do not stimulate cAMP, had no effect on UBP41 expression."

reach
"Parathyroid hormone (hPTH 1-38) stimulates the expression of UBP41, an ubiquitin specific protease, in bone."

reach
"A further study revealed that PTH (1-34) could upregulate the expression of USP2 and promote PTHR deubiquitination as well as stabilization [108]."
PTH activates USP2.
| 1 3
PTH activates USP2. 4 / 4
| 1 3

reach
"USP2 was found to be stimulated by PTH in the bone."

reach
"Together, these results demonstrate that UBP41, an ubiquitin specific protease, is selectively upregulated in bone by the osteotropic agents PTH, PTHrP, and PGE2, possibly via the PKA and cAMP pathway."

reach
"In contrast, PTH (7-34) does not increase USP2."

eidos
"A further study revealed that PTH ( 1-34 ) could upregulate the expression of USP2 and promote PTHR deubiquitination as well as stabilization [ 108 ] ."
EBF2 affects USP2
| 3 5
EBF2 binds USP2. 5 / 5
| 5

sparser
"Moreover, immunofluorescence staining confirmed the interaction of USP2 and EBF2 in primary brown adipocytes ( xref K)."

sparser
"Inhibition of the USP2EBF2 axis significantly reduced the half-life of EBF2 protein and suppressed adipocyte thermogenesis."

sparser
"Mechanistically, we demonstrated that USP2 directly interacts with EBF2 and reduces K63-linked polyubiquitination of EBF2 at residue K321, subsequently enhancing the EBF2-mediated adaptive thermogenesis."

sparser
"Overall, we revealed a USP2-EBF2 axis in BAT thermogenesis, expanding our understanding of how ubiquitination dynamics influence metabolic adaptations, and identified USP2 as a potential therapeutic target for metabolic diseases."

sparser
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."
| 3

reach
"Moreover, immunofluorescence staining confirmed the interaction of USP2 and EBF2 in primary brown adipocytes (Figure 6K)."

reach
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."

reach
"The association between Flag-USP2 and HA-EBF2 was detected in these cells (Figure 6I,J)."
2,3,7,8-tetrachlorodibenzodioxine increases the amount of USP2.
4 |
4 |

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2,3,7,8-tetrachlorodibenzodioxine decreases the amount of USP2.
4 |
4 |

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Disulfiram affects USP2
| 5 2
| 5 2

reach
"Notably, addition of a detergent, Triton X-100, showed no effect to the inhibitory function of disulfiram (Fig. S1), indicating the specificity of disulfiram-mediated inhibition.Next, to characterize [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The results showed that pre-treatment of disulfiram diminished nearly 96% and 84% DUB activities of USP2 and USP21, respectively ( Fig. 4 )."

sparser
"Next, to characterize the disulfiram-induced inhibitions of USP2 and USP21, kinetic measurements were carried out with a range of concentrations of inhibitor, and raw data were fitted to available inh[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Consistent with the previous study in papain-like proteases, we found that the disulfiram-induced inhibitions of USP2 and USP21 were best fitted to the competitive inhibition model ( Fig. 2 )."

reach
"Furthermore, disulfiram and 6 TG synergistically inhibited the enzymatic activity of USP2 (Lin et al., 2021), but whether these two compounds suppress the activity of other DUBs remains to be studied further."

reach
"Recently, disulfiram and 6-thioguanine were reported to synergistically inhibit USP2 and USP21 but with low affinity in the micromolar range."

reach
"We also showed that disulfiram and 6TG can synergistically inhibit the activities of USP2 and USP21, suggesting a repurposing application of disulfiram and 6TG in treating USP-related tumors.The expre[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects NBN
2 | 2 3
USP2 binds NBN.
2 | 3
2 | 3

sparser
"However, USP2 can interact with NBS1 protein when both proteins are overexpressed ( xref C ), and direct interaction between USP2 and SKP2 was also observed in a previous study ( xref )."

No evidence text available

No evidence text available

sparser
"On the other hand, USP2 stably interacted with NBS1 upon NCS treatment, and their interaction persisted regardless of NCS treatment ( xref C )."

sparser
"As shown in this study, USP2 stably interacts with NBS1 ( xref ) and affects ubiquitination of NBS1 ( xref and xref )."
USP2 deubiquitinates NBN.
| 2
USP2 leads to the deubiquitination of NBN. 2 / 2
| 2

reach
"Taken together, these results suggest that USP2 prevents premature disassembly of the MRN complex in RECQL4-defective cells by counteracting the ubiquitination of NBS1."

reach
"As SKP2 E3 ligase is responsible for NBS1 ubiquitination during the DSB response, USP2 may also be able to reduce NBS1 ubiquitination through interaction with SKP2, although the importance of this mode of action in stabilizing the MRN complex during the DSB response has not yet been determined.The MRN complex is one of the key factors in the DSB response, and its recruitment to DSB sites and its roles in the DSB response have been well documented in many previous studies."
USP2 affects MYLIP
1 | 5
USP2 deubiquitinates MYLIP.
| 3
USP2 deubiquitinates MYLIP. 3 / 3
| 3

reach
"In summary, USP2 activity leads to the deubiquitylation and stabilization of cell surface LDLR in and IDOL dependent manner, but it remains uncertain exactly why such a phenomenon is accompanied by an extended IDOL protein half-life."

reach
"Since overexpression of Usp2 promotes the deubiquitination of IDOL, USP2 was considered to suppress LDL uptake via ubiquitin-dependent degradation of LDL [212]."

reach
"The authors attempt to answer this question by introducing a tripartite complex model, in which USP2 interacts with LDLR at the plasma membrane in an IDOL dependent manner to deubiquitylate and stabilize both LDLR and IDOL."
USP2 binds MYLIP.
1 |
1 |

No evidence text available
USP2 activates MYLIP.
| 2
USP2 activates MYLIP. 2 / 2
| 2

reach
"Such a temporal expression pattern would leave little opportunity to alleviate the USP2 mediated antagonism of IDOL, and the tripartite complex model would imply that IDOL is constantly engaged in a futile reaction, in which its ubiquitylation of LDLR is immediately reversed by USP2."

reach
"As expected of an interacting DUB, USP2 decreased the ubiquitylation and increased the protein stability of IDOL."
USP2 affects KRAS
| 5 2
USP2 binds KRAS.
| 2 2
| 2 2

reach
"Taken together, these results suggest that USP2 may enhance the protein stability of KRAS.Based on the above results, we hypothesized that USP2 may interact with and deubiquitinate KRAS."

reach
"Endogenous interaction of KRAS and USP2 was further verified in MM.1S cells (Fig. 4J)."

sparser
"In the present study, we demonstrated that USP2 is a DUB that directly controls KRAS ubiquitination, as supported by the following evidence: 1) overexpression of USP2 stabilizes KRAS; 2) USP2 interacts with KRAS in cells as indicated by the immunoprecipitation and immunofluorescence assays; 3) USP2 directly removes ubiquitin from ubiquitylated KRAS, which leads to KRAS stabilization, whereas USP2 inhibition results in the proteasomal degradation of KRAS."

sparser
"Endogenous interaction of KRAS and USP2 was further verified in MM."
USP2 deubiquitinates KRAS.
| 2
USP2 deubiquitinates KRAS. 2 / 2
| 2

reach
"Taken together, these results suggest that USP2 may enhance the protein stability of KRAS.Based on the above results, we hypothesized that USP2 may interact with and deubiquitinate KRAS."

reach
"To determine whether USP2 deubiquitinates KRAS, USP2 , USP2 and its active-site mutant, USP2 , were separately transfected into HEK293T cells, which were subsequently lysed and subjected to the immunoprecipitation assay."
USP2 activates KRAS.
| 1
USP2 activates KRAS. 1 / 1
| 1

reach
"These data collectively suggest that the USP2 plays a critical role in the proliferation and survival of MM cells in vitro and in vivo.3.8 GA induces the apoptosis of MM cells by targeting USP2 and destabilizing KRAS."
USP2 affects KCNQ1
2 | 3 1
USP2 binds KCNQ1.
2 | 1 1
2 | 1 1

No evidence text available

No evidence text available

reach
"A reduced level of ubiquitylation leads to an increased surface density of KCNQ1 channels and thereby explains the rescued current level observed in the functional experiments.Whether or not Nedd4-2 i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Whether or not Nedd4-2 is coexpressed does not seem to influence the binding of USP2 to KCNQ1, as both USP2-45 and USP-69 bind equally well under the 2 conditions ( Figure 3 )."
USP2 activates KCNQ1.
| 1
USP2 activates KCNQ1. 1 / 2
| 1

reach
"In contrast to previous observations with the ENaC, coexpression of the 2 isoforms of USP2 in the present study did not increase the KCNQ1 and KCNE1 mediated current in Xenopus oocytes per se."
USP2 increases the amount of KCNQ1.
| 1
USP2 increases the amount of KCNQ1. 1 / 1
| 1

reach
"The catalytically inactive USP2 (CA) did not restore the KCNQ1 level."
USP2 affects Interferon
| 3
USP2 activates Interferon.
| 1
| 1

reach
"We further found that USP2 inhibits the proliferation of myeloid cells to suppress the activation and expansion of CD4 T cells and CD8 T cells and the production of IL-22 and IFNγ."
USP2 inhibits Interferon.
| 2
| 2

reach
"These data together suggest that inhibition of USP2 promotes the proliferation of macrophages and dendritic cells to activate IL-22- and IFNγ-producing T cells during DSS-induced colitis.We also examined whether targeting USP2 by ML364 inhibited the dysregulation of ECM and promoted tissue repair."

reach
"Taken together, these data suggest that USP2 restricts the proliferation of myeloid cells and thereby inhibits the activation and expansion of IL-22- and IFNγ-producing T cells in the lamina propria during DSS-induced colitis.2.6 USP2 in myeloid cells inhibits immune responses to bacterial infections in the gut."

reach
"Ubiquitin-specific peptidase 2-mediated inhibition of epithelial-mesenchymal transition in clear cell renal cell carcinoma cells is dependent on the nuclear factor-kappaB pathway."

reach
"Ubiquitin-specific peptidase 2 downregulation in clear cell renal cell carcinoma is associated with worse clinical stage and prognosis."

reach
"Ubiquitin-specific peptidase 2 inhibited the proliferation, migration, and invasion of clear cell renal cell carcinoma cells."

reach
"These data showed that overexpression of ubiquitin-specific peptidase 2 downregulates the metastatic ability of clear cell renal cell carcinoma cells."

reach
"Overexpression of ubiquitin-specific peptidase 2 suppressed metastasis of clear cell renal cell carcinoma cells in vivo."

reach
"Hence, ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition of clear cell renal cell carcinoma by downregulating the NF-κB pathway."

reach
"Ubiquitin-specific peptidase 2 inhibition promoted clear cell renal cell carcinoma cell proliferation, migration, and invasion."
7 |
Particulate Matter increases the amount of USP2.
5 |
Particulate Matter increases the amount of USP2. 5 / 5
5 |

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Particulate Matter decreases the amount of USP2.
2 |
Particulate Matter decreases the amount of USP2. 2 / 2
2 |

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KRAS affects USP2
| 5 2
KRAS binds USP2.
| 2 2
| 2 2

reach
"Taken together, these results suggest that USP2 may enhance the protein stability of KRAS.Based on the above results, we hypothesized that USP2 may interact with and deubiquitinate KRAS."

reach
"Endogenous interaction of KRAS and USP2 was further verified in MM.1S cells (Fig. 4J)."

sparser
"In the present study, we demonstrated that USP2 is a DUB that directly controls KRAS ubiquitination, as supported by the following evidence: 1) overexpression of USP2 stabilizes KRAS; 2) USP2 interacts with KRAS in cells as indicated by the immunoprecipitation and immunofluorescence assays; 3) USP2 directly removes ubiquitin from ubiquitylated KRAS, which leads to KRAS stabilization, whereas USP2 inhibition results in the proteasomal degradation of KRAS."

sparser
"Endogenous interaction of KRAS and USP2 was further verified in MM."
KRAS activates USP2.
| 3
KRAS activates USP2. 3 / 3
| 3

reach
"Overexpression of KRAS (Fig. 6D) could partially abrogate the USP2 knockout-induced cell growth inhibition (Fig. 6E and F)."

reach
"These data suggest that GA can induce the proteasomal degradation of KRAS, which in turn impairs the downstream signal transduction in MM cells.3.2 Identification of USP2 as a novel target of GA."

reach
"Building on the observations that GA decreases KRAS protein levels in MM cells and inhibits USP2 activity, we hypothesized that USP2 may regulate the stability of KRAS."
CD274 affects USP2
4 | 3
4 | 3

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sparser
"USP2’s interaction with PD-L1 is a testament to its significance."

sparser
"A recent study by Kuang et al ( xref ) reported that Thr288, Arg292 and Asp293 of USP2 bind to PD-L1 through the resolution of K48 link polyubiquitination at the PD-L1 lysine 270 site."

sparser
"In colorectal cancer and prostate cancer cells, USP2 can directly interact with PD-L1 and promote the K48 deubiquitination of PD-L1."
7 |
Air Pollutants increases the amount of USP2.
5 |
Air Pollutants increases the amount of USP2. 5 / 5
5 |

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Air Pollutants decreases the amount of USP2.
2 |
Air Pollutants decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
Catenin affects USP2
| 6
Catenin binds USP2. 6 / 6
| 6

sparser
"However, inhibiting the Wnt /β‐catenin canonical pathway and down‐regulating β‐catenin can achieve the aim of improving the ventricular remodelling. xref Actually, one of the major mechanisms of β‐catenin regulation is ubiquitination, beside which DUBs, such as USP47, USP20, and USP2, have also been shown to deubiquitinate and stabilize β‐catenin directly. xref , xref , xref In this study, β‐catenin was down‐regulated by USP2 inhibition, which prompted us to assess the interaction between USP2 and β‐catenin."

sparser
"As expected, USP2 interacted with β‐catenin, and regulated its ubiquitination and degradation in CFs."

sparser
"Furthermore, USP2 interacted with β‐catenin, thereby regulating its deubiquitination and stabilization in CFs."

sparser
"To further detect the interaction between β‐catenin and USP2, Co‐IP was performed."

sparser
"To investigate the potential interaction between β‐catenin and USP2, co‐immunoprecipitation (Co‐IP) was performed."

sparser
"As expected, we found that USP2 notably interacted with β‐catenin in CFs (Figure xref )."
Bisphenol A affects USP2
6 |
Bisphenol A decreases the amount of USP2.
4 |
Bisphenol A decreases the amount of USP2. 4 / 4
4 |

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Bisphenol A increases the amount of USP2.
2 |
Bisphenol A increases the amount of USP2. 2 / 2
2 |

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6 |
Benzo[a]pyrene increases the amount of USP2. 6 / 6
6 |

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VCP affects USP2
| 2 4
VCP binds USP2.
| 4
| 4

sparser
"Direct physical interaction of VCP with USP2 was observed in 293 T cells, demonstrating that domain 2 of VCP and the amino acid sequence (200-403) of USP2 were critical for their interactions (Fig. xref )."

sparser
"To further clarify the interaction between VCP and USP2, we introduced an inhibitor targeting the D2 domain of VCP."

sparser
"To confirm whether CB-5083 disrupted the binding of USP2 to the D2 domain of VCP, we conducted CO-IP experiments."

sparser
"The results exhibited that CB-5083 disrupted the VCP-USP2 interaction (Supplementary Fig. xref )."
VCP activates USP2.
| 2
VCP activates USP2. 2 / 2
| 2

reach
"VCP enhances autophagy-related osteosarcoma progression by recruiting USP2 to inhibit ubiquitination and degradation of FASN."

reach
"Valosin-containing protein (VCP) enhanced the FASN stability by recruiting ubiquitin specific peptidase-2 (USP2) to remove the K48-linked ubiquitin chains from FASN; domain 2 of VCP and the amino acid sequence () of USP2 were critical for their interactions."
USP2 affects ferroptosis
| 6
USP2 activates ferroptosis.
| 4
| 4

reach
"Moreover, USP2 overexpression could promote ferroptosis through p53 deubiquitination."

reach
"The evidence suggested that USP2 overexpression promoted ferroptosis in NSCLC cells.Moreover, p53 is also involved in regulating ferroptosis [10–12]."

reach
"Further research revealed that USP2 overexpression suppressed cell viability in DDP-resistant NSCLC and promoted ferroptosis."

reach
"USP2 promoted ferroptosis through the nuclear transfer of p53."
USP2 inhibits ferroptosis.
| 2
| 2

reach
"Pharmacological USP2 targeting suppresses ovarian cancer growth by potentiating apoptosis and ferroptosis."

reach
"Moreover, USP2 silencing enhanced the iron ion concentration and lipid peroxidation accumulation as well as suppressed ferroptosis, while upregulating USP2 promoted ferroptotic cell death in ESCC cells."
USP2 affects catenin
| 6
Catenin binds USP2. 6 / 6
| 6

sparser
"However, inhibiting the Wnt /β‐catenin canonical pathway and down‐regulating β‐catenin can achieve the aim of improving the ventricular remodelling. xref Actually, one of the major mechanisms of β‐catenin regulation is ubiquitination, beside which DUBs, such as USP47, USP20, and USP2, have also been shown to deubiquitinate and stabilize β‐catenin directly. xref , xref , xref In this study, β‐catenin was down‐regulated by USP2 inhibition, which prompted us to assess the interaction between USP2 and β‐catenin."

sparser
"As expected, USP2 interacted with β‐catenin, and regulated its ubiquitination and degradation in CFs."

sparser
"Furthermore, USP2 interacted with β‐catenin, thereby regulating its deubiquitination and stabilization in CFs."

sparser
"To further detect the interaction between β‐catenin and USP2, Co‐IP was performed."

sparser
"To investigate the potential interaction between β‐catenin and USP2, co‐immunoprecipitation (Co‐IP) was performed."

sparser
"As expected, we found that USP2 notably interacted with β‐catenin in CFs (Figure xref )."
USP2 affects TRAF6
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

reach
"Since USP2 stabilizes PGC1α by deubiquitination, USP2 might eliminate mitochondrial ROS by activating the PGC1α–UCP2 axis."

reach
"In agreement with this, our preliminary experiments showed that the inhibition of USP2 also potentiated mitochondrial ROS production in myotubes."

reach
"Thus, USP2 seems to prevent ROS production in mitochondria in the muscle cell lineage."

reach
"Mechanistically, an ML364 treatment experiment suggests that USP2 inhibits the accumulation of ROS in VMH neurons, and thereby leads to continued ATP production in the mitochondria."
| 2

reach
"Among the ferroptosis markers, the results showed that USP2 overexpression promoted LDH release, Fe 2+ level, MDA and Lipid ROS, while inhibited GPX4 activity and GSH levels."

reach
"With regards to myoblasts, the chemical and genetic ablation of USP2 rapidly induces ROS accumulation, with accompanying defects in mitochondrial ATP synthesis [33]."
USP2 affects RIPK1
1 4 | 1
USP2 binds RIPK1.
4 | 1
4 | 1

No evidence text available

No evidence text available

No evidence text available

sparser
"For example, when the N-terminus of USP2 binds to receptor interacting protein-1 (RIP1) protein, it removes the ubiquitinated molecules on RIP1 protein and increases RIP1 expression, which in turn promotes apoptosis ( xref )."

