IndraLab

Statements


ADRM1 affects UCHL5
24 5 | 2 43 105
ADRM1 binds UCHL5.
24 5 | 25 72
24 4 | 24 65

reach
"The hRpn13 and UCH37 complex hydrolyzes large Ub conjugates with incorporation into the 19S complex."

sparser
"In contrast to a previous report, we find that RPN13 binds ubiquitin with an affinity similar to that of other proteasome-associated ubiquitin receptors and that RPN2, ubiquitin, and the deubiquitylase UCH37 bind to RPN13 with independent energetics."

No evidence text available

sparser
"With the exception of the shorter Rpn13 protein from S. cerevisiae , Rpn13 binds the DUB Uch37 and thereby recruits it to the proteasome's RP xref - xref ."

No evidence text available

No evidence text available

sparser
"Because of their shared domain architecture, our current understanding of PR-DUB catalytic activity is built upon both recent crystal structures of the Drosophila PR-DUB complex and prior structures of the UCHL5Rpn13 complex."

reach
"Thus, hRpn13 activates hINO80 associated Uch37 without displacing it from the hINO80 complex, and activation is a consequence of transient interactions between Uch37 and hRpn13."

sparser
"As expected, hRpn13 (253-407) binds Uch37 and intermolecular NOE interactions ( xref ) defined a compact binding surface composed of residues from H2, the H2-H3 loop, and H9 ( xref )."

sparser
"Therefore, the external stimuli of exogenous rHcADRM1 may hamper the ADRM1-Uch37 interaction in host T cells, disrupt the balance of cell cycle protein degradation and interfere with ADRM1 substrate IκBα signaling along with its downstream effectors [ xref , xref ]."
| PMC
| 2

sparser
"Because hRpn13 and NFRKB interact with Uch37 in the proteasome and hINO80, respectively, we asked whether the isolated proteins bind to Uch37 in a mutually exclusive fashion."

sparser
"Biological information indicated that both RPN13 and INO80G bind to a long fragment at the C-terminal of the UCH-L5."
1 | 1

sparser
"We tested this possibility through pharmacological inhibition of the ADRM1-PSMD1-UCH37 interaction by using RA190 xref ."

No evidence text available
| 2

sparser
"Structures of UCH-L5-Rpn13 bound to ubiquitin are available."

sparser
"RPN13 binding to ubiquitin chains and UCH37 might simultaneously facilitate UCH37's distal end xref deubiquitylating activity xref - xref by orienting the ubiquitin moieties in a configuration favorable to hydrolysis."
ADRM1 binds UCHL5 and monoubiquitin. 1 / 1
| 1

sparser
"Additionally, there has been recent development of another competitive FP assay that includes fluorescently-labeled monoubiquitin bound to the Uch37:Rpn13 complex and then competed off with unlabeled ubiquitin chains."
ADRM1 binds UCHL5 and NOS2. 1 / 1
| 1

sparser
"We previously found that the cisplatin treatment induced Rpn13 transcription by p-ΔNp63α and subsequently increased the physical interaction of Rpn13, UCH37 and NOS2 proteins leading to an essential degradation of the latter through a proteasome-dependent mechanism in SCC cells [ xref ]."
ADRM1 binds UCHL5 and UCH domain. 1 / 1
| 1

reach
"Binding of Adrm1 to UCH37 relieved auto-inhibition by the UCH domain and activated its deubiquitination activity [XREF_BIBR]."
ADRM1 activates UCHL5.
| 2 15 33
ADRM1 activates UCHL5. 10 / 67
| 2 15 33

sparser
"Consistent with our previous report that hRpn13 activates Uch37, the Uch37-hRpn13 complex reacts with UbVS more rapidly than Uch37 alone ( Figure 3 A)."

reach
"We further demonstrated that the carboxyl-terminal tail of Uch37 is auto-inhibitory and that hRpn13 activates the Uch37 deubiquitinating activity through interactions with this tail domain."

sparser
"Furthermore, ADRM1 activates the deubiquitinase UCH37 and avoids its auto-inhibition."

sparser
"However, this mutant was only marginally activated by the addition of RPN13 DEU (2.7-fold instead of 7-fold in WT), indicating that UCH-L5 activation by RPN13 DEU requires an intact CL ( xref F; xref and xref )."

sparser
"Stimulation of deubiquitylating activity can now be added to this list of proteasomal functions, but neither ATPase nor proteolytic function of the proteasome is required, as Rpn13 alone can activate Uch37."

sparser
"Their interaction with ubiquitin is not essential for RPN13 activation of UCH37 for ubiquitin-AMC, as their mutation to alanine produced a 2.5-fold reduction ( xref ) or negligible effect ( xref ); wild-type RPN13 increases this activity 8-fold ( xref ; xref )."

sparser
"RPN13 and INO80G have opposing effects on UCH37; RPN13 activates UCH37 toward model mono-Ub substrates and INO80G acts as an inhibitor."

eidos
"Rpn13 , the proteasomal receptor for Uch37 in the proteasome 19S regulatory particle , can activate UCH37 by disrupting dimerization.9 USP14 binds to the regulatory particle Rpn1 to release its catalytic USP domain and polyubiquitin chains of substrate protein.10 Overexpression of Rpn13 or Rpn1 upregulated the COPS5 protein levels and downregulated the p53 protein levels ( Supplementary Fig. S2p ) ."

sparser
"Ubiquitin receptors play an integral role in substrate capture and apparently contribute to ubiquitin chain deconjugation, as Rpn13 binds and activates deubiquitinating enzyme Uch37 ( xref ; xref ; xref )."

sparser
"This relocation simultaneously precludes Trp36 availability for ubiquitin binding and impedes the formation of the intricate polar interaction network in the ULD anchor that is required for UCH-L5 activation by RPN13 DEU ( xref C)."
ADRM1 inhibits UCHL5.
| 1
ADRM1 inhibits UCHL5. 1 / 6
| 1

reach
"We propose that RA190 targets hRpn13 and Uch37 through parallel mechanisms and at proteasomes, RA190 inactivated Uch37 can not disassemble hRpn13 bound ubiquitin chains."
ADRM1 increases the amount of UCHL5.
| 1
ADRM1 increases the amount of UCHL5. 1 / 1
| 1

reach
"Intriguingly, hRpn13 knockdown leads to reduced levels of Uch37 in cells XREF_BIBR XREF_BIBR and similar phenotypes are observed by hRpn13 or Uch37 knockdown XREF_BIBR; however, the mechanism linking hRpn13 to Uch37 protein levels remain unknown."
ADRM1 deubiquitinates UCHL5.
| 1
ADRM1 deubiquitinates UCHL5. 1 / 1
| 1

reach
"HRpn13 Pru domain binds proteasomes and ubiquitin whereas its DEUBAD domain binds deubiquitinating enzyme UCHL5."
UCHL5 affects USP14
2 | 6 71
UCHL5 binds USP14.
2 | 3 71
2 | 3 47

No evidence text available

sparser
"This is the first report to define the effects and underlying mechanisms associated with inhibition of USP14 and UCHL5 DUB activity in WM tumour cells."

sparser
"We hypothesize that ITCs, as electrophiles, can interact with the catalytic triads (CYS, HIS and ASP) of the proteasomal cysteine deubiquitinases USP14 and UCHL5, ultimately inhibiting their activities."

sparser
"As with USP14, UCH37 is associated with oncogenesis, although this DUB has been also implicated in many different cellular processes [ xref ]."

reach
"GST-pulldown and co-immunoprecipitation (co-IP) assays have been applied to investigate the interactions between USP14, UCHL5, and Skp2."

sparser
"Docking results suggest that benzyl isothiocyanate (BITC), phenethyl isothiocyanate (PEITC) and DL-Sulforaphane (SFN) are more potent inhibitors of UCHL5 than USP14, and these ITCs could interact with the catalytic triads of UCHL5 and USP14."

sparser
"b-AP15, a compound that has a similar structure as P1 and P2, has been found to inhibit the proteasome by specifically interacting with UCHL5 and USP14, resulting in an accumulation of high molecular weight poly-ubiquitinated proteins [ xref ]."

sparser
"We hypothesized that several electrophilic optical brighteners can interact with the catalytic triads (CYS, HIS, and ASP) of the proteasomal cysteine deubiquitinases (DUBs) UCHL5 and USP14 ( xref ; xref ), ultimately leading to inhibition of their activities."

sparser
"Recently, we have reported that the anti-cancer activity of copper (II) pyrithione CuPT and gold (I) complex auranofin is associated with targeting the 19S proteasome-associated deubiquitinases (DUBs), UCHL5 and USP14."

sparser
"Incubation of ZnPT at 5μM with 26S proteasomes substantially reduced UbVS binding to UCHL5 and abolished UbVS binding to USP14, and when ZnPt is increased to 50μM, UbVS binding to both UCHL5 and USP14 was abolished (Figure xref )."
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
RPN1 binds USP14 and UCHL5. 1 / 1
| 1

sparser
"Interestingly, USP14 and UCH37 bind to RPN1 and to RPN13, respectively, which are, in addition to the RPN10, substrate receptors of the 19S regulatory particle [ xref ], which provide a versatile binding platform for various ubiquitin chains [ xref ]."
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
UCHL5 inhibits USP14.
| 2
UCHL5 inhibits USP14. 2 / 2
| 2

reach
"b-AP15/VLX1570 are small molecule inhibitors of the ubiquitin specific peptidase 14 (USP14) and ubiquitin carboxyl-terminal hydrolase 5 (UCHL5) deubiquitinases (DUBs) of the 19S proteasome."

reach
"We also found that inhibition of USP-14 and UCHL5 activities by the ITCs caused increased levels of USP14 and UCHL5 proteins, but not the third 19S deubiquitinating enzyme (DUB), POH1 and RPN11, suggesting feedback loop activation and further supporting that ITCs are inhibitors of proteasomal cysteine DUBs."
UCHL5 ubiquitinates USP14.
| 1
UCHL5-C88A leads to the ubiquitination of USP14-C114A. 1 / 1
| 1

reach
"Besides beta-catenin, we also verified with immunoblotting that UCHL5 C88A inhibits its own deubiquitination and USP14 C114A inhibits deubiquitination of two proteasomal subunits PSMC1 and PSMD4."
USP14 affects UCHL5
2 | 5 71
USP14 binds UCHL5.
2 | 3 71
2 | 3 47

No evidence text available

sparser
"This is the first report to define the effects and underlying mechanisms associated with inhibition of USP14 and UCHL5 DUB activity in WM tumour cells."

sparser
"We hypothesize that ITCs, as electrophiles, can interact with the catalytic triads (CYS, HIS and ASP) of the proteasomal cysteine deubiquitinases USP14 and UCHL5, ultimately inhibiting their activities."

sparser
"As with USP14, UCH37 is associated with oncogenesis, although this DUB has been also implicated in many different cellular processes [ xref ]."

reach
"GST-pulldown and co-immunoprecipitation (co-IP) assays have been applied to investigate the interactions between USP14, UCHL5, and Skp2."

