IndraLab

Statements


USP13 affects PTEN
15 1 | 69 19
USP13 binds PTEN.
15 | 13 18
15 | 13 18

sparser
"To further validate the interaction between USP13 and PTEN, an immunofluorescent confocal microscopic analysis was conducted."

sparser
"Moreover, purified GST-USP13, but not the GST-GFP control, was able to bind to FLAG-tagged PTEN under cell-free conditions ( xref ), which demonstrated a direct interaction between USP13 and PTEN."

sparser
"In contrast, USP13 directly binds and deubiquitylates PTEN to suppress tumorigenesis and glycolysis in PTEN-positive breast cancer cells ( xref )."

sparser
"Additionally, USP13 interacted with the tumor suppressor phosphatase and tensin homolog (PTEN)."

reach
"In contrast, USP13 directly binds and deubiquitylates PTEN to suppress tumorigenesis and glycolysis in PTEN-positive breast cancer cells (35)."

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USP13 deubiquitinates PTEN.
1 | 18
USP13 deubiquitinates PTEN. 10 / 19
1 | 18

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"However, Zhang et al. revealed that USP13 can deubiquitinate and stabilize the PTEN protein, thus inhibiting the Akt pathway and suppressing breast cancer progression [18]."

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"Wild-type USP13 purified from either bacteria or 293T cells, but not its catalytically inactive mutant C345A, decreased PTEN poly-ubiquitination by 64-70% in vitro (XREF_FIG)."

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"Deubiquitylation and stabilization of PTEN by USP13."

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"In addition to its oncogenic roles, USP13 exerts a tumor-suppressive role by deubiquitinating PTEN in different types of cancers."

"Deubiquitylation and stabilization of <span class="match term1">PTEN</span> by <span class="match term0">USP13</span>"

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"USP13, another PTEN deubiquitinase, different with USP7-mediated deubiquitination of PTEN monoubiquitination, it reverses PTEN polyubiquitination, thus promoting the PTEN stabilization and tumor suppr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"In these tumors, USP13 interacts with and deubiquitinates the tumor suppressor PTEN protein, thereby stabilizing and increasing the expression of PTEN [16–18]."

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"Ubiquitin-specific peptidase 13 (USP13) can deubiquitinate the PTEN protein and stabilize it."

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"Our findings revealed that USP13 alleviates MASH by directly binding to and deubiquitinating PTEN."

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"The decreased expression of USP13 promoted PTEN ubiquitylation, leading to PTEN degradation, thereby contributing to fibroblast activation and pathogenesis of IPF [101]."
USP13 increases the amount of PTEN.
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USP13 increases the amount of PTEN. 10 / 10
| 10

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"Furthermore, restoration of USP13 rescued PTEN expression, cell phenotypes and xenograft tumor growth which is altered by NF-kB activation."

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"In these tumors, USP13 interacts with and deubiquitinates the tumor suppressor PTEN protein, thereby stabilizing and increasing the expression of PTEN [16–18]."

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"USP13 upregulated PTEN expression and restrained oxidative stress, apoptosis, and inflammation in OA via AKT signaling [8]."

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"Results indicated that USP13 knockdown notably decreased PTEN protein expression and increased phosphor-AKT levels in BC cells."

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"Results suggested that the overexpression of NF-kB p65 significantly decreased the expression levels of USP13 and PTEN, and re-introduction of USP13 rescued PTEN expression."

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"Moreover, reintroduction of USP13 repressed the signaling transduction of NF-kB and largely rescued PTEN expression in NF-kB activated or miR-130b/301b overexpressed BC cells."

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"6 In OSCC tissues, PTEN expression is downregulated and negatively associated with activated AKT.7 10 Consistent with earlier findings, USP13 increased PTEN protein levels and decreased p-AKT levels in OSCC cells, and overexpression of PTEN partially reversed the USP13-knockdown-induced glucose uptake and lactate production in the present study.Loss of PTEN and abnormal AKT activation influences the regulation of various downstream effectors, thus contributing to tumorigenesis.11 13 GLUT1 and HK2 are important glycolysis regulators downstream of the AKT pathway.26 27 AKT activity is essential for insulin-induced GLUT1 gene expression."

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"Although each of these five DUBs could interact with endogenous PTEN (XREF_SUPPLEMENTARY), only one of them, USP13, significantly increased endogenous PTEN protein expression (XREF_SUPPLEMENTARY)."

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"In addition, reintroduction of USP13 partially rescued PTEN expression as well as the oncogenesis trend caused by NF-kB."

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"Our data revealed that knockdown of USP13 by short hairpin RNA (shRNA) significantly decreased PTEN protein expression in BC cells."
Modified USP13 increases the amount of PTEN. 2 / 2
| 2

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"Collectively, these data suggest that loss of USP13 may contribute to loss of PTEN in a substantial fraction of human tumors, whereas in other tumors PTEN can be inactivated by different mechanisms, including genetic alterations and upregulation of PTEN ubiquitin ligases (such as NEDD4-1 14 and WWP2 15)."

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"Additionally, we noticed that in fibroblasts, the loss of USP13 down-regulated PTEN protein expression by increasing PTEN ubiquitination."
USP13 bound to PTEN increases the amount of PTEN. 1 / 1
| 1

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"Mechanistically, USP13 directly bound to phosphatase and tensin homolog (PTEN) and deubiquitinated it, thereby elevating the PTEN expression and improving the MASH phenotype."
USP13 inhibits PTEN.
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USP13 inhibits PTEN. 8 / 10
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"Down-regulation of USP13 mediates PTEN protein loss and fibroblast phenotypic change, and thereby plays a crucial role in IPF pathogenesis."

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"Zhang et al. reported that USP13 suppresses tumorigenesis and glycolysis in PTEN-positive breast cancer through the deubiquitination of PTEN [18]."

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"In addition to its role in breast cancer, USP13 has been demonstrated to suppress tumor development via PTEN in oral squamous cell cancer [36] and bladder cancer [19]."

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"USP13 increases PTEN stability [126] , whereas USP7 reverses monoubiquitylation and nuclear accumulation of PTEN [125] ."

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"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

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"Conversely, overexpression of USP13 suppresses tumorigenesis and glycolysis in PTEN positive but not PTEN-null breast cancer cells."

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"The coding sequence of USP13 or PTEN was amplified with the following primers: USP13 (NM_003940.2), forward 5′-CGGAATTCATGCAGCGCCGGG-3′ and reverse 5′-CGGGATCCTTAGCTTGGTATCCTGCGG-3′; and PTEN (CR450306.1), forward 5′-CGGAATTCATGACAGCCATCATCAAAGAG-3′ and reverse 5′-CGGGATCCCGATCTCTTTGATGATGGCTG-3′."

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"Previous research showed that depletion of USP13 in breast cancer led to increased Akt phosphorylation, cell proliferation on 2D and 3D, glycolysis, and tumor growth by downregulating PTEN tumor suppressor proteins (25)."
USP13 activates PTEN.
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USP13 activates PTEN. 6 / 10
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"USP13 mediates PTEN to ameliorate osteoarthritis by restraining oxidative stress, apoptosis and inflammation via AKT dependent manner."

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"Previous studies have shown that knockdown of USP13 reduces PTEN, and over-expression of USP13 results in higher levels of PTEN [18]."

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"These data supported our notion that USP13 suppresses tumorigenesis by blocking the NF-kB-driven PTEN downregulation in BC."

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"Researchers attributed a tumor suppressor role to USP13 gene by showing that USP13 mediates PTEN signaling in different types of cancers (15–17)."

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"USP13 increases PTEN stability [126] , whereas USP7 reverses monoubiquitylation and nuclear accumulation of PTEN [125] ."

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"Similarly, Jing Geng et al. found that inhibition of USP13-mediated PTEN protein loss caused phenotypic changes in fibroblasts and impacted the pathogenesis of IPF."
USP13 decreases the amount of PTEN.
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USP13 decreases the amount of PTEN. 7 / 8
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"Furthermore, USP13 overexpression induced phosphatase and tensin homolog deleted on chromosome 10 (PTEN) expression and repressed the activation of AKT as well as the expression of the downstream effectors glucose transporter-1 (GLUT1) and hexokinase-2 (HK2)."

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"Reintroduction of USP13 could block the pathway and rescue PTEN protein expression, and thereby inhibited the carcinogenesis of bladder cells."

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"In these tumors, USP13 interacts with and deubiquitinates the tumor suppressor PTEN protein, thereby stabilizing and increasing the expression of PTEN [16–18]."

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"For example, USP13 overexpression induces an increase in the expression of the tumor suppressor PTEN in BC cells through deubiquitylation and stabilization of PTEN, thereby inhibiting tumor growth [ 1[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Knockdown of USP13 by USP13 shRNA, miR-130b/301b overexpression or NF-kB activation in BC cells leads to the loss of PTEN expression, which was subsequently demonstrated to be reversed by reintroduction of USP13."

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"Restoring USP13 levels partially inhibits the invasion and migration of tumor cells by stabilizing the expression of PTEN (84)."

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"Two independent USP13 shRNAs both decreased PTEN protein expression by 80% and increased phospho-AKT and phospho-FOXO1/3 levels by 3- to 5-fold in SUM159 breast cancer cells, while restoration of PTEN or expression of an RNAi resistant ' silence mutant ' (i.e., no amino acid change) of USP13 (USP13-RE) in USP13 depleted SUM159 cells completely reversed the effect of USP13 shRNA on upregulating the phosphorylation of AKT and FOXO (XREF_FIG and XREF_SUPPLEMENTARY)."
USP13 ubiquitinates PTEN.
| 4 1
USP13 ubiquitinates PTEN. 5 / 5
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sparser
"In order to determine whether PTEN is a direct substrate of USP13, we purified USP13 and ubiquitinated PTEN and then incubated them in a cell-free system."

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"For example, USP13 prevents tumor cell growth by deubiquitinating PTEN in breast cancer, OSCC and bladder cancer."

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"Ubiquitin C-terminal hydrolase 13 (USP13) has been found to reverse PTEN ubiquitination thus leading to its stabilization in breast cancer cells [ 92 ]."

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"Furthermore, several DUBs, including HAUSP/USP7, USP10, USP11, USP13, OTUD3, and Ataxin-3 have been shown to reverse the ubiquitination of PTEN, a key antagonist in the PI3K growth-promoting pathway, implicating its function in cancer-specific contexts [15]."

reach
"USP13, another PTEN deubiquitinase, different with USP7-mediated deubiquitination of PTEN monoubiquitination, it reverses PTEN polyubiquitination, thus promoting the PTEN stabilization and tumor suppr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP13 is modified
2 | 72 10
USP13 is phosphorylated.
2 | 62 10
USP13 is phosphorylated. 10 / 21
| 19 2

sparser
"As shown in xref , T196 was phosphorylated after DNA damage, and the T196A mutation inhibited USP13 phosphorylation after DNA damage."

sparser
"Together, these results suggest that USP13 phosphorylation by ATM is important for its recruitment to DSBs, DDR and chemo sensitivity."

sparser
"Next, we investigated how USP13 phosphorylation regulates its localization."

sparser
"Here the authors report that phosphorylated USP13 deubiquitinates RAP80 after DNA damage, prompting recruitment to the break site."

sparser
"Taken together, these results demonstrate that USP13 phosphorylation by ATM is important for USP13 recruitment to DSB sites following DNA damage and proper DDR."

sparser
"Identification of USP13 Phosphorylation in Ovarian Cancer Cells by LC-MS-MS Analysis."

sparser
"The activity, expression, and cellular localization of CK2 components could further impact USP13 phosphorylation in response to intrinsic or extrinsic cellular stresses in the tumor microenvironment."

sparser
"Following DNA damage, phosphorylated USP13 by ATM is recruited to DNA damage sites to cleave the ubiquitin chains from more than three sites of RAP80 (K75, K90, and K112), releasing their restriction on the K63-linked ubiquitin chain, improving the focus formation of the RAP80-BRCA1 complex, and eventually facilitating DDR."

sparser
"To further confirm these results, we generated site-specific antibody against p-Thr196 and examined the USP13 phosphorylation following DNA damage."

sparser
"The anti-pT122 antibody specifically detected a phosphorylated USP13 in HeyA8 and COV318 expressing wild-type USP13, not T122A mutant USP13 ( xref C,D)."
USP13 is phosphorylated on T122. 10 / 18
2 | 16

sparser
"Our data suggest that inhibiting Thr122 phosphorylation of USP13 may provide an effective therapeutic strategy for targeting USP13 in USP13-amplified ovarian cancer."

No evidence text available

sparser
"To understand the biological role of Thr122 phosphorylation of USP13 in ovarian cancer, we examined the effect of inhibiting T122 phosphorylation on cell proliferation."

sparser
"This study revealed that USP13 is phosphorylated at Thr122, which upregulates the stability of USP13 at a post-translation level."

sparser
"Here, we found that USP13 is phosphorylated at Thr122 in ovarian cancer cells."

sparser
"We also reveal that phosphorylation at Threonine 122 of USP13 promotes the proliferation of ovarian cancer cells."

sparser
"Inhibition of CK2 Decreased Thr122 Phosphorylation on USP13."

sparser
"To study the function of phosphorylation on Thr122 of USP13 in ovarian cancer, we generated an anti-phospho-Thr122 USP13 antibody (anti-pT122 antibody)."

sparser
"The treatment with CK2 inhibitor (CX-4945) dramatically reduced the signal of Thr122 phosphorylation of USP13 in HEK293T cells ( xref A)."

sparser
"To validate the activity of the anti-pT122 antibody, we generated the point mutation of Thr122 by using a site-directed substitution of threonine to alanine on USP13 to abolish USP13 phosphorylation on Thr122 residue."
USP13 is phosphorylated on Y708. 10 / 16
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sparser
"The Q607R mutant also up-regulated USP13 phosphorylation at Y708 ( xref Bii)."

sparser
"Further analysis found that tacrine hydrochloride significantly decreased CLK3-Q607R–enhanced purine production and USP13-Y708 phosphorylation in CCA cells ( xref )."

sparser
"Similar data were also observed in the EGF-induced increase in CLK3 expression, USP13 phosphorylation at Y708, and proliferation in CCA cells ( xref )."

sparser
"We demonstrated that (1) a recurrent Q607R somatic substitution in CLK3 was identified in 8% of 100 human CCAs, particularly in patients with CCA metastasis; (2) the expression of CLK3 was significantly up-regulated in CCA compared with matched control tissues; (3) CLK3 knockdown significantly inhibited CCA aggressiveness in vitro and in vivo; (4) gene ontology term enrichment and MS assays indicated that high CLK3 expression in CCA patients mainly regulated nucleotide metabolism, especially purine biosynthesis; (5) mechanistically, CLK3 directly phosphorylated USP13 at Y708, which promoted its binding to c-Myc, a critical purine synthesis–associated transcription factor, thereby preventing Fbxl14-mediated c-Myc ubiquitination and activating the transcription of purine metabolic genes; (6) the CCA-associated CLK3-Q607R mutant induced USP13-Y708 phosphorylation and enhanced the activity of c-Myc; (7) in turn, c-Myc transcriptionally up-regulated CLK3; (8) importantly, levels of CLK3 significantly correlated with the expression of phospho-USP13-Y708, c-Myc, and ATIC in human CCA specimens; and (9) tacrine hydrochloride was identified as a potential compound to inhibit the aberrant CLK3-enhanced CCA invasiveness."

sparser
"Thus, CLK3-mediated phosphorylation of USP13 at Tyr708 promotes cholangiocarcinoma progression by activating c-Myc-induced purine synthesis, providing a new and viable therapeutic target for the treatment of cholangiocarcinoma associated with CLK3 mutations ( xref )."

sparser
"Aurora B and ataxia telangiectasia mutated (ATM) regulate the expression of USP13 via phosphorylation of USP13 at Ser114 and Thr196 to promote the interaction between them, respectively, while the CDC-like kinase 3 (CLK3)-Q607R mutation induces the phosphorylation of USP13 at Tyr708 and promotes its association with c-Myc ( xref )."

sparser
"USP13 stabilizes its substrate, and at the same time, the activity of USP13 is upregulated by CLK3 via promoting phosphorylation of USP13 at Tyr708, which increases its deubiquitinating enzyme activity."

sparser
"Notably, the CCA-associated CLK3-Q607R mutant induced USP13-Y708 phosphorylation and enhanced the activity of c-Myc."

sparser
"CLK3-mediated phosphorylation of USP13 at Tyr708 promotes cholangiocarcinoma progression by activating c-Myc-induced purine synthesis, providing a new and viable therapeutic target for treating cholangiocarcinoma associated with CLK3 mutations [ xref ]."

sparser
"In cholangiocarcinoma, TGF-β signaling triggers the phosphorylation of CLK3, a serine/threonine kinase that directly phosphorylates USP13 at Y708 and facilitates USP13 interaction with c-Myc ( xref ); in GSCs, USP13 can enhance the stability through deubiquitinating c-Myc, activating purine synthesis mediated by c-Myc and inducing the tumorigenesis of GSCs ( xref ); in hepatocellular carcinoma, knockdown of USP13 by shRNA can markedly downregulate c-Myc expression, resisting xenograft tumor growth of HCC ( xref )."
USP13 is phosphorylated on T196. 10 / 11
| 5 6

rlimsp
"To further demonstrate the direct interaction between the MDC1 FHA domain and phosphorylated Thr 196 of USP13, T196 or p-T196 peptide was incubated with purified GST-MDC1 FHA domain in vitro."

rlimsp
"To further characterize the biological significance of USP13 phosphorylation, we stably expressed USP13 WT and T196A mutant in USP13-deficient cells."

sparser
"Recently, some scientists have reported that ATM is activated to phosphorylate USP13 at Thr196 site in response to DNA damage, stabilizing RAP80 to assemble BRCA1 to repair DSBs xref ."

sparser
"Aurora B and ataxia telangiectasia mutated (ATM) regulate the expression of USP13 via phosphorylation of USP13 at Ser114 and Thr196 to promote the interaction between them, respectively, while the CDC-like kinase 3 (CLK3)-Q607R mutation induces the phosphorylation of USP13 at Tyr708 and promotes its association with c-Myc ( xref )."

sparser
"Recently, researchers from Mayo Clinic uncover that ATM is activated in response to IR damage, and then phosphorylates USP13 at Thr196 site to stabilize RAP80, leading to the efficient recruitment of BRCA1 to the damage sites xref ."

sparser
"To further demonstrate the direct interaction between the MDC1 FHA domain and phosphorylated Thr 196 of USP13, T196 or p-T196 peptide was incubated with purified GST-MDC1 FHA domain in vitro ."

rlimsp
"Recently, researchers from Mayo Clinic uncover that ATM is activated in response to IR damage, and then phosphorylates USP13 at Thr196 site to stabilize RAP80, leading to the efficient recruitment of BRCA1 to the damage sites."

rlimsp
"To further confirm these results, we generated site-specific antibody against p-Thr196 and examined the USP13 phosphorylation following DNA damage."

rlimsp
"As shown in Supplementary Fig. 8c, T196 was phosphorylated after DNA damage, and the T196A mutation inhibited USP13 phosphorylation after DNA damage."

rlimsp
"As shown in Supplementary Fig. 8c, T196 was phosphorylated after DNA damage, and the T196A mutation inhibited USP13 phosphorylation after DNA damage. In addition, USP13 was phosphorylated on Thr196 following DNA damage in ATM-proficient cells but not ATM-deficient cells (Fig. 5c), suggesting ATM phosphorylates USP13 on T196 following DNA damage."
USP13 is phosphorylated on S114. 6 / 6
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"Specifically, Esposito et al. found that Aurora kinase B, crucial for mitosis, phosphorylates USP13 at the serine 114 and that this phosphorylation enhances the interaction of both proteins, thereby maintaining the stability of Aurora kinase B. Intriguingly, USP13 stabilizes Aurora kinase B by modulating its ubiquitination, a process that occurs independently of its DUB activity."

sparser
"Studies have shown that Aurora B phosphorylates USP13 at Ser114 and promotes its interaction with USP13."

sparser
"Aurora B also phosphorylates USP13 at Ser114 to promote the interaction between them and stabilizes the expression of Aurora B ( xref )."

sparser
"Aurora B and ataxia telangiectasia mutated (ATM) regulate the expression of USP13 via phosphorylation of USP13 at Ser114 and Thr196 to promote the interaction between them, respectively, while the CDC-like kinase 3 (CLK3)-Q607R mutation induces the phosphorylation of USP13 at Tyr708 and promotes its association with c-Myc ( xref )."

sparser
"Additionally, kinase Aurora B induced phosphorylation of the Ser114 residue in USP13 prevents several major human cancers by promoting its interaction with partners and stability ( xref )."

sparser
"Aurora B phosphorylates USP13 at Serine 114, prompting their interaction between USP13 and Aurora, showing a potential direct link between abnormalities occurring during the cell cycle and USP13 in several human cancers [ xref ]."
USP13 is phosphorylated on T196. 2 / 2
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rlimsp
"After 1 h, USP13 was immunoprecipitated and blots probed with the antibody specifically recognizing phosphorylation of USP13 at T196."

rlimsp
"(e) Control, USP13 knockout, and USP13 knockout cells stably expressing the indicated constructs were subjected to colony formation assay to assess the sensitivity of cells to cisplatin. Error bars represent±s.e.m. from three independent experiments. (f) HEK293T cells transfected with HA-MDC1 were left untreated or treated with cisplatin. Cell lysates were subjected to immunoprecipitation with HA antibody. The immunoprecipitates were then blotted with the indicated antibodies. (g) HEK293T cells transfected with deletion mutants of HA-MDC1 were subjected to co-immunoprecipitation as in f. (h) HEK293T cells transfected with WT or T196A USP13 were treated with cisplatin, and cell lysates were incubated with Sepharose coupled with GST or GST-MDC1-FHA domain. After washing, proteins bound on Sepharose were blotted with the indicated antibodies. (i) Nonphosphorylated or phosphorylated Thr 196 peptide (T196 or p-T196, respectively) was conjugated to Sepharose beads and incubated with purified GST-MDC1-FHA domain in NETN buffer."
USP13 is ubiquitinated.
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USP13 is ubiquitinated. 10 / 10
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sparser
"NEDD4 undergoes K29-linked auto-ubiquitination at K1279 and serves as a scaffold for recruiting USP13 to form a NEDD4-USP13 deubiquitination complex, which subsequently cleaves the K48-linked poly-ubiquitin chains from PIK3C3 at K419, thus stabilizing PIK3C3 to promote autophagy. the ubiquitination of USP13; however, NEDD4 appears to be its own substrate for ubiquitination, and the catalytic activity of NEDD4 is required for its auto-ubiquitination."

sparser
"On the other hand, coexpression of USP10 with USP13 has a much less pronounced effect on ubiquitination of USP13 (data not shown)."

sparser
"Protein interaction experiments demonstrated that USP13 could co-immunoprecipitate with endogenous TopBP1, and in vitro deubiquitination enzyme experiments showed that WT-USP13 could deubiquitinate TopBP1, while CA-USP13 could not, highlighting the requirement for USP13 ubiquitination activity ( xref )."

sparser
"USP13 and gp78 control ubiquitination of Ubl4A."

sparser
"We noticed that NEDD4-1 did not affect the ubiquitination of USP13, suggesting that USP13 is not a direct substrate of NEDD4-1 ( Figure S5 A)."

sparser
"LPS-induced USP13 degradation was attenuated by actinomycin D ( xref ), suggesting a new protein induced by LPS regulates USP13 ubiquitination."

sparser
"To investigate the molecular regulation of the unknown protein, which regulates USP13 ubiquitination and degradation, we examined several factors related to transcriptional regulation."

sparser
"We found that short time exposure of LPS did not trigger USP13 degradation, suggesting that an immediate LPS signaling through TLR4 does not contribute to USP13 ubiquitination and degradation."

sparser
"From the time points of LPS’ effect, we hypothesized that activation of JNK by LPS does not directly affect USP13 ubiquitination."

sparser
"To investigate whether LPS regulates USP13 degradation through ubiquitination, we examined the effect of LPS on USP13 ubiquitination."
USP13 affects MCL1
7 1 | 55 12
USP13 binds MCL1.
7 | 12 12
7 | 12 12

sparser
"We further conducted the in vitro immunoprecipitation experiment, the result of which suggested that the interaction between USP13 and MCL1 was direct and independent on MCL1 ubiquitination status (Fig.  xref )."

sparser
"Second, USP13 interacted with MCL1 and decreased MCL1 polyubiquitination in a DUB activity-dependent manner."

No evidence text available

sparser
"We first performed co-immunoprecipitation assays to confirm the interaction between USP13 and Mcl-1 by transfecting HEK293T cells with FLAG-USP13 and V5-hMcl-1."

sparser
"These findings suggest that the USP13-MCL-1 axis represents a potential therapeutic target for AKI management [ xref ]."

sparser
"USP13 interacts with MCL1 and stabilizes its expression by deubiquitinating MCL1."

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"However, thrombin treatment did not have any notable effect on the interaction between USP13 and MCL1."
USP13 activates MCL1.
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USP13 activates MCL1. 10 / 16
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"Our data shows that USP13 and Mcl-1 protein expression correlates, suggesting that USP13 mediated Mcl-1 stabilisation is clinically relevant."

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"USP13 promotes Mcl-1 stability."

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"The deubiquitinase (DUB) USP13 promotes Mcl-1 stabilisation in cervical cancer."

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"USP13 promotes MCL1 protein stability."

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"As a complimentary approach, we lentivirally expressed ectopic USP13 into OVCAR5, PC9 and NCI-H2170 cells, whose endogenous USP13 levels were relatively low, and found that overexpression of USP13 increased MCL1 protein, but not mRNA, abundance."

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"Taken together, our results indicated that pharmacological inhibition of USP13 by spautin-1 reduced MCL1 protein abundance and increased tumor cell sensitivity to ABT-263."

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"USP13 knockdown by siRNA markedly reduced the half-life of MCL1 in TOV-21G and SW-1573 cells."

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"In lung infection, USP13 are aberrantly expressed, inhibition of USP13 reduces the abundance of anti-apoptotic protein MCL1 in the lung [42]."

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"Conversely, exogenous USP13 overexpression evidently prolonged the half-life of endogenous MCL1 in NCI-H2170 and PC9 cells."

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"We concluded that USP13 promoted MCL1 stability at the post transcriptional level in lung and ovarian cancer cells."
USP13 deubiquitinates MCL1.
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USP13 deubiquitinates MCL1. 10 / 12
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"USP13, on the other hand, can deubiquitinate and stabilize MCL1, which is not sensitive to MCL-2 family inhibitors, and render tumor cells highly resistant to BH3-type chemotherapy drugs (Oltersdorf et al., 2005; Delbridge et al., 2016; Kotschy et al., 2016)."

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"USP13 interacts with and deubiquitinates Mcl-1 in cervical cancer cells."

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"Next, we confirmed that USP13 deubiquitinated Mcl-1."

"Collectively, we nominate <span class="match term0">USP13</span> as a novel deubiquitinase which regulates <span class="match term1">MCL1</span> turnover in diverse solid tumors and propose that <span class="match term0">USP13</span> may be a potential therapeutic target for the treatment of various malignancies."

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"Two DUBs USP9X and USP13 deubiquitinate and stabilise MCL1, and hypomorphic mutations in both have been linked to neurodevelopmental disorders and neurodegenerative disease [XREF_BIBR, XREF_BIBR]."

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"USP13 interacts with and deubiquitinates MCL1."

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"Another study has shown that USP13 is essential for HPV-positive cervical cancer cells to proliferate, at least in part by deubiquitinating and stabilizing the prosurvival protein Mcl-1 [122]."

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"Therefore, USP13 interacted with and deubiquitinated MCL1 to preserve its protein stability."

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"USP13 interacts with MCL1 and stabilizes its expression by deubiquitinating MCL1."

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"Morgan et al. (2021) demonstrated that USP13 deubiquitinates and stabilizes Mcl-1, promoting the proliferation in cervical cancer."
USP13 increases the amount of MCL1.
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USP13 increases the amount of MCL1. 9 / 9
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"We confirmed the siRNA screen results by transfecting each of the four oligos against USP13 into HEK293T cells, and indeed found that USP13 knockdown decreased the endogenous protein levels of MCL1, while modulating USP9X or OTUB2 with different siRNAs had no consistent effects on MCL1 expression."

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"Interestingly, Zhang et al. showed that pharmacological inhibition of USP13 by a small-molecule inhibitor, spautin-1, markedly downregulated MCL1 protein levels and sensitized ovarian and lung cancer cells to ABT-263, a selective antagonist of BCL-2 and BCL-XL.We selected DUB3 as an ideal target in ovarian cancer due to its oncogenic functions in various cancer subtypes ."

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"USP13 knockdown downregulates the expression of MCL-1 and enhances the sensitivity of tumor cells to ABT-263."

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"Loss of USP13 significantly reduced Mcl-1 protein expression, without effecting MCL1 mRNA expression."

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"Interestingly, over-expression or depletion of USP13 did not modulate Mcl-1 expression in HPV- C33A cells, suggesting that Mcl-1 expression is not regulated by USP13 in these cells."

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"Critically, treatment of USP13 depleted cells with the proteasome inhibitor MG132 rescued Mcl-1 protein levels, suggesting that USP13 protects Mcl-1 from proteasomal degradation."

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"In line with these observations, MCL1 protein levels were downregulated by depletion of USP9X but not USP13 (Figure S6D)."

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"In addition, we explored whether pharmacological inhibition of USP13 downregulated MCL1 protein expression and synergistically kill tumor cells in combination with ABT-263."

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"Specifically, one studied showed that USP13 depletion led to decreased MCL1 protein levels in lung cancer and ovarian cancer cells, therefore sensitizing tumor cells toward BH3 mimetic inhibitors (38)."
USP13 decreases the amount of MCL1.
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USP13 decreases the amount of MCL1. 3 / 3
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"Interestingly, although USP13 depletion reduced MCL1 protein levels, neither cell proliferation nor cell migration was significantly changed under normal growth conditions."

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"Therefore, if USP13 inhibition reduces the protein level of MCL1, it significantly elevates cancer cell apoptosis."

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"In addition, genetic depletion of USP13 using clustered regularly interspaced palindromic repeats (CRISPR)/Cas9, or pharmacological inhibition of USP13 by a small-molecule inhibitor spautin-1, markedly downregulates MCL1 protein expression and shows synergistic effects against tumor cells in combination with ABT-263, a selective antagonist of BCL-2 and BCL-XL."
USP13 ubiquitinates MCL1.
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USP13 leads to the ubiquitination of MCL1. 2 / 2
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"Finally, we confirmed that depletion of USP13 from both HeLa and SiHa cells increased the ubiquitination of endogenous Mcl-1, suggesting USP13 mediated deubiquitination of Mcl-1 is relevant in cervical cancer cells."

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"Conversely, ubiquitination of Mcl1 can be reversed by the deubiquitinase USP9X and the more recently identified USP13 ."
USP13 inhibits MCL1.
| 2
USP13 inhibits MCL1. 2 / 2
| 2

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"Hence, pharmaceutical intervention of USP13 activity is expected to antagonize the tumorigenic potential of MCL1 oncoprotein, and combined administration of USP13 inhibitors with clinically approved venetoclax therapy may represent a promising targeting strategy for the treatment of human cancer by inducing tumor cell death."

reach
"Usp13 knockdown obviously reduces MCL1 protein abundance and significantly increases the sensitivity of tumor cell to BH3 mimetic inhibitors targeting BCL-2 and BCL-XL [ 57 ]."

reach
"Consistently, knockout or pharmacological inhibition of USP13 impeded tumor cell proliferation and enhanced sensitivity to chemotherapeutic agents in both cell lines and mouse models (Han et al., 2016; Zhang et al., 2018)."

reach
"Inhibition of the phosphorylation of USP13 downregulated USP13 protein stability and reduced the proliferation of ovarian cancer cells, which may lead to novel therapeutics targeting USP13 in USP13-amplified cancers."

reach
"These results demonstrate that USP13 indirectly activates the Akt/mTOR pathway through WWP1.CCK-8 and colony formation assays revealed that WWP1 rescued the suppression of HCC cell proliferation caused by USP13 silencing (Fig. 5C, D)."

reach
"Our results indicate that spautin-1 as well as knockdown of its downstream targets, USP10 and USP13, reduced the proliferation and migration of glioblastoma cells."

reach
"Conversely, WWP1 knockdown reduced the increase in cell proliferation, migration, and stemness induced by USP13 overexpression (Supplementary Fig. S4C–I)."

reach
"Stable knockdown (KD) of USP13 by lentiviral short hairpin RNAs (shRNAs) markedly inhibited the proliferation of the USP13 amplified cell lines (CAOV3 and HeyA8) and the USP13 overexpressing cell line (OVCAR8)."

reach
"USP13 promotes the proliferation of small cell lung cancer (SCLC) (44)."

reach
"USP13 signaling in cell proliferation and apoptosis."

reach
"Ectopically expressed K488R of METTL3 significantly promoted cell proliferation and glycolysis, while USP13 knockdown decreased cell proliferation and glycolysis with WT METTL3 rather than K488R METTL3, which indicated that mutation of K488 of METTL3 promoted cell proliferation and glycolysis and abolished USP13 depletion-induced proliferation and glycolysis suppression (Fig. 4i-n)."

reach
"USP13 has been shown to upregulate fatty acid synthase (FASN) to promote the proliferation of SCLC cells and Twist-related protein 1 (Twist1) to promote the migration and invasion of breast cancer cells, while it inhibits cell proliferation via downregulation of glycolysis in oral squamous cell carcinoma (47,48)."

reach
"USP13 knockdown in BC cells promoted their proliferation, invasion and migration."

reach
"By contrast, KD of USP13 only modestly reduced proliferation of cell lines with low USP13 expression (SKOV3 and IGROV1) (XREF_FIG; XREF_SUPPLEMENTARY)."

reach
"In two other USP13 high HGSC cell lines (OAW28 and OVCAR3) XREF_BIBR, silencing USP13 also profoundly inhibited cell proliferation (XREF_SUPPLEMENTARY)."

reach
"Overexpression of miR-130b/301b or USP13 knockdown by shRNA delivery decreases USP13 expression, and further reduces PTEN protein expression and leads to enhancement of cell proliferation, invasion and migration."

reach
"Functional studies demonstrated that overexpression of USP13 suppressed OSCC cell proliferation, glucose uptake and lactate production in vitro and inhibited tumor growth in vivo."

reach
"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

reach
"Treatment with afatinib alone diminished the number of proliferating cells to approximately 36%, while USP13 shRNA only slightly decreased cell proliferation."

reach
"USP13 overexpression inhibited the proliferation of and glycolysis in OSCC cells in vitro and suppressed tumorigenesis in nude mice."

reach
"USP13 overexpression repressed cell proliferation and the Warburg effect in OSCC cells."

reach
"USP13 loss leads to decreased ccRCC cell proliferation in vitro and tumor growth in vivo."

reach
"Loss of USP13 in breast cancer cells promotes AKT phosphorylation, cell proliferation, anchorage independent growth, glycolysis and tumor growth through downregulation of PTEN."

reach
"Overexpression of miR-130b/301b or USP13 knockdown by shRNA delivery decreases USP13 expression, and further reduces PTEN protein expression and leads to enhancement of cell proliferation, invasion and migration."
USP13 affects MYC
2 2 1 | 38 14
USP13 binds MYC.
2 | 8 14
2 | 8 10

sparser
"The MYC-USP13 group also significantly reduced PTEN ubiquitination in comparison with control group in HCT116 cells ( xref )."

reach
"We showed a strong molecular association between USP13 and c-MYC, leading to the upregulation of squamous programs in murine and human lung cancer cells."

sparser
"By coexpressing USP13Myc and Bcl‐XL‐Flag in HEK293T cells, we discovered that USP13 could be detected in Bcl‐XL immunoprecipitates after pull down by anti‐Flag beads ( Figure   xref )."

reach
"USP13 and FBXL14 interact with c-Myc in glioma cells."

reach
"However, overexpression of CLK3 in HCCC9810 cells promoted the binding of c-Myc to WT-USP13, not USP13 Y708F mutant (XREF_FIG)."

reach
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag tagged USP13 or FBXL14."

reach
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

reach
"Computational modeling of structures predicted in ZDOCK and Pymol software showed that Y708 phosphorylation was necessary for the direct binding of USP13 to c-Myc (XREF_FIG)."

reach
"As USP13 is a deubiquitinase, the interaction between USP13 and c-Myc led us to hypothesize that USP13 may stabilize c-Myc protein in GSCs through deubiquitination."

No evidence text available
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
USP13 activates MYC.
| 6
USP13 activates MYC. 4 / 10
| 4

reach
"In HCC cell lines, knockdown of USP13 by shRNAs significantly reduced cell growth, and decreased levels of c‐Myc."

reach
"We found a positive correlation between TRF1 levels and all the stem cell markers with the exception of MYC, although TRF1 was positively correlated with the MYC modulator USP13 (Fang et al., 2017)."

reach
"Collectively, these data demonstrate that the ubiquitination insensitive mutant of c-Myc is able to rescue the phenotypes caused by USP13 knockdown or FBXL14 overexpression in vitro and in vivo, which supports that USP13 and FBXL14 mediate posttranslational regulation of c-Myc to control the stem cell like phenotype and tumorigenic potential of GSCs."

reach
"Aurora B and ataxia telangiectasia mutated (ATM) regulate the expression of USP13 via phosphorylation of USP13 at Ser114 and Thr196 to promote the interaction between them, respectively, while the CDC-like kinase 3 (CLK3)-Q607R mutation induces the phosphorylation of USP13 at Tyr708 and promotes its association with c-Myc (43)."
USP13 activates MYC. 2 / 2
| 2

reach
"In addition, IHC staining of Ki-67 and cleaved caspase 3 confirmed that USP13 disruption reduced c-Myc protein in the GSC derived xenografts and led to a significant decrease in cell proliferation and an increase in cell apoptosis within the tumor (not depicted)."

reach
"CHX chase assays indicated that overexpression of WT-USP13, particularly a phosphomimetic USP13-Y708E, but not USP-Y708F, significantly up-regulated the half-time of the c-Myc protein in HEK293 cells (XREF_FIG)."
USP13 deubiquitinates MYC.
1 1 | 9
USP13 deubiquitinates MYC. 9 / 9
1 1 | 7

reach
"In cholangiocarcinoma, TGF-β signaling triggers the phosphorylation of CLK3, a serine/threonine kinase that directly phosphorylates USP13 at Y708 and facilitates USP13 interaction with c-Myc (Zhou et al., 2020); in GSCs, USP13 can enhance the stability through deubiquitinating c-Myc, activating purine synthesis mediated by c-Myc and inducing the tumorigenesis of GSCs (Fang et al., 2017); in hepatocellular carcinoma, knockdown of USP13 by shRNA can markedly downregulate c-Myc expression, resisting xenograft tumor growth of HCC (Huang et al., 2020)."

reach
"Moreover, USP13 knockdown increased c-Myc ubiquitination, whereas FBXL14 knockdown reduced c-Myc ubiquitination (XREF_FIG)."

"In this study, we demonstrate that the deubiquitinase <span class="match term0">USP13</span> stabilizes c-<span class="match term1">Myc</span> by antagonizing FBXL14-mediated ubiquitination to maintain GSC self-renewal and tumorigenic potential."

reach
"In these tumors, USP13 deubiquitinates and stabilizes some oncogenes, including MCL1 [19, 20], Myc [8], MITF [21] and ZHX2 [22]."

"In this study, we demonstrate that the deubiquitinase USP13 stabilizes c-Myc by antagonizing FBXL14-mediated ubiquitination to maintain GSC self-renewal and tumorigenic potential. USP13 was preferentially expressed in GSCs, and its depletion potently inhibited GSC proliferation and tumor growth by promoting c-Myc ubiquitination and degradation."

reach
"The ubiquitination assay demonstrated that disruption of USP13 by shRNA markedly increased c-Myc ubiquitination and reduced c-Myc protein level in GSCs (XREF_FIG)."

reach
"Furthermore, USP13 can deubiquitinate and stabilise ACLY and OGDH and c-Myc in ovarian cancer and glioblastoma, respectively, thereby functioning as an oncogene [XREF_BIBR, XREF_BIBR]."

reach
"Our data have demonstrated that USP13 mediates deubiquitination of c-Myc, whereas FBXL14 facilitates c-Myc ubiquitination in glioma cells."

reach
"It has been reported that c-Myc can be de-ubiquitinated by USP13 [33]."
Modified USP13 leads to the deubiquitination of MYC. 2 / 2
| 2

reach
"Overexpression of USP13 reduced c-Myc ubiquitination, and forced expression of FBXL14 increased c-Myc ubiquitination (XREF_FIG)."

reach
"Importantly, overexpression of USP13 together with FBXL14 abolished the increased c-Myc ubiquitination caused by FBXL14 overexpression (XREF_FIG, last lane)."
USP13 increases the amount of MYC.
1 | 8
USP13 increases the amount of MYC. 5 / 5
| 5

reach
"As USP13 functions as a deubiquitinase that also interacts with c-Myc (XREF_FIG) and the preferential expression of USP13 in GSCs positively regulates c-Myc protein levels (XREF_FIG), we hypothesized that USP13 might mediate deubiquitination of c-Myc protein to prevent its degradation and stabilize c-Myc in GSCs."

reach
"The ubiquitination assay demonstrated that disruption of USP13 by shRNA markedly increased c-Myc ubiquitination and reduced c-Myc protein level in GSCs (XREF_FIG)."

reach
"In this study, we demonstrated that USP13 mediated deubiquitination and FBXL14 mediated ubiquitination regulate c-Myc protein levels in glioma cells and play a critical role in controlling the maintenance or differentiation of GSCs (XREF_FIG)."

reach
"Targeting USP13 reduces c-Myc protein levels and impairs GSC maintenance."

reach
"In vitro analysis showed inducible disruption of USP13 in GSCs by 70% reduced c-Myc protein levels and inhibited GSC proliferation (XREF_FIG)."
USP13 increases the amount of MYC. 4 / 4
1 | 3

reach
"The knockdown of USP13 by short hairpin RNAs (shRNAs) markedly decreases cell growth in HCC by reducing c-Myc expression."

"In this study, we demonstrate that the deubiquitinase USP13 stabilizes c-Myc by antagonizing FBXL14-mediated ubiquitination to maintain GSC self-renewal and tumorigenic potential. USP13 was preferentially expressed in GSCs, and its depletion potently inhibited GSC proliferation and tumor growth by promoting c-Myc ubiquitination and degradation."

reach
"In HCC cell lines, knockdown of USP13 by shRNAs markedly decreased HCC cell growth, and mechanistic investigations revealed that USP13 knockdown could markedly downregulate the expression levels of c-Myc."

reach
"In cholangiocarcinoma, TGF-β signaling triggers the phosphorylation of CLK3, a serine/threonine kinase that directly phosphorylates USP13 at Y708 and facilitates USP13 interaction with c-Myc (Zhou et al., 2020); in GSCs, USP13 can enhance the stability through deubiquitinating c-Myc, activating purine synthesis mediated by c-Myc and inducing the tumorigenesis of GSCs (Fang et al., 2017); in hepatocellular carcinoma, knockdown of USP13 by shRNA can markedly downregulate c-Myc expression, resisting xenograft tumor growth of HCC (Huang et al., 2020)."
USP13 ubiquitinates MYC.
| 5
USP13 leads to the ubiquitination of MYC. 5 / 5
| 5

reach
"Further, USP13 and FBXL14 reciprocally modulate MYC polyubiquitination in GSCs ."

reach
"The phosphorylation of USP13 at Tyrosine 708 by CLK3 is required for USP13′s binding to c-Myc, which prevents its Fbxl14-mediated ubiquitination of c-Myc in glioma stem cells [10]."

reach
"Silencing FBXL14 alongside USP13 disruption attenuated the increased c-Myc ubiquitination caused by USP13 disruption (XREF_FIG)."

reach
"USP13 mediates ubiquitination of c-Myc to antagonize FBXL14 mediated ubiquitination and thus maintains the stem cell like phenotype and tumorigenic potential of GSCs."

reach
"20 In addition, USP13 is aberrantly overexpressed in ovarian cancers, and USP13 drives ovarian cancer metabolism by deubiquitinating ATP citrate lyase and oxoglutarate dehydrogenase.21 USP13 disruption inhibited the proliferation of glioma stem cells by promoting the ubiquitination and degradation of c-Myc."
USP13 inhibits MYC.
| 2
USP13 inhibits MYC. 2 / 3
| 2

reach
"USP13, FBXL14 and c-Myc can interact with each other; USP13 inhibits the degradation of c-Myc, hence inhibition of USP13 promotes the apoptosis of glioma stem cells."

reach
"Moreover, treatment with a proteasome inhibitor blocked the c-Myc loss caused by USP13 knockdown (not depicted), but overexpression of USP13 delayed c-Myc turnover in the presence of cycloheximide (not depicted), suggesting a critical role of USP13 in preventing proteasomal degradation of c-Myc in GSCs."
USP13 affects BECN1
2 1 | 33 21
USP13 binds BECN1.
| 10 21
| 10 19

sparser
"USP13 directly interacts with Beclin-1 and regulates de-ubiquitination of Beclin-1."

sparser
"USP13 can interact with Beclin1, deubiquitinate, and stabilize it (Liu et al. xref )."

sparser
"Importantly, we identified a novel interaction between USP13 and Beclin-1, where USP13 facilitates the deubiquitination of Beclin-1, thereby promoting autophagy."

reach
"Furthermore, Beclin1 also interacts with USP13 and regulates the stability of USP13."

sparser
"USP13 interacts with Beclin 1 and stabilizes its expression, further regulating the expression of the Vps34 complex and p53."

sparser
"The Vps34 complexes can upregulate their expression by stabilizing the interaction between USP13 and Beclin 1, which in turn upregulates the expression of Beclin 1 by deubiquitination."

reach
"At the molecular level, the authors convincingly demonstrate that USP13 directly interacts with and deubiquitinates Beclin-1 (Figure 1B)."

sparser
"Mechanistically, USP13 interacted with Beclin-1, facilitating its deubiquitination and promoting autophagic flux, as shown by increased LC3 II/I ratios and decreased p62 levels."

sparser
"Intriguingly, a previous study revealed that USP13 and Beclin1 could interact with each other."

reach
"Moreover, USP13 directly interacts with Beclin 1 in untreated cells, while it dissociates from Beclin 1 after addition of Spautin-1 (XREF_FIG a)."
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
USP13 deubiquitinates BECN1.
1 1 | 11
USP13 deubiquitinates BECN1. 10 / 13
1 1 | 11

reach
"Furthermore, the presence of spautin-1 inhibited the deubiquitination of Beclin1 mediated by USP10 and USP13."

reach
"It promotes the degradation of Vps34 complexes by inhibiting USP10 and USP13, two ubiquitin specific peptidases that target the deubiquitination of Beclin-1."

reach
"In addition, USP10 or USP13 can deubiquitinate and stabilize Beclin 1 in promoting autophagy ."

reach
"Specific and potent autophagy inhibitor-1 (Spautin-1) was identified to inhibit USP10 and USP13, which deubiquitinate the Beclin 1 subunit of Vsp34 complex, and thus promoted the degradation of Vsp34 PI3 kinase complex."

reach
"Likewise, USP13, which deubiquitinates BECN1 and stabilizes PIK3C3 complexes, is amplified in LUSQ."

reach
"We demonstrate that USP10 and USP13 can both mediate the deubiquitination of Beclin1."

reach
"USP10 and USP13 have been shown to mediate the deubiquitination of Beclin 1, thereby stabilizing the Vps34 complex XREF_BIBR."

reach
"Recent studies indicated that USP13 can deubiquitinate and regulate protein levels of Beclin-1, microphathalmia associated transcription factor, Siah2, phosphatase and tensin homolog, and STAT-1, and the deubiquitination process of USP13 could also be orchestrated by Beclin-1."

reach
"In 2011, Yuan et al. found that USP13 could interact with the C-terminal domain of Beclin-1 subunit in the VPS34 complex and deubiquitinate Beclin-1, thereby enhancing the stability of the VPS34 complex, which would contribute to the formation of autophagosomes (Liu et al., 2011)."

reach
"Conversely, it was show that Beclin 1 is deubiquitinated by USP10 and USP13 and adding complexity, Beclin 1 itself controlled the protein stabilities of USP10 and USP13 by regulating their deubiquitinating activities, in turn regulating the levels of tumor suppressor p53 [XREF_BIBR]."
USP13 activates BECN1.
| 6
USP13 activates BECN1. 6 / 6
| 6

reach
"Spautin-1, a potent small molecule, can promote the degradation of PI3KCIII and Vps34 complexes by inhibiting two ubiquitin specific peptidases, USP10 and USP13 which target the Beclin-1 subunit of Vp[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"This molecule promotes the degradation of Vps34-PI 3 kinase complexes by inhibiting two ubiquitin specific peptidases, USP10 and USP13, that target the Beclin-1 subunit of Vps34 complexes."

reach
"Spautin-1 has been recently reported to prevent autophagy by inhibiting the deubiquitinases USP10 and USP13, which leads to beclin 1 and Vps34 (or PI3KIII) degradation [XREF_BIBR]."

reach
"XREF_BIBR Spautin-1 promotes the degradation of BECN1 by inhibiting 2 USPs (ubiquitin specific peptidases), USP10 and USP13, which target BECN1."

reach
"Overexpression of USP13 in PC12 cells almost reversed the Spautin‐1‐mediated decrease in NLRP3, USP13, Beclin1 and GSDMD‐N expression levels (Figure 3D,E)."

reach
"The function of USP10 and USP13 is reversed by a chemical compound, spautin-1, which inhibits autophagy by promoting Beclin-1 degradation."
USP13 ubiquitinates BECN1.
| 4
USP13 leads to the ubiquitination of BECN1. 4 / 4
| 4

reach
"Chemical Inhibition of the Deubiquitinating Enzymes USP10 and USP13 Causes Ubiquitination and Degradation of BECLIN 1."

reach
"For instance, USP10 and USP13 reduce Beclin-1 ubiquitination to prevent its degradation."

reach
"Inhibition of the Deubiquitinating Enzymes USP10 and USP13 Causes Ubiquitination of Beclin 1."

reach
"On the other hand, autophagy can be also inhibited blocking PI3KC3 complex formation; Spautin-1 indirectly inhibits the activity of VPS34 by proteosomal degradation of proteins that form VPS34 complexes through reduction of Beclin-1 deubiquitination mediated by USP10 and USP13 [120]."
USP13 increases the amount of BECN1.
1 | 1
USP13 increases the amount of BECN1. 2 / 2
1 | 1

"Similarly, the overexpression of USP13 reduced the levels of ubiquitinated Beclin1 which was inhibited by spautin-1 (Figure 4E)"

reach
"Moreover, beclin-1 and USP10 are involved in a potential feedforward mechanism in which beclin-1 stabilizes USP13, which in turn deubiquitinates and stabilizes USP10, leading to increased beclin-1 levels and activity."
USP13 decreases the amount of BECN1.
| 1
USP13 decreases the amount of BECN1. 1 / 2
| 1

reach
"Deubiquitination experiments demonstrated that overexpression of USP13 could reduce the ubiquitination level of Beclin-1, and this effect could be counteracted by the USP13 inhibitor spautin-1."
PTEN affects USP13
15 | 15 18
PTEN binds USP13.
15 | 13 18
15 | 13 18

sparser
"To further validate the interaction between USP13 and PTEN, an immunofluorescent confocal microscopic analysis was conducted."

sparser
"Moreover, purified GST-USP13, but not the GST-GFP control, was able to bind to FLAG-tagged PTEN under cell-free conditions ( xref ), which demonstrated a direct interaction between USP13 and PTEN."

sparser
"In contrast, USP13 directly binds and deubiquitylates PTEN to suppress tumorigenesis and glycolysis in PTEN-positive breast cancer cells ( xref )."

sparser
"Additionally, USP13 interacted with the tumor suppressor phosphatase and tensin homolog (PTEN)."

reach
"In contrast, USP13 directly binds and deubiquitylates PTEN to suppress tumorigenesis and glycolysis in PTEN-positive breast cancer cells (35)."