No evidence text available
USP2 deubiquitinates RIPK1.
1 |
USP2 deubiquitinates RIPK1 on K377. 1 / 1
1 |

No evidence text available
USP2 affects PEX5
1 | 3 1
1 | 3 1

reach
"By mammalian two-hybrid analysis, a clear binding between PEX5 and USP2 was observed, whereas no interaction was detected when the PTS1-sequence is deleted (DeltaSRM) (XREF_FIG)."

reach
"The results of the two-hybrid analysis indicate that the interaction between USP2 and PEX5 depends on the quality of the PTS1 and is significantly increased by conversion of the sequence."

reach
"Taking into account that USP2 interacts with the import receptor PEX5, deubiquitination of monoubiquitinated PEX5 could have been a peroxisome related function of the enzyme."

No evidence text available

sparser
"The results of the two-hybrid analysis indicate that the interaction between USP2 and PEX5 depends on the quality of the PTS1 and is significantly increased by conversion of the sequence."
USP2 affects NHERF4
| 3
| 3

sparser
"We observed that USP2-45 interacts with NHERF4 ( xref ) and endogenous Clathrin Heavy Chain (CLH) ( xref ), whereas the non-circadian isoform USP2-69 does not."

sparser
"Finally, USP2-45 interacts in vitro with NHERF4 and endogenous Clathrin Heavy Chain."

sparser
"Accordingly, USP2-45 interacts with NHERF4 and endogenous Clathrin Heavy Chain (CLH) in transfected HEK293 cells ( xref )."
USP2 affects MMP2
1 | 4
USP2 increases the amount of MMP2.
| 2
USP2 increases the amount of MMP2. 2 / 3
| 2

reach
"USP2 was found to promote MMP2 expression and its upregulation is associated with enhanced invasiveness of breast cancer cell lines [ 105 ]."

reach
"This study also showed that USP2 upregulates MMP2 expression and induces the motility and invasiveness of TNBC cells [XREF_BIBR]; however, it remains unknown whether MMP2 is a direct substrate of USP2."
USP2 deubiquitinates MMP2.
1 | 1
USP2 leads to the deubiquitination of MMP2. 2 / 2
1 | 1

reach
"It has been proven that USP2 promotes breast cancer metastasis by deubiquitinating MMP2 XREF_BIBR."
USP2 decreases the amount of MMP2.
| 1
USP2 decreases the amount of MMP2. 1 / 1
| 1

reach
"Silencing of USP2 expression decreased migration and invasion in LM2-4175 and SCP46 cells in association with the downregulation of matrix metalloproteinase-2 (MMP2) expression, whereas overexpression of USP2 in MDA-MB-468 and MDA-MB-231 cells enhanced migration and invasion and upregulated the expression of MMP2."
USP2 affects GST
| 6
| 6

sparser
"We immunoprecipitated GPx4 complexes and incubated with GST-USP2, a DUB that can remove different types of ubiquitin chains ( xref )."

sparser
"The result showed that the recruitment of HOIP to GPx4 was eliminated by incubation with purified GST-USP2 ( xref and)."

sparser
"Additionally, the interaction between HOIP and GPx4 was also disrupted after in vitro deubiquitination assay mediated by GST-USP2 ()."

sparser
"Conversely, the recombinant GST-CD47, but not GST alone, bound to USP2 (Fig.  xref )."

sparser
"This interaction appears to be direct since purified Flag-VPRBP binds a purified GST-USP2 fusion protein but not GST alone (Fig.  xref )."

sparser
"The glutathione S-transferase (GST) pull-down assay showed that bacterially purified recombinant GST-USP2, but not GST alone, interacted with CD47 (Fig.  xref )."
USP2 affects DLL1
| 6
| 6

sparser
"These results suggest that USP2-45 binding to α 2 δ-1 promotes the de-ubiquitylation of both Ca v 1.2 and α 2 δ-1 subunits, in order to regulate the expression of Ca v 1.2 channels at the plasma membrane."

sparser
"This result suggests that the first critical event is the binding of USP2-45 to the auxiliary α 2 δ-1 subunit which may act as an anchor allowing for USP2-45 to de-ubiquitylate Ca v 1.2 channels."

sparser
"In addition, the binding of USP2-45 to α 2 δ-1 may disrupt the chaperone role of this subunit towards Ca v 1.2, leading to the reduction of Ca v 1.2 surface expression."

sparser
"However, USP2-45 specifically interacts with α 2 δ-1 rather than with the Ca V 1.2 channel or β 2 -subunit, thus it is not appropriate to define USP2-45 as a deubiquitinase of the Ca V 1.2 channel."

sparser
"Unexpectedly, the α 2 δ-1-USP2-45 protein interaction appears to downregulate the total expression of Ca V 1.2, α 2 δ-1 and β 2 subunits, and simultaneously to inhibit plasma membrane insertion and Ca 2+ current density of Ca V 1.2 channels in transfected HEK 293 cells."

sparser
"Though the reason for this peculiarity is presently unknown, it may lie in the fact that the USP2-45 binding to Ca V α 2 δ-1 might disrupt the chaperone role of the auxiliary subunit, leading to a reduction of the Ca V 1.2α 1 pore-forming subunit trafficked to the plasma membrane [ xref ]."
USP2 affects CEBPA
| 6
USP2 deubiquitinates CEBPA.
| 3
USP2 deubiquitinates CEBPA. 3 / 3
| 3

reach
"Importantly, by deubiquitinating CCAAT/enhancer binding protein alpha (C/EBPα), USP2 also regulates 11beta-hydroxysteroid dehydrogenase type 1 (11β-HSD1), thereby playing a vital role in the homeostasis of glucocorticoids within liver cells16.In this study, we found that high dietary fructose consumption promotes MASLD development and progression in C57BL/6J mice, identified Usp2 as a specific fructose-responsive gene, and revealed that the USP2-mediated C/EBPα/11β-HSD1 signaling is involved in disrupting cortisol homeostasis."

reach
"Lin’s team reported that USP2 mediates the deubiquitylation of Cebpa and induces the expression of the 11β-hydroxysteroid dehydrogenase 1 (Hsd1) gene, thereby enhancing glucocorticoid signaling and stimulating hepatic gluconeogenesis [44]."

reach
"Importantly, by deubiquitinating CCAAT/enhancer binding protein alpha (C/EBPα), USP2 also regulates 11beta-hydroxysteroid dehydrogenase type 1 (11β-HSD1), thereby playing a vital role in the homeostasis of glucocorticoids within liver cells16.In this study, we found that high dietary fructose consumption promotes MASLD development and progression in C57BL/6J mice, identified Usp2 as a specific fructose-responsive gene, and revealed that the USP2-mediated C/EBPα/11β-HSD1 signaling is involved in disrupting cortisol homeostasis."
USP2 activates CEBPA.
| 3
USP2 activates CEBPA. 3 / 3
| 3

reach
"To validate that USP2 mediated the C/EBPα/ 11β-HSD1 pathway, we detected their expression in the liver of HFCS-drinking mice."

reach
"Collectively, our findings revealed the role of dietary fructose in MASLD pathogenesis and identified the USP2-mediated C/EBPalpha/ 11beta-HSD1 signaling as a potential target for the management of MASLD."

reach
"Our research indicates that fructose exposure leads to the upregulation of C/EBPα and 11β-HSD1, unveiling a mechanism by which USP2-mediated activation of C/EBPα and 11β-HSD1 is implicated in the onset and progression of MASLD.The liver is the central viscus for converting inactive cortisone into active cortisol via 11β-HSD132-34."
TWIST1 affects USP2
| 3 3
| 3 3

reach
"We carried out a co-immunoprecipitation experiment and showed a strong interaction between endogenous USP2 and Twist in BT549 cells."

sparser
"We carried out a co-immunoprecipitation experiment and showed a strong interaction between endogenous USP2 and Twist in BT549 cells (Fig.  xref )."

reach
"Further characterization of the interaction between USP2 and Twist uncovered that the N-terminal domain of USP2 mediates its binding to Twist."

reach
"We demonstrate that TJP1 functions as a scaffold for the interaction between USP2 and TWIST1 and this may provide potential therapeutic targets in bladder cancer."

sparser
"We demonstrate that TJP1 functions as a scaffold for the interaction between USP2 and TWIST1 and this may provide potential therapeutic targets in bladder cancer."

sparser
"Further characterization of the interaction between USP2 and Twist uncovered that the N-terminal domain of USP2 mediates its binding to Twist (Supplementary Figure  xref )."
TRAF6 affects USP2
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
SPOP affects USP2
| 6
SPOP binds USP2.
| 5
| 5

sparser
"Moreover, we demonstrated that SPOP interacts with USP2, promoting its ubiquitination and subsequent degradation, which in turn facilitates CD47 degradation ( xref xref )."

sparser
"Notably, deletion of degron 2 in USP2, but not degron 1, could reduce the interaction between SPOP and USP2 in this experimental setting ( xref )."

sparser
"Consistent with the fact that the N-terminal MATH (meprin and TRAF-C homology) domain of SPOP recognizes substrates, xref we found that deletion of the MATH domain eliminated SPOP binding to USP2."

sparser
"Depletion of the BTB domain, which was previously reported to largely mediate the interaction with the cullin 3 complex, xref did not affect SPOP binding to USP2 ( xref )."

sparser
"In further support of degron 2 as the critical motif recognized by SPOP, mutation of degron 2 also decreased the interaction between SPOP and USP2 ( xref )."
SPOP ubiquitinates USP2.
| 1
SPOP ubiquitinates USP2. 1 / 1
| 1

sparser
"We further confirmed that SPOP could promote the ubiquitination of wild-type USP2, but not the degron 2 mutant form of USP2 ( xref )."
SMAD7 affects USP2
4 | 2
4 | 2

No evidence text available

No evidence text available

No evidence text available

No evidence text available

sparser
"Specifically, USP2 interacted with SMAD7 and prevented SMAD7 ubiquitination."

sparser
"USP2 interacts with SMAD7 and prevents SMAD7 ubiquitination in Glioblastoma, while SMAD7 is a major inhibitor of TGFβ signaling xref ."
PEX5 affects USP2
1 | 3 1
1 | 3 1

reach
"By mammalian two-hybrid analysis, a clear binding between PEX5 and USP2 was observed, whereas no interaction was detected when the PTS1-sequence is deleted (DeltaSRM) (XREF_FIG)."

reach
"The results of the two-hybrid analysis indicate that the interaction between USP2 and PEX5 depends on the quality of the PTS1 and is significantly increased by conversion of the sequence."

reach
"Taking into account that USP2 interacts with the import receptor PEX5, deubiquitination of monoubiquitinated PEX5 could have been a peroxisome related function of the enzyme."

No evidence text available

sparser
"The results of the two-hybrid analysis indicate that the interaction between USP2 and PEX5 depends on the quality of the PTS1 and is significantly increased by conversion of the sequence."
NHERF4 affects USP2
| 3
| 3

sparser
"We observed that USP2-45 interacts with NHERF4 ( xref ) and endogenous Clathrin Heavy Chain (CLH) ( xref ), whereas the non-circadian isoform USP2-69 does not."

sparser
"Finally, USP2-45 interacts in vitro with NHERF4 and endogenous Clathrin Heavy Chain."

sparser
"Accordingly, USP2-45 interacts with NHERF4 and endogenous Clathrin Heavy Chain (CLH) in transfected HEK293 cells ( xref )."
LEF1 affects USP2
| 4 2
LEF1 binds USP2.
| 2 2
| 2 2

reach
"LEF1 protein stability analysis and co-immunoprecipitation (Co-IP) assay were performed to test the USP2/LEF1 interaction."

reach
"Through Co-IP assays using an anti-USP2 or anti-LEF1 antibody, we demonstrated the interaction between USP2 and LEF1 in hBMSCs, as indicated by LEF1 enrichment in USP2-associating precipitates and USP2 enrichment in LEF1-related precipitates (Fig. 4D)."

sparser
"Through Co-IP assays using an anti-USP2 or anti-LEF1 antibody, we demonstrated the interaction between USP2 and LEF1 in hBMSCs, as indicated by LEF1 enrichment in USP2-associating precipitates and USP2 enrichment in LEF1-related precipitates (Fig.  xref D)."

sparser
"These findings reveal a novel mechanism by which USP2 and LEF1 interact to regulate hBMSC osteogenic differentiation."
LEF1 activates USP2.
| 2
LEF1 activates USP2. 2 / 2
| 2

reach
"Increased expression of LEF1 reverses USP2 knockdown-imposed suppression on proliferation and osteogenic differentiation of hBMSCs."

reach
"Increased expression of LEF1 reversed USP2 knockdown-imposed suppression on proliferation and osteogenic differentiation of hBMSCs."
JUN affects USP2
6 |
JUN decreases the amount of USP2. 6 / 6
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
GST affects USP2
| 6
| 6

sparser
"We immunoprecipitated GPx4 complexes and incubated with GST-USP2, a DUB that can remove different types of ubiquitin chains ( xref )."

sparser
"The result showed that the recruitment of HOIP to GPx4 was eliminated by incubation with purified GST-USP2 ( xref and)."

sparser
"Additionally, the interaction between HOIP and GPx4 was also disrupted after in vitro deubiquitination assay mediated by GST-USP2 ()."

sparser
"Conversely, the recombinant GST-CD47, but not GST alone, bound to USP2 (Fig.  xref )."

sparser
"This interaction appears to be direct since purified Flag-VPRBP binds a purified GST-USP2 fusion protein but not GST alone (Fig.  xref )."

sparser
"The glutathione S-transferase (GST) pull-down assay showed that bacterially purified recombinant GST-USP2, but not GST alone, interacted with CD47 (Fig.  xref )."
DLL1 affects USP2
| 6
| 6

sparser
"These results suggest that USP2-45 binding to α 2 δ-1 promotes the de-ubiquitylation of both Ca v 1.2 and α 2 δ-1 subunits, in order to regulate the expression of Ca v 1.2 channels at the plasma membrane."

sparser
"This result suggests that the first critical event is the binding of USP2-45 to the auxiliary α 2 δ-1 subunit which may act as an anchor allowing for USP2-45 to de-ubiquitylate Ca v 1.2 channels."

sparser
"In addition, the binding of USP2-45 to α 2 δ-1 may disrupt the chaperone role of this subunit towards Ca v 1.2, leading to the reduction of Ca v 1.2 surface expression."

sparser
"However, USP2-45 specifically interacts with α 2 δ-1 rather than with the Ca V 1.2 channel or β 2 -subunit, thus it is not appropriate to define USP2-45 as a deubiquitinase of the Ca V 1.2 channel."

sparser
"Unexpectedly, the α 2 δ-1-USP2-45 protein interaction appears to downregulate the total expression of Ca V 1.2, α 2 δ-1 and β 2 subunits, and simultaneously to inhibit plasma membrane insertion and Ca 2+ current density of Ca V 1.2 channels in transfected HEK 293 cells."

sparser
"Though the reason for this peculiarity is presently unknown, it may lie in the fact that the USP2-45 binding to Ca V α 2 δ-1 might disrupt the chaperone role of the auxiliary subunit, leading to a reduction of the Ca V 1.2α 1 pore-forming subunit trafficked to the plasma membrane [ xref ]."
5 |
Sodium arsenite increases the amount of USP2.
3 |
Sodium arsenite increases the amount of USP2. 3 / 3
3 |

No evidence text available

No evidence text available

No evidence text available
Sodium arsenite decreases the amount of USP2.
2 |
Sodium arsenite decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
Monoubiquitin affects USP2
| 4 1
Monoubiquitin binds USP2. 5 / 5
| 4 1

reach
"As a support of this notion, the binding of monoubiquitin to the USP2 catalytic core is monophasic and lacks a conformational change step following the bimolecular binding step."

sparser
"Kinetics of monoubiquitin binding to USP2."