sparser
"Docking results suggest that benzyl isothiocyanate (BITC), phenethyl isothiocyanate (PEITC) and DL-Sulforaphane (SFN) are more potent inhibitors of UCHL5 than USP14, and these ITCs could interact with the catalytic triads of UCHL5 and USP14."

sparser
"b-AP15, a compound that has a similar structure as P1 and P2, has been found to inhibit the proteasome by specifically interacting with UCHL5 and USP14, resulting in an accumulation of high molecular weight poly-ubiquitinated proteins [ xref ]."

sparser
"We hypothesized that several electrophilic optical brighteners can interact with the catalytic triads (CYS, HIS, and ASP) of the proteasomal cysteine deubiquitinases (DUBs) UCHL5 and USP14 ( xref ; xref ), ultimately leading to inhibition of their activities."

sparser
"Recently, we have reported that the anti-cancer activity of copper (II) pyrithione CuPT and gold (I) complex auranofin is associated with targeting the 19S proteasome-associated deubiquitinases (DUBs), UCHL5 and USP14."

sparser
"Incubation of ZnPT at 5μM with 26S proteasomes substantially reduced UbVS binding to UCHL5 and abolished UbVS binding to USP14, and when ZnPt is increased to 50μM, UbVS binding to both UCHL5 and USP14 was abolished (Figure xref )."
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
RPN1 binds USP14 and UCHL5. 1 / 1
| 1

sparser
"Interestingly, USP14 and UCH37 bind to RPN1 and to RPN13, respectively, which are, in addition to the RPN10, substrate receptors of the 19S regulatory particle [ xref ], which provide a versatile binding platform for various ubiquitin chains [ xref ]."
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
USP14 inhibits UCHL5.
| 2
USP14 inhibits UCHL5. 2 / 2
| 2

reach
"By using overexpression model of chicken UCH-L5 and b-AP15 inhibitor of UCH-L5 and USP14 [XREF_BIBR], which inhibited UCH-L5 at low molar concentration, but it did not significantly inhibit other DUBs in HD11 cells (XREF_SUPPLEMENTARY), we observed the overexpression of UCH-L5 had a negative effect on cell viability, which depended on its catalytic activity as a DUB, since if UCH-L5 overexpressing cells were treated with b-AP15 inhibitor, the cell viability was partially restored (XREF_FIG)."

reach
"We also found that inhibition of USP-14 and UCHL5 activities by the ITCs caused increased levels of USP14 and UCHL5 proteins, but not the third 19S deubiquitinating enzyme (DUB), POH1 and RPN11, suggesting feedback loop activation and further supporting that ITCs are inhibitors of proteasomal cysteine DUBs."
NFRKB affects UCHL5
12 | 37 21
NFRKB binds UCHL5.
12 | 18 19
12 | 18 17

sparser
"The activity of UCH-L5 is enhanced when UCH-L5 combines with proteasome 19S regulatory subunit by Rpn13/Admr1 receptor and inhibited when UCH-L5 interacts with NFRKB."

sparser
"And UCH-L5 also interacts with NFRKB in INO80 complex, but it remains unknown that whether UCH-L5 interacts with other components of INO80 complex or not."

reach
"UCHL5 interacts with and protects NFRKB (also known as INO80G) that is a component of INO80 chromatin remodeling complex from proteasomal degradation by removing K48 linked ubiquitin chain."

sparser
"Importantly, two-hybrid assays detected an interaction between NFRKB and full-length Uch37, but not a truncated Uch37 that retains only the UCH domain."

reach
"However, the effect of DRAIC on the combination of UCHL5 and NFRKB was still unclarified, so we detected this combination in oeDRAIC and shDRAIC cell lines, whose results showed that oeDRAIC can significantly reduce the level of NFRKB coprecipitated by UCHL5, while shDRAIC can increase NFRKB binding with UCHL5, which confirmed the speculation that DRAIC may indirectly down-regulate the expression of NFRKB through affecting the deubiquitination induced by UCHL5."

No evidence text available

sparser
"While the activity of UCH-L5 is inhibited when UCH-L5 interacts with NFRKB, a component of INO80 complex, by combination and deubiquitination [ xref ]."

sparser
"Considering the deubiquitination function of UCHL5, we speculated that DRAIC might regulate the ubiquitination level of NFRKB by influencing the binding of UCHL5 and NFRKB, and then affect the protein expression of NFRKB."

reach
"Interestingly, immunoprecipitation studies from MG132 treated cells revealed that NFRKB interacted with UCHL5 less in the chromatin fraction than in the nucleoplasm, despite its interactions with INO80 being essentially equivalent in these two fractions (XREF_FIG; for input fractions, see XREF_SUPPLEMENTARY)."

No evidence text available
| 2

sparser
"Because hRpn13 and NFRKB interact with Uch37 in the proteasome and hINO80, respectively, we asked whether the isolated proteins bind to Uch37 in a mutually exclusive fashion."

sparser
"Biological information indicated that both RPN13 and INO80G bind to a long fragment at the C-terminal of the UCH-L5."
NFRKB inhibits UCHL5.
| 11 2
| 11 2

sparser
"In contrast, NFRKB inhibits UCH37 by blocking the ubiquitin-binding site and by disrupting the enzyme active site."

reach
"Uch37 activity can be inhibited by NFRKB alone or by incorporation into hINO80."

sparser
"Mechanism of UCH-L5 Inhibition by INO80G DEU ."

reach
"Interestingly, UCHL5 is activated by the proteasome subunit RPN13 and inhibited by the INO80G subunit ."

reach
"By contrast, UCH37 is inhibited by the chromatin remodeling complex component INO80G mediated by the N-terminal domain of NFRKB (nuclear factor related to kappaB, NFRKB)."

reach
"The deubiquitinating enzyme UCH-L5 can be inhibited and activated by regulatory proteins INO80G and RPN13, respectively."

reach
"Our binding assays showed that INO80G DEU decreases the affinity of UCH-L5 for substrates (XREF_FIG A and 2B)."

reach
"The related DEUBAD domain in INO80G inhibits UCH-L5 by exploiting similar structural elements in UCH-L5 to promote a radically different conformation, and employs molecular mimicry to block ubiquitin docking."

reach
"NFRKB NTD inhibits UCH37 because of two distinct contacts that it makes with the UCH domain."

reach
"Surprisingly, the truncated protein INO80G DEU Deltaalpha6 still inhibited UCH-L5 (XREF_FIG B), demonstrating that helix alpha6 is not required for inhibition under these conditions."
NFRKB activates UCHL5.
| 8
NFRKB activates UCHL5. 8 / 8
| 8

reach
"To determine whether NFRKB also modulates Uch37 activity, we affinity purified from insect cells recombinant Uch37 in complexes with NFRKB, various NFRKB derivatives, or hRpn13 and assayed Uch37 for i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Why does NFRKB DEU fail to activate Uch37?"

reach
"Enzyme kinetics analysis confirmed that INO80G short activates UCH-L5 on Ub-AMC (XREF_FIG B)."

reach
"To determine whether NFRKB also modulates Uch37 activity, we affinity purified from insect cells recombinant Uch37 in complexes with NFRKB, various NFRKB derivatives, or hRpn13 (XREF_SUPPLEMENTARY) and assayed Uch37 for its ability to react with ubiquitin vinylsulfone (UbVS)."

reach
"INO80G short, containing only the helices alpha2-alpha4 of the DEUBAD domain, can activate UCH-L5, demonstrating that the core DEUBAD fold is already sufficient to bind and provide modest activation."

reach
"Our data show how UCH-L5 activity can be modulated by DEUBAD domains present in RPN13 and INO80G through remarkably large conformational changes."

reach
"Our observations are consistent with the findings that RPN13 competes with NFRKB 1-101 for binding to UCH37 and that NFRKB residues 1-101 activate UCH37 but that the longer full-length and 1-465 constructs inhibit UCH37, as do the 39-156 and 1-117 constructs used in this study."

reach
"Intriguingly, an artificial shorter version of INO80G was found to activate UCH-L5 invitro."
RPN13 affects UCHL5
| 66
RPN13 binds UCHL5.
| 34
| 33

reach
"As with the case of Uch37 binding to Rpn13, Ubp6 is activated by association with Rpn1, and Ubp6 also seems to modify 19S RP structure because its binding delays proteolysis by a mechanism that is independent of its catalytic activity."

reach
"Furthermore, we discovered that IkappaB-alpha, a protein whose proteasomal degradation activates the transcription factor NF-kappaB, is also a substrate for the Rpn13 and UCH37 complex."

reach
"RPN13 presumably interacts with the ubiquitin carboxyl-terminal hydrolase UCH-L5 and UCH37 (## 20, 21 in Fig. 2 A) [34,35]."

reach
"A recent publication has provided further proof of the selective role of Rpn13 in proteasomal function by identifying nitric oxide synthase and IkappaB-alpha as two specific substrates of the Rpn13 and Uch37 complex XREF_BIBR."

reach
"Rpn13 can interact with Uch37 and recruit it to the proteasome via its C-terminal 46 residues (also called the KEKE motif) and activates Uch37."

reach
"The binding of Rpn13 to the KEKE motif of Uch37 likely perturbs the hydrophobic interactions of tetrameric Hc, resulting in disassembly of the oligomer into a monomer via the formation of a stable Rpn13 and Uch37 complex that provides a means to keep Uch37 in the monomeric state."

reach
"While the UCH-L5 and Rpn13 complex has provided a template for PR-DUB catalysis and modelling of some cancer derived mutations XREF_BIBR, XREF_BIBR, XREF_BIBR, many outstanding questions remain unanswered."

reach
"Therefore, we used this Rpn13C for further analysis of the Rpn13 and Uch37 complex."

reach
"While the UCH-L5 and RPN13 complex is equimolar at 1:1, the ligand number in our ITC analysis (between 0.25 and 0.5) suggests that the BAP1 and ASXL1 DEU complex is asymmetric and consists of multiple BAP1 molecules bound to one ASXL1 DEU molecule."

reach
"In vitro binding of Uch37 with Rpn13 was shown to promote the hydrolysis of ubiquitin-7-amido-4-methylcoumarin (Ub-AMC) and Uch37 's reactivity with suicide inhibitors such as ubiquitin vinylsulfone (UbVS) and ubiquitin aldehyde (Ubal)."
UCHL5 binds RPN13 and (ULD) domain. 1 / 1
| 1

reach
"Rpn13 also binds to the C-terminal UCH37 like domain (ULD) domain of the UCH, locking the ULD into a favorable conformation for ubiquitin binding."
RPN13 activates UCHL5.
| 27
RPN13 activates UCHL5. 10 / 25
| 25

reach
"Stimulation of deubiquitylating activity can now be added to this list of proteasomal functions, but neither ATPase nor proteolytic function of the proteasome is required, as Rpn13 alone can activate Uch37."

reach
"In line with previous data, we found that the DEUBAD domain of RPN13 activates UCH-L5 (UR)."

reach
"Therefore, it appears that Rpn13 can transiently activate Uch37 in a " hit-and-run " manner without disrupting its association with INO80."