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PTEN activates USP13.
| 2
PTEN activates USP13. 2 / 2
| 2

reach
"These data suggested that the PTEN/AKT pathway mediated the effects of USP13 on the glycolysis in OSCC cells."

reach
"A comprehensive analysis revealed that bergenin augmented PTEN protein level by promoting the association between PTEN and ubiquitin-specific protease 13 (USP13), thereby boosting the stability of PTEN."
USP13 affects Flag
| 41
| 41

sparser
"Co-transfection of NIH/3T3 cells with FlagUSP13, HA–Ub/K63, and His–STING showed that USP13 removed the ubiquitin chain of STING, especially the K63-linked ubiquitin chain ( xref A)."

sparser
"Next, we generated Flag-USP13 Mut, the USP13 enzymatically inactive triple mutant containing C345A, M664E, and M739E, to observe the restoration of ubiquitination."

sparser
"HEK293T cells were co-transfected with GFP-tagged HMGB1 (GFP-HMGB1), Flag-USP13, or empty plasmids."

sparser
"To determine whether USP13 influences the stability of HMGB1, we overexpressed Myc-HMGB1 in HEK293T cells and then increased the number of Flag-USP13 plasmids."

sparser
"Flag-USP13 Mut, however, failed to stabilize HMGB1 protein levels (Fig.  xref C), indicating that USP13 regulates HMGB1 protein stability."

sparser
"Next, we transfected Myc-tagged WWP1 (Myc-WWP1), along with Flag-tagged USP13 (Flag-USP13) or an empty vector, into HCC and HEK293T cells, followed by a Co-IP assay to validate the binding between USP13 and WWP1."

sparser
"Co-IP results further corroborated the exogenous interaction of USP13 and STING in NIH/3T3 cells co-transfected with FlagUSP13 and His–STING ( xref D)."

sparser
"To determine the ubiquitin linkage patterns of WWP1 by USP13, we co-transfected Myc-WWP1 with HA-Ub mutants with all lysine mutations (K0), or all lysine mutations but one (K6, K11, K27, K29, K33, K48, or K63), along with or without Flag-USP13, into HEK293T cells (Fig.  xref H)."

sparser
"To further verify the involvement of K29- and K48-linked polyubiquitination of WWP1 by USP13, we co-transfected Myc-WWP1 with wild-type (WT) HA-Ub or HA-Ub with a single lysine mutation (K6R, K11R, K27R, K29R, K33R, K48R, or K63R), along with or without Flag-USP13, into HEK293T cells (Fig.  xref I)."

sparser
"To further confirm the role of USP13 on METTL3 ubiquitination, HEK 293T cells were co-transfected with Flag-USP13 (either WT or C345A mutant), Myc-METTL3 and HA-Ub."
BECN1 affects USP13
2 | 15 21
BECN1 binds USP13.
| 10 21
| 10 19

sparser
"USP13 directly interacts with Beclin-1 and regulates de-ubiquitination of Beclin-1."

sparser
"USP13 can interact with Beclin1, deubiquitinate, and stabilize it (Liu et al. xref )."

sparser
"Importantly, we identified a novel interaction between USP13 and Beclin-1, where USP13 facilitates the deubiquitination of Beclin-1, thereby promoting autophagy."

reach
"Furthermore, Beclin1 also interacts with USP13 and regulates the stability of USP13."

sparser
"USP13 interacts with Beclin 1 and stabilizes its expression, further regulating the expression of the Vps34 complex and p53."

sparser
"The Vps34 complexes can upregulate their expression by stabilizing the interaction between USP13 and Beclin 1, which in turn upregulates the expression of Beclin 1 by deubiquitination."

reach
"At the molecular level, the authors convincingly demonstrate that USP13 directly interacts with and deubiquitinates Beclin-1 (Figure 1B)."

sparser
"Mechanistically, USP13 interacted with Beclin-1, facilitating its deubiquitination and promoting autophagic flux, as shown by increased LC3 II/I ratios and decreased p62 levels."

sparser
"Intriguingly, a previous study revealed that USP13 and Beclin1 could interact with each other."

reach
"Moreover, USP13 directly interacts with Beclin 1 in untreated cells, while it dissociates from Beclin 1 after addition of Spautin-1 (XREF_FIG a)."
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
BECN1 activates USP13.
| 4
BECN1 activates USP13. 4 / 4
| 4

reach
"Taken together, these results suggest that Beclin1 is the primary target of USP10 and USP13."

reach
"Another interesting observation made in the Spautin-1 study is that knock-down of VPS34 and Beclin 1 causes instability of USP10 and USP13, which indicates the existence of a regulatory feedback loop [XREF_BIBR]."

reach
"Interestingly, Beclin-1 knockout also reduced USP13 protein abundance in turn, indicating the two proteins are regulated reciprocally."

reach
"The close functional interconnection between the central PI3K-III subunits and the DUBs is demonstrated by the interesting observation that knock-down of VPS34 and Beclin 1 causes instability of USP10 and USP13, which strongly indicates the existence of a regulatory feedback loop [XREF_BIBR]."
BECN1 increases the amount of USP13.
1 | 1
BECN1 increases the amount of USP13. 2 / 2
1 | 1

reach
"Unexpectedly, we found that the knockdown of Beclin1 or Vps34 could also reduce the endogenous levels of USP10 and USP13 (XREF_FIG)."

"We found that endogenous Beclin1 can interact with USP13 and the interaction was reduced in the presence of spautin-1 (Figure 5C). Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."
BECN1 deubiquitinates USP13.
1 |
BECN1 deubiquitinates USP13. 1 / 1
1 |

"We found that endogenous Beclin1 can interact with USP13 and the interaction was reduced in the presence of spautin-1 (Figure 5C). Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."
USP13 affects STING1
1 1 | 18 19
USP13 binds STING1.
1 | 8 19
1 | 8 19

sparser
"Co-IP results further corroborated the exogenous interaction of USP13 and STING in NIH/3T3 cells co-transfected with Flag–USP13 and His–STING ( xref D)."

sparser
"In contrast, the truncate (aa301-863) that preserves DUB activity and UBA motifs inhibited cGAS and STING-mediated activation of IFN-β promoter and catalysed deubiquitination of STING ( xref ), indicating that USP13-mediated deubiquitination of STING is associated with its suppression of antiviral signalling."

No evidence text available

sparser
"Interestingly, a previous study found that USP13 interacts with STING and promotes the deubiquitination of STING, thereby lessening the antiviral responses by decreasing the activation of STING–TBK1 signaling xref ."

sparser
"Here, we showed the interaction between STING and USP13 in cardiomyocytes during DIC, and the K63-linked deubiquitination of STING by USP13 resulted in the STING degradation via a P62-dependent autophagic way."

sparser
"USP13, a deubiquitinating enzyme, interacts with STING and catalyzed removal of K27 O- or K33 O-linked but not K27 R-linked polyubiquitin chains from STING."

sparser
"Moreover, USP22 significantly promoted the interaction between USP13 and STING ( xref )."

sparser
"Similar to WT USP22, both USP22-C185A and USP22-UCH facilitated the interaction between STING and USP13, but USP22-ZNF failed to do so ( xref )."

sparser
"Here, we perform an unbiased screen by coimmunoprecipitation assays in cells cotransfected with FLAG-tagged DUBs and HA-STING, and find that USP13 interacts with STING."

reach
"Here, we perform an unbiased screen by coimmunoprecipitation assays in cells cotransfected with FLAG tagged DUBs and HA STING, and find that USP13 interacts with STING."
USP13 deubiquitinates STING1.
1 | 4
USP13 deubiquitinates STING1. 5 / 5
1 | 4

"Here, we show that ubiquitin-specific protease 13 (<span class="match term0">USP13</span>) is a STING-interacting protein that catalyses deubiquitination of <span class="match term1">STING</span>."

reach
"The USP family of DUBs is very important, with USP13, USP18, USP20, USP21, USP44, and USP49 shown to mediate the deubiquitination of STING (Chen et al., 2021)."

reach
"Consistently, reconstitution of USP13 but not USP13 (AE) into Usp13 m/m MEFs inhibited HSV-1-induced ubiquitination of STING (XREF_FIG)."

reach
"For example, USP13 regulates antiviral responses by deubiquitinating STAT1 and STING (Sun et al. 2017; Yeh et al. 2013) and is related to PTEN and MITF in the modulation of tumorigenesis (Zhang et al. 2013; Zhao et al. 2011)."

reach
"In addition, USP13 negatively modulates antiviral immunity by deubiquitinating STING, and targeting USP13 consequently triggers STING-interferon signaling and strengthens innate immunity [26]."
USP13 ubiquitinates STING1.
| 4
USP13 ubiquitinates STING1. 4 / 4
| 4

reach
"Nevertheless, USP13 deficiency enhances antiviral responses through deubiquitinating stimulator of interferon (STING) [56]."

reach
"Consistently, reconstitution of USP13 but not USP13 (AE) into Usp13 m/m MEFs inhibited HSV-1-induced ubiquitination of STING (XREF_FIG)."

reach
"USP13 negatively regulates antiviral responses by deubiquitinating STING."

reach
"These data together suggest that USP13 inhibits the recruitment of TBK1 to STING by deubiquitinating STING."
USP13 inhibits STING1.
| 2
USP13 inhibits STING1. 2 / 2
| 2

reach
"In this sense, USP13 inhibition may activate the STING pathway along with innate immunity, which may provide a synergetic effect with ICIs."

reach
"The K27 linked-polyubiquitin chains are removed by the deubiquitinating enzyme (DUB) ubiquitin-specific protease 13 (USP13), which inhibits the basal activity of MITA in a manner dependent on blocking its recruitment of TBK1 (Sun et al., 2017)."
USP13 affects UIMC1
4 1 | 33
USP13 deubiquitinates UIMC1.
1 | 23
USP13 deubiquitinates UIMC1. 10 / 24
1 | 23

reach
"USP13 deubiquitinates RAP80, regulating RAP80-BRCA1 foci formation (55)."

reach
"Deubiquitination of RAP80 by USP13 plays an important role in the ability of RAP80 to bind polyubiquitin, which is important for RAP80 recruitment and DDR."

reach
"Our results suggest that USP13 deubiquitinates RAP80 following DNA damage, which in turn facilitates RAP80 recruitment to DSBs."

reach
"USP13 can deubiquitinate RAP80 (receptor-associated protein 80) and promote the recruitment of the RAP80-BRCA1 complex to damage sites, fine-tuning the DNA repair system ."

reach
"USP13, in turn, deubiquitinates RAP80 and promotes RAP80 recruitment and proper DDR."

reach
"These results suggest that deubiquitination of RAP80 by USP13 following DNA damage facilitates the binding between RAP80 and K63- linked polyubiquitin chain."

reach
"USP13 interacts with and deubiquitinates RAP80."

reach
"USP13 deubiquitinates receptor associated protein 80 (RAP80) and promotes DNA damage response."

reach
"Since USP13 deubiquitinates RAP80 and regulates DDR, we next examined the role of USP13 in cancer."

reach
"Also, USP13 interacts to and deubiquitylates receptor-associated protein 80 (RAP80) following DNA damage to activate its ability, promoting DNA-damage responses ."
USP13 binds UIMC1.
4 | 6
4 | 6

reach
"To test our hypothesis, we first examined whether USP13 interacts with RAP80."

reach
"Also, USP13 interacts with and deubiquitylates RAP80 to activate its function ."

No evidence text available

No evidence text available

No evidence text available

reach
"USP13 interacts with and deubiquitinates RAP80."

No evidence text available

reach
"Also, USP13 interacts to and deubiquitylates receptor-associated protein 80 (RAP80) following DNA damage to activate its ability, promoting DNA-damage responses ."

reach
"To evaluate the recruitment of USP13 and RAP80 at DSBs, we analysed the kinetics of GFP-USP13 and GFP-RAP80 accumulation on stripes after laser microirradiation."

reach
"Consistent with their reports, we also found that RAP80 binded with USP13 in EC cells treated with or without cisplatin."
USP13 ubiquitinates UIMC1.
| 3
USP13 ubiquitinates UIMC1. 3 / 3
| 3

reach
"We found that mutating the 3 sites (K75, K90 and K112, 3KR) near or at the UIM domains, but not the other sites, abolished USP13 deficiency induced RAP80 ubiquitination (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"As shown in XREF_FIG, USP13 deficiency dramatically elevated the ubiquitination of RAP80 in vivo."

reach
"Taken together, USP13 has an impact on ubiquitination of RAP80, instead of its protein degradation, to regulate its focus formation and DDR-related function."
USP13 inhibits UIMC1.
| 1
USP13 inhibits UIMC1. 1 / 1
| 1

reach
"USP13, on the other hand, reverses an inhibitory ubiquitylation on RAP80 close to its UIM, which disrupts its binding to K63-linked polyUb, enabling BRCA1-A recruitment to DSBs [97]."
Spautin-1 affects USP13
| 2 31
Spautin-1 inhibits USP13.
| 2 19
Spautin-1 inhibits USP13. 10 / 21
| 2 19

reach
"Spautin-1 inhibits autophagy by reducing the activity of USP10 and USP13, which promotes a major aspect of autophagy: degradation by Vps34 complexes."

reach
"Spautin-1 promotes the degradation of PIK3C3 (phosphatidylinositol 3-kinase catalytic subunit type 3, also termed Vps34 in yeast) complexes by inhibiting USP10 (ubiquitin specific peptidase 10) and USP13 (ubiquitin specific peptidase 13)."

reach
"Spautin-1 inhibits USP10 and USP13, two deubiquitinating enzymes that regulate the stability of BECN1, a key autophagy protein [291]."

reach
"A study by Liu et al. (Liu et al., 2011) has demonstrated that Spautin-1 is a potent small-molecule inhibitor of USP10 and USP13, and treatment with Spautin-1 inhibits the deubiquitinase activity of USP10 and USP13, resulting in an increasing ubiquitination and accelerating degradation of Beclin1 in Vps34 complexes, ultimately inhibiting autophagy."

reach
"Spautin-1 was shown to inhibit USP-13, a de-ubiquitinase of the host cell hijacked by the SARS-CoV-2 which led to successful ubiquitination of nsp-13 [109]."

eidos
"In both HeLa and SiHa cells , inhibition of USP13 by Spautin-1 resulted in a dose-dependent decrease in Mcl-1 protein expression ( Fig. 8A ) ."

reach
"Spautin-1, which effectively inhibits two ubiquitin-specific peptidases, USP10 and USP13, that target Beclin-1, was reportedly shown to effectively inhibit autophagy [17] ."

eidos
"Spautin-1 also reduced USP13 expression , as has been previously observed in other cell lines ( Fig. 8A ) ."

reach
"Spautin-1 inhibits the activity of two ubiquitin specific peptidases, USP10 and USP13, causing an increase in proteasomal degradation of class III PI3 kinase complexes, which have been shown to regulate autophagy [XREF_BIBR]."

reach
"Moreover, treatment of Spautin-1 almost abolished the ability of USP13 to cleave ubiquitin moieties from polyubiquitinated METTL3 in 143B cells and HEK 293T cells (Fig. 8c, d)."
Spautin-1 activates USP13.
| 9
Spautin-1 activates USP13. 9 / 9
| 9

reach
"Specifically, Spautin-1 promotes Beclin-1 degradation, by acting as an antagonist for two ubiquitin-specific peptidases, USP10/USP13, which are known to regulate Beclin-1 deubiquitination in Vps34 complexes."

reach
"Spautin-1 reportedly inhibits autophagosomal initiation and maturation by inhibiting deubiquitylases USP10 and USP13, which stabilize the Beclin-1 and VPS34 complex (26)."

reach
"The effect of nutrient deprivation on axonal maximal length was attenuated by the specific autophagy inhibitor Spautin-1, which prevents the deubiquitinating activity of USP10 and USP13 ( Liu et al., [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The Autophagy Inhibitor Spautin-1 Antagonizes Rescue of Mutant CFTR Through an Autophagy-Independent and USP13-Mediated Mechanism."

reach
"Spautin-1 has been recently reported to prevent autophagy by inhibiting the deubiquitinases USP10 and USP13, which leads to beclin 1 and Vps34 (or PI3KIII) degradation [XREF_BIBR]."

reach
"USP13 inhibitor, Spautin-1, treatment also impaired HR in control but not USP13 deficient cells (XREF_SUPPLEMENTARY)."

reach
"Spautin-1 reportedly suppresses autophagy by inhibiting the USP10 and USP13 deubiquitinases."

reach
"Spautin-1 targets the activity of two ubiquitin specific peptidases, USP10 and USP13 and causes PI3K complex degradation."

reach
"Spautin-1 promotes PINK1-PRKN-mediated mitophagy independent of USP10 and USP13.."
Spautin-1 decreases the amount of USP13.
| 2
Spautin-1 decreases the amount of USP13. 2 / 2
| 2

reach
"Spautin-1 inhibits USP13 expression, but the effect is not specific; its role in the treatment of diseases deserves further investigation."

reach
"Spautin-1 also reduced USP13 expression, as has been previously observed in other cell lines."
Spautin-1 ubiquitinates USP13.
| 1
Spautin-1 ubiquitinates USP13. 1 / 1
| 1

reach
"Spautin-1 inhibits deubiquitinating enzymes USP10 and USP13 and promotes degradation of the BECN1-PI3K autophagy initiation complex, which is required for canonical autophagy induction [30]."
MCL1 affects USP13
7 | 14 12
MCL1 binds USP13.
7 | 12 12
7 | 12 12

sparser
"We further conducted the in vitro immunoprecipitation experiment, the result of which suggested that the interaction between USP13 and MCL1 was direct and independent on MCL1 ubiquitination status (Fig.  xref )."

sparser
"Second, USP13 interacted with MCL1 and decreased MCL1 polyubiquitination in a DUB activity-dependent manner."

No evidence text available

sparser
"We first performed co-immunoprecipitation assays to confirm the interaction between USP13 and Mcl-1 by transfecting HEK293T cells with FLAG-USP13 and V5-hMcl-1."

sparser
"These findings suggest that the USP13-MCL-1 axis represents a potential therapeutic target for AKI management [ xref ]."

sparser
"USP13 interacts with MCL1 and stabilizes its expression by deubiquitinating MCL1."

No evidence text available

No evidence text available

No evidence text available

reach
"However, thrombin treatment did not have any notable effect on the interaction between USP13 and MCL1."
MCL1 activates USP13.
| 2
MCL1 activates USP13. 2 / 2
| 2

reach
"Interestingly, restoration of Mcl-1 expression only partially restored cell proliferation in USP13 depleted cervical cancer cells."

reach
"Here, we provided several lines of evidence to support that MCL1 was a deubiquitination target of USP13 in lung and ovarian cancer."
ATM affects USP13
| 13 11 9
ATM phosphorylates USP13. 10 / 23
| 8 8 7

sparser
"Recent research indicates that ATM phosphorylation of USP13, following DNA damage, enables USP13 to deubiquitinate RAP80."

sparser
"In response to DNA damage, USP13 is phosphorylated by ATM, which in turn facilitates USP13 recruitment to DSBs, RAP80 deubiquitination, and triggers DDR signalling."

reach
"This result suggests that USP13 may be phosphorylated by ATM following DNA damage."

sparser
"Taken together, these results demonstrate that USP13 phosphorylation by ATM is important for USP13 recruitment to DSB sites following DNA damage and proper DDR."

sparser
"Together, these results suggest that USP13 phosphorylation by ATM is important for its recruitment to DSBs, DDR and chemo sensitivity."

sparser
"Mechanistically, USP13 is phosphorylated by ATM following DNA damage which, in turn, facilitates its DSB localization."

sparser
"This result suggests that USP13 may be phosphorylated by ATM following DNA damage."

rlimsp
"Mechanistically, USP13 is phosphorylated by ATM following DNA damage which, in turn, facilitates its DSB localization."

reach
"USP13 is phosphorylated by ATM following DNA damage."

reach
"Mechanistically, USP13 is phosphorylated by ATM following DNA damage which, in turn, facilitates its DSB localization."
ATM phosphorylates USP13 on T196. 9 / 9
| 5 3 1

reach
"In addition, USP13 was phosphorylated on Thr196 following DNA damage in ATM-proficient cells but not ATM deficient cells (XREF_FIG), suggesting ATM phosphorylates USP13 on T196 following DNA damage."

reach
"Additionally, ATM phosphorylates USP13 at Thr196, which is important for the recruitment of USP13 to double-strand breaks and the induction of DNA damage."

reach
"Recently, researchers from Mayo Clinic uncover that ATM is activated in response to IR damage, and then phosphorylates USP13 at Thr196 site to stabilize RAP80, leading to the efficient recruitment of BRCA1 to the damage sites 17."

rlimsp
"In addition, USP13 was phosphorylated on Thr196 following DNA damage in ATM-proficient cells but not ATM-deficient cells (Fig. 5c), suggesting ATM phosphorylates USP13 on T196 following DNA damage."

sparser
"Additionally, ATM phosphorylates USP13 at Thr196, which is important for the recruitment of USP13 to double-strand breaks and the induction of DNA damage."

sparser
"Mechanistically, USP13 is phosphorylated at Thr196 by ATM following DNA damage, which facilitates the localization of USP13 at the double-strand breaks."

sparser
"USP13 is phosphorylated at Threonine 196 by ATM in response to the DNA damage response, which facilitates its recruitment to DNA double-strand breaks in HEK293T cells [ xref ]."

reach
"Recently, some scientists have reported that ATM is activated to phosphorylate USP13 at Thr196 site in response to DNA damage, stabilizing RAP80 to assemble BRCA1 to repair DSBs 17."

reach
"USP13 is phosphorylated at Threonine 196 by ATM in response to the DNA damage response, which facilitates its recruitment to DNA double-strand breaks in HEK293T cells [15]."
ATM phosphorylates USP13 on T196. 1 / 1
| 1

rlimsp
"Recently, some scientists have reported that ATM is activated to phosphorylate USP13 at Thr196 site in response to DNA damage, stabilizing RAP80 to assemble BRCA1 to repair DSBs."
| 3 29
| 3 15

reach
"LPS induces USP13 degradation by the ubiquitin-proteasome system in Kupffer cells.."

reach
"MG-132, but not leupeptin, attenuated LPS-induced USP13 degradation (Fig. 4B–D)."

reach
"Inhibition of JNK had no effect on LPS reduction of USP13 in Raw264.7 cells, while JNK inhibitor attenuates USP13 degradation in Kupffer cells."

reach
"Inhibition of proteasome, but not lysosome or immunoproteasome, attenuates LPS-induced USP13 degradation, suggesting USP13 degradation is mediated by the ubiquitin-proteasome system."

reach
"Inhibition of NF-κB, JNK, and p38 had no effect on LPS-induced USP13 reduction (L. Li et al., 2019)."

reach
"However, in contrast to USP13 mRNA changes in Raw264.7 cells, LPS reduces USP13 stability, but not mRNA levels, in Kupffer cells."

eidos
"LPS induces USP13 degradation by the ubiquitin-proteasome system in Kupffer cells ."

eidos
"LPS diminishes USP13 levels , but not USP13 mRNA , in Kupffer cells ."

reach
"Catalytic activity of USP13 is not necessary for LPS-reduced USP13.."

reach
"Thus, catalytic activity of USP13 is a minor factor for regulating its half-life, while it is not necessary for LPS-induced USP13 degradation."
Lipopolysaccharide decreases the amount of USP13.
| 10
Lipopolysaccharide decreases the amount of USP13. 10 / 10
| 10

reach
"We have shown that LPS reduced USP13 mRNA expression in Raw264.7 cells."

reach
"Interestingly, USP13 expression was decreased in human-FLSs after stimulation by LPS, TNF-α and IL-1β."

reach
"Furthermore, LPS reduced USP13 mRNA levels in Raw264.7 cells (L. Li et al., 2019)."

reach
"To further investigate molecular mechanisms of USP13 reduction in an inflammatory condition, we confirmed our published data and showed that intratracheal LPS challenge diminished USP13 protein levels in murine lungs (Fig. 1A and B)."

reach
"We found that LPS treatment of Raw264.7 cells diminished USP13 levels in both dose- and time-dependent manners (Fig. 1 H–K), consistent with our previous study."

reach
"To investigate the molecular mechanisms by which LPS reduces USP13 levels in macrophages in the liver, we examined the effect of LPS on USP13 levels in Kupffer cells."

reach
"LPS treatment starting from 10 ng/ml decreased USP13 levels after 24 h (Fig. 2A–D)."

reach
"The western blotting further confirmed that LPS reduces USP13 protein levels without altering its transcription (Fig. 3B), suggesting that LPS may affect USP13 protein stability."

reach
"We revealed that LPS reduces USP13 protein levels in both cell types, while USP13 mRNA is reduced in Raw264.7 cells, but not in Kupffer cells."

reach
"We have shown that LPS treatment decreased USP13 levels in mouse macrophage-like RAW264.7 cells ; however, the molecular regulation of USP13 during the development of pulmonary fibrosis has not been investigated."
Lipopolysaccharide ubiquitinates USP13.
| 2
Lipopolysaccharide leads to the ubiquitination of USP13. 2 / 2
| 2

reach
"We found that short time exposure of LPS did not trigger USP13 degradation, suggesting that an immediate LPS signaling through TLR4 does not contribute to USP13 ubiquitination and degradation."

reach
"Furthermore, LPS increases USP13 polyubiquitination."
| 2

reach
"LPS-induced USP13 degradation was attenuated by actinomycin D (Fig. 6), suggesting a new protein induced by LPS regulates USP13 ubiquitination."

reach
"As shown in Fig. 4A, LPS increased USP13 lysine (K)48-linked polyubiquitination."
USP13 affects STAT1
2 | 12 14
USP13 binds STAT1.
2 | 2 14
2 | 2 14

sparser
"The ability of USP13(C343A) to remove ubiquitin molecules from STAT1 was decreased (Fig.  xref ), although USP13(C343A) still binds to STAT1 (Supplementary Fig.  xref )."

sparser
"Additionally, we reveal the protective role of the cardiomyocyte-specific USP13-STAT1 axis in myocardial hypertrophy, indicating that cardiac-specific gene therapy targeting USP13 could represent a promising therapeutic strategy for cardiac hypertrophy."

sparser
"These observations validated that USP13 directly binding STAT1 protein in cardiomyocytes."

sparser
"Usp13 mainly interacted with the nonactivated form of Stat1 ( xref )."

sparser
"This study illustrated a beneficial effect of USP13 in hypertrophic cardiomyocytes and identified a cardiomyocyte-specific USP13-STAT1 axis in regulating cardiac hypertrophy."

sparser
"Here, the authors show the beneficial effects of a cardiomyocyte-derived deubiquitinating enzyme, USP13, in hypertrophic cardiomyocytes and identify a cardiomyocyte-specific USP13-STAT1 axis in regulating cardiac hypertrophy."

No evidence text available

No evidence text available

reach
"Moreover, we observed substantial attenuation of the interaction between STAT1 and USP13 upon overexpression of Mbtps1 (Fig. 7f)."

sparser
"Thus, we hypothesize that USP13-STAT1 axis protects against cardiac hypertrophy through upregulating Nppb transcription and BNP expression."
USP13 deubiquitinates STAT1.
| 6
USP13 deubiquitinates STAT1. 6 / 6
| 6

reach
"A recent report suggested that the deubiquitinase USP13 decreased STAT1 ubiquitination, which however was independent of IFNs treatment [XREF_BIBR]."

reach
"For example, USP13 regulates antiviral responses by deubiquitinating STAT1 and STING (Sun et al. 2017; Yeh et al. 2013) and is related to PTEN and MITF in the modulation of tumorigenesis (Zhang et al. 2013; Zhao et al. 2011)."

reach
"This finding was consistent with the findings of a recent study showing that USP13 deubiquitinates STAT1 ."

reach
"We detected that overexpression of USP13 attenuated the ubiquitination of STAT1, reversing the MBTPS1-induced increase in STAT1 ubiquitination, reduction in stability, and subsequent decrease in STAT1 protein levels (Fig. 7i–k)."

reach
"This competition subsequently interfered with the deubiquitination of STAT1 by USP13, promoting its degradation via the ubiquitin‒proteasome pathway."

reach
"Congruently, STAT1 ubiquitination is reduced in cells by USP13 overexpression and increased with USP13 knockdown regardless of IFNα treatment (115)."
USP13 activates STAT1.
| 1
USP13 activates STAT1. 1 / 3
| 1

reach
"In this study, we detected an interaction between MBTPS1 and STAT1, which disrupted the USP13-dependent deubiquitination mediated STAT1 stabilization."
USP13 ubiquitinates STAT1.
| 2
USP13 ubiquitinates STAT1. 2 / 2
| 2

reach
"We detected that overexpression of USP13 attenuated the ubiquitination of STAT1, reversing the MBTPS1-induced increase in STAT1 ubiquitination, reduction in stability, and subsequent decrease in STAT1 protein levels (Fig. 7i–k)."

reach
"Other targets of interest within the UPS include USP13, which has been reported to regulate IFN signaling via its ubiquitylation of STAT1 [108]."
USP13 inhibits STAT1.
| 1
USP13 inhibits STAT1. 1 / 2
| 1

reach
"Mechanistic studies reveal that tumor cell-intrinsic MBTPS1 competes with USP13 for binding to STAT1, thereby disrupting the USP13-dependent deubiquitination-mediated STAT1 stabilization."
USP13 affects HMGB1
4 | 18 9
USP13 binds HMGB1.
4 | 7 8
4 | 7 8

sparser
"In this study, we demonstrated, based on LC–MS/MS, immunoprecipitation, and PLA studies, that the deubiquitinase USP13 interacts with HMGB1."

sparser
"Further study including the binding of USP13 to HMGB1 by proinflammatory stimulation is under investigation."

reach
"To confirm the interaction between HMGB1 and USP13, PLA was performed using the Duolink in situ fluorescence kit (Sigma-Aldrich)."

reach
"In this study, we investigated the effect of isopeptidase T-3, also called USP13, on HMGB1, since HMGB1 ubiquitination has rarely been reported.To confirm whether USP13 binds to HMGB1, both Myc-HMGB1 and Flag-tagged USP13 (Flag-USP13) or USP10 and USP15 plasmids were overexpressed in HEK293T cells for co-immunoprecipitation."

reach
"As shown in Fig. 1B and 1C, HMGB1 binds to USP13 but not to either USP10 or USP15 after immunoprecipitation with Myc."

reach
"HEK293T cells were immunoprecipitated with an anti-USP13 antibody for endogenous binding, and the binding of endogenous HMGB1 to USP13 is shown (Fig. 1D)."

reach
"As expected, Spautin-1 treatment decreased the binding of HMGB1 to USP13, suggesting that inhibition of the deubiquitinase activity of USP13 affects its binding to HMGB1 (Fig. 1F)."

reach
"The interaction between HMGB1 and USP13 was also observed using a proximity ligation assay (PLA) in HEK293T cells overexpressing HMGB1 and USP13 (Fig. 1G)."

reach
"Further study including the binding of USP13 to HMGB1 by proinflammatory stimulation is under investigation."

sparser
"USP13 Interacts with HMGB1."
USP13 increases the amount of HMGB1.
| 4
USP13 increases the amount of HMGB1. 4 / 4
| 4

reach
"In addition, the level of HMGB1 protein was significantly increased by USP13 overexpression and decreased by shUSP13 or Spautin-1 treatment, showing that HMGB1 levels are regulated by the deubiquitinase USP13."

reach
"Knockdown of USP13 expression by shUSP13 plasmid or inhibition of USP13 activity by Spautin-1 treatment decreased the stability of HMGB1 protein (Fig. 4D and E), demonstrating that USP13 regulates HMGB1 protein levels."

reach
"HMGB1 secretion was promoted by USP13 overexpression in a dose-dependent manner (Fig. 6A)."

reach
"Collectively, USP13 positively regulates HMGB1 secretion."
USP13 deubiquitinates HMGB1.
| 4
USP13 deubiquitinates HMGB1. 4 / 4
| 4

reach
"We demonstrated that USP13 deubiquitinates and stabilizes HMGB1."

reach
"USP13 deubiquitinates HMGB1."

reach
"Subsequently, we tested whether USP13 could deubiquitinate HMGB1."

reach
"Collectively, our results suggest that USP13 deubiquitinates HMGB1 to maintain its stability and is a key molecule that regulates HMGB1 secretion (Fig. 7)."
USP13 ubiquitinates HMGB1.
| 1 1
USP13 ubiquitinates HMGB1. 2 / 2
| 1 1

reach
"Thus, HMGB1 ubiquitination and deubiquitination by SYVN1 and USP13, respectively, are important mechanisms for controlling the homeostasis of HMGB1."

sparser
"Thus, HMGB1 ubiquitination and deubiquitination by SYVN1 and USP13, respectively, are important mechanisms for controlling the homeostasis of HMGB1."
USP13 activates HMGB1.
| 2
USP13 activates HMGB1. 2 / 2
| 2

reach
"Deubiquitination by USP13 increases HMGB1 stability."

reach
"These results suggest that USP13 increases HMGB1 nucleocytoplasmic translocation via a CRM1-dependent nuclear export pathway.We further studied the effects of USP13 on HMGB1 secretion."
USP13 affects Neoplasms
| 28
USP13 activates Neoplasms.
| 21
| 21

reach
"Conversely, overexpression of USP13 promoted xenografted tumor growth, size and weight in vivo (Fig. S2j, k)."

reach
"We further demonstrated that USP13 obviously promoted OS cells invasion using a 3D tumor spheroid cell-invasion assay (Fig. 2c, d)."

reach
"Finally, miR‐367 silencing also significantly inhibited 3D cell invasion in tumor spheroids of both USP13‐MED and USP7‐ATRT cells (Fig. 4), suggesting that miR‐367 silencing inhibits two key features of tumor aggressiveness, namely, stem‐like properties and invasive behavior."

reach
"Importantly, overexpression of Usp13 accelerates tumorigenesis, enhances tumor metastasis, and causes poor outcomes in transgenic mouse models of ovarian cancer , underscoring its importance in promoting tumorigenesis in vivo."

reach
"USP13 promoted the growth of HCC tumors, since tumors in the USP13 group were larger than those in the NC group."

reach
"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

reach
"In hepatocellular carcinoma, the tumor growth and metastasis are promoted by USP13 through its regulation of TLR4 and subsequent activation of the TLR4/MyD88/NF-κB signaling cascade [23]."

reach
"The in vivo rescue experiments also indicate that re-expression of USP13 or PTEN can partially restore tumor growth due to PTEN or USP13 knockout, respectively (36)."

reach
"Nude mouse experiments indicated that depletion of USP13 in GC cells dramatically suppressed tumor growth in vivo."

reach
"USP13 signaling in tumor metastasis."
USP13 inhibits Neoplasms.
| 7
| 7

reach
"The reduced volume and weight of tumor nodules caused by USP13 silence was found to be counteracted by ectopic overexpression of METTL3 in 143B cells (Fig. 5a, b)."

reach
"Our results showed that ZHX2 overexpression could at least partially rescue decreased tumor volume caused by USP13 depletion (Fig. 4 M and N)."

reach
"For example, USP13 overexpression induces an increase in the expression of the tumor suppressor PTEN in BC cells through deubiquitylation and stabilization of PTEN, thereby inhibiting tumor growth [ 1[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In conclusion, USP13 either promotes or inhibits the development of different tumors."

reach
"In addition to its role in breast cancer, USP13 has been demonstrated to suppress tumor development via PTEN in oral squamous cell cancer [36] and bladder cancer [19]."

reach
"For example, Han et al. (95) reported that inhibiting ubiquitin-specific peptidase 13 (USP13), which upregulates enzymes critical for mitochondrial respiration and glutaminolysis, suppressed tumor progression and sensitized ovarian tumor cells to a PI3K/AKT inhibitor."

reach
"Targeting USP13 may inhibit tumor growth and provide additional benefits in cooperation with DDR inhibitors and immunotherapy."
USP13 affects ACLY
1 1 | 15 10
USP13 binds ACLY.
1 | 10
1 | 8

No evidence text available

sparser
"Mechanistically, Spautin-1 blocks the USP13-ACLY axis, thereby causing mitochondrial damage and releasing DAMPs and cytokines, which promotes the infiltration and activation of CD8 + T cells in DLBCL."

sparser
"These data indicate that Spautin-1 induces mitochondrial damage by inhibiting the USP13-ACLY axis, thereby suppressing DLBCL growth."

sparser
"USP13 directly binds to ACLY."

sparser
"First, the interaction between USP13 and ACLY was confirmed through co-immunoprecipitation analysis."

sparser
"Mechanistically, Spautin-1 promotes PANoptosis by inducing mitochondrial damage mediated by USP13-ACLY inhibition in DLBCL cells, which in turn induces the release of injury-related molecular patterns and cytokines."

sparser
"Although our research results indicate that Spautin-1 exerts anti-tumor activity dependent on the induction of PANoptosis, there are still three key limitations that need to be addressed: (1) The mechanism by which Spautin-1 regulates the USP13-ACLY axis has not been thoroughly verified. (2) The expression pattern and prognostic value of USP13 have not yet been verified in clinical samples."

sparser
"Furthermore, to explore the precise regions responsible for the interaction between USP13 and ACLY, we conducted deletion analyses."

sparser
"Likewise, we discovered that ACLY’s CoA ligase domain binds to USP13 (Fig. xref )."
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."
USP13 activates ACLY.
| 6
USP13 activates ACLY. 6 / 9
| 6

reach
"USP13 led to the upregulation of ACLY and oxoglutarate dehydrogenase (OGDH), which is one of the subunit of α-ketoglutarate dehydrogenase complex, an enzymatic complex involved in glutaminolysis."

reach
"USP13 primarily deubiquitinates and therefore upregulates ATP citrate lyase and oxoglutarate dehydrogenase, two crucial enzymes that control glutaminolysis and fatty acid synthesis and also mitochondrial respiration."

reach
"USP13 upregulates ACLY and OGDH, two key regulators that drive glutaminolysis, ATP generation and lipid synthesis in cancer metabolism."

reach
"USP13-KD also significantly decreased ACLY activity, while USP13 overexpression enhanced ACLY activity in the ACLY activity assay (XREF_SUPPLEMENTARY)."

reach
"In ovarian cancer, USP13 upregulates ATP citrate lyase (ACLY) via deubiquitination, enhancing fatty acid synthesis and accelerating tumour growth [16]."

reach
"Next, we examined whether USP13 augments the stability of ACLY and OGDH."
USP13 deubiquitinates ACLY.
1 | 7
USP13 deubiquitinates ACLY. 8 / 8
1 | 7

reach
"In ovarian tumors, upregulation of USP13 enhances deubiquitination and stabilization of ACLY (ATP citrate lyase) and OGDH (oxoglutarate dehydrogenase), two key enzymes that drive glutaminolysis, ATP generation, and lipid synthesis in cancer metabolism , and MCL1, a pivotal member of the antiapoptotic BCL-2 family of proteins ."

"<span class="match term0">USP13</span>, the main regulator of ovarian cancer energy metabolism, specifically deubiquitinates and stabilizes oxoglutarate dehydrogenase and ATP citrate lyase, which can catalyze fatty acid synthesis, glutaminolysis and mitochondrial respiration."

reach
"Furthermore, USP13 can deubiquitinate and stabilise ACLY and OGDH and c-Myc in ovarian cancer and glioblastoma, respectively, thereby functioning as an oncogene [XREF_BIBR, XREF_BIBR]."

reach
"As it was aforementioned, USP13 deubiquitinates OGDH and ACLY."

reach
"Taken together, these studies suggest that USP14 is involved in the occurrence and progression of multiple malignant tumors.Other USP family members: USP13, the main regulator of ovarian cancer energy metabolism, specifically deubiquitinates and stabilizes oxoglutarate dehydrogenase and ATP citrate lyase, which can catalyze fatty acid synthesis, glutaminolysis and mitochondrial respiration."

reach
"USP13 influences cell cycle progression in gastric cancer by stabilizing cyclin D1 [15] and enhances the deubiquitination and stabilization of ATP citrate lyase (ACLY) and oxoglutarate dehydrogenase (OGDH) in ovarian tumors [16]."

reach
"USP13 directly deubiquitinates ACLY and OGDH."

reach
"Mechanistically, USP13 promoted the energy metabolism of tumor cells, and provided precursor substances for the synthesis of sugar, lipids and non-essential amino acids in cancer cells, through deubiquitinating and stabilizing ACLY and OGDH (Han et al., 2016)."
USP13 inhibits ACLY.
| 2
USP13 inhibits ACLY. 2 / 2
| 2

reach
"These data suggest that USP13-KD induces mitochondrial dysfunction and lipogenic dysfunction by reducing OGDH and ACLY activity, thereby reducing glutamine 's reductive carboxylation and glucose 's oxidation for lipid synthesis (XREF_FIG)."

reach
"Depletion of USP13 markedly decreased protein stability of ACLY and OGDH (XREF_FIG; XREF_SUPPLEMENTARY), but had no effect on the transcription of ACLY and OGDH (XREF_FIG)."
NEDD4 affects USP13
| 11 19
NEDD4 binds USP13.
| 10 19
| 10 19

sparser
"NEDD4 undergoes K29-linked auto-ubiquitination at K1279 and serves as a scaffold for recruiting USP13 to form a NEDD4-USP13 deubiquitination complex, which subsequently cleaves the K48-linked poly-ubiquitin chains from PIK3C3 at K419, thus stabilizing PIK3C3 to promote autophagy. the ubiquitination of USP13; however, NEDD4 appears to be its own substrate for ubiquitination, and the catalytic activity of NEDD4 is required for its auto-ubiquitination."

sparser
"Since knockdown of VPS34 did not affect the association between NEDD4-1 and USP13 ( Figure 4 D), we hypothesized that NEDD4-1 may first bind to USP13 to form an NEDD4-1-USP13 complex."

sparser
"In addition, KO of BECN1 could not affect the interaction of NEDD4-1 with USP13 or VPS34, indicating that the NEDD4-1-USP13 complex binds to VPS34 in a Beclin-1-independent manner ( Figure S4 E)."

sparser
"Immunoblot analysis showed that NEDD4-1 interacted with USP13 at 1 h post-EBSS incubation, while the apparent interaction between NEDD4-1-USP13 and VPS34 could be detected at a later time point (at 2 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"These results suggest that the formation of NEDD4-1-USP13 complex is an essential prerequisite for VPS34 deubiquitination."

reach
"We observed that NEDD4 interacts with USP13 upon autophagy induction, whereas NEDD4USP13 interacts with PIK3C3 at a later time point, indicating that NEDD4 might recruit USP13 to form a NEDD4 and USP13 complex."

sparser
"Vps34, a unique member of the class III phosphoinositide 3-kinase, is an autophagy-associated protein that controls autophagosome generation, and its ubiquitination is downregulated by the NEDD4-USP13 complex."

sparser
"NEDD4-1 recruits USP13 to form a NEDD4-USP13 complex, thereby increasing the interaction between USP13 and Vps34 and inhibiting K48-linked ubiquitination of Vps34 on K419."

sparser
"USP13 could not affect the ubiquitination and protein stability of the VPS34 K419R mutant ( Figure 4 K), further confirming that the NEDD4-1-USP13 complex mainly cleaves K48-linked poly-ubiquitin chai[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Subsequently, the NEDD4-USP13 axis cleaves the K48-linked poly-ubiquitin chains on PIK3C3 at K419 and promotes autophagy through suppressing the proteasomal degradation of PIK3C3."
NEDD4 activates USP13.
| 1
NEDD4 activates USP13. 1 / 1
| 1

reach
"A recent study identified a role of USP13 in VPS34 deubiquitination and stabilization and NEDD4-1 autoubiquitination promotes the recruitment of USP13 to VPS34 35."
USP13 affects Cohesin
| 1 10 18
USP13 binds Cohesin.
| 1 9 17
| 1 9 17

reach
"USP13 interacts with cohesin and regulates its ubiquitination in human cells."

sparser
"As shown in xref D , the interaction between cohesin and USP13 was enhanced in cells arrested in the S phase when compared with asynchronous cycling cells, or cells arrested at other points in the cell cycle."

sparser
"These data indicate that the interaction of USP13 with cohesin is cell cycle dependent—it is enhanced during DNA replication"

sparser
"We next sought to determine which (if either) of these domains was required for the interaction of USP13 with cohesin."

sparser
"These data demonstrate that the UBA1/2 domains are required for the interaction of USP13 with cohesin and that the ZnF domain is dispensable for the interaction of USP13 with cohesin."

sparser
"We wondered whether ubiquitinated cohesin subunits were present primarily in the soluble fraction (where the USP13cohesin complex resides), in the chromatin fraction, or both."

sparser
"Mass spectrometry analysis revealed that USP13 could interact with Cohesin, and USP13 functioned as both ubiquitinase and deubiquitinase for Cohesin subunits."

sparser
"USP13 and cohesin interact especially during DNA replication, and USP13 regulates the ubiquitination/deubiquitination of the interacting subunit of cohesion through its UBA1/2 domain ( xref )."

sparser
"Interaction of endogenous USP13 with endogenous cohesin in cultured human cells."

sparser
"To further confirm the interaction between USP13 and cohesin, endogenous cohesin was immunoprecipitated with SMC3 antibodies from whole cell lysates prepared from genetically unmodified HCT116 cells and Western blot performed with USP13 antibodies (and STAG2 antibodies as a positive control)."
USP13 ubiquitinates Cohesin.
| 1 1
USP13 ubiquitinates Cohesin. 2 / 2
| 1 1

reach
"Finally, USP13 dependent ubiquitination of cohesin is required for the dissociation of cohesin from chromatin during mitosis."

sparser
"Finally, USP13-dependent ubiquitination of cohesin is required for the dissociation of cohesin from chromatin during mitosis."
USP13 affects TWIST1
4 | 16 8
USP13 binds TWIST1.
4 | 4 8
4 | 4 8

sparser
"Here, we investigated a potential physical/functional interaction between Twist1 and USP13 in human breast cancer."

sparser
"USP13 directly interacts with Twist1."

reach
"Although the biochemical and molecular functions of USP13 have been explored in various human cancers, including breast cancer, the oncogenic or tumor suppressor roles of USP13 in different studies has remained controversial.Here, we investigated a potential physical/functional interaction between Twist1 and USP13 in human breast cancer."

reach
"In accordance with the observed association between USP13 and Twist1, we detected an efficient co-precipitation between endogenous USP13 and Twist1 in SUM159PT and MDA-MB231 cells (Fig. 2a)."

sparser
"We found that USP13 and Twist1 can form a negative feedback loop to jointly regulate the migration and invasion of breast cancer cells."

sparser
"We also found that USP13 can directly interact with Twist1 and specifically cleave the K48-linked polyubiquitin chains of Twist1 induced by FBXL14."

sparser
"The interaction between USP13 and Twist1 was further confirmed by IF analysis since we found that USP13 co-localizes with Twist1 in the nuclei of MDA-MB231 cells (Fig.  xref )."

reach
"The interaction between USP13 and Twist1 was further confirmed by IF analysis since we found that USP13 co-localizes with Twist1 in the nuclei of MDA-MB231 cells (Fig. 2d)."

reach
"Therefore, the above findings indicate a direct interaction between USP13 and Twist1."

sparser
"For instance, USP13 directly interacts with Twist1 and cleaves FBXL14-induced K48-linked polyubiquitin chains, increasing Twist1 protein levels and facilitating BC cell migration and tumor metastasis in vitro and in vivo xref ."
USP13 deubiquitinates TWIST1.
| 5
USP13 deubiquitinates TWIST1. 5 / 5
| 5

reach
"We found that USP13 de-ubiquitinates Twist1 by binding to the WR region."

reach
"We found that Twist1 can be de-ubiquitinated by USP13."

reach
"USP13 deubiquitinates Twist1."

reach
"USP13 inhibits FBXL14-induced Twist1 ubiquitination."

reach
"USP13, which acts as an oncogene, was shown to promote breast cancer metastasis by deubiquitinating Twist 1 [35]."
USP13 increases the amount of TWIST1.
| 3
USP13 increases the amount of TWIST1. 3 / 3
| 3

reach
"Moreover, we found that the level of Twist1 increased in a USP13 dose-dependent manner, but not that of USP13AE, when Twist1 was co-expressed with increasing amounts of USP13 or USP13AE in HEK293T cells (Fig. 1f)."

reach
"We found that USP13, when co-expressed with Twist1 in HEK293T cells, remarkably increased the protein, but not the mRNA level of Twist1, an effect comparable to that observed after treatment with 20 μM proteasome inhibitor MG132 (Fig. 1d and Fig. S2a)."

reach
"We found that in two cell lines, T47D and ZR-75–30, which represent typical luminal cells with undetected endogenous Twist1 expression, the expression of USP13 significantly increased the levels of Twist1 and its downstream proteins (Fig. 4f)."
USP13 ubiquitinates TWIST1.
| 2
USP13 ubiquitinates TWIST1. 2 / 2
| 2

reach
"USP13 promotes breast cancer metastasis through FBXL14-induced Twist1 ubiquitination."

reach
"Moreover, USP13 had no effect on the ubiquitination of Twist1 by β-Trcp1, presumably due to the ineffectiveness of USP13 in Twist1 phosphorylation.As a typical transcription factor, Twist1 can recognize and bind to the E-box region of downstream gene promoters to activate or inhibit the transcription of target genes [68]."
USP13 activates TWIST1.
| 2
USP13 activates TWIST1. 2 / 2
| 2

reach
"A possible, but one-sided explanation may be that normal physiological concentrations of Twist1 individually inhibit USP13 promoter activation, whereas abnormal expression of USP13 may induce Twist1 to recruit co-factors to promote the transcription of its target genes including USP13."

reach
"USP13 has been shown to upregulate fatty acid synthase (FASN) to promote the proliferation of SCLC cells and Twist-related protein 1 (Twist1) to promote the migration and invasion of breast cancer cells, while it inhibits cell proliferation via downregulation of glycolysis in oral squamous cell carcinoma (47,48)."
USP13 affects PIK3C3
1 | 17 10
USP13 binds PIK3C3.
| 10 10
| 10 10

sparser
"Likewise, we observed that neither the NEDD4H1284A,C1286A nor the NEDD4K1279R mutant reduces the K48-linked ubiquitination of PIK3C3, or interacts with PIK3C3 and USP13."

sparser
"Since previous reports indicated that the knockdown of USP10 led to a reduction in the level of USP13 ( Liu et al., 2011 ), we further examined the interaction between USP13 and VPS34, as well as the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We found that USP13 could still interact with VPS34 and stabilize VPS34 in USP10 KO cells, indicating that USP10 is not required for USP13 to interact and stabilize VPS34 ( Figures S4 B and S4C)."

sparser
"Consistently, knockdown of NEDD4-1 resulted in impaired interaction between VPS34 and USP13 ( Figure S4 D)."

sparser
"As expected, KO of NEDD4-1 completely abrogated the interaction between USP13 and VPS34 ( Figure 4 E)."

sparser
"To confirm whether USP13 could directly bind to VPS34, we performed in vitro His-pull-down assays and found that purified NEDD4-1 could directly promote the association between USP13 and VPS34 ( Figur[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"NEDD4-1 recruits USP13 to form a NEDD4-USP13 complex, thereby increasing the interaction between USP13 and Vps34 and inhibiting K48-linked ubiquitination of Vps34 on K419."

reach
"Interestingly, although knockdown of PIK3C3 does not affect the association between NEDD4 and USP13, the association between USP13 and PIK3C3 is completely abrogated in NEDD4 deficient cells."

reach
"We found that NEDD4-1 markedly enhanced the association between VPS34 and USP13, but not USP10 ( Figures 4 A and S4 A)."

reach
"Since previous reports indicated that the knockdown of USP10 led to a reduction in the level of USP13 ( Liu et al., 2011 ), we further examined the interaction between USP13 and VPS34, as well as the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP13 deubiquitinates PIK3C3.
1 | 3
USP13 deubiquitinates PIK3C3. 4 / 4
1 | 3

reach
"Moreover, spautin-1, is another effective inhibitor, which impedes the initiation step of autophagy, by suppressing the crucial for the process ubiquitin-specific peptidases USP13, USP10, as well as Beclin-1, which is deubiquitinated in Vps34 complex[99]."

reach
"Meanwhile, an in vitro deubiquitination assay showed that USP13 inhibits the K48linked ubiquitination of PIK3C3 in the presence of NEDD4."

reach
"Interestingly, NEDD4-1 undergoes auto-ubiquitination that serves it as a scaffold for engaging the ubiquitin-specific protease 13 (USP13) to form a NEDD4-1/USP13 deubiquitination complex, which subsequently deubiquitinates and stabilizes VPS34 to induce phagophore nucleation in cancer cells."