reach
"While a simple one-step binding of monoubiquitin to USP2 was observed, a biphasic binding was evident for diubiquitin."

reach
"Kinetics of monoubiquitin binding to USP2."

reach
"Here we developed two methods to follow the binding of monoubiquitin to USP2 catalytic core."
USP7 affects USP2
| 2 3
| 2 3

sparser
"Eleven of these water molecules are found at identical positions in the USP2 and USP7 complexes ( xref F)."

reach
"Indeed, the 76 ubiquitin Cα positions of the USP2 and USP7 complexes differ with an rmsd of only 0."

sparser
"It is possible that upon mutating Leu8 to Ala in Ub, the absence of the hydrophobic interaction shifts the conformation of the β1–β2 loop in Ub and reduced its binding to USP7 and USP2."

sparser
"Indeed, the 76 ubiquitin Cα positions of the USP2 and USP7 complexes differ with an rmsd of only 0.74 Å. This high structural similarity shows that the different protocols for the preparation of USP/ubiquitin complexes have no influence on the overall ubiquitin binding mode."

reach
"Eleven of these water molecules are found at identical positions in the USP2 and USP7 complexes (Figure 2F)."
USP2-AS1 affects USP2
| 5
USP2-AS1 activates USP2.
| 3
USP2-AS1 activates USP2. 3 / 3
| 3

reach
"Our subsequent assays were to verify whether USP2-AS1 positively regulated USP2 through modulating ETS1."

reach
"To conclude, USP2-AS1 promotes the osteogenic differentiation of HBMSCs by positively regulating USP2."

reach
"However, the results of luciferase reporter assay indicated that USP2-AS1 upregulated USP2 by activating its transcription."
USP2-AS1 increases the amount of USP2.
| 2
USP2-AS1 increases the amount of USP2. 2 / 2
| 2

reach
"In this study, we found that USP2-AS1 could positively regulate the expression of its nearby gene USP2."

reach
"Further exploration indicated that USP2-AS1 positively regulated the expression of its nearby gene USP2."
USP2 affects proteolysis
| 5
USP2 inhibits proteolysis.
| 3
| 3

reach
"The ubiquitin dependent proteolysis of cyclin D1 is antagonized by the deubiquitinating enzyme USP2, thus increasing cyclin D1 half-life."

reach
"Knockdown of USP2 or USP14 accelerated protein degradation of TNF-alpha, and abolished the effect of miR-124 on TNF-alpha protein stability."

reach
"Seventh, alcohol induced dramatic reduction of USP2, the major deubiquitinating enzyme may provide mechanistic perspectives underlying caveolar protein depletion and/or degradation as USP2 depletion has been documented to significantly enhance protein degradation."
USP2 activates proteolysis.
| 2
| 2

reach
"Other studies also indicated that USP2 augments the cancer cell cycle by interacting with cyclin D1 and reducing its protein degradation by ubiquitination [19]."

reach
"Functional experiments suggested USP2 inhibited lung cancer cell invasion and migration by reducing ARID2 protein degradation."
USP2 affects monoubiquitin
| 4 1
Monoubiquitin binds USP2. 5 / 5
| 4 1

reach
"As a support of this notion, the binding of monoubiquitin to the USP2 catalytic core is monophasic and lacks a conformational change step following the bimolecular binding step."

sparser
"Kinetics of monoubiquitin binding to USP2."

reach
"While a simple one-step binding of monoubiquitin to USP2 was observed, a biphasic binding was evident for diubiquitin."

reach
"Kinetics of monoubiquitin binding to USP2."

reach
"Here we developed two methods to follow the binding of monoubiquitin to USP2 catalytic core."
USP2 affects cisplatin
| 5
| 5

reach
"Our findings suggested that USP2 inhibited cisplatin resistance through promoting ferroptosis."

reach
"Overall, USP2 inhibited cisplatin resistance through promoting ferroptosis in NSCLC.USP2, a deubiquitinating enzyme, performs crucial biological functions related to ubiquitination."

reach
"38 PR-619 and ML364, inhibitors of pan-DUBs and USP2, effectively enhance the anti-cancer effect of cisplatin in metastatic bladder urothelial carcinoma patients ( Table 2 )."

reach
"Therefore, we hypothesized that USP2 might inhibit cisplatin resistance of NSCLC cells through promoting ferroptosis."

reach
"USP2 overexpression inhibited cisplatin resistance through promoting ferroptosis in A549/DDP and H1299/DDP cells."
USP2 affects USP7
| 2 3
| 2 3

sparser
"Eleven of these water molecules are found at identical positions in the USP2 and USP7 complexes ( xref F)."

reach
"Indeed, the 76 ubiquitin Cα positions of the USP2 and USP7 complexes differ with an rmsd of only 0."

sparser
"It is possible that upon mutating Leu8 to Ala in Ub, the absence of the hydrophobic interaction shifts the conformation of the β1–β2 loop in Ub and reduced its binding to USP7 and USP2."

sparser
"Indeed, the 76 ubiquitin Cα positions of the USP2 and USP7 complexes differ with an rmsd of only 0.74 Å. This high structural similarity shows that the different protocols for the preparation of USP/ubiquitin complexes have no influence on the overall ubiquitin binding mode."

reach
"Eleven of these water molecules are found at identical positions in the USP2 and USP7 complexes (Figure 2F)."
USP2 affects UCP2
| 5
USP2 inhibits UCP2.
| 2
USP2 inhibits UCP2. 2 / 2
| 2

reach
"Since USP2 stabilizes PGC1α by deubiquitination, USP2 might eliminate mitochondrial ROS by activating the PGC1α–UCP2 axis."

reach
"Based on this idea, a putative protein digestion mechanism might reduce the excessive accumulation of UCP2 in C2C12 cells.As found in the current study, the inhibition of USP2 interrupts mitochondrial ATP synthesis in myoblasts, leading to defects in proliferation and differentiation [33,42]."
USP2 increases the amount of UCP2.
| 2
USP2 increases the amount of UCP2. 2 / 2
| 2

reach
"Considering that the tissue distribution of USP2 is also relatively ubiquitous [50], USP2 might prevent oxidative stress in various tissues through the preservation of UCP2.The lack of USP2 dramatically repressed UCP2 at the mRNA level, implying that USP2 stimulates the transcription of the Ucp2 gene."

reach
"Since we did not demonstrate the involvement of PGC1α in USP2-modulated UCP2 expression, we cannot exclude the possibility that USP2 promotes Ucp2 expression via other transcriptional regulators."
USP2 activates UCP2.
| 1
USP2 activates UCP2. 1 / 1
| 1

reach
"Comprehensive RT-qPCR screening and following protein analysis indicated that both the genetic and chemical inhibition of USP2 elicited a decrease in uncoupling protein 2 (UCP2) at mRNA and protein levels."
USP2 affects SPOP
| 5
| 5

sparser
"Moreover, we demonstrated that SPOP interacts with USP2, promoting its ubiquitination and subsequent degradation, which in turn facilitates CD47 degradation ( xref xref )."

sparser
"Notably, deletion of degron 2 in USP2, but not degron 1, could reduce the interaction between SPOP and USP2 in this experimental setting ( xref )."

sparser
"Consistent with the fact that the N-terminal MATH (meprin and TRAF-C homology) domain of SPOP recognizes substrates, xref we found that deletion of the MATH domain eliminated SPOP binding to USP2."

sparser
"Depletion of the BTB domain, which was previously reported to largely mediate the interaction with the cullin 3 complex, xref did not affect SPOP binding to USP2 ( xref )."

sparser
"In further support of degron 2 as the critical motif recognized by SPOP, mutation of degron 2 also decreased the interaction between SPOP and USP2 ( xref )."
USP2 affects INS
| 5
USP2 activates INS.
| 3
USP2 activates INS. 3 / 3
| 3

reach
"Ubiquitin specific peptidase 2 (USP2) expression is decreased in patients with type 2 diabetes and mouse models of diabetes-induced muscle atrophy.USP2 can enhance insulin sensitivity and improve muscle mass and function in diabetic mice.USP2 stabilizes PPAR-gamma via deubiquitination to regulate muscle atrophy, uncovering a key mechanism in this process.The findings indicate the USP2-PPAR-gamma axis has potential as a therapeutic target for metabolic disorders and sarcopenia."

reach
"Correspondingly, infection with an adenovirus encoding Usp2 shRNA significantly lowered blood glucose, blood insulin, hepatic glycogen content, and insulin sensitivity in high fat diet–fed mice, suggesting that USP2 plays a role in aggravating T2DM pathogenesis [102]."

reach
"USP2 modulates glucose and lipid metabolism, insulin sensitivity, and inflammatory processes."
USP2 inhibits INS.
| 2
USP2 inhibits INS. 2 / 2
| 2

reach
"Our current cellular experiments suggest that macrophage USP2 inhibits the insulin resistance of myocytes in an adipocyte dependent manner."

reach
"Macrophage USP2 may therefore selectively prevent age- and/or adiposity related chronic inflammation and subsequent insulin resistance."
USP2 affects Hypertrophy
| 5
USP2 activates Hypertrophy.
| 3
| 3

reach
"The above data suggested that USP2 overexpression alleviated Ang II-induced hypertrophy of cardiomyocytes.After the determination of calcium overload in vitro , it was found that Ang II induced increa[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Collectively, we hypothesized that Mycn possibly regulates USP2 transcription, which modulates JUP protein stability and the Akt/β-catenin signaling to affect cardiac hypertrophy progression."

reach
"Accordingly, our results revealed that USP2 overexpression improved cardiac functions, decreased the ratios of HW/BW and HW/TL, and reduced the degree of cardiac hypertrophy and fibrosis along with di[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 inhibits Hypertrophy.
| 2
| 2

reach
"In addition, by comparing the GEO dataset and the mRNA level of USP2 in hypertrophic heart induced by trans-arterial coarctation (TAC), it was determined that the expression of USP2 was downregulated in hypertrophic heart, and the overexpression of USP2 could relieve the cardiac hypertrophy induced by TAC."

reach
"Altogether, we initially demonstrated that USP2 overexpression suppressed hypertrophy, calcium overload, and mitochondrial dysfunction and increased protein levels of MFN2 via deubiquitination in Ang [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects FOXC1
| 3 2
USP2 binds FOXC1.
| 2 2
| 2

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"After Ubibrowser online database prediction, the interaction between USP2 and FOXC1 was verified using Co-immunoprecipitation (CoIP) assay."

reach
"Mechanistically, USP2 directly interacted with FOXC1 and deubiquitinated FOXC1, therefore enhancing FOXC1 protein stability."
USP2 binds FOXC1. 2 / 2
| 2

sparser
"After Ubibrowser online database prediction, the interaction between USP2 and FOXC1 was verified using Co-immunoprecipitation (CoIP) assay."

sparser
"Mechanistically, USP2 directly interacted with FOXC1 and deubiquitinated FOXC1, therefore enhancing FOXC1 protein stability."
USP2 deubiquitinates FOXC1.
| 1
USP2 deubiquitinates FOXC1. 1 / 1
| 1

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"Mechanistically, USP2 directly interacted with FOXC1 and deubiquitinated FOXC1, therefore enhancing FOXC1 protein stability."
| 5
USP2 activates Cell Survival.
| 3
| 3

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"Knockdown or inhibition of USP2 alone was found to reduce TNBC cell viability."

reach
"For example, USP2 and USP7 modulate p53 and MDM2, impacting cell survival and tumor development, while USP22 stabilizes c-Myc, driving cancer proliferation and metastasis [9–13]."

reach
"USP2 knockdown decreases MyLa2000 cell viability after PUVA by 50% but not Hut-78, suggesting that the function of USP2 in CTCL cells is p53 dependent."
USP2 inhibits Cell Survival.
| 2
| 2

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"USP2 overexpression inhibited cell viability in cisplatin-resistant cells."

reach
"Further research revealed that USP2 overexpression suppressed cell viability in DDP-resistant NSCLC and promoted ferroptosis."
USP2 affects AURKA
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RIPK1 affects USP2
4 | 1
4 | 1

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sparser
"For example, when the N-terminus of USP2 binds to receptor interacting protein-1 (RIP1) protein, it removes the ubiquitinated molecules on RIP1 protein and increases RIP1 expression, which in turn promotes apoptosis ( xref )."

No evidence text available
NBN affects USP2
2 | 3
2 | 3

sparser
"However, USP2 can interact with NBS1 protein when both proteins are overexpressed ( xref C ), and direct interaction between USP2 and SKP2 was also observed in a previous study ( xref )."

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sparser
"On the other hand, USP2 stably interacted with NBS1 upon NCS treatment, and their interaction persisted regardless of NCS treatment ( xref C )."

sparser
"As shown in this study, USP2 stably interacts with NBS1 ( xref ) and affects ubiquitination of NBS1 ( xref and xref )."
MYCN affects USP2
| 3 2
MYCN increases the amount of USP2.
| 3
MYCN increases the amount of USP2. 3 / 3
| 3

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"Here, we observed that the USP2 expression was decreased in mice with HCM and restored by Mycn overexpression."

reach
"Collectively, we hypothesized that Mycn possibly regulates USP2 transcription, which modulates JUP protein stability and the Akt/β-catenin signaling to affect cardiac hypertrophy progression."

reach
"Mycn activates USP2 transcription."
MYCN binds USP2.
| 2
| 2

sparser
"Mycn bound to the USP2 promoter to activate its transcription."

sparser
"Collectively, this evidence suggests that the Mycn binds to the USP2 promoter for its transcription activation."
MYC affects USP2
2 1 | 2
MYC binds USP2.
1 | 2
1 | 2

sparser
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure xref ) and interactive Flag-ARID2 (Supplementary Figure xref ) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

sparser
"Immunoprecipitation assay showed that Flag-KRAS interacted with Myc-USP2 ( xref I)."

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MYC decreases the amount of USP2.
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MYC decreases the amount of USP2. 2 / 2
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MEF2A affects USP2
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MEF2A decreases the amount of USP2. 5 / 5
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Imd affects USP2
| 5
| 5

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"The control of Imd homeostasis by USP2 is associated with the hydrolysis of Imd linked K48-ubiquitin chains and the synergistic binding of USP2 and Imd to the proteasome, as evidenced by both mass-spectrometry analysis of USP2 partners and by co-immunoprecipitation experiments."

reach
"Focusing on USP2 biochemical function, we show that USP2 binds to Imd and promotes the cleavage of Ub K48 chains from the protein in both cultured cells and flies."

reach
"USP2 interacts with Imd and promotes cleavage of Imd linked Ub K48."

reach
"By using GST pull-down assays with GST tagged full length or truncated forms of USP2 (USP2-N-ter [AA :1-531] and USP2-C-ter [AA :475-856]), we finally showed that USP2 preferentially interacts with Imd through its non catalytic N-ter domain."

reach
"Thus, the Imd and USP2 complex apparently binds more efficiently to the proteasome than do Imd or USP2 alone."
AURKA affects USP2
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ADIPOQ affects USP2
| 5
ADIPOQ increases the amount of USP2. 5 / 5
| 5

reach
"Herein, we present the evidence that changes in USP-2 expression by adiponectin and leptin contribute to the regulation of cyclin D1 degradation by proteasomes that impacts cell cycle modulation in ca[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"As expected from our microarray data, adiponectin did not trigger elevated transcript levels of TR2, CASP1 and USP2 in MCF-7 breast cancer cells, but a significantly reduced USP2 expression by approximately 60%."

reach
"Although USP-2 seems to play a critical role in USP-2 expression modulated by adiponectin and leptin, the only little information regarding molecular mechanisms underlying USP-2 expression is availabl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"It would be interesting to examine the role of AMPK and JAK and STAT signaling in the modulation of USP-2 expression by adiponectin and leptin for the future study.Cyclin D1, an important regulator of[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In the present study, we clearly demonstrated that adiponectin and leptin modulates expression of USP-2 expression in the opposite manner, which in turn directly impacts cyclin D1 degradation (Figs. 3[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
4 |
Valproic acid increases the amount of USP2.
3 |
Valproic acid increases the amount of USP2. 3 / 3
3 |

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Valproic acid decreases the amount of USP2.
1 |
Valproic acid decreases the amount of USP2. 1 / 1
1 |

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| 2

reach
"Consistently, we found that PPAR-γ knockdown increased and Usp2 overexpression decreased the expression of FoxO signaling pathway-related genes, including cyclin G2 (Ccng2), cyclin-dependent kinase inhibitor 1B (Cdkn1b), retinoblastoma-like protein 2 (Rbl2), and Bcl-2 interacting protein 3 (Bnip3) (Fig. 6B)."

reach
"Conversely, USP2 inhibits glioblastoma development by suppressing the TGF-B signaling pathway through the deubiquitination of SMAD7 [19]."
| 2

reach
"This study provides an understanding of the USP2-mediated signaling pathway in TNBC, emphasizing the role of USP2 and its substrates with apoptotic genes."

reach
"Afterwards, we observed from TOP/FOP flash assay that USP2 silencing significantly inhibited the activity of Wnt/β-catenin signalling pathway (Figure 6G)."

reach
"Thus, it is plausible to consider that disruptions in the cell cycle regulation caused by USP2 inhibition could substantially influence viral replication."

reach
"In addition, USP2 inhibition primarily led to cell cycle arrest, and numerous viruses exploit the host cell cycle machinery to support their replication."

reach
"ML364, a small molecule inhibitor of USP2, causes cell cycle arrest through degradation of cyclin D1 in colorectal cancer [13]."