reach
"We show how the DEUBAD domain in RPN13 activates UCH-L5 by tuning the conformation of structural elements in UCH-L5, and inhibits in INO80G, where it exploits molecular mimicry and UCH-L5 conformational plasticity to prevent ubiquitin docking and catalysis."

reach
"That Uch37 binds and is activated by Rpn13 raises the potential for its selective activity on substrates docked via Rpn13."

reach
"Rpn13, the proteasomal receptor for Uch37 in the proteasome 19S regulatory particle, can activate UCH37 by disrupting dimerization."

reach
"Interestingly, a recent study has shown that the proteasomal subunit RPN13 acts as an accessory protein to enhance the activity of the DUB UCH37 toward K48 containing, branched triubiquitin, and this could provide new ways to further explore the assembly and disassembly of these more complex ubiquitin chain types."
| PMC

reach
"Mechanism of the Rpn13 induced activation of Uch37."

reach
"The DEUBAD Domain of RPN13 Activates UCH-L5 by ULD and CL Positioning."

reach
"In this context, the Rpn13 dependent activation of the UCH37 deubiquitinase is noteworthy (cf. XREF_FIG) XREF_BIBR XREF_BIBR, since it would provide SGTA bound substrates with an opportunity for selective deubiquitination XREF_BIBR."
RPN13 bound to UCHL5 activates UCHL5. 2 / 2
| 2

reach
"Binding of Rpn13 to Uch37 increases the isopeptidase activity of Uch37; therefore it may facilitate the rescue of ubiquitinated substrates from proteolysis XREF_BIBR, XREF_BIBR, XREF_BIBR."

reach
"Rpn13 can interact with Uch37 and recruit it to the proteasome via its C-terminal 46 residues (also called the KEKE motif) and activates Uch37."
RPN13 inhibits UCHL5.
| 3
RPN13 inhibits UCHL5. 3 / 3
| 3

reach
"RPN13 disrupts a UCH37 dimer at high concentration."

reach
"Using a combination of mutagenesis, biochemical, and small-angle X-ray scattering (SAXS) techniques, we demonstrated that Rpn13 activated the auto-inhibited Uch37 by de-oligomerizing it and sequestering it to form a 1:1 stoichiometric complex."

reach
"The deubiquitinating enzyme UCH-L5 can be inhibited and activated by regulatory proteins INO80G and RPN13, respectively."
RPN13 increases the amount of UCHL5.
| 1
Modified RPN13 increases the amount of UCHL5. 1 / 1
| 1

reach
"Loss of Rpn13 caused concurrent loss of Uch37, a deubiquitinating enzyme bound to Rpn13, consistent with our previous work [XREF_BIBR]."
RPN13 deubiquitinates UCHL5.
| 1
RPN13 deubiquitinates UCHL5. 1 / 1
| 1

reach
"RP ubiquitin receptors S5a and Rpn10 and Rpn13 capture substrates by recognizing their covalently attached ubiquitin chains, which are removed and disassembled by three deubiquitinating enzymes Rpn11, Ubp6 and Usp14 and Uch37 and UCHL5."
UCHL5 affects NFRKB
12 | 28 19
UCHL5 binds NFRKB.
12 | 18 19
12 | 18 17

sparser
"The activity of UCH-L5 is enhanced when UCH-L5 combines with proteasome 19S regulatory subunit by Rpn13/Admr1 receptor and inhibited when UCH-L5 interacts with NFRKB."

sparser
"And UCH-L5 also interacts with NFRKB in INO80 complex, but it remains unknown that whether UCH-L5 interacts with other components of INO80 complex or not."

reach
"UCHL5 interacts with and protects NFRKB (also known as INO80G) that is a component of INO80 chromatin remodeling complex from proteasomal degradation by removing K48 linked ubiquitin chain."

sparser
"Importantly, two-hybrid assays detected an interaction between NFRKB and full-length Uch37, but not a truncated Uch37 that retains only the UCH domain."

reach
"However, the effect of DRAIC on the combination of UCHL5 and NFRKB was still unclarified, so we detected this combination in oeDRAIC and shDRAIC cell lines, whose results showed that oeDRAIC can significantly reduce the level of NFRKB coprecipitated by UCHL5, while shDRAIC can increase NFRKB binding with UCHL5, which confirmed the speculation that DRAIC may indirectly down-regulate the expression of NFRKB through affecting the deubiquitination induced by UCHL5."

No evidence text available

sparser
"While the activity of UCH-L5 is inhibited when UCH-L5 interacts with NFRKB, a component of INO80 complex, by combination and deubiquitination [ xref ]."

sparser
"Considering the deubiquitination function of UCHL5, we speculated that DRAIC might regulate the ubiquitination level of NFRKB by influencing the binding of UCHL5 and NFRKB, and then affect the protein expression of NFRKB."

reach
"Interestingly, immunoprecipitation studies from MG132 treated cells revealed that NFRKB interacted with UCHL5 less in the chromatin fraction than in the nucleoplasm, despite its interactions with INO80 being essentially equivalent in these two fractions (XREF_FIG; for input fractions, see XREF_SUPPLEMENTARY)."

No evidence text available
| 2

sparser
"Because hRpn13 and NFRKB interact with Uch37 in the proteasome and hINO80, respectively, we asked whether the isolated proteins bind to Uch37 in a mutually exclusive fashion."

sparser
"Biological information indicated that both RPN13 and INO80G bind to a long fragment at the C-terminal of the UCH-L5."
UCHL5 deubiquitinates NFRKB.
| 5
UCHL5 deubiquitinates NFRKB. 5 / 5
| 5

reach
"LncRNA DRAIC inhibits proliferation and metastasis of gastric cancer cells through interfering with NFRKB deubiquitination mediated by UCHL5."

reach
"Combining the above results, it can be concluded that DRAIC mediates the ubiquitylation degradation of NFRKB by interfering with deubiquitination of NFRKB induced by UCHL5, and then exerts an anti-cancer effect in GC."

reach
"On the other hand, Zhang et al. found that lncRNA DRAIC could suppress GC metastasis of GC cells via through influencing NFRKB de-ubiquitination induced by UCHL5 27.EMT is considered to be a core factor of tumor metastasis, and it is clear that a variety of lncRNAs participated in GC development by regulating this cell program."

reach
"So we further tested the ubiquitination level of NFRKB, and found that the NFRKB ubiquitination level increased significantly after oeDRAIC, which could demonstrate that DRAIC weakens the deubiquitination of NFRKB mediated by UCHL5, and maintains the ubiquitination level of NFRKB and boost the degradation of NFRKB via the ubiquitination-proteasome pathway."

reach
"UCHL5 is also required for BLM/EXO1 dependent end resection through de-ubiquitinating the INO80 subunit NFRKB, although the function of this subunit in regulating end resection is not currently understood [37]."
UCHL5 decreases the amount of NFRKB.
| 2
UCHL5 decreases the amount of NFRKB. 2 / 2
| 2

reach
"UCHL5 depletion did not, however, reduce NFRKB mRNA levels, nor protein levels of other INO80-complex subunits with suggested roles in DSB repair (XREF_SUPPLEMENTARY) XREF_BIBR - XREF_BIBR."

reach
"We found that UCHL5 depletion reduced the steady-state level of NFRKB but not hRPN13 (XREF_FIG and data not shown); and time-course studies in cells treated with cycloheximide, to prevent de novo protein synthesis, revealed that UCHL5 depletion reduced NFRKB protein half-life (XREF_FIG)."
UCHL5 activates NFRKB.
| 2
UCHL5 activates NFRKB. 2 / 2
| 2

reach
"UCHL5 aids resection by protecting NFRKB from proteasomal degradation."

reach
"Further studies showed that DRAIC can promote the ubiquitination degradation of NFRKB mediated by UCHL5, and confirmed the inhibitory effect of DRAIC on proliferation and metastasis of GC cells, which could be rescued by oeNFRKB."
UCHL5 inhibits NFRKB.
| 1
UCHL5 bound to NFRKB inhibits NFRKB. 1 / 1
| 1

reach
"DRAIC combined with UCHL5 and attenuated binding of UCHL5 and NFRKB, meanwhile promoting the degradation of NFRKB via ubiquitination, and then inhibited the proliferation and metastasis of GC cells, which can be rescued by oeNFRKB."
UCHL5 affects Ubiquitin
| 18 27
UCHL5 binds Ubiquitin.
| 1 27
| 1 19

sparser
"The FRF hairpin inserts into the pocket of Uch37 that normally binds ubiquitin and is clasped in place by binding of the long, C-terminal helix of NFRKB DEU to the catalytic domain of Uch37, which fo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"An inspection of 39 DUB-ubiquitin structures in the protein data bank reveals the uniqueness of the salt bridge in ubiquitin bound to UCH37, an interaction that disappears when the ULD is deleted, as revealed in the structure of the catalytic domain alone bound to UbVME."

sparser
"Interestingly, the association of a Ub conjugate with Uch37 can also allosterically activate peptide entry."

sparser
"The FRF hairpin inserts into the pocket of Uch37 that normally binds ubiquitin, and is clasped in place by binding of the long, C-terminal helix of NFRKB DEU to the catalytic domain of Uch37, which forces the second helix in the Uch37 ULD domain to bend [ xref , xref , xref ] ( xref )."

sparser
"For targets 99 and 100 the ubiquitin-propargyl was attached to UCH-L5 using ubiquitin bound to another deubiquitinase UCH37 (PDB ID 4I6N xref ) as a template."

sparser
"The X-ray structure of the Ubiquitin (Ub) bound to UCH-L5 shows a β-sheet hairpin in Ub that contains a crucial hydrophobic patch involved in the interaction with UCH-L5."

sparser
"Undoubtedly, DUB activity is a promising strategy for cancer therapy. xref  VLX1570 targets the Ub‐USP14 or UbUCHL5 conjugates and is a reversible non‐selective competitive inhibitor of ubiquitin peptidase 14 (USP14) and ubiquitin carboxyl‐terminal hydrolase 5 (UCHL5) in multiple myeloma. xref  VLX1570 has been shown to have significant anti‐cancer efficacy with high potency and solubility."

sparser
"We cannot, however, exclude the possibility that assembly of Uch37 into hINO80 affects DUB activity by a different mechanism, since our attempts to measure the binding of ubiquitin to Uch37 within hINO80 have been unsuccessful."

sparser
"To delineate the structural basis of the auto-inhibition of Uch37 and its activation by Rpn13C, we used a Uch37 dimer model derived from its crystal structure and compared it with the structure of the binary complex of ubiquitin-bound Uch37 (PDB ID 4IG7) and its homolog UCH-L3 (PDB ID 1XD3)."

sparser
"However, how ubiquitin binds to UCH37 has not been structurally characterized."
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
| 2

sparser
"Structures of UCH-L5-Rpn13 bound to ubiquitin are available."