"Auto-ubiquitination of NEDD4-1 Recruits <span class="match term0">USP13</span> to Facilitate Autophagy through Deubiquitinating <span class="match term1">VPS34</span>"
USP13 inhibits PIK3C3.
| 2
USP13 inhibits PIK3C3. 2 / 2
| 2

reach
"Since USP13 could regulate several substrates in other cancers including MCL1, SKP2, MITF, PTEN, TOPBP1, and c-MYC as described above, ZHX2 could be one of downstream targets for USP13 and other substrates could contribute to the lack of complete rescue in ccRCC, which remains to be further explored.Spautin-1 was previously found to be the inhibitor for both USP10 and USP13 by promoting the degradation of Vps34 (39)."

reach
"USP10 and USP13, two ubiquitin‐specific peptidases, can target the Beclin1 subunit of Vps34‐Beclin1 and promote the degradation of the Vps34 complex."
USP13 activates PIK3C3.
| 2
USP13 activates PIK3C3. 2 / 2
| 2

reach
"Interestingly, Spautin-1-mediated specific and potent inhibition of deubiquitinating activity of USP10 and USP13 was shown to promote proteasomal degradation of PIK3C3 and thereby inhibit autophagy [179]."

reach
"We propose that USP10 and USP13 mediate the deubiqutination of Vps34 complexes to regulate the levels of class III PI3 kinase."
| 28
| 23

reach
"USP13 promotes breast cancer cell migration, invasion and lung metastasis through Twist1."

reach
"These data underscore a key role of Twist1 in USP13-induced enhancement of lung metastasis in breast cancer."

reach
"USP13 promotes OS cells metastasis."

reach
"Notably, USP13 greatly enhanced peritoneal metastasis of ovarian tumors with frequent development of hemorrhagic ascites."

reach
"Taken together, these data indicated that USP13 accelerated OS metastasis in vitro and in vivo."

reach
"Deubiquitinase USP13 promotes the epithelial-mesenchymal transition and metastasis in gastric cancer by maintaining Snail protein."

reach
"Importantly, overexpression of Usp13 accelerates tumorigenesis, enhances tumor metastasis, and causes poor outcomes in transgenic mouse models of ovarian cancer , underscoring its importance in promoting tumorigenesis in vivo."

reach
"In hepatocellular carcinoma, the tumor growth and metastasis are promoted by USP13 through its regulation of TLR4 and subsequent activation of the TLR4/MyD88/NF-κB signaling cascade [23]."

reach
"Snail can be maintained by USP13 to trigger EMT and metastasis, resulting in the poor prognosis of GC patients indirectly [58]."

reach
"USP13 signaling in tumor metastasis."
| 5

reach
"Androgen receptors decrease the expression of USP13 by upregulating miRNA-539-3P, thus increasing the degradation of MITF protein by ubiquitination and promoting melanoma metastasis (85,86)."

reach
"In pulmonary metastatic models, a reduction in pulmonary metastasis caused by USP13 knockdown was abolished in 143B cells with METTL3 overexpression, while silencing METTL3 in HOS cells markedly counteracted the pro-metastatic influence of USP13 upregulation on pulmonary metastasis, as evidenced by histological analysis (Fig. 5f, g)."

reach
"Furthermore, we show in bladder injected-liver metastasis xenograft model that USP13 inhibits bladder cancer metastasis through destabilizing cytoplasmic KDM3A."

reach
"Thus, targeting USP13 would suppress metastasis and progression in RAS -mutant tumors.USP13 has been implicated in cancer development through deubiquitinating both oncogenic and tumor-suppressive prot[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Non-classic deubiquitinase USP13 inhibits bladder cancer metastasis through destabilizing cytoplasmic KDM3A."
STING1 affects USP13
1 | 8 19
1 | 8 19

sparser
"Co-IP results further corroborated the exogenous interaction of USP13 and STING in NIH/3T3 cells co-transfected with Flag–USP13 and His–STING ( xref D)."

sparser
"In contrast, the truncate (aa301-863) that preserves DUB activity and UBA motifs inhibited cGAS and STING-mediated activation of IFN-β promoter and catalysed deubiquitination of STING ( xref ), indicating that USP13-mediated deubiquitination of STING is associated with its suppression of antiviral signalling."

No evidence text available

sparser
"Interestingly, a previous study found that USP13 interacts with STING and promotes the deubiquitination of STING, thereby lessening the antiviral responses by decreasing the activation of STING–TBK1 signaling xref ."

sparser
"Here, we showed the interaction between STING and USP13 in cardiomyocytes during DIC, and the K63-linked deubiquitination of STING by USP13 resulted in the STING degradation via a P62-dependent autophagic way."

sparser
"USP13, a deubiquitinating enzyme, interacts with STING and catalyzed removal of K27 O- or K33 O-linked but not K27 R-linked polyubiquitin chains from STING."

sparser
"Moreover, USP22 significantly promoted the interaction between USP13 and STING ( xref )."

sparser
"Similar to WT USP22, both USP22-C185A and USP22-UCH facilitated the interaction between STING and USP13, but USP22-ZNF failed to do so ( xref )."

sparser
"Here, we perform an unbiased screen by coimmunoprecipitation assays in cells cotransfected with FLAG-tagged DUBs and HA-STING, and find that USP13 interacts with STING."

reach
"Here, we perform an unbiased screen by coimmunoprecipitation assays in cells cotransfected with FLAG tagged DUBs and HA STING, and find that USP13 interacts with STING."
MYC affects USP13
2 | 9 14
MYC binds USP13.
2 | 8 14
2 | 8 10

sparser
"The MYC-USP13 group also significantly reduced PTEN ubiquitination in comparison with control group in HCT116 cells ( xref )."

reach
"We showed a strong molecular association between USP13 and c-MYC, leading to the upregulation of squamous programs in murine and human lung cancer cells."

sparser
"By coexpressing USP13Myc and Bcl‐XL‐Flag in HEK293T cells, we discovered that USP13 could be detected in Bcl‐XL immunoprecipitates after pull down by anti‐Flag beads ( Figure   xref )."

reach
"USP13 and FBXL14 interact with c-Myc in glioma cells."

reach
"However, overexpression of CLK3 in HCCC9810 cells promoted the binding of c-Myc to WT-USP13, not USP13 Y708F mutant (XREF_FIG)."

reach
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag tagged USP13 or FBXL14."

reach
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

reach
"Computational modeling of structures predicted in ZDOCK and Pymol software showed that Y708 phosphorylation was necessary for the direct binding of USP13 to c-Myc (XREF_FIG)."

reach
"As USP13 is a deubiquitinase, the interaction between USP13 and c-Myc led us to hypothesize that USP13 may stabilize c-Myc protein in GSCs through deubiquitination."

No evidence text available
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
MYC activates USP13.
| 1
MYC activates USP13. 1 / 3
| 1

reach
"We found a positive correlation between TRF1 levels and all the stem cell markers with the exception of MYC, although TRF1 was positively correlated with the MYC modulator USP13 (Fang et al., 2017)."
USP13 affects ZHX2
2 | 23 2
USP13 deubiquitinates ZHX2.
| 12
USP13 deubiquitinates ZHX2. 10 / 12
| 12

reach
"USP13 promotes deubiquitination of ZHX2 and tumorigenesis in kidney cancer."

reach
"Interestingly, USP13 has also been implicated in renal cancer, but rather than altering HIF activity, USP13 deubiquitinates another VHL target gene and oncogenic driver, ZHX2 [141]."

reach
"However, a deubiquitinase USP13 inhibits the ubiquitination of ZHX2 and enhances its stability (15)."

reach
"Next, we tried to examine whether USP13 can act as a DUB for ZHX2 and promote ZHX2 deubiquitination and protein stability in ccRCC cells."

reach
"Under denaturing conditions, we found that USP13 WT overexpression led to decreased ZHX2 ubiquitination in ccRCC cell lines (UMRC-2 and 786-O) (Fig. 1M)."

reach
"In these two cells, USP13 depletion by two USP13 short hairpin RNA increased the ZHX2 ubiquitination level (Fig. 2 J and K)."

reach
"In conclusion, our results suggest that USP13 depletion down-regulated the ZHX2 level by increasing ZHX2 ubiquitination and degradation."

reach
"In addition, USP13 promotes tumorigenesis of clear cell renal cell carcinoma through deubiquitinating ZHX2 ."

reach
"19 Similarly, the process of ZHX2 deubiquitination catalyzed by USP13 is the committed step in the tumorigenesis of ccRCC."

reach
"It was found that USP13 could deubiquitinate zinc fingers and homeoboxes 2 (ZHX2) and stabilize ZHX2 protein."
USP13 binds ZHX2.
2 | 2 2
2 | 2 2

No evidence text available

No evidence text available

reach
"Next, we validated semiendogenous interaction between ZHX2 and USP13 by using reciprocal immunoprecipitations (Fig. 1 C and D)."

reach
"With reciprocal immunoprecipitations, we confirmed the endogenous binding between ZHX2 and USP13 in all of these cell lines (Fig. 1 E and F)."

sparser
"Together, this suggests a significant clinical association of ZHX2-USP13 in RCC patients ( P < 0.001) ( xref )."

sparser
"Next, we validated semiendogenous interaction between ZHX2 and USP13 by using reciprocal immunoprecipitations ( xref )."
USP13 increases the amount of ZHX2.
| 5
USP13 increases the amount of ZHX2. 5 / 5
| 5

reach
"Among them, we found that depletion of USP13, but not the other DUBs, down-regulated ZHX2 protein levels in ccRCC cell lines (UMRC-2) (Fig. 1B)."

reach
"First, we overexpressed USP13 in 293T cells and found that USP13 led to increased ZHX2 protein levels in 293T cells or HKC cells while VHL overexpression caused down-regulation of ZHX2 (Fig. 1 G and H), reinforcing the role of pVHL as the E3 ligase for ZHX2 as shown previously (4)."

reach
"WT, but not catalytically inactive USP13 (CD), up-regulated ZHX2 protein levels (Fig.1 I and J), revealing that USP13 overexpression leads to the up-regulation of the ZHX2 protein level in an enzymatically dependent way."

reach
"Accordingly, USP13 depletion by single guide RNAs down-regulated ZHX2 protein levels in 786-O, the phenotype ameliorated by concurrent treatment with the proteasomal inhibitor MG132 (Fig. 2E)."

reach
"In conclusion, our results suggest that USP13 depletion down-regulated the ZHX2 level by increasing ZHX2 ubiquitination and degradation."
USP13 decreases the amount of ZHX2.
| 2
USP13 decreases the amount of ZHX2. 2 / 2
| 2

reach
"In these two cells, USP13 depletion by two USP13 short hairpin RNA increased the ZHX2 ubiquitination level (Fig. 2 J and K)."

reach
"Our results showed that USP13 short hairpin RNA inhibited ZHX2 expression and ccRCC cell growth, while these changes were rescued by the USP13 cDNA (short hairpin RNAs resistant) (SI Appendix, Fig. S4C and Fig. 3 D–G), revealing that the effect of USP13 short hairpin RNA on ZHX2 and soft agar growth was on-target."
USP13 activates ZHX2.
| 2
USP13 activates ZHX2. 2 / 2
| 2

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"USP13 Depletion Leads to Decreased ZHX2 Stability in ccRCC.."

reach
"Thus, the depletion of USP13 decreases ZHX2 and inhibits the proliferation of ccRCC (68)."
| 2 25
| 2 19

reach
"Downregulation of USP13 significantly attenuated the invasiveness of 143B cells, while overexpression of USP13 promoted the invasiveness of HOS cells by using the transwell invasion assay (Fig. 2a, b)."

reach
"USP13 promotes breast cancer cell migration, invasion and lung metastasis through Twist1."

reach
"USP13 promotes melanoma cell invasion by stabilizing the microphthalmia-associated transcription factor."

reach
"We further demonstrated that USP13 obviously promoted OS cells invasion using a 3D tumor spheroid cell-invasion assay (Fig. 2c, d)."

reach
"USP13 knockdown prominently suppressed MGC-803 cell migration and invasion."

eidos
"USP13 knockdown prominently suppressed MGC-803 cell migration and invasion ."

reach
"Next, USP13 knockdown prominently reduced the proliferation, epithelial-mesenchymal transition (EMT), migration, and invasion of Hep3B and Huh7 cells, while USP13 overexpression enhanced these biological behaviors of HepG2 and LO2 cells."

reach
"Although USP13 assumes an antiviral role during the pathogenesis of SARS-CoV-2 (Ravindran et al., 2022), some contend that in specific scenarios, USP13 not only fails to impart antiviral effects but instead stimulates viral invasion."

reach
"High expression of USP13 promotes the migration and invasion of tumors such as breast cancer through Twist1 (79)."

reach
"USP13 promotes migration and invasion of GC cells by maintaining the stability of Snail."

reach
"In addition, USP13 was found to be enriched in the hypoxia response via HIF activation, suggesting that targeting USP13 may suppress hypoxia and thereby promote immune infiltration and the response to immunotherapy in prostate cancer [34]."

reach
"On the other hand, although USP13 arm-level gains and high amplifications significantly increased lymphocyte infiltration in HNSC, OV and STAD, higher USP13 levels were correlated with shorter surviva[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

eidos
"It is also noteworthy to point out that USP13 amplifications did not seem to increase B-cell infiltration in LUSC , which has the opposite effect on disease outcome [ 38 ] ."
| PMC

reach
"It is also noteworthy to point out that USP13 amplifications did not seem to increase B-cell infiltration in LUSC, which has the opposite effect on disease outcome [38]."
| PMC

reach
"On the other hand, although USP13 arm-level gains and high amplifications significantly increased lymphocyte infiltration in HNSC, OV and STAD, higher USP13 levels were correlated with shorter survival in ovarian and gastric cancers."
| PMC

reach
"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

reach
"Downregulation of USP13 significantly attenuated the invasiveness of 143B cells, while overexpression of USP13 promoted the invasiveness of HOS cells by using the transwell invasion assay (Fig. 2a, b)."

reach
"Results suggested that overexpression of NF-kB p65 greatly increased the number of migrated and invaded cells, however knockdown of miR-130b/301b or restoration of USP13 rescued the increased cell invasion and migration caused by NF-kB activation."

reach
"USP13 knockdown in BC cells promoted their proliferation, invasion and migration."

reach
"Overexpression of miR-130b/301b or USP13 knockdown by shRNA delivery decreases USP13 expression, and further reduces PTEN protein expression and leads to enhancement of cell proliferation, invasion and migration."

reach
"Meanwhile, silencing of USP13 greatly enhanced proliferation, invasion and migration of the tested cells."
Cohesin affects USP13
| 1 9 17
| 1 9 17

reach
"USP13 interacts with cohesin and regulates its ubiquitination in human cells."

sparser
"As shown in xref D , the interaction between cohesin and USP13 was enhanced in cells arrested in the S phase when compared with asynchronous cycling cells, or cells arrested at other points in the cell cycle."

sparser
"These data indicate that the interaction of USP13 with cohesin is cell cycle dependent—it is enhanced during DNA replication"

sparser
"We next sought to determine which (if either) of these domains was required for the interaction of USP13 with cohesin."

sparser
"These data demonstrate that the UBA1/2 domains are required for the interaction of USP13 with cohesin and that the ZnF domain is dispensable for the interaction of USP13 with cohesin."

sparser
"We wondered whether ubiquitinated cohesin subunits were present primarily in the soluble fraction (where the USP13cohesin complex resides), in the chromatin fraction, or both."

sparser
"Mass spectrometry analysis revealed that USP13 could interact with Cohesin, and USP13 functioned as both ubiquitinase and deubiquitinase for Cohesin subunits."

sparser
"USP13 and cohesin interact especially during DNA replication, and USP13 regulates the ubiquitination/deubiquitination of the interacting subunit of cohesion through its UBA1/2 domain ( xref )."

sparser
"Interaction of endogenous USP13 with endogenous cohesin in cultured human cells."

sparser
"To further confirm the interaction between USP13 and cohesin, endogenous cohesin was immunoprecipitated with SMC3 antibodies from whole cell lysates prepared from genetically unmodified HCT116 cells and Western blot performed with USP13 antibodies (and STAG2 antibodies as a positive control)."
USP13 affects autophagy
| 3 23
USP13 activates autophagy.
| 3 16
| 3 16

reach
"Spautin-1 inhibits autophagy by reducing the activity of USP10 and USP13, which promotes a major aspect of autophagy: degradation by Vps34 complexes."

eidos
"Our data showed that USP13 knockdown inhibited autophagy , which is consistent with the previous report that USP13 promotes autophagy through deubiquitinating BECN1 to stabilize the protein ( Liu et al ., 2011 ) ."

reach
"USP13 also promotes the expression of autophagy-related proteins by interacting with the Vps34 complex (37,40,41).4."

reach
"Moreover, USP13 promoted autophagy in AML cells."

reach
"USP13 stabilizes the beclin1 subunit of Vps34-beclin1 and promote autophagy."

reach
"USP13 promotes acute myeloid leukemia cell proliferation and autophagy by promoting ATG5."

reach
"USP13 promotes AML cell growth and autophagy by upregulating ATG5."

reach
"A recent study showed that inhibition of USP13 reduced autophagy and ROS accumulation, and attenuated pyroptosis."

reach
"However, these effects were greatly blocked by USP13 overexpression which causes increases in autophagy as well as ferroptosis [21]."

reach
"In all, these results indicated that USP13 promoted autophagy dependent on METTL3 in OS cells (Fig. 7)."
USP13 inhibits autophagy.
| 7
| 7

reach
"Because the reductions in the levels of USP10 and USP13 in H4-LC3-GFP cells treated with spautin-1 appeared later than the reductions in the levels of Vps34 complexes and autophagy (XREF_FIG), the reduced levels of USP10 and USP13 are unlikely to be the primary reason for the ability of spautin-1 to reduce the levels of PtdIns3P and inhibit autophagy."

reach
"As expected, USP13 overexpression inhibited autophagy and survival of PC12 cells under OGD/R conditions, and Spautin‐1 treatment counteracted these effects."

reach
"ROR1, after binding to Wnt ligands, triggers non-canonical signaling cascades that increase the level of calcium or decrease the level of cGMP within the cell, and these changes are closely associated with COPD onset and progression.51 SEC14L4 is known to be differentially expressed in COPD.52 USP13, which inhibits autophagy, may be involved in COPD pathogenesis.53,54Among the 11 feature genes, RS1 has not previously been linked to COPD, and our study justifies further exploration of this potential link."

reach
"Taken together, overexpression of METTL3 significantly recovered the formation of autophagosome and inhibition of autophagy flux caused by depletion of USP13 (Fig. 7a-f)."

reach
"Spautin-1 inhibits deubiquitinating enzymes USP10 and USP13 and promotes degradation of the BECN1-PI3K autophagy initiation complex, which is required for canonical autophagy induction [30]."

reach
"Interestingly, Spautin-1-mediated specific and potent inhibition of deubiquitinating activity of USP10 and USP13 was shown to promote proteasomal degradation of PIK3C3 and thereby inhibit autophagy [179]."

reach
"The function of USP10 and USP13 is reversed by a chemical compound, spautin-1, which inhibits autophagy by promoting Beclin-1 degradation."
USP13 affects METTL3
3 | 19 4
USP13 binds METTL3.
3 | 3 4
3 | 3 4

No evidence text available

reach
"Taken together, our results confirm that USP13 interacts with METTL3.Since USP13 binds to METTL3, we then explored the effect of downregulating or upregulating USP13 on METTL3 stability in OS cells."

sparser
"In this study, we found that USP13 interacts with METTL3 and decreases METTL3 ubiquitination, resulting in reduced METTL3 degradation in OS."

sparser
"Moreover, the interaction between USP13 and METTL3 was verified by Co-IP analysis in HEK 293T cells with exogenously transfection of Flag-tagged USP13 and Myc-tagged METTL3 (Fig. xref c)."

sparser
"Taken together, our results confirm that USP13 interacts with METTL3."

sparser
"Since USP13 binds to METTL3, we then explored the effect of downregulating or upregulating USP13 on METTL3 stability in OS cells."

No evidence text available

No evidence text available

reach
"USP13 interacts with and stabilizes METTL3."

reach
"Moreover, the interaction between USP13 and METTL3 was verified by Co-IP analysis in HEK 293T cells with exogenously transfection of Flag-tagged USP13 and Myc-tagged METTL3 (Fig. 3c)."
USP13 deubiquitinates METTL3.
| 8
USP13 leads to the deubiquitination of METTL3. 5 / 5
| 5

reach
"USP13 was shown to positively promote glycolysis and tumor progression by stabilizing and deubiquitinating METTL3, which is a well-known “writer” for m A modification, at K488 by removing K48-linked ubiquitin chains."

reach
"In this study, we found that USP13 interacts with METTL3 and decreases METTL3 ubiquitination, resulting in reduced METTL3 degradation in OS."

reach
"Depletion of USP13 significantly promoted METTL3 ubiquitination level compared with shNC in 143B cells (Fig. 4a)."

reach
"In contrast, overexpression of USP13 significantly decreased ubiquitination of METTL3 in HOS cells (Fig. S4a)."

reach
"Overexpression of WT USP13 decreased METTL3 ubiquitination while C345A mutant USP13 showed no significant effect (Fig. 4b)."
USP13 deubiquitinates METTL3 on K488. 3 / 3
| 3

reach
"Ubiquitin-specific proteases 13 (USP13) regulates glycolytic reprogramming and proliferation in osteosarcoma by deubiquitinating METTL3 at K488."

reach
"Our results indicated that USP13 promotes glycolytic reprogramming and progression in OS by stabilizing as well as deubiquitinating METTL3 protein at K488 by removing K48-linked ubiquitin chains, which further stabilizes ATG5 mRNA and activates oncogenic autophagy."

reach
"USP13 deubiquitinates METTL3 at K488 by removing K48-linked ubiquitin chains."
USP13 activates METTL3.
| 6
USP13 activates METTL3. 6 / 6
| 6

reach
"Collectively, our study results indicate that USP13 promotes glycolysis and tumor progression in OS by stabilizing METTL3, thereby stabilizing ATG5 mRNA and facilitating autophagy in OS."

reach
"Silencing of USP13 significantly accelerated METTL3 degradation while upregulation of USP13 obviously inhibited METTL3 degradation (Fig. 3h, i)."

reach
"Wang et al. reveal that injecting Spautin-1 into mice bearing xenografts effectively inhibits the USP13-enhanced stabilization of METTL3, subsequently reducing the stability of ATG5 mRNA and thereby curbing tumor proliferation and metastasis 130."

reach
"Ectopically expressed K488R of METTL3 significantly promoted cell proliferation and glycolysis, while USP13 knockdown decreased cell proliferation and glycolysis with WT METTL3 rather than K488R METTL3, which indicated that mutation of K488 of METTL3 promoted cell proliferation and glycolysis and abolished USP13 depletion-induced proliferation and glycolysis suppression (Fig. 4i-n)."

reach
"Collectively, our results demonstrated that pharmacological inhibition of USP13 effectively inhibit the malignancy of OS by promoting METTL3 destabilization."

reach
"USP13 promotes glycolysis and progression in OS by binding to and stabilizing METTL3."
USP13 inhibits METTL3.
| 2
USP13 inhibits METTL3. 2 / 2
| 2

reach
"Using a small-molecule inhibitor named Spautin-1 to pharmacologically inhibit USP13 induced METTL3 degradation and exhibited significant therapeutic efficacy both in vitro and in vivo."

reach
"Knockdown or overexpression of USP13 markedly decreased or increased METTL3 protein but not mRNA levels (Fig. 3e, S3)."
USP13 affects AMFR
3 | 6 13
USP13 binds AMFR.
3 | 4 13
3 | 4 11

sparser
"It appears that the second UBA domain is crucial for USP13 to bind gp78."

sparser
"The requirement of gp78 interaction for ERAD inhibition by ectopic USP13 indicates that the interaction of USP13 with gp78 is functionally relevant to ERAD."

No evidence text available

No evidence text available

No evidence text available

sparser
"Importantly, USP13 associates with AMFR to prevent polyubiquitylation of UBL4A, protecting the BAG6 complex from proteasomal degradation and maintaining proper ERAD pathway function [ xref ]."

sparser
"Thus, the association of USP13 with AMFR prevents collateral damage in the ERAD pathway by promoting the distinction between bona fide ERAD substrates and protein complexes that ERAD pathway E3s must engage to ensure proper substrate shuttling for degradation."

sparser
"Interestingly, USP13 associates with gp78 to attenuate polyubiquitination of ubiquitin-like protein 4A (UBL4A), a part of the Bag6 complex that helps chaperone retrotranslocated ERAD substrates to the proteasome, thereby protecting the Bag6 complex from proteasomal degradation and maintaining an adequate ERAD pathway, highlighting the importance of the associated DUB activity during ERAD [ xref ]."

sparser
"Recently, cleaved Bag6 induced by caspase three activation was reported to interact with LC3-Ⅰ and pro-LC3 under ER stress conditions such as gp78 overexpression and USP13 downregulation, leading to autophagy inhibition and apoptosis activation ( xref )."

reach
"USP13 interacts directly with gp78 in ERAD."
USP13 binds AMFR and BAG6. 2 / 2
| 2

sparser
"USP13 forms a multiprotein complex with gp78 and Bag6."

sparser
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."
USP13 inhibits AMFR.
| 2
USP13 inhibits AMFR. 2 / 5
| 2

reach
"Because the accumulation of Bag6 * upon USP13 depletion is significantly reduced in cells co-depleted of USP13 and gp78 (XREF_FIG), we propose that USP13 is required to antagonize a promiscuous activity of gp78 towards Ubl4A, which would otherwise impair the function of the Bag6 complex by altering its interaction pattern and/or increasing its cleavage by a cellular protease."

reach
"USP13 antagonizes gp78 by removing ubiquitin."
TWIST1 affects USP13
4 | 14 8
TWIST1 binds USP13.
4 | 4 8
4 | 4 8

sparser
"Here, we investigated a potential physical/functional interaction between Twist1 and USP13 in human breast cancer."

sparser
"USP13 directly interacts with Twist1."

reach
"Although the biochemical and molecular functions of USP13 have been explored in various human cancers, including breast cancer, the oncogenic or tumor suppressor roles of USP13 in different studies has remained controversial.Here, we investigated a potential physical/functional interaction between Twist1 and USP13 in human breast cancer."

reach
"In accordance with the observed association between USP13 and Twist1, we detected an efficient co-precipitation between endogenous USP13 and Twist1 in SUM159PT and MDA-MB231 cells (Fig. 2a)."

sparser
"We found that USP13 and Twist1 can form a negative feedback loop to jointly regulate the migration and invasion of breast cancer cells."

sparser
"We also found that USP13 can directly interact with Twist1 and specifically cleave the K48-linked polyubiquitin chains of Twist1 induced by FBXL14."

sparser
"The interaction between USP13 and Twist1 was further confirmed by IF analysis since we found that USP13 co-localizes with Twist1 in the nuclei of MDA-MB231 cells (Fig.  xref )."

reach
"The interaction between USP13 and Twist1 was further confirmed by IF analysis since we found that USP13 co-localizes with Twist1 in the nuclei of MDA-MB231 cells (Fig. 2d)."

reach
"Therefore, the above findings indicate a direct interaction between USP13 and Twist1."

sparser
"For instance, USP13 directly interacts with Twist1 and cleaves FBXL14-induced K48-linked polyubiquitin chains, increasing Twist1 protein levels and facilitating BC cell migration and tumor metastasis in vitro and in vivo xref ."
TWIST1 inhibits USP13.
| 4
TWIST1 inhibits USP13. 4 / 4
| 4

reach
"We found that the effect of USP13 in promoting the migration and invasion capacities of breast cancer cells can at least partly be achieved through its regulation of Twist1, while Twist1 can inhibit the transcriptional activity of USP13."

reach
"However, this finding is contradictory to the notion that Twist1 inhibits USP13 promoter activation."

reach
"In addition, we found that Twist1 could inhibit the activity of the USP13 promoter in a dose-dependent manner (Fig. 7c), and directly bind to the USP13 promoter in SUM159PT cells as demonstrated by ChIP (Fig. 7d)."

reach
"A possible, but one-sided explanation may be that normal physiological concentrations of Twist1 individually inhibit USP13 promoter activation, whereas abnormal expression of USP13 may induce Twist1 to recruit co-factors to promote the transcription of its target genes including USP13."
TWIST1 increases the amount of USP13.
| 3
TWIST1 increases the amount of USP13. 3 / 3
| 3

reach
"Twist1 inhibits USP13 promoter transcription."

reach
"In addition, we found that the protein levels of Twist1 dose-dependently increased with USP13 overexpression, while USP13 knockdown resulted in a decreased expression of endogenous Twist1."

reach
"These results underscore a key role of Twist1 in a USP13-induced enhancement of migration and invasion capacities of breast cancer cells.We also assessed the effects of USP13 expression on lung metastasis of breast cancer to further substantiate the above in vitro findings."
TWIST1 decreases the amount of USP13.
| 3
TWIST1 decreases the amount of USP13. 3 / 3
| 3

reach
"Twist1 inhibits USP13 promoter transcription."

reach
"Drawn from the combined results from the ChIP and luciferase reporter assays, we found that Twist1 can inhibit the transcription of USP13 by directly binding to these three E-box regions."

reach
"Twist1 overexpression decreased USP13 mRNA levels in T47D and ZR-75–30 cells, whereas Twist1 knockdown increased USP13 mRNA levels in SUM159PT and MDA-MB231 cells (Fig. 7e)."
CLK3 affects USP13
1 2 4 | 9 10
CLK3 phosphorylates USP13.
1 1 | 6 8
CLK3 phosphorylates USP13 on Y708. 10 / 16
1 1 | 6 8

sparser
"Therefore, we propose that CLK3-induced Y708 phosphorylation of USP13 promotes CCA progression by activating c-Myc–mediated purine synthesis."

reach
"In cholangiocarcinoma, TGF-β signaling triggers the phosphorylation of CLK3, a serine/threonine kinase that directly phosphorylates USP13 at Y708 and facilitates USP13 interaction with c-Myc (Zhou et al., 2020); in GSCs, USP13 can enhance the stability through deubiquitinating c-Myc, activating purine synthesis mediated by c-Myc and inducing the tumorigenesis of GSCs (Fang et al., 2017); in hepatocellular carcinoma, knockdown of USP13 by shRNA can markedly downregulate c-Myc expression, resisting xenograft tumor growth of HCC (Huang et al., 2020)."

sparser
"We demonstrated that (1) a recurrent Q607R somatic substitution in CLK3 was identified in 8% of 100 human CCAs, particularly in patients with CCA metastasis; (2) the expression of CLK3 was significantly up-regulated in CCA compared with matched control tissues; (3) CLK3 knockdown significantly inhibited CCA aggressiveness in vitro and in vivo; (4) gene ontology term enrichment and MS assays indicated that high CLK3 expression in CCA patients mainly regulated nucleotide metabolism, especially purine biosynthesis; (5) mechanistically, CLK3 directly phosphorylated USP13 at Y708, which promoted its binding to c-Myc, a critical purine synthesis–associated transcription factor, thereby preventing Fbxl14-mediated c-Myc ubiquitination and activating the transcription of purine metabolic genes; (6) the CCA-associated CLK3-Q607R mutant induced USP13-Y708 phosphorylation and enhanced the activity of c-Myc; (7) in turn, c-Myc transcriptionally up-regulated CLK3; (8) importantly, levels of CLK3 significantly correlated with the expression of phospho-USP13-Y708, c-Myc, and ATIC in human CCA specimens; and (9) tacrine hydrochloride was identified as a potential compound to inhibit the aberrant CLK3-enhanced CCA invasiveness."

sparser
"Moreover, a recent study also found that CDC-like kinase 3 (CLK3) or the cholangiocarcinoma-associated CLK3-Q607R mutant can directly phosphorylate USP13 at Tyr708, and promote its binding to c-Myc ( xref ; xref ) ( xref )."

reach
"USP13 stabilizes its substrate, and at the same time, the activity of USP13 is upregulated by CLK3 via promoting phosphorylation of USP13 at Tyr708, which increases its deubiquitinating enzyme activity."

sparser
"CLK3 directly phosphorylated USP13 at Y708, which promoted its binding to c-Myc, thereby preventing Fbxl14-mediated c-Myc ubiquitination and activating the transcription of purine metabolic genes."

sparser
"The phosphorylation of USP13 at Tyrosine 708 by CLK3 is required for USP13′s binding to c-Myc, which prevents its Fbxl14-mediated ubiquitination of c-Myc in glioma stem cells [ xref ]."

No evidence text available

sparser
"Notably, the CCA-associated CLK3-Q607R mutant induced USP13-Y708 phosphorylation and enhanced the activity of c-Myc."

reach
"CLK3 mediated the phosphorylation of USP13 at Y708, promoting its binding to c-Myc, which is an important transcription factor and also known as an oncogene in many cancers [71]."
CLK3 binds USP13.
4 | 1 2
4 | 1 2

No evidence text available

sparser
"CLK3 interacts with USP13, phosphorylates it at the Tyr708 residue and inhibits the ubiquitination of c-Myc to activate c-Myc-mediated purine biosynthesis, thus promoting the progression of bile duct cell carcinoma ( xref )."

reach
"A pulldown experiment suggests a direct binding of USP13 to CLK3 (XREF_FIG)."

No evidence text available

No evidence text available

sparser
"A pulldown experiment suggests a direct binding of USP13 to CLK3 ( xref )."

No evidence text available
CLK3 activates USP13.
1 | 2
CLK3 activates USP13. 3 / 3
1 | 2

reach
"Moreover, CLK3 specifically has been shown to activate the ubiquitin-specific peptidase 13 (USP13) through phosphorylation [66], and although they did not further investigate it, coIP-MS experiments published by Funnel et al. showed that the protein most significantly associated with CLK2 was no splicing factor, but USP7 [49]."

reach
"XREF_FIG indicates that CLK3 knockdown in HuCCT1 cells strongly impaired the interaction of USP13 with c-Myc."

"CLK3 directly phosphorylated USP13 at Y708, which promoted its binding to c-Myc, thereby preventing Fbxl14-mediated c-Myc ubiquitination and activating the transcription of purine metabolic genes."
USP13 affects OGDH
2 1 | 15 3
USP13 deubiquitinates OGDH.
1 | 7
USP13 deubiquitinates OGDH. 8 / 8
1 | 7

reach
"Mechanistically, USP13 promoted the energy metabolism of tumor cells, and provided precursor substances for the synthesis of sugar, lipids and non-essential amino acids in cancer cells, through deubiquitinating and stabilizing ACLY and OGDH (Han et al., 2016)."

reach
"Furthermore, USP13 can deubiquitinate and stabilise ACLY and OGDH and c-Myc in ovarian cancer and glioblastoma, respectively, thereby functioning as an oncogene [XREF_BIBR, XREF_BIBR]."

reach
"As it was aforementioned, USP13 deubiquitinates OGDH and ACLY."

"<span class="match term0">USP13</span>, the main regulator of ovarian cancer energy metabolism, specifically deubiquitinates and stabilizes oxoglutarate dehydrogenase and ATP citrate lyase, which can catalyze fatty acid synthesis, glutaminolysis and mitochondrial respiration."

reach
"Taken together, these studies suggest that USP14 is involved in the occurrence and progression of multiple malignant tumors.Other USP family members: USP13, the main regulator of ovarian cancer energy metabolism, specifically deubiquitinates and stabilizes oxoglutarate dehydrogenase and ATP citrate lyase, which can catalyze fatty acid synthesis, glutaminolysis and mitochondrial respiration."

reach
"USP13 influences cell cycle progression in gastric cancer by stabilizing cyclin D1 [15] and enhances the deubiquitination and stabilization of ATP citrate lyase (ACLY) and oxoglutarate dehydrogenase (OGDH) in ovarian tumors [16]."

reach
"USP13 directly deubiquitinates ACLY and OGDH."

reach
"In ovarian tumors, upregulation of USP13 enhances deubiquitination and stabilization of ACLY (ATP citrate lyase) and OGDH (oxoglutarate dehydrogenase), two key enzymes that drive glutaminolysis, ATP generation, and lipid synthesis in cancer metabolism , and MCL1, a pivotal member of the antiapoptotic BCL-2 family of proteins ."
USP13 activates OGDH.
| 4
USP13 activates OGDH. 4 / 6
| 4

reach
"USP13 led to the upregulation of ACLY and oxoglutarate dehydrogenase (OGDH), which is one of the subunit of α-ketoglutarate dehydrogenase complex, an enzymatic complex involved in glutaminolysis."

reach
"USP13 primarily deubiquitinates and therefore upregulates ATP citrate lyase and oxoglutarate dehydrogenase, two crucial enzymes that control glutaminolysis and fatty acid synthesis and also mitochondrial respiration."

reach
"USP13 upregulates ACLY and OGDH, two key regulators that drive glutaminolysis, ATP generation and lipid synthesis in cancer metabolism."

reach
"Next, we examined whether USP13 augments the stability of ACLY and OGDH."
USP13 binds OGDH.
2 | 3
2 | 1

No evidence text available

sparser
"The N terminus including a ubiquitin-specific protease-type zinc-finger domain (residues 1–336) was essential for USP13 to interact with OGDH, whereas the ubiquitin carboxyl-terminal hydrolase domain (residues 336–863) of USP13 is required for its binding with ACLY ( xref )."

No evidence text available
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."
USP13 inhibits OGDH.
| 2
USP13 inhibits OGDH. 2 / 3
| 2

reach
"These data suggest that USP13-KD induces mitochondrial dysfunction and lipogenic dysfunction by reducing OGDH and ACLY activity, thereby reducing glutamine 's reductive carboxylation and glucose 's oxidation for lipid synthesis (XREF_FIG)."

reach
"Depletion of USP13 markedly decreased protein stability of ACLY and OGDH (XREF_FIG; XREF_SUPPLEMENTARY), but had no effect on the transcription of ACLY and OGDH (XREF_FIG)."
USP13 decreases the amount of OGDH.
| 2
USP13 decreases the amount of OGDH. 2 / 3
| 2

reach
"Consistently, tissue microarray detected that USP13 knockdown could reduce the ACLY and OGDH protein levels in cells, reducing the synthesis of fatty acids from the source."

reach
"We proposed that KD of USP13 reduces OGDH expression thereby limiting the synthesis of TCA cycle intermediates from oxidative pathways (XREF_FIG)."
PIK3C3 affects USP13
| 15 10
PIK3C3 binds USP13.
| 10 10
| 10 10

sparser
"Likewise, we observed that neither the NEDD4H1284A,C1286A nor the NEDD4K1279R mutant reduces the K48-linked ubiquitination of PIK3C3, or interacts with PIK3C3 and USP13."

sparser
"Since previous reports indicated that the knockdown of USP10 led to a reduction in the level of USP13 ( Liu et al., 2011 ), we further examined the interaction between USP13 and VPS34, as well as the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We found that USP13 could still interact with VPS34 and stabilize VPS34 in USP10 KO cells, indicating that USP10 is not required for USP13 to interact and stabilize VPS34 ( Figures S4 B and S4C)."

sparser
"Consistently, knockdown of NEDD4-1 resulted in impaired interaction between VPS34 and USP13 ( Figure S4 D)."

sparser
"As expected, KO of NEDD4-1 completely abrogated the interaction between USP13 and VPS34 ( Figure 4 E)."

sparser
"To confirm whether USP13 could directly bind to VPS34, we performed in vitro His-pull-down assays and found that purified NEDD4-1 could directly promote the association between USP13 and VPS34 ( Figur[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"NEDD4-1 recruits USP13 to form a NEDD4-USP13 complex, thereby increasing the interaction between USP13 and Vps34 and inhibiting K48-linked ubiquitination of Vps34 on K419."

reach
"Interestingly, although knockdown of PIK3C3 does not affect the association between NEDD4 and USP13, the association between USP13 and PIK3C3 is completely abrogated in NEDD4 deficient cells."

reach
"We found that NEDD4-1 markedly enhanced the association between VPS34 and USP13, but not USP10 ( Figures 4 A and S4 A)."

reach
"Since previous reports indicated that the knockdown of USP10 led to a reduction in the level of USP13 ( Liu et al., 2011 ), we further examined the interaction between USP13 and VPS34, as well as the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
PIK3C3 inhibits USP13.
| 3
PIK3C3 inhibits USP13. 3 / 3
| 3

reach
"It is interesting to note that the knock-down of VPS34 and BECLIN 1 also causes the degradation of USP10 and USP13, which indicates the existence of a regulatory feedback loop [XREF_BIBR]."

reach
"Interestingly, Vps34 acts as a regulator for autophagy, and Vps34 complexes down-regulate USP10 and USP13 by Beclin1, suggesting that they are downstream of autophagy activation ( Liu et al., 2011 )."

reach
"Using an imaging-based screen, Liu et al. (2011) recently identified a highly potent small molecule inhibitor of autophagy they named spautin-1 (specific and potent autophagy inhibitor 1), which promotes degradation of the Vps34 complexes via inhibiting ubiquitin-specific processing protease 10 (USP10) and USP13, two ubiquitin-specific peptidases that target the deubiquitination of Beclin1.2."
PIK3C3 activates USP13.
| 2
PIK3C3 activates USP13. 2 / 2
| 2

reach
"The close functional interconnection between the central PI3K-III subunits and the DUBs is demonstrated by the interesting observation that knock-down of VPS34 and Beclin 1 causes instability of USP10 and USP13, which strongly indicates the existence of a regulatory feedback loop [XREF_BIBR]."

reach
"Another interesting observation made in the Spautin-1 study is that knock-down of VPS34 and Beclin 1 causes instability of USP10 and USP13, which indicates the existence of a regulatory feedback loop [XREF_BIBR]."
USP13 affects WWP1
| 16 8
USP13 binds WWP1.
| 5 7
| 5 7

reach
"Immunoprecipitation with WWP1 antibody further validated the interaction between WWP1 and USP13 (Fig. 3B)."

reach
"Next, we transfected Myc-tagged WWP1 (Myc-WWP1), along with Flag-tagged USP13 (Flag-USP13) or an empty vector, into HCC and HEK293T cells, followed by a Co-IP assay to validate the binding between USP13 and WWP1."

reach
"Overall, there was an interaction between USP13 and WWP1."

reach
"Since our findings described above demonstrated the interaction between USP13 and WWP1, we wondered whether USP13 regulated WWP1 through deubiquitination."

reach
"According to our results, USP13 truncations containing either the N-terminal region (aa1–300 and aa1–624) or the C-terminal region (aa301–863 and aa625–863) could bind to WWP1."

sparser
"Finally, our functional experiments in HCC cells and a subcutaneous xenograft model further revealed the oncogenic role of the USP13-WWP1 axis both in vitro and in vivo."

sparser
"Furthermore, USP13 can interact with WWP1 and then remove the K29- and K48-linked polyubiquitination chains from WWP1 to stabilize the WWP1 protein via the ubiquitin–proteasome pathway."

sparser
"USP13 interacted with WWP1 in HCC cells."

sparser
"Immunoprecipitation with WWP1 antibody further validated the interaction between WWP1 and USP13 (Fig.  xref B)."

sparser
"To determine the specific region of USP13 that interacts with WWP1, we constructed five Flag-tagged USP13 truncation plasmids."
USP13 deubiquitinates WWP1.
| 5
USP13 deubiquitinates WWP1. 5 / 5
| 5

reach
"YY1 induced USP13 transcriptional activation drives the malignant progression of hepatocellular carcinoma by deubiquitinating WWP1."

reach
"USP13 deubiquitinated and regulated WWP1 in a proteasomal-dependent way.."

reach
"As shown in Fig. 4I, the HA-Ub mutants K27R, K48R, and K63R attenuated the deubiquitination of WWP1 by USP13."

reach
"By acting as a DUB, USP13 can bind to and then deubiquitinate and stabilize the WWP1 protein, thus activating the Akt/mTOR pathway."

reach
"Here, we revealed that USP13, which acts as a DUB, deubiquitinates WWP1, which was identified as a new substrate of USP13."
USP13 increases the amount of WWP1.
| 3 1
USP13 increases the amount of WWP1. 3 / 3
| 2 1

reach
"The results revealed that USP13 knockdown significantly reduced the protein levels of WWP1, p-Akt, and p-mTOR, which was reversed by WWP1 overexpression (Fig. 5A)."

sparser
"Conversely, we also transfected WWP1 siRNA or negative control siRNA into HCC cells overexpressing USP13 and found that WWP1 knockdown attenuated USP13-induced WWP1 expression and Akt/mTOR pathway activation (Fig.  xref B)."

reach
"Moreover, USP13 increased the protein level (Fig. 4B) but not the RNA level of WWP1 (Supplementary Fig. S3B and S3D)."
USP13 increases the amount of ubiquitinated WWP1. 1 / 1
| 1

reach
"The results indicated that USP13 knockdown increased, while USP13 overexpression reduced, the level of ubiquitinated WWP1 protein (Fig. 4C, D)."
USP13 activates WWP1.
| 3
USP13 activates WWP1. 3 / 3
| 3

reach
"At the same time, the detection of various ubiquitination-related proteins, including PTEN ubiquitination inhibitory proteins USP7 [36] and USP13 [37] and PTEN ubiquitination-promoting protein NEDD4-1 [38], WWP1 [34], WWP2 [39] and FBXO22 [35], showed that only WWP1 and FBXO22 were significantly down-regulated, while others were not significantly changed (Fig. 4e)."

reach
"In contrast, WWP1 was elevated in the USP13 group, and I3C suppressed the upregulation of WWP1 mediated by USP13 overexpression (Fig. 6L)."

reach
"Our results indicated that MG132 could restrain the degradation of WWP1 induced by USP13 knockdown, whereas HCQ could not (Fig. 4E)."
USP13 affects USP10
1 | 14 6
USP13 binds USP10.
| 4 6
| 4 4

sparser
"USP13 also interacts with USP10, and the interaction increases USP13 deubiquitinating activity ( xref )."

sparser
"In addition, whether and how the complex relationship between PTEN DUBs (e.g., HAUSP−USP10 xref , HAUSP−USP11 xref and USP10USP13 xref ) strengthens their activity toward PTEN requires further study."

reach
"USP13 also interacts with USP10, and the interaction increases USP13 deubiquitinating activity."

sparser
"Although no structural information for USP13 and USP10 is currently available, the enhanced DUB activity when USP13 interacts with USP10 or when they interact with Beclin1 suggest that the interaction of USP13 and USP10 with each other or with their substrates can lead to changes in their conformation which may be critical for the catalytic activities."

reach
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

reach
"Using spautin-1 as a tool, we explored the interaction of USP10 and USP13 with Vps34 complexes."

sparser
"No orthogonal biophysical methods, such as the surface plasmon response (SPR) or isothermal titration calorimetry (ITC), were used to validate the binding of spautin-1 with USP10 or USP13."

reach
"Although no structural information for USP13 and USP10 is currently available, the enhanced DUB activity when USP13 interacts with USP10 or when they interact with Beclin1 suggest that the interaction of USP13 and USP10 with each other or with their substrates can lead to changes in their conformation which may be critical for the catalytic activities."
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
USP13 deubiquitinates USP10.
1 | 8
USP13 deubiquitinates USP10. 9 / 9
1 | 8

reach
"As previously mentioned, USP10 mediates the deubiquitination of Beclin1, and USP13 can directly regulate the deubiquitination of USP10 to promote the formation of autophagosomes ."

reach
"A previous study indicated that USP13 affects p53 by deubiquitinating USP10 [33]."