eidos
"Since leptin and adiponectin have opposite effects on USP2 expression in these cells , USP2 likely contributes to cell cycle regulation via adipokines [ 41 ] ."
| 4
USP2 inhibits muscle atrophy.
| 2

reach
"Taken together, these results revealed a previously unrecognized yet critical role of USP2 in the regulation of PPAR-γ signaling and in the pathogenesis of muscle atrophy.Our mechanistic investigations revealed that USP2 prevents muscle atrophy by regulating the stability of PPAR-γ, a key TF implicated in muscle metabolism and insulin sensitivity (31–33)."

reach
"The global deletion of USP2 aggravated diabetes- and DEX-induced muscle atrophy and insulin resistance in mice."
USP2 activates muscle atrophy.
| 2
| 2

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"We investigated the changes in endogenous USP2 expression in diabetes-induced muscle atrophy mouse models and found that the Gas and TA muscles of diabetic mice (STZ + HFD; spontaneous diabetes mouse model) had lower USP2 levels than their respective Ctrl groups (Supplementary Fig. 1B and C)."

reach
"USP2 Overexpression Protects Against Diabetes-Induced Muscle Atrophy in Mice."
| 4

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"USP2 increases hepatic gluconeogenesis and exacerbates glucose intolerance in diet-induced obese mice (Molusky et al., 2012)."

reach
"Both circadian and USP2 deregulations accelerate the process of hepatic gluconeogenesis by increasing glucose secretion."

reach
"Thus, USP7 prevents T2DM by (1) improving insulin sensitivity, and (2) inhibiting gluconeogenesis.Previous reports have demonstrated that two additional USPs, namely USP2 and USP14, modulate hepatic gluconeogenesis [102,103]."

reach
"USP2 and 14 enhance gluconeogenesis, leading to high blood glucose levels."

reach
"Taken together, these data suggest that pharmacological inhibition of USP2 prevents dysregulation of ECM and promotes the integrity of the epithelial barrier in the gut during inflammation, and that targeting USP2 might provide a potential strategy for therapeutic intervention of inflammatory diseases in the gut.3 Discussion."

reach
"In addition, results from Masson trichromatic staining showed that collagens (major components of ECM in the gut) (Rieder et al., 2013) accumulated more intensively in the colon tissues from Usp2 mice than Lyz2-Cre;Usp2 mice after colitis induction (Fig. 5D), indicating that knockout of USP2 impairs the ECM remodeling in the colon inflammation."

reach
"These data together suggest that knockout of USP2 in myeloid cells inhibits dysregulation of ECM in the colon after DSS treatment.To further confirm that the myeloid cells trained the epithelium to induce dysregulation of ECM, we isolated RFP  cells from the LPMCs of Lyz2-Cre;Usp2 Ai14 mice Lyz2-Cre;Ai14 mice that were treated with DSS and cultured these cells for 18 h."

reach
"In addition, knockout of USP2 in myeloid cells inhibits the production of pro-inflammatory cytokines to relieve the dysregulation of extracellular matrix (ECM) network and promote the gut epithelial integrity after DSS treatment."
USP2 affects cell growth
| 3
USP2 activates cell growth.
| 2
| 2

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"To completely exclude the possibility that the cell growth repression induced by USP2 knockdown is p53 dependent, we performed the same experiments in isogenic p53 knockout cells (HCT116 p53-/-)."

reach
"Again, knockdown of USP2 in MCF-7 and PC3 cells significantly reduced the steady state levels of cyclin D1 protein (XREF_FIG, also in XREF_SUPPLEMENTARY) and also induced cell growth suppression (XREF_FIG)."
USP2 inhibits cell growth.
| 1
| 1

reach
"These data demonstrate that inactivation of USP2 downregulates the levels of endogenous cyclin D1 and suppresses cell growth by delaying G1/S progression."
USP2 affects VCP
| 4
| 4

sparser
"Direct physical interaction of VCP with USP2 was observed in 293 T cells, demonstrating that domain 2 of VCP and the amino acid sequence (200-403) of USP2 were critical for their interactions (Fig. xref )."

sparser
"To further clarify the interaction between VCP and USP2, we introduced an inhibitor targeting the D2 domain of VCP."

sparser
"To confirm whether CB-5083 disrupted the binding of USP2 to the D2 domain of VCP, we conducted CO-IP experiments."

sparser
"The results exhibited that CB-5083 disrupted the VCP-USP2 interaction (Supplementary Fig. xref )."
USP2 affects TRIM27
3 | 1
USP2 binds TRIM27.
3 |
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USP2 activates TRIM27.
| 1
USP2 activates TRIM27. 1 / 1
| 1

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"Importantly, when RFP was immunoprecipitated in the presence of NEM and then treated with the deubiquitinating enzyme Usp2 the slower migrating band was lost in both RFP and HA western blots (XREF_FIG, lane 12), confirming that " JMD-Ub " was ubiquitinated."
USP2 affects Proteasome
| 4
USP2 binds Proteasome.
| 2

reach
"USP2 and Imd synergistically interact with the proteasome."

reach
"Taken together, our data suggest that USP2 and Imd synergistically bind to the proteasome and that USP2 contributes to Imd degradation at the proteasome level."
USP2 activates Proteasome.
| 2
| 2

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"At the molecular level, the simplest explanation for this result is that USP2 drives the Ub K48 -Imd molecules to the proteasome and/or ensures the removal of Ub K48 from Imd at the level of the proteasome, then allowing for Imd degradation by the proteasome machinery."

reach
"USP2 directly interacts with cyclin D1 to perform deubiquitylation and thereby antagonize proteasome-dependent degradation of cyclin D1."
USP2 affects PPARGC1B
| 4
USP2 inhibits PPARGC1B.
| 2
| 2

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"Indeed, the overexpression of the Usp2 gene decreased the quantity of K48-linked polyubiquitin chains on PGC1α, suggesting that USP2 is a direct stabilizing enzyme for PGC1α through deubiquitination."

reach
"Since USP2 stabilizes PGC1α by deubiquitination, USP2 might eliminate mitochondrial ROS by activating the PGC1α–UCP2 axis."
USP2 activates PPARGC1B.
| 2
USP2 activates PPARGC1B. 2 / 2
| 2

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"Accordingly, an overexpression study using 293FT cells demonstrated that the introduction of USP2 increased the accumulation of Halo-tagged PGC1α in the nucleus (Figure 5C)."

reach
"Given that the protein levels of USP2 or C276A were comparable in USP2- and C276A-transfected cells, USP2 modulates PGC1α levels via ubiquitin isopeptidase activity.Finally, we examined whether USP2 directly digested the lysine48 (K48)-linked polyubiquitin chains on PGC1α, since K48-linked polyubiquitin chanins represent a canonical signal for proteasome degradation [41]."
USP2 affects NFkappaB
| 3
| 3

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"Ubiquitin-specific peptidase 2 inhibits epithelial-mesenchymal transition in clear cell renal cell carcinoma metastasis by downregulating the NF-kappaB pathway."

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"The ubiquitin #specific proteases system 2 (USP2) stabilizes the ubiquitin-E3-ligase ITCH and lowers NF-kappaB basal activity, which leads to reduced c-FLIP mRNA production; proteasomal degradation of c- FLIP isoforms is also elevated by its negative regulator proteasome ITCH [XREF_BIBR]."

reach
"USP2a, a 69-kDa splice variant of USP2, also known as USP9 and UBP41, negatively regulates NF-kappaB activation and proinflammatory cytokine production by interacting with TRAF6."
USP2 affects NEDD4L
| 4
USP2 inhibits NEDD4L.
| 2
USP2 inhibits NEDD4L. 2 / 2
| 2

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"In cells coexpressing USP2-45/-69, the amount of ubiquitylated KCNQ1 was significantly decreased and the total amount of KCNQ1 was increased, demonstrating that USP2 antagonizes the effect of Nedd4-2 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The biochemical experiments show an augmented surface expression and reduced ubiquitylation level of KCNQ1 when catalytically active USP2 is coexpressed with Nedd4-2, suggesting that USP2 may counter [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 activates NEDD4L.
| 2
USP2 activates NEDD4L. 2 / 2
| 2

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"However, coexpression of USP2 together with Nedd4-2 and KCNQ1 augments the binding efficiency of Nedd4-2, indicating that either both USP2-45 and USP-69 stabilize the binding to the KCNQ1 protein or N[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Interestingly, Nedd4-2 coimmunoprecipitation with wild-type KCNQ1 was increased when USP2 was coexpressed, suggesting that USP2 may augment the binding of Nedd4-2 to the channel complex."
USP2 affects MYC
1 | 1 2
USP2 binds MYC.
1 | 2
1 | 2

sparser
"Using anti-Flag or anti-Myc antibody precipitated protein as input, interactive Myc-USP2 protein (Supplementary Figure xref ) and interactive Flag-ARID2 (Supplementary Figure xref ) were, respectively, detected; the above findings provided direct evidence of the interaction of USP2 and ARID2 in cell line models."

sparser
"Immunoprecipitation assay showed that Flag-KRAS interacted with Myc-USP2 ( xref I)."

No evidence text available
USP2 increases the amount of MYC.
| 1
USP2 increases the amount of MYC. 1 / 1
| 1

reach
"The deubiquitinating enzyme USP2 has been found to enhance MYC levels through the modulation of specific subsets of microRNAs in prostate cancer [XREF_BIBR]."
USP2 affects MFN2
| 4
USP2 increases the amount of MFN2.
| 2
USP2 increases the amount of MFN2. 2 / 2
| 2

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"Our subsequent experiments showed that the protein level of MFN2 was decreased both in vivo and in vitro models, which is in consistent with its expression pattern in hypertrophic cardiomyocytes repor[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In the current study, our findings supported that USP2 overexpression attenuates calcium overload, upregulates MFN2 expression through deubiquitination, and further alleviates mitochondrial dysfunctio[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 deubiquitinates MFN2.
| 2
USP2 deubiquitinates MFN2. 2 / 2
| 2

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"However, whether USP2 induces the deubiquitination of MFN2 to regulate cardiac hypertrophy has not been reported before and warrants further exploration.In light of the above associations, we raised a[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"USP2 overexpression decreases the concentration of Ca and increases SERCA2a by deubiquitinating mitochondrial protein Mitofusin-2 (Mfn2), alleviating mitochondrial dysfunction, and apoptosis in the Ang II-induced cardiac hypertrophy mouse model [150]."
USP2 affects KRT15
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USP2 affects JUP
| 4
USP2 activates JUP.
| 3
USP2 activates JUP. 3 / 3
| 3

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"This effect is achieved through the activation of USP2, which enhances the stability of JUP protein via deubiquitination modification, subsequently blocking the Akt/β-catenin pathway (Fig. 8)."

reach
"Indeed, we found that the USP2 overexpression in cardiomyocytes enhanced the protein stability of JUP, which is a novel interaction that has been reported before."

reach
"Collectively, we hypothesized that Mycn possibly regulates USP2 transcription, which modulates JUP protein stability and the Akt/β-catenin signaling to affect cardiac hypertrophy progression."
USP2 deubiquitinates JUP.
| 1
USP2 leads to the deubiquitination of JUP. 1 / 1
| 1

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"We hypothesized that USP2 possibly mediates JUP deubiquitination and inactivates the Akt/β-catenin cascade to ameliorate cardiac hypertrophy."
USP2 affects HR3
| 4
USP2 inhibits HR3.
| 2
USP2 inhibits HR3. 2 / 2
| 2

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"Expression of either EcR-A ( Hiruma and Riddiford, 2003 ) or USP-2 ( Lan et al., 1999 ) together with EcR-B1 and USP-1 in the GV1 cells reduced the activation of MHR3 by 20E about two-fold and also th[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In the cell line, high constitutive expression of the EcR-B1/USP-1 complex increases this 20E-induced activation by binding to the ecdysone response element in the promoter region of the MHR3 gene, bu[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 activates HR3.
| 2
USP2 activates HR3. 2 / 2
| 2

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"In M. sexta, the MHR3 was activated by the application of EcRB1 and USP1 but not by EcRB1 and USP2 (Lan et al., 1999)."

reach
"A delayed-early gene encoding MHR3 could be activated by EcRB1 and USP1 but not by EcRB1 and USP2 in Manduca GV1 cells (Lan et al., 1999)."
USP2 affects BMAL1
| 2 2
USP2 deubiquitinates BMAL1.
| 2
USP2 deubiquitinates BMAL1. 2 / 2
| 2

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"Accordingly, other investigators have shown that UBP41, a truncated form of USP2, can deubiquitinate BMAL1 in vitro, and that in the presence of USP2b, BMAL1 appears less ubiquitinated and more stable."

reach
"Further, USP2 is able to deubiquitinate the ubiquitinated forms of the clock proteins BMAL1 (Scoma et al. 2011), PER1 (Yang et al. 2012) and CRY1 (Tong et al. 2012)."
USP2 binds BMAL1.
| 2
| 2

sparser
"Consistently, our work revealed that DCAF7 served as a scaffold protein to promote the interaction between USP2 and BMAL1 and thus to stabilize BMAL1."

sparser
"Moreover, DCAF7 notably increased the interaction between USP2 and BMAL1 (Fig. xref and Fig. xref ), whereas DCAF7 knockdown markedly reduced the interaction of BMAL1 with both USP2 isoforms (USP2a and USP2b) (Fig. xref )."
TNF affects USP2
| 4
TNF inhibits USP2.
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TNF inhibits USP2. 2 / 2
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"TNF-α treatment downregulated USP2 (Fig. 5D and F)."

reach
"However, tumour necrosis factor alpha (TNFalpha) has been reported to downregulate a USP2 isoform in hepatocytes 8, so possibly, USP2 inhibited degradation of LDLR is a pathway that is affected in inflammatory conditions."
TNF activates USP2.
| 2
TNF activates USP2. 2 / 2
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"Here we investigated the role of TNFalpha induced down-modulation of the de-ubiquitinating enzyme USP2 for hepatocyte survival."

reach
"Our results indicate that TNFalpha induced USP2 down-regulation is an effective cytoprotective mechanism in hepatocytes."
PUVA affects USP2
| 4
PUVA increases the amount of USP2.
| 2
PUVA increases the amount of USP2. 2 / 2
| 2

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"PUVA caused a significant (P < 0.01) upregulation of USP2 mRNA expression already after 8 hours with an even stronger increase after 24 and 48 hours post-treatment."

reach
"More importantly, PUVA did not upregulate USP2 mRNA expression in Hut-78 cells and only slightly in SeAx cells."
PUVA activates USP2.
| 2
PUVA activates USP2. 2 / 2
| 2

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"USP2 is upregulated by PUVA and a p53 activator, nutlin3a."

reach
"We propose that a negative regulatory loop in which USP2 is induced by PUVA via p53 increases cell resistance to apoptosis via deubiquitination of Mdm2."
KRT15 affects USP2
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KCNQ1 affects USP2
2 | 1 1
2 | 1 1

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"A reduced level of ubiquitylation leads to an increased surface density of KCNQ1 channels and thereby explains the rescued current level observed in the functional experiments.Whether or not Nedd4-2 i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Whether or not Nedd4-2 is coexpressed does not seem to influence the binding of USP2 to KCNQ1, as both USP2-45 and USP-69 bind equally well under the 2 conditions ( Figure 3 )."
FOXC1 affects USP2
| 2 2
| 2

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"After Ubibrowser online database prediction, the interaction between USP2 and FOXC1 was verified using Co-immunoprecipitation (CoIP) assay."

reach
"Mechanistically, USP2 directly interacted with FOXC1 and deubiquitinated FOXC1, therefore enhancing FOXC1 protein stability."
USP2 binds FOXC1. 2 / 2
| 2

sparser
"After Ubibrowser online database prediction, the interaction between USP2 and FOXC1 was verified using Co-immunoprecipitation (CoIP) assay."

sparser
"Mechanistically, USP2 directly interacted with FOXC1 and deubiquitinated FOXC1, therefore enhancing FOXC1 protein stability."
CRY1 affects USP2
3 | 1
3 | 1

sparser
"Targeting the CRY1-USP2 feedback loop ( xref ) offers a promising avenue for therapeutic intervention in Group 3 MB."

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BMAL1 affects USP2
| 2 2
BMAL1 binds USP2.
| 2
| 2

sparser
"Consistently, our work revealed that DCAF7 served as a scaffold protein to promote the interaction between USP2 and BMAL1 and thus to stabilize BMAL1."

sparser
"Moreover, DCAF7 notably increased the interaction between USP2 and BMAL1 (Fig. xref and Fig. xref ), whereas DCAF7 knockdown markedly reduced the interaction of BMAL1 with both USP2 isoforms (USP2a and USP2b) (Fig. xref )."
BMAL1 activates USP2.
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BMAL1 activates USP2. 2 / 2
| 2

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"This suggests a scenario in USP2 -/- mice in which increased turnover of BMAL1 coincides with increased CLOCK and BMAL1 driven gene expression in the USP2 -/- SCN."

reach
"The circadian rhythm of Usp2 is blunted in Clock mutant and Bmal1 KO mice, and the Usp2 promoter is activated by CLOCK and BMAL1, indicating that Usp2 is a direct target of these transcription factors."
3 |
Tetrachloromethane decreases the amount of USP2. 3 / 3
3 |

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Epithelial sodium channel affects USP2
| 3
USP2 binds epithelial sodium channel. 3 / 3
| 3

sparser
"The interaction of Usp2-45 with ENaC exogenously expressed in HEK293 cells was tested by coimmunoprecipitation."

sparser
"Here, in HEK293 cells stably transfected with ENaC, Usp2-45 interacted with ENaC, leading to deubiquitylation of the channel and stimulation of ENaC activity >20-fold."

sparser
"They report that Usp2-45 interacts directly with ENaC, although not with the PY-motif, to alter both cell surface expression and furin, but not prostasin, cleavage."
ZEB1 affects USP2
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ZEB1 decreases the amount of USP2. 3 / 3
3 |

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UbV.2.6 affects USP2
| 3
UbV.2.6 inhibits USP2. 3 / 3
| 3

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"To better understand the basis for the potent inhibition of USP2 by UbV.2.6, we introduced each of the four substitutions from UbV.2.6 individually into the sequence of Ub.wt and assessed effects on U[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"The substitutions at positions 6 and 11 had the greatest effects, and indeed, a UbV bearing substitutions at both positions was only 6-fold less potent than Ub.V.2.6.To better understand the basis for[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Taken together, the mutational and structural analyses revealed that a triad of substituted residues in UbV.2.6 forms the center of the interface with USP2, and the side chains of Cys 6 and Arg 11 sub[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Ub-AMC affects USP2
| 3
Ub-AMC binds USP2. 3 / 3
| 3

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"Notably, both native ubiquitin and Ub-AMC bind to USP2 with monophasic kinetics."