sparser
"RPN13 binding to ubiquitin chains and UCH37 might simultaneously facilitate UCH37's distal end xref deubiquitylating activity xref - xref by orienting the ubiquitin moieties in a configuration favorable to hydrolysis."
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
| 1

sparser
"Out of these ten proposed residues S10, E31, Y33, I214, F228, and L230, positions were chosen based on Ts UCHL5ubiquitin interactions."
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
UCHL5 activates Ubiquitin.
| 10
| 10

reach
"RPN11 and POH1 has been demonstrated to cleave at the base of the ubiquitin chain, removing ubiquitin en masse, while UCH37 and USP14 mediate a stepwise removal of ubiquitin from the substrate starting from the distal end."

reach
"RPN11 cleaves at the base of the ubiquitin chain where it is linked to the substrate, whereas UCH37 and apparently USP14 mediate a stepwise removal of ubiquitin from the substrate by disassembling the chain from its distal tip."

reach
"UPS14 and UCH37 are thought to mediate stepwise disassembly of the Ub chain from the distal end 6 ."

reach
"However, as UCHL5 and USP14 are proposed to mediate a stepwise removal of ubiquitin from the substrate by trimming the chain from its distal tip, this leaves the opportunity for MGRN1 to reach a monou[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We, therefore, wanted to determine the type of the ubiquitin linkage targeted on Tcf7 by Uch37."

reach
"It appears that POH1 cleaves at the base of the ubiquitin chain where it is linked to the target protein, whereas USP14 and UCHL5 mediate a stepwise removal of ubiquitin from the protein by disassembling the chain from its distal tip [XREF_BIBR]."

reach
"The deeper mechanism may be that the degradation of β-catenin depends on the UCHL5-mediated ubiquitin–proteasome pathway."

reach
"The western blot results showed that inhibition of both UCHL5 and USP14 by b-AP15 significantly increased the accumulation of polyubiquitinated proteins by 31.80 +/- 6.25% (P = 0.008, n = 5) and reduc[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"RPN11 appears to cleave at the base of the ubiquitin chain, where it is linked to the substrate, whereas USP14 and UCHL5 are known to mediate stepwise removal of ubiquitin from the substrate by disassembling the chain from its distal end [XREF_BIBR]."

reach
"Once bound by the proteasome, deubiquitinating enzymes, like the 19S subunit Rpn11 [XREF_BIBR, XREF_BIBR] or proteasome associated Ubp6 [XREF_BIBR, XREF_BIBR] and Uch37 [XREF_BIBR, XREF_BIBR], remove the ubiquitin moiety to allow recycling of ubiquitin before degradation of the substrate."
UCHL5 inhibits Ubiquitin.
| 7
| 7

reach
"By contrast, USP14 and UCHL5 are located further from the 20S core and antagonize degradation by removing Ub in a stepwise manner from the distal end, promoting substrate dissociation from the proteasome 17.The human genome encodes for approximately 90 DUBs, which fall into six classes 18, 19."

reach
"It has been reported that both USP14 and UCH37 prevent substrate degradation by removing ubiquitin chains and promoting proteasomal substrate dissociation."

reach
"Functional studies suggest that UCH-37 inhibits the degradation of ubiquitinated proteins by the proteasome by removing ubiquitin from the distal end of the polyubiquitin chain."

reach
"By contrast, USP14 and UCHL5 are located further from the 20S core and antagonize degradation by removing Ub in a stepwise manner from the distal end, promoting substrate dissociation from the proteasome [17] ."

reach
"Since Uch37 is expected to disassemble ubiquitin chains at hRpn13 in the proteasome, we hypothesized that RA190 inactivation of Uch37 could impair the disassembly and clearance of ubiquitin chains from the proteasome."

reach
"Mechanistically, loss of Uch37 activity at the proteasome would cause ubiquitin chains to become stalled at hRpn13 in the proteasome (XREF_FIG, right panel)."

reach
"The degradation of Ub itself is closely related to the activities of DUBs associated with the proteasome since the loss of USP14 (or Ubp6 in yeast) or UCH37 has been shown to trigger degradation of Ub along with its target substrates, leading to depletion of the free Ub pool XREF_BIBR - XREF_BIBR."
UCHL5 affects Proteasome
1 | 30 13
UCHL5 binds Proteasome.
1 | 17 13
1 | 16 8

reach
"Binding of UCHL5 to the proteasome repositions a crossover loop, thereby relieving an auto-inhibitory effect and allowing substrate access to the active site 48."

reach
"Uch37 and Usp14 associate reversibly with the proteasome, whereas Rpn11 is a stoichiometric subunit XREF_BIBR."

reach
"Two additional DUBs, USP14 and UCHL5 (also known as and hereafter referred to as UCH37), also associate with the proteasome, although their precise roles are unclear (D'Arcy and Linder, 2012; Komander and Rape, 2012; Lee et al., 2011)."

reach
"Considering that UCH-L5 is associated with the proteasome, where the local concentration of ubiquitin is much higher than the average cellular concentration, UCH-L5 might effectively deubiquitinate its substrates for editing purposes before the substrates are degraded by the proteasome."

reach
"17 Of the three, USP14 and UCHL5 reversibly associate with the proteasome, whereas the third, RPN11 and POH1, is an intrinsic component of the lid subcomplex of the 19S cap."

sparser
"UCHL5 reversibly associates with the 26S proteasome and prevents target proteins degradation by hydrolyzing ubiquitin chains ( xref )."

sparser
"As PSMD1 (scRpn2) encodes for the largest non-ATPase subunit of the 19S regulator lid of the 26S proteasome and this is involved in binding to the deubiquitinase UCH37 (scUCHL5) via the adapter protein ADRM1 (scRpn13), we hypothesized that knockout of PSMD1 may be disrupting the binding of UCH37 to the 26S proteasome, thereby inhibiting its deubiquitinase activity and leading to HIV-1 reactivation xref (Fig.  xref )."

sparser
"UCHL5 is associated with the 26S proteasome, where it serves to remove distal ubiquitin moieties from polyubiquitylated proteins."

"It was initially believed that the presence of DUBs at the proteasome (such as Uch37/UchL5 and Usp14) would promote the degradation of proteins through facilitating substrate processing (see also the next section)"

reach
"Proteasome bound UCH37 is thought to behave as an " editor ", relieving poorly ubiquitinated substrates from degradation by sequentially dismantling their K48 linked polyubiquitin chains from the very distal end, removing one ubiquitin at a time."
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
UCHL5 activates Proteasome.
| 6
| 6

reach
"In addition, the fact that UCH37 activation in the proteasome complex is reversed in the INO80 complex may imply a unique compartmental role or functional partitioning of this enzyme."

reach
"In a more targeted approach, an active-site ubiquitin probe (HA-Ub-VMS) has been used to demonstrate that USP14/UCHL5 inhibition by a small molecule (b-AP15) inhibits the 19S proteasome in a reconstituted biochemical assay."

reach
"The inhibition of UCHL5 and USP14 deubiquitinase activity by WP1130 is expected to block the function of the proteasome in tumor tissue cells, but this still needs to be tested (D'Arcy et al., 2015)."

reach
"The C. elegans homolog of UCHL5, UBH-4, tissue-specifically modulates proteasome activity, also affecting the health and lifespan of these animals [XREF_BIBR]."

reach
"UCHL5 can suppress proteasome degradation through disassembly of distal polyubiquitin moieties (Lam et al., 1997; Schreiner et al., 2008; Koulich et al., 2008; Jacobson et al., 2009)."

reach
"In contrast, siRNA of either UCHL5 or USP14 alone did not affect proteasome composition but did increase the rate of proteasome activity, supporting previous studies."
UCHL5 inhibits Proteasome.
| 4
| 4

reach
"For elderly patients with reduced proteostasis capacity, high UCHL5 levels may further inhibit proteasome activity, thereby promoting apoptosis in cancer cells."

reach
"UCH-L5 can suppress proteasome mediated degradation via the disassembly of distal polyubiquitin moieties."

reach
"In particular, deubiquitinating enzymes Uch37 and Usp14, being physically associated with the proteasome, may suppress proteasome activity through deubiquitinating a subset of ubiquitinated substrates, once the substrate is docked at the proteasome."

reach
"It is possible that excess UCHL5 expression substantially impairs proteasome activity, causing abnormal accumulation of proteasomal substrates, and harming tumor cells."
UCHL5 deubiquitinates Proteasome.
| 2
UCHL5 deubiquitinates Proteasome on S26. 1 / 1
| 1

reach
"Thus Uch37 and Usp14 may both deubiquitinate the 26S proteasome."
UCHL5 deubiquitinates Proteasome. 1 / 1
| 1

reach
"Another proposal is that UCH37 deubiquitylates proteasome subunits that can undergo regulatory ubiquitylation."
UCHL5 ubiquitinates Proteasome.
| 1
UCHL5 leads to the ubiquitination of Proteasome on S26. 1 / 1
| 1

reach
"Thus the DUB activities of Uch37 and/or Usp14 appear to antagonize ubiquitination of the 26S proteasome."
Proteasome affects UCHL5
1 | 27 13
Proteasome binds UCHL5.
1 | 17 13
1 | 16 8

reach
"Binding of UCHL5 to the proteasome repositions a crossover loop, thereby relieving an auto-inhibitory effect and allowing substrate access to the active site 48."

reach
"Uch37 and Usp14 associate reversibly with the proteasome, whereas Rpn11 is a stoichiometric subunit XREF_BIBR."

reach
"Two additional DUBs, USP14 and UCHL5 (also known as and hereafter referred to as UCH37), also associate with the proteasome, although their precise roles are unclear (D'Arcy and Linder, 2012; Komander and Rape, 2012; Lee et al., 2011)."

reach
"Considering that UCH-L5 is associated with the proteasome, where the local concentration of ubiquitin is much higher than the average cellular concentration, UCH-L5 might effectively deubiquitinate its substrates for editing purposes before the substrates are degraded by the proteasome."

reach
"17 Of the three, USP14 and UCHL5 reversibly associate with the proteasome, whereas the third, RPN11 and POH1, is an intrinsic component of the lid subcomplex of the 19S cap."

sparser
"UCHL5 reversibly associates with the 26S proteasome and prevents target proteins degradation by hydrolyzing ubiquitin chains ( xref )."

sparser
"As PSMD1 (scRpn2) encodes for the largest non-ATPase subunit of the 19S regulator lid of the 26S proteasome and this is involved in binding to the deubiquitinase UCH37 (scUCHL5) via the adapter protein ADRM1 (scRpn13), we hypothesized that knockout of PSMD1 may be disrupting the binding of UCH37 to the 26S proteasome, thereby inhibiting its deubiquitinase activity and leading to HIV-1 reactivation xref (Fig.  xref )."

sparser
"UCHL5 is associated with the 26S proteasome, where it serves to remove distal ubiquitin moieties from polyubiquitylated proteins."