No evidence text available

reach
"Consistent with this possibility, the ubiquitination levels of USP10 were reduced when cells were cotransfected with an expression vector of USP13 and the addition of spautin-1 inhibited the deubiquitination of USP10 by USP13 (XREF_FIG)."

reach
"These results suggest that USP13 may directly regulate the deubiquitination of USP10; however, USP10 may regulate USP13 indirectly perhaps by affecting the levels of Vps34 complexes."

reach
"Moreover, beclin-1 and USP10 are involved in a potential feedforward mechanism in which beclin-1 stabilizes USP13, which in turn deubiquitinates and stabilizes USP10, leading to increased beclin-1 levels and activity."

reach
"Since USP13 can also deubiquitinate USP10, regulating the stability of USP13 by Beclin1 provides a mechanism for Beclin1 to control the stability of USP10."

reach
"Very interestingly, they also pointed out that beclin-1 controlled the stability of USP10 by regulating the stability of USP13, which can deubiquitinate USP10 (also see Figure 4) [53]."

reach
"USP13 has been found to stabilize p53, a typical tumor suppressor by de-ubiquitinating its de-ubiquitinating protease USP10 [24]."
USP13 inhibits USP10.
| 2
USP13 inhibits USP10. 2 / 2
| 2

reach
"It will be interesting to elucidate if the stability of the other PI3K and III complex members is also affected, as in the case of Spautin-1 -- mediated inhibition of USP10 and USP13 [XREF_BIBR] and if the control of Beclin 1 stability is mediated by HSP90 in general or whether this represents a regulatory event specifically developed in phagocytic cells."

reach
"USP13 further modulates this network by regulating USP10 degradation, as demonstrated in the heart, lung and liver of newborn mice [104]."
AMFR affects USP13
3 | 5 13
AMFR binds USP13.
3 | 4 13
3 | 4 11

sparser
"It appears that the second UBA domain is crucial for USP13 to bind gp78."

sparser
"The requirement of gp78 interaction for ERAD inhibition by ectopic USP13 indicates that the interaction of USP13 with gp78 is functionally relevant to ERAD."

No evidence text available

No evidence text available

No evidence text available

sparser
"Importantly, USP13 associates with AMFR to prevent polyubiquitylation of UBL4A, protecting the BAG6 complex from proteasomal degradation and maintaining proper ERAD pathway function [ xref ]."

sparser
"Thus, the association of USP13 with AMFR prevents collateral damage in the ERAD pathway by promoting the distinction between bona fide ERAD substrates and protein complexes that ERAD pathway E3s must engage to ensure proper substrate shuttling for degradation."

sparser
"Interestingly, USP13 associates with gp78 to attenuate polyubiquitination of ubiquitin-like protein 4A (UBL4A), a part of the Bag6 complex that helps chaperone retrotranslocated ERAD substrates to the proteasome, thereby protecting the Bag6 complex from proteasomal degradation and maintaining an adequate ERAD pathway, highlighting the importance of the associated DUB activity during ERAD [ xref ]."

sparser
"Recently, cleaved Bag6 induced by caspase three activation was reported to interact with LC3-Ⅰ and pro-LC3 under ER stress conditions such as gp78 overexpression and USP13 downregulation, leading to autophagy inhibition and apoptosis activation ( xref )."

reach
"USP13 interacts directly with gp78 in ERAD."
USP13 binds AMFR and BAG6. 2 / 2
| 2

sparser
"USP13 forms a multiprotein complex with gp78 and Bag6."

sparser
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."
AMFR activates USP13.
| 1
AMFR activates USP13. 1 / 2
| 1

reach
"Since purified gp78 is insufficient to activate USP13 (Liu, Y unpublished results), further experiments using gp78 containing retrotranslocation complexes reconstituted in proteoliposome are required to elucidate how USP13 is regulated in the context of retrotranslocation."
Ubiquitin affects USP13
| 4 10
Ubiquitin binds USP13.
| 1 9
| 1 6

sparser
"The GST pull-down experiment indicates that full-length USP13 can bind with Ub, but not with ISG15, SUMO1 and SUMO2 ( xref )."

sparser
"Although the experimental results showed that USP13 could bind ubiquitin, bacterial-purified recombinant USP13 still exhibited only weak deubiquitination enzyme activity [ xref , xref , xref ]."

sparser
"The ZnF domain of USP13 cannot bind with ubiquitin, so that USP13 loses its ability to be activated by free ubiquitin ( xref )."

sparser
"We firstly examined whether the ZnF domain of USP13 (USP13-ZnF) interacts with Ub by isothermal titration calorimetry (ITC) experiment ( xref )."

reach
"In addition, NMR titration data reflected that M664, F666, M739, and F741 might be the key residues responsible for the binding of USP13-UBA to Ub (Zhang et al., 2011)."

sparser
"Overall, the USP13-ZnF domain cannot bind to Ub to activate USP13, whereas USP13-UBA can bind, which may partially explain the reason why USP13 displays only weak basal deubiquitination enzyme activity: the binding sites of USP13 to Ub are less than that other USP members, thus providing weaker binding affinity and consequent cleavage activity towards ubiquitin chains; There possibly exists constitutive self-inhibition for full-length USP13 supported by the interaction of UBA with ZnF domain, which is hypothesized to be released by recruitment of other proteins or modification, such as phosphorylation."

sparser
"The ZnF domain of USP13 cannot bind with Ub."
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
Ubiquitin activates USP13.
| 3 1
| 3 1

reach
"It was shown that transfection of Lys48R ubiquitin reversed the influence of silencing of USP13 in decreasing IκBα protein level (Fig. 6h)."

sparser
"Since the ZnF domain of USP13 is not the Ub receptor, it is crucial to examine whether USP13 could be catalytically activated by Ub."

reach
"Since the ZnF domain of USP13 is not the Ub receptor, it is crucial to examine whether USP13 could be catalytically activated by Ub."

reach
"Domain swapping confers USP13 the ability to be activated by Ub."
USP13 affects Ubiquitin
| 5 9
USP13 binds Ubiquitin.
| 1 9
| 1 6

sparser
"The GST pull-down experiment indicates that full-length USP13 can bind with Ub, but not with ISG15, SUMO1 and SUMO2 ( xref )."

sparser
"Although the experimental results showed that USP13 could bind ubiquitin, bacterial-purified recombinant USP13 still exhibited only weak deubiquitination enzyme activity [ xref , xref , xref ]."

sparser
"The ZnF domain of USP13 cannot bind with ubiquitin, so that USP13 loses its ability to be activated by free ubiquitin ( xref )."

sparser
"We firstly examined whether the ZnF domain of USP13 (USP13-ZnF) interacts with Ub by isothermal titration calorimetry (ITC) experiment ( xref )."

reach
"In addition, NMR titration data reflected that M664, F666, M739, and F741 might be the key residues responsible for the binding of USP13-UBA to Ub (Zhang et al., 2011)."

sparser
"Overall, the USP13-ZnF domain cannot bind to Ub to activate USP13, whereas USP13-UBA can bind, which may partially explain the reason why USP13 displays only weak basal deubiquitination enzyme activity: the binding sites of USP13 to Ub are less than that other USP members, thus providing weaker binding affinity and consequent cleavage activity towards ubiquitin chains; There possibly exists constitutive self-inhibition for full-length USP13 supported by the interaction of UBA with ZnF domain, which is hypothesized to be released by recruitment of other proteins or modification, such as phosphorylation."

sparser
"The ZnF domain of USP13 cannot bind with Ub."
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
USP13 inhibits Ubiquitin.
| 4
| 4

reach
"USP13 interacted with TLR4 and inhibited the ubiquitin mediated degradation of TLR4."

reach
"3.5 USP13 inhibits the ubiquitin mediated degradation of mutant EGFR."

reach
"In a cell study, USP5 and USP13 were found to degrade ubiquitin chains inside stress granules, defined as clumps of protein or RNA created when cells are stressed [XREF_BIBR]."

reach
"USP13 antagonizes gp78 by removing ubiquitin."
| 21
| 13

reach
"In this study, we identified USP13 as the first deubiquitinase that reverses PTEN poly-ubiquitination and stabilizes PTEN protein, and found that USP13 suppresses tumorigenesis and glycolysis through PTEN."

reach
"These data suggested that USP13 overexpression inhibited glycolysis in OSCC cells."

reach
"Indeed, knockdown of USP13 increased glucose uptake and glycolysis, which could be fully reversed by restoration of PTEN, as gauged by lactate production and glucose incorporation assays (XREF_FIG)."

reach
"Then, we explored the effects of USP13 on aerobic glycolysis and found that silencing USP13 in OS cells decreased levels of extracellular acidification rate (ECAR, Fig. 1e) and reduced glycolysis and glycolytic capacity (Fig. 1f)."

reach
"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

reach
"Our data together with previous reports suggest the important context-specific role of USP13 in tumorigenesis.To explore the mechanism by which USP13 inhibited glycolysis in OSCC cells, we examined its effects on AKT signaling, which is known to be activated by USP13."

reach
"Ectopic expression of USP13 suppressed cell proliferation, glycolysis, and tumorigenicity."

reach
"miR-135b promoted cell proliferation and glycolysis that could be reversed by the overexpression of USP13 or PTEN."

reach
"Accordingly, we propose that USP13 inhibits the proliferation and glycolysis of OSCC cells via the upregulation of PTEN and the downregulation of AKT activity."

reach
"Zhang et al. reported that USP13 suppresses tumorigenesis and glycolysis in PTEN-positive breast cancer through the deubiquitination of PTEN [18]."
| 8

reach
"USP13 promotes glycolysis and progression in OS by binding to and stabilizing METTL3."

reach
"Ectopically expressed K488R of METTL3 significantly promoted cell proliferation and glycolysis, while USP13 knockdown decreased cell proliferation and glycolysis with WT METTL3 rather than K488R METTL3, which indicated that mutation of K488 of METTL3 promoted cell proliferation and glycolysis and abolished USP13 depletion-induced proliferation and glycolysis suppression (Fig. 4i-n)."

reach
"Next, we investigated the role of METTL3 in the mechanism by which USP13 promotes glycolysis and progression of OS cells via lentiviral-mediated METTL3 overexpression in USP13-silenced 143B cells and METTL3 silencing in USP13-upregulated HOS cells."

reach
"Using various in vitro and in vivo functional experiments, USP13 was demonstrated to promote glycolysis and tumor progression in OS."

reach
"Despite this, the diverse roles of autophagy in tumor biology should be explored more comprehensively.Collectively, our findings in the present study showed that USP13 promotes aerobic glycolysis and OS development by binding to, stabilizing and reducing ubiquitination and degradation of the m A writer METTL3 at K488 by removing K48-linked ubiquitin chains., which subsequently increases the m A modification of ATG5 mRNA and stabilizes it, thus promoting oncogenic autophagy."

reach
"USP13 was found to upregulate in osteosarcoma (OS) specimens and promote OS progression through regulating aerobic glycolysis."

reach
"Collectively, our study results indicate that USP13 promotes glycolysis and tumor progression in OS by stabilizing METTL3, thereby stabilizing ATG5 mRNA and facilitating autophagy in OS."

reach
"USP13 promotes glycolysis and cell proliferation in OS."
USP13 affects EGFR
| 14 7
USP13 binds EGFR.
| 4 7
| 3 7

sparser
"We then recapitulated the interaction between exogenous USP13 and EGFR in HEK293T cells."

sparser
"The interaction of wild‐type EGFR with USP13 was hardly detectable, whereas USP13 coimmunoprecipitated with the EGFR ΔE746‐A750 and L858R mutants (Figure xref ), indicating that USP13 associates much more tightly with mutant than wild‐type EGFR."

sparser
"By employing the USP13‐C345A mutant or versions mutated in UBA1, UBA2 or both, we established that the UBA domains' integrity is necessary for USP13 to bind to mutant EGFR (Figure xref )."

reach
"We then recapitulated the interaction between exogenous USP13 and EGFR in HEK293T cells."

sparser
"The fraction of mutant EGFR bound to USP13 was also enriched in those two types of ubiquitin chains (Figure xref )."

reach
"We found that USP13 inhibits mutant EGFR degradation by a process that leads to the accumulation of ubiquitinated (K48 and K63) EGFR, which is bound to c-Cbl and USP13."

sparser
"The association is dependent on the presence of c‐Cbl, as we observed that increased expression of c‐Cbl enhanced USP13‐mutant EGFR interaction."

sparser
"The isopeptidase‐independent mechanism of action proposed so far for USP13 relies on its UBA domains, which bind to ubiquitinated substrates, possibly inducing structural changes that stabilize them. xref Coincidentally, K63‐linked ubiquitin chains are required for EGFR sorting and degradation. xref This type of chains has also been reported as the preferred substrates for the USP13 UBA domains. xref We found that the UBA domains of USP13 were required for the binding to and stabilizing EGFR, as mutated UBA domains in USP13 resulted in a loss of USP13EGFR interaction and of EGFR stabilization."

sparser
"Moreover, degradation of mutant EGFR, mediated by the ubiquitination system, is inhibited by this enzyme, resulting in an accumulation of polyubiquitinated-EGFR bound to USP13."

reach
"USP13 binds more tightly to the mutant EGFR than to wild-type EGFR."
UBA domains binds USP13 and mutated EGFR. 1 / 1
| 1

reach
"USP13 inhibits the degradation of mutant EGFR via deubiquitination, binds to mutant EGFR via its UBA domains, and inhibits EGFR endosomal sorting and lysosomal degradation."
USP13 inhibits EGFR.
| 5
USP13 inhibits mutated EGFR. 3 / 3
| 3

reach
"Overall, we showed that USP13 abrogates sorting and degradation of ubiquitinated mutant EGFR, independent of its catalytic activity but requiring its UBA domains that are known to bind to K63-type ubiquitin chains preferentially."

reach
"23 In HEK293T cells expressing EGFR-DeltaE740-A746, USP13 antagonized the destabilization of mutant EGFR mediated by c-Cbl or Cbl-b."

reach
"We demonstrate that USP13 targeting leads to an increased sensitivity toward EGFR inhibition by both osimertinib and afatinib in NSCLC harboring the most common EGFR mutations (DeltaE746-A750 and L858R and T790M) while sparing EGFR wild-type cells and tumors."
USP13 inhibits EGFR. 2 / 2
| 2

reach
"Moreover, USP13 inhibition and silencing lead to a decreased half-life of the mutant EGFR protein in NSCLC cells, indicating that USP13 likely stabilizes EGFR."

reach
"38, 39 Given that USP13 targeting caused EGFR destabilization, some of the effects may be independent of the EGFR kinase activity as EGFR can suppress cell death in manners that are independent of its kinase activity."
USP13 activates EGFR.
| 5
USP13 activates mutated EGFR. 5 / 5
| 5

reach
"37 USP13 is likely to stabilize mutant EGFR in a ubiquitinated state, and inhibition of USP13 destabilizes mutant EGFR and abrogates signaling of mutant EGFR in lung cancer cells."

reach
"Targeting USP13 mediated drug tolerance increases the efficacy of EGFR inhibition of mutant EGFR in non small cell lung cancer."

reach
"We found that USP13 inhibits mutant EGFR degradation by a process that leads to the accumulation of ubiquitinated (K48 and K63) EGFR, which is bound to c-Cbl and USP13."

reach
"Combined targeting of USP13 and EGFR strongly reduces the viability of EGFR mutant lung cancer cells in vitro and in vivo while sparing non malignant (EGFR wild-type) lung epithelial cells."

reach
"Targeting USP13 increased the sensitivity of EGFRm NSCLC cells to EGFR inhibition with small molecules in vitro and in vivo, suggesting that USP13 is a strong mutant EGFR-specific co-target that could improve the treatment efficacy of EGFR-targeted therapies."
USP13 affects RHBDF2
4 | 6 10
4 | 4 10

reach
"Direct protein binding between USP13 and IRHOM2 was carried out via the GST pull-down analyses according to our previous reports methods13 24."

sparser
"Mechanistically, we further determined that USP13 directly interacts with IRHOM2 and removes its K63-linked ubiquitination induced by ubiquitin-conjugating enzyme E2N (UBC13), a ubiquitin E2 conjugating enzyme, and thus prevents its activation of downstream cascade."

reach
"Also, the potential protein interaction of USP13 and IRHOM2 in mice were predicted and showed by GeneMANIA and STRING (Fig. S7B)."

sparser
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection ( xref )."

reach
"In light of the function of USP13 on regulation of IRHOM2-related signaling pathway, the in vivo and in vitro data forced us to put forward another crucial concerns by which whether the USP13 binds with IRHOM2 in the context of NASH pathologies."

reach
"The results actually indicate that USP13 directly binds with IRHOM2."

No evidence text available

sparser
"Also, the potential protein interaction of USP13 and IRHOM2 in mice were predicted and showed by GeneMANIA and STRING ( xref )."

sparser
"Inactive rhomboid protein 2 (IRHOM2) was a crucial and positive regulator of inflammation-associated diseases and hepatocyte USP13 interacted with IRHOM2 and removed its K63-linked ubiquitination in response to metabolic stresses."

sparser
"Thus, to identify potential inhibitory function of USP13 on Irhom2 signaling, we accordingly generated hepatocyte-specific Irhom2 -knockout (Hep Irhom2 KO) and hepatocyte-specific Usp13 and Irhom2 double deletion (Hep Usp13-Irhom2 DKO) mice (Supporting Information xref ), as indicated in Methods section."
USP13 binds RHBDF2 and L02. 2 / 2
| 2

reach
"Finally, the interaction of USP13, UBC13 and IRHOM2 was detected in the L02 cells."

reach
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection (Fig. S3C–S3E)."
USP13 affects FlaG
| 20
| 20

sparser
"Forced expression of FLAG-USP13 partially reversed the U0126-caused vimentin reduction ( Figures 4 H and S6 D)."

sparser
"Results showed that when HA-Ub-K48 and FLAG-USP13 were co-transfected, the ubiquitination level of MYC-HIF-1α was markedly reduced."

sparser
"However, when K48 was mutated to arginine (K48R), the ubiquitination level of HIF-1α remained unchanged despite the presence of FLAG-USP13."

sparser
"As expected, substitution of FLAG-USP13 with FLAG-USP13-AE resulted in a pronounced increase in HIF-1α ubiquitination, closely resembling the levels observed in the absence of FLAG-USP13 (Fig.  xref G)."

sparser
"FLAG-USP13 was transfected into 293 T cells in a gradient manner, and the protein level of HIF-1α was assessed using Western blot analysis."

sparser
"We first performed co-immunoprecipitation assays to confirm the interaction between USP13 and Mcl-1 by transfecting HEK293T cells with FLAG-USP13 and V5-hMcl-1."

sparser
"We also observed that re-expression of wild-type (WT) FLAG-USP13, but not catalytically inactive mutant (C345A) USP13, in H1299 USP13 -KO cells could rescue invasive phenotype in vitro ( Figure S1 H)."

sparser
"Immunoprecipitation with anti-FLAG antibody demonstrated that FLAG-USP13 interacted with over-expressed Mcl-1 (Fig. xref )."

sparser
"Consistent with in vitro transwell assay, we observed that depletion of USP13 significantly reduced the number of metastatic nodules, whereas re-expression of WT but not C345A FLAG-USP13 increased the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"HEK293T cells were transfected with FLAG-USP13, V5-Mcl-1 and HA-Ubiquitin, and anti-V5 immunoprecipitates were probed for the level of ubiquitination using the HA-antibody."
RHBDF2 affects USP13
4 | 6 10
4 | 4 10

reach
"Direct protein binding between USP13 and IRHOM2 was carried out via the GST pull-down analyses according to our previous reports methods13 24."

sparser
"Mechanistically, we further determined that USP13 directly interacts with IRHOM2 and removes its K63-linked ubiquitination induced by ubiquitin-conjugating enzyme E2N (UBC13), a ubiquitin E2 conjugating enzyme, and thus prevents its activation of downstream cascade."

reach
"Also, the potential protein interaction of USP13 and IRHOM2 in mice were predicted and showed by GeneMANIA and STRING (Fig. S7B)."

sparser
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection ( xref )."

reach
"In light of the function of USP13 on regulation of IRHOM2-related signaling pathway, the in vivo and in vitro data forced us to put forward another crucial concerns by which whether the USP13 binds with IRHOM2 in the context of NASH pathologies."

reach
"The results actually indicate that USP13 directly binds with IRHOM2."

No evidence text available

sparser
"Also, the potential protein interaction of USP13 and IRHOM2 in mice were predicted and showed by GeneMANIA and STRING ( xref )."

sparser
"Inactive rhomboid protein 2 (IRHOM2) was a crucial and positive regulator of inflammation-associated diseases and hepatocyte USP13 interacted with IRHOM2 and removed its K63-linked ubiquitination in response to metabolic stresses."

sparser
"Thus, to identify potential inhibitory function of USP13 on Irhom2 signaling, we accordingly generated hepatocyte-specific Irhom2 -knockout (Hep Irhom2 KO) and hepatocyte-specific Usp13 and Irhom2 double deletion (Hep Usp13-Irhom2 DKO) mice (Supporting Information xref ), as indicated in Methods section."
USP13 binds RHBDF2 and L02. 2 / 2
| 2

reach
"Finally, the interaction of USP13, UBC13 and IRHOM2 was detected in the L02 cells."

reach
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection (Fig. S3C–S3E)."
HMGB1 affects USP13
4 | 8 8
HMGB1 binds USP13.
4 | 7 8
4 | 7 8

sparser
"In this study, we demonstrated, based on LC–MS/MS, immunoprecipitation, and PLA studies, that the deubiquitinase USP13 interacts with HMGB1."

sparser
"Further study including the binding of USP13 to HMGB1 by proinflammatory stimulation is under investigation."

reach
"To confirm the interaction between HMGB1 and USP13, PLA was performed using the Duolink in situ fluorescence kit (Sigma-Aldrich)."

reach
"In this study, we investigated the effect of isopeptidase T-3, also called USP13, on HMGB1, since HMGB1 ubiquitination has rarely been reported.To confirm whether USP13 binds to HMGB1, both Myc-HMGB1 and Flag-tagged USP13 (Flag-USP13) or USP10 and USP15 plasmids were overexpressed in HEK293T cells for co-immunoprecipitation."

reach
"As shown in Fig. 1B and 1C, HMGB1 binds to USP13 but not to either USP10 or USP15 after immunoprecipitation with Myc."

reach
"HEK293T cells were immunoprecipitated with an anti-USP13 antibody for endogenous binding, and the binding of endogenous HMGB1 to USP13 is shown (Fig. 1D)."

reach
"As expected, Spautin-1 treatment decreased the binding of HMGB1 to USP13, suggesting that inhibition of the deubiquitinase activity of USP13 affects its binding to HMGB1 (Fig. 1F)."

reach
"The interaction between HMGB1 and USP13 was also observed using a proximity ligation assay (PLA) in HEK293T cells overexpressing HMGB1 and USP13 (Fig. 1G)."

reach
"Further study including the binding of USP13 to HMGB1 by proinflammatory stimulation is under investigation."

sparser
"USP13 Interacts with HMGB1."
HMGB1 increases the amount of USP13.
| 1
HMGB1 increases the amount of USP13. 1 / 1
| 1

reach
"HMGB1 expression increased with high USP13 expression (Fig. 4A)."
FlaG affects USP13
| 20
| 20

sparser
"Forced expression of FLAG-USP13 partially reversed the U0126-caused vimentin reduction ( Figures 4 H and S6 D)."

sparser
"Results showed that when HA-Ub-K48 and FLAG-USP13 were co-transfected, the ubiquitination level of MYC-HIF-1α was markedly reduced."

sparser
"However, when K48 was mutated to arginine (K48R), the ubiquitination level of HIF-1α remained unchanged despite the presence of FLAG-USP13."

sparser
"As expected, substitution of FLAG-USP13 with FLAG-USP13-AE resulted in a pronounced increase in HIF-1α ubiquitination, closely resembling the levels observed in the absence of FLAG-USP13 (Fig.  xref G)."

sparser
"FLAG-USP13 was transfected into 293 T cells in a gradient manner, and the protein level of HIF-1α was assessed using Western blot analysis."

sparser
"We first performed co-immunoprecipitation assays to confirm the interaction between USP13 and Mcl-1 by transfecting HEK293T cells with FLAG-USP13 and V5-hMcl-1."

sparser
"We also observed that re-expression of wild-type (WT) FLAG-USP13, but not catalytically inactive mutant (C345A) USP13, in H1299 USP13 -KO cells could rescue invasive phenotype in vitro ( Figure S1 H)."

sparser
"Immunoprecipitation with anti-FLAG antibody demonstrated that FLAG-USP13 interacted with over-expressed Mcl-1 (Fig. xref )."

sparser
"Consistent with in vitro transwell assay, we observed that depletion of USP13 significantly reduced the number of metastatic nodules, whereas re-expression of WT but not C345A FLAG-USP13 increased the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"HEK293T cells were transfected with FLAG-USP13, V5-Mcl-1 and HA-Ubiquitin, and anti-V5 immunoprecipitates were probed for the level of ubiquitination using the HA-antibody."
| 15

reach
"We have shown that USP13 deficiency promotes lung inflammation."

reach
"For example, USP13 inhibited lung inflammation via stabilization of the anti-inflammatory receptor IL-1R8/single immunoglobin interleukin-1-related receptor (Sigirr) (116)."

reach
"USP13 reduces septic mediated cardiomyocyte oxidative stress and inflammation by inducing Nrf2."

reach
"Upregulation of USP13 repressed inflammatory response in H-FLSs upon TNF-α or IL-1β challenge due to the improvement of PTEN and reduction of AKT phosphorylation as well as inactivation of NF-κB pathway (102)."

reach
"Furthermore, the deubiquitinating enzyme USP13 stabilizes the anti-inflammatory receptor IL-1R8/Sigirr to suppress lung inflammation [40,41,42]."

reach
"The deubiquitinase USP13 stabilizes the anti-inflammatory receptor IL-1R8 and Sigirr to suppress lung inflammation."

reach
"For example, the deubiquitinating enzyme Ubiquitin Specific Peptidase 13 (USP13) stabilizes the anti-inflammatory receptor immunoglobin interleukin-1-related receptor (IL-1R8/Sigirr) to inhibit lung inflammation (162)."

reach
"Further studies showed that USP13 reduced LPS-induced oxidative stress and inflammation by inducing Nrf2."

reach
"In a series of mice models with NASH phenotype, hepatocyte-specific deficiency of USP13 dramatically potentiated hepatosteatosis, hepatic inflammation, and hepatofibrosis, whereas USP13 overexpression in hepatocytes efficiently suppressed NASH diet-induced steatohepatitis."

reach
"To summarize, USP13 suppressed liver steatosis, inflammation, and insulin resistance to demonstrate efficacy in treating ob/ob mice."
| 4

reach
"The roles of USP13 and the newly synthesized protein in liver inflammation, injury, and repair will be the future focuses."

reach
"We found that after a long-term HFHC/WTDF administration, dysfunctional USP13 in hepatocytes significantly promoted hepatic steatosis, liver inflammation, and hepatofibrosis."

reach
"Hepatocyte-specific loss of the Usp13 disrupts liver metabolic homeostasis, followed by glycometabolic disorder, lipid deposition, increased inflammation, and markedly promotes NASH development."

reach
"In collagen-mediated arthritis (CIA) mice, USP13 attenuated synovial hyperplasia, cartilage damage and inflammation and bone loss (102)."
USP13 affects USP13
| 1 17
USP13 activates USP13.
| 1 8
USP13 activates USP13. 9 / 10
| 1 8

reach
"Inhibition of the phosphorylation of USP13 downregulated USP13 protein stability and reduced the proliferation of ovarian cancer cells, which may lead to novel therapeutics targeting USP13 in USP13-amplified cancers."

reach
"Phosphorylation of USP13 at Thr122 by CK2 increases the stability of USP13, which in turn promotes ovarian cancer proliferation."

eidos
"( D ) Hypoxia increased USP13 , TLR4 , MyD88 , and P-NF-kappaB p65 proteins , which was subsequently reduced by USP13 knockdown ."

reach
"This study revealed that USP13 is phosphorylated at Thr122, which upregulates the stability of USP13 at a post-translation level."

reach
"Overexpression of USP13 in PC12 cells almost reversed the Spautin‐1‐mediated decrease in NLRP3, USP13, Beclin1 and GSDMD‐N expression levels (Figure 3D,E)."

reach
"We further demonstrated that Casein kinase 2 (CK2) directly interacts with and phosphorylates USP13 at Thr122, which promotes the stability of USP13 protein."

reach
"Moreover, low USP13-expressing HOS cells were chosen to overexpress USP13 stably (Fig. S2a)."

reach
"Genes most strongly decreased in ALS included MAP kinase activating death domain (MADD) and ubiquitin specific peptidase 13 (USP13) (Figures 2H–N)."

reach
"CK2 phosphorylates the Thr122 residue of USP13, which upregulates the stability of USP13 at a post-translation level."
USP13 increases the amount of USP13.
| 6
USP13 increases the amount of USP13. 6 / 6
| 6

reach
"In addition, Guo et al. reported that RREB1 is a TF of USP13, that promotes USP13 expression [49]."

reach
"The lentivirus overexpressing USP13 effectively upregulated the expression of USP13 compared with the control lentivirus (vector) and the control cells that did not undergo any treatment (Figure 2A)."

reach
"In addition, knockdown of USP13 reduced vimentin expression and invasion capacity in A549 cells expressing WT β-catenin but failed to inhibit vimentin and invasion in cells expressing K508R β-catenin [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"To generate the LV-Usp13 or LV-shUsp13 vectors, the mouse Usp13 cDNA sequences or shRNA targeting mouse Usp13 sequences were loaded into pLenti-GFP-Puro-CMV or pLenti-U6-GFP (Addgene) to upregulate or downregulate USP13 expression in vivo experiments.2.9 Metabolic factors and serum cytokines parameters."

reach
"In addition, via bioinformatic analyses and molecular biology experiments, we revealed that Yin Yang 1 (YY1) is a new transcription factor of USP13 that promotes USP13 transcription through binding to its promoter region."

reach
"These results underscore a key role of Twist1 in a USP13-induced enhancement of migration and invasion capacities of breast cancer cells.We also assessed the effects of USP13 expression on lung metastasis of breast cancer to further substantiate the above in vitro findings."
USP13 decreases the amount of USP13.
| 2
USP13 decreases the amount of USP13. 2 / 2
| 2

reach
"Moreover, treatment with a proteasome inhibitor blocked the c-Myc loss caused by USP13 knockdown (not depicted), but overexpression of USP13 delayed c-Myc turnover in the presence of cycloheximide (not depicted), suggesting a critical role of USP13 in preventing proteasomal degradation of c-Myc in GSCs."

reach
"We found that both USP13 shRNAs reduced USP13 expression in vivo compared to the control shRNA although with different efficiencies."
USP13 dephosphorylates USP13.
| 1
USP13 leads to the dephosphorylation of USP13 on T122. 1 / 1
| 1

reach
"To validate the activity of the anti-pT122 antibody, we generated the point mutation of Thr122 by using a site-directed substitution of threonine to alanine on USP13 to abolish USP13 phosphorylation on Thr122 residue."
USP13 affects HYCC1
| 19
USP13 activates HYCC1.
| 17
USP13 activates HYCC1. 10 / 17
| 17

reach
"In addition, implanting USP13 knockdown cells to nude mice showed that knockdown of USP13 significantly inhibited HCC tumor growth."

reach
"These results suggest that the USP13‐PRPF6 axis promotes HCC cell proliferation by regulating AKT‐mTOR signalling.3.6 USP13 Expression Is Correlated With PRPF6 in Human HCC Tissues."

reach
"Thus, USP13 deficiency suppressed HCC cell proliferation, migration, and stemness in vitro.To further validate the function of USP13 in HCC, we established USP13-overexpressing HCC cells via lentivirus (Supplementary Fig. S2A, B)."

reach
"68 While inhibiting USP13 significantly suppresses HCC and ovarian tumor progression is unknown whether these effects are mediated by ATP citrate lyase and oxoglutarate dehydrogenase."

reach
"Further investigation revealed that the knockout of USP13 inhibited HCC cell proliferation, whereas overexpression of USP13 had the opposite effect."

reach
"Further experimental results confirmed that the USP13‐PRPF6 axis promoted the proliferation of HCC cells by modulating the AKT‐mTOR signalling pathway.In conclusion, we have demonstrated that USP13 can decrease the K48/63‐linked polyubiquitination of PRPF6, thereby stabilising this protein and promoting HCC cell proliferation via regulation of the AKT‐mTOR pathway (Figure 8C)."

reach
"However, there is no evidence that the regulation of WWP1 deubiquitination serves as a substrate.In this study, we confirmed that USP13 was elevated in HCC and acted as an oncogene to enhance the malignant characteristics of HCC cells both in vitro and in vivo."

reach
"According to the results of the Transwell assay (Supplementary Fig. S2F) and wound healing assay (Supplementary Fig. S2G), USP13 promoted HCC cell migration."

reach
"3.1 USP13 Is Up-Regulated in HCC and Associated With Poor Prognosis."

reach
"Moreover, USP13 stabilized WWP1 by removing the K29- and K48-linked polyubiquitination chains from WWP1 and then activated the Akt/mTOR pathway in HCC."
USP13 inhibits HYCC1.
| 2
USP13 inhibits HYCC1. 2 / 2
| 2

reach
"These results demonstrate that USP13 indirectly activates the Akt/mTOR pathway through WWP1.CCK-8 and colony formation assays revealed that WWP1 rescued the suppression of HCC cell proliferation caused by USP13 silencing (Fig. 5C, D)."

reach
"Moreover, our results demonstrated that downregulation of USP13 inhibited HCC cell proliferation, while its overexpression promoted cellular proliferation."
STAT1 affects USP13
2 | 2 14
2 | 2 14

sparser
"The ability of USP13(C343A) to remove ubiquitin molecules from STAT1 was decreased (Fig.  xref ), although USP13(C343A) still binds to STAT1 (Supplementary Fig.  xref )."

sparser
"Additionally, we reveal the protective role of the cardiomyocyte-specific USP13-STAT1 axis in myocardial hypertrophy, indicating that cardiac-specific gene therapy targeting USP13 could represent a promising therapeutic strategy for cardiac hypertrophy."

sparser
"These observations validated that USP13 directly binding STAT1 protein in cardiomyocytes."

sparser
"Usp13 mainly interacted with the nonactivated form of Stat1 ( xref )."

sparser
"This study illustrated a beneficial effect of USP13 in hypertrophic cardiomyocytes and identified a cardiomyocyte-specific USP13-STAT1 axis in regulating cardiac hypertrophy."

sparser
"Here, the authors show the beneficial effects of a cardiomyocyte-derived deubiquitinating enzyme, USP13, in hypertrophic cardiomyocytes and identify a cardiomyocyte-specific USP13-STAT1 axis in regulating cardiac hypertrophy."

No evidence text available

No evidence text available

reach
"Moreover, we observed substantial attenuation of the interaction between STAT1 and USP13 upon overexpression of Mbtps1 (Fig. 7f)."

sparser
"Thus, we hypothesize that USP13-STAT1 axis protects against cardiac hypertrophy through upregulating Nppb transcription and BNP expression."
USP13 affects MDM2
| 11 6
USP13 binds MDM2.
| 2 6
| 2 6

sparser
"USP13 is associated with MDM2."

sparser
"These experimental results indicate that USP13 interacts with MDM2 in the nucleus."

sparser
"The Co-IP results demonstrate that USP13 is associated with MDM2 ( xref and xref )."

sparser
"To further confirm the interaction between USP13 and MDM2, Co-IP experiments after over-expression of exogenous genes were conducted."

sparser
"USP13 interacts with MDM2 and removes K63-linked ubiquitin chains on MDM2."

sparser
"The results demonstrate that exogenously expressed USP13 and MDM2 proteins are associated each other ( xref and xref )."

reach
"To further confirm the interaction between USP13 and MDM2, Co-IP experiments after over-expression of exogenous genes were conducted."

reach
"Considering the direct association between USP13 and MDM2 and the role of USP13 in the degradation of MDM2, we examined if USP13 deubiquitinates MDM2."
USP13 deubiquitinates MDM2.
| 4
USP13 deubiquitinates MDM2. 4 / 4
| 4

reach
"This study is the first to report that USP13 is involved in aging by deubiquitinating MDM2."

reach
"Considering the direct association between USP13 and MDM2 and the role of USP13 in the degradation of MDM2, we examined if USP13 deubiquitinates MDM2."

reach
"The results indicate that USP13 deubiquitinates MDM2."

reach
"We also used in vitro experiments to examine the deubiquitination of MDM2 by USP13 using a purified human recombinant protein USP13 (rhUSP13) and MDM2 autoubiquitination kit."
USP13 inhibits MDM2.
| 3
USP13 inhibits MDM2. 3 / 3
| 3

reach
"USP13 negatively regulates MDM2 through K63-linked ubiquitination."

reach
"Taken together, this data indicates that over-expression of USP13 causes the reduction and inhibition of MDM2 and thus promotes the senescence of cells."

reach
"The degradation of MDM2 caused by USP13 was prevented by the proteasome inhibitor MG132."
USP13 decreases the amount of MDM2.
| 2
USP13 decreases the amount of MDM2. 2 / 2
| 2

reach
"Knockdown of USP13 increased MDM2 levels in lung cells, while overexpression of USP13 reduced it."

reach
"The results indicate that exogenous MDM2 levels were decreased by over-expression of USP13."
USP13 affects BAG6
4 | 4 9
4 | 4 5

sparser
"We agree with the reviewers that the physical interaction of USP13 with the Bag6 complex makes it hard to distinguish whether the effect of USP13 knockdown on Bag6-SGTA interaction is due to lack of deubiquitination by USP13 or its scaffolding function."

reach
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."

No evidence text available

No evidence text available

reach
"We next tested whether USP13 could physically interact with Bag6 and/or its cofactors."

reach
"Because depletion of gp78 did not affect the interaction of USP13 with Bag6 (XREF_FIG, lane 7 vs 5), USP13 might bind Bag6 directly."

No evidence text available

reach
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."

No evidence text available

sparser
"By contrast, the cytosolic chaperone Hsp90 was not co-precipitated ( xref ), demonstrating a specific interaction between USP13 and the Bag6 complex."
USP13 binds AMFR and BAG6. 2 / 2
| 2

sparser
"USP13 forms a multiprotein complex with gp78 and Bag6."

sparser
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."
UBL binds USP13 and BAG6. 2 / 2
| 2

sparser
"On the other hand, the binding of USP13 with the UBL domain of Bag6 can be demonstrated using pulldown experiments ( xref )."

sparser
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."
USP13 affects AKT
| 1 13 1
USP13 activates AKT.
| 1 5
USP13 activates AKT. 6 / 7
| 1 5

reach
"Further experimental results confirmed that the USP13‐PRPF6 axis promoted the proliferation of HCC cells by modulating the AKT‐mTOR signalling pathway.In conclusion, we have demonstrated that USP13 can decrease the K48/63‐linked polyubiquitination of PRPF6, thereby stabilising this protein and promoting HCC cell proliferation via regulation of the AKT‐mTOR pathway (Figure 8C)."

reach
"Overall, USP13 facilitates the malignant progression of HCC cells through the upregulation of WWP1.Thus, we concluded that USP13 stabilized WWP1, and then activated the Akt/mTOR pathway."

reach
"Moreover, USP13 stabilized WWP1 by removing the K29- and K48-linked polyubiquitination chains from WWP1 and then activated the Akt/mTOR pathway in HCC."

reach
"By acting as an oncogene, USP13 affects HCC malignant progression through the WWP1-mediated Akt/mTOR pathway."

eidos
"Wu et al demonstrate that amplification of USP13 in some NSCLC leads to enhanced AKT and MAPK signaling that sustains cancer progression ."

reach
"Importantly, USP13 modulated the AKT‐mTOR signalling pathway in a manner analogous to PRPF6 (Figures 6D,E)."
USP13 dephosphorylates AKT.
| 4
USP13 leads to the dephosphorylation of AKT. 2 / 2
| 2

reach
"The deubiquitinating enzyme USP13 increases PTEN protein stability and inhibits Akt phosphorylation, suppressing tumourigenesis and glycolysis in breast cancer (BC) [42] ."

reach
"6 The PTEN protein level was increased and the phosphorylation of AKT (p-AKT) was reduced by the overexpression of USP13 (Figure 2F)."
Modified USP13 leads to the dephosphorylation of AKT. 2 / 2
| 2

reach
"Loss of USP13 in breast cancer cells promotes AKT phosphorylation, cell proliferation, anchorage independent growth, glycolysis and tumor growth through downregulation of PTEN."

reach
"This overexpression upregulated PTEN protein and downregulated AKT and FOXO1/3 phosphorylation in the MDAMB-231 cell line (XREF_FIG) which expresses moderate but detectable levels of endogenous PTEN (XREF_SUPPLEMENTARY), while knockdown of PTEN in USP13 overexpressing MDA-MB-231 cells rescued the phosphorylation of both AKT and FOXO1/3 (XREF_SUPPLEMENTARY)."
USP13 phosphorylates AKT.
| 3
USP13 leads to the phosphorylation of AKT. 3 / 3
| 3

reach
"Depletion of USP13 in breast cancer cells promoted AKT phosphorylation, cell proliferation, glycolysis, and tumor growth through downregulation of PTEN."

reach
"Similarly, depletion of USP13 downregulated PTEN protein and upregulated AKT and FOXO1/3 phosphorylation in MCF10A (XREF_FIG) and MCF7 (XREF_FIG) mammary epithelial cells and in HCT116 colon cancer cells (XREF_FIG), but not in the isogenic PTEN-null HCT116 cells (XREF_FIG)."

reach
"USP13 also protected BMSCs from MPS-induced apoptosis by modulating PTEN and reducing AKT phosphorylation."
USP13 inhibits AKT.
| 1 1
USP13 inhibits AKT. 2 / 2
| 1 1

reach
"We conclude from these data that USP13 inhibits AKT signaling through positive regulation of PTEN protein."

sparser
"Some studies indicate that USP13 inhibits AKT by stabilizing PTEN in bladder cancer, thereby reducing tumor proliferation, invasion, and migration [ xref ]."
Spautin-1 affects USP13
1 | 2 12
Spautin-1 inhibits USP13.
1 | 2 7
Spautin-1 inhibits USP13. 7 / 7
| 7

reach
"It has been reported that spautin-1 suppresses autophagy via inhibition of USP10 and USP13, the two deubiquitinases to degrade PIK3C3/VPS34 and BECN1 [12]."

reach
"To directly test if spautin-1 can inhibit the deubiquitinating activity of USP10 and USP13, we tested the activity of isolated USP10 and USP13 on ubiquitinated Beclin1 in vitro."

reach
"Using this assay, we found that spautin-1 inhibited USP10 and USP13 with IC 50 of ~ 0.6-0.7 microM while having no inhibitory activity toward CYLD which is also a member of ubiquitin specific peptidase family (XREF_FIG)."

reach
"Further, inhibition of USP13 by spautin-1 caused a decrease in TXNIP levels in MLE12 cells (Fig. 4)."

reach
"Consistently, knockdown or pharmacological inhibition of USP13 by spautin-1 retards the growth, differentiation and invasion of many tumors, providing a possibility for antagonizing the drug resistance of tumor cells."

reach
"Another autophagy inhibitor, spautin-1, degrades class III PI3K by inhibiting the activity of the ubiquitin specific peptidases, USP10 and USP13 (Mateo et al., 2013, Liu et al., 2011)."

reach
"Among them, spautin-1, a well-established autophagy inhibitor, is known to promote the degradation of BECN1 and PIK3C3/VPS34 by targeting the ubiquitin-specific peptidases, USP10 and USP13 [12]."
1 | 2

eidos
"Spautin-1 was shown to inhibit USP-13 , a de-ubiquitinase of the host cell hijacked by the SARS-CoV-2 which led to successful ubiquitination of nsp-13 [ 109 ] ."

eidos
"Further , inhibition of USP13 by spautin-1 caused a decrease in TXNIP levels in MLE12 cells ( Fig. 4 ) ."

No evidence text available
Spautin-1 decreases the amount of USP13.
| 3
Spautin-1 decreases the amount of USP13. 3 / 3
| 3

reach
"In addition, spautin-1 also downregulated the expression of USP10 and USP13 at the protein level [53]."

reach
"Cerebral I/R significantly reduced the expressions of NLRP3, USP13, beclin 1, and GSDMD-N, which was abolished by spautin-1, indicating that spautin-1 inhibited autophagy and NLRP3 inflammasome-mediated pyroptosis."

reach
"Interestingly, the treatment of spautin-1 also reduced the levels of USP10 and USP13, but not USP14, a DUB involved in regulating proteasome function, or Rubicon, a negative regulator of type III PI3 kinase (XREF_FIG)."
Spautin-1 activates USP13.
| 2
Spautin-1 activates USP13. 2 / 2
| 2

reach
"Because the reductions in the levels of USP10 and USP13 in H4-LC3-GFP cells treated with spautin-1 appeared later than the reductions in the levels of Vps34 complexes and autophagy (XREF_FIG), the reduced levels of USP10 and USP13 are unlikely to be the primary reason for the ability of spautin-1 to reduce the levels of PtdIns3P and inhibit autophagy."

reach
"Briefly, spautin-1 targets two ubiquitin specific peptidases (USP10 and USP13), both of which mediate deubiquitination of Beclin1 and p53, thus leading to their proteasomal degradation [93]."
USP13 affects Cyclin
| 15
USP13 deubiquitinates Cyclin.
| 5
USP13 deubiquitinates Cyclin on D1. 5 / 5
| 5

reach
"USP13 deubiquitinates and stabilizes cyclin D1 to promote gastric cancer cell cycle progression and cell proliferation."

reach
"Taken together, our data demonstrate that USP13 deubiquitinates and stabilizes cyclin D1, thereby promoting cell cycle progression and cell proliferation in GC."

reach
"USP13 interacted with and deubiquitinated cyclin D1."

reach
"To further define whether USP13 deubiquitinated cyclin D1, we transfected USP13 siRNAs or negative control siRNA together with Myc-cyclin D1 and HA-Ub into HEK293T cells and performed Co-IP assays."

reach
"In this study, we reported for the first time that USP13 stabilized and deubiquitinated cell cycle protein cyclin D1 in GC."
USP13 increases the amount of Cyclin.
| 3
USP13 increases the amount of Cyclin. 3 / 3
| 3

reach
"Furthermore, Maria et al. (2022) showed that USP13 silencing by siRNAs decreased expression of Cyclin D1, leading to cell cycle arrest [26]."

reach
"As shown in Fig. 4A, B, USP13 siRNAs remarkably decreased the cyclin D1 protein level, but had no obvious effect on other cell cycle related proteins."

reach
"Enforced expression of USP13 increased cyclin D1 protein level, while knockdown of USP13 reduced cyclin D1 protein level."
USP13 activates Cyclin.
| 3
USP13 activates Cyclin. 3 / 3
| 3

reach
"USP13 increases the stability of cyclin D1 in GC cells."

reach
"Enforced expression of USP13 increased cyclin D1 protein level, while knockdown of USP13 reduced cyclin D1 protein level."

reach
"Besides, USP13 protein was positively associated with cyclin D1 protein abundance in human GC tissue samples, indicating that USP13-mediated cyclin D1 regulation was clinically relevant."
USP13 inhibits Cyclin.
| 2
USP13 inhibits Cyclin. 2 / 2
| 2

reach
"Our results showed that USP13 knockdown decreased the half-life of cyclin D1 and facilitated the degradation of cyclin D1 in GC cells (Fig. 4E, F), whereas enforced expression of WT USP13, but not the USP13-C345A mutant, prolonged the half-life of cyclin D1 and delayed its degradation (Fig. 4G, H)."

reach
"To further explore whether proteasome degradation pathway is involved in USP13-mediated stability of cyclin D1, we treated the transfected cells with proteasome inhibitor MG132 and found that MG132 alleviated USP13 siRNA-induced cyclin D1 downregulation (Fig. 4I)."
USP13 decreases the amount of Cyclin.
| 2
USP13 decreases the amount of Cyclin. 2 / 2
| 2

reach
"As shown in Fig. 6A, knockdown of USP13 with siRNA increased the ubiquitination level of cyclin D1 in HEK293T cells."

reach
"In contrast, overexpression of Flag-USP13 decreased the ubiquitination level of cyclin D1 in HEK293T cells and GC cells (Fig. 6B)."
USP13 affects COL9A2
| 15
| 15

sparser
"We noted that VAPB KO clones in the USP13-Med line displayed a remarkably altered morphology, as they became more flattened and more elongated (Sup."

sparser
"The therapeutic potential of miR‐367 was further confirmed in vivo , in an orthotopic xenograft model of aggressive embryonal CNS tumors initiated by OCT4A‐overexpressing cells (Daoy, USP13MED, or USP7‐ATRT) in Balb/C nude mice."

sparser
"Furthermore, USP13-Med and DAOY VAPB KO cells showed decreased proliferation rates compared to controls (Fig.  xref B and C)."

sparser
"The overall survival rate of animals harboring Daoy, USP13MED, or USP7‐ATRT‐initiated tumors was also significantly improved by treatment with miR‐367 inhibitor (Fig. xref D)."

sparser
"In order to assess any potential impact of doxycycline on cell proliferation, we conducted additional experiments applying doxycycline in DAOY and USP13-Med wild-type and VAPB KO cells."

sparser
"Next, an in vivo study was conducted with a intracerebroventricular injection of ZIKV in BALB/c nude mice after period of CNS tumor establishment (1 to 2 weeks for DAOY, USP13-MED and USP7-ATRT cell lines)."

sparser
"We noted that the number of differentially expressed genes between VAPB WT and VAPB KO cells was much higher in the USP13-Med background than in DAOY cells (Fig.  xref A)."

sparser
"This finding agrees with the pronounced morphology change observed in USP13-Med VAPB KO s, an effect that was not detected in the DAOY VAPB KO cells."

sparser
"In this setting, both DAOY as well as USP13-Med cell lines formed tumors in vivo and overall survival is displayed in Fig.  xref G and H. Histologic analysis of the brain of terminated mice confirmed tumors in brain tissue in every group (Fig. xref ), although VAPB KO tumors were smaller."

sparser
"The DAOY cell line, representative of the Sonic Hedgehog (SHH) subgroup, and the USP13-Med cell line, which was obtained from a 3-year-old boy with classic medulloblastoma, subgroup 4 xref ."
USP13 affects CCND1
3 | 12
3 | 12

No evidence text available

sparser
"Unexpectedly, compared with the full-length USP13, deletion of the UBP domain obviously decreased the interation with cyclin D1, indicating that the UBP domain might play some positive role in the interaction between USP13 and cyclin D1."

sparser
"Mechanistically, USP13 binds physically to the N-terminal domain of cyclin D1 and selectively deubiquitinates its K48-linked polyubiquitination chain while leaving K63-linked chains intact ( xref )."

sparser
"To detect the direct interaction between USP13 and cyclin D1, we performed GST-pulldown assays with the recombinant GST-USP13 protein."

sparser
"Third, Co-IP, GST pull down and immunofluorescence assays validated the physical interaction between USP13 and cyclin D1."

sparser
"In this study, we tried to map the regions that mediated the interaction between USP13 and cyclin D1."

sparser
"The results indicated that the N-terminal but not the C-terminal of cyclin D1 could interact with USP13 (Fig. xref )."

sparser
"Our results showed that the N-terminal domain of cyclin D1 (aa1–153) and the UBA (aa 625–863) domain of USP13 mediated the interaction between USP13 and cyclin D1."

sparser
"USP13 was positively associated with cyclin D1 in human GC specimens."

sparser
"USP13 directly interacts with cyclin D1 in GC cells."
USP13 affects ATG5
1 4 | 8 2
USP13 activates ATG5.
| 7
USP13 activates ATG5. 7 / 7
| 7

reach
"On the contrary, IGF2BP2 elevates the stability of a deubiquitinase ubiquitin specific peptidase 13 (USP13) to prolong the half-life of autophagy-related protein 5 (ATG5), inducing pro-survival autophagy and drug resistance in gastrointestinal stromal tumors 60."

reach
"Taken together, these findings demonstrated that USP13 upregulated ATG5 via an m A-dependent mechanism through binding with and stabilizing the m A writer METTL3 in OS."

reach
"USP13 promotes oncogenic autophagy dependent on METTL3-mediated upregulation of ATG5 in OS cells."

reach
"Collectively, our study results indicate that USP13 promotes glycolysis and tumor progression in OS by stabilizing METTL3, thereby stabilizing ATG5 mRNA and facilitating autophagy in OS."

reach
"USP13 promotes AML cell growth and autophagy by upregulating ATG5."

reach
"USP13 promotes acute myeloid leukemia cell proliferation and autophagy by promoting ATG5."

reach
"In gastrointestinal mesenchymal tumors, inhibitors of USP13 attenuate ATG5 and inhibit m6A modification-mediated autophagy."
USP13 binds ATG5.
4 | 2
4 | 2

No evidence text available

No evidence text available

sparser
"USP13 preferentially is induced in GIST cells by IM and interacts with ATG5, which leads to stabilization of ATG5 through deubiquitination."

sparser
"Activation of PAK1 promoted phosphorylation of ATG5 thereby enhancing the interaction of ATG5 with USP13."