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"Binding of Ub-AMC to USP2 under an equilibrium condition."

reach
"We also determined the equilibrium binding constant of Ub-AMC binding to USP2 by measuring the decrease in Ub-AMC fluorescence in the presence of increasing concentrations of C276A USP2."
USP25 affects USP2
| 2 1
| 2 1

sparser
"We found that Ubvs that bind to USP2 or USP21 contain a remarkably similar core functional epitope, or &quot;hot spot,&quot; consisting mainly of positions that are conserved as the wild type sequence, but also some positions that prefer mutant sequences."

reach
"In the USP2 and USP21 complex structure, the side chain of Ub Lys48 does not occupy the nearby pocket on USP (see XREF_FIG)."

reach
"We first screened a panel of DUBs and found that both USP2 and USP21 bound to endogenous SKP2, but only USP2 deubiquitylated and stabilized SKP2 protein."
USP2 affects epithelial sodium channel
| 3
USP2 binds epithelial sodium channel. 3 / 3
| 3

sparser
"The interaction of Usp2-45 with ENaC exogenously expressed in HEK293 cells was tested by coimmunoprecipitation."

sparser
"Here, in HEK293 cells stably transfected with ENaC, Usp2-45 interacted with ENaC, leading to deubiquitylation of the channel and stimulation of ENaC activity >20-fold."

sparser
"They report that Usp2-45 interacts directly with ENaC, although not with the PY-motif, to alter both cell surface expression and furin, but not prostasin, cleavage."

reach
"USP2a, a 69-kDa splice variant of USP2, also known as USP9 and UBP41, negatively regulates NF-kappaB activation and proinflammatory cytokine production by interacting with TRAF6."

reach
"In LPS stimulated macrophage like cell line J774.1, the knockdown of USP2 promoted cytokine production, whereas the ectopic expression of USP2 repressed cytokine production."

reach
"In this study, we demonstrated that USP2KD promotes 25 of 104 cytokines in macrophage like cells after LPS stimulation, while ectopic expression of USP2 represses most cytokine production following this stimulation."
| 3
| 3

reach
"As shown in Figures 4(a) and (b), USP2 shRNA transfection uniformly enhanced H1299 and A549 cell migration and invasion, while USP2 overexpression vector transfection significantly suppressed the migrative and invasive capability of cancer cells.Subsequent wound healing assay suggested that H1299 cells with USP2 shRNAs transfection elevated cancer cell migration and wound healing after 48 h of incubation (Supplementary Figure 2 A-B)."

reach
"It is upregulated in various malignancies, such as triple negative breast cancer [17] and choroidal melanoma [18] and USP2 overexpression stimulates cell migration and invasion."

reach
"USP2 promotes cell migration and invasion in triple negative breast cancer cell lines."

reach
"Our findings provide the first demonstration of the USP2/LEF1 cascade that enhances the osteogenic differentiation of hBMSCs, broadening the field for the development of BMSCs as effective agents in osteoporosis therapy."

reach
"By studying USP2 and LEF1 expression alterations, we observed their roles in osteoporosis pathogenesis.Collectively, our study provides the first demonstration of the USP2/LEF1 cascade that enhances the osteogenic differentiation of hBMSCs, as illustrated in Fig. 7."

reach
"Moreover, increased USP2 expression reduced bone loss and enhanced osteogenic differentiation in OVX mice."
USP2 affects cell death
| 2
| 2

reach
"Accordingly, overexpression of a catalytically active UBP41 allele was shown to induce cell death in several cancer cell lines [41] ."

reach
"Moreover, USP2 silencing enhanced the iron ion concentration and lipid peroxidation accumulation as well as suppressed ferroptosis, while upregulating USP2 promoted ferroptotic cell death in ESCC cells."
USP2 affects calcium(2+)
| 3
| 3

reach
"Our subsequent results revealed that Ang II induced increases in Ca 2+ content and expression levels of calcium overload-related protein t-CaMKⅡ, and p-CaMKⅡ and a reduction in SERCA2 expression, wher[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Altogether, we initially demonstrated that USP2 overexpression suppressed hypertrophy, calcium overload, and mitochondrial dysfunction and increased protein levels of MFN2 via deubiquitination in Ang [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In summary, we found that the inactivation of Usp2 in mouse and of its orthologue CG14619 in Drosophila does not affect the circadian free running period, but impairs bodily Ca 2+ homeostasis in both species, especially in dietary Ca 2+ absorption in mouse small intestine."
USP2 affects Ub-AMC
| 3
Ub-AMC binds USP2. 3 / 3
| 3

reach
"Notably, both native ubiquitin and Ub-AMC bind to USP2 with monophasic kinetics."

reach
"Binding of Ub-AMC to USP2 under an equilibrium condition."

reach
"We also determined the equilibrium binding constant of Ub-AMC binding to USP2 by measuring the decrease in Ub-AMC fluorescence in the presence of increasing concentrations of C276A USP2."
USP2 affects USP25
| 2 1
| 2 1

sparser
"We found that Ubvs that bind to USP2 or USP21 contain a remarkably similar core functional epitope, or &quot;hot spot,&quot; consisting mainly of positions that are conserved as the wild type sequence, but also some positions that prefer mutant sequences."

reach
"In the USP2 and USP21 complex structure, the side chain of Ub Lys48 does not occupy the nearby pocket on USP (see XREF_FIG)."

reach
"We first screened a panel of DUBs and found that both USP2 and USP21 bound to endogenous SKP2, but only USP2 deubiquitylated and stabilized SKP2 protein."
USP2 affects USP1
| 3
| 3

sparser
"The two EcRs could interact with either USP1 or USP2 in vitro."

sparser
"Additionally, EMSAs analyzing the interaction between RosP and Usp1 and Usp2 promoter proteins demonstrate specific binding of RosP-His to the 340-bp promoter site of Usp1 and Usp2."

sparser
"The purified EcRs (EcRA and EcRB1) interacted with both USP1 and USP2 (Fig. 6)."
USP2 affects TNFSF10
| 3
USP2 inhibits TNFSF10. 3 / 3
| 3

reach
"The pharmacological inhibitor (ML364) and siRNA targeting USP2 enhanced TNF-related apoptosis-inducing ligand (TRAIL)-induced cancer cell death, but not normal cells."

reach
"These data suggested that survivin contributes to depletion of USP2-mediated TRAIL sensitization."

reach
"Inhibition of USP2 Enhances TRAIL-Mediated Cancer Cell Death through Downregulation of Survivin."
USP2 affects TNF
| 3
USP2 inhibits TNF. 3 / 3
| 3

reach
"Knockdown of USP2 or USP14 accelerated protein degradation of TNF-alpha, and abolished the effect of miR-124 on TNF-alpha protein stability."

reach
"Usp2 overexpression also suppressed the TNF-α–induced expression of atrogenes (Fig. 5F)."

reach
"Collectively, these results suggest USP2 prevents TNF-α– and DEX-induced myotube atrophy."
USP2 affects TGFBR1
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USP2 affects STAT1
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USP2 affects SNAI1
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3 |

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USP2 affects SLC2A4
| 3
USP2 increases the amount of SLC2A4. 3 / 3
| 3

reach
"Usp2 knockdown reduced the expression of PPAR-γ, GLUT4, and IRS1, and increased atrogene expression under DEX stimulation (Supplementary Fig. 4A)."

reach
"Knockdown of PPAR-γ reduced and Usp2 overexpression increased the levels of PPAR-γ, IRS1, and GLUT4 in C2C12 cells (Fig. 6A)."

reach
"In contrast, Usp2 overexpression induced the expression of PPAR-γ, GLUT4, and IRS1, and reduced the expression of atrogenes under DEX stimulation (Supplementary Fig. 4B)."
USP2 affects SLC22A2
| 3
USP2 activates SLC22A2. 3 / 3
| 3

reach
"Therefore, USP2 potentiates nuclear accumulation of Oct-1 and Oct-2 proteins."

reach
"USP2 Modulates Oct-1 and Oct-2 in LPS Stimulated HL-60 Cells."

reach
"Overexpression of USP2 did not cause modification of the K48 linked polyubiquitination of Oct-2 protein, suggesting that USP2 is unlikely to modulate Oct-2 digestion in a direct manner."
USP2 affects SCNN1A
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USP2 affects RAB1A
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3 |

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USP2 affects PCNA
3 |
3 |

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USP2 affects MYOD1
| 3
| 3

sparser
"Furthermore, TNF-α and DEX treatment inhibited the binding of MYOD1 to the promoter of USP2 in the C2C12 cells ( xref )."

sparser
"Two putative MYOD1 binding motif sequences (MBE) were predicted in the USP2 promoter region (JASPAR analysis), and ChIP-qPCR assays further demonstrated the binding of MYOD1 to the promoter of the USP2 ( xref )."

sparser
"Our study further showed that TNF-α–induced inflammation significantly reduced MYOD1 binding to the USP2 promoter ( xref )."
USP2 affects MALT1
3 |
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USP2 affects IRS1
| 3
USP2 increases the amount of IRS1. 3 / 3
| 3

reach
"In contrast, Usp2 overexpression induced the expression of PPAR-γ, GLUT4, and IRS1, and reduced the expression of atrogenes under DEX stimulation (Supplementary Fig. 4B)."

reach
"Knockdown of PPAR-γ reduced and Usp2 overexpression increased the levels of PPAR-γ, IRS1, and GLUT4 in C2C12 cells (Fig. 6A)."

reach
"Usp2 knockdown reduced the expression of PPAR-γ, GLUT4, and IRS1, and increased atrogene expression under DEX stimulation (Supplementary Fig. 4A)."
USP2 affects IL22
| 3
USP2 inhibits IL22. 3 / 3
| 3

reach
"Knockout or pharmacologic inhibition of USP2 promotes the proliferation of myeloid cells to activate IL-22 and IFNgamma production of T cells."

reach
"These data together suggest that inhibition of USP2 promotes the proliferation of macrophages and dendritic cells to activate IL-22- and IFNγ-producing T cells during DSS-induced colitis.We also examined whether targeting USP2 by ML364 inhibited the dysregulation of ECM and promoted tissue repair."

reach
"Taken together, these data suggest that USP2 restricts the proliferation of myeloid cells and thereby inhibits the activation and expansion of IL-22- and IFNγ-producing T cells in the lamina propria during DSS-induced colitis.2.6 USP2 in myeloid cells inhibits immune responses to bacterial infections in the gut."
USP2 affects HOOK2
2 1 |
2 1 |

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USP2 affects GTF2E1
| 3
USP2 increases the amount of GTF2E1.
| 2
USP2 increases the amount of GTF2E1. 2 / 2
| 2

reach
"As anticipated, USP2 overexpression promoted LDH release, Fe level, MDA and lipid ROS, while inhibited GPX4, GSH and SLC7A11 levels."

reach
"Among the ferroptosis markers, the results showed that USP2 overexpression promoted LDH release, Fe 2+ level, MDA and Lipid ROS, while inhibited GPX4 activity and GSH levels."
USP2 activates GTF2E1.
| 1
USP2 activates GTF2E1. 1 / 1
| 1

reach
"Moreover, USP2 overexpression promoted Fe (Fig. 3F) levels in cisplatin-resistant cells."
USP2 affects Flag
| 3
| 3

sparser
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."

sparser
"The association between Flag-USP2 and HA-EBF2 was detected in these cells ( xref I,J)."

sparser
"To explore the proteins that were potentially regulated by USP2, we then performed immunoprecipitation-based mass spectrometry (IP-MS) following ectopic expression of Flag-USP2 and immunoprecipitation."
USP2 affects FAS
| 2 1
| 2 1

reach
"In 2004, Graner and colleagues showed that USP2 binds to and stabilizes FAS [XREF_BIBR], a protein known to be upregulated in many cancers including prostate cancer [XREF_BIBR]."

reach
"In prostate cancer cells, USP2 interacts with and stabilizes FAS, which is often over-expressed in biologically aggressive human tumors."

sparser
"In 2004, Graner and colleagues showed that USP2 binds to and stabilizes FAS [ xref ], a protein known to be upregulated in many cancers including prostate cancer [ xref ]."
USP2 affects ETS1
| 1 2
| 1 2

sparser
"In our study, we carried out assays and validated that ETS1 could bind to USP2 promoter region and activate USP2 transcription."

reach
"Therefore, we further conducted mechanism experiment to prove the interaction between ETS1 and USP2."

sparser
"Therefore, we further conducted mechanism experiment to prove the interaction between ETS1 and USP2."
USP2 affects ERBB2
1 | 2
1 | 2

sparser
"Additionally, in the subsequent co-immunoprecipitation analysis to examine the association of USP2 with ErbB2, USP2 was readily pulled down by immunoprecipitated ErbB2 especially in cells treated with 17-AAG, suggesting stronger association between USP2 and ubiquitylated ErbB2 (Fig.  xref )."

sparser
"USP2 associates with internalized ErbB2 and prevents its lysosomal sorting and degradation via exerting deubiquitylase activity."

No evidence text available
USP2 affects Colitis
| 3
USP2 activates Colitis. 3 / 3
| 3

reach
"These data suggest that USP2 is upregulated in the inflamed colon tissues in DSS-treated mice and in UC patients.We next investigated the roles of USP2 in DSS-induced colitis in mice."

reach
"USP2 promotes experimental colitis and bacterial infections by inhibiting the proliferation of myeloid cells and remodeling the extracellular matrix network."

reach
"These data together suggest that knockout of USP2 increases myeloid cells in the lamina propria of mice after DSS treatment.We next sorted the RFP cells from the CD45 LPMCs of Lyz2-Cre;Ai14 and Lyz2-Cre;Usp2 Ai14 mice that were induced colitis and performed transcriptome sequencing assays (Fig. 2A)."
USP2 affects BMI1
| 1 1 1
USP2 activates BMI1. 3 / 3
| 1 1 1

sparser
"In addition, USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats-containing proteins (β-TrCP), a cellular E3 ubiquitin ligase ( xref )."

eidos
"In addition , USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats-containing proteins ( beta-TrCP ) , a cellular E3 ubiquitin ligase ( 104 ) ."

reach
"In addition, USP2 activates Bmi1 and EMT by stabilizing Twist through removing ubiquitylation mediated by beta-transducin repeats containing proteins (beta-TrCP), a cellular E3 ubiquitin ligase."
USP2 affects ATP
| 1 2
USP2 activates ATP. 3 / 3
| 1 2

eidos
"Since the OCR is mainly determined by OXPHOS , USP2 deficiency seems to inhibit ATP synthesis in the mitochondria of cultured myoblasts ."

reach
"Mechanistically, an ML364 treatment experiment suggests that USP2 inhibits the accumulation of ROS in VMH neurons, and thereby leads to continued ATP production in the mitochondria."

reach
"In myoblasts, USP2 knockout decreased the accumulation of intracellular ATP and oxygen consumption."
USP1 affects USP2
| 3
| 3

sparser
"The two EcRs could interact with either USP1 or USP2 in vitro."

sparser
"Additionally, EMSAs analyzing the interaction between RosP and Usp1 and Usp2 promoter proteins demonstrate specific binding of RosP-His to the 340-bp promoter site of Usp1 and Usp2."

sparser
"The purified EcRs (EcRA and EcRB1) interacted with both USP1 and USP2 (Fig. 6)."
TRIM27 affects USP2
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TGFBR1 affects USP2
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TFDP1 affects USP2
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TFDP1 decreases the amount of USP2. 3 / 3
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TCF3 affects USP2
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TCF3 decreases the amount of USP2. 3 / 3
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STAT1 affects USP2
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SNAI1 affects USP2
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SCNN1A affects USP2
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RAB1A affects USP2
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No evidence text available

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RA-9 affects USP2
| 3
RA-9 inhibits USP2. 3 / 3
| 3

reach
"RA-9, a chalcone derivative with a structure similar to b-AP15, was reported to inhibit proteasomal DUBs [XREF_BIBR] as well as UCHL1, UCHL3, USP2, USP5, and USP8 [XREF_BIBR]."