"It was initially believed that the presence of DUBs at the proteasome (such as Uch37/UchL5 and Usp14) would promote the degradation of proteins through facilitating substrate processing (see also the next section)"

reach
"Proteasome bound UCH37 is thought to behave as an " editor ", relieving poorly ubiquitinated substrates from degradation by sequentially dismantling their K48 linked polyubiquitin chains from the very distal end, removing one ubiquitin at a time."
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
Proteasome deubiquitinates UCHL5.
| 5
Proteasome deubiquitinates UCHL5. 5 / 5
| 5

reach
"Furthermore, direct interaction of PSMD8 has been reported with the proteasome associated deubiquitinating enzyme UCH37 (Li et al., 2001)."

reach
"Additional targets which play a role in WM cell survival include TLR7, 8 and 9, proteasome associated deubiquitinating enzymes (USP14 and UCHL5), XPO1 and CRM1 and AURKA."

reach
"Furthermore, since deubiquitinating enzymes associated with the proteasome are responsible for ubiquitin trimming from substrates targeted to the proteasome for degradation, and in light of growing evidence that the manner in which proteasome associated deubiquitinating enzymes, USP14 and UCH37, deubiquitinate substrates can in fact suppress and delay degradation and modulate proteasome function, we decided to next analyze the functional activity of the proteasome associated deubiquitinating enzyme USP14."

reach
"B-AP15 simultaneously inhibits the proteasome associated deubiquitinating enzymes UchL5 and Usp14, whereas RA190 binds and inhibits the ubiquitin receptor subunit Rpn13 [XREF_BIBR, XREF_BIBR]."

reach
"Recently, we reported that the proteasome associated deubiquitinating enzyme UCHL5 and Uch37 is a new prognostic marker in both rectal cancer and pancreatic ductal adenocarcinoma."
Proteasome ubiquitinates UCHL5.
| 1
Proteasome ubiquitinates UCHL5. 1 / 3
| 1

reach
"Here we report that the human 26S proteasome is ubiquitinated, by which the ubiquitin receptors Adrm1 and S5a, the ATPase subunit Rpt5, and the deubiquitinating enzyme Uch37 are ubiquitinated in situ by proteasome associating ubiquitination enzymes."
Proteasome activates UCHL5.
| 2
| 2

reach
"However, only recruitment into the proteasome activates Uch37."

reach
"First, does proteasome activated Uch37 affect INO80 mediated nucleosome remodeling activity?"
Proteasome inhibits UCHL5.
| 2
| 2

reach
"HA-UbVS pretreatment of auranofin could bind the HA tagged UbVS in the purified 26S proteasome, supporting that auranofin inhibits UCHL5 and USP14."

reach
"Activity probe assays with either the whole 26S proteasome or the 19S regulatory particle showed that the compound blocked the reaction of both USP14 and UCHL5 with haemagglutinin (HA)-tagged ubiquitin vinyl methyl sulfone (VMS)."
Ubiquitin affects UCHL5
| 4 27
Ubiquitin binds UCHL5.
| 1 27
| 1 19

sparser
"The FRF hairpin inserts into the pocket of Uch37 that normally binds ubiquitin and is clasped in place by binding of the long, C-terminal helix of NFRKB DEU to the catalytic domain of Uch37, which fo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"An inspection of 39 DUB-ubiquitin structures in the protein data bank reveals the uniqueness of the salt bridge in ubiquitin bound to UCH37, an interaction that disappears when the ULD is deleted, as revealed in the structure of the catalytic domain alone bound to UbVME."

sparser
"Interestingly, the association of a Ub conjugate with Uch37 can also allosterically activate peptide entry."

sparser
"The FRF hairpin inserts into the pocket of Uch37 that normally binds ubiquitin, and is clasped in place by binding of the long, C-terminal helix of NFRKB DEU to the catalytic domain of Uch37, which forces the second helix in the Uch37 ULD domain to bend [ xref , xref , xref ] ( xref )."

sparser
"For targets 99 and 100 the ubiquitin-propargyl was attached to UCH-L5 using ubiquitin bound to another deubiquitinase UCH37 (PDB ID 4I6N xref ) as a template."

sparser
"The X-ray structure of the Ubiquitin (Ub) bound to UCH-L5 shows a β-sheet hairpin in Ub that contains a crucial hydrophobic patch involved in the interaction with UCH-L5."

sparser
"Undoubtedly, DUB activity is a promising strategy for cancer therapy. xref  VLX1570 targets the Ub‐USP14 or UbUCHL5 conjugates and is a reversible non‐selective competitive inhibitor of ubiquitin peptidase 14 (USP14) and ubiquitin carboxyl‐terminal hydrolase 5 (UCHL5) in multiple myeloma. xref  VLX1570 has been shown to have significant anti‐cancer efficacy with high potency and solubility."

sparser
"We cannot, however, exclude the possibility that assembly of Uch37 into hINO80 affects DUB activity by a different mechanism, since our attempts to measure the binding of ubiquitin to Uch37 within hINO80 have been unsuccessful."

sparser
"To delineate the structural basis of the auto-inhibition of Uch37 and its activation by Rpn13C, we used a Uch37 dimer model derived from its crystal structure and compared it with the structure of the binary complex of ubiquitin-bound Uch37 (PDB ID 4IG7) and its homolog UCH-L3 (PDB ID 1XD3)."

sparser
"However, how ubiquitin binds to UCH37 has not been structurally characterized."
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
| 2

sparser
"Structures of UCH-L5-Rpn13 bound to ubiquitin are available."

sparser
"RPN13 binding to ubiquitin chains and UCH37 might simultaneously facilitate UCH37's distal end xref deubiquitylating activity xref - xref by orienting the ubiquitin moieties in a configuration favorable to hydrolysis."
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
| 1

sparser
"Out of these ten proposed residues S10, E31, Y33, I214, F228, and L230, positions were chosen based on Ts UCHL5ubiquitin interactions."
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
Ubiquitin activates UCHL5.
| 2
| 2

reach
"Recently it was reported that UCH37 activity is stimulated by branched ubiquitin chain architectures."

reach
"It associates with the 19S RP via the Rpn13 subunit, the proteasomal ubiquitin receptor that activates Uch37 deubiquitylating activity."
Ubiquitin inhibits UCHL5.
| 1
| 1

reach
"Herein, we designed and developed both a Ub sequence-based linear- and cyclic- β-sheet hairpin peptide that was found to preferably inhibit UCH-L5."
UCHL5 affects SMAD2
4 1 | 1 12 2
UCHL5 activates SMAD2.
2 |
UCHL5 activates SMAD2. 2 / 10
2 |

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"
UCHL5 binds SMAD2.
1 | 1 3 2
1 | 1 3 1

trips
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."

reach
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY - motif in Smad7 that interacts with Smurf ubiquitin ligases."

No evidence text available

sparser
"UCH37 also weakly binds to SMAD2 and 3, however it only deubiquitylates ALK5 and hence modifies TGFβ-induced transcription xref ."

reach
"UCH37 binds strongly to Smad7 and weakly to Smad2 and Smad3."

reach
"UCH37 also weakly binds to SMAD2 and 3, however it only deubiquitylates ALK5 and hence modifies TGFbeta induced transcription XREF_BIBR."
| 1

sparser
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."
UCHL5 ubiquitinates SMAD2.
| 1
UCHL5 leads to the ubiquitination of SMAD2. 1 / 7
| 1

reach
"Consistent with the finding from using b-AP15, UCHL5 overexpression decreased poly-ubiquitination of Smad2 and Smad3 (XREF_FIG), while down-regulation of UCHL5 (~ 73%) by UCHL5 shRNA increased poly-ubiquitination of Smad2 and Smad3 (XREF_FIG)."
UCHL5 deubiquitinates SMAD2.
2 | 4
UCHL5 deubiquitinates SMAD2. 6 / 6
2 | 4

reach
"UCH5/UCH37 deubiquitinates both smad2 and smad3 to promote TGFβ-1 induced lung fibrosis [70]."

reach
"In the present study, we demonstrate that UCHL5 de-ubiquitinates and stabilizes Smad2 and Smad3, thereby promoting TGFbeta-1 signaling."

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"

reach
"Here, we demonstrate that UCHL5 de-ubiquitinates and stabilizes Smad2 and Smad3, thereby promoting TGFbeta-1 signaling and contributes to the pathogenesis of pulmonary fibrosis."

reach
"Also, OTUB1 regulates only phosphorylated Smad2 and Smad3 under TGFbeta-1 treatment XREF_BIBR, while UCHL5 de-ubiquitinates Smad2 and Smad3 regardless of TGFbeta-1 treatment."
UCHL5 decreases the amount of SMAD2.
| 2
UCHL5 decreases the amount of SMAD2. 2 / 2
| 2

reach
"UCHL5 shRNA (# 2, # 3, not # 1, # 4) diminished protein levels of Smad2 and Smad3 (XREF_FIG)."

reach
"In the present study, we confirmed that the blockade of UCHL5 activity by the DUB inhibitor bAP15 inhibited expression of phospho-Smad2/Smad3 in a concentration-dependent manner and induced apoptosis in ovarian cancer cells."
UCHL5 increases the amount of SMAD2.
| 1
UCHL5 increases the amount of SMAD2. 1 / 1
| 1

reach
"Inhibition or down-regulation of UCHL5 reduced Smad2 and Smad3 levels and TGFbeta-1-induced the expression of FN and alpha-SMA in human lung fibroblast."
UCHL5 dephosphorylates SMAD2.
| 1
UCHL5 leads to the dephosphorylation of SMAD2. 1 / 1
| 1

reach
"Beyond that, dephosphorylation of Smad2 could also be mediated by UCHL5, which plays a crucial role in suppressing cell apoptosis and sustaining cell survival [XREF_BIBR]."
UCHL5 affects RPN13
| 35
UCHL5 binds RPN13.
| 34
| 33

reach
"As with the case of Uch37 binding to Rpn13, Ubp6 is activated by association with Rpn1, and Ubp6 also seems to modify 19S RP structure because its binding delays proteolysis by a mechanism that is independent of its catalytic activity."

reach
"Furthermore, we discovered that IkappaB-alpha, a protein whose proteasomal degradation activates the transcription factor NF-kappaB, is also a substrate for the Rpn13 and UCH37 complex."

reach
"RPN13 presumably interacts with the ubiquitin carboxyl-terminal hydrolase UCH-L5 and UCH37 (## 20, 21 in Fig. 2 A) [34,35]."

reach
"A recent publication has provided further proof of the selective role of Rpn13 in proteasomal function by identifying nitric oxide synthase and IkappaB-alpha as two specific substrates of the Rpn13 and Uch37 complex XREF_BIBR."

reach
"Rpn13 can interact with Uch37 and recruit it to the proteasome via its C-terminal 46 residues (also called the KEKE motif) and activates Uch37."

reach
"The binding of Rpn13 to the KEKE motif of Uch37 likely perturbs the hydrophobic interactions of tetrameric Hc, resulting in disassembly of the oligomer into a monomer via the formation of a stable Rpn13 and Uch37 complex that provides a means to keep Uch37 in the monomeric state."

reach
"While the UCH-L5 and Rpn13 complex has provided a template for PR-DUB catalysis and modelling of some cancer derived mutations XREF_BIBR, XREF_BIBR, XREF_BIBR, many outstanding questions remain unanswered."

reach
"Therefore, we used this Rpn13C for further analysis of the Rpn13 and Uch37 complex."