No evidence text available

No evidence text available
USP13 increases the amount of ATG5.
1 | 1
USP13 increases the amount of ATG5. 2 / 2
1 | 1

reach
"Also, our results indicated that upregulation of METTL3 in USP13-knockdown 143B cells rescued the downregulated ATG5 mRNA, while silencing of METTL3 in USP13-overexpressing HOS cells decreased the upregulated ATG5 mRNA level (Fig. 6l)."

"Here, we identified USP13 as an essential deubiquitinase that stabilizes ATG5 in a process that depends on the PAK1 serine/threonine-protein kinase and which enhances autophagy and promotes IM resistance in GIST cells."
COL9A2 affects USP13
| 15
| 15

sparser
"We noted that VAPB KO clones in the USP13-Med line displayed a remarkably altered morphology, as they became more flattened and more elongated (Sup."

sparser
"The therapeutic potential of miR‐367 was further confirmed in vivo , in an orthotopic xenograft model of aggressive embryonal CNS tumors initiated by OCT4A‐overexpressing cells (Daoy, USP13MED, or USP7‐ATRT) in Balb/C nude mice."

sparser
"Furthermore, USP13-Med and DAOY VAPB KO cells showed decreased proliferation rates compared to controls (Fig.  xref B and C)."

sparser
"The overall survival rate of animals harboring Daoy, USP13MED, or USP7‐ATRT‐initiated tumors was also significantly improved by treatment with miR‐367 inhibitor (Fig. xref D)."

sparser
"In order to assess any potential impact of doxycycline on cell proliferation, we conducted additional experiments applying doxycycline in DAOY and USP13-Med wild-type and VAPB KO cells."

sparser
"Next, an in vivo study was conducted with a intracerebroventricular injection of ZIKV in BALB/c nude mice after period of CNS tumor establishment (1 to 2 weeks for DAOY, USP13-MED and USP7-ATRT cell lines)."

sparser
"We noted that the number of differentially expressed genes between VAPB WT and VAPB KO cells was much higher in the USP13-Med background than in DAOY cells (Fig.  xref A)."

sparser
"This finding agrees with the pronounced morphology change observed in USP13-Med VAPB KO s, an effect that was not detected in the DAOY VAPB KO cells."

sparser
"In this setting, both DAOY as well as USP13-Med cell lines formed tumors in vivo and overall survival is displayed in Fig.  xref G and H. Histologic analysis of the brain of terminated mice confirmed tumors in brain tissue in every group (Fig. xref ), although VAPB KO tumors were smaller."

sparser
"The DAOY cell line, representative of the Sonic Hedgehog (SHH) subgroup, and the USP13-Med cell line, which was obtained from a 3-year-old boy with classic medulloblastoma, subgroup 4 xref ."
CCND1 affects USP13
3 | 12
3 | 12

No evidence text available

sparser
"Unexpectedly, compared with the full-length USP13, deletion of the UBP domain obviously decreased the interation with cyclin D1, indicating that the UBP domain might play some positive role in the interaction between USP13 and cyclin D1."

sparser
"Mechanistically, USP13 binds physically to the N-terminal domain of cyclin D1 and selectively deubiquitinates its K48-linked polyubiquitination chain while leaving K63-linked chains intact ( xref )."

sparser
"To detect the direct interaction between USP13 and cyclin D1, we performed GST-pulldown assays with the recombinant GST-USP13 protein."

sparser
"Third, Co-IP, GST pull down and immunofluorescence assays validated the physical interaction between USP13 and cyclin D1."

sparser
"In this study, we tried to map the regions that mediated the interaction between USP13 and cyclin D1."

sparser
"The results indicated that the N-terminal but not the C-terminal of cyclin D1 could interact with USP13 (Fig. xref )."

sparser
"Our results showed that the N-terminal domain of cyclin D1 (aa1–153) and the UBA (aa 625–863) domain of USP13 mediated the interaction between USP13 and cyclin D1."

sparser
"USP13 was positively associated with cyclin D1 in human GC specimens."

sparser
"USP13 directly interacts with cyclin D1 in GC cells."
USP13 affects PRPF6
| 8 6
USP13 binds PRPF6.
| 1 6
| 1 6

sparser
"Furthermore, the USP13PRPF6 axis facilitates HCC cell proliferation via modulation of the mTOR signalling pathway."

sparser
"Further experimental results confirmed that the USP13PRPF6 axis promoted the proliferation of HCC cells by modulating the AKT‐mTOR signalling pathway."

reach
"The co‐IP experiment was initially employed to validate the interaction between USP13 and PRPF6."

sparser
"The co‐IP experiment was initially employed to validate the interaction between USP13 and PRPF6."

sparser
"These results suggest that the USP13PRPF6 axis promotes HCC cell proliferation by regulating AKT‐mTOR signalling."

sparser
"To further explore the signalling pathways underlying the effect of the USP13PRPF6 axis on HCC cell proliferation, bioinformatic analysis was performed."

sparser
"Ultimately, we demonstrated that the USP13PRPF6 axis promoted HCC cell proliferation was closely associated with the activation of the AKT‐mTOR signalling pathway."
USP13 activates PRPF6.
| 3
USP13 activates PRPF6. 3 / 3
| 3

reach
"Our subsequent analyses revealed that USP13 enhanced the stability of PRPF6 by reducing its K48/63‐linked polyubiquitination level, a process mediated by the deubiquitinating activity of USP13."

reach
"These results suggest that overexpression of USP13 promotes the proliferation of HCC cells.3.4 USP13 Stabilises PRPF6 Protein by Decreasing Its K48/63-Linked Polyubiquitination Levels."

reach
"In contrast, USP13 overexpression markedly increased PRPF6 protein stability (Figure 5G)."
USP13 deubiquitinates PRPF6.
| 2
USP13 deubiquitinates PRPF6. 2 / 2
| 2

reach
"We subsequently determined that USP13 significantly reduced the polyubiquitination of PRPF6, with a specific focus on the K48 and K63 chain linkages (Figures 5D,E)."

reach
"The results above indicate that USP13 deubiquitinates PRPF6, thereby stabilising its protein levels and promoting the proliferation of HCC cells via modulation of the AKT‐mTOR signalling pathway."
USP13 increases the amount of PRPF6.
| 1
USP13 increases the amount of PRPF6. 1 / 1
| 1

reach
"Encouragingly, the USP13‐C345A mutant exhibited a loss of its ability to modulate PRPF6 protein levels and failed to reduce the polyubiquitination levels of PRPF6 (Figures 5B,C)."
USP13 decreases the amount of PRPF6.
| 1
USP13 decreases the amount of PRPF6. 1 / 1
| 1

reach
"Our results revealed that USP13 overexpression significantly attenuated the polyubiquitination levels of PRPF6 (Figure 5A)."
USP13 affects MITF
1 1 | 9 1
USP13 activates MITF.
| 4
USP13 activates MITF. 4 / 5
| 4

reach
"USP13 promotes melanoma cell invasion by stabilizing the microphthalmia-associated transcription factor."

reach
"On the other hand, overexpression of the deubiquitinase USP13, which has been identified as a key regulator of MITF turnover, can enhance the basal half-life of the MITF protein to up to 4h (Zhao etal., 2011)."

reach
"More recently, it was shown that MITF is targeted by the de-ubiquitinase USP13, a theoretically drug-able protease whose suppression results in strong downregulation of MITF protein levels."

reach
"Consequently, ectopic expression of USP13 stabilized MITF and up-regulated MITF target genes, whereas USP13 knockdown decreased MITF protein, but not mRNA, and blocked melanoma cell proliferation.A range of additional phosphorylation events have been detected in various high-throughput proteomic screens (PhosphSitePlus, http://www.phosphosite.org/proteinAction?id=1001&showAllSites=true); however, the significance of these modifications is yet to be determined."
USP13 increases the amount of MITF.
| 2
USP13 increases the amount of MITF. 2 / 3
| 2

reach
"By modulate the expression of MITF downstream genes, USP13 appeared to be essential for melanoma growth both in vivo and in vitro, indicating the possibility of targeting USP13 for melanoma therapy."

reach
"USP13 has been identified as a deubiquitination enzyme of MITF to modulate the ubiquitination level of MITF, affecting the survival of melanoma cells (Zhao et al., 2011)."
USP13 binds MITF.
1 | 1 1
1 | 1 1

sparser
"USP13 binds to MITF and cleaves the ubiquitin chain ( xref )."

reach
"USP13 binds to MITF and cleaves the ubiquitin chain (65)."

No evidence text available
USP13 deubiquitinates MITF.
1 | 1
USP13 deubiquitinates MITF. 2 / 2
1 | 1

reach
"In these tumors, USP13 deubiquitinates and stabilizes some oncogenes, including MCL1 [19, 20], Myc [8], MITF [21] and ZHX2 [22]."

"However, deubiquitinase <span class="match term0">USP13</span> can stabilize <span class="match term1">MITF</span> expression and prolong its half-life."
USP13 inhibits MITF.
| 1
USP13 inhibits MITF. 1 / 1
| 1

reach
"Androgen receptors decrease the expression of USP13 by upregulating miRNA-539-3P, thus increasing the degradation of MITF protein by ubiquitination and promoting melanoma metastasis (85,86)."
USP13 affects SNAI1
1 | 11 1
USP13 deubiquitinates SNAI1.
| 5
USP13 deubiquitinates SNAI1. 5 / 5
| 5

reach
"The report from Zhang et al. revealed that USP13 promotes the epithelial-mesenchymal transition (EMT) and metastasis in GC by deubiquitinating and stabilizing Snail in GC [10]."

reach
"The transcriptional activator PLAGL2 activates the ubiquitin-specific peptidase 13 (USP13), which deubiquitinates and maintains Snail to influence the metastatic power of GC cells [64]."

reach
"USP13 interacted with Snail to deubiquitinate and stabilize Snail in GC cells."

reach
"USP13, USP29, and USP37 were reported to promote the metastasis of GC by deubiquitinating and stabilizing Snail [35,36,54,59]."

reach
"Further evidence showed that EMT might be the primary mechanism for the enhanced metastatic property in GC cells: USP13 and USP29 promoted TGF-β1-induced EMT through interacting with and deubiquitinating Snail, while USP3 facilitated this process by deubiquitinating SUZ12 (Wu et al. 2021; Zhang et al. 2022a, b; Qian et al. 2020)."
USP13 activates SNAI1.
| 4
USP13 activates SNAI1. 4 / 4
| 4

reach
"Deubiquitinase USP13 promotes the epithelial-mesenchymal transition and metastasis in gastric cancer by maintaining Snail protein."

reach
"USP13 promotes migration and invasion of GC cells by maintaining the stability of Snail."

reach
"In conclusion, USP13 overexpression was frequently detected in GC and contributed to the EMT and metastasis of GC by stabilizing Snail."

reach
"Mechanistically, USP13 knockdown promoted Snail degradation, which could be blocked by the proteasome inhibitor MG132."
USP13 increases the amount of SNAI1.
| 2
USP13 increases the amount of SNAI1. 2 / 2
| 2

reach
"Among these DUBs, USP29 and USP13 have been reported to increase Snail1 protein levels in GC cells."

reach
"Interestingly, USP13 remarkably enhanced Snail protein expression but did not affect its mRNA levels in GC cells."
USP13 binds SNAI1.
1 | 1
1 | 1

No evidence text available

sparser
"USP13 interacted with Snail to deubiquitinate and stabilize Snail in GC cells."
USP13 affects NLRP3
1 | 12
USP13 binds NLRP3.
1 | 11
1 | 11

sparser
"USP13 binds to the PYD of NLRP3, which prevents the interaction of NLRP3 with its E3 ubiquitin ligase TRIM31."

sparser
"In response to LPS treatment, the interaction between USP13 and NLRP3 rapidly increases in the early phase, while the interaction between TRIM31 and NLRP3 potently increases at relatively later time points."

sparser
"Co-IP assays showed that NLRP3 and USP13 interacted with each other (Fig.  xref f, g)."

No evidence text available

sparser
"USP13 only binds to NLRP3 but not to ASC or pro–caspase-1, the other two subunits of the NLRP3 inflammasome, when coexpressed together in HEK293T cells ( xref )."

sparser
"In murine BMDMs, LPS priming induced a rapid binding of USP13 to NLRP3 but a relatively later binding of TRIM31 to NLRP3; however, USP13 deficiency resulted in an early and robust interaction between TRIM31 and NLRP3 ( xref )."

sparser
"The results showed that the truncated USP13 lacking the N-terminal region (amino acid residues 625 to 863) almost lost its ability to interact with NLRP3, indicating that NLRP3 binds to the N-terminal region of USP13 ( xref )."

sparser
"Moreover, the domains of USP13 required for USP13-NLRP3 interaction and USP13-Paxillin association were assessed by constructing and analyzing plasmids encoding for WT USP13(1–863) and four truncated proteins (Fig.  xref i)."

sparser
"Like WT USP13(1–863), USP13(301–863) and USP13(625–863) interacted with NLRP3, but USP13(1–300) or USP13(301–624) failed to interact with NLRP3 (Fig.  xref j), indicating that 625aa–863aa of USP13 are involved in USP13-NLRP3 interaction."

sparser
"We first confirmed the interaction between NLRP3 and USP13 in human embryonic kidney (HEK) 293T cells overexpressed with Myc-tagged human NLRP3 and Flag-tagged human USP13 by coimmunoprecipitation assay ( xref )."
USP13 ubiquitinates NLRP3.
| 1
USP13 ubiquitinates NLRP3. 1 / 1
| 1

sparser
"Unexpectedly, we found that treatment with spautin-1, an inhibitor of USP13 enzyme activity, could not reverse the inhibition of NLRP3 ubiquitination by USP13 ( xref )."
USP13 affects HSPA5
| 13
| 13

sparser
"Our data show that both LPS and Tm could induce the formation of the GRP78 and USP13 complex (Figure  xref )."

sparser
"Similarly, endogenous USP13 and GRP78 formed a complex in DSS‐induced colon tissues (Figure  xref )."

sparser
"Also, the interaction between exogenous USP13 and GRP78 was confirmed in NIH/3T3 cells co‐transfected with Flag‐USP13 and His‐GRP78 plasmids (Figure  xref ; Figure xref , Supporting Information) in a dose‐dependent manner (Figure  xref )."

sparser
"We also demonstrated the protective role of the intestinal epithelial cell‐specific USP13GRP78 axis in IBD, suggesting that targeting USP13 for intestinal epithelial cell‐specific gene therapy may be a potential treatment strategy for IBD."

sparser
"Mechanistically, USP13 interacted with  GRP78, attenuating ER stress‐induced apoptosis and maintaining intestinal barrier integrity by selectively removing K63‐linked ubiquitin at lysine 327."

sparser
"These observations validated that USP13 interacts with GRP78 protein both in vitro and in vivo."

sparser
"These findings underscore the protective role of the USP13GRP78 axis in IBD and suggest that targeting USP13 through intestinal epithelial‐specific gene therapy may be a potential therapeutic strategy for IBD."

sparser
"USP13 interacts with GRP78 and reduces its K63‐linked ubiquitination to alleviate epithelial injury."

sparser
"DUBs control cellular functions by altering the ubiquitination status of proteins, which influences protein stability and regulates signaling pathways. [ xref ] The interaction between USP13 and GRP78 raises the question of whether USP13 plays a regulatory role in GRP78."

sparser
"Anti‐ HA (51064‐2‐AP), anti‐Flag (66008‐4‐Ig), anti‐6×His (His, 10001‐0‐AP), anti‐USP13 (16840‐1‐AP), anti‐GRP78 (11587‐1‐AP, 66574‐1‐lg), anti‐eukaryotic translation initiation factor 2‐alpha kinase 3 (PERK, 24390‐1‐AP), anti‐tight junction protein (ZO‐1, 21773‐1‐AP), anti‐Claudin‐1 (28674‐1‐AP), anti‐Occludin (27260‐1‐AP), anti‐DNA‐damage‐inducible transcript 3 (CHOP, 15204‐1‐AP), anti‐Cleaved Caspase‐3 (25128‐1‐AP), anti‐Caspase‐3 (19677‐1‐AP), anti‐activating transcription factor 4 (ATF4, 10835‐1‐AP), anti‐E‐cadherin (60335‐1‐Ig), and anti‐epithelial cell adhesion molecule (EpCAM, 21050‐1‐AP) antibodies were purchased from Proteintech (Wuhan, China)."
| 11
USP13 activates Carcinogenesis.
| 9

reach
"In our genetically engineered mouse model, USP13 overexpression promoted ovarian tumorigenesis and metastasis [14]."

reach
"Ubiquitin-specific Peptidase 13 (USP13) is highly amplified and promotes tumorigenesis and metastasis in ovarian cancer."

reach
"USP13 is highly amplified in human ovarian cancer, and elevated expression of USP13 promotes tumorigenesis and metastasis of ovarian cancer."

reach
"USP13 knockdown impaired the tumorigenesis of GC cells in vivo."

reach
"To investigate whether USP13 knockdown suppressed the tumorigenesis of GC cells in vivo, we injected MKN-45 GC cells which were stably knocked-down USP13 (shUSP13) or the negative control (shNC) into the subcutaneous of the nude mice."

reach
"Importantly, overexpression of Usp13 accelerates tumorigenesis, enhances tumor metastasis, and causes poor outcomes in transgenic mouse models of ovarian cancer , underscoring its importance in promoting tumorigenesis in vivo."

reach
"In vitro and in vivo depletion of USP13 inhibited SCLC cancer stem cells (CSCs) properties and tumorigenesis, and this inhibitory effect was rescued by reconstituted expression of wide type (WT) USP13 but not the enzyme-inactive USP13 mutant."

reach
"More importantly, the small molecule inhibitor of FASN, TVB-2640, significantly inhibits lipogenic phenotype and attenuates self-renewal ability, chemotherapy resistance and USP13-mediated tumorigenesis in SCLC."

reach
"Overexpression of USP13 with deletion of Trp53 and Pten in murine ovarian surface epithelium accelerated ovarian tumorigenesis and led to decreased survival in mice."
USP13 inhibits Carcinogenesis.
| 2

reach
"Zhang et al. reported that USP13 suppresses tumorigenesis and glycolysis in PTEN-positive breast cancer through the deubiquitination of PTEN [18]."

reach
"USP13 overexpression inhibited the proliferation of and glycolysis in OSCC cells in vitro and suppressed tumorigenesis in nude mice."
USP10 affects USP13
| 4 6
| 4 4

sparser
"USP13 also interacts with USP10, and the interaction increases USP13 deubiquitinating activity ( xref )."

sparser
"In addition, whether and how the complex relationship between PTEN DUBs (e.g., HAUSP−USP10 xref , HAUSP−USP11 xref and USP10USP13 xref ) strengthens their activity toward PTEN requires further study."

reach
"USP13 also interacts with USP10, and the interaction increases USP13 deubiquitinating activity."

sparser
"Although no structural information for USP13 and USP10 is currently available, the enhanced DUB activity when USP13 interacts with USP10 or when they interact with Beclin1 suggest that the interaction of USP13 and USP10 with each other or with their substrates can lead to changes in their conformation which may be critical for the catalytic activities."

reach
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

reach
"Using spautin-1 as a tool, we explored the interaction of USP10 and USP13 with Vps34 complexes."

sparser
"No orthogonal biophysical methods, such as the surface plasmon response (SPR) or isothermal titration calorimetry (ITC), were used to validate the binding of spautin-1 with USP10 or USP13."

reach
"Although no structural information for USP13 and USP10 is currently available, the enhanced DUB activity when USP13 interacts with USP10 or when they interact with Beclin1 suggest that the interaction of USP13 and USP10 with each other or with their substrates can lead to changes in their conformation which may be critical for the catalytic activities."
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
METTL3 affects USP13
1 3 | 5 4
METTL3 binds USP13.
3 | 3 4
3 | 3 4

No evidence text available

reach
"Taken together, our results confirm that USP13 interacts with METTL3.Since USP13 binds to METTL3, we then explored the effect of downregulating or upregulating USP13 on METTL3 stability in OS cells."

sparser
"In this study, we found that USP13 interacts with METTL3 and decreases METTL3 ubiquitination, resulting in reduced METTL3 degradation in OS."

sparser
"Moreover, the interaction between USP13 and METTL3 was verified by Co-IP analysis in HEK 293T cells with exogenously transfection of Flag-tagged USP13 and Myc-tagged METTL3 (Fig. xref c)."

sparser
"Taken together, our results confirm that USP13 interacts with METTL3."

sparser
"Since USP13 binds to METTL3, we then explored the effect of downregulating or upregulating USP13 on METTL3 stability in OS cells."

No evidence text available

No evidence text available

reach
"USP13 interacts with and stabilizes METTL3."

reach
"Moreover, the interaction between USP13 and METTL3 was verified by Co-IP analysis in HEK 293T cells with exogenously transfection of Flag-tagged USP13 and Myc-tagged METTL3 (Fig. 3c)."
METTL3 activates USP13.
| 2
METTL3 activates USP13. 2 / 2
| 2

reach
"Furthermore, N 6 -methyladenosine methyltransferase-like 3 (METTL3) mediated stabilization of USP13 mRNA that required the m 6 A reader IGF2BP2."

reach
"Mechanically, USP13’s m A modification is catalyzed by METTL3, and IGF2BP2 promotes USP13 mRNA translation by identifying m A residues on USP13 transcripts, ultimately triggering autophagy [169]."
METTL3 increases the amount of USP13.
1 |
METTL3 increases the amount of USP13. 1 / 1
1 |

"Furthermore, N6-methyladenosine methyltransferase-like 3 (METTL3) mediated stabilization of USP13 mRNA that required the m6A reader IGF2BP2."
HSPA5 affects USP13
| 13
| 13

sparser
"Our data show that both LPS and Tm could induce the formation of the GRP78 and USP13 complex (Figure  xref )."

sparser
"Similarly, endogenous USP13 and GRP78 formed a complex in DSS‐induced colon tissues (Figure  xref )."

sparser
"Also, the interaction between exogenous USP13 and GRP78 was confirmed in NIH/3T3 cells co‐transfected with Flag‐USP13 and His‐GRP78 plasmids (Figure  xref ; Figure xref , Supporting Information) in a dose‐dependent manner (Figure  xref )."

sparser
"We also demonstrated the protective role of the intestinal epithelial cell‐specific USP13GRP78 axis in IBD, suggesting that targeting USP13 for intestinal epithelial cell‐specific gene therapy may be a potential treatment strategy for IBD."

sparser
"Mechanistically, USP13 interacted with  GRP78, attenuating ER stress‐induced apoptosis and maintaining intestinal barrier integrity by selectively removing K63‐linked ubiquitin at lysine 327."

sparser
"These observations validated that USP13 interacts with GRP78 protein both in vitro and in vivo."

sparser
"These findings underscore the protective role of the USP13GRP78 axis in IBD and suggest that targeting USP13 through intestinal epithelial‐specific gene therapy may be a potential therapeutic strategy for IBD."

sparser
"USP13 interacts with GRP78 and reduces its K63‐linked ubiquitination to alleviate epithelial injury."

sparser
"DUBs control cellular functions by altering the ubiquitination status of proteins, which influences protein stability and regulates signaling pathways. [ xref ] The interaction between USP13 and GRP78 raises the question of whether USP13 plays a regulatory role in GRP78."

sparser
"Anti‐ HA (51064‐2‐AP), anti‐Flag (66008‐4‐Ig), anti‐6×His (His, 10001‐0‐AP), anti‐USP13 (16840‐1‐AP), anti‐GRP78 (11587‐1‐AP, 66574‐1‐lg), anti‐eukaryotic translation initiation factor 2‐alpha kinase 3 (PERK, 24390‐1‐AP), anti‐tight junction protein (ZO‐1, 21773‐1‐AP), anti‐Claudin‐1 (28674‐1‐AP), anti‐Occludin (27260‐1‐AP), anti‐DNA‐damage‐inducible transcript 3 (CHOP, 15204‐1‐AP), anti‐Cleaved Caspase‐3 (25128‐1‐AP), anti‐Caspase‐3 (19677‐1‐AP), anti‐activating transcription factor 4 (ATF4, 10835‐1‐AP), anti‐E‐cadherin (60335‐1‐Ig), and anti‐epithelial cell adhesion molecule (EpCAM, 21050‐1‐AP) antibodies were purchased from Proteintech (Wuhan, China)."
WWP1 affects USP13
| 5 7
| 5 7

reach
"Immunoprecipitation with WWP1 antibody further validated the interaction between WWP1 and USP13 (Fig. 3B)."

reach
"Next, we transfected Myc-tagged WWP1 (Myc-WWP1), along with Flag-tagged USP13 (Flag-USP13) or an empty vector, into HCC and HEK293T cells, followed by a Co-IP assay to validate the binding between USP13 and WWP1."

reach
"Overall, there was an interaction between USP13 and WWP1."

reach
"Since our findings described above demonstrated the interaction between USP13 and WWP1, we wondered whether USP13 regulated WWP1 through deubiquitination."

reach
"According to our results, USP13 truncations containing either the N-terminal region (aa1–300 and aa1–624) or the C-terminal region (aa301–863 and aa625–863) could bind to WWP1."

sparser
"Finally, our functional experiments in HCC cells and a subcutaneous xenograft model further revealed the oncogenic role of the USP13-WWP1 axis both in vitro and in vivo."

sparser
"Furthermore, USP13 can interact with WWP1 and then remove the K29- and K48-linked polyubiquitination chains from WWP1 to stabilize the WWP1 protein via the ubiquitin–proteasome pathway."

sparser
"USP13 interacted with WWP1 in HCC cells."

sparser
"Immunoprecipitation with WWP1 antibody further validated the interaction between WWP1 and USP13 (Fig.  xref B)."

sparser
"To determine the specific region of USP13 that interacts with WWP1, we constructed five Flag-tagged USP13 truncation plasmids."
| 2 10
| 2 8

eidos
"Results Depletion of USP13 Induces Apoptosis and Suppresses Autophagy Impaired ERAD pathway causes prolonged ER stress , which activates alternative signal pathways and promotes apoptosis ( Hetz , 2012 ; Hetz and Papa , 2018 ; Tabas and Ron , 2011 ) ."

reach
"USP13, FBXL14 and c-Myc can interact with each other; USP13 inhibits the degradation of c-Myc, hence inhibition of USP13 promotes the apoptosis of glioma stem cells."

reach
"In addition, terminal deoxynucleotidyl transferase deoxyuridine triphosphate nick-end labeling (TUNEL) assay demonstrated that expression of the mutant c-Myc diminished the increased apoptosis caused by USP13 disruption in GSC derived xenografts (not depicted)."

eidos
"USP13 suppression induces apoptosis upon EGFR inhibition of mutant NSCLC cells USP13 downregulation or inhibition severely compromised the viability of PC9 and HCC827 cells when cotreated with afatinib ."

reach
"In conclusion, this study demonstrated that USP13 was downregulated in septic myocardial injury tissues, and USP13 overexpression increased Nrf2 levels and reduced apoptosis."

reach
"3.3 USP13 suppression induces apoptosis upon EGFR inhibition of mutant NSCLC cells."

reach
"We then showed that over-expressing USP13 markedly suppressed inflammatory response, oxidative stress and apoptosis in H-FLSs upon IL-1beta or TNF-alpha challenge, whereas USP13 knockdown exhibited detrimental effects."

reach
"Additionally, it is found that USP13 abrogates cell apoptosis by preventing excessive ubiquitin-proteasome degradation of Bcl-XL."

reach
"However, in the context of hypoxic or oxidative stresses, USP13 loss caused a substantial decrease of cell viability and a significant increase of cell apoptosis in both TOV-21G and SW-1573 tumor models."

reach
"USP13 depletion induced cell apoptosis in ccRCC cells (SI Appendix, Fig. S4 P and Q)."
| 2

reach
"Downregulation of USP13 resulted in reduced WISP1 protein stability, decreased cell proliferation, migration, and EMT, and increased apoptosis in vitro."

reach
"USP13 signaling in cell proliferation and apoptosis."
USP13 affects SIAH2
2 | 6 1
USP13 activates SIAH2.
| 4
USP13 activates SIAH2. 4 / 7
| 4

reach
"Conversely, inhibition of USP13 expression with corresponding shRNA decreases the stability of both Siah2 and its substrate Spry2."

reach
"Different forms of stress modulate these activities : hypoxia reduces USP13 expression enabling increased Siah2 activity, while cellular stress induces p38 activity and alters Siah localization, affecting its recognition and targeting of substrates 174."

reach
"In melanoma cells, USP13 attenuates the autodegradation of Siah2, a RING (really interesting new gene) finger family E3 ubiquitin ligase."

reach
"Overexpression of USP13 increases Siah2 stability by attenuating its autodegradation."
USP13 binds SIAH2.
2 | 1
2 | 1

sparser
"Overall, this study identifies a new layer of Siah2 regulation mediated by USP13 binding to ubiquitinated Siah2 protein with a concomitant inhibitory effect on its activity under normoxia."

No evidence text available

No evidence text available
USP13 deubiquitinates SIAH2.
| 2
USP13 deubiquitinates SIAH2. 2 / 2
| 2

reach
"The DUB USP13 regulates Siah2 availability and activity as deubiquitination of Siah2 by USP13 results in a more stable, albeit less active, ubiquitin ligase."

reach
"Besides phosphorylation, Siah2 activity can be modulated by the deubiquitinating enzyme USP13, which binds to and deubiquitinates Siah2, increasing its stability but diminishing its activity toward its substrates."
UIMC1 affects USP13
4 | 6 2
UIMC1 binds USP13.
4 | 6
4 | 6

reach
"To test our hypothesis, we first examined whether USP13 interacts with RAP80."

reach
"Also, USP13 interacts with and deubiquitylates RAP80 to activate its function ."

No evidence text available

No evidence text available

No evidence text available

reach
"USP13 interacts with and deubiquitinates RAP80."

No evidence text available

reach
"Also, USP13 interacts to and deubiquitylates receptor-associated protein 80 (RAP80) following DNA damage to activate its ability, promoting DNA-damage responses ."

reach
"To evaluate the recruitment of USP13 and RAP80 at DSBs, we analysed the kinetics of GFP-USP13 and GFP-RAP80 accumulation on stripes after laser microirradiation."

reach
"Consistent with their reports, we also found that RAP80 binded with USP13 in EC cells treated with or without cisplatin."
UIMC1 ubiquitinates USP13.
| 2
UIMC1 ubiquitinates USP13. 2 / 2
| 2

sparser
"We found that mutating the 3 sites (K75, K90 and K112, 3KR) near or at the UIM domains, but not the other sites, abolished USP13 deficiency-induced RAP80 ubiquitination ( xref and xref )."

sparser
"In addition, we found all the single mutations of these three sites cannot abolish the USP13 deficiency-induced RAP80 ubiquitination ( xref )."
NLRP3 affects USP13
1 | 11
1 | 11

sparser
"USP13 binds to the PYD of NLRP3, which prevents the interaction of NLRP3 with its E3 ubiquitin ligase TRIM31."

sparser
"In response to LPS treatment, the interaction between USP13 and NLRP3 rapidly increases in the early phase, while the interaction between TRIM31 and NLRP3 potently increases at relatively later time points."

sparser
"Co-IP assays showed that NLRP3 and USP13 interacted with each other (Fig.  xref f, g)."

No evidence text available

sparser
"USP13 only binds to NLRP3 but not to ASC or pro–caspase-1, the other two subunits of the NLRP3 inflammasome, when coexpressed together in HEK293T cells ( xref )."

sparser
"In murine BMDMs, LPS priming induced a rapid binding of USP13 to NLRP3 but a relatively later binding of TRIM31 to NLRP3; however, USP13 deficiency resulted in an early and robust interaction between TRIM31 and NLRP3 ( xref )."

sparser
"The results showed that the truncated USP13 lacking the N-terminal region (amino acid residues 625 to 863) almost lost its ability to interact with NLRP3, indicating that NLRP3 binds to the N-terminal region of USP13 ( xref )."

sparser
"Moreover, the domains of USP13 required for USP13-NLRP3 interaction and USP13-Paxillin association were assessed by constructing and analyzing plasmids encoding for WT USP13(1–863) and four truncated proteins (Fig.  xref i)."

sparser
"Like WT USP13(1–863), USP13(301–863) and USP13(625–863) interacted with NLRP3, but USP13(1–300) or USP13(301–624) failed to interact with NLRP3 (Fig.  xref j), indicating that 625aa–863aa of USP13 are involved in USP13-NLRP3 interaction."

sparser
"We first confirmed the interaction between NLRP3 and USP13 in human embryonic kidney (HEK) 293T cells overexpressed with Myc-tagged human NLRP3 and Flag-tagged human USP13 by coimmunoprecipitation assay ( xref )."
EGFR affects USP13
| 5 7
EGFR binds USP13.
| 4 7
| 3 7

sparser
"We then recapitulated the interaction between exogenous USP13 and EGFR in HEK293T cells."

sparser
"The interaction of wild‐type EGFR with USP13 was hardly detectable, whereas USP13 coimmunoprecipitated with the EGFR ΔE746‐A750 and L858R mutants (Figure xref ), indicating that USP13 associates much more tightly with mutant than wild‐type EGFR."

sparser
"By employing the USP13‐C345A mutant or versions mutated in UBA1, UBA2 or both, we established that the UBA domains' integrity is necessary for USP13 to bind to mutant EGFR (Figure xref )."

reach
"We then recapitulated the interaction between exogenous USP13 and EGFR in HEK293T cells."

sparser
"The fraction of mutant EGFR bound to USP13 was also enriched in those two types of ubiquitin chains (Figure xref )."

reach
"We found that USP13 inhibits mutant EGFR degradation by a process that leads to the accumulation of ubiquitinated (K48 and K63) EGFR, which is bound to c-Cbl and USP13."

sparser
"The association is dependent on the presence of c‐Cbl, as we observed that increased expression of c‐Cbl enhanced USP13‐mutant EGFR interaction."

sparser
"The isopeptidase‐independent mechanism of action proposed so far for USP13 relies on its UBA domains, which bind to ubiquitinated substrates, possibly inducing structural changes that stabilize them. xref Coincidentally, K63‐linked ubiquitin chains are required for EGFR sorting and degradation. xref This type of chains has also been reported as the preferred substrates for the USP13 UBA domains. xref We found that the UBA domains of USP13 were required for the binding to and stabilizing EGFR, as mutated UBA domains in USP13 resulted in a loss of USP13EGFR interaction and of EGFR stabilization."

sparser
"Moreover, degradation of mutant EGFR, mediated by the ubiquitination system, is inhibited by this enzyme, resulting in an accumulation of polyubiquitinated-EGFR bound to USP13."

reach
"USP13 binds more tightly to the mutant EGFR than to wild-type EGFR."
UBA domains binds USP13 and mutated EGFR. 1 / 1
| 1

reach
"USP13 inhibits the degradation of mutant EGFR via deubiquitination, binds to mutant EGFR via its UBA domains, and inhibits EGFR endosomal sorting and lysosomal degradation."
EGFR activates USP13.
| 1
EGFR activates USP13. 1 / 1
| 1

reach
"In addition, the combined effect of USP13 downregulation by EGFR inhibition and USP13 siRNA also increased the negative effect on EGFR stability."
CK2 affects USP13
| 5 7
CK2 phosphorylates USP13.
| 4 5
CK2 phosphorylates USP13. 5 / 5
| 2 3

reach
"We then addressed whether CK2 phosphorylates USP13 in the cellular context by inhibiting CK2 kinase activity in cells with CK2 inhibitors, silmitasertib (CX-4945) and 4,5,6,7-tetrabromobenzotriazole (TBB) [33,34,35]."

sparser
"Western blotting with an anti-pT122 antibody showed that CK2 phosphorylated GST-USP13 protein but not GST, indicating that USP13 is a substrate of CK2 in vitro ( xref D), suggesting that USP13 is a substrate for CK2."

reach
"Further studies will determine whether CK2-mediated Thr122 phosphorylation affects the deubiquitinase activity of USP13, USP13-interactome network, or other PTMs on USP13."

sparser
"We then addressed whether CK2 phosphorylates USP13 in the cellular context by inhibiting CK2 kinase activity in cells with CK2 inhibitors, silmitasertib (CX-4945) and 4,5,6,7-tetrabromobenzotriazole (TBB) [ xref , xref , xref ]."

sparser
"Previous studies have identified various regulatory mechanisms of USP13, including METTL3 and IGF2BP2-dependent N6-methyladenosine modifications of USP13 [ xref ] and CK2-induced phosphorylation of USP13 [ xref ]."
CK2 phosphorylates USP13 on T122. 4 / 4
| 2 2

sparser
"Phosphorylation of USP13 at Thr122 by CK2 increases the stability of USP13, which in turn promotes ovarian cancer proliferation."

reach
"CK2 phosphorylates the Thr122 residue of USP13, which upregulates the stability of USP13 at a post-translation level."

reach
"We further demonstrated that Casein kinase 2 (CK2) directly interacts with and phosphorylates USP13 at Thr122, which promotes the stability of USP13 protein."

sparser
"CK2 phosphorylates the Thr122 residue of USP13, which upregulates the stability of USP13 at a post-translation level."
CK2 binds USP13.
| 1 2
| 1 2

sparser
"To determine whether USP13 interacts with CK2 in ovarian cancer cells, we examined CK2USP13 binding in HeyA8 cells expressing FLAG-tagged USP13."

sparser
"This experiment showed that CK2 was co-immunoprecipitated with USP13, indicating that USP13 interacts with CK2 in ovarian cancer cells ( xref C)."

reach
"Interaction between USP13 and CK2 was verified with western blotting."
Rep affects USP13
8 | 3
8 | 3

No evidence text available

No evidence text available

sparser
"We found that nsp13 interacts with the deubiquitinase USP13, which deubiquitinates and stabilizes nsp13."

No evidence text available
| DOI

No evidence text available

sparser
"Nsp13 also interacts with the deubiquitinating enzyme USP13, which is proposed to stabilize Nsp13 via its deubiquitylation ( xref )."

No evidence text available
| DOI

No evidence text available
| DOI

No evidence text available
| DOI

sparser
"The interaction with the highest pDockQ score (0.433) was the NSP13-USP13 interaction (Fig.  xref ), involving the NSP13 viral helicase and the human deubiquitinase USP13 xref ."
USP13 affects UBC
11 |
11 |

No evidence text available

No evidence text available

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No evidence text available
USP13 affects TOPBP1
2 1 | 4 4
USP13 binds TOPBP1.
2 | 1 4
2 | 1 4

No evidence text available

No evidence text available

sparser
"USP13 binds to and stabilizes TopBP1 through deubiquitination and orchestrates the replication stress response [ xref ]."

sparser
"Mechanistically, USP13 binds to TopBP1 and stabilizes TopBP1 by deubiquitination."

reach
"Mechanistically, USP13 binds to TopBP1 and stabilizes TopBP1 by deubiquitination."

sparser
"To test whether USP13 is a deubiquitinase for TopBP1, we first examined whether USP13 interacts with TopBP1."

sparser
"Our study shows that USP13 interacts with TopBP1."
USP13 deubiquitinates TOPBP1.
1 | 3
USP13 deubiquitinates TOPBP1. 4 / 4
1 | 3

"Mechanistically, <span class="match term0">USP13</span> binds to <span class="match term1">TopBP1</span> and stabilizes <span class="match term1">TopBP1</span> by deubiquitination"

reach
"Protein interaction experiments demonstrated that USP13 could co-immunoprecipitate with endogenous TopBP1, and in vitro deubiquitination enzyme experiments showed that WT-USP13 could deubiquitinate TopBP1, while CA-USP13 could not, highlighting the requirement for USP13 ubiquitination activity (Kim et al., 2021)."

reach
"USP13 can deubiquitinate DNA topoisomerase 2 binding protein 1 (TopBP1), influencing DNA chain breakage and repair processes."

reach
"For instance, studies have shown that TopBP1 is ubiquitinated by the E3 ubiquitin ligase hHYD and deubiquitinated by USP13, while its phosphorylation enhances its recruitment to DNA damage sites [18, 19]."
USP13 affects MAP3K7
| 3 8
USP13 binds MAP3K7.
| 1 8
| 1 8

sparser
"USP13 interacts with TAK1, and represses TAK1 activation though removing ubiquitination."

sparser
"USP13 interacts with TAK1, inhibits TAK1 activation by removing ubiquitination of TAK1, and subsequently inhibits the NF-κB & MAPK signaling pathway activation (Fig.  xref ), thereby treating NAFLD as a latent molecular target."

sparser
"We then investigated whether USP13 physically interacts with TAK1."

sparser
"In the present study, we observed that USP13 interacted with TAK1 and inhibited its activation."

sparser
"Using molecular docking and co-immunoprecipitation techniques, we demonstrated that USP13 binds to TAK1 (Fig.  xref E and F)."

sparser
"Secondly, we did not specifically investigate whether the interaction between USP13 and TAK1 changes under HFD versus ND conditions."

sparser
"Therefore, we deduced that USP13 interacted with TAK1, removed the ubiquitin chain on TAK1, and subsequently reduced TAK1 phosphorylation, followed by activation and downstream events."

reach
"Using molecular docking and co-immunoprecipitation techniques, we demonstrated that USP13 binds to TAK1 (Fig. 6E and F)."

sparser
"Mechanistically, USP13 interacted with TAK1, repressing its K63-linked ubiquitination and phosphorylation, thereby modulating downstream inflammatory and insulin pathways."
USP13 inhibits MAP3K7.
| 2
USP13 inhibits MAP3K7. 2 / 2
| 2

reach
"Because TAK1 activation was induced by K63-linked polyubiquitination [23, 45], we assumed that USP13 inhibited TAK1 activation by reducing K63-associated polyubiquitination."

reach
"USP13 interacts with TAK1, and represses TAK1 activation though removing ubiquitination."
USP13 affects FASN
2 | 3 6
USP13 binds FASN.
2 | 4
2 | 4

sparser
"Mechanistically, USP13 interacts with fatty acid synthase (FASN) and enhances FASN protein stability."

sparser
"USP13 interacts with FASN to enhance its protein stability to promote fatty acid synthesis, cancer stemness and sphere formation. xref Inhibition of FASN resulted in the decrease of lipogenesis, self‐renewal ability and chemotherapy resistance in the cancer cells."

sparser
"We confirmed the interaction between endogenous USP13 and FASN by the co-immunoprecipitation (Co-IP) of H1048 cell lysates ( xref )."

sparser
"To detect the role of USP13-FASN axis in regulating cancer stemness and lipogenesis in SCLC, we stably depleted FASN after WT USP13 reconstitution in USP13 knockdown cells."

No evidence text available

No evidence text available
USP13 increases the amount of FASN.
| 1 2
USP13 increases the amount of FASN. 3 / 3
| 1 2

reach
"USP13 promotes the growth of SCLC cells by maintaining the expression of FASN through the ubiquitin-proteasome pathway."

sparser
"USP13-Dependent FASN Expression Promotes SCLC Stemness and Lipogenesis."

sparser
"In addition, IHC analysis and oil red O staining of tumor tissues indicated that USP13-dependent FASN expression increases lipogenesis ( xref )."
USP13 activates FASN.
| 2
USP13 activates FASN. 2 / 2
| 2

reach
"USP13 has been shown to upregulate fatty acid synthase (FASN) to promote the proliferation of SCLC cells and Twist-related protein 1 (Twist1) to promote the migration and invasion of breast cancer cells, while it inhibits cell proliferation via downregulation of glycolysis in oral squamous cell carcinoma (47,48)."

reach
"Mechanistically, USP13 interacts with fatty acid synthase (FASN) and enhances FASN protein stability."
UBC affects USP13
11 |
11 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

No evidence text available
MAP3K7 affects USP13
| 3 8
MAP3K7 binds USP13.
| 1 8
| 1 8

sparser
"USP13 interacts with TAK1, and represses TAK1 activation though removing ubiquitination."

sparser
"USP13 interacts with TAK1, inhibits TAK1 activation by removing ubiquitination of TAK1, and subsequently inhibits the NF-κB & MAPK signaling pathway activation (Fig.  xref ), thereby treating NAFLD as a latent molecular target."

sparser
"We then investigated whether USP13 physically interacts with TAK1."

sparser
"In the present study, we observed that USP13 interacted with TAK1 and inhibited its activation."

sparser
"Using molecular docking and co-immunoprecipitation techniques, we demonstrated that USP13 binds to TAK1 (Fig.  xref E and F)."

sparser
"Secondly, we did not specifically investigate whether the interaction between USP13 and TAK1 changes under HFD versus ND conditions."

sparser
"Therefore, we deduced that USP13 interacted with TAK1, removed the ubiquitin chain on TAK1, and subsequently reduced TAK1 phosphorylation, followed by activation and downstream events."

reach
"Using molecular docking and co-immunoprecipitation techniques, we demonstrated that USP13 binds to TAK1 (Fig. 6E and F)."

sparser
"Mechanistically, USP13 interacted with TAK1, repressing its K63-linked ubiquitination and phosphorylation, thereby modulating downstream inflammatory and insulin pathways."
MAP3K7 activates USP13.
| 2
MAP3K7 activates USP13. 2 / 2
| 2

reach
"Therefore, inhibiting the TAK1 activation could reverse the exacerbation of NAFLD brought from the deficiency of USP13, indicating that targeting TAK1 is necessary for USP13 to alleviate NAFLD."

reach
"TAK1 is necessary for USP13 to alleviate NAFLD."
ACLY affects USP13
1 | 10
1 | 8

No evidence text available

sparser
"Mechanistically, Spautin-1 blocks the USP13-ACLY axis, thereby causing mitochondrial damage and releasing DAMPs and cytokines, which promotes the infiltration and activation of CD8 + T cells in DLBCL."

sparser
"These data indicate that Spautin-1 induces mitochondrial damage by inhibiting the USP13-ACLY axis, thereby suppressing DLBCL growth."

sparser
"USP13 directly binds to ACLY."

sparser
"First, the interaction between USP13 and ACLY was confirmed through co-immunoprecipitation analysis."

sparser
"Mechanistically, Spautin-1 promotes PANoptosis by inducing mitochondrial damage mediated by USP13-ACLY inhibition in DLBCL cells, which in turn induces the release of injury-related molecular patterns and cytokines."

sparser
"Although our research results indicate that Spautin-1 exerts anti-tumor activity dependent on the induction of PANoptosis, there are still three key limitations that need to be addressed: (1) The mechanism by which Spautin-1 regulates the USP13-ACLY axis has not been thoroughly verified. (2) The expression pattern and prognostic value of USP13 have not yet been verified in clinical samples."

sparser
"Furthermore, to explore the precise regions responsible for the interaction between USP13 and ACLY, we conducted deletion analyses."

sparser
"Likewise, we discovered that ACLY’s CoA ligase domain binds to USP13 (Fig. xref )."
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."
USP13 affects UBL4A
4 | 3 2
USP13 ubiquitinates UBL4A.
| 3 1
USP13 ubiquitinates UBL4A. 4 / 5
| 3 1

reach
"Interestingly, USP13 counteracts the GP78-mediated ubiquitination of Ubl4A through the deubiquitination and stabilization of Ubl4A, thus playing a positive role in the degradation of misfolded proteins [36]."

reach
"USP13 and gp78 control ubiquitination of Ubl4A."

reach
"USP19 deubiquitinates HRD1, rescuing it from UPS-mediated degradation [153], while USP13 works in concert with the ERAD E3 ligase GP78 to maintain precise ubiquitination of the cytosolic ERAD component UBL4A [154]."

sparser
"USP13 and gp78 control ubiquitination of Ubl4A."
USP13 binds UBL4A.
4 | 1
4 | 1

No evidence text available

No evidence text available

No evidence text available

sparser
"Additionally, as catalytic-inactive DUBs often trap their substrates, can a catalytic-inactive USP13 bind preferentially to Ubl4A?"