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."
PCNA affects USP2
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3 |

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Mice affects USP2
| 3
Mice inhibits USP2.
| 2
Mice inhibits USP2. 2 / 2
| 2

reach
"Consistently, both immunohistochemistry and immunoblotting analyses showed that UCP1 abundance was lower in Usp2 knockdown mice compared to controls, which was rebounded by overexpression of EBF2 (Figure 7E,F)."

reach
"Of note, overexpression of EBF2 in BAT-specific USP2 knockdown mice not only reduced lipid droplet accumulation but also restored BAT mass, indicating that the dual effects of USP2 in BAT are at least partially mediated through the regulation of EBF2."
Mice activates USP2.
| 1
Mice activates USP2. 1 / 1
| 1

reach
"In this study, the cardiac hypertrophy mouse model was established using Ang II induction and Ang II-induced mice were injected with AAV9-oe-USP2 to overexpress USP2."
MYH9 affects USP2
| 3
MYH9 activates USP2. 2 / 2
| 2

reach
"The present study revealed that MYH9 prevented β-catenin degradation and enhanced its nuclear activity by recruiting the deubiquitinase USP2."

reach
"In summary, our results indicated that ENKUR recruits FBXW7 to promote ubiquitination and degradation of MYH9 protein.2.4 MYH9 attenuates ENKUR-induced EMT signaling inhibition by recruiting deubiquitinase USP2 to stabilize beta-catenin protein."
MYH9 bound to CTNNB1 activates USP2. 1 / 1
| 1

reach
"In this process, MYH9 binds to β-catenin and increases its protein level by recruiting the deubiquitinase USP2 to form a complex."
MFN2 affects USP2
| 3
MFN2 decreases the amount of USP2. 3 / 3
| 3

reach
"The above data suggested that MFN2 downregulation attenuated the suppressive role of USP2 overexpression in hypertrophy of NMCMs.To probe into the impact of MFN2 in cardiac hypertrophy in vivo , Ang I[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The above data suggested that MFN2 downregulation attenuated the protective role of USP2 overexpression in mitochondrial functions of NMCMs.For the role of MFN2 in USP2-regulated hypertrophy of NMCMs,[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In vivo , MFN2 downregulation also reversed the protective role of USP2 overexpression against mitochondrial dysfunction and cardiac hypertrophy."
MALT1 affects USP2
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HOOK2 affects USP2
2 1 |
2 1 |

No evidence text available

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Flag affects USP2
| 3
| 3

sparser
"To further determine the interaction between USP2 and EBF2, we constructed Flag-USP2 and HA-EBF2 plasmids and transfected them individually or together into HEK293T cells."

sparser
"The association between Flag-USP2 and HA-EBF2 was detected in these cells ( xref I,J)."

sparser
"To explore the proteins that were potentially regulated by USP2, we then performed immunoprecipitation-based mass spectrometry (IP-MS) following ectopic expression of Flag-USP2 and immunoprecipitation."
FAS affects USP2
| 2 1
| 2 1

reach
"In 2004, Graner and colleagues showed that USP2 binds to and stabilizes FAS [XREF_BIBR], a protein known to be upregulated in many cancers including prostate cancer [XREF_BIBR]."

reach
"In prostate cancer cells, USP2 interacts with and stabilizes FAS, which is often over-expressed in biologically aggressive human tumors."

sparser
"In 2004, Graner and colleagues showed that USP2 binds to and stabilizes FAS [ xref ], a protein known to be upregulated in many cancers including prostate cancer [ xref ]."
ERBB2 affects USP2
1 | 2
1 | 2

sparser
"Additionally, in the subsequent co-immunoprecipitation analysis to examine the association of USP2 with ErbB2, USP2 was readily pulled down by immunoprecipitated ErbB2 especially in cells treated with 17-AAG, suggesting stronger association between USP2 and ubiquitylated ErbB2 (Fig.  xref )."

sparser
"USP2 associates with internalized ErbB2 and prevents its lysosomal sorting and degradation via exerting deubiquitylase activity."

No evidence text available
EGFR affects USP2
1 | 1
1 | 1

sparser
"For example, USP2 interacts with early EGFR transporters and performs deubiquitination modifications, which prevent EGFR from being transported to lysosomes for degradation."

No evidence text available
Dietary Fats affects USP2
3 |
Dietary Fats increases the amount of USP2. 3 / 3
3 |

No evidence text available

No evidence text available

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CLOCK affects USP2
| 3
CLOCK activates USP2. 3 / 3
| 3

reach
"This suggests a scenario in USP2 -/- mice in which increased turnover of BMAL1 coincides with increased CLOCK and BMAL1 driven gene expression in the USP2 -/- SCN."

reach
"However, the free running period (tau), a key intrinsic characteristic of the central circadian clock, was significantly increased in Usp2 KO mice (XREF_FIG; XREF_TABLE) (also reproduced in an additional group of mice, data not shown), suggesting the involvement of USP2 in the molecular clockwork."

reach
"The circadian rhythm of Usp2 is blunted in Clock mutant and Bmal1 KO mice, and the Usp2 promoter is activated by CLOCK and BMAL1, indicating that Usp2 is a direct target of these transcription factors."
Tioguanine affects USP2
| 1 1
| 1 1

sparser
"Previous studies indicated that 6-TG could non-competitively inhibit human USP2 ( xref ) thus plays a critical role in tumor cells survival."

reach
"Recently, disulfiram and 6-thioguanine were reported to synergistically inhibit USP2 and USP21 but with low affinity in the micromolar range."
Tetraphene affects USP2
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Tetraphene increases the amount of USP2. 2 / 2
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Succimer affects USP2
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Succimer decreases the amount of USP2. 2 / 2
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2 |
Pirinixic acid decreases the amount of USP2. 2 / 2
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Phenylmercury acetate decreases the amount of USP2. 2 / 2
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2 |
Phenobarbital increases the amount of USP2. 2 / 2
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Magnetite nanoparticle decreases the amount of USP2. 2 / 2
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No evidence text available

No evidence text available
HPTH 1-38 affects USP2
| 2
HPTH 1-38 increases the amount of USP2. 2 / 2
| 2

reach
"Parathyroid hormone (hPTH 1-38) stimulates the expression of UBP41, an ubiquitin specific protease, in bone."

reach
"Parathyroid hormone (hPTH 1-38) stimulates the expression of UBP41, an ubiquitin specific protease, in bone."
GAcrp affects USP2
| 2
GAcrp decreases the amount of USP2. 2 / 2
| 2

reach
"As depicted in Fig. 1C and 1D, gAcrp significantly decreased USP-2 protein levels in a dose- and time dependent manner."

reach
"Similarly, gAcrp decreased USP-2 protein levels in a time dependent manner in MCF-7 cells."
Fructose affects USP2
| 2
Fructose increases the amount of USP2. 2 / 2
| 2

reach
"High fructose intake upregulates hepatic USP2 expression."

reach
"Fructose increases lipid accumulation and USP2 expression in hepatocytes."
Folic acid affects USP2
2 |
Folic acid increases the amount of USP2. 2 / 2
2 |

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No evidence text available
Ezogabine affects USP2
| 2
Ezogabine increases the amount of USP2. 2 / 2
| 2

reach
"Indeed, we found that retigabine (8 mg/kg) significantly increased the protein levels of USP2 in hippocampal CA1 area of stressed rats."

reach
"On the other hand, the expression and coexpression of GluA1 and USP2 in hippocampal CA1 area were down-regulated after acute stress, and increased by retigabine treatment."
Cisplatin affects USP2
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Cisplatin decreases the amount of USP2. 1 / 2
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Chrysene affects USP2
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Chrysene increases the amount of USP2. 2 / 2
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No evidence text available
Cannabidiol affects USP2
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Cannabidiol decreases the amount of USP2. 2 / 2
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Benzo[b]fluoranthene increases the amount of USP2. 2 / 2
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VPS52 affects USP2
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VDAC2 affects USP2
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No evidence text available
| 2

reach
"Similarly, Similarly, USP2 may directly modulate BAT thermogenesis-related gene expression through DNA binding or transcriptional regulation [58]."

reach
"Collectively, these data suggest that USP2 in myeloid cells promotes colon inflammation after DSS treatment.2.3 Knockout of USP2 alters gene expression profile in myeloid cells after DSS treatment."
USP2 affects doxorubicin
| 2
| 2

reach
"In light of the vital role of USP2 in CSC regulation, we sought to determine whether USP2 inhibition increases TNBC sensitivity to doxorubicin and paclitaxel."

reach
"Moreover, USP2 inhibition by ML364 considerably enhanced the cytotoxicity of doxorubicin or paclitaxel in multiple TNBC cell models."
USP2 affects Wnt/β-catenin
| 2
USP2 activates Wnt/β-catenin. 2 / 2
| 2

reach
"Afterwards, we observed from TOP/FOP flash assay that USP2 silencing significantly inhibited the activity of Wnt/β-catenin signalling pathway (Figure 6G)."

reach
"Furthermore, the USP2-activated Wnt/β-catenin pathway by mediating deubiquitination of β-catenin protein contributes to the osteogenic differentiation of hBMSCs [37]."
USP2 affects VPS52
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USP2 affects VDAC2
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USP2 affects TRIM23
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USP2 affects TNFRSF1A
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USP2 affects TGFBR2
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USP2 affects TCP1
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USP2 affects TBK1
1 |
1 |

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USP2 affects TARDBP
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USP2 affects SLC7A11
| 2
USP2 decreases the amount of SLC7A11. 2 / 2
| 2

reach
"In our study, USP2 overexpression significantly suppressed SLC7A11 and GPX4 expression (Fig. 4A, B and C)."

reach
"The WB results revealed that USP2 overexpression inhibited GPX4, SLC7A11 and cytoplasm p53 protein expression, while promoted the nucleus p53 protein expression."
USP2 affects SLC22A1
| 2
USP2 activates SLC22A1. 2 / 2
| 2

reach
"USP2 Modulates Oct-1 and Oct-2 in LPS Stimulated HL-60 Cells."

reach
"Therefore, USP2 potentiates nuclear accumulation of Oct-1 and Oct-2 proteins."
USP2 affects SIRT2
| 2
| 2

sparser
"We found that the interaction between SIRT2 and USP2, as determined by both endogenous and exogenous co-immunoprecipitation (Co-IP), was present under TGFβ1 stimulation (Fig. xref )."

sparser
"To explore and confirm whether TGFβ1 influences the interaction between USP2 and SIRT2, we detected the binding of USP2 to SIRT2 under TGFβ1 stimulation (Fig. xref )."
USP2 affects SCNN1G
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No evidence text available
USP2 affects RUVBL1
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2 |

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USP2 affects PTH1R
| 1
USP2 inhibits PTH1R. 1 / 2
| 1

reach
"Furthermore, overexpression of USP2 in ROS cells rescued PTHR degradation induced by PTH (7-34) (XREF_FIG)."
USP2 affects PTH
| 2
USP2 activates PTH. 2 / 2
| 2

reach
"Overexpression of USP2 prevented PTH (7-34)-induced PTHR degradation."

reach
"Recently, research data have demonstrated that USP2 is necessary for PTH (1-34) to induce osteoblast proliferation [109]."
USP2 affects Obesity
| 2
USP2 activates Obesity. 2 / 2
| 2

reach
"Taken together, these results suggest that USP2 modulates thermogenic activation in brown adipocytes in a cell-autonomous manner.3.6 BAT-specific overexpression of USP2 promotes thermogenesis and improves HFD-induced obesity."

reach
"Conversely, overexpression of Usp2 in BAT conferred protection against high-fat diet-induced obesity and associated metabolic disorders."
USP2 affects NFKB2
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1 1 |

No evidence text available

No evidence text available
USP2 affects NEFL
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No evidence text available

No evidence text available
USP2 affects NDEL1
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No evidence text available

No evidence text available
USP2 affects NCOA4
| 2
| 2

sparser
"Hypoxia inhibits ferritinophagy-mediated ferroptosis in esophageal squamous cell carcinoma via the USP2-NCOA4 axis."

sparser
"Correction: Hypoxia inhibits ferritinophagy-mediated ferroptosis in esophageal squamous cell carcinoma via the USP2-NCOA4 axis."
USP2 affects MYCN
| 2
| 2

sparser
"Mycn bound to the USP2 promoter to activate its transcription."

sparser
"Collectively, this evidence suggests that the Mycn binds to the USP2 promoter for its transcription activation."
USP2 affects MTUS2
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No evidence text available

No evidence text available
USP2 affects ML364
| 2
ML364 binds USP2. 2 / 2
| 2

reach
"Direct binding of ML364 to USP2 was demonstrated using microscale thermophoresis."

reach
"ML364 directly binds to USP2 and selectively inhibits its enzymatic activity 31."
USP2 affects LZTS2
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No evidence text available

No evidence text available
USP2 affects LDL
| 2
USP2 inhibits LDL. 2 / 2
| 2

reach
"Since overexpression of Usp2 promotes the deubiquitination of IDOL, USP2 was considered to suppress LDL uptake via ubiquitin-dependent degradation of LDL [212]."

reach
"Transfection of a Usp2-expressing construct restored IDOL-repressed LDL uptake in cultured cells, whereas Usp2 knockdown reduced LDL uptake by ~50% in HepG2 hepatocytes [212]."
USP2 affects LDH
| 2
USP2 increases the amount of LDH. 2 / 2
| 2

reach
"As anticipated, USP2 overexpression promoted LDH release, Fe level, MDA and lipid ROS, while inhibited GPX4, GSH and SLC7A11 levels."

reach
"Among the ferroptosis markers, the results showed that USP2 overexpression promoted LDH release, Fe 2+ level, MDA and Lipid ROS, while inhibited GPX4 activity and GSH levels."
USP2 affects KRT40
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No evidence text available

No evidence text available
USP2 affects KIFC3
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No evidence text available

No evidence text available
USP2 affects KAZN
1 | 1
1 | 1

No evidence text available

sparser
"These results provide insights into the molecular mechanisms that KAZN may interact with USP2 to affect lung cancer pathogenesis and highlights potential targets for further investigation."
| 2

reach
"Moreover, macrophage USP2 represses adipose tissue inflammation and subsequent insulin resistance [79]."

reach
"The global deletion of USP2 aggravated diabetes- and DEX-induced muscle atrophy and insulin resistance in mice."
USP2 affects Infections
| 2
| 2

reach
"USP2 inhibition prevents infection with ACE2-dependent coronaviruses in vitro and is protective against SARS-CoV-2 in mice."

reach
"These results demonstrate that the targeted inhibition of USP2 with ML364 effectively prevents the infection of ACE2-dependent SARS-CoV-2, irrespective of S protein mutations."
USP2 affects IMD
| 2
USP2 inhibits IMD. 2 / 2
| 2

reach
"Interestingly, the ubiquitin carboxyl-terminal hydrolase Usp2 , which in Drosophila suppresses the activation of the Toll pathway and prevents the activation of the IMD pathway ( Engel et al., 2014 ),[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The D. melanogaster ubiquitin-specific protease 36 (USP36) and USP2 negatively regulate the IMD pathway after infection by Gram-negative pathogens (42, 43)."
| 2
| 2

reach
"USP22 in pancreatic beta-cells, USP2 in adipose tissue macrophages, USP9X, 20, and 33 in myocytes, USP4, 7, 10, and 18 in hepatocytes, and USP2 in hypothalamus improve hyperglycemia, whereas USP19 in adipocytes, USP21 in myocytes, and USP2, 14, and 20 in hepatocytes promote hyperglycemia."

reach
"Given that HSD1 converts circulating inactive cortisone to active cortisol [107], hepatic USP2 causes hyperglycemia by activating glucocorticoid signaling.Sustained low-grade endoplasmic reticulum (ER) stress induced by low doses of tunicamycin (200 μg/kg) treatment for two weeks increased hepatic gluconeogenesis and resulted in hyperglycemia [103]."
USP2 affects HSPA1A
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No evidence text available

No evidence text available
USP2 affects HOMER3
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No evidence text available

No evidence text available
| 1 1

eidos
"Given that USP2-4 positively regulates the expression of several glucocorticoid-regulated genes in the liver , USP2 appears to exacerbate glucose intolerance by stimulating glucocorticoid signaling [ 11 ] ."

reach
"Several USPs, including USP2, USP10, USP14, USP15, USP18, and USP22, have been associated with obesity and related metabolic disorders.In diet-induced obese mice, it is reported that USP2 can increase the production of hepatic glucose and promote glucose intolerance (143)."
USP2 affects Glioblastoma
| 2
| 2

reach
"Loss of deubiquitylase USP2 triggers development of glioblastoma via TGF-β signaling."

reach
"Conversely, USP2 inhibits glioblastoma development by suppressing the TGF-B signaling pathway through the deubiquitination of SMAD7 [19]."
USP2 affects GPX4
| 2
USP2 decreases the amount of GPX4. 2 / 2
| 2

reach
"In our study, USP2 overexpression significantly suppressed SLC7A11 and GPX4 expression (Fig. 4A, B and C)."

reach
"The WB results revealed that USP2 overexpression inhibited GPX4, SLC7A11 and cytoplasm p53 protein expression, while promoted the nucleus p53 protein expression."
USP2 affects GOLGA2
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No evidence text available

No evidence text available
USP2 affects GATA2
| 2
USP2 activates GATA2. 2 / 2
| 2

sparser
"Although patient SJINF066 was an infant (9-month-old) and SJBALL021549_D1 was a 9-year-old child, the expression levels of GATA2 in these two cases were comparable, indicating that trans-activation of GATA2 by KMT2A-USP2 is not influenced by age but rather by the function of this fusion."

sparser
"Further checking confirmed that these patients were the only two carrying this fusion in the cohort analyzed ( xref ), indicating that GATA2 could be trans-activated by KMT2A-USP2."
USP2 affects EZH2
| 2
| 2

sparser
"Additionally, as a deubiquitinase, USP2 interacted with EZH2 to downregulate SOX1 expression, thereby promoting BLCA tumorigenesis."