reach
"While the UCH-L5 and RPN13 complex is equimolar at 1:1, the ligand number in our ITC analysis (between 0.25 and 0.5) suggests that the BAP1 and ASXL1 DEU complex is asymmetric and consists of multiple BAP1 molecules bound to one ASXL1 DEU molecule."

reach
"In vitro binding of Uch37 with Rpn13 was shown to promote the hydrolysis of ubiquitin-7-amido-4-methylcoumarin (Ub-AMC) and Uch37 's reactivity with suicide inhibitors such as ubiquitin vinylsulfone (UbVS) and ubiquitin aldehyde (Ubal)."
UCHL5 binds RPN13 and (ULD) domain. 1 / 1
| 1

reach
"Rpn13 also binds to the C-terminal UCH37 like domain (ULD) domain of the UCH, locking the ULD into a favorable conformation for ubiquitin binding."
UCHL5 inhibits RPN13.
| 1
UCHL5 inhibits RPN13. 1 / 1
| 1

reach
"Uch37 knockdown in the same cell line led to decreased deubiquitination activity XREF_BIBR, XREF_BIBR and expression of the C-terminal domain of Rpn13, that competes for the binding to Uch37, reduced proteolytic activity XREF_BIBR, XREF_BIBR."
Rpn13C affects UCHL5
| 26 7
Rpn13C binds UCHL5.
| 25 6
UCHL5 binds Rpn13C. 10 / 31
| 25 6

reach
"Furthermore, the atomic model of the Uch37 and Rpn13C complex generated by MASSHA could fit the ab initio low-resolution shape well."

reach
"In sharp contrast, the elution volume of the Uch37 and Rpn13C complex was independent of the concentration."

reach
"Although the overall SAXS based Uch37 and Rpn13C complex model is in agreement with the model reported by Chen et al., the distance between the Uch37 binding surface of Rpn13 and the Uch37 molecule is slightly larger."

reach
"Uch37 bound to Rpn13C tightly with a low dissociation constant (K d = 5.35 nmol/L, Fig."

reach
"The MWs calculated from the SAXS P (r) functions for Uch37 and Uch37 and Rpn13C complex were approximately 117.7 and 64.6 kDa, respectively."

sparser
"A moderate increase in the I 0 implied that Rpn13C interacted with the oligomerized Uch37 [(Uch37)n], forming a larger intermediate (Uch37)n/Rpn13C complex."

sparser
"Interestingly, 4 mol/L NaCl could not reverse the interaction between Uch37 and Rpn13C, indicating a role for hydrophobic interactions in the formation of the complex (Fig.  xref E)."

reach
"Reconstruction of the Uch37 and Rpn13C complex structure by SAXS."

sparser
"To confirm that the interaction of Rpn13C with Uch37 was specific, we performed surface plasmon resonance (SPR) experiments and measured the affinity between Rpn13C and Uch37."

reach
"In this SAXS based Uch37 and Rpn13C complex model, the Uch37 binding surface of Rpn13C reported by Chen et al. was facing toward the Uch37 molecule."
Rpn13C activates UCHL5.
| 1 1
Rpn13C activates UCHL5. 2 / 2
| 1 1

sparser
"To further demonstrate that Rpn13C activated Uch37 through de-oligomerization, we performed Ub-AMC activity assays for Uch37 and its C-terminal region deletion truncations in the presence or absence of Rpn13."

reach
"To further demonstrate that Rpn13C activated Uch37 through de-oligomerization, we performed Ub-AMC activity assays for Uch37 and its C-terminal region deletion truncations in the presence or absence of Rpn13."
INO80 affects UCHL5
4 | 9 17
INO80 binds UCHL5.
4 | 5 17
4 | 5 16

sparser
"Interestingly, UCH37 also associates in the nucleus with the human Ino80 chromatin-remodeling complex, where it is held in an inactive state compared to the free enzyme( xref )."

No evidence text available

sparser
"Uch37 is bound to hINO80 via its C-terminal tail and the N-terminal domain of NFRKB."

No evidence text available

sparser
"Further studies have suggested that UCH37 is associated with the human Ino80 chromatin-remodeling complex (hINO80) in the nucleus and can be activated via transient association of 19S regulatory parti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Our dissection of interactions between hINO80 and Uch37 shows that one polypeptide, NFRKB, has elements that can inhibit and activate deubiquitinating activity."

reach
"In contrast, INO80 associated Uch37 or Uch37 in complex with the NFRKB 1-465 fragment is either inhibited (as assessed by UbVS reactivity) or held in a relatively inactive state similar to that of free Uch37 (as assessed by its ability to hydrolyze UbAMC)."

reach
"Considering that INO80 functions in transcription and DNA repair through chromatin remodeling [65], histones and transcription factors are among likely candidates for substrates of INO80 bound Uch37."

sparser
"The transient association of the INO80 complex with UCH37 regulates the deubiquitylating activity of this subunit."

No evidence text available
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
INO80 activates UCHL5.
| 4
INO80 activates UCHL5. 4 / 4
| 4

reach
"A possibility is that INO80 controls UCH-L5 in a temporal manner, where in some circumstances UCH-L5 is inhibited while under other circumstances post-translational modifications (PTMs) and/or conformational changes release the inhibition and activate UCH-L5, allowing for additional layers of regulation."

reach
"Curiously, INO80 associated UCH37 can be activated by transient incubation with either RPN13 or 26S proteasome lacking UCH37, without UCH37 dissociating from the INO80 complex."

reach
"In the nucleus, UCHL5 is also associated with human Ino80 chromatin remodeling complex and kept in inactive state, and then activated by transient interaction of the Ino80 complex with proteasome, suggesting that it may cooperate to regulate transcription or DNA repair XREF_BIBR."

reach
"Intriguingly, Uch37 is held in an inactive state in the INO80 complex but, at least in vitro, can be activated by transient interaction of the INO80 complex with proteasomes [26]."
UCHL5 affects SMAD3
2 2 | 1 11 5
UCHL5 binds SMAD3.
2 | 1 2 5
2 | 1 2 4

sparser
"As shown in xref , UCHL5 was not associated with phosphorylated Smad3."

reach
"To examine whether TGFbeta-1 treatment enhances the association between phospho-Smad3 and UCHL5, HLF cells were treated with TGFbeta-1 for 30min, and co-IP were performed."

trips
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."

sparser
"As shown in xref , UCHL5 was associated with Smad3, not Smad2."

sparser
"In the study by Wicks and colleagues xref , the authors detected weak association between UCHL5 and Smad2/Smad3, while we show that UCHL5 associates with Smad3 on Thr66, not Smad2."

sparser
"We show that b-AP15 increased Smad2/3 poly-ubiquitination and that UCHL5 is associated with Smad3."

No evidence text available

reach
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY - motif in Smad7 that interacts with Smurf ubiquitin ligases."

No evidence text available
| 1

sparser
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."
UCHL5 deubiquitinates SMAD3.
1 | 5
UCHL5 deubiquitinates SMAD3. 5 / 5
1 | 4

reach
"UCH5/UCH37 deubiquitinates both smad2 and smad3 to promote TGFβ-1 induced lung fibrosis [70]."

reach
"Here, we demonstrate that UCHL5 de-ubiquitinates and stabilizes Smad2 and Smad3, thereby promoting TGFbeta-1 signaling and contributes to the pathogenesis of pulmonary fibrosis."

reach
"In the present study, we demonstrate that UCHL5 de-ubiquitinates and stabilizes Smad2 and Smad3, thereby promoting TGFbeta-1 signaling."

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"

reach
"Also, OTUB1 regulates only phosphorylated Smad2 and Smad3 under TGFbeta-1 treatment XREF_BIBR, while UCHL5 de-ubiquitinates Smad2 and Smad3 regardless of TGFbeta-1 treatment."
Modified UCHL5 leads to the deubiquitination of SMAD3. 1 / 1
| 1

reach
"We demonstrate that Smad2 and Smad3 ubiquitination was diminished by over-expression of UCHL5, while it was enhanced by inhibition or down-regulation of UCHL5."
UCHL5 ubiquitinates SMAD3.
| 1
UCHL5 leads to the ubiquitination of SMAD3. 1 / 5
| 1

reach
"Consistent with the finding from using b-AP15, UCHL5 overexpression decreased poly-ubiquitination of Smad2 and Smad3 (XREF_FIG), while down-regulation of UCHL5 (~ 73%) by UCHL5 shRNA increased poly-ubiquitination of Smad2 and Smad3 (XREF_FIG)."
UCHL5 activates SMAD3.
1 |
UCHL5 activates SMAD3. 1 / 5
1 |

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases"
UCHL5 decreases the amount of SMAD3.
| 2
UCHL5 decreases the amount of SMAD3. 2 / 2
| 2

reach
"In the present study, we confirmed that the blockade of UCHL5 activity by the DUB inhibitor bAP15 inhibited expression of phospho-Smad2/Smad3 in a concentration-dependent manner and induced apoptosis in ovarian cancer cells."

reach
"UCHL5 shRNA (# 2, # 3, not # 1, # 4) diminished protein levels of Smad2 and Smad3 (XREF_FIG)."
UCHL5 increases the amount of SMAD3.
| 1
UCHL5 increases the amount of SMAD3. 1 / 1
| 1

reach
"Inhibition or down-regulation of UCHL5 reduced Smad2 and Smad3 levels and TGFbeta-1-induced the expression of FN and alpha-SMA in human lung fibroblast."
HAUS7 affects UCHL5
3 1 | 9 13
HAUS7 binds UCHL5.
3 1 | 8 13
3 1 | 7 7

reach
"Furthermore, it appears that UCH37 binds UIP1 more strongly compared to its binding of S14 as judged by the beta-galactosidase activity.The transcription of the UIP1 gene in human tissues was studied [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"As the results of experiments using the yeast two-hybrid assay showed that the interaction between UIP1 and UCH37 was much stronger than that between S14 and UCH37, it was therefore of interest to det[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

reach
"This indicates that UCH37 binds UIP1 preferentially and that the binding of S14 by UCH37 may be blocked by UIP1.When using anti-Myc antibody to precipitate the lysates from COS-1 cells co-transfected [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"However, UIP1 did not interact with the N-terminal part of UCH37 (the part of UCH37 that includes the UCH catalytic domain)."

sparser
"The interaction of UCH37 with UIP1 was confirmed by an in vitro binding assay ( Fig. 4 ) and in vivo co-immunoprecipitation analysis ( Fig. 5 )."

sparser
"This finding suggests that the UIP1 could bind UCH37 via its C-terminal extension in vivo."

sparser
"The interaction of UIP1 with UCH37 is stronger than that of UCH37 with S14 as shown by the results of the yeast two-hybrid assay and the in vitro competitive binding assay."
| 6

sparser
"These results indicate that S14 and UIP1 could bind UCH37 specifically in vitro in a C-terminal extension dependent manner."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The Y187 yeast strain harbouring S14/pGAD-GH or UIP1/pGAD-GH was mated with PJ69-2A strain transformed with pU1, pU2 or pU3 respectively to test the interaction between S14 or UIP1 with UCH37, UCH37ΔC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report the identification of two proteins, S14 and UCH37 interacting protein 1 (UIP1), which interact with UCH37."

sparser
"The interaction of UCH37 with S14 or UIP1 was confirmed by in vitro binding assay and in vivo co-immunoprecipitation analysis."
UCHL5 binds HAUS7 and S14. 1 / 1
| 1

reach
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
HAUS7 inhibits UCHL5.
| 1
HAUS7 inhibits UCHL5. 1 / 2
| 1

reach
"UIP1 may modulate the ubiquitinated protein turnover by blocking the association of UCH37 with the 26S proteasome."
UCHL5 affects INO80
4 | 5 17
4 | 5 16

sparser
"Interestingly, UCH37 also associates in the nucleus with the human Ino80 chromatin-remodeling complex, where it is held in an inactive state compared to the free enzyme( xref )."