No evidence text available
JUN affects USP13
9 | 1
JUN decreases the amount of USP13.
9 |
JUN decreases the amount of USP13. 9 / 9
9 |

No evidence text available

No evidence text available

No evidence text available

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JUN inhibits USP13.
| 1
JUN inhibits USP13. 1 / 1
| 1

reach
"USP13 is also downregulated by the autophagy inducer C-JUN/AP-1, and inhibition of apoptosis-related MAPK/JNK signaling effectively downregulates the expression of USP13."
| 5 4
| 5

eidos
"To investigate the molecular mechanisms by which USP13 modulates ECM expression , we focused on the role of USP13 on Smad4 expression ."

eidos
"The results indicate that USP13 promotes ECM expression by stabilizing Smad4 in lung fibroblasts and plays a role in the maintenance of the extracellular matrix in lungs ."

eidos
"Taken together , USP13 deficiency causes loss in extracellular matrix levels in lungs , thereby enlarging alveolar space in response to CS exposure ."

eidos
"28 In this study , we identify that USP13 promotes ECM expression by stabilizing Smad4 in lung fibroblast cells ."

eidos
"We used a mild CS exposure system and found that USP13 deficiency caused loss of extracellular matrix , increased lung inflammation and enlargement of alveolar spaces in response to CS exposure ."
USP13 increases the amount of extracellular matrix.
| 4
USP13 increases the amount of extracellular matrix. 4 / 4
| 4

reach
"In summary, USP13 is a positive regulator of Smad4 stability and promotes ECM expression in lung fibroblast cells."

reach
"To investigate the molecular mechanisms by which USP13 modulates ECM expression, we focused on the role of USP13 on Smad4 expression."

reach
"The results indicate that USP13 promotes ECM expression by stabilizing Smad4 in lung fibroblasts and plays a role in the maintenance of the extracellular matrix in lungs."

reach
"In this study, we identify that USP13 promotes ECM expression by stabilizing Smad4 in lung fibroblast cells."
USP13 affects TXNIP
| 8 1
USP13 deubiquitinates TXNIP.
| 4
USP13 deubiquitinates TXNIP. 4 / 4
| 4

reach
"Treatment with a TXNIP inhibitor, spautin-1, increased TXNIP polyubiquitination (Figure 7B), while overexpression of USP13 decreased TXNIP polyubiquitination (Figure 7C), suggesting USP13 targets and deubiquitinates TXNIP, leading to stabilization of TXNIP.4."

reach
"USP13 deubiquitinates and stabilizes TXNIP in lung fibroblasts."

reach
"This study is the first to reveal that USP13 may deubiquitinate and stabilize TXNIP."

reach
"Mechanistic studies revealed that USP13 targeted and deubiquitinated TXNIP."
USP13 increases the amount of TXNIP.
| 3
USP13 increases the amount of TXNIP. 3 / 3
| 3

reach
"Knockout or knockdown of USP13 in lungs or lung fibroblast cells significantly reduced TXNIP levels."

reach
"Knockdown of USP13 significantly reduced TXNIP levels in lungs or lung fibroblast cells."

reach
"USP13 Deficiency Reduces TXNIP Levels in Lung Cells."
USP13 binds TXNIP.
| 1 1
| 1 1

sparser
"As shown in xref , USP13 was detected in the TXNIP antibody-precipitated complex, suggesting that USP13 is associated with TXNIP."

reach
"The study may therefore provide a molecular basis to develop new small molecules to degrade TXNIP based on TXNIP/USP13 interaction.Two independent studies demonstrated that TXNIP exhibits a pro-fibrotic property in renal fibrosis [25,26], but has been shown to exhibit an anti-fibrotic property in the liver [27]."
USP13 affects SMAD4
1 | 1 5 2
USP13 binds SMAD4.
1 | 1 2
1 | 1 2

sparser
"In addition, we demonstrate that USP13 is associated with Smad4, and the association is not affected by TGF- β 1."

reach
"Interestingly, we also found that while Smad4 is translocated into the nucleus under the stimulation of TGF-β1 (10 ng/mL, 1 hour), USP13-V5 was also observed in the nucleus (Fig 6B, C), suggesting that USP13 is associated with Smad4 and the USP13/Smad4 complex is translocated into nucleus in response to TGF-β1 treatment."

sparser
"Interestingly, we also found that while Smad4 is translocated into the nucleus under the stimulation of TGF- β 1 (10 ng/mL, 1 hour), USP13-V5 was also observed in the nucleus ( xref , C), suggesting that USP13 is associated with Smad4 and the USP13/Smad4 complex is translocated into nucleus in response to TGF- β 1 treatment."

No evidence text available
USP13 activates SMAD4.
| 1 2
USP13 activates SMAD4. 3 / 3
| 1 2

eidos
"Knockdown of USP13 augmented Smad4 degradation and reduced Smad4 half-life ( Fig 6A ) ."

reach
"Knockdown of USP13 augmented Smad4 degradation and reduced Smad4 half-life (Fig 6A)."

reach
"Deubiquitinase USP13 promotes extracellular matrix expression by stabilizing Smad4 in lung fibroblast cells."
USP13 deubiquitinates SMAD4.
| 2
USP13 deubiquitinates SMAD4. 2 / 2
| 2

reach
"Knockdown of USP13 decreased Smad4 half-life and promoted Smad4 ubiquitination."

reach
"USP13 stabilizes Smad4 by deubiquitinating Smad4.."
USP13 affects RAF1
| 6 3
USP13 deubiquitinates RAF1.
| 5
USP13 deubiquitinates RAF1. 5 / 5
| 5

reach
"Sp-1 functions mainly by inhibiting the activity of USP13, which associates with and deubiquitinates RAF1."

reach
"USP13 deubiquitinates Raf1."

reach
"The abovementioned results indicated that USP13 deubiquitinates Raf1 to increase its stability, while Sp-1 antagonizes USP13 function and promotes Raf1 degradation."

reach
"Finally, the ubiquitination of HA-hRAF1 was examined in FLAG-hUSP13-expressing hiPSCs and the western blot results showed that USP13 suppressed RAF1 ubiquitination (Fig. 8H)."

reach
"Sp-1 mainly functions by inhibiting the deubiquitinating protein USP13, which interacts with and thereby deubiquitinates the Raf1 protein in the cells; the latter is a key component in the activation of MEK/ERK signaling (Fig. 9)."
USP13 binds RAF1.
| 1 3
| 1 3

sparser
"USP13 interacts with Raf1."

sparser
"A coimmunoprecipitation assay was performed in Flag-tagged Raf1 (Flag-Raf1)-overexpressing 46C mESCs carrying the HA or HA-USP13 transgene with a Flag antibody-affinity gel, which was used to bind the Flag fusion protein for immunoprecipitation analysis, to determine whether USP13 interacts with Raf1 ( xref A )."

reach
"Mechanistically, we demonstrate that USP13 physically interacts with, deubiquitinates, and stabilizes serine/threonine kinase Raf1 and thereby sustains Raf1 protein at the posttranslational level to activate the FGF/MEK/ERK prodifferentiation signaling pathway in naive mouse ESCs."

sparser
"Co-IP was performed and showed that RAF1 interacted with USP13 in hiPSCs ( xref G )."
USP13 affects cell cycle
| 1 7
USP13 activates cell cycle.
| 1 6
| 1 6

reach
"In addition, reconstitution of WT USP13, but not the T196A mutant, fully rescued the foci formation of the BRCA1-A complex, DNA repair and cell cycle checkpoint in USP13 deficient cells and reversed hypersensitivity to cisplatin or olaparib induced by USP13 deficiency (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"USP13 knockdown inhibits cell cycle progression and cell proliferation in GC."

reach
"USP13 overexpression promotes cell cycle progression and cell proliferation in a DUB enzyme activity dependent manner."

reach
"Furthermore, USP13 was downregulated in both cancer and healthy tissues.3.11 KDELR3 promotes malignant behaviours and modulates the cell cycle of PC cells."

reach
"However, the role of USP13 in modulating cell cycle proteins has not been reported yet."

eidos
"This result indicates that USP13 deficiency or inhibition prevents cell cycle arrest in S phase in response to replication stress ."

reach
"The elevated expression of USP13 promotes the cell cycle progression and cell proliferation by maintaining the stability of cyclin D1."
USP13 inhibits cell cycle.
| 1
| 1

reach
"Cotargeting of EGFR and USP13 led to a robust increase in treatment efficacy both in vitro and in vivo, mainly due to a strong induction of apoptosis rather than enhancing the cell cycle arrest caused by afatinib alone."
USP13 affects UBA1
| 8
| 5

sparser
"Although the experimental results showed that USP13-UBA could bind ubiquitin, USP13 still exhibited only weak deubiquitination enzyme activity, which is incompatible with the findings that USP13 can deubiquitinate various important substrates implicated in disease and tumor development ( xref ; xref )."

sparser
"In 2011, Hu et al. reported the solution structure of USP13-UBA (PDB 2LBC)."

sparser
"Albeit no ubiquitin-bound UBA structure is resolved, sequence alignment revealed that USP13-UBA contains an MGF motif that presumably can bind Ub ( xref ), consistent with results from pull-down and ITC experiments ( xref )."

sparser
"In addition, NMR titration data reflected that M664, F666, M739, and F741 might be the key residues responsible for the binding of USP13-UBA to Ub ( xref )."

sparser
"Overall, the USP13-ZnF domain cannot bind to Ub to activate USP13, whereas USP13-UBA can bind, which may partially explain the reason why USP13 displays only weak basal deubiquitination enzyme activity: the binding sites of USP13 to Ub are less than that other USP members, thus providing weaker binding affinity and consequent cleavage activity towards ubiquitin chains; There possibly exists constitutive self-inhibition for full-length USP13 supported by the interaction of UBA with ZnF domain, which is hypothesized to be released by recruitment of other proteins or modification, such as phosphorylation."
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
USP13 affects TLR4
1 | 4 3
USP13 binds TLR4.
1 | 1 3
1 | 1 3

sparser
"USP13 interacts with TLR4 and inhibits the ubiquitin-mediated degradation of TLR4 ( xref )."

reach
"USP13 interacted with TLR4 and inhibited the ubiquitin mediated degradation of TLR4."

No evidence text available

sparser
"USP13 interacted with TLR4 and inhibited the ubiquitin-mediated degradation of TLR4."

sparser
"The co-IP assay revealed that USP13 directly interacted with TLR4 in Hep3B cells ( xref )."
USP13 deubiquitinates TLR4.
| 3
USP13 deubiquitinates TLR4. 3 / 3
| 3

reach
"In more detail, Gao et al. demonstrated that USP13 potentiated the deubiquitination and the stabilization of TLR4, activating the TLR4/MyD88/NF-κB pathway under hypoxic conditions."

reach
"Hypoxia-induced ubiquitin-specific peptidase 13 (USP13) deubiquitinated and stabilized toll-like receptor 4 (TLR4) to activate the myeloid differentiation primary response gene 88/nuclear factor-κB (MyD88/NF-κB) pathway, thereby contributing to HCC progression (Gao et al., 2020)."

reach
"USP13 knockdown increases TLR4 ubiquitination, leading to its decreased expression, which results in the decreased expression of MYD88, phosphorylated-NF-κB and p65, thus inhibiting the proliferation and invasion of HCC cells."
USP13 affects SMC3
7 | 1
USP13 binds SMC3.
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
USP13 decreases the amount of SMC3.
| 1
USP13 decreases the amount of ubiquitinated SMC3. 1 / 1
| 1

reach
"Ectopic overexpression of USP13 significantly decreased the levels of endogenous ubiquitinated SMC3 and RAD21, indicating that USP13 can function as a deubiquitinase for cohesin subunits."
USP13 affects SKP2
1 1 | 3 3
USP13 binds SKP2.
1 | 1 3
1 | 1 3

sparser
"UFD1 was reported to function as a scaffold for the USP13-Skp2 (the F-box adaptor of the E3 ubiquitin ligase SCF Skp2 ) interaction, maintaining Skp2 stability."

reach
"We found that SKP2 interacted with USP10, USP13, UCHL5, USP14, and USP7 in KBM5-T315I cells."

sparser
"In HeLa cells, the ubiquitin-recognition protein Ufd1 was reported as a scaffold protein for SKP2USP13 interaction to facilitate SKP2 deubiquitylation by USP13."

sparser
"Prolonged endoplasmic reticulum stress downregulated Ufd1 to attenuate the USP13SKP2 interaction, leading to SKP2 destabilization and p27 accumulation."

No evidence text available
USP13 deubiquitinates SKP2.
1 | 2
USP13 deubiquitinates SKP2. 3 / 3
1 | 2

reach
"We reported here that USP2 is a new deubiquitylase of SKP2 E3 ligase, which interestingly stabilized both SKP2 and SKP2 substrates.Two DUBs, USP10 and USP13, were previously reported to deubiquitylating SKP2."

No evidence text available

reach
"On the other hand, USP10 (28) and USP13 (29) were reported to promote SKP2 deubiquitylation."
USP13 affects SIGIRR
| 5 3
USP13 binds SIGIRR.
| 3 3
| 3 3

reach
"To determine molecular mechanisms by which GSK3β promotes Sigirr degradation, we examined the effect of GSK3β-mediated phosphorylation on the association between Sigirr and USP13."

reach
"Overexpression of GSK3β reduced the association between Sigirr and USP13 (Fig. 7B), suggesting that GSK3β-mediated phosphorylation prevents Sigirr degradation from Sigirr interaction with its stabilizer, USP13."

sparser
"USP13 is associated with Sigirr."

sparser
"Co-immunoprecipitation (Co-IP) also revealed USP13's association with Sigirr ( xref b, c)."

reach
"Indeed, the T372 site is within the USP13 binding site, thus GSK3β-mediated phosphorylation of this residue modifies the conformation preventing IL-1R8/USP13 interaction and IL-1R8 stabilization ( Fig[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Overexpression of GSK3β reduced the association between Sigirr and USP13 ( xref ), suggesting that GSK3β-mediated phosphorylation prevents Sigirr degradation from Sigirr interaction with its stabilizer, USP13."
USP13 deubiquitinates SIGIRR.
| 2
USP13 deubiquitinates SIGIRR. 2 / 2
| 2

reach
"We have shown that deubiquitination of Sigirr by USP13 increased Sigirr half-life (L. Li et al., 2019)."

reach
"The ubiquitination and degradation of Sigirr is negatively regulated by a deubiquitinase, USP13 (L. Li et al., 2019)."
USP13 affects MTOR
| 8
USP13 activates MTOR. 8 / 8
| 8

reach
"To confirm USP13 accelerates the proliferation of HCC cells through regulating PRPF6‐AKT‐mTOR signalling, we performed the rescue experiments by silencing PRPF6 in USP13 up‐regulated cells."

reach
"It was found that knock down of PRPF6 effectively inhibited the cell proliferation and also reduced the expressions of p‐AKT and p‐mTOR induced by overexpression of USP13 (Figures 7A,B)."

reach
"These results suggest that the USP13‐PRPF6 axis promotes HCC cell proliferation by regulating AKT‐mTOR signalling.3.6 USP13 Expression Is Correlated With PRPF6 in Human HCC Tissues."

reach
"Overall, USP13 facilitates the malignant progression of HCC cells through the upregulation of WWP1.Thus, we concluded that USP13 stabilized WWP1, and then activated the Akt/mTOR pathway."

reach
"Further experimental results confirmed that the USP13‐PRPF6 axis promoted the proliferation of HCC cells by modulating the AKT‐mTOR signalling pathway.In conclusion, we have demonstrated that USP13 can decrease the K48/63‐linked polyubiquitination of PRPF6, thereby stabilising this protein and promoting HCC cell proliferation via regulation of the AKT‐mTOR pathway (Figure 8C)."

reach
"Moreover, USP13 stabilized WWP1 by removing the K29- and K48-linked polyubiquitination chains from WWP1 and then activated the Akt/mTOR pathway in HCC."

reach
"By acting as an oncogene, USP13 affects HCC malignant progression through the WWP1-mediated Akt/mTOR pathway."

reach
"Importantly, USP13 modulated the AKT‐mTOR signalling pathway in a manner analogous to PRPF6 (Figures 6D,E)."
USP13 affects GST
| 8
| 8

sparser
"To detect the direct interaction between USP13 and cyclin D1, we performed GST-pulldown assays with the recombinant GST-USP13 protein."

sparser
"Moreover, we found that GST-USP13 mainly interacted with the full length (FL) and the 169–202 amino acid fragment (Twist1 WR domain) of GFP-Twist1 rather than with other Twist1 fragments (Fig.  xref )."

sparser
"Meanwhile, the results from the pull-down assay showed that purified GST-USP13 directly interacted with endogenous Raf1 in 46C ESC lysates ( xref E )."

sparser
"The recombinant GST or GST-USP13 full-length was expressed from E. coli BL21 and immobilized on glutathione-Sepharose beads."

sparser
"In vitro kinase assays were performed directly on the GST or GST-USP13 beads with active CK2 enzyme and ATP."

sparser
"Western blotting with an anti-pT122 antibody showed that CK2 phosphorylated GST-USP13 protein but not GST, indicating that USP13 is a substrate of CK2 in vitro ( xref D), suggesting that USP13 is a substrate for CK2."

sparser
"The results showed that cyclin D1 can be pulled down by purified GST-USP13 protein (Fig. xref )."

sparser
"Using GST-pulldown assays, we found that Flag-Twist1 co-precipitated with GST-USP13, but not with GST (Fig.  xref )."
UBA1 affects USP13
| 8
| 5

sparser
"Although the experimental results showed that USP13-UBA could bind ubiquitin, USP13 still exhibited only weak deubiquitination enzyme activity, which is incompatible with the findings that USP13 can deubiquitinate various important substrates implicated in disease and tumor development ( xref ; xref )."

sparser
"In 2011, Hu et al. reported the solution structure of USP13-UBA (PDB 2LBC)."

sparser
"Albeit no ubiquitin-bound UBA structure is resolved, sequence alignment revealed that USP13-UBA contains an MGF motif that presumably can bind Ub ( xref ), consistent with results from pull-down and ITC experiments ( xref )."

sparser
"In addition, NMR titration data reflected that M664, F666, M739, and F741 might be the key residues responsible for the binding of USP13-UBA to Ub ( xref )."

sparser
"Overall, the USP13-ZnF domain cannot bind to Ub to activate USP13, whereas USP13-UBA can bind, which may partially explain the reason why USP13 displays only weak basal deubiquitination enzyme activity: the binding sites of USP13 to Ub are less than that other USP members, thus providing weaker binding affinity and consequent cleavage activity towards ubiquitin chains; There possibly exists constitutive self-inhibition for full-length USP13 supported by the interaction of UBA with ZnF domain, which is hypothesized to be released by recruitment of other proteins or modification, such as phosphorylation."
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
MDM2 affects USP13
| 2 6
| 2 6

sparser
"USP13 is associated with MDM2."

sparser
"These experimental results indicate that USP13 interacts with MDM2 in the nucleus."

sparser
"The Co-IP results demonstrate that USP13 is associated with MDM2 ( xref and xref )."

sparser
"To further confirm the interaction between USP13 and MDM2, Co-IP experiments after over-expression of exogenous genes were conducted."

sparser
"USP13 interacts with MDM2 and removes K63-linked ubiquitin chains on MDM2."

sparser
"The results demonstrate that exogenously expressed USP13 and MDM2 proteins are associated each other ( xref and xref )."

reach
"To further confirm the interaction between USP13 and MDM2, Co-IP experiments after over-expression of exogenous genes were conducted."

reach
"Considering the direct association between USP13 and MDM2 and the role of USP13 in the degradation of MDM2, we examined if USP13 deubiquitinates MDM2."
GST affects USP13
| 8
| 8

sparser
"To detect the direct interaction between USP13 and cyclin D1, we performed GST-pulldown assays with the recombinant GST-USP13 protein."

sparser
"Moreover, we found that GST-USP13 mainly interacted with the full length (FL) and the 169–202 amino acid fragment (Twist1 WR domain) of GFP-Twist1 rather than with other Twist1 fragments (Fig.  xref )."

sparser
"Meanwhile, the results from the pull-down assay showed that purified GST-USP13 directly interacted with endogenous Raf1 in 46C ESC lysates ( xref E )."

sparser
"The recombinant GST or GST-USP13 full-length was expressed from E. coli BL21 and immobilized on glutathione-Sepharose beads."

sparser
"In vitro kinase assays were performed directly on the GST or GST-USP13 beads with active CK2 enzyme and ATP."

sparser
"Western blotting with an anti-pT122 antibody showed that CK2 phosphorylated GST-USP13 protein but not GST, indicating that USP13 is a substrate of CK2 in vitro ( xref D), suggesting that USP13 is a substrate for CK2."

sparser
"The results showed that cyclin D1 can be pulled down by purified GST-USP13 protein (Fig. xref )."

sparser
"Using GST-pulldown assays, we found that Flag-Twist1 co-precipitated with GST-USP13, but not with GST (Fig.  xref )."
USP13 affects SQSTM1
4 | 3
USP13 binds SQSTM1.
4 | 1
4 | 1

No evidence text available

reach
"In this study, we found that ubiquitin-specific protease 13 (USP13), a member of the USP family, directly binds p62 and removes ubiquitin at Lys7 (K7) of the PB1 domain."

No evidence text available

No evidence text available

No evidence text available
USP13 deubiquitinates SQSTM1.
| 2
USP13 deubiquitinates SQSTM1. 2 / 2
| 2

reach
"USP13 deubiquitinates p62/SQSTM1 to induce autophagy and Nrf2 release for activating antioxidant response genes."

reach
"USP13 deubiquitinates p62/SQSTM1 to induce autophagy and Nrf2 release for activating antioxidant response genes."
USP13 affects SMC1A
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
USP13 affects PXN
2 | 5
2 | 5

sparser
"Furthermore, Paxillin and Paxillin(Y118A) interacted with USP13, but Paxillin(Y31A) failed to associate with USP13 (Fig.  xref l), implicating that Paxillin Y31 phosphorylation is important for USP13-Paxillin interaction."

sparser
"USP13 interacts with Paxillin and NLRP3, phosphorylation of Paxillin at Y31 is important for Paxillin-USP13 interaction, and USP13 is essential for Paxillin-mediated NLRP3 deubiquitination upon ATP treatment."

sparser
"Moreover, Co-IP assays showed that Paxillin and USP13 interacted with each other (Fig.  xref d, e)."

sparser
"Moreover, the domains of USP13 required for USP13-NLRP3 interaction and USP13-Paxillin association were assessed by constructing and analyzing plasmids encoding for WT USP13(1–863) and four truncated proteins (Fig.  xref i)."

No evidence text available

sparser
"Additionally, USP13(1–863) and USP13(1–300) interacted with Paxillin, but USP13(301–624), USP13(301–863), and USP13(625–863) could not interact with Paxillin (Fig.  xref k), indicating that 1aa–300aa of USP13 is involved in USP13-Paxillin interaction."

No evidence text available
USP13 affects MDC1
3 | 1 3
3 | 1 3

sparser
"In addition, USP13 interacted with MDC1 and the interaction between USP13 and MDC1 was dramatically increased following cisplatin treatment ( xref )."

No evidence text available

sparser
"Moreover, we examined the interaction between USP13 and different deletion mutants of MDC1 and found that deletion of FHA domain of MDC1 abolishes the USP13-MDC1 interaction ( xref )."

reach
"In addition, USP13 interacted with MDC1 and the interaction between USP13 and MDC1 was dramatically increased following cisplatin treatment (XREF_FIG)."

No evidence text available

sparser
"To further demonstrate that the FHA domain of MDC1 is critical for USP13-MDC1 interaction, we purified GST-MDC1-FHA fragment from bacteria and performed pull-down assay."

No evidence text available
USP13 affects MAP2K3
| 7
| 7

sparser
"Our understanding of the USP13-MKK3 axis in this study primarily focuses on CRC, while its role in other cancers remains insufficiently explored."

sparser
"Meanwhile, blocking ubiquitination at the K32 site of MKK3 (K32R mutation) mimics the USP13 overexpression phenotype, further confirming that this site is the core target of the USP13-MKK3 regulatory axis."

sparser
"Additionally, whether the USP13-MKK3 axis can be effectively combined with existing targeted therapies (such as MEK, PI3K, and BRAF inhibitors) or immunotherapies (such as PD-1/PD-L1 antibodies) to enhance therapeutic efficacy requires further clinical trial data."

sparser
"USP13 interact with MKK3."

sparser
"Specifically, USP13 directly interacts with the C-terminal domain of MKK3 through its UBA domain and, relying on its deubiquitinase activity, selectively removes K48-linked ubiquitination (especially at the K32 site) on MKK3, thereby inhibiting its degradation via the proteasome."

sparser
"Finally, we demonstrated that p38 activation is both sufficient and necessary for the tumor-promoting effects downstream of the USP13-MKK3 axis."

sparser
"Therefore, further investigation is needed to elucidate the specific roles and mechanisms of the USP13-MKK3 axis in different cancers."
USP13 affects HIF1A
| 7
| 7

sparser
"Luciferase assays revealed that HIF-1α specifically binds to Region #2 of the USP13 promoter (Fig. xref )."

sparser
"USP13 directly bound HIF-1α via its C-terminal UBA domain, cleaving K48-linked polyubiquitin chains to stabilize HIF-1α and enhance its transcriptional activity."

sparser
"USP13-HIF-1α interaction was assessed via co-immunoprecipitation (co-IP), immunofluorescence, and domain mapping."

sparser
"In this study, we found that the HIF-1α K532R mutation significantly reduced the interaction between HIF-1α and USP13 (Fig.  xref B)."

sparser
"USP13 interacts with HIF-1α."

sparser
"Subsequently, we carried out co-IP experiments to verify the interaction between HIF-1α and USP13."

sparser
"The interaction between USP13 and HIF-1α was also validated in tumor tissues formed by MC38 cells and in CRC model (Fig. S5)."
USP13 affects CTNNB1
4 | 3
USP13 binds CTNNB1.
4 | 1
4 | 1

No evidence text available

No evidence text available

No evidence text available

No evidence text available

reach
"We demonstrated that USP13 interacts with and deubiquitinates β-catenin, which promotes NSCLC metastasis."
USP13 activates CTNNB1.
| 2
USP13 activates CTNNB1. 2 / 2
| 2

reach
"Furthermore, knockdown of USP13 or WISP1 impaired the activation of the Wnt/CTNNB1 signaling pathway and reduced immune checkpoint marker expression, indicating a mechanism by which USP13 promotes immune evasion in ESCC."

reach
"As shown in Figures 2 F and 2G, luciferase activity of TOP reporter, but not FOP reporter, was significantly decreased in USP13-depleted cells, indicating USP13 might positively regulate β-catenin act[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP13 affects CLK3
4 | 1 2
4 | 1 2

No evidence text available

sparser
"CLK3 interacts with USP13, phosphorylates it at the Tyr708 residue and inhibits the ubiquitination of c-Myc to activate c-Myc-mediated purine biosynthesis, thus promoting the progression of bile duct cell carcinoma ( xref )."

reach
"A pulldown experiment suggests a direct binding of USP13 to CLK3 (XREF_FIG)."

No evidence text available

No evidence text available

sparser
"A pulldown experiment suggests a direct binding of USP13 to CLK3 ( xref )."

No evidence text available
USP13 affects CDH1
| 1 3 3
USP13 binds CDH1.
| 2 3
| 2 3

sparser
"We performed immunoprecipitation assays followed by western-blotting to assess the interaction between USP13 and CDH1."

sparser
"We found that USP13 binds to the APC/C adaptor CDH1."

reach
"We performed immunoprecipitation assays followed by western-blotting to assess the interaction between USP13 and CDH1."

reach
"We found that USP13 binds to the APC/C adaptor CDH1."

sparser
"Shortly after this finding, the interaction of USP13 with CDH1 was also confirmed by the same team ( xref )."
USP13 activates CDH1.
| 1 1
USP13 activates CDH1. 2 / 2
| 1 1

reach
"USP13 modulates the stability of the APC/C adaptor CDH1."

eidos
"In addition , we report for the first time that USP13 controls CDH1 protein levels in cells : overexpression of USP13 increased CDH1 levels , whereas depletion of USP13 decreased CDH1 levels ."
USP13 affects AE
| 7
AE binds USP13. 7 / 7
| 7

sparser
"In HCT15 cells, USP13 overexpression increased DNA synthesis, but the USP13-AE mutant did not (Fig.  xref g)."

sparser
"Knockdown of USP13 increased MKK3 ubiquitination (Fig.  xref a), and wild-type USP13, but not the catalytically inactive USP13-AE mutant, effectively reduced this modification (Fig.  xref b)."

sparser
"In HCT116 cells, USP13 overexpression enhanced migration, but the USP13-AE mutant did not (Fig.  xref i)."

sparser
"To determine whether USP13 contributes to malignant phenotypes, we established HCT116 and HCT15 cell lines stably overexpressing wild-type USP13 (USP13-WT) or its catalytically inactive mutant USP13-AE (C345A/M664/739E) (Fig.  xref a), and generated USP13-knockout RKO and SW48 cells via CRISPR/Cas9 (Fig.  xref b)."

sparser
"Furthermore, The interaction was dependent on USP13 catalytic activity, as the USP13-AE mutant displayed reduced binding (Fig.  xref g)."

sparser
"USP13 overexpression promoted colony formation, whereas the USP13-AE mutant had no such effect (Fig.  xref e-f)."

sparser
"Moreover, CHX chase assay demonstrated that overexpression of wild-type USP13, but not the USP13-AE mutant, resulted in an extension of MKK3's half-life and a delay in its degradation (Fig.  xref k)."
TOPBP1 affects USP13
2 | 1 4
2 | 1 4

No evidence text available

No evidence text available

sparser
"USP13 binds to and stabilizes TopBP1 through deubiquitination and orchestrates the replication stress response [ xref ]."

sparser
"Mechanistically, USP13 binds to TopBP1 and stabilizes TopBP1 by deubiquitination."

reach
"Mechanistically, USP13 binds to TopBP1 and stabilizes TopBP1 by deubiquitination."

sparser
"To test whether USP13 is a deubiquitinase for TopBP1, we first examined whether USP13 interacts with TopBP1."

sparser
"Our study shows that USP13 interacts with TopBP1."
SMC3 affects USP13
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
SMC1A affects USP13
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PXN affects USP13
2 | 5
2 | 5

sparser
"Furthermore, Paxillin and Paxillin(Y118A) interacted with USP13, but Paxillin(Y31A) failed to associate with USP13 (Fig.  xref l), implicating that Paxillin Y31 phosphorylation is important for USP13-Paxillin interaction."

sparser
"USP13 interacts with Paxillin and NLRP3, phosphorylation of Paxillin at Y31 is important for Paxillin-USP13 interaction, and USP13 is essential for Paxillin-mediated NLRP3 deubiquitination upon ATP treatment."

sparser
"Moreover, Co-IP assays showed that Paxillin and USP13 interacted with each other (Fig.  xref d, e)."

sparser
"Moreover, the domains of USP13 required for USP13-NLRP3 interaction and USP13-Paxillin association were assessed by constructing and analyzing plasmids encoding for WT USP13(1–863) and four truncated proteins (Fig.  xref i)."

No evidence text available

sparser
"Additionally, USP13(1–863) and USP13(1–300) interacted with Paxillin, but USP13(301–624), USP13(301–863), and USP13(625–863) could not interact with Paxillin (Fig.  xref k), indicating that 1aa–300aa of USP13 is involved in USP13-Paxillin interaction."

No evidence text available
PRPF6 affects USP13
| 1 6
| 1 6

sparser
"Furthermore, the USP13PRPF6 axis facilitates HCC cell proliferation via modulation of the mTOR signalling pathway."

sparser
"Further experimental results confirmed that the USP13PRPF6 axis promoted the proliferation of HCC cells by modulating the AKT‐mTOR signalling pathway."

reach
"The co‐IP experiment was initially employed to validate the interaction between USP13 and PRPF6."

sparser
"The co‐IP experiment was initially employed to validate the interaction between USP13 and PRPF6."

sparser
"These results suggest that the USP13PRPF6 axis promotes HCC cell proliferation by regulating AKT‐mTOR signalling."

sparser
"To further explore the signalling pathways underlying the effect of the USP13PRPF6 axis on HCC cell proliferation, bioinformatic analysis was performed."

sparser
"Ultimately, we demonstrated that the USP13PRPF6 axis promoted HCC cell proliferation was closely associated with the activation of the AKT‐mTOR signalling pathway."
MDC1 affects USP13
3 | 1 3
3 | 1 3

sparser
"In addition, USP13 interacted with MDC1 and the interaction between USP13 and MDC1 was dramatically increased following cisplatin treatment ( xref )."

No evidence text available

sparser
"Moreover, we examined the interaction between USP13 and different deletion mutants of MDC1 and found that deletion of FHA domain of MDC1 abolishes the USP13-MDC1 interaction ( xref )."

reach
"In addition, USP13 interacted with MDC1 and the interaction between USP13 and MDC1 was dramatically increased following cisplatin treatment (XREF_FIG)."

No evidence text available

sparser
"To further demonstrate that the FHA domain of MDC1 is critical for USP13-MDC1 interaction, we purified GST-MDC1-FHA fragment from bacteria and performed pull-down assay."

No evidence text available
MAP2K3 affects USP13
| 7
| 7

sparser
"Our understanding of the USP13-MKK3 axis in this study primarily focuses on CRC, while its role in other cancers remains insufficiently explored."

sparser
"Meanwhile, blocking ubiquitination at the K32 site of MKK3 (K32R mutation) mimics the USP13 overexpression phenotype, further confirming that this site is the core target of the USP13-MKK3 regulatory axis."

sparser
"Additionally, whether the USP13-MKK3 axis can be effectively combined with existing targeted therapies (such as MEK, PI3K, and BRAF inhibitors) or immunotherapies (such as PD-1/PD-L1 antibodies) to enhance therapeutic efficacy requires further clinical trial data."

sparser
"USP13 interact with MKK3."

sparser
"Specifically, USP13 directly interacts with the C-terminal domain of MKK3 through its UBA domain and, relying on its deubiquitinase activity, selectively removes K48-linked ubiquitination (especially at the K32 site) on MKK3, thereby inhibiting its degradation via the proteasome."

sparser
"Finally, we demonstrated that p38 activation is both sufficient and necessary for the tumor-promoting effects downstream of the USP13-MKK3 axis."

sparser
"Therefore, further investigation is needed to elucidate the specific roles and mechanisms of the USP13-MKK3 axis in different cancers."
HIF1A affects USP13
| 7
| 7

sparser
"Luciferase assays revealed that HIF-1α specifically binds to Region #2 of the USP13 promoter (Fig. xref )."

sparser
"USP13 directly bound HIF-1α via its C-terminal UBA domain, cleaving K48-linked polyubiquitin chains to stabilize HIF-1α and enhance its transcriptional activity."

sparser
"USP13-HIF-1α interaction was assessed via co-immunoprecipitation (co-IP), immunofluorescence, and domain mapping."

sparser
"In this study, we found that the HIF-1α K532R mutation significantly reduced the interaction between HIF-1α and USP13 (Fig.  xref B)."

sparser
"USP13 interacts with HIF-1α."

sparser
"Subsequently, we carried out co-IP experiments to verify the interaction between HIF-1α and USP13."

sparser
"The interaction between USP13 and HIF-1α was also validated in tumor tissues formed by MC38 cells and in CRC model (Fig. S5)."
AE affects USP13
| 7
AE binds USP13. 7 / 7
| 7

sparser
"In HCT15 cells, USP13 overexpression increased DNA synthesis, but the USP13-AE mutant did not (Fig.  xref g)."

sparser
"Knockdown of USP13 increased MKK3 ubiquitination (Fig.  xref a), and wild-type USP13, but not the catalytically inactive USP13-AE mutant, effectively reduced this modification (Fig.  xref b)."

sparser
"In HCT116 cells, USP13 overexpression enhanced migration, but the USP13-AE mutant did not (Fig.  xref i)."

sparser
"To determine whether USP13 contributes to malignant phenotypes, we established HCT116 and HCT15 cell lines stably overexpressing wild-type USP13 (USP13-WT) or its catalytically inactive mutant USP13-AE (C345A/M664/739E) (Fig.  xref a), and generated USP13-knockout RKO and SW48 cells via CRISPR/Cas9 (Fig.  xref b)."

sparser
"Furthermore, The interaction was dependent on USP13 catalytic activity, as the USP13-AE mutant displayed reduced binding (Fig.  xref g)."

sparser
"USP13 overexpression promoted colony formation, whereas the USP13-AE mutant had no such effect (Fig.  xref e-f)."

sparser
"Moreover, CHX chase assay demonstrated that overexpression of wild-type USP13, but not the USP13-AE mutant, resulted in an extension of MKK3's half-life and a delay in its degradation (Fig.  xref k)."
Barium(2+) affects USP13
| 6
Barium(2+) activates USP13.
| 3
| 3

reach
"The results showed that BA significantly induced apoptosis in both EV and USP13‐OE MM cells (Figure 6E)."

reach
"These findings suggest that combining BA with a PARP inhibitor can effectively counteract the resistance to BA induced by USP13 overexpression.3.7 BA Hampers MM Progression in Mouse Models."

reach
"CCK‐8 assay and flow cytometry analyses demonstrated that the combination of BA and olaparib effectively inhibited cell proliferation and enhanced cell apoptosis in USP13‐OE MM cells compared to cells treatment with BA alone (Figure 6F,G)."
Barium(2+) inhibits USP13.
| 2
| 2

reach
"More importantly, BA effectively suppresses the growth and progression of MM in vitro and in vivo by targeting USP13/PARP1 signalling, implying its potential application in the treatment of MM solely or in combination with current treatment options."

reach
"Our study provides novel insights into the anticancer effects of BA on MM, as it appears to disrupt DDR in MM cells by targeting the USP13/PARP1 axis and inducing cellular apoptosis."
Barium(2+) decreases the amount of USP13.
| 1
Barium(2+) decreases the amount of USP13. 1 / 1
| 1

reach
"The results showed that BA significantly reduced the protein expression of USP13 (Figure 6A)."
ZHX2 affects USP13
2 | 2 2
2 | 2 2

No evidence text available

No evidence text available

reach
"Next, we validated semiendogenous interaction between ZHX2 and USP13 by using reciprocal immunoprecipitations (Fig. 1 C and D)."

reach
"With reciprocal immunoprecipitations, we confirmed the endogenous binding between ZHX2 and USP13 in all of these cell lines (Fig. 1 E and F)."

sparser
"Together, this suggests a significant clinical association of ZHX2-USP13 in RCC patients ( P < 0.001) ( xref )."

sparser
"Next, we validated semiendogenous interaction between ZHX2 and USP13 by using reciprocal immunoprecipitations ( xref )."
USP13 affects cell growth
| 6
| 5

reach
"USP13 promoted HCC cell growth and metastasis by regulating the TLR4/MyD88/NF-κB pathway [37]."

reach
"In HCC cell lines, knockdown of USP13 by shRNAs significantly reduced cell growth, and decreased levels of c‐Myc."

reach
"Colony formation assay revealed that loss of USP13 slightly decreased cell growth capacity ( Figure S1 C), whereas we detected that USP13 knockout (KO) or silence remarkably inhibited cell invasion in[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"USP13 promotes AML cell growth and autophagy by upregulating ATG5."

reach
"In contrast, MHY effectively increased HCC cell proliferation and rescued the cell growth inhibition induced by USP13 knockdown (Supplementary Fig. S5B–D)."
Mutated USP13 activates cell growth. 1 / 1
| 1

reach
"This enhancement was dependent on USP13 deubiquitinase activity, as a catalytically inactive mutant of USP13 (USP13 C345A) failed to increase cell growth."
USP13 affects VCL
| 4 2
USP13 increases the amount of VCL.
| 4
USP13 increases the amount of VCL. 4 / 4
| 4

reach
"Among the identified proteins, the vinculin protein level was upregulated by USP13 through an enzymatically mediated mechanism, which was confirmed by western blot analysis and RT-qPCR."

reach
"These results suggest that USP13 upregulates vinculin protein levels, but not the mRNA levels, through an enzymatically mediated mechanism."

reach
"USP13 upregulates vinculin protein levels, but not mRNA levels."

reach
"To further validate the upregulation of vinculin protein levels by USP13, western blot analysis and RT-qPCR analysis were performed."
USP13 binds VCL.
| 2
| 2

sparser
"Among the identified proteins, the altered expression level of vinculin was associated with the DUB activity of USP13, which was confirmed by western blot analysis and reverse transcription-quantitative polymerase chain reaction (RT-qPCR)."

sparser
"However, sufficient follow-up validation studies are required in order to determine whether vinculin protein directly interacts with USP13."
| 6
| 4

reach
"USP13 promotes migration and invasion of GC cells by maintaining the stability of Snail."

reach
"To investigate whether USP13 knockdown suppressed the tumorigenesis of GC cells in vivo, we injected MKN-45 GC cells which were stably knocked-down USP13 (shUSP13) or the negative control (shNC) into the subcutaneous of the nude mice."

reach
"USP13 knockdown impaired the tumorigenesis of GC cells in vivo."

reach
"USP13 knockdown inhibits cell cycle progression and cell proliferation in GC."
USP13 deubiquitinates Stomach Neoplasms.
| 2
| 2

reach
"The report from Zhang et al. revealed that USP13 promotes the epithelial-mesenchymal transition (EMT) and metastasis in GC by deubiquitinating and stabilizing Snail in GC [10]."

reach
"USP13 interacted with Snail to deubiquitinate and stabilize Snail in GC cells."
USP13 affects NFKBIA
| 1 5
| 1 5

reach
"USP13 binds to and stabilizes IkappaBalpha."

sparser
"The findings showed that USP13 interacts with IκBα and inhibits the ubiquitination of IκBα, leading to reduced degradation of IκBα in vitro and in vivo."

sparser
"USP13 binds to and stabilizes IκBα."

sparser
"Furthermore, the interaction between exogenous USP13 and IκBα was investigated and confirmed in HEK 293T cells (Fig.  xref c, d)."

sparser
"In all, above findings show that USP13 binds to IκBα."

sparser
"USP13 binds to IκBα, therefore, the effect of downregulating USP13 in BV2 microglia on IκBα expression level was further investigated."
USP13 affects FBXL14
| 1 5
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
| 1 1

sparser
"The antagonistic actions between ubiquitin ligases and DUBs are responsible for protein homeostasis as exemplified that USP13-FBXL14 regulated reversible ubiquitination of c-Myc in glioblastoma stem cells [ xref ], and USP15-SMURF2 cooperatively modulated TGF-β receptor in glioblastoma [ xref ]."

reach
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag tagged USP13 or FBXL14."
USP13 affects ALDOA
| 6
| 6

sparser
"In conclusion, these results show that USP13 binds to ALDOA directly via its USP domain."

sparser
"The interaction of USP13 with the ALDOA prompts the question of whether USP13 exerts a regulatory effect on the ALDOA."

sparser
"USP13 directly interacts with ALDOA."

sparser
"The interaction between ALDOA and USP13 was also validated using the heart tissue ( xref E)."

sparser
"Next, we replace the lysine at the three sites with arginine and observed that ALDOA K13R , ALDOA K87R and ALDOA K139R were still bound with USP13 ( xref J)."

sparser
"Subsequently, to further explore the interaction between USP13 and ALDOA, we transfected plasmids encoding Flag-USP13 and Ha-ALDOA into 293T cells and found the interaction between USP13 and ALDOA with co-immunoprecipitation approach ( xref F and G)."
SIGIRR affects USP13
| 3 3
| 3 3

reach
"To determine molecular mechanisms by which GSK3β promotes Sigirr degradation, we examined the effect of GSK3β-mediated phosphorylation on the association between Sigirr and USP13."

reach
"Overexpression of GSK3β reduced the association between Sigirr and USP13 (Fig. 7B), suggesting that GSK3β-mediated phosphorylation prevents Sigirr degradation from Sigirr interaction with its stabilizer, USP13."

sparser
"USP13 is associated with Sigirr."

sparser
"Co-immunoprecipitation (Co-IP) also revealed USP13's association with Sigirr ( xref b, c)."

reach
"Indeed, the T372 site is within the USP13 binding site, thus GSK3β-mediated phosphorylation of this residue modifies the conformation preventing IL-1R8/USP13 interaction and IL-1R8 stabilization ( Fig[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Overexpression of GSK3β reduced the association between Sigirr and USP13 ( xref ), suggesting that GSK3β-mediated phosphorylation prevents Sigirr degradation from Sigirr interaction with its stabilizer, USP13."
NFKBIA affects USP13
| 1 5
| 1 5

reach
"USP13 binds to and stabilizes IkappaBalpha."

sparser
"The findings showed that USP13 interacts with IκBα and inhibits the ubiquitination of IκBα, leading to reduced degradation of IκBα in vitro and in vivo."

sparser
"USP13 binds to and stabilizes IκBα."

sparser
"Furthermore, the interaction between exogenous USP13 and IκBα was investigated and confirmed in HEK 293T cells (Fig.  xref c, d)."

sparser
"In all, above findings show that USP13 binds to IκBα."

sparser
"USP13 binds to IκBα, therefore, the effect of downregulating USP13 in BV2 microglia on IκBα expression level was further investigated."
JNK affects USP13
| 3 3
JNK activates USP13.
| 3 1
JNK activates USP13. 4 / 4
| 3 1

reach
"Inhibition of c-Jun N-terminal kinase (JNK) attenuates USP13 degradation, indicating that a JNK-dependent new protein synthesis is necessary for USP13 degradation."

eidos
"Furthermore , among the inhibitors of matrix metalloproteinases ( MMPs ) , ROCK , p38 MAPK , and JNK , only JNK inhibitor II attenuated USP13 degradation ( Fig. 7D ) ."

eidos
"Inhibition of c-Jun N-terminal kinase ( JNK ) attenuates USP13 degradation , indicating that a JNK-dependent new protein synthesis is necessary for USP13 degradation ."

eidos
"Inhibition of JNK had no effect on LPS reduction of USP13 in Raw264.7 cells , while JNK inhibitor attenuates USP13 degradation in Kupffer cells ."
JNK increases the amount of USP13.
| 2
JNK increases the amount of USP13. 2 / 2
| 2

reach
"JNK inhibition also decreased USP13 mRNA expression, demonstrating that JNK mediated c-Jun activity may regulate USP13 at the transcriptional level."

reach
"USP13 is also downregulated by the autophagy inducer C-JUN/AP-1, and inhibition of apoptosis-related MAPK/JNK signaling effectively downregulates the expression of USP13."
FBXL14 affects USP13
| 1 5
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
| 1 1

sparser
"The antagonistic actions between ubiquitin ligases and DUBs are responsible for protein homeostasis as exemplified that USP13-FBXL14 regulated reversible ubiquitination of c-Myc in glioblastoma stem cells [ xref ], and USP15-SMURF2 cooperatively modulated TGF-β receptor in glioblastoma [ xref ]."

reach
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag tagged USP13 or FBXL14."
FASN affects USP13
2 | 4
2 | 4

sparser
"Mechanistically, USP13 interacts with fatty acid synthase (FASN) and enhances FASN protein stability."

sparser
"USP13 interacts with FASN to enhance its protein stability to promote fatty acid synthesis, cancer stemness and sphere formation. xref Inhibition of FASN resulted in the decrease of lipogenesis, self‐renewal ability and chemotherapy resistance in the cancer cells."

sparser
"We confirmed the interaction between endogenous USP13 and FASN by the co-immunoprecipitation (Co-IP) of H1048 cell lysates ( xref )."

sparser
"To detect the role of USP13-FASN axis in regulating cancer stemness and lipogenesis in SCLC, we stably depleted FASN after WT USP13 reconstitution in USP13 knockdown cells."

No evidence text available

No evidence text available
ATG5 affects USP13
4 | 2
4 | 2

No evidence text available

No evidence text available

sparser
"USP13 preferentially is induced in GIST cells by IM and interacts with ATG5, which leads to stabilization of ATG5 through deubiquitination."

sparser
"Activation of PAK1 promoted phosphorylation of ATG5 thereby enhancing the interaction of ATG5 with USP13."