sparser
"Co-IP assay demonstrated that endogenous USP2 and EZH2 interacted with each other in BLCA cells ( xref )."
USP2 affects ERK
| 2
USP2 activates ERK. 2 / 2
| 2

reach
"How exactly USP2 modulates this ERK1/2‐Per1 pathway should be a focus of subsequent research."

reach
"Thus, the blunting of photic induction of Per1 in the absence of USP2 is particularly pronounced in the retinorecipient, ventral part of the SCN.3.11 USP2 Modulates the Photic Activation of the ERK/MAPK Pathway."
USP2 affects E6
1 1 |
E6 binds USP2. 2 / 2
1 1 |

No evidence text available

No evidence text available
USP2 affects DTX3
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No evidence text available

No evidence text available
USP2 affects DNAJA1
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No evidence text available

No evidence text available

reach
"We next investigated whether USP2 activates transcription of the cytokine genes by modifying chromatin accessibility after TLR stimulation."

sparser
"We next investigated whether USP2 activates transcription of the cytokine genes by modifying chromatin accessibility after TLR stimulation."

reach
"Overexpression of USP2 restores the DSB response in RECQL4-defective cells."

reach
"Taken together, these results suggest that overexpression of USP2 prevented premature disassembly of the MRN complex from DSB sites and restored the DSB response in RECQL4-defective cells."
USP2 affects D1
| 2
USP2 binds Cyclin and D1. 2 / 2
| 2

reach
"USP2 binds to Cyclin D1 and Mdm2 and induces their deubiquitination and stabilization, and its silencing causes downregulation of Cyclin D1 and Mdm2, leading to inhibiting cell proliferation."

reach
"By using co-immunoprecipitation approaches, we tried to determine whether elevation of O GlcNAcylation favors the binding of USP2 to cyclin D1, as described for BAP1 and PGC1alpha."
USP2 affects CFLAR
| 1 1
USP2 inhibits CFLAR. 2 / 2
| 1 1

reach
"USP2 knockdown via siRNA inhibits actinomycin and TNF-α-induced apoptosis via c-FLIP upregulation in hepatocytes [27]."

eidos
"Compared to its parental cells , the sorafenib-resistant clones express lower levels of USP2 , which presumably caused the aforementioned increase in cFLIP levels and decrease in ITCH levels [ 13,81 ] ."
USP2 affects CDKN1A
| 2
USP2 decreases the amount of CDKN1A. 2 / 2
| 2

reach
"As shown in Figure XREF_FIG, USP2 knockdown significantly increased p21 mRNA expression compared with MyLa2000 cells with control siRNA (p < 0.05)."

reach
"Given that USP2 knockdown reduced the levels of both SKP2 and SKP2 substrates, p21 and p27 (Fig. 6B), this growth inhibitory effect appears to be mediated by other substrates of USP2 in a manner largely independent of SKP2."
USP2 affects CCDC85B
1 1 |
1 1 |

No evidence text available

No evidence text available
USP2 affects CCDC136
2 |
2 |

No evidence text available

No evidence text available
USP2 affects CARD11
2 |
2 |

No evidence text available

No evidence text available
| 1 1

reach
"USP2 promotes experimental colitis and bacterial infections by inhibiting the proliferation of myeloid cells and remodeling the extracellular matrix network."

eidos
"USP2 promotes experimental colitis and bacterial infections by inhibiting the proliferation of myeloid cells and remodeling the extracellular matrix network ."
USP2 affects BRCA1
2 |
2 |

No evidence text available

No evidence text available
USP2 affects BEND5
1 1 |
1 1 |

No evidence text available

No evidence text available
USP2 affects BEGAIN
1 1 |
1 1 |

No evidence text available

No evidence text available
USP2 affects ASAH1
| 1
USP2 activates ASAH1. 1 / 2
| 1

reach
"USP2 siRNA decreased ACDase protein, whereas USP2 overexpression increased ACDase protein of LNCaP cells."
USP2 affects APP
2 |
2 |

No evidence text available

No evidence text available
TRIM23 affects USP2
2 |
2 |

No evidence text available

No evidence text available
TNFRSF1A affects USP2
2 |
2 |

No evidence text available

No evidence text available
TCP1 affects USP2
2 |
2 |

No evidence text available

No evidence text available
TBK1 affects USP2
1 |
1 |

No evidence text available
TARDBP affects USP2
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2 |

No evidence text available

No evidence text available
Smoke affects USP2
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Smoke increases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
SJB3-019A affects USP2
| 1 1
SJB3-019A inhibits USP2. 2 / 2
| 1 1

eidos
"Interestingly , USP2 and USP8 , which were both inhibited by SJB3-019A in a previous report , were also inhibited by 42 % and 68 % , respectively , in our evaluation ."
| PMC

reach
"Interestingly, USP2 and USP8, which were both inhibited by SJB3-019A in a previous report, were also inhibited by 42% and 68%, respectively, in our evaluation."
| PMC
SIRT2 affects USP2
| 2
| 2

sparser
"We found that the interaction between SIRT2 and USP2, as determined by both endogenous and exogenous co-immunoprecipitation (Co-IP), was present under TGFβ1 stimulation (Fig. xref )."

sparser
"To explore and confirm whether TGFβ1 influences the interaction between USP2 and SIRT2, we detected the binding of USP2 to SIRT2 under TGFβ1 stimulation (Fig. xref )."
SCNN1G affects USP2
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No evidence text available

No evidence text available

sparser
"While USP2 was readily inhibited by ROS, the activity of AMSH was unaffected, even in the presence of STAM, the binding partner and catalytic activator of AMSH ( xref )."

reach
"While USP2 was readily inhibited by ROS, the activity of AMSH was unaffected, even in the presence of STAM, the binding partner and catalytic activator of AMSH (XREF_FIG)."
RUVBL1 affects USP2
2 |
2 |

No evidence text available

No evidence text available
RAG2 affects USP2
| 2
RAG2 inhibits USP2. 2 / 2
| 2

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."
Proteasome affects USP2
| 2

reach
"USP2 and Imd synergistically interact with the proteasome."

reach
"Taken together, our data suggest that USP2 and Imd synergistically bind to the proteasome and that USP2 contributes to Imd degradation at the proteasome level."
2 |
Nanotubes, Carbon decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
NFKB2 affects USP2
1 1 |
1 1 |

No evidence text available

No evidence text available
NFIL3 affects USP2
2 |
NFIL3 decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
NEFL affects USP2
2 |
2 |

No evidence text available

No evidence text available
NDEL1 affects USP2
2 |
2 |

No evidence text available

No evidence text available
NCOA4 affects USP2
| 2
| 2

sparser
"Hypoxia inhibits ferritinophagy-mediated ferroptosis in esophageal squamous cell carcinoma via the USP2-NCOA4 axis."

sparser
"Correction: Hypoxia inhibits ferritinophagy-mediated ferroptosis in esophageal squamous cell carcinoma via the USP2-NCOA4 axis."
MYLIP affects USP2
1 |
1 |

No evidence text available
MTUS2 affects USP2
2 |
2 |

No evidence text available

No evidence text available
MECOM affects USP2
2 |
MECOM decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
LZTS2 affects USP2
2 |
2 |

No evidence text available

No evidence text available
LDLR affects USP2
1 | 1
1 | 1

No evidence text available

reach
"The authors attempt to answer this question by introducing a tripartite complex model, in which USP2 interacts with LDLR at the plasma membrane in an IDOL dependent manner to deubiquitylate and stabilize both LDLR and IDOL."
KRT40 affects USP2
2 |
2 |

No evidence text available

No evidence text available
KIFC3 affects USP2
2 |
2 |

No evidence text available

No evidence text available
KAZN affects USP2
1 | 1
1 | 1

No evidence text available

sparser
"These results provide insights into the molecular mechanisms that KAZN may interact with USP2 to affect lung cancer pathogenesis and highlights potential targets for further investigation."
HSPA1A affects USP2
2 |
2 |

No evidence text available

No evidence text available
HOMER3 affects USP2
2 |
2 |

No evidence text available

No evidence text available
GOLGA2 affects USP2
2 |
2 |

No evidence text available

No evidence text available
EZH2 affects USP2
| 2
| 2

sparser
"Additionally, as a deubiquitinase, USP2 interacted with EZH2 to downregulate SOX1 expression, thereby promoting BLCA tumorigenesis."

sparser
"Co-IP assay demonstrated that endogenous USP2 and EZH2 interacted with each other in BLCA cells ( xref )."
E6 affects USP2
1 1 |
E6 binds USP2. 2 / 2
1 1 |

No evidence text available

No evidence text available
E2F1 affects USP2
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E2F1 decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
DTX3 affects USP2
2 |
2 |

No evidence text available

No evidence text available
DNAJA1 affects USP2
2 |
2 |

No evidence text available

No evidence text available
DIM affects USP2
| 2
DIM inhibits USP2. 2 / 2
| 2

reach
"Additionally, DIM suppressed the G1 phase (early stage of adipogenesis) and induced post-translational degradation of cyclin D1 by inhibiting ubiquitin-specific peptidase 2 (USP2) activities."

reach
"Additionally, it has been shown that DIM suppresses adipogenesis by inhibiting the ubiquitin-specific peptidase 2, but it also can inhibits high-fat diet-induced obesity in high-fat diet-fed obese mice."
D1 affects USP2
| 2
USP2 binds Cyclin and D1. 2 / 2
| 2

reach
"USP2 binds to Cyclin D1 and Mdm2 and induces their deubiquitination and stabilization, and its silencing causes downregulation of Cyclin D1 and Mdm2, leading to inhibiting cell proliferation."

reach
"By using co-immunoprecipitation approaches, we tried to determine whether elevation of O GlcNAcylation favors the binding of USP2 to cyclin D1, as described for BAP1 and PGC1alpha."
CEBPA affects USP2
| 2
CEBPA activates USP2. 2 / 2
| 2

reach
"C/EBPα is a potent transcriptional factor of 11β-HSD1 in the liver, and activation of the USP2/ C/EBPα/ 11β-HSD1 signaling may account for fructose effects on MASLD (Fig. 6A)."

reach
"We detected the potential downstream molecules of USP2 in primary hepatocytes treated with fructose, the expression of C/EBPα (C/ebpα) and 11β-HSD1 (Hsd11b1) was significantly increased both at mRNA (Fig. 6B&C) and protein level (Fig. 6D)."
CCDC85B affects USP2
1 1 |
1 1 |

No evidence text available

No evidence text available
CCDC136 affects USP2
2 |
2 |

No evidence text available

No evidence text available
CARD11 affects USP2
2 |
2 |

No evidence text available

No evidence text available
BRCA1 affects USP2
2 |
2 |

No evidence text available

No evidence text available
BEND5 affects USP2
1 1 |
1 1 |

No evidence text available

No evidence text available
BEGAIN affects USP2
1 1 |
1 1 |

No evidence text available

No evidence text available
BAAT affects USP2
| 2
BAAT inhibits USP2. 2 / 2
| 2

reach
"Intriguingly, we noticed that there were 16 proteins that were completely absent in the Usp2 knockdown BAT (Figure 6D)."

reach
"Among these proteins, EBF2 has been demonstrated as the pivotal regulatory role in the development of brown adipose and adaptive thermogenesis [32], which piqued our interest.Indeed, RT-qPCR and western blot analysis of EBF2 showed that the protein levels of EBF2 were significantly decreased in the Usp2 knockdown BAT compared to controls, with comparable mRNA levels of Ebf2 (Figure 6E,F)."
ATF2 affects USP2
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ATF2 decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
APP affects USP2
2 |
2 |

No evidence text available

No evidence text available
AM146 affects USP2
| 2
AM146 inhibits USP2. 2 / 2
| 2

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."
2 |
17beta-estradiol decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
2 |
1,2-dichloroethane decreases the amount of USP2. 2 / 2
2 |

No evidence text available

No evidence text available
2 |

No evidence text available

No evidence text available
1 |
Trovafloxacin decreases the amount of USP2. 1 / 1
1 |

No evidence text available
Streptozocin affects USP2
1 |
Streptozocin increases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |
SandostatinLAR decreases the amount of USP2. 1 / 1
1 |

No evidence text available
Resveratrol affects USP2
1 |
Resveratrol increases the amount of USP2. 1 / 1
1 |

No evidence text available
Perfluorooctanoic acid decreases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |

No evidence text available
1 |
Nickel sulfate decreases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |

No evidence text available
| 1

eidos
"LPS suppressed the expression of USP2 in macrophages ( human myeloid HL-60 cell line ) [ 21 ] ."
Isoprenaline affects USP2
1 |
Isoprenaline decreases the amount of USP2. 1 / 1
1 |

No evidence text available
| 1

reach
"Interestingly, preincubation of iodoacetic acid (IAA), a cysteine alkylating agent, with USP2 completely inhibited the binding of biotin-GA to USP2 (Supporting Information Fig. S8), suggesting that GA may covalently bind to the cysteine(s) of USP2, which is consistent with previous reports that GA can covalently bind to its targets through cysteine residues56."
Inulin affects USP2
1 |
Inulin increases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-493-5p decreases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4667-5p decreases the amount of USP2. 1 / 1
1 |

No evidence text available
1 |
Graphene oxide increases the amount of USP2. 1 / 1
1 |

No evidence text available
Doxorubicin affects USP2
1 |
Doxorubicin decreases the amount of USP2. 1 / 1
1 |

No evidence text available
Dicrotophos affects USP2
1 |
Dicrotophos increases the amount of USP2. 1 / 1
1 |

No evidence text available
Decabromodiphenyl ether increases the amount of USP2. 1 / 1
1 |

No evidence text available
Azoxystrobin affects USP2
1 |
Azoxystrobin decreases the amount of USP2. 1 / 1
1 |

No evidence text available
Antithrombin affects USP2
| 1
Antithrombin activates USP2. 1 / 1
| 1

reach
"Given that ML364 inhibits USP2, we next investigated the role of USP2 in antithrombin turnover."
Amiodarone affects USP2
1 |
Amiodarone increases the amount of USP2. 1 / 1
1 |

No evidence text available
All-trans-retinoic acid decreases the amount of USP2. 1 / 1
1 |

No evidence text available
YAP1 affects USP2
| 1
| 1

sparser
"Interestingly, we identified three rare gene fusions (USP2YAP1, YAP1–FAM118B and KIRREL3–SIK3) in a case of pilocytic astrocytoma that have not been previously reported."
VPS28 affects USP2
1 |
1 |

No evidence text available
Usp2 affects USP2
1 |
1 |

No evidence text available
USP8 affects USP2
| 1
USP2 binds USP8 and KCNN4. 1 / 1
| 1

sparser
"Channel-DUB interactions were verified after 90 min as there were notable interactions of KCa3.1 with USP2 and USP8 and a weaker association with AMSH (associated molecule with the SH3 domain of STAM; Figure xref )."
USP2 affects pro-degradation HSP90 ErbB2 respective therapeutic windows
| 1
USP2 activates pro-degradation HSP90 ErbB2 respective therapeutic windows. 1 / 1
| 1

eidos
"Through promoting ErbB2 ubiquitylation , USP2 inhibitor is expected to augment the pro-degradation effects of HSP90 inhibitors toward ErbB2 within their respective therapeutic windows , thus eliciting potent anti-tumor effects with tolerable adverse side effects , which warrants further investigations and clinical evaluations ."
USP2 affects polyubiquitin chain
| 1
USP2 activates polyubiquitin chain. 1 / 1
| 1

reach
"It is possible that USP2 deficiency caused aberrant digestion of the polyubiquitin chain, resulting in remarkably reduced Oct-2 in the nucleus."
| 1
USP2 leads to the deubiquitination of lipopolysaccharide. 1 / 1
| 1

reach
"Otten and colleagues did, however, show that the promiscuous deubiquitinase USP2 can reverse LPS ubiquitination in vitro ( Otten et al., 2021 )."

reach
"A decrease in USP2 expression reduces the keratinocyte proliferation rate and increases the keratinocyte differentiation, demonstrated by a rise in the mRNA expression level of differentiation markers (keratin 1410 and involucrin)."

eidos
"In addition , USP2 inhibition suppressed CFs proliferation , collagen synthesis and cell cycle progression ."

reach
"mi125b-related FGFR2 and USP2 pathways may contribute to only small arms for miRNA125b underlying cellular metabolism, as shown in Figure 2."
USP2 affects alpha-induced apoptosis
| 1
USP2 activates alpha-induced apoptosis. 1 / 1
| 1

eidos
"In addition to cancer cells , USP2 has been also postulated to modulate tumor necrosis factor ( TNF ) alpha-induced apoptosis in hepatocytes ."
USP2 affects acute respiratory syndrome coronavirus papain-like protease
| 1
USP2 activates acute respiratory syndrome coronavirus papain-like protease. 1 / 1
| 1

eidos
"Meanwhile , drug specificity should be taken into consideration , because many chemical inhibitors have unexpected molecular targets [ 159 ] ; for example , a recent paper reported that a USP2 blocker also inhibits severe acute respiratory syndrome coronavirus 2 papain-like protease [ 160 ] ."
USP2 affects abundance Na H exchange regulatory cofactor
| 1
USP2 inhibits abundance Na H exchange regulatory cofactor. 1 / 1
| 1

eidos
"Proteomics data from the intestine revealed that the lack of USP2 strongly increased the abundance of Na + / H + exchange regulatory cofactor 4 ( NHERF4 ) [ 90 ] , which is a known regulator of a calcium ion channel [ 91 ] ."
USP2 affects YAP1
| 1
| 1

sparser
"Interestingly, we identified three rare gene fusions (USP2YAP1, YAP1–FAM118B and KIRREL3–SIK3) in a case of pilocytic astrocytoma that have not been previously reported."
USP2 affects VPS28
1 |
1 |