No evidence text available

sparser
"Uch37 is bound to hINO80 via its C-terminal tail and the N-terminal domain of NFRKB."

No evidence text available

sparser
"Further studies have suggested that UCH37 is associated with the human Ino80 chromatin-remodeling complex (hINO80) in the nucleus and can be activated via transient association of 19S regulatory parti[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Our dissection of interactions between hINO80 and Uch37 shows that one polypeptide, NFRKB, has elements that can inhibit and activate deubiquitinating activity."

reach
"In contrast, INO80 associated Uch37 or Uch37 in complex with the NFRKB 1-465 fragment is either inhibited (as assessed by UbVS reactivity) or held in a relatively inactive state similar to that of free Uch37 (as assessed by its ability to hydrolyze UbAMC)."

reach
"Considering that INO80 functions in transcription and DNA repair through chromatin remodeling [65], histones and transcription factors are among likely candidates for substrates of INO80 bound Uch37."

sparser
"The transient association of the INO80 complex with UCH37 regulates the deubiquitylating activity of this subunit."

No evidence text available
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
UCHL5 affects HAUS7
3 1 | 8 13
3 1 | 7 7

reach
"Furthermore, it appears that UCH37 binds UIP1 more strongly compared to its binding of S14 as judged by the beta-galactosidase activity.The transcription of the UIP1 gene in human tissues was studied [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"As the results of experiments using the yeast two-hybrid assay showed that the interaction between UIP1 and UCH37 was much stronger than that between S14 and UCH37, it was therefore of interest to det[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

reach
"This indicates that UCH37 binds UIP1 preferentially and that the binding of S14 by UCH37 may be blocked by UIP1.When using anti-Myc antibody to precipitate the lysates from COS-1 cells co-transfected [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

No evidence text available

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"However, UIP1 did not interact with the N-terminal part of UCH37 (the part of UCH37 that includes the UCH catalytic domain)."

sparser
"The interaction of UCH37 with UIP1 was confirmed by an in vitro binding assay ( Fig. 4 ) and in vivo co-immunoprecipitation analysis ( Fig. 5 )."

sparser
"This finding suggests that the UIP1 could bind UCH37 via its C-terminal extension in vivo."

sparser
"The interaction of UIP1 with UCH37 is stronger than that of UCH37 with S14 as shown by the results of the yeast two-hybrid assay and the in vitro competitive binding assay."
| 6

sparser
"These results indicate that S14 and UIP1 could bind UCH37 specifically in vitro in a C-terminal extension dependent manner."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The Y187 yeast strain harbouring S14/pGAD-GH or UIP1/pGAD-GH was mated with PJ69-2A strain transformed with pU1, pU2 or pU3 respectively to test the interaction between S14 or UIP1 with UCH37, UCH37ΔC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report the identification of two proteins, S14 and UCH37 interacting protein 1 (UIP1), which interact with UCH37."

sparser
"The interaction of UCH37 with S14 or UIP1 was confirmed by in vitro binding assay and in vivo co-immunoprecipitation analysis."
UCHL5 binds HAUS7 and S14. 1 / 1
| 1

reach
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2 21
| 2 10

reach
"UCHL5 is also overexpressed in hepatocellular carcinoma, and was shown to promote cell migration and invasion 57 ."

reach
"We believe that UCH37 could be a good diagnostic and therapeutic target for controlling HCC recurrence, and further investigations in this field are needed.This study provided evidence for the first t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In vitro, we discovered that UCH37 could promote cell migration and invasion."

reach
"Intriguingly, knock-down of PRP19 reduced the capability of cell migration and invasion, compare with the control cells, suggesting that UCH37 exerted its effects at least in part by deubiquitinating [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"UCH37-knockdown reduced the cell capability to invade extracellular matrix (Huh7-pLKO.1, 23 +/- 3 vs. Huh7-shUCH37, 3 +/- 1, p < 0.05), whereas forced expression of UCH37 increased invasiveness (L02-p[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"UCHL5 is also overexpressed in hepatocellular carcinoma, and was shown to promote cell migration and invasion ."

reach
"A high UCHL5 expression level predicts early recurrence and promotes cell migration and invasion in hepatocellular carcinoma (HCC) XREF_BIBR."

eidos
"A high UCHL5 expression level predicts early recurrence and promotes cell migration and invasion in hepatocellular carcinoma ( HCC ) 12 ."

reach
"Furthermore, we discovered that UCH37 could promote cell migration and invasion in HCC cell lines through interacting and deubiquitinating PRP19, an essential RNA splicing factor.Tumor specimens used [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

eidos
"UCHL5 is also overexpressed in hepatocellular carcinoma , and was shown to promote cell migration and invasion 57 ."
| 11

reach
"Overexpression of UCH-L5 by Lentivirus infection inhibits migration and invasion of human glioma cells."

reach
"In vitro analysis revealed that UCHL5 can inhibit migration and invasion of glioma cells mediated via a downregulation of SNRPF [142]."

reach
"On the other hand, UCH37, the deubiquitinase activated by ADRM1, inhibits glioma cell migration and invasion [42], suggesting that ADRM1 inhibition could have a positive or negative effect on tumor progression, depending on the stage of the tumor.The discovery of a new role for HDAC8 and ADRM1 in MGMT regulation expands the possibilities of the development of new therapies to overcome TMZ resistance in GBM, although several questions about the mechanisms remain to be answered."

reach
"In vitro, we found that UCH-L5 could inhibit migration and invasion of U87MG and U251 cells."

reach
"Furthermore, we discovered that UCH-L5 could inhibit cell migration and invasion of glioma cell lines through downregulating SNRPF, a factor of Sm protein ring in the spliceosome."

reach
"These results suggest that UCH-L5 may inhibit the migration and invasion of glioma cells to inhibit the occurrence of glioma."

reach
"Furthermore, we found that knockdown of UCH-L5 expression promotes the migration and invasion of U87MG and U251 cells."

reach
"And overexpression of UCH-L5 inhibits the migration and invasion of U87MG and U251 cells."

reach
"So UCH-L5 could inhibit glioma cell migration and invasion via downregulating SNRPF."

reach
"Our study showed that UCH-L5 could inhibit migration and invasion of glioma cells via down regulating expression of SNRPF."
UCHL5 affects PSMD8
5 1 | 16
5 1 | 8

sparser
"The finding of the interaction between S14 and UCH37 may provide insights into the subunit arrangement of PA700, in particular the subunits of PA700 that associate with UCH37."

sparser
"This indicates that UCH37 binds UIP1 preferentially and that the binding of S14 by UCH37 may be blocked by UIP1."

No evidence text available

No evidence text available

sparser
"Support for the fact that the two proteins occupy the same binding site came from the observation that UIP1 blocked the interaction between UCH37 and S14 in vitro [ xref ]."

No evidence text available

sparser
"The results of the yeast two-hybrid assay indicates that S14 may interact with UCH37 and that the interaction may be dependent on both the N-terminal UCH domain and the C-terminal extension of UCH37."

No evidence text available

sparser
"This suggests that UIP1 may modulate UCH37’s interaction with S14 in the 26S proteasome."

sparser
"The interaction of UCH37 with S14 was further confirmed by an in vitro binding assay."
| 6

sparser
"These results indicate that S14 and UIP1 could bind UCH37 specifically in vitro in a C-terminal extension dependent manner."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The Y187 yeast strain harbouring S14/pGAD-GH or UIP1/pGAD-GH was mated with PJ69-2A strain transformed with pU1, pU2 or pU3 respectively to test the interaction between S14 or UIP1 with UCH37, UCH37ΔC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report the identification of two proteins, S14 and UCH37 interacting protein 1 (UIP1), which interact with UCH37."

sparser
"The interaction of UCH37 with S14 or UIP1 was confirmed by in vitro binding assay and in vivo co-immunoprecipitation analysis."
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
UCHL5 affects SMAD7
2 2 | 6 8
UCHL5 binds SMAD7.
1 2 | 4 8
1 2 | 4 8

reach
"UCH37 binds strongly to Smad7 and weakly to Smad2 and Smad3."

sparser
"UCH37 also interacts with SMAD7 through the SMAD7 N-terminal domain (1–260 aa), and not via the PY motif, a region that mediates SMAD7’s binding to SMURF (Wicks et al., xref )."

sparser
"Using GST pull down assays, UCH37 was shown to specifically interact with Smad7 in vitro and in vivo via co-immunoprecipitation with HA-tagged UCH37 in transfected HEK-293 cells."

sparser
"However, ubiquitination is reversed by the deubiquitinating enzyme UCH37, which competitively binds Smad7 ( xref )."

reach
"The deubiquitinating enzyme (DUB) UCH37 can bind to Smad7."

sparser
"Endogenous Smad7 and UCH37 formed a stable complex in U4A/JAK1 cells, and FLAG-Smad7 co-immunoprecipitated with HA-UCH37 in transfected HEK-293 cells."