No evidence text available

No evidence text available
ALDOA affects USP13
| 6
| 6

sparser
"In conclusion, these results show that USP13 binds to ALDOA directly via its USP domain."

sparser
"The interaction of USP13 with the ALDOA prompts the question of whether USP13 exerts a regulatory effect on the ALDOA."

sparser
"USP13 directly interacts with ALDOA."

sparser
"The interaction between ALDOA and USP13 was also validated using the heart tissue ( xref E)."

sparser
"Next, we replace the lysine at the three sites with arginine and observed that ALDOA K13R , ALDOA K87R and ALDOA K139R were still bound with USP13 ( xref J)."

sparser
"Subsequently, to further explore the interaction between USP13 and ALDOA, we transfected plasmids encoding Flag-USP13 and Ha-ALDOA into 293T cells and found the interaction between USP13 and ALDOA with co-immunoprecipitation approach ( xref F and G)."
P97 affects USP13
| 5
P97 binds USP13. 5 / 5
| 5

sparser
"The data also suggest that the interaction of USP13 with p97 is mediated by several cellular factors with gp78 being a major one."

sparser
"Compared to the control (cells transfected with an empty vector), expression of several UBX domain-containing p97 cofactors including ASPL, SAK1, and UbxD1 reduced the p97-USP13 interaction ( xref , lanes 3, 5, 7 vs lane 1), suggesting that these factors may compete with USP13 for p97 binding."

sparser
"By contrast, several other factors enhanced the p97-USP13 interaction."

sparser
"Secondly, depletion of gp78 by shRNA-mediated gene knockdown or gene deletion reduced the USP13-p97 interaction by threefold to fourfold ( xref , xref )."

sparser
"We used the GST pull-down assay to screen a collection of known p97 interactors in order to identify factor(s) that promotes the USP13-p97 interaction."
YY1 affects USP13
| 4 1
YY1 increases the amount of USP13.
| 3
YY1 increases the amount of USP13. 3 / 3
| 3

reach
"Together, our findings revealed that YY1 transcriptionally upregulated USP13 expression, indirectly increased WWP1 protein expression and promoted HCC progression."

reach
"Next, a dual luciferase reporter assay demonstrated that YY1 bound to the AAAAATAGCGTC sequence in motif 3 and activated USP13 transcription."

reach
"Our results revealed that YY1 depletion in HCC cells led to a decrease in both the mRNA and protein levels of USP13 (Fig. 7B, C) and that YY1 overexpression upregulated USP13 expression (Supplementary Fig. S7C, D)."
YY1 activates USP13.
| 1 1
YY1 activates USP13. 2 / 2
| 1 1

reach
"YY1 transcriptionally activated USP13 and formed a carcinogenic YY1/USP13/WWP1 axis in HCC."

sparser
"YY1 transcriptionally activated USP13 and formed a carcinogenic YY1/USP13/WWP1 axis in HCC."
USP13 affects p97
| 5
P97 binds USP13. 5 / 5
| 5

sparser
"The data also suggest that the interaction of USP13 with p97 is mediated by several cellular factors with gp78 being a major one."

sparser
"Compared to the control (cells transfected with an empty vector), expression of several UBX domain-containing p97 cofactors including ASPL, SAK1, and UbxD1 reduced the p97-USP13 interaction ( xref , lanes 3, 5, 7 vs lane 1), suggesting that these factors may compete with USP13 for p97 binding."

sparser
"By contrast, several other factors enhanced the p97-USP13 interaction."

sparser
"Secondly, depletion of gp78 by shRNA-mediated gene knockdown or gene deletion reduced the USP13-p97 interaction by threefold to fourfold ( xref , xref )."

sparser
"We used the GST pull-down assay to screen a collection of known p97 interactors in order to identify factor(s) that promotes the USP13-p97 interaction."

reach
"This finding further supports the conclusion that USP13 promotes cellular senescence."

reach
"In this study, we found that USP13 promotes cellular senescence."

reach
"Overexpression of USP13 leads to cellular senescence."

reach
"This result suggests that over-expression of MDM2 may rescue cellular senescence caused by over-expression of USP13."

reach
"Additionally, over-expression of USP13 in human vascular smooth muscle cells promotes cellular senescence."
USP13 affects cell
| 5
USP13 activates cell. 5 / 5
| 5

eidos
"Significantly , USP13 knockdown inhibited hypoxia-induced activation of the TLR4 / MyD88 / NF-kappaB pathway in HCC cells ."

eidos
"Interestingly , the USP13 knockdown repressed the hypoxia-induced activation of the TLR4 / MyD88 / NF-kappaB pathway in HCC cells ( Figure 7D ) ."

eidos
"USP13 knockdown inhibits the proliferation of HCC cells ."

eidos
"Conversely , suppression of USP13 led to GC cell cycle arrest in G1 phase and the inhibition of cell proliferation ."

eidos
"Notably , USP13 knockdown repressed hypoxia-induced activation of the TLR4 / MyD88 / NF-kappaB pathway in HCC cells ."
USP13 affects USP
| 5
USP binds USP13. 5 / 5
| 5

sparser
"178–301) and USP13-USP domains (aa."

sparser
"The HMGB1 B-box mainly binds to USP13 WT and the USP13-USP domain binds to HMGB1."

sparser
"IP analysis showed that the USP13-USP domain was weakly bound to HMGB1 WT (Fig.  xref B)."

sparser
"In addition, the endogenous HMGB1 binding to the USP13-USP domain was observed by HMGB1 IP (Fig.  xref C)."

sparser
"The USP13-USP domain, which binds to HMGB1 (Fig.  xref B and C), mainly plays a role in HMGB1 deubiquitination (Fig.  xref B)."
USP13 affects RAD21
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
USP13 affects PARP1
| 2 3
| 2 3

sparser
"Our study reveals a new regulatory mechanism by which the USP13PARP1 complex can be activated to promote MM progression by enhancing DDR, contributing to a comprehensive understanding of the regulatory spectrum governed by USP13."

sparser
"Then, Co‐IP and reverse Co‐IP experiments further determined the physiological interaction between USP13 and PARP1 (Figure  xref )."

reach
"Our study reveals a new regulatory mechanism by which the USP13PARP1 complex can be activated to promote MM progression by enhancing DDR, contributing to a comprehensive understanding of the regulatory spectrum governed by USP13.There has been a growing interest in bioactive compounds derived from Traditional Chinese Medicine Materials (TCMMs), and their potential to combat MM has been extensively studied [39]."

reach
"To clarify the interaction between USP13 and PARP1, we conducted a WB examination."

sparser
"To clarify the interaction between USP13 and PARP1, we conducted a WB examination."
USP13 affects Interferon
| 1 4
| 1 4

reach
"Thus, USP13 positively regulates type I and type II IFN signaling by deubiquitinating and stabilizing STAT1 protein."

eidos
"It has also been reported that USP13 promotes interferon signaling by stabilizing STAT1 [ 25 ] ."

reach
"USP13 has been reported to deubiquitinate and stabilize STAT1 and promote interferon (IFN)-induced signalling XREF_BIBR."

reach
"USP13 is considered to promote IFN signaling and play an antiviral role by stabilizing STAT1 [55]."

reach
"However, USP13, USP19, and USP22 inhibit virus-induced IFN production by removing K27-linked polyubiquitin chains from STING (40, 81) or TRIF (90)."
USP13 affects FN1
| 5
USP13 increases the amount of FN1. 5 / 5
| 5

reach
"It has been shown that USP13 knockout reduced fibronectin expression in lungs."

reach
"Our previous study showed that USP13 reduction decreased fibronectin expression in lung fibroblast cells [18]."

reach
"We have shown that USP13 knockout reduced fibronectin expression in lungs."

reach
"Overexpression of USP13 increased FN and Smad4 protein levels in lung fibroblasts, while downregulation of USP13 reduced Smad4 protein levels, without altering Smad4 mRNA expression, suggesting that USP13 regulates Smad4 protein stability."

reach
"Though TXNIP has an anti-TGF-β1 signaling effect in lung cells, USP13 has been shown to increase FN levels by stabilizing Smad4 [35], suggesting that the effect of USP13 on TGF-β1 signaling is complex."
USP13 affects ERAD
| 1 4
USP13 activates ERAD.
| 1 2
USP13 activates ERAD. 3 / 3
| 1 2

reach
"Despite having opposing activities, USP13 can cooperate with gp78 to promote ERAD."

eidos
"On the other hand , USP13 antagonizes AMFR by deubiquitinating UBL4A to maintain the integrity of BAG6 complex and therefore promotes ERAD ( Liu et al ., 2014 ) ."

reach
"By contrast, USP13 depletion did not affect deglycosylation of TCRalpha (lane 2 vs 1), suggesting that USP13 promotes ERAD downstream of p97."
USP13 inhibits ERAD.
| 2
USP13 inhibits ERAD. 2 / 2
| 2

reach
"The fact that similar ERAD phenotypes were detected under both USP13 overexpression and depletion conditions further confirmed that USP13 overexpression inhibits ERAD by a dominant negative manner."

reach
"The requirement of gp78 interaction for ERAD inhibition by ectopic USP13 indicates that the interaction of USP13 with gp78 is functionally relevant to ERAD."
USP13 affects DDR
| 5
USP13 activates DDR. 4 / 4
| 4

reach
"In response to DNA damage, USP13 is phosphorylated by ATM, which in turn facilitates USP13 recruitment to DSBs, RAP80 deubiquitination, and triggers DDR signalling."

reach
"Therefore, we hypothesized that USP13/PARP1 may enhance MM cell proliferation by mediating DDR."

reach
"These findings demonstrate that BA can retard the development of MM by targeting USP13/PARP1‐mediated DDR and triggering cell apoptosis, providing a novel mechanism of action for BA in MM (Figure 7H).4 Discussion."

reach
"These findings confirm that the USP13/PARP1 axis positively regulates the DDR of MM cells, promoting MM cell proliferation.3.6 BA Inhibits MM Cell Proliferation in Vitro by Targeting the USP13/PARP1 Axis."
USP13 bound to PARP1 activates DDR. 1 / 1
| 1

reach
"Our study reveals a new regulatory mechanism by which the USP13‐PARP1 complex can be activated to promote MM progression by enhancing DDR, contributing to a comprehensive understanding of the regulatory spectrum governed by USP13.There has been a growing interest in bioactive compounds derived from Traditional Chinese Medicine Materials (TCMMs), and their potential to combat MM has been extensively studied [39]."
USP13 affects Aging
| 5
USP13 activates Aging. 5 / 5
| 5

reach
"Furthermore, overexpression of USP13 promoted cell senescence."

reach
"Increase in USP13 promoted cell senescence."

reach
"The results indicate that overexpression of USP13 promotes cell senescence."

reach
"In order to study if the overexpression of MDM2 can rescue the cell senescence caused by USP13, HVSMC cells were co-transfected with Flag-tagged USP13 and HA-tagged MDM2 plasmids."

reach
"The results showed that over-expression of USP13 induced cell senescence."
USP affects USP13
| 5
USP binds USP13. 5 / 5
| 5

sparser
"178–301) and USP13-USP domains (aa."

sparser
"The HMGB1 B-box mainly binds to USP13 WT and the USP13-USP domain binds to HMGB1."

sparser
"IP analysis showed that the USP13-USP domain was weakly bound to HMGB1 WT (Fig.  xref B)."

sparser
"In addition, the endogenous HMGB1 binding to the USP13-USP domain was observed by HMGB1 IP (Fig.  xref C)."

sparser
"The USP13-USP domain, which binds to HMGB1 (Fig.  xref B and C), mainly plays a role in HMGB1 deubiquitination (Fig.  xref B)."
UBL4A affects USP13
4 | 1
4 | 1

No evidence text available

No evidence text available

No evidence text available

sparser
"Additionally, as catalytic-inactive DUBs often trap their substrates, can a catalytic-inactive USP13 bind preferentially to Ubl4A?"

No evidence text available
TLR4 affects USP13
1 | 1 3
1 | 1 3

sparser
"USP13 interacts with TLR4 and inhibits the ubiquitin-mediated degradation of TLR4 ( xref )."

reach
"USP13 interacted with TLR4 and inhibited the ubiquitin mediated degradation of TLR4."

No evidence text available

sparser
"USP13 interacted with TLR4 and inhibited the ubiquitin-mediated degradation of TLR4."

sparser
"The co-IP assay revealed that USP13 directly interacted with TLR4 in Hep3B cells ( xref )."
SQSTM1 affects USP13
4 | 1
4 | 1

No evidence text available

reach
"In this study, we found that ubiquitin-specific protease 13 (USP13), a member of the USP family, directly binds p62 and removes ubiquitin at Lys7 (K7) of the PB1 domain."

No evidence text available

No evidence text available

No evidence text available
SP1 affects USP13
| 4 1
SP1 inhibits USP13. 5 / 5
| 4 1

sparser
"We also showed that USP13 directly removes classical K48-linked ubiquitin chains from the Raf1 protein ( xref G ), while inhibition of USP13 by Sp-1 promotes mESC self-renewal in an autophagy-independent manner ( xref )."

reach
"Sp-1 simultaneously inhibits the activity of USP10 and USP13 (19)."

reach
"Because USP13 is a deubiquitinated protein, we hypothesized that Sp-1 will accelerate the degradation of target proteins by inhibiting USP13."

reach
"We also showed that USP13 directly removes classical K48-linked ubiquitin chains from the Raf1 protein (Fig. 6G), while inhibition of USP13 by Sp-1 promotes mESC self-renewal in an autophagy-independent manner (Fig. S4)."

reach
"Moreover, Sp-1 functions by inhibiting the activity of USP10 and USP13, while PD0325901 is a specific inhibitor of MEK (5, 19)."
SKP2 affects USP13
1 | 1 3
1 | 1 3

sparser
"UFD1 was reported to function as a scaffold for the USP13-Skp2 (the F-box adaptor of the E3 ubiquitin ligase SCF Skp2 ) interaction, maintaining Skp2 stability."

reach
"We found that SKP2 interacted with USP10, USP13, UCHL5, USP14, and USP7 in KBM5-T315I cells."

sparser
"In HeLa cells, the ubiquitin-recognition protein Ufd1 was reported as a scaffold protein for SKP2USP13 interaction to facilitate SKP2 deubiquitylation by USP13."

sparser
"Prolonged endoplasmic reticulum stress downregulated Ufd1 to attenuate the USP13SKP2 interaction, leading to SKP2 destabilization and p27 accumulation."

No evidence text available
RAD21 affects USP13
5 |
5 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PARP1 affects USP13
| 2 3
| 2 3

sparser
"Our study reveals a new regulatory mechanism by which the USP13PARP1 complex can be activated to promote MM progression by enhancing DDR, contributing to a comprehensive understanding of the regulatory spectrum governed by USP13."

sparser
"Then, Co‐IP and reverse Co‐IP experiments further determined the physiological interaction between USP13 and PARP1 (Figure  xref )."

reach
"Our study reveals a new regulatory mechanism by which the USP13PARP1 complex can be activated to promote MM progression by enhancing DDR, contributing to a comprehensive understanding of the regulatory spectrum governed by USP13.There has been a growing interest in bioactive compounds derived from Traditional Chinese Medicine Materials (TCMMs), and their potential to combat MM has been extensively studied [39]."

reach
"To clarify the interaction between USP13 and PARP1, we conducted a WB examination."

sparser
"To clarify the interaction between USP13 and PARP1, we conducted a WB examination."
OGDH affects USP13
2 | 3
2 | 1

No evidence text available

sparser
"The N terminus including a ubiquitin-specific protease-type zinc-finger domain (residues 1–336) was essential for USP13 to interact with OGDH, whereas the ubiquitin carboxyl-terminal hydrolase domain (residues 336–863) of USP13 is required for its binding with ACLY ( xref )."

No evidence text available
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."
Mice affects USP13
| 5
Mice activates USP13.
| 3
Mice activates USP13. 3 / 3
| 3

reach
"In contrast, the fasting blood glucose levels in USP13 overexpression mice were significantly reduced after being fed HFD for 16 weeks when compared with the control group (Fig. 3B)."

reach
"The Rosa mice subjected to AAV8-TBG-Cre injection were used to specifically overexpress USP13 in hepatocytes, followed by a 24-weeks administration with HFHC diet (Fig. 4A) (hereunder named as Hep OE HFHC)."

reach
"Accordingly, the Rosa mice injected with AAV-TBG-Cre vectors were used to overexpress USP13 abundance only in hepatocytes, followed by a HFMCD fodder feeding for 8 weeks (Fig. 7A) (hereafter referred to as Hep OE HFMCD)."
Mice increases the amount of USP13.
| 2
Mice increases the amount of USP13. 2 / 2
| 2

reach
"To obtain the conditional Usp13 gain-of-function mice, the HFHC or WTDF diet-fed Rosa mice were injected with a 1.5 × 10 genome copies (gc) dose of an AAV8-TBG-Cre vectors via the tail vein to induce hepatocytes-specific Usp13 overexpression (Hep OE HFHC, Hep OE WTDF)."

reach
"The Rosa mice were injected with 1.5 × 10 genome copies (gc) of AAV8-TBG-Cre vectors via the tail vein to induce hepatocytes-specific Usp13 overexpression (Hep OE HFMCD)."
CTNNB1 affects USP13
4 | 1
4 | 1

No evidence text available

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reach
"We demonstrated that USP13 interacts with and deubiquitinates β-catenin, which promotes NSCLC metastasis."
CDH1 affects USP13
| 2 3
| 2 3

sparser
"We performed immunoprecipitation assays followed by western-blotting to assess the interaction between USP13 and CDH1."

sparser
"We found that USP13 binds to the APC/C adaptor CDH1."

reach
"We performed immunoprecipitation assays followed by western-blotting to assess the interaction between USP13 and CDH1."

reach
"We found that USP13 binds to the APC/C adaptor CDH1."

sparser
"Shortly after this finding, the interaction of USP13 with CDH1 was also confirmed by the same team ( xref )."
Ubiquitin-proteasome affects USP13
| 4
Ubiquitin-proteasome inhibits USP13. 4 / 4
| 4

reach
"Bortezomib treatment increased accumulation of USP13 (Fig. 4E), indicating that USP13 degradation is mediated by the ubiquitin-proteasome system."

reach
"Here, we report that USP13 degradation is mediated by the ubiquitin-proteasome system."

reach
"However, despite the requirement of IL-1R8 of IL-37 to exert its function, upon IL-37 treatment IL-1R8 is degraded by the ubiquitin-proteasome system, a pathway that is negatively regulated by the deu[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Inhibition of proteasome, but not lysosome or immunoproteasome, attenuates LPS-induced USP13 degradation, suggesting USP13 degradation is mediated by the ubiquitin-proteasome system."
MiR-130b/301b affects USP13
| 4
Modified miR-130b/301b decreases the amount of USP13. 4 / 4
| 4

reach
"Overexpression of miR-130b/301b or USP13 knockdown by shRNA delivery decreases USP13 expression, and further reduces PTEN protein expression and leads to enhancement of cell proliferation, invasion and migration."

reach
"We reported a potential regulatory loop that the NF-kB-induced miR-130b/301b overexpression decreased USP13 expression and subsequently resulted in the downregulation of PTEN protein and promoted tumorigenesis of bladder cancer."

reach
"Results showed that overexpression of miR-130b/301b dramatically decreased protein expression of USP13 and PTEN, while restoration of USP13 completely reversed the effect."

reach
"Results showed that overexpression of miR-130b/301b significantly decreased USP13 expression in transcription and translational levels, and knockdown of miR-130b/301b revealed an opposite effect on USP13 expression."
WAPL affects USP13
4 |
4 |

No evidence text available

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No evidence text available
VCP affects USP13
2 | 2
2 | 2

reach
"In addition, USP13 interacts with the P97 and valosin containing protein complex, a key chaperone in ER associated degradation, which increases the protein level of CD3delta, an ER associated degradation substrate."

No evidence text available

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."

No evidence text available
Ufd1 affects USP13
| 1 3
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
| 1 1

sparser
"USP13 also interacts with the ubiquitin-recognition protein Ufd1 and facilitates endoplasmic reticulum (ER)-associated degradation (ERAD) of misfolded proteins [ xref ]."

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."
| 2

reach
"Consistently, it has been reported that deletion of USP13 can activate the NF-κB signaling pathway in response to herpesvirus infection, increasing resistance to the virus (Sun et al., 2017)."

reach
"Furthermore, knockdown of USP13 upregulated expression level of nuclear accumulation of p65 in BV2 microglia, implying that it activated the NF-κB signaling pathway (Additional file 1: Fig. S6a, b)."
| 2

reach
"Importantly, USP13 modulated the AKT‐mTOR signalling pathway in a manner analogous to PRPF6 (Figures 6D,E)."

reach
"Furthermore, knockdown of USP13 or WISP1 impaired the activation of the Wnt/CTNNB1 signaling pathway and reduced immune checkpoint marker expression, indicating a mechanism by which USP13 promotes immune evasion in ESCC."
USP13 affects pyroptosis
| 4
| 4

reach
"However, the overexpression of USP13 counteracted spautin-1 improvement of cerebral I/R injury and significantly abolished spautin-1 suppression of autophagy and NLRP3 inflammasome-dependent pyroptosis."

reach
"Cerebral I/R significantly reduced the expressions of NLRP3, USP13, beclin 1, and GSDMD-N, which was abolished by spautin-1, indicating that spautin-1 inhibited autophagy and NLRP3 inflammasome-mediated pyroptosis."

reach
"A recent study showed that inhibition of USP13 reduced autophagy and ROS accumulation, and attenuated pyroptosis."

reach
"However, USP13 overexpression abolished these beneficial effects of spautin-1 and notably counteracted the spautin-1 inhibition of autophagy and NLRP3 inflammasome-mediated pyroptosis."
USP13 affects nsp13
| 4
USP13 ubiquitinates nsp13.
| 2
USP13 leads to the ubiquitination of nsp13. 2 / 2
| 2

reach
"Loss of USP13 led to the increase of ubiquitinated nsp13."

reach
"Predicting hijacking of host de-ubiquitination by the virus, Guo et al. checked for interactions between host de-ubiquitinase and nsp13 and found USP13 to interact with it and observed that loss of USP13 led to greater ubiquitinated nsp13."
USP13 deubiquitinates nsp13.
| 2
USP13 deubiquitinates nsp13. 2 / 2
| 2

reach
"We found that nsp13 interacts with the deubiquitinase USP13, which deubiquitinates and stabilizes nsp13."

reach
"Loss of USP13 enhances ubiquitination of nsp13 and destabilizes nsp13 protein."
USP13 affects nilotinib
| 4
| 4

reach
"Moreover, overexpression of USP13 inhibits the effect of the tyrosine kinase inhibitor, Nilotinib, and USP13 knockdown enhances the effect of Nilotinib on α-synuclein clearance [ 49 ]."

reach
"Conclusively, the above findings confirmed that USP13 interacted with and stabilized Bcl‐XL under both physiological and pathological conditions, thus involving in nilotinib‐induced nephrotoxicity.2.5 CQ Relieves Nilotinib-Induced Nephrotoxicity In Vitro."

reach
"Overexpression of USP13 inhibits the effects of small molecules, including the tyrosine kinase inhibitor, Nilotinib; conversely, USP13 knockdown enhances the effects of Nilotinib on α-synuclein cleara[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Additionally, it was shown that overexpression of USP13 could antagonize the positive effects of the drug Nilotinib, which is known to increase α-synuclein clearance in this animal model, whereas USP13 KD augmented the effects of the drug [64]."
USP13 affects glioblastoma multiforme cancer stem cell
| 4
USP13 activates glioblastoma multiforme cancer stem cell. 4 / 4
| 4

reach
"In vivo bioluminescent analysis confirmed that induced disruption of USP13 by doxycycline treatment significantly inhibited GSC tumor growth in mouse intracranial xenografts (XREF_FIG)."

reach
"In addition, USP13 disruption significantly reduced GSC proliferation (XREF_FIG) but showed little effects on NSTCs and NPCs (not depicted)."

reach
"Disrupting USP13 abrogates GSC tumor growth."

reach
"In addition, IHC staining of Ki-67 and cleaved caspase 3 confirmed that USP13 disruption reduced c-Myc protein in the GSC derived xenografts and led to a significant decrease in cell proliferation and an increase in cell apoptosis within the tumor (not depicted)."

reach
"Snail can be maintained by USP13 to trigger EMT and metastasis, resulting in the poor prognosis of GC patients indirectly [58]."

reach
"Deubiquitinase USP13 promotes the epithelial-mesenchymal transition and metastasis in gastric cancer by maintaining Snail protein."

reach
"Downregulation of USP13 resulted in reduced WISP1 protein stability, decreased cell proliferation, migration, and EMT, and increased apoptosis in vitro."

reach
"Furthermore, USP13 positively regulates the epithelial-mesenchymal transition (EMT) of GC cells."
| 4

reach
"Finally, Snail knockdown significantly blocked USP13-induced SGC-7901 cell migration and invasion."

reach
"USP13 knockdown prominently suppressed MGC-803 cell migration and invasion."

reach
"USP13 promotes breast cancer cell migration, invasion and lung metastasis through Twist1."

reach
"According to the results of the Transwell assay (Supplementary Fig. S2F) and wound healing assay (Supplementary Fig. S2G), USP13 promoted HCC cell migration."
USP13 affects WAPL
4 |
4 |

No evidence text available

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USP13 affects VCP
2 | 2
2 | 2

reach
"In addition, USP13 interacts with the P97 and valosin containing protein complex, a key chaperone in ER associated degradation, which increases the protein level of CD3delta, an ER associated degradation substrate."

No evidence text available

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."

No evidence text available
USP13 affects Ufd1
| 1 3
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
| 1 1

sparser
"USP13 also interacts with the ubiquitin-recognition protein Ufd1 and facilitates endoplasmic reticulum (ER)-associated degradation (ERAD) of misfolded proteins [ xref ]."

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."
USP13 affects UBL
| 4
USP13 binds UBL. 2 / 2
| 2

sparser
"On the other hand, USP13 can bind a UBL domain in Bag6 ( xref ), which explains why the catalytically inactive USP13 binds the Bag6-Ubl4A complex with similar affinity as the wild type protein."

sparser
"Furthermore, USP13 may be associated with other ubiquitin-like (UbL) proteins such as ISG15, SUMO1 and SUMO2 [ xref , xref ]."
UBL binds USP13 and BAG6. 2 / 2
| 2

sparser
"On the other hand, the binding of USP13 with the UBL domain of Bag6 can be demonstrated using pulldown experiments ( xref )."

sparser
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."
USP13 affects TRIM63
2 1 | 1
2 1 | 1

reach
"MURF-1 also binds to ubiquitin conjugating enzyme-9 and isopeptidase T-3, enzymes involved in small ubiquitin related modifier mediated nuclear import, and with glucocorticoid modulatory element binding protein-1 (GMEB-1), a transcriptional regulator."

No evidence text available

No evidence text available

No evidence text available
USP13 affects PDS5A
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
USP13 affects NPLOC4
1 | 1 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
1 | 1

No evidence text available

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."
USP13 affects NFE2L2
| 4
USP13 inhibits NFE2L2.
| 2
USP13 inhibits NFE2L2. 2 / 2
| 2

reach
"Further studies showed that USP13 reduced LPS-induced oxidative stress and inflammation by inducing Nrf2."

reach
"USP13 reduces septic mediated cardiomyocyte oxidative stress and inflammation by inducing Nrf2."
USP13 increases the amount of NFE2L2.
| 2
USP13 increases the amount of NFE2L2. 2 / 2
| 2

reach
"In conclusion, this study demonstrated that USP13 was downregulated in septic myocardial injury tissues, and USP13 overexpression increased Nrf2 levels and reduced apoptosis."

reach
"USP13 deubiquitinates p62/SQSTM1 to induce autophagy and Nrf2 release for activating antioxidant response genes."
USP13 affects NASH
| 4
USP13 activates NASH. 4 / 4
| 4

reach
"These findings further reveal that adipocyte proliferation might links to the increased weight of visceral fat in NASH diet-challenged mice.3.4 Hepatocyte-specific Usp13 overexpression mitigates HFHC/WTDF-triggered NASH."

reach
"Given the close-knit correlation of IRHOM2 deactivation with ubiquitin-associated domains activity of USP13, to thoroughly study whether USP13–IRHOM2 interaction is necessary for the inhibitory function of USP13 against NASH progression, AAV8-TBG-loaded WT full-length Usp13 sequences vectors (AAV-TBG-Usp13) and AAV8-TBG-loaded Usp13 with UBA1/2 domain deletion vectors (AAV-TBG-Usp13 (ΔUBA)) was generated and used again to further explore the influence of USP13 with UBA1/2 mutants on HFHC-induced NASH pathologies in vivo (Fig. S18N)."

reach
"Also, we further indicated that UBC13-deletion dramatically decreased IRHOM2 ubiquitination, and USP13 could not exhibit suppressive effect on IRHOM2 ubiquitination in these in vitro experiments (Fig. 6M).3.9 Hepatocyte-specific Usp13 overexpression alleviates HFMCD-induced NASH."

reach
"The obtained results indicated that UBA domain of USP13 is essential for its suppressive effects in regulation of NASH development via IRHOM2 pathway.3.7 USP13-IRHOM2 interaction is necessary for USP13-triggered mitigation of NASH."
USP13 affects HK2
| 4
USP13 increases the amount of HK2.
| 3
USP13 increases the amount of HK2. 3 / 3
| 3

reach
"As shown in Figure 2E, the Western blotting analysis also illustrated that USP13 overexpression led to an obvious reduction in the protein levels of GLUT1 and HK2."

reach
"Furthermore, USP13 overexpression induced phosphatase and tensin homolog deleted on chromosome 10 (PTEN) expression and repressed the activation of AKT as well as the expression of the downstream effectors glucose transporter-1 (GLUT1) and hexokinase-2 (HK2)."

reach
"Furthermore, USP13 overexpression induced phosphatase and tensin homolog deleted on chromosome 10 (PTEN) expression and repressed the activation of AKT as well as the expression of the downstream effectors glucose transporter-1 (GLUT1) and hexokinase-2 (HK2)."
USP13 decreases the amount of HK2.
| 1
USP13 decreases the amount of HK2. 1 / 1
| 1

reach
"27 In our experiments, USP13 overexpression reduced the levels of GLUT1 and HK2 in OSCC cells, while knockdown of USP13 had the opposite effects."
USP13 affects GEMIN4
| 4
| 4

reach
"USP13 interacts with p97, UFD1, NPL4, and UBXD8, and upon USP13 deletion, cells are more susceptible to ER stress."

reach
"By contrast, neither GST-USP13 nor GST-USP5 bound p97 (XREF_FIG)."

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."

reach
"In addition, USP13 interacts with the P97 and valosin containing protein complex, a key chaperone in ER associated degradation, which increases the protein level of CD3delta, an ER associated degradation substrate."
USP13 affects DNA Damage
| 4
USP13 inhibits DNA Damage.
| 2
| 2

reach
"Similarly, USP10 and USP13 silencing could induce ROS generation and cause DNA damage."
| PMC

reach
"However, overexpression of USP13 reduced the degree of DNA damage, effectively reversing the DNA damage caused by MMS treatment to the same level as the control group."
USP13 activates DNA Damage.
| 2
| 2

reach
"Additionally, USP13 inhibition potentially suppresses DNA damage repair and protein neddylation of PCa cells, thereby facilitating the cellular response to stimuli and the modification of TME."

reach
"However, overexpression of USP13 reduced the degree of DNA damage, effectively reversing the DNA damage caused by MMS treatment to the same level as the control group."
| 2 2
| 2 2

eidos
"( A ) USP13 knockdown decreases cell viability with or without MG132 treatment ."

reach
"MTS assay indicated that USP13 knockdown inhibited cell viability (Fig. S1, A and B)."

reach
"However, in the context of hypoxic or oxidative stresses, USP13 loss caused a substantial decrease of cell viability and a significant increase of cell apoptosis in both TOV-21G and SW-1573 tumor models."

eidos
"The knockdown of USP13 also led to a substantial cell viability reduction upon osimertinib treatment ( Figure 2E ) ."
USP13 affects CK2
| 2 2
USP13 binds CK2.
| 1 2
| 1 2

sparser
"To determine whether USP13 interacts with CK2 in ovarian cancer cells, we examined CK2USP13 binding in HeyA8 cells expressing FLAG-tagged USP13."

sparser
"This experiment showed that CK2 was co-immunoprecipitated with USP13, indicating that USP13 interacts with CK2 in ovarian cancer cells ( xref C)."

reach
"Interaction between USP13 and CK2 was verified with western blotting."
USP13 inhibits CK2.
| 1
USP13 inhibits CK2. 1 / 1
| 1

reach
"We then addressed whether CK2 phosphorylates USP13 in the cellular context by inhibiting CK2 kinase activity in cells with CK2 inhibitors, silmitasertib (CX-4945) and 4,5,6,7-tetrabromobenzotriazole (TBB) [33,34,35]."
USP13 affects ATG7
| 4
| 4

sparser
"USP13 Interacts With ATG7 in the Granulosa Cells of Chicken."

sparser
"These findings show that USP13 and ATG7 interact in chicken granulosa cells."

sparser
"Given the involvement of diverse physiological processes in ferroptosis, gaining a comprehensive understanding of the role of the USP13-ATG7 axis in chicken follicle growth and development can provide valuable insights into molecular-assisted breeding strategies and serve as a foundation for enhancing poultry yield."

sparser
"In conclusion, this study elucidates the crucial role of the USP13-ATG7 axis in regulating ferroptosis in chicken follicle granulosa cells, thereby presenting a novel avenue for molecular breeding research in chickens."
USP13 affects AAV9
| 4
USP13 binds AAV9. 4 / 4
| 4

sparser
"To achieve cardiomyocyte-specific overexpression of USP13 in vivo, we employed an AAV9 containing a cardiac-specific cTnT promoter component under which the vector of USP13 was placed (AAV9-USP13 oe ) while the AAV9-cTnT-vector used as a control (AAV9-Ctrl)."

sparser
"The results revealed a pronounced increase in GPX4 and SLC7A11, along with a significant decrease in PTGS2, TNF and IL-6, in the AAV9-USP13 oe mice compared to the AAV9-Ctrl mice, while there was no substantial discrepancy in the expression of cleaved caspase-3. ( xref B through 7D and xref through S14D)."

sparser
"Furthermore, Masson's trichrome staining revealed that AAV9-USP13 oe mice exhibited significantly reduced MI scar sizes compared with those of AAV9-Ctrl mice ( xref K and L)."

sparser
"Ultrasound-guided, in situ multi-point injections of AAV9-USP13 oe were administered to mouse hearts."
UBL affects USP13
| 4
USP13 binds UBL. 2 / 2
| 2

sparser
"On the other hand, USP13 can bind a UBL domain in Bag6 ( xref ), which explains why the catalytically inactive USP13 binds the Bag6-Ubl4A complex with similar affinity as the wild type protein."

sparser
"Furthermore, USP13 may be associated with other ubiquitin-like (UbL) proteins such as ISG15, SUMO1 and SUMO2 [ xref , xref ]."
UBL binds USP13 and BAG6. 2 / 2
| 2

sparser
"On the other hand, the binding of USP13 with the UBL domain of Bag6 can be demonstrated using pulldown experiments ( xref )."

sparser
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."
TRIM63 affects USP13
2 1 | 1
2 1 | 1

reach
"MURF-1 also binds to ubiquitin conjugating enzyme-9 and isopeptidase T-3, enzymes involved in small ubiquitin related modifier mediated nuclear import, and with glucocorticoid modulatory element binding protein-1 (GMEB-1), a transcriptional regulator."

No evidence text available

No evidence text available

No evidence text available
SMAD4 affects USP13
1 | 1 2
1 | 1 2

sparser
"In addition, we demonstrate that USP13 is associated with Smad4, and the association is not affected by TGF- β 1."

reach
"Interestingly, we also found that while Smad4 is translocated into the nucleus under the stimulation of TGF-β1 (10 ng/mL, 1 hour), USP13-V5 was also observed in the nucleus (Fig 6B, C), suggesting that USP13 is associated with Smad4 and the USP13/Smad4 complex is translocated into nucleus in response to TGF-β1 treatment."

sparser
"Interestingly, we also found that while Smad4 is translocated into the nucleus under the stimulation of TGF- β 1 (10 ng/mL, 1 hour), USP13-V5 was also observed in the nucleus ( xref , C), suggesting that USP13 is associated with Smad4 and the USP13/Smad4 complex is translocated into nucleus in response to TGF- β 1 treatment."

No evidence text available
RAF1 affects USP13
| 1 3
| 1 3

sparser
"USP13 interacts with Raf1."

sparser
"A coimmunoprecipitation assay was performed in Flag-tagged Raf1 (Flag-Raf1)-overexpressing 46C mESCs carrying the HA or HA-USP13 transgene with a Flag antibody-affinity gel, which was used to bind the Flag fusion protein for immunoprecipitation analysis, to determine whether USP13 interacts with Raf1 ( xref A )."

reach
"Mechanistically, we demonstrate that USP13 physically interacts with, deubiquitinates, and stabilizes serine/threonine kinase Raf1 and thereby sustains Raf1 protein at the posttranslational level to activate the FGF/MEK/ERK prodifferentiation signaling pathway in naive mouse ESCs."

sparser
"Co-IP was performed and showed that RAF1 interacted with USP13 in hiPSCs ( xref G )."
PDS5A affects USP13
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
NPLOC4 affects USP13
1 | 1 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
1 | 1

No evidence text available

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."
MYC affects FBXL14
| 4
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
MITF affects USP13
1 | 2 1
MITF binds USP13.
1 | 1 1
1 | 1 1

sparser
"USP13 binds to MITF and cleaves the ubiquitin chain ( xref )."

reach
"USP13 binds to MITF and cleaves the ubiquitin chain (65)."

No evidence text available
MITF activates USP13.
| 1
MITF activates USP13. 1 / 1
| 1

reach
"More importantly, USP13 deficiency resulted in impaired melanoma growth both in vitro and in vivo, and reintroduction of MITF reversed the inhibitory effect of USP13, demonstrating that MITF contributed to melanoma progression under the ubiquitination modification of USP13 95."
KRAS affects USP13
| 4
KRAS increases the amount of USP13.
| 2
KRAS increases the amount of USP13. 2 / 2
| 2

reach
"In agreement with previous observations that blockage of β-catenin/TCF-4 interaction leads to cytoplasmic retention of β-catenin and reduction of its nuclear localization, 70 , 71 we also found that d[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These results demonstrate that KRAS activation induces USP13 expression to promote NSCLC metastasis.To explore the mechanism by which KRAS induces USP13 expression, we analyzed the promoter region of [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
KRAS activates USP13.
| 2
KRAS activates USP13. 2 / 2
| 2

reach
"Besides, U0126-mediated inhibition of KRAS effector MAP kinase kinase (MEK) decreased USP13 and vimentin protein and mRNA levels in A549 and H460 cells, both of which carry mutant KRAS ( Figures 4 G a[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These results demonstrate that KRAS signaling upregulates USP13 expression through transcription factor RREB1.To investigate the clinical relevance of USP13 in NSCLC, we analyzed USP13 expression stat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Hypoxia affects USP13
| 1 3
Hypoxia inhibits USP13.
| 1 1
| 1 1

reach
"However, these changes seem to be cell type-specific, since only melanoma cells but not HEK293 or HeLa cells displayed the hypoxia-mediated USP13 reduction."

eidos
"( D ) Hypoxia increased USP13 , TLR4 , MyD88 , and P-NF-kappaB p65 proteins , which was subsequently reduced by USP13 knockdown ."
Hypoxia increases the amount of USP13.
| 2
Hypoxia increases the amount of USP13. 2 / 2
| 2

reach
"In HCC, hypoxia induces USP13 expression, enhancing TLR4 deubiquitination and activating the TLR4/MyD88/NF‐κB pathway, which accelerates tumour growth [18]."

reach
"By contrast, USP13 expression induced by hypoxia activates TLR4/MYD88/NF-κB signaling and promotes cell proliferation (81)."
GEMIN4 affects USP13
| 4
| 4

reach
"USP13 interacts with p97, UFD1, NPL4, and UBXD8, and upon USP13 deletion, cells are more susceptible to ER stress."

reach
"By contrast, neither GST-USP13 nor GST-USP5 bound p97 (XREF_FIG)."

reach
"USP13 interacts with p97 and VCP, Ufd1, Npl4, and UBXD8 [XREF_BIBR]."

reach
"In addition, USP13 interacts with the P97 and valosin containing protein complex, a key chaperone in ER associated degradation, which increases the protein level of CD3delta, an ER associated degradation substrate."
FBXL14 affects MYC, and USP13
| 4
USP13 binds FBXL14 and MYC. 4 / 4
| 4

sparser
"To further confirm the interaction between endogenous c-Myc and USP13 or FBXL14 in GSCs, c-Myc was immunoprecipitated from cell lysate of partially differentiated GSCs that were induced by serum acutely (2 d)."

sparser
"USP13 and FBXL14 interact with c-Myc in glioma cells."

sparser
"These results demonstrate that c-Myc interacts with USP13 and FBXL14 in glioma cells, indicating that deubiquitinase USP13 and the ubiquitin E3 ligase FBXL14 are potential regulators of c-Myc protein stability in GBM."

sparser
"The interaction between c-Myc and USP13 or FBXL14 was validated in HEK293 cells coexpressing c-Myc and Flag-tagged USP13 or FBXL14."
ATG7 affects USP13
| 4
| 4

sparser
"USP13 Interacts With ATG7 in the Granulosa Cells of Chicken."

sparser
"These findings show that USP13 and ATG7 interact in chicken granulosa cells."

sparser
"Given the involvement of diverse physiological processes in ferroptosis, gaining a comprehensive understanding of the role of the USP13-ATG7 axis in chicken follicle growth and development can provide valuable insights into molecular-assisted breeding strategies and serve as a foundation for enhancing poultry yield."

sparser
"In conclusion, this study elucidates the crucial role of the USP13-ATG7 axis in regulating ferroptosis in chicken follicle granulosa cells, thereby presenting a novel avenue for molecular breeding research in chickens."
AAV9 affects USP13
| 4
USP13 binds AAV9. 4 / 4
| 4

sparser
"To achieve cardiomyocyte-specific overexpression of USP13 in vivo, we employed an AAV9 containing a cardiac-specific cTnT promoter component under which the vector of USP13 was placed (AAV9-USP13 oe ) while the AAV9-cTnT-vector used as a control (AAV9-Ctrl)."

sparser
"The results revealed a pronounced increase in GPX4 and SLC7A11, along with a significant decrease in PTGS2, TNF and IL-6, in the AAV9-USP13 oe mice compared to the AAV9-Ctrl mice, while there was no substantial discrepancy in the expression of cleaved caspase-3. ( xref B through 7D and xref through S14D)."

sparser
"Furthermore, Masson's trichrome staining revealed that AAV9-USP13 oe mice exhibited significantly reduced MI scar sizes compared with those of AAV9-Ctrl mice ( xref K and L)."

sparser
"Ultrasound-guided, in situ multi-point injections of AAV9-USP13 oe were administered to mouse hearts."
Ubiquitin-proteasome system affects USP13
| 3
Ubiquitin-proteasome system inhibits USP13. 3 / 3
| 3

eidos
"Inhibition of proteasome , but not lysosome or immunoproteasome , attenuates LPS-induced USP13 degradation , suggesting USP13 degradation is mediated by the ubiquitin-proteasome system ."

eidos
"Bortezomib treatment increased accumulation of USP13 ( Fig. 4E ) , indicating that USP13 degradation is mediated by the ubiquitin-proteasome system ."

eidos
"Here , we report that USP13 degradation is mediated by the ubiquitin-proteasome system ."
Ubiquitin affects UBA1
| 3
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
USP13 affects α-synuclein
| 3
USP13 ubiquitinates α-synuclein. 3 / 3
| 3

reach
"Here, it was shown that USP13 negatively regulates Parkin ubiquitination, function, and solubility (Figure 5), negatively regulates the level, ubiquitination, and clearance of α-synuclein, and regulates proteasome activity independently of Parkin, which affects dopaminergic neuronal viability and motor performances in animal models."

reach
"The reduced ubiquitination and clearance of α-synuclein by USP13 was verified in vivo in transgenic mice with expression of mutant α-synuclein."

reach
"This perspective suggests that the monoubiquitinated α-synuclein should be extensively managed.Inhibition of USP13 enhances ubiquitination of α-synuclein, and causes the removal of α-synuclein through[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"For example, overexpression of USP13 blocks the AKT signaling pathway and suppresses tumor cell proliferation, invasion, and glycolysis by upregulating PTEN, while USP13 levels are downregulated in breast, bladder, and oral squamous tumors, in correlation with PTEN levels (Fig. 3)."

reach
"It is assumed that overexpression of USP13 can block AKT signaling pathway, suppressing tumor cell proliferation, invasion, and glycolysis through up-regulating PTEN protein levels (Zhang et al., 2013; Xiang et al., 2015; Zhang et al., 2018; Man et al., 2019)."

reach
"For instance, in non-small-cell lung cancer (NSCLC), downregulation of USP13 inhibits MAPK/AKT signaling (Han et al., 2016)."
USP13 affects miR-301b-3p
| 3
USP13 activates miR-301b-3p. 3 / 3
| 3

reach
"Furthermore, we discovered that USP13 was a common target of miR-130b-3p and miR-301b-3p, and the miR-130b/301b cluster, which could be transcriptionally upregulated by NF-kB."

reach
"Man et al. [XREF_BIBR] reported that USP13 is a direct target of miR-301b-3p, and it can be downregulated by miR-301b-3p overexpression, leading to decreased protein expression of PTEN, consequently promoting the occurrence of bladder cancer."

reach
"USP13 is a common target of mR-130b-3p and miR-301b-3p, and is down-regulated in human bladder cancer tumors."
USP13 affects ferroptosis
| 3
| 3

reach
"However, these effects were greatly blocked by USP13 overexpression which causes increases in autophagy as well as ferroptosis [21]."

reach
"Overexpression of USP13 results in depletion of glutathione (GSH), peroxidation of lipids, accumulation of iron, and activation of ferroptosis in chicken granulosa cells."

reach
"In contrast, USP13 knockdown significantly inhibited ferroptosis events."
USP13 affects cell death
| 3
| 3

reach
"Our study provides a potential therapeutic strategy in which targeting USP13 blocks biosynthesis of metabolic intermediates and lipids thereby simultaneously inducing energy stress and cell death."

reach
"Compound Spautin-1 targets the USP13, which promotes cell death and inhibits autophagic cell death under starvation conditions [173]."

reach
"Similar to Beclin 1 knockdown, we found that suppression of the ubiquitin specific peptidase, USP10, or a small molecule inhibitor of the deubiquitinases USP10 and USP13, i.e., spautin-1 XREF_BIBR, can increase radiation induced DSBs and promote tumor cell death."
USP13 affects barium(2+)
| 3
| 3

reach
"These findings suggest that combining BA with a PARP inhibitor can effectively counteract the resistance to BA induced by USP13 overexpression.3.7 BA Hampers MM Progression in Mouse Models."

reach
"In order to overcome the resistance to BA induced by USP13 overexpression, we combined BA with the PARP inhibitor olaparib."

reach
"Additionally, our findings show that combining BA with a PARP inhibitor, olaparib, can effectively counteract the resistance to BA induced by USP13 overexpression."
USP13 affects afatinib
| 1 2
| 1 2

eidos
"Similar results were obtained in HCC827 cells ( Figure 2D ) , confirming that USP13 knockdown strongly enhances afatinib in treatment-naive lung cancer cell lines ."

reach
"Overall, these data indicate that targeting of USP13 strongly potentiates the anticancer effects of afatinib in vivo."

reach
"Similar results were obtained in HCC827 cells, confirming that USP13 knockdown strongly enhances afatinib in treatment-naive lung cancer cell lines."
USP13 affects UFD1
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USP13 affects TRIM55
2 1 |
2 1 |

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USP13 affects SYK
| 1 2
| 1 2

reach
"Protein-protein interactions assays showed that USP13 interacted with the co-immunoprecipitated protein spleen tyrosine kinase (SYK) and deubiquitinated SYK."

sparser
"Protein-protein interactions assays showed that USP13 interacted with the co-immunoprecipitated protein spleen tyrosine kinase (SYK) and deubiquitinated SYK."

sparser
"The USP13-SYK axis may serve as a therapeutic target for treating FcεRI-mediated activation of MCs and associated inflammatory responses."
USP13 affects STAG2
3 |
3 |

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USP13 affects SCCPDH
3 |
3 |

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USP13 affects SARM1
| 2 1
USP13 deubiquitinates SARM1.
| 2
USP13 deubiquitinates SARM1. 2 / 2
| 2

reach
"Further investigation revealed that USP13 deubiquitinated SARM1, which increased the inhibitory interaction between the N-terminal armadillo repeat motif (ARM) and C-terminal Toll/interleukin-1 receptor (TIR) domains of the SARM1 protein, thereby suppressing SARM1 activation in axon injury."

reach
"Deubiquitination of SARM1 by USP13 regulates SARM1 activation and axon degeneration."
USP13 binds SARM1.
| 1
| 1

sparser
"USP13 interacts with SARM1 and regulates axonal integrity via SARM1."
USP13 affects RPP25
3 |
3 |

No evidence text available

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USP13 affects Proteasome
| 3
| 3

reach
"USP13 is upregulated in Alzheimer's disease (AD) and Parkinson's disease (PD), and USP13 knockdown via shRNA reduces neurotoxic proteins and increases proteasome activity in models of neurodegeneration."

reach
"Notably, USP13 not only bound Bag6, but also interacted with MMS1 and alpha6, subunits of the proteasome (lane 5), further implicating USP13 in delivery pathways that targets substrates to the proteasome."

reach
"In the study by Liu et al. [64], it was also shown that USP13 could regulate proteasome activity independently of Parkin, where USP13 expression, even with KD, could increase proteasome activity in α-synuclein-expressing cells irrespective of Parkin expression."
USP13 affects MAPK
| 3
USP13 inhibits MAPK. 3 / 3
| 3

reach
"As expected, USP13 inhibited inflammatory pathways, including MAPK and NF-κB signaling pathways, in the livers of NAFLD mice."

reach
"In light of the above findings that USP13 inhibits the MAPK and NF-κB signaling pathways, we considered TAK1, a known upstream kinase of these pathways, as a potential downstream target of USP13."

reach
"For instance, in non-small-cell lung cancer (NSCLC), downregulation of USP13 inhibits MAPK/AKT signaling (Han et al., 2016)."
USP13 affects HMOX1
| 3
USP13 decreases the amount of HMOX1. 3 / 3
| 3

reach
"Overexpression of USP13 reduced HMOX1 protein levels in lung fibroblast cells, suggesting that modulation of USP13 levels may affect oxidative stress."

reach
"Knockdown of USP13 by siRNA increased HMOX1 levels in lung fibroblast cells (Mrc5) and epithelial cells (A549 and BEAS-2B) (Fig. 2B, D, E), while HMOX1 levels were decreased in V5-tagged USP13-overexpressed Mrc5 cells (Fig. 2C), indicating that USP13 regulates HMOX1 levels in lung cells."

reach
"Knockdown of USP13 increased HMOX1 basal levels but did not rescue the HMOX1 reduction by TGF-β1 (Fig. 2B)."
| 1
| 1

reach
"Our results indicate that spautin-1 as well as knockdown of its downstream targets, USP10 and USP13, reduced the proliferation and migration of glioblastoma cells."
USP13 affects EIF5B
| 3
| 3

sparser
"USP13 interacts with eIF5B and negatively regulates its post-transcriptional expression."

sparser
"First, USP13 binds to eIF5B and enhances its interaction with SQSTM1/P62, promoting autophagy-lysosomal degradation of eIF5B. Second, previous studies show the functions of USP13 in SQSTM1/P62–USP13 stabilizes SQSTM1/P62 through depubiquitination, thus promoting the oligomerization of SQSTM1/P62 [ xref ], augmenting SQSTM1/P62’s substrate recruitment."

sparser
"Co-immunoprecipitation and immunoblot analysis confirmed the interaction between exogenous USP13 and endogenous eIF5B in NRVMs, with similar observations in the HEK−293T cell line (Fig. xref C, E)."
| 3

reach
"As shown in Fig. 5i, j, USP13 overexpression significantly suppressed IκBα degradation while USP13 downregulation accelerated IκBα degradation.USP13 is a deubiquitinase (DUB) and regulates IκBα stability, therefore, analyses were performed to explore if USP13 regulates IκBα ubiquitination."

reach
"As shown in Fig. 5i, j, USP13 overexpression significantly suppressed IκBα degradation while USP13 downregulation accelerated IκBα degradation.USP13 is a deubiquitinase (DUB) and regulates IκBα stability, therefore, analyses were performed to explore if USP13 regulates IκBα ubiquitination."

reach
"Thr122 phosphorylation could affect the ubiquitination status of USP13 by inhibiting the binding of E3 ubiquitin ligase or promoting the binding of deubiquitinase on USP13."
USP13 affects Beclin1 subunit
| 3
USP13 activates Beclin1 subunit. 3 / 3
| 3

reach
"Our second treatment was with Spautin-1 which promotes the degradation of Vps34 PI3 kinase complexes by inhibiting two ubiquitin specific peptidases, USP10 and USP13, which target the Beclin1 subunit of Vps34 complexes."

reach
"We treated TBEV infected neuroblastoma cells with rapamycin (inducer of autophagy, as inhibition of mTOR mimics cellular starvation by blocking signals required for cell growth and proliferation), and spautin-1 (highly specific and potent autophagy inhibitor in mammalian cells, which promotes the degradation of Vps34 PI3 kinase complexes by inhibiting two ubiquitin specific peptidases, USP10 and USP13 that target the Beclin1 subunit of Vps34 complexes) and investigated the effect of the treatment of TBEV growth."

reach
"Spautin-1 promotes the degradation of Vps34 PI3 kinase complexes by inhibiting two ubiquitin specific peptidases, USP10 and USP13, that target the Beclin1 subunit of Vps34 complexes."
USP13 affects BCL2L1
| 3
USP13 activates BCL2L1.
| 2
USP13 activates BCL2L1. 2 / 2
| 2

reach
"We further report that genetic or pharmacological inhibition of USP13 considerably reduces MCL1 protein abundance and significantly increases tumor cell sensitivity to BH3 mimetic inhibitors targeting BCL-2 and BCL-XL."

reach
"Additionally, it is found that USP13 abrogates cell apoptosis by preventing excessive ubiquitin-proteasome degradation of Bcl-XL."
USP13 inhibits BCL2L1.
| 1
USP13 inhibits BCL2L1. 1 / 1
| 1

reach
"We further report that genetic or pharmacological inhibition of USP13 considerably reduces MCL1 protein abundance and significantly increases tumor cell sensitivity to BH3 mimetic inhibitors targeting BCL-2 and BCL-XL."
USP13 affects Aloe emodin
| 3

sparser
"Compared to the control, USP13 overexpression increased HIF-1α levels, while the enzymatically inactive mutant USP13-AE (C345A/M664/739 [ xref ]) failed to maintain them, indicating that USP13 stabilizes HIF-1α via its deubiquitinase activity."

sparser
"Notably, USP13-AE mutants exerted no significant effects on migratory, invasive, or wound healing capacities, demonstrating that USP13-mediated enhancement of cellular motility strictly requires its deubiquitinase activity (Fig.  xref I, S4C, S4D)."

sparser
"Given previous findings that USP13 stabilizes HIF-1α through its deubiquitinase activity, stable USP13-AE overexpressing cells exhibited comparable proliferation rates to control groups (Fig. S4A), suggesting that USP13 promotes CRC cell proliferation in a deubiquitinase activity-dependent manner."
USP13 affects ASNS
| 3
| 3

sparser
"Molecular mechanistic studies revealed that ASNS was highly bound to USP13 protein and increased USP13 expression."

sparser
"We found that ASNS was highly bound to USP13 proteins and co‐localized in the cytoplasm."

sparser
"To find the specific structural domains of ASNS binding to protein USP13, truncation of different regions of protein USP13 was constructed to co‐transfect ARPE‐19 cells with ASNS."
UFD1 affects USP13
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2 |

No evidence text available

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UBA1 affects Ubiquitin
| 3
| 3

sparser
"Moreover, the individual UBA domains (UBA1 and UBA2) of USP13 can also bind with Ub ( xref )."

sparser
"The tandem UBA domains of USP13 interact with Ub."

sparser
"Notably, the tandem UBA domains of USP13 could bind to Ub and catalyze the hydrolysis of the polyUb chains (Lys-63/48/27) ( xref ; xref ; xref )."
TRIM55 affects USP13
2 1 |
2 1 |

No evidence text available

No evidence text available

No evidence text available
Spautin‐1 affects USP13
| 3
Spautin‐1 inhibits USP13. 3 / 3
| 3

reach
"Spautin‐1 is an inhibitor of USP10 and USP13."

reach
"26 The role of autophagy in cerebral ischaemia‐reperfusion needs to be further investigated.Spautin‐1 was further used to investigate the mechanism of autophagy activation on cerebral ischaemia‐reperfusion injury, and Spautin‐1 is as an inhibitor of USP13."

reach
"Overexpression of USP13 in PC12 cells almost reversed the Spautin‐1‐mediated decrease in NLRP3, USP13, Beclin1 and GSDMD‐N expression levels (Figure 3D,E)."
SYK affects USP13
| 1 2
| 1 2

reach
"Protein-protein interactions assays showed that USP13 interacted with the co-immunoprecipitated protein spleen tyrosine kinase (SYK) and deubiquitinated SYK."

sparser
"Protein-protein interactions assays showed that USP13 interacted with the co-immunoprecipitated protein spleen tyrosine kinase (SYK) and deubiquitinated SYK."

sparser
"The USP13-SYK axis may serve as a therapeutic target for treating FcεRI-mediated activation of MCs and associated inflammatory responses."
STAG2 affects USP13
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No evidence text available

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No evidence text available
SIAH2 affects USP13
2 | 1
2 | 1

sparser
"Overall, this study identifies a new layer of Siah2 regulation mediated by USP13 binding to ubiquitinated Siah2 protein with a concomitant inhibitory effect on its activity under normoxia."