No evidence text available
USP2 affects Usp2
1 |
1 |

No evidence text available
USP2 affects Ub chain
| 1
USP2 activates Ub chain. 1 / 1
| 1

reach
"No significant inhibitory effect of SJB was noted in USP7 or USP2 mediated Ub chain disassembly (XREF_SUPPLEMENTARY)."
USP2 affects USP8
| 1
USP2 binds USP8 and KCNN4. 1 / 1
| 1

sparser
"Channel-DUB interactions were verified after 90 min as there were notable interactions of KCa3.1 with USP2 and USP8 and a weaker association with AMSH (associated molecule with the SH3 domain of STAM; Figure xref )."
USP2 affects USP2 activation
| 1
USP2 inhibits USP2 activation. 1 / 1
| 1

eidos
"Because USP2 gene was transcriptionally activated in splenic B-lymphocytes in response to stimulation with IL-4 , the role of USP2 in lymphocyte activation may not be restricted to T-lymphocytes , but may also encompass B-lymphocytes [ 147 ] ."
USP2 affects TRIP6
1 |
1 |

No evidence text available
USP2 affects TRIM54
1 |
1 |

No evidence text available
USP2 affects TNF receptor
| 1
USP2 activates TNF receptor. 1 / 1
| 1

reach
"We have previously shown that the 41 kDa USP2 isoform induces apoptosis [3] and that the 69 kDa USP2a isoform mediates the apoptosis signal of the TNF receptor [5]."
USP2 affects TLE5
1 |
1 |

No evidence text available
USP2 affects TEKT4
1 |
1 |

No evidence text available
USP2 affects SRRT
1 |
1 |

No evidence text available
USP2 affects SORBS3
1 |
1 |

No evidence text available
USP2 affects SMAD2
1 |
1 |

No evidence text available
USP2 affects SKA3
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No evidence text available
USP2 affects SH3BP5
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"SAB can directly bind to ubiquitin-carboxy-terminal hydrolase 2 (USP2) in colon cancer cell RKO, inhibit its deubiquitination activity, enhance the killing activity of T cells in tumor cells, and finally play an anti-tumor role [46]."
USP2 affects SCN.3 Results
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USP2 increases the amount of SCN.3 Results. 1 / 1
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"We found that USP2 modulates photic entrainment and light‐dependent signaling pathways to dusk light cues at the level of the SCN.3 Results."
USP2 affects RUXN2
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USP2 increases the amount of RUXN2. 1 / 1
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"Inhibition of USP2 reduced the levels of RUXN2 and OPN proteins in hBMSCs (Fig. 3H) and hindered their proliferation (Fig. 3I) under the induction conditions."
USP2 affects RUNDC3A
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No evidence text available
USP2 affects RNF126
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No evidence text available
USP2 affects RNF125
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No evidence text available
USP2 affects RAD18
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No evidence text available
USP2 affects RAC
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EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
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sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
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"However, when we repeated the PABPC1 IP with the addition of the nonspecific deubiquitinating enzyme USP2, which was sufficient to eliminate poly-ubiquitin, we saw no change in the PABPC1 interactome (fig."
USP2 affects PTEN
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No evidence text available
USP2 affects PRKAA1
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No evidence text available
USP2 affects PPARGC1A
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sparser
"Further experiment showed that there was an interaction between USP2 and the transcription coactivator—PGC-1α which is related with various liver fasting reactions."
USP2 affects POU5F1
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sparser
"In contrast to negligible differences in NF- κ B binding activity between USP2 KD and control cells, we found a slight increase in Oct binding activity in USP2-deficient conditions."
USP2 affects PDCD1
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USP2 activates PDCD1. 1 / 1
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"These data demonstrate that CD8 + T cells, but not CD4 + T cells, are the primary effector cells underlying the combination treatment in these tumor models.Finally, to ascertain whether USP2 inhibition potentiates the efficacy of PD-1/PD-L1 blockade by enhancing p53 function in a tumor cell autonomous manner, we also examined the effects of combination therapy in Balb/c mice bearing p53-null EMT6 tumors."

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"Since our study is the first to uncover USP2 's novel function in CSCs, future studies will be needed to delineate the contribution of cyclin D1 to USP2 mediated CSC expansion in TNBC."
USP2 affects NF-kappaB basal
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USP2 inhibits NF-kappaB basal. 1 / 1
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eidos
"The ubiquitin #specific proteases system 2 ( USP2 ) stabilizes the ubiquitin-E3-ligase ITCH and lowers NF-kappaB basal activity , which leads to reduced c-FLIP mRNA production ; proteasomal degradation of c - FLIP isoforms is also elevated by its negative regulator proteasome ITCH [ 31 ] ."
USP2 affects Myoblasts
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"It can , therefore , be concluded that depletion of USP2 decreases proliferation of myoblasts ."
USP2 affects MPZ
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USP2 inhibits MPZ. 1 / 1
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"USP2 attenuated MPP + -triggered SK-N-SH cell injury through stabilizing FOXC1, providing a promising therapeutic target for PD treatment."
USP2 affects MID1IP1
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No evidence text available
USP2 affects MAVS
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No evidence text available
USP2 affects MAPK
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USP2 activates MAPK. 1 / 1
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"Thus, the blunting of photic induction of Per1 in the absence of USP2 is particularly pronounced in the retinorecipient, ventral part of the SCN.3.11 USP2 Modulates the Photic Activation of the ERK/MAPK Pathway."
USP2 affects LNX2
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No evidence text available
USP2 affects LMNA
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No evidence text available
USP2 affects KCNN4
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USP2 binds USP8 and KCNN4. 1 / 1
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sparser
"Channel-DUB interactions were verified after 90 min as there were notable interactions of KCa3.1 with USP2 and USP8 and a weaker association with AMSH (associated molecule with the SH3 domain of STAM; Figure xref )."
USP2 affects K63-linked polyubiquitination TRAF2
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USP2 inhibits K63-linked polyubiquitination TRAF2. 1 / 1
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"On the contrary , USP2 knockdown augmented the K63-linked polyubiquitination of both RIP1 and TRAF2 [ 77 ] ."
USP2 affects INSRbeta-GFP
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USP2 activates INSRbeta-GFP. 1 / 1
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"The INSRbeta or INSRbeta-GFP immunoprecipitates treated with USP2 enzyme were analysed for INSRbeta or INSRbeta-GFP."
USP2 affects IL32
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USP2 deubiquitinates IL32. 1 / 1
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"USP2 deubiquitinated and stabilized tAIF, thus promoting AIF mediated cell death."
USP2 affects IKZF3
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No evidence text available
USP2 affects IKBKG
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No evidence text available
USP2 affects IHO1
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"In addition, macrophage Usp2 ablation led to a decrease in the sperm population exhibiting high intracellular pH, calcium influx, and mitochondrial membrane potential."
USP2 affects HW/BW
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USP2 inhibits HW/BW. 1 / 1
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"Accordingly, our results revealed that USP2 overexpression improved cardiac functions, decreased the ratios of HW/BW and HW/TL, and reduced the degree of cardiac hypertrophy and fibrosis along with di[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects HSD1 mRNA
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USP2 activates HSD1 mRNA. 1 / 1
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"Hepatic overexpression of USP2 increases HSD1 mRNA and protein levels, leading to the induction of gluconeogenic genes as well as several other glucocorticoid targets, whereas RNAi knockdown of USP2 significantly lowers HSD1 gene expression in the liver."
USP2 affects GRIA
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USP2 leads to the deubiquitination of GRIA. 1 / 1
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"Given the same type of deubiquitinating enzyme (USP46) is able to deubiquitinate AMPA receptors, and knockdown of USP46 elevated AMPA receptor ubiquitination and reduced AMPA receptor expression in hi[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects GOLGA6L9
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USP2 affects Fibrosis
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USP2 activates Fibrosis. 1 / 1
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"Accordingly, our results revealed that USP2 overexpression improved cardiac functions, decreased the ratios of HW/BW and HW/TL, and reduced the degree of cardiac hypertrophy and fibrosis along with di[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects EcR-A
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EcR-A binds USP2. 1 / 1
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"Proliferation and differentiation processes of midgut stem cells are carried out via EcR-A and USP-2 heterodimer complex."
USP2 affects EEF1A2
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EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
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"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
USP2 affects E3_Ub_ligase
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USP2 deubiquitinates E3_Ub_ligase. 1 / 1
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"USP2 deubiquitinates the E3 ligase MDM2 and prevents its degradation [23] , and thus, we explored the effects of UbVs on MDM2 ubiquitination."
USP2 affects DUSP21
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No evidence text available
USP2 affects DSCR4
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No evidence text available
USP2 affects DISC1
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No evidence text available
USP2 affects Chemokine
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USP2 increases the amount of Chemokine. 1 / 1
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"These data together suggest that knockout of USP2 in myeloid cells inhibits the NF-κB signaling pathway and the expression of pro-inflammatory cytokines and chemokines and promotes the expression of cell cycle-related genes in the myeloid cells of lamina propria from mice treated with DSS.2.4 USP2 inhibits the proliferation of macrophages and dendritic cells in the lamina propria during DSS-induced colitis."
USP2 affects CUE5
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"The increased binding of Cue5 to inactivated proteasomes, and its reduced binding upon ubiquitin removal with USP2, is reminiscent of that seen with Arabidopsis RPN10, where its proteasome occupancy s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2 affects CRY2
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USP2 deubiquitinates CRY2. 1 / 1
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"USP2 is essential to deubiquitinating PER1, CRY1 and CRY2 in vivo [54–57]."
USP2 affects CRBN
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"Mechanistically, we find that USP2 directly interacts with CRBN, leading to the deubiquitination and stabilization of CRBN in senescent liver cancer cells."
USP2 affects CEP70
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No evidence text available
USP2 affects CCR8
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No evidence text available
USP2 affects CCDC57
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No evidence text available
USP2 affects CADPS
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No evidence text available
USP2 affects CACNA2D2
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No evidence text available
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No evidence text available
USP2 affects Aurora-A
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USP2 binds Aurora-A. 1 / 1
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"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."
USP2 affects Ang II-induced
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USP2 activates Ang II-induced. 1 / 1
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"To determine whether USP2 inhibition may reverse the effects of Ang II-induced proliferation , CFs were administered various ML364 amounts for distinct times , followed by the MTS assay ."
USP2 affects ACTN2
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No evidence text available
USP2 affects ABI2
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No evidence text available
TRIP6 affects USP2
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No evidence text available
TRIM54 affects USP2
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No evidence text available
TLE5 affects USP2
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No evidence text available
TEKT4 affects USP2
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No evidence text available
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Sleep Deprivation decreases the amount of USP2. 1 / 1
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"SD also significantly decreased expression levels of Rbm3 and Hnrpdl (mRNA metabolism and trafficking (Kawamura et al. 2002; Smart et al. 2007)) and Usp2 (deubiquitination and control of circadian rhythms (Metzig et al. 2011; Scoma et al. 2011; Yang et al. 2012)) (Fig. 3B; two-way ANOVA for each, P < 0.05)."
SRRT affects USP2
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No evidence text available
SORBS3 affects USP2
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No evidence text available
SMAD2 affects USP2
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No evidence text available
SKA3 affects USP2
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No evidence text available
SH3BP5 affects USP2
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"SAB can directly bind to ubiquitin-carboxy-terminal hydrolase 2 (USP2) in colon cancer cell RKO, inhibit its deubiquitination activity, enhance the killing activity of T cells in tumor cells, and finally play an anti-tumor role [46]."
S-nitrosoglutathione increases the amount of USP2. 1 / 1
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No evidence text available
RUNDC3A affects USP2
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No evidence text available
RNF126 affects USP2
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No evidence text available
RNF125 affects USP2
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No evidence text available
RAD18 affects USP2
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No evidence text available
RAC affects USP2
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EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
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sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
PTEN affects USP2
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No evidence text available
PRKAA1 affects USP2
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No evidence text available
PPARGC1A affects USP2
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sparser
"Further experiment showed that there was an interaction between USP2 and the transcription coactivator—PGC-1α which is related with various liver fasting reactions."
POU5F1 affects USP2
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"In contrast to negligible differences in NF- κ B binding activity between USP2 KD and control cells, we found a slight increase in Oct binding activity in USP2-deficient conditions."
NFkappaB affects USP2
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NFkappaB activates USP2. 1 / 1
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"TRAF6 and NF-kappaB Are Not Targets of USP2 in LPS Stimulated HL-60 Cells."
N-nitrosodiethylamine decreases the amount of USP2. 1 / 1
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No evidence text available

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"Indeed, in multiple lung cancer cell lines, ML364 reduced the levels of SKP2, as well as MDM2 and Aurora-A, two known substrates of USP2 (23, 31), serving as the positive controls, in a dose- (Fig. 3A) and time-dependent manner (Fig. 3B), which can be completely rescued by MG132 (Fig. 3C)."
MID1IP1 affects USP2
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No evidence text available
MEIS1 affects USP2
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MEIS1 decreases the amount of USP2. 1 / 1
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No evidence text available
MDM4 affects RAC
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EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
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sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
MAVS affects USP2
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No evidence text available
Lipopolysaccharides increases the amount of USP2. 1 / 1
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No evidence text available
LNX2 affects USP2
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No evidence text available
LMNA affects USP2
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No evidence text available
L-methionine affects USP2
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L-methionine increases the amount of USP2. 1 / 1
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No evidence text available
KCNN4 affects USP8
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USP2 binds USP8 and KCNN4. 1 / 1
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"Channel-DUB interactions were verified after 90 min as there were notable interactions of KCa3.1 with USP2 and USP8 and a weaker association with AMSH (associated molecule with the SH3 domain of STAM; Figure xref )."
IKZF3 affects USP2
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No evidence text available
IKBKG affects USP2
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No evidence text available
IHO1 affects USP2
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No evidence text available
HSD11B1 affects USP2
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HSD11B1 activates USP2. 1 / 1
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"We detected the potential downstream molecules of USP2 in primary hepatocytes treated with fructose, the expression of C/EBPα (C/ebpα) and 11β-HSD1 (Hsd11b1) was significantly increased both at mRNA (Fig. 6B&C) and protein level (Fig. 6D)."
GOLGA6L9 affects USP2
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No evidence text available
FOXJ2 affects USP2
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FOXJ2 decreases the amount of USP2. 1 / 1
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No evidence text available
EcR-A affects USP2
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EcR-A binds USP2. 1 / 1
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"Proliferation and differentiation processes of midgut stem cells are carried out via EcR-A and USP-2 heterodimer complex."
EEF1A2 affects USP2
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EEF1A2 binds RAC, USP2, and MDM4. 1 / 1
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sparser
"In multivariate analysis high level of EEF1A2, pAKT, USP2 and MDM4 were clearly associated with a shorter patients' survival ( xref )."
DUSP21 affects USP2
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No evidence text available
DSCR4 affects USP2
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No evidence text available
DISC1 affects USP2
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No evidence text available
CUE5 affects USP2
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"The increased binding of Cue5 to inactivated proteasomes, and its reduced binding upon ubiquitin removal with USP2, is reminiscent of that seen with Arabidopsis RPN10, where its proteasome occupancy s[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
CRBN affects USP2
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sparser
"Mechanistically, we find that USP2 directly interacts with CRBN, leading to the deubiquitination and stabilization of CRBN in senescent liver cancer cells."
CEP70 affects USP2
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No evidence text available
CCR8 affects USP2
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No evidence text available
CCDC57 affects USP2
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No evidence text available
CADPS affects USP2
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No evidence text available
CACNA2D2 affects USP2
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No evidence text available

sparser
"In addition to PER1 and BMAL1, USP2 deubiquitinates CRY1 in cultured cells in response to a serum shock, and in the mouse liver, Usp2 knockdown increases CRY ubiquitination and decreases CRY1 protein levels ( xref )."
Benzo[k]fluoranthene increases the amount of USP2. 1 / 1
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No evidence text available
BMP3 affects USP2
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BMP3 inhibits USP2. 1 / 1
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"These data indicate that MXD3 direct transcriptional regulation could account for both phenotypes described in this paper, namely proliferation (upregulation of putative direct targets IRF4, FBL, HTR1B) and growth arrest and apoptosis (upregulation of RUNX1T1 and BMP3; downregulation of USP2 and MYCL1) as shown in XREF_TABLE."
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No evidence text available
Aurora-A affects USP2
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USP2 binds Aurora-A. 1 / 1
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"For example, USP2 interacts and stabilizes fatty acid synthase [8], MDM2 [21], MDMX [22], Cyclin D1 [23], Aurora-A [24], Smad4 [25], and β-catenin [26]."
ATM affects USP2
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ATM phosphorylates USP2 on T137. 1 / 1
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"Collectively, these data suggest that ATM is a key regulator for USP2 recruitment to DSB sites by phosphorylating USP2 at S2 and T137, which is essential for the interaction between USP2 and RECQL4."
ASAH1 affects USP2
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ASAH1 increases the amount of USP2. 1 / 1
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"Thus, ACDase regulation by androgen in androgen sensitive LNCaP cells is mainly due to its prolonged protein half-life by androgen stimulated USP2 expression."
ACTN2 affects USP2
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No evidence text available
ABI2 affects USP2
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No evidence text available

No evidence text available
1H-pyrazole affects USP2
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1H-pyrazole decreases the amount of USP2. 1 / 1
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No evidence text available
1,2-dimethylhydrazine decreases the amount of USP2. 1 / 1
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No evidence text available