No evidence text available

No evidence text available
SCCPDH affects USP13
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
RPP25 affects USP13
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
EIF5B affects USP13
| 3
| 3

sparser
"USP13 interacts with eIF5B and negatively regulates its post-transcriptional expression."

sparser
"First, USP13 binds to eIF5B and enhances its interaction with SQSTM1/P62, promoting autophagy-lysosomal degradation of eIF5B. Second, previous studies show the functions of USP13 in SQSTM1/P62–USP13 stabilizes SQSTM1/P62 through depubiquitination, thus promoting the oligomerization of SQSTM1/P62 [ xref ], augmenting SQSTM1/P62’s substrate recruitment."

sparser
"Co-immunoprecipitation and immunoblot analysis confirmed the interaction between exogenous USP13 and endogenous eIF5B in NRVMs, with similar observations in the HEK−293T cell line (Fig. xref C, E)."
CDCA8 affects USP13
| 3
CDCA8 increases the amount of USP13. 3 / 3
| 3

reach
"TPM3, NECAP2, and USP13 expression were inhibited by CDCA8 knockdown in four HCC cell lines, suggesting that they were likely downstream molecules."

reach
"Next-generation sequencing and laboratory validation showed that CDCA8 silencing inhibited the expression of TPM3, NECAP2, and USP13."

reach
"The above results that CDCA8 may activate MAPK signaling through mediating USP13 levels."
Aloe emodin affects USP13
| 3

sparser
"Compared to the control, USP13 overexpression increased HIF-1α levels, while the enzymatically inactive mutant USP13-AE (C345A/M664/739 [ xref ]) failed to maintain them, indicating that USP13 stabilizes HIF-1α via its deubiquitinase activity."

sparser
"Notably, USP13-AE mutants exerted no significant effects on migratory, invasive, or wound healing capacities, demonstrating that USP13-mediated enhancement of cellular motility strictly requires its deubiquitinase activity (Fig.  xref I, S4C, S4D)."

sparser
"Given previous findings that USP13 stabilizes HIF-1α through its deubiquitinase activity, stable USP13-AE overexpressing cells exhibited comparable proliferation rates to control groups (Fig. S4A), suggesting that USP13 promotes CRC cell proliferation in a deubiquitinase activity-dependent manner."
ASNS affects USP13
| 3
| 3

sparser
"Molecular mechanistic studies revealed that ASNS was highly bound to USP13 protein and increased USP13 expression."

sparser
"We found that ASNS was highly bound to USP13 proteins and co‐localized in the cytoplasm."

sparser
"To find the specific structural domains of ASNS binding to protein USP13, truncation of different regions of protein USP13 was constructed to co‐transfect ARPE‐19 cells with ASNS."
2 |
Valproic acid decreases the amount of USP13. 2 / 2
2 |

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No evidence text available
Phenylmercury acetate decreases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
MiR-145-5p affects USP13
| 2
MiR-145-5p inhibits USP13. 2 / 2
| 2

reach
"MicroRNA profiling reveals miR-145-5p inhibits goat myoblast differentiation by targeting the coding domain sequence of USP13."

reach
"They also observed that miR-145-5p inhibited the development and apoptosis of goat myoblasts by suppressing USP13."
Imidazolide affects USP13
| 1 1
| 1 1

eidos
"USP13 preferentially is induced in GIST cells by IM and interacts with ATG5 , which leads to stabilization of ATG5 through deubiquitination ."

reach
"USP13 preferentially is induced in GIST cells by IM and interacts with ATG5, which leads to stabilization of ATG5 through deubiquitination."
Doxycycline affects USP13
| 2
| 2

reach
"To further confirm the clinical relevance of targeting USP13 in established GBM tumors, we applied the Tet-on inducible knockdown system to examine whether inducible disruption of USP13 by doxycycline affects the growth of established xenograft tumors and animal survival."

reach
"In vivo bioluminescent analysis confirmed that induced disruption of USP13 by doxycycline treatment significantly inhibited GSC tumor growth in mouse intracranial xenografts (XREF_FIG)."
Bisphenol A affects USP13
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Bisphenol A decreases the amount of USP13. 2 / 2
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No evidence text available

No evidence text available
2 |
Benzo[a]pyrene decreases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
AurK affects USP13
| 2
| 2

sparser
"Aurora B phosphorylates USP13 at Serine 114, prompting their interaction between USP13 and Aurora, showing a potential direct link between abnormalities occurring during the cell cycle and USP13 in several human cancers [ xref ]."

sparser
"Overall our study unveils the molecular and cellular connections of the USP13-Aurora B axis, which potentially participates in the rewiring of the cell cycle happening in cancer cells."
All-trans-retinoic acid decreases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
ZEB1 affects USP13
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VCL affects USP13
| 2
| 2

sparser
"Among the identified proteins, the altered expression level of vinculin was associated with the DUB activity of USP13, which was confirmed by western blot analysis and reverse transcription-quantitative polymerase chain reaction (RT-qPCR)."

sparser
"However, sufficient follow-up validation studies are required in order to determine whether vinculin protein directly interacts with USP13."
Ufd1 affects NPLOC4
| 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
USP5 affects USP13
2 |
2 |

No evidence text available

No evidence text available
USP22 affects USP13
| 2
| 2

sparser
"Coimmunoprecipitation and immunoblot analyses revealed that USP22 could interact with USP13, which could specifically cleave K27-linked ubiquitin chains on STING ( xref ), and the interaction between USP22 and USP13 became stronger after HSV-1 stimulation ( xref )."

sparser
"During virus infection, USP22 interacts with USP13 and specifically cleaves K27-linked ubiquitin chains on STING ( xref )."
USP15 affects USP13
| 1 1
| 1 1

sparser
"Furthermore, CRAF protein stability is maintained by physical interaction with USP13, USP15 and inhibitors of apoptosis proteins (IAPs) at the post-translational level [ xref – xref ]."

reach
"We next evaluated protein–protein interactions between USP13, USP15 and USP30 and ubiquitin/Parkin."
USP13 affects tumor growth
| 2
USP13 activates tumor growth. 2 / 2
| 2

eidos
"Moreover , the DUB USP7 , USP13 , USP22 , USP28 , USP36 and USP37 stabilize c-Myc , thereby stimulating tumor growth [ 155 , 157 , 171-174 ] ."
| PMC

eidos
"Previous studies showed that two deubiquitinases , USP9X and USP13 , stabilize MCL-1 protein levels and promote tumor growth 5 , 7 ."
USP13 affects stabilization TLR4
| 2
USP13 activates stabilization TLR4. 2 / 2
| 2

eidos
"Here , we demonstrated that USP13 enhanced the stabilization of TLR4 via direct binding and deubiquitylation of TLR4 ."

eidos
"USP13 knockdown reduced the stabilization of TLR4 ."
USP13 affects stabilisation Mcl-1
| 2
USP13 activates stabilisation Mcl-1. 2 / 2
| 2

eidos
"Taken together , these data suggest that USP13 mediated stabilisation of Mcl-1 may be a potential mechanism for the resistance of cervical cancer cells to BH3 mimetics ."

eidos
"These results suggest that the effect of USP13 on cell proliferation in cervical cancer cells is at least partially dependent on the stabilisation of Mcl-1 expression ."

reach
"USP13 was also showed to interact with aurora kinase B and disrupt cell cycle regulation (36)."

reach
"Furthermore, re-expression of cyclin D1 partially reversed the cell cycle arrest and cell proliferation inhibition induced by USP13 depletion in GC cells."
USP13 affects recruitment
| 1 1
USP13 inhibits recruitment. 2 / 2
| 1 1

sparser
"These data together suggest that USP13 inhibits the recruitment of TBK1 to STING by deubiquitinating STING."

eidos
"Previously , it has been demonstrated that USP13 removes K27-linked polyubiquitin chains from MITA and impairs its recruitment of TBK1 , contributing to inhibition of innate immune response to DNA viruses [ 36 ] ."
| 2
| 2

reach
"Collectively, our above findings confirm that USP13 is a strong blocker of NASH pathogenesis and its related metabolism dysregulation in mice.3.5 USP13 alleviates NASH pathogenesis by repressing IRHOM2-MAP3K7 signal cascade."

reach
"To explore the mechanism of attenuated virulence caused by usp13 deletion, we examined the effect of usp13 mutation on the ability to block activation of host phagocytes."
USP13 affects navitoclax
| 2
| 2

reach
"As anticipated, USP13 knockdown by siRNAs or knockout by CRISPR and Cas9 increased sensitivity to ABT-263 in SW-1573 and TOV-21G cells."

reach
"Conversely, USP13 overexpression in NCI-H2170 cells modestly impaired the inhibitory effects of ABT-263."
| 2

reach
"For example, USP13 stimulates melanoma cell invasion,20 drives ovarian cancer metabolism,21 and promotes the proliferation of glioma stem cells,22 which demonstrates the oncogenic roles of USP13 in these cancers."

reach
"However, studies have also shown that USP13 may act as an oncogene by promoting Mcl-1 stability in cervical cancer [36], and USP13 also drives the progression of ovarian cancer by promoting cancer metabolism [37]."
| 2

reach
"Edwardsiella piscicida USP13 is essential for pathogenicity and can help block the host immune response to pathogen infection [80]."

reach
"For example, USP3, USP25, and USP18 played a positive regulation of the antiviral immune response [ 5–8 ], while USP5, USP4, USP13, USP15, USP14, USP2b, and USP19 negatively regulated antiviral immune[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP13 affects half-life MITF-M
| 2
USP13 activates half-life MITF-M. 2 / 2
| 2

eidos
"As per the outcomes , USP13 upregulation mitigated the MITF-M degradation and significantly increased the half-life of MITF-M ( Figure 5C ) ."

eidos
"In addition , USP13 upregulation mitigated the MITF-M degradation and significantly increased the half-life of MITF-M ."
USP13 affects glucose
| 2
| 2

reach
"Indeed, knockdown of USP13 increased glucose uptake and glycolysis, which could be fully reversed by restoration of PTEN, as gauged by lactate production and glucose incorporation assays (XREF_FIG)."

reach
"Therefore, USP13-KD also decreases glucose oxidation into TCA cycle metabolites."
| 2

reach
"Functional studies demonstrated that overexpression of USP13 suppressed OSCC cell proliferation, glucose uptake and lactate production in vitro and inhibited tumor growth in vivo."

reach
"USP13 deficiency promotes glucose uptake and glycolysis (51)."
USP13 affects glioma stem cells
| 2
USP13 activates glioma stem cells. 2 / 2
| 2

eidos
"In addition , USP13 stabilizes microphthalmia-associated transcription factor and promotes cell invasion in melanoma [ 20 ] ; and , USP13 inhibition decreases proliferation of glioma stem cells by inducing c-Myc degradation [ 21 ] ."

eidos
"Moreover , USP13 depletion represses the proliferation of glioma stem cells ( GSCs ) and restrains tumor growth in mice ( Fang et al ., 2017 ) ."

reach
"N6-methyladenosine-modified USP13 induces pro-survival autophagy and imatinib resistance via regulating the stabilization of autophagy-related protein 5 in gastrointestinal stromal tumors."

reach
"On the contrary, IGF2BP2 elevates the stability of a deubiquitinase ubiquitin specific peptidase 13 (USP13) to prolong the half-life of autophagy-related protein 5 (ATG5), inducing pro-survival autophagy and drug resistance in gastrointestinal stromal tumors 60."
USP13 affects aurK
| 2
| 2

sparser
"Aurora B phosphorylates USP13 at Serine 114, prompting their interaction between USP13 and Aurora, showing a potential direct link between abnormalities occurring during the cell cycle and USP13 in several human cancers [ xref ]."

sparser
"Overall our study unveils the molecular and cellular connections of the USP13-Aurora B axis, which potentially participates in the rewiring of the cell cycle happening in cancer cells."
USP13 affects ZEB1
2 |
2 |

No evidence text available

No evidence text available
| 2

eidos
"Our results demonstrate that USP13 is hijacked to maintain nsp13 expression and the inhibitory role of nsp13 in regulating type I IFN production , and USP13 inhibitor could be employed to suppress virus replication by targeting nsp13 for degradation thereby disrupting its inhibitory role in regulating type I IFN production ."

eidos
"Moreover , depletion of USP13 or treatment with USP13 inhibitor relieves the inhibitory role of nsp13 for type I IFN response and suppresses virus replication in host cells , suggesting that USP13 inhibitor could be employed to suppress virus replication by targeting nsp13 for degradation ."
USP13 affects USP5
2 |
2 |

No evidence text available

No evidence text available
USP13 affects USP22
| 2
| 2

sparser
"Coimmunoprecipitation and immunoblot analyses revealed that USP22 could interact with USP13, which could specifically cleave K27-linked ubiquitin chains on STING ( xref ), and the interaction between USP22 and USP13 became stronger after HSV-1 stimulation ( xref )."

sparser
"During virus infection, USP22 interacts with USP13 and specifically cleaves K27-linked ubiquitin chains on STING ( xref )."
USP13 affects USP15
| 1 1
| 1 1

sparser
"Furthermore, CRAF protein stability is maintained by physical interaction with USP13, USP15 and inhibitors of apoptosis proteins (IAPs) at the post-translational level [ xref – xref ]."

reach
"We next evaluated protein–protein interactions between USP13, USP15 and USP30 and ubiquitin/Parkin."
USP13 affects TXN
2 |
2 |

No evidence text available

No evidence text available
USP13 affects TP53
| 2
| 2

sparser
"Several studies have reported that the expression of PTEN and that of the apoptosis-inducing protein p53 is positively associated with USP13 ( xref – xref )."

sparser
"Additionally, USP13 binds to P53, impacting the expression of the P53 protein."
USP13 affects TCRalpha
| 2
USP13 activates TCRalpha. 2 / 2
| 2

reach
"Immunoblotting confirmed that USP13 depletion caused more TCRalpha to partition into the NP40-insoluble fractions (XREF_FIG) compared to control cells."

reach
"When the degradation of the model ERAD substrate TCRalpha was examined in cells overexpressing either USP13 or the USP13 variants, we found that wild-type USP13 significantly increased the half-life of TCRalpha (from 1.6 hr to 4.6 hr)."
USP13 affects SMURF1
| 2
| 2

sparser
"BRCC36 modulates the ubiquitination status and stability of STAT1 by forming a regulatory complex with USP13 and Smurf1 ( xref C), which fine-tunes its activity in immune signaling [ xref ]."

sparser
"However, both Smurf1 and USP13 do not interact with pY701-STAT1 and ubiquitination regulation of STAT1 by them is independent of IFNs treatment [ xref , xref ]."
USP13 affects SMAD3
| 2
| 2

sparser
"USP13 binds to the MH2 domain of SMAD3 and deubiquitinates SMAD3."

sparser
"This study illustrates an HSC-specific USP13-SMAD3 axis in regulating liver fibrosis and presents USP13 as a potential target for the treatment of liver fibrosis."
USP13 affects SLC2A1
| 2
USP13 decreases the amount of SLC2A1. 2 / 2
| 2

reach
"Furthermore, USP13 overexpression induced phosphatase and tensin homolog deleted on chromosome 10 (PTEN) expression and repressed the activation of AKT as well as the expression of the downstream effectors glucose transporter-1 (GLUT1) and hexokinase-2 (HK2)."

reach
"27 In our experiments, USP13 overexpression reduced the levels of GLUT1 and HK2 in OSCC cells, while knockdown of USP13 had the opposite effects."
USP13 affects SI
| 2
USP13 activates SI. 2 / 2
| 2

reach
"Among them, 17.7% (69 of 390) overlapped with USP13 knockdown down-regulated genes (SI Appendix, Fig. S5E and Table S2)."

reach
"Similarly, our results showed that depletion of USP13 by single guide RNAs also inhibited cell proliferation and colony growth (SI Appendix, Fig. S4 J and K and Fig. 3 K and L)."

reach
"Silencing of USP13 in Exos evidently suppressed the M2 shift and upregulated ROS production in microglia in comparison with the control by qRT-PCR and flow cytometry analysis (Additional file 1: Fig. S5a–g)."

reach
"Similarly, USP10 and USP13 silencing could induce ROS generation and cause DNA damage."
| PMC
USP13 affects PRKN
| 2
USP13 leads to the deubiquitination of PRKN. 2 / 2
| 2

reach
"Here, it was shown that USP13 negatively regulates Parkin ubiquitination, function, and solubility (Figure 5), negatively regulates the level, ubiquitination, and clearance of α-synuclein, and regulates proteasome activity independently of Parkin, which affects dopaminergic neuronal viability and motor performances in animal models."

reach
"Second, USP13 knockdown can increase Parkin ubiquitination and activity, thereby reducing the level of α-Syn; the excessive expression of USP13 increases the expression of α-Syn [68,69]."
| 2
| 2

reach
"However, USP13-upregulated HOS cells were more sensitive to the FBS gradient and exhibited a stronger directed migration (Fig. 2f)."

reach
"Downregulation of USP13 significantly attenuated the invasiveness of 143B cells, while overexpression of USP13 promoted the invasiveness of HOS cells by using the transwell invasion assay (Fig. 2a, b)."
USP13 affects NFkappaB
| 1 1
| 1 1

eidos
"USP13 suppresses NF-kB to promote PTEN expression , resulting in antitumor activity [ 65 ] ."

reach
"USP13 suppresses NF-kB to promote PTEN expression, resulting in antitumor activity [XREF_BIBR]."
USP13 affects Mice
| 2
USP13 activates Mice. 2 / 2
| 2

reach
"Knockdown of Twist1 reversed the migration and invasion capacities of breast cancer cells and the occurrence of lung metastasis in mice caused by overexpression of USP13."

reach
"Notably, in HFHC-induced mice NASH model, liver USP13 protein expressive abundance was inversely correlated with the serum ALT, AST, liver Tnfa mRNA levels, and liver IRHOM2 protein levels (Fig. S6E–S6J)."
USP13 affects Melanoma
| 2
| 2

reach
"For example, USP13 stimulates melanoma cell invasion,20 drives ovarian cancer metabolism,21 and promotes the proliferation of glioma stem cells,22 which demonstrates the oncogenic roles of USP13 in these cancers."

reach
"USP13 promotes melanoma cell invasion by stabilizing the microphthalmia-associated transcription factor."
USP13 affects MYO15A
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects MTUS2
| 2
USP13 activates MTUS2. 2 / 2
| 2

reach
"Hence, targeting USP13 may lead to DDR deficiency and high MSI as well as activation of innate immune and antitumor immunity, and may promote a synergetic effect with DDR inhibitors and ICIs."

reach
"To date, no further studies have been carried out to demonstrate the potential role of USP13 in anticancer therapy, nor revealing that targeting USP13 may synergize with DDR inhibitors or ICIs."
USP13 affects MRPL20
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects MITF-M protein
| 2
USP13 activates MITF-M protein. 2 / 2
| 2

eidos
"Furthermore , Exogenous USP13 significantly upregulated the endogenous MITF-M protein level , downregulated USP13 significantly inhibited MITF-M protein levels , without altering MITF-M mRNA expression ."

eidos
"Conversely , USP13 upregulation promoted MITF-M protein levels , without influencing MITF-M mRNA levels ."
USP13 affects LDH
| 2
USP13 increases the amount of LDH. 2 / 2
| 2

reach
"In addition, specific inhibitors of mTOR, ULK1, USP10/USP13, and ULK1/ULK2 involved in the process of autophagy, i.e., temsirolimus, SBI-0206965, spautin-1, and MRT68921 dihydrochloride, decreased LDH release to 304%, 301%, 273%, and 271% of the control, respectively (Fig. 1b)."

reach
"Except for temsirolimus (mTOR inhibitor), specific inhibitors of kinases ULK1 and ULK1/ULK2 (SBI-0206965 and MRT68921 dihydrochloride) as well as hydrolases USP10/USP13 (Spautin-1) involved in the process of autophagy decreased LDH release to 138%, 141%, and 134% of the control, respectively (Fig. 1a)."
USP13 affects LAPTM5
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects L02
| 2
USP13 binds RHBDF2 and L02. 2 / 2
| 2

reach
"Finally, the interaction of USP13, UBC13 and IRHOM2 was detected in the L02 cells."

reach
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection (Fig. S3C–S3E)."
USP13 affects KDM3A
| 1 1
| 1 1

sparser
"Mechanistically, we identify that non-classic deubiquitinase USP13 interacts with KDM3A to promote its degradation in cytoplasm via the proteasome-specific pathway."

reach
"Collectively, our findings identify KDM3A is an important regulator of bladder cancer cell growth and metastasis and targeting USP13/KDM3A complex could be a valuable strategy to ameliorate bladder cancer progression and metastasis."
USP13 affects IRHOM2
| 2
USP13 binds IRHOM2. 2 / 2
| 2

sparser
"Currently, we determined, for the first time, that liver USP13 interacts with IRHOM2 and restrains its K63-linked ubiquitination of IRHOM2 by UBC13."

sparser
"Mechanistically, liver USP13 could directly interact with IRHOM2 and dismantle its metabolic stress-triggered K63-linked ubiquitination by ubiquitin-conjugating enzyme UBC13."
USP13 affects IRF3
1 | 1
1 | 1

No evidence text available

reach
"Although USP13 interacted with cGAS and IRF3 in an overexpression system, USP13 did not affect the ubiquitination of cGAS or IRF3 when coexpressed in HEK293 cells (XREF_SUPPLEMENTARY)."
2 |

No evidence text available

No evidence text available
USP13 affects G3BP1
2 |
2 |

No evidence text available

No evidence text available
USP13 affects FUNDC2
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects FNBP1L
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects FAF2
| 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
| 2
| 2

reach
"In melanoma cells, USP13 attenuates the autodegradation of Siah2, a RING (really interesting new gene) finger family E3 ubiquitin ligase."

reach
"Thus, several ubiquitin-conjugating E2 enzymes (UBE2L3, UBE2L5), the E3 ubiquitin ligases (TRIM21, TRIM38, ARIH2) and the ubiquitin specific protease or deconjugases (USP13) are 1.5-3.2-fold induced in the infected cells, while other E2, E3 enzymes and deconjugases (e.g."
| DOI
USP13 affects DVL1
1 1 |
1 1 |

No evidence text available

No evidence text available
USP13 affects CSNK2A1
2 |
2 |

No evidence text available

No evidence text available
USP13 affects CCN4
| 1 1
| 1 1

sparser
"Mass spectrometry, immunoprecipitation, and in vitro functional assays were conducted to explore the interaction between USP13 and WISP1 and to assess the effects of USP13 downregulation on cell proliferation, migration, invasion, epithelial-mesenchymal transition (EMT), and apoptosis."

reach
"Mass spectrometry, immunoprecipitation, and in vitro functional assays were conducted to explore the interaction between USP13 and WISP1 and to assess the effects of USP13 downregulation on cell proliferation, migration, invasion, epithelial-mesenchymal transition (EMT), and apoptosis."
USP13 affects ATXN3
1 |
1 |

No evidence text available
USP13 affects ATP
| 2
USP13 activates ATP. 2 / 2
| 2

reach
"USP13 specifically deubiquitinates and thus upregulates ATP citrate lyase and oxoglutarate dehydrogenase, two key enzymes that determine mitochondrial respiration, glutaminolysis and fatty acid synthesis."

reach
"Therefore, USP13 suppression decreases ATP production by repressing the TCA cycle and inhibiting de novo lipogenesis [99]."
USP13 affects 7H-purine
| 2
| 2

reach
"CLK3 dependent phosphorylation of USP13 at Y708 promotes CCA progression by activating c-Myc-mediated purine synthesis."

reach
"Therefore, we propose that CLK3 induced Y708 phosphorylation of USP13 promotes CCA progression by activating c-Myc-mediated purine synthesis."
USP10 affects BECN1
| 2
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
UBL affects BAG6
| 2
UBL binds USP13 and BAG6. 2 / 2
| 2

sparser
"On the other hand, the binding of USP13 with the UBL domain of Bag6 can be demonstrated using pulldown experiments ( xref )."

sparser
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."
TXNIP affects USP13
| 1 1
| 1 1

sparser
"As shown in xref , USP13 was detected in the TXNIP antibody-precipitated complex, suggesting that USP13 is associated with TXNIP."

reach
"The study may therefore provide a molecular basis to develop new small molecules to degrade TXNIP based on TXNIP/USP13 interaction.Two independent studies demonstrated that TXNIP exhibits a pro-fibrotic property in renal fibrosis [25,26], but has been shown to exhibit an anti-fibrotic property in the liver [27]."
TXN affects USP13
2 |
2 |

No evidence text available

No evidence text available
TP53 affects USP13
| 2
| 2

sparser
"Several studies have reported that the expression of PTEN and that of the apoptosis-inducing protein p53 is positively associated with USP13 ( xref – xref )."

sparser
"Additionally, USP13 binds to P53, impacting the expression of the P53 protein."
SNAI1 affects USP13
1 | 1
1 | 1

No evidence text available

sparser
"USP13 interacted with Snail to deubiquitinate and stabilize Snail in GC cells."
SMURF1 affects USP13
| 2
| 2

sparser
"BRCC36 modulates the ubiquitination status and stability of STAT1 by forming a regulatory complex with USP13 and Smurf1 ( xref C), which fine-tunes its activity in immune signaling [ xref ]."

sparser
"However, both Smurf1 and USP13 do not interact with pY701-STAT1 and ubiquitination regulation of STAT1 by them is independent of IFNs treatment [ xref , xref ]."
SMAD3 affects USP13
| 2
| 2

sparser
"USP13 binds to the MH2 domain of SMAD3 and deubiquitinates SMAD3."

sparser
"This study illustrates an HSC-specific USP13-SMAD3 axis in regulating liver fibrosis and presents USP13 as a potential target for the treatment of liver fibrosis."
RREB1 affects USP13
| 2
RREB1 increases the amount of USP13. 2 / 2
| 2

reach
"Similarly, overexpression of RREB1 significantly increased the protein and mRNA level of USP13 ( Figures 5 D and 5E)."

reach
"These results suggest that RREB1 might drive USP13 transcription."
RHBDF2 affects L02
| 2
USP13 binds RHBDF2 and L02. 2 / 2
| 2

reach
"Finally, the interaction of USP13, UBC13 and IRHOM2 was detected in the L02 cells."

reach
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection (Fig. S3C–S3E)."
Particulate Matter increases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
OGDH affects ACLY
| 2
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."
NPLOC4 affects Ufd1
| 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
NFE2 affects USP13
2 |
NFE2 decreases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
MYO15A affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
MRPL20 affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
MEF2A affects USP13
2 |
MEF2A decreases the amount of USP13. 2 / 2
2 |

No evidence text available

No evidence text available
LAPTM5 affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
L02 affects USP13
| 2
USP13 binds RHBDF2 and L02. 2 / 2
| 2

reach
"Finally, the interaction of USP13, UBC13 and IRHOM2 was detected in the L02 cells."

reach
"Indeed, significant increase in ubiquitination of IRHOM2, and occurrence of protein binding of USP13 and IRHOM2 in corresponding vectors-transfected L02 cells were further confirmed by co-immunoprecipitation (Co-IP) detection (Fig. S3C–S3E)."
KDM3A affects USP13
| 1 1
| 1 1

sparser
"Mechanistically, we identify that non-classic deubiquitinase USP13 interacts with KDM3A to promote its degradation in cytoplasm via the proteasome-specific pathway."

reach
"Collectively, our findings identify KDM3A is an important regulator of bladder cancer cell growth and metastasis and targeting USP13/KDM3A complex could be a valuable strategy to ameliorate bladder cancer progression and metastasis."
IRHOM2 affects USP13
| 2
USP13 binds IRHOM2. 2 / 2
| 2

sparser
"Currently, we determined, for the first time, that liver USP13 interacts with IRHOM2 and restrains its K63-linked ubiquitination of IRHOM2 by UBC13."

sparser
"Mechanistically, liver USP13 could directly interact with IRHOM2 and dismantle its metabolic stress-triggered K63-linked ubiquitination by ubiquitin-conjugating enzyme UBC13."
IRF3 affects USP13
1 | 1
1 | 1

No evidence text available

reach
"Although USP13 interacted with cGAS and IRF3 in an overexpression system, USP13 did not affect the ubiquitination of cGAS or IRF3 when coexpressed in HEK293 cells (XREF_SUPPLEMENTARY)."
IGF2BP2 affects USP13
| 2
IGF2BP2 activates USP13. 2 / 2
| 2

reach
"Mechanically, USP13’s m A modification is catalyzed by METTL3, and IGF2BP2 promotes USP13 mRNA translation by identifying m A residues on USP13 transcripts, ultimately triggering autophagy [169]."

reach
"On the contrary, IGF2BP2 elevates the stability of a deubiquitinase ubiquitin specific peptidase 13 (USP13) to prolong the half-life of autophagy-related protein 5 (ATG5), inducing pro-survival autophagy and drug resistance in gastrointestinal stromal tumors 60."
Hep KO mice affects USP13
| 2
Hep KO mice activates USP13. 2 / 2
| 2

reach
"The Hep KO mice were injected with AAV-TBG-Usp13 (Hep GOF) or AAV-TBG-Usp13 (ΔUBA) (Hep GOF) to induce corresponding USP13 gain-of-function (GOF) expression, followed by 24-weeks NASH-provoking HFHC diet feeding."

reach
"The Hep KO mice were then injected with a 1.5 × 10 genome copies (gc) dose of corresponding vectors to produce Usp13 gain-of-function mice."
2 |

No evidence text available

No evidence text available
HYCC1 affects USP13
| 2
HYCC1 activates USP13. 2 / 2
| 2

reach
"Thus, CDCA8 promoted HCC proliferation and invasion by up-regulating TPM3, NECAP2, and USP13 via MEK/ERK pathway."

reach
"Thus, USP13 deficiency suppressed HCC cell proliferation, migration, and stemness in vitro.To further validate the function of USP13 in HCC, we established USP13-overexpressing HCC cells via lentivirus (Supplementary Fig. S2A, B)."
G3BP1 affects USP13
2 |
2 |

No evidence text available

No evidence text available
FUNDC2 affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
FNBP1L affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
FAF2 affects Ufd1
| 2
| 2

sparser
"USP13 interacts with p97/VCP, Ufd1, Npl4, and UBXD8 [ xref ]."

sparser
"It was shown that USP13 associates with p97/VCP, Ufd1, Npl4, and UBXD8, and that depletion of USP13 leads to the accumulation of ERAD substrates [ xref ]."
DVL1 affects USP13
1 1 |
1 1 |

No evidence text available

No evidence text available
CSNK2A1 affects USP13
2 |
2 |

No evidence text available

No evidence text available
CCN4 affects USP13
| 1 1
| 1 1

sparser
"Mass spectrometry, immunoprecipitation, and in vitro functional assays were conducted to explore the interaction between USP13 and WISP1 and to assess the effects of USP13 downregulation on cell proliferation, migration, invasion, epithelial-mesenchymal transition (EMT), and apoptosis."

reach
"Mass spectrometry, immunoprecipitation, and in vitro functional assays were conducted to explore the interaction between USP13 and WISP1 and to assess the effects of USP13 downregulation on cell proliferation, migration, invasion, epithelial-mesenchymal transition (EMT), and apoptosis."
BECN1 affects USP10, and USP13
| 2
| 2

sparser
"Interestingly, the DUB activities were significantly increased when USP13 and USP10 coincubated together or with Beclin1 or all 3 proteins together, suggesting the DUB activity can be significantly enhanced when USP13 interacts with its substrate Beclin1 or USP10."

sparser
"To determine the mechanism by which Vps34 complexes regulate the stability of USP13 and USP10, we examined the possibility that Beclin1 may interact with USP10 and USP13."
BAG6 affects UBL, and USP13
| 2
UBL binds USP13 and BAG6. 2 / 2
| 2

sparser
"On the other hand, the binding of USP13 with the UBL domain of Bag6 can be demonstrated using pulldown experiments ( xref )."

sparser
"Taken together, these results suggest a direct interaction between USP13 and Bag6 that is mediated by the Bag6 UBL domain."
BAG6 affects AMFR
| 2
USP13 binds AMFR and BAG6. 2 / 2
| 2

sparser
"USP13 forms a multiprotein complex with gp78 and Bag6."

sparser
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."
AURKB affects USP13
1 | 1
AURKB phosphorylates USP13 on S114. 2 / 2
1 | 1

reach
"Specifically, Esposito et al. found that Aurora kinase B, crucial for mitosis, phosphorylates USP13 at the serine 114 and that this phosphorylation enhances the interaction of both proteins, thereby maintaining the stability of Aurora kinase B. Intriguingly, USP13 stabilizes Aurora kinase B by modulating its ubiquitination, a process that occurs independently of its DUB activity."

No evidence text available
ATXN3 affects USP13
1 |
1 |

No evidence text available
AMFR affects BAG6, and USP13
| 2
USP13 binds AMFR and BAG6. 2 / 2
| 2

sparser
"USP13 forms a multiprotein complex with gp78 and Bag6."

sparser
"Since Bag6 can interact with both gp78 and USP13, we asked whether these proteins could form a multiprotein complex."
ACLY affects OGDH, and USP13
| 2
ACLY binds OGDH and USP13. 2 / 2
| 2

sparser
"Western blot analyses of HEK293T cell lysates immunoprecipitated with USP13 antibody showed the interaction of USP13 with ACLY and OGDH ( xref )."

sparser
"Reciprocal analyses in which immunoprecipitated ACLY or OGDH were probed for USP13 confirmed the USP13ACLY and USP13OGDH interactions ( xref )."

reach
"As shown in Figure 7 D, we detected that 2-methoxyestradiol significantly inhibited USP13-mediated cleavage of K63-Ub4 ( Figure 7 D)."

reach
"Then, we used K63-linked tetra-Ub (K63-Ub4) as substrate to further confirm whether 2-methoxyestradiol could inhibit catalytic activity of USP13 toward K63-linked polyubiquitination."
Saxitoxin affects USP13
1 |
Saxitoxin increases the amount of USP13. 1 / 1
1 |

No evidence text available
Perfluorooctanesulfonamide decreases the amount of USP13. 1 / 1
1 |

No evidence text available
Isobutanol affects USP13
1 |
Isobutanol increases the amount of USP13. 1 / 1
1 |

No evidence text available
Indometacin affects USP13
1 |
Indometacin increases the amount of USP13. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548f-5p decreases the amount of USP13. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-539-3p decreases the amount of USP13. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3118 decreases the amount of USP13. 1 / 1
1 |

No evidence text available
Estrogen affects USP13
1 |
Estrogen decreases the amount of USP13. 1 / 1
1 |

No evidence text available
Adeno-associated virus serotype-8 affects USP13
| 1
Adeno-associated virus serotype-8 increases the amount of USP13. 1 / 1
| 1

reach
"In the indicated experiments, the conditional overexpression of Usp13 in hepatocytes (Hep OE) were triggered by injection of adeno-associated virus serotype-8 (AAV8)-thyroxine-binding globulin promoter (TBG)-recombinase Cre vector (AAV8-TBG-Cre) via intravenous injection and then determined by immunoblotting assay."
Aconitine affects USP13
1 |
Aconitine increases the amount of USP13. 1 / 1
1 |

No evidence text available
VIM affects USP13
| 1
USP13 binds VIM and H460. 1 / 1
| 1

reach
"We failed to detect any change of vimentin protein degradation rate in USP13 -KO cells ( Figure S3 E), as well as USP13/vimentin interaction in H460 and H1299 cells ( Figures S3 F and S3G)."
1 |
USP13 phosphorylated on Y708 is active. 1 / 1
1 |

"CLK3 directly phosphorylated USP13 at Y708, which promoted its binding to c-Myc, thereby preventing Fbxl14-mediated c-Myc ubiquitination and activating the transcription of purine metabolic genes."
USP13 affects tumor initiation
| 1
USP13 activates tumor initiation. 1 / 1
| 1

eidos
"Some findings suggest that USP13 may promote tumor initiation or progression ."
USP13 affects transporter-1
| 1
USP13 decreases the amount of transporter-1. 1 / 1
| 1

reach
"Furthermore, USP13 overexpression induced phosphatase and tensin homolog deleted on chromosome 10 (PTEN) expression and repressed the activation of AKT as well as the expression of the downstream effectors glucose transporter-1 (GLUT1) and hexokinase-2 (HK2)."
USP13 affects stemness-related protein
| 1
USP13 increases the amount of stemness-related protein. 1 / 1
| 1

reach
"Furthermore, USP13 overexpression enhanced the sphere formation ability (Supplementary Fig. S2H) and stemness-related protein expression (Supplementary Fig. S2I) of HCC cells."
USP13 affects sensitivity cervical cancer cells BH3 mimetic ABT-263
| 1
USP13 activates sensitivity cervical cancer cells BH3 mimetic ABT-263. 1 / 1
| 1

eidos
"Finally , we demonstrated that depletion of USP13 expression or inhibition of USP13 enzymatic activity increased the sensitivity of cervical cancer cells to the BH3 mimetic inhibitor ABT-263 ."
USP13 affects pathways
| 1
USP13 inhibits pathways. 1 / 1
| 1

sparser
"As expected, USP13 inhibited inflammatory pathways, including MAPK and NF-κB signaling pathways, in the livers of NAFLD mice."
USP13 affects p38
| 1
USP13 phosphorylates p38. 1 / 1
| 1

sparser
"In summary, USP13 selectively activates p38 phosphorylation by binding to MKK3 in a catalytically and phosphorylation dependent manner, indicating a functional USP13–MKK3–p38 axis in CRC."
USP13 affects p-RPA32 S33 staining
| 1
USP13 activates p-RPA32 S33 staining. 1 / 1
| 1

eidos
"As shown in Fig. 3A-B and Fig. S2 , depletion of USP13 diminished p-RPA32 ( S33 ) staining after hydroxyurea ( HU ) treatment ."

reach
"N6-methyladenosine-modified USP13 induces pro-survival autophagy and imatinib resistance via regulating the stabilization of autophagy-related protein 5 in gastrointestinal stromal tumors."
USP13 affects lyase
| 1
USP13 activates lyase. 1 / 1
| 1

reach
"USP13 specifically deubiquitinates and thus upregulates ATP citrate lyase and oxoglutarate dehydrogenase, two key enzymes that determine mitochondrial respiration, glutaminolysis and fatty acid synthesis."
| 1
| 1

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"Different forms of stress modulate these activities : hypoxia reduces USP13 expression enabling increased Siah2 activity, while cellular stress induces p38 activity and alters Siah localization, affecting its recognition and targeting of substrates 174."
USP13 affects hiPSCs
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USP13 activates hiPSCs. 1 / 1
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"After seeding in hiPSC basal medium without Activin A and bFGF, USP13 overexpressing hiPSCs expressed OCT4, PRDM14, SOX2, and NANOG at higher levels (Fig. 8, D and E), but exhibited a slower differentiation rate than PB control cells (Fig. 8F)."
USP13 affects half-life mutant protein NSCLC cells
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USP13 activates half-life mutant protein NSCLC cells. 1 / 1
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"Moreover , USP13 inhibition and silencing lead to a decreased half-life of the mutant EGFR protein in NSCLC cells , indicating that USP13 likely stabilizes EGFR ( Figure 3D ) ."
USP13 affects growth metastasis mice
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USP13 activates growth metastasis mice. 1 / 1
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"USP13 knockdown inhibits HCC growth and metastasis in mice ."
USP13 affects cervical cancer cell
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USP13 activates cervical cancer cell. 1 / 1
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"Depletion of USP13 inhibited cervical cancer cell proliferation ."
USP13 affects active caspase
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USP13 inhibits active caspase. 1 / 1
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"The downregulation of USP13 also increased active caspase 3 , indicating tumor apoptosis ( Figure 5A ) ."
USP13 affects abundance anti-apoptotic protein MCL1 lung
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USP13 activates abundance anti-apoptotic protein MCL1 lung. 1 / 1
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"In lung infection , USP13 are aberrantly expressed , inhibition of USP13 reduces the abundance of anti-apoptotic protein MCL1 in the lung [ 42 ] ."
USP13 affects VIM
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USP13 binds VIM and H460. 1 / 1
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"We failed to detect any change of vimentin protein degradation rate in USP13 -KO cells ( Figure S3 E), as well as USP13/vimentin interaction in H460 and H1299 cells ( Figures S3 F and S3G)."
USP13 affects VHL
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USP13 deubiquitinates VHL. 1 / 1
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"Interestingly, USP13 has also been implicated in renal cancer, but rather than altering HIF activity, USP13 deubiquitinates another VHL target gene and oncogenic driver, ZHX2 [141]."
USP13 affects UBA domains
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UBA domains binds USP13 and mutated EGFR. 1 / 1
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"USP13 inhibits the degradation of mutant EGFR via deubiquitination, binds to mutant EGFR via its UBA domains, and inhibits EGFR endosomal sorting and lysosomal degradation."
USP13 affects TRPC4
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USP13 affects TRIM69
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USP13 activates TRIM69. 1 / 1
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"However, USP13, USP19, and USP22 inhibit virus-induced IFN production by removing K27-linked polyubiquitin chains from STING (40, 81) or TRIF (90)."
USP13 affects TCAF1
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USP13 affects TBK1
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USP13 inhibits TBK1. 1 / 1
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"To determine the specificity of the USP38-mediated TBK1 degradation, we used other USP family members USP3 and USP13 as a control, and we found that USP38, but not USP3 or USP13, induced TBK1 degradat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Like NLRP3, the NLRP3 NACHT domain and LRR domain interacted with USP13, but the NLRP3 PYRIN domain failed to interact with USP13 (Fig.  xref h)."
USP13 affects Protease
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USP13 leads to the ubiquitination of Protease. 1 / 1
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"16 For instance, USP13 was able to indirectly mediate the protein stability of p53 by regulating its deubiquitinating protease USP10."
USP13 affects NEDD8
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USP13 affects MAVS
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USP13 ubiquitinates MAVS. 1 / 1
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"Similarly, the enhanced K11-linked ubiquitination of MAVS caused by USP13 led to the increase of total ubiquitination."

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"In addition, upregulation of USP13 is proved to attenuate intervertebral disc degeneration (IVDD) through promoting autophagy (Dai et al., 2021)."
USP13 affects ITCH
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USP13 affects IL1B
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USP13 inhibits IL1B. 1 / 1
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"Both miR-19a-3p knockdown and USP13 overexpression in MH-S cells inhibited the LPS induced production of TNF-alpha, IL-6 and IL-1beta."
USP13 affects Hypoxia
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USP13 activates Hypoxia. 1 / 1
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"USP13 knockdown repressed hypoxia-induced activation of the TLR4/MyD88/NF-κB pathway in HCC cells [25] ."
USP13 affects HSPA4
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USP13 affects H460
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USP13 binds VIM and H460. 1 / 1
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"We failed to detect any change of vimentin protein degradation rate in USP13 -KO cells ( Figure S3 E), as well as USP13/vimentin interaction in H460 and H1299 cells ( Figures S3 F and S3G)."
USP13 affects Glioma
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USP13 inhibits Glioma. 1 / 1
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"USP13, FBXL14 and c-Myc can interact with each other; USP13 inhibits the degradation of c-Myc, hence inhibition of USP13 promotes the apoptosis of glioma stem cells."
USP13 affects Death
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USP13 activates Death. 1 / 1
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"In brief, USP13 and USP8 promote α-Syn-related pathology, and the knockdown of USP13 and USP8 reduces α-Syn-induced DA neuronal death."

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"USP13 activates mitochondrial energy metabolism, autophagy, DNA damage response, glutaminolysis, and fatty acid synthesis [73]."
USP13 affects DAZAP2
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USP13 affects CWC15
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USP13 affects CPT
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USP13 inhibits CPT. 1 / 1
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"Of the three genes, SENP2 had the greatest influence on IR resistance, while increased SENP2 and USP13 reduced CPT resistance (Supplemental Fig. S6F–H)Since our data show a critical role for SENP2 in DSB repair, we explored whether increased SENP2 expression alters repair."
USP13 affects CD4
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USP13 activates CD4. 1 / 1
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"In addition , in B-cells and CD4 + T-cells , high amplifications and arm level gains of USP13 resulted in increased tumor infiltration ."
| PMC
USP13 affects BCL2
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USP13 activates BCL2. 1 / 1
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"We further report that genetic or pharmacological inhibition of USP13 considerably reduces MCL1 protein abundance and significantly increases tumor cell sensitivity to BH3 mimetic inhibitors targeting BCL-2 and BCL-XL."
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"In addition , in B-cells and CD4 + T-cells , high amplifications and arm level gains of USP13 resulted in increased tumor infiltration ."
| PMC
USP13 affects ATR
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USP13 inhibits ATR. 1 / 1
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"Depletion of USP13 impedes ATR activation and hypersensitizes cells to replication stress inducing agents."
USP13 affects ATM
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USP13 activates ATM. 1 / 1
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"Unexpectedly, USP13 could not reverse the inhibition of ATM activity in shRAP80 transfected cells."
USP13 affects AKT inhibitors
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USP13 activates AKT inhibitors. 1 / 1
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"The results suggest that inhibition of USP13 may enhance the effects of PI3K and AKT inhibitors on killing OVCA cells."
USP13 affects 135b
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USP13 decreases the amount of 135b. 1 / 1
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"Furthermore, the suppression of USP13 increased endogenous miR‐135b expression."
USP-Y708F affects USP13
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USP-Y708F activates USP13. 1 / 1
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"CHX chase assays indicated that overexpression of WT-USP13, particularly a phosphomimetic USP13-Y708E, but not USP-Y708F, significantly up-regulated the half-time of the c-Myc protein in HEK293 cells (XREF_FIG)."
UBA domains affects USP13
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UBA domains binds USP13 and mutated EGFR. 1 / 1
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"USP13 inhibits the degradation of mutant EGFR via deubiquitination, binds to mutant EGFR via its UBA domains, and inhibits EGFR endosomal sorting and lysosomal degradation."
TRPC4 affects USP13
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TCAF1 affects USP13
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No evidence text available
Soot affects USP13
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Soot increases the amount of USP13. 1 / 1
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SARM1 affects USP13
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"USP13 interacts with SARM1 and regulates axonal integrity via SARM1."

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"26 The role of autophagy in cerebral ischaemia‐reperfusion needs to be further investigated.Spautin‐1 was further used to investigate the mechanism of autophagy activation on cerebral ischaemia‐reperfusion injury, and Spautin‐1 is as an inhibitor of USP13."

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"Like NLRP3, the NLRP3 NACHT domain and LRR domain interacted with USP13, but the NLRP3 PYRIN domain failed to interact with USP13 (Fig.  xref h)."
NF-kB p65 affects USP13
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Modified NF-kB p65 decreases the amount of USP13. 1 / 1
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"Results suggested that the overexpression of NF-kB p65 significantly decreased the expression levels of USP13 and PTEN, and re-introduction of USP13 rescued PTEN expression."
NEDD8 affects USP13
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MAPK affects USP13
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MAPK inhibits USP13. 1 / 1
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"Besides, U0126-mediated inhibition of KRAS effector MAP kinase kinase (MEK) decreased USP13 and vimentin protein and mRNA levels in A549 and H460 cells, both of which carry mutant KRAS ( Figures 4 G a[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
JNK inhibitor affects USP13
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JNK inhibitor activates USP13. 1 / 1
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"Furthermore, among the inhibitors of matrix metalloproteinases (MMPs), ROCK, p38 MAPK, and JNK, only JNK inhibitor II attenuated USP13 degradation (Fig. 7D)."
Infections affects USP13
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Infections increases the amount of USP13. 1 / 1
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"qRT-PCR and western blot results confirmed that infection with lentivirus harboring USP13 shRNA obviously decreased the expression level of USP13 in GC cells (Fig. 2B, C)."
ITCH affects USP13
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HSPA4 affects USP13
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H460 affects VIM
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USP13 binds VIM and H460. 1 / 1
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"We failed to detect any change of vimentin protein degradation rate in USP13 -KO cells ( Figure S3 E), as well as USP13/vimentin interaction in H460 and H1299 cells ( Figures S3 F and S3G)."
EGFR affects UBA domains, and USP13
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UBA domains binds USP13 and mutated EGFR. 1 / 1
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"USP13 inhibits the degradation of mutant EGFR via deubiquitination, binds to mutant EGFR via its UBA domains, and inhibits EGFR endosomal sorting and lysosomal degradation."

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"The addition of an MEK inhibitor impaired the effect of USP13 upregulation in these cells."
DAZAP2 affects USP13
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No evidence text available
CWC15 affects USP13
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No evidence text available
CBLB affects USP13
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CBLB activates USP13. 1 / 1
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"c-Cbl enhanced the interaction between EGFR-DeltaE740-A746 and USP13, while Cbl-b increased the binding of both c-Cbl and USP13 to mutant EGFR."