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases."

sparser
"UCH37 is associated with SMAD7 and influences TGF-β-mediated transcription during the early phase of TGF-β receptor activation."

sparser
"We have demonstrated specific interactions between UCH37 and inhibitory Smad7, as well as weaker associations with Smad2 and Smad3."

reach
"Wicks and colleagues reported UCH37 interacts with Smad7 to control TGF-beta and Smad signaling activity, suggesting that UCH37 mediated deubiquitination might contribute to tumorigenesis."
UCHL5 deubiquitinates SMAD7.
| 2
UCHL5 deubiquitinates SMAD7. 2 / 2
| 2

reach
"Type 1 TGF-beta receptor (TGF-betaR1) is degraded by Smad7 dependent ubiquitination-proteasomal pathway, which is deubiquitinated by ubiquitin C-terminal hydrolase-L5 (UCHL5)."

reach
"Type I TGF-beta receptor (TGF-betaR I) is degraded by Smad7 dependent ubiquitination-proteasomal pathway, which is deubiquitinated by ubiquitin C-terminal hydrolase-L5 (UCHL5)."
UCHL5 activates SMAD7.
1 |
UCHL5 activates SMAD7. 1 / 1
1 |

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases."
UCHL5 is modified
| 18 1
UCHL5 is ubiquitinated.
| 18
UCHL5 is ubiquitinated. 10 / 18
| 18

sparser
"In situ ubiquitination of Adrm1, S5a, Rpt5, and Uch37 is accomplished by proteasome-associating UbE1 and E3 Ub ligases together with the UbE2D family of E2s."

sparser
"While ubiquitinated UCHL5 was identified to be more abundant by mass spectrometry in UCHL5 C88A but not USP14 C114A samples ( xref )."

sparser
"Our study also shows that binding of polyUb chains on the 26S proteasome inhibits ubiquitination of Adrm1, S5a, Rpt5, and Uch37 in vitro."

sparser
"Furthermore UCHL5 C88A expression led to a high molecular weight laddering of UCHL5 itself as detected by IB with a UCHL5 antibody, presumably representing ubiquitinated UCHL5 ( xref ) [ xref – xref ]."

sparser
"In support of this, we found that Adrm1 preferred to bind ubiquitinated Uch37 compared with nonubiquitinated Uch37 (Tian and Liu, unpublished result)."

sparser
"We next examined the effect of these polyUb chains on S5a, Adrm1, Uch37, and Rpt5 ubiquitination by supplementing increasing concentrations of these chains into in vitro ubiquitination assays."

sparser
"Ubiquitination of S5a and Uch37 on PA700 was strongly promoted by addition of UbE3A, whereas Rpt5 ubiquitination was only mildly increased ( xref , lane 6)."

sparser
"S5a, Adrm1, Uch37, and Rpt5 are ubiquitinated on the 26S proteasome."

sparser
"While the IP-MS data suggest USP14 is required for deubiquitination of multiple proteasomal subunits ( xref ), USP14 C114A did not affect UCHL5 ubiquitination as shown by the lack of the UCHL5 laddering ( xref )."

sparser
"The robust accumulation of ubiquitin conjugates, ubiquitinated β-catenin, and ubiquitinated UCHL5 were clearly observed in this set of experiments as seen previously (Figs xref – xref )."
UCHL5 is phosphorylated.
| 1
UCHL5 is phosphorylated. 1 / 1
| 1

rlimsp
"To examine whether TGFβ-1 treatment enhances the association between phospho-Smad3 and UCHL5, HLF cells were treated with TGFβ-1 for 30 min, and co-IP were performed."
UCHL5 affects TGFB
| 16
UCHL5 activates TGFB.
| 9
UCHL5 activates TGFB. 9 / 11
| 9

reach
"We show that UCH37 knockdown significantly inhibits the activity of a TGFbeta dependent gene reporter and selectively decreases levels of some TGFbeta dependent target genes, notably p21 and PAI-1, but only during the early phase of TGFbeta receptor activation."

reach
"The up-regulation of TGF-beta signaling by UCH37 could also be partly explained by the deubiquitination and stabilization of Smad3 [67,68]."

reach
"The up-regulation of TGF-beta signaling by UCH37 could also be partly explained by the deubiquitination and stabilization of Smad3 [32,33]."

reach
"In the TGFbeta pathway, UCH37 is capable of deubiquitinating the TGFbeta Type I receptor via a complex that includes Smad7 in mammalian cells and UCH37 activity serves to promote TGFbeta signaling."

reach
"In contrast, UCH37 demonstrated a minimal increase in CAGA-Luc activity suggesting that UCH37 may also target the TGF-beta pathway downstream of the TbetaR complex to regulate overall TGF-beta signaling (XREF_SUPPLEMENTARY)."

reach
"We have shown that UCHL5 promotes TGFbeta signaling via stabilization of Smad2 and Smad3 and may be a potential therapeutic target to block the differentiation of myofibroblasts and the expression of matrix in IPF."

reach
"Finally, AMSH-LP and UCHL5 promote TGF-beta responses through their interaction with inhibitory I-SMADs [XREF_BIBR, XREF_BIBR]."

reach
"The DUBs USP4, USP11, USP15, and UCH37 have previously been demonstrated to modulate TGF-beta pathway activity by directly deubiquitinating the TbetaRI, resulting in increased TbetaRI stability (XREF_FIG) XREF_BIBR XREF_BIBR XREF_BIBR XREF_BIBR."

reach
"In addition, UCH37 stabilizes Smad2/3 and promotes TGF-β1 signal transduction (39)."
UCHL5 deubiquitinates TGFB.
| 4
UCHL5 deubiquitinates TGFB. 3 / 3
| 3

reach
"In addition, we show that UCH37 can deubiquitinate and stabilize the type I TGF-beta receptor."

reach
"The study hypothesized that Smad7 could act as an adaptor to recruit UCH37 to the type I TGF-beta receptor and showed that UCH37 dramatically up-regulates TGF-beta-dependent gene expression by deubiquitinating and stabilizing the type I TGF-beta receptor [XREF_BIBR]."

reach
"Wicks et al. reported that UCH37 can deubiquitinate and stabilize type I TGFbeta receptor and augment TGFbeta signaling [XREF_BIBR] (XREF_FIG)."
UCHL5 deubiquitinates TGFB on R1. 1 / 1
| 1

reach
"UCHL5 can deubiquitinate and stabilize Smads as well as TGF-beta receptor 1 (TGF-beta R1) and therefore activate TGF-beta signaling [XREF_BIBR]."
UCHL5 increases the amount of TGFB.
| 2
UCHL5 increases the amount of TGFB. 2 / 2
| 2

reach
"UCH37 knockdown inhibits transcription of TGF-beta target genes and slows lateral cell migration (Cutts et al., 2011)."

reach
"UCH37 knockdown decreases transcription of TGFbeta dependent target genes and also slows lateral cell migration XREF_BIBR."
UCHL5 binds TGFB.
| 1
| 1

reach
"Smad6 is thought to function preferentially as an inhibitor of BMP, while Smad7 is a general inhibitor of TGFbeta family induced signals (mainly by binding to the TGF-beta type I receptors and preventing phosphorylation of Smads2 and 3, or recruiting ligases that degrade the TGF-beta type I and 2 complexes); however, Smad7 also may enhance TGF-beta, when binding Smad7 to the deubiquitinating enzyme UCH37 [XREF_BIBR])."
B-AP15 affects UCHL5
| 2 15
B-AP15 inhibits UCHL5. 10 / 17
| 2 15

reach
"It is known that b-AP15 and PtPT can inhibit the activity of both USP14 and UCHL5 simultaneously, implying that the downregulation of ERα by b-AP15 and PtPT may attribute to the suppression of USP14 and UCHL5."

reach
"In line with this, Tian et al. reported that b-AP15 selectively blocks deubiquitylating activity of USP14 and UCHL5, leading to the inhibition of cell growth and overcoming bortezomib resistance in MM cells [XREF_BIBR]."

reach
"XREF_BIBR These results have also been further supported by the findings of the Feng et al XREF_BIBR study which have shown that b-AP15 inhibits the deubiquitylating activity of USP14 and UCHL5 enzymes, triggering apoptosis in MM cell lines in a time dependent and dose dependent manner."

reach
"Thus, b-AP15 can inhibit the Ub chain trimming enzymes Uch37 and Usp14 of the 26S proteasome, functions similarly to proteasome inhibitors when used to treat cells, and has potential uses in cancer therapy."

reach
"VLX1570 and b-AP15 both inhibit USP14 and UCHL5 activity of 19S regulatory particles with the inhibition of USP14 being more pronounced (XREF_FIG)."

reach
"Finally, b-AP15 specifically inhibits two proteasome associated DUBs : UCH-L5 and USP14, and thus blocks proteasome function leading to the accumulation of polyubiquitin in treated cells [XREF_BIBR]."

reach
"Inhibition of USP14 and UCH37 deubiquitinating activity by b-AP15 as a potential therapy for tumors with"

reach
"Both studies have clearly shown that IU1 inhibits Usp14, b-AP15 inhibits Usp14 and Uch37, and neither inhibitor abolishes the activity of several other tested DUBs."

reach
"In contrast to inhibitors of the 20S proteasome, b-AP15 blocks the deubiquitylating activity of USP14 and UCHL5, which are associated with the 19S regulatory particle, without affecting proteolytic activities of the 20S proteasome [XREF_BIBR, XREF_BIBR, XREF_BIBR]."

eidos
"B-AP15 inhibits both UCH37 and USP14 , and induces accumulation of ubiquitinated substrates [ 214 ] ."
| 1 15
| 1 12

reach
"Targeted inhibition of the deubiquitinating enzymes, USP14 and UCHL5, induces proteotoxic stress and apoptosis in Waldenstrom macroglobulinaemia tumour cells."

reach
"Moreover, UCHL5 overexpression decreased the percentage of apoptosis cells and sub-G1 population (Figures 4D,E)."

reach
"UCH37 depletion also decreases both cell proliferation and apoptosis induction in functional assays."

reach
"UCHL5 knockdown descended the relative viability of HCE-1-A cells measured by CCK-8 kits (Figure 3C) and augmented the percentage of apoptosis cells (Figure 3D)."

reach
"A novel small molecule inhibitor of deubiquitylating enzyme USP14 and UCHL5 induces apoptosis in multiple myeloma and overcomes bortezomib resistance."

reach
"Furthermore, cyclin-dependent kinase 4/6 (CDK4/6) inhibition decreased UCHL5 expression, suppressed OTX015-R cell proliferation, and induced apoptosis."

reach
"Inhibition of USP14 and UCHL5 activates caspase and triggers apoptosis of ER + BCa cells."

reach
"Along with others, we previously have been reported that inhibition of USP14 and UCHL5 of the 19S proteasome induces apoptosis."

reach
"Recently, novel small molecule inhibitors of the deubiquitylating enzymes USP14 and UCHL5 were developed to overcome bortezomib resistance and induce cell apoptosis of multiple myeloma XREF_BIBR, XREF_BIBR."

reach
"Small molecule inhibitors of UCHL5 could induce apoptosis of multiple myeloma cells and reverse bortezomib resistance [XREF_BIBR]."
| 3

reach
"Other studies have also found that b-AP15 inhibition of USP14 and UCHL5 triggers apoptosis in MM cell lines in a time dependent and dose dependent manner.77 Similar cytotoxic effects, as those seen with b-AP15 treatment, occur within myeloma cells when treated with an alternative DUB inhibitor, copper pyrithione.77 Targeting E1 ubiquitin activating enzyme and E3 ubiquitin ligases has synergistic activity with bortezomib on cell lines.78, 79 An inhibitor of USP7, P5091, can interfere with ubiquitin binding and overcome bortezomib resistance invitro and invivo.80 NIMA related kinase 2 (NEK2) overexpression is associated with resistance to multiple drugs and poor prognosis in MM, and NEK2 inhibitor has been shown to decrease proteasome activity.81 Further investigation into its role in Bortezomib resistance is required."

reach
"Our recent study exemplifies the feasibility of such an approach : specifically, we showed that blockade of 19S associated DUBs USP14 and UCHL5 with a small-molecule inhibitor (bAP15 and VLX1570) induces apoptosis in MM cells and overcome bortezomib resistance, with a favorable toxicity profile."

reach
"b-AP15 is a specific UCHL5 and USP14 inhibitor identified from a screening of small molecules that induce the lysosomal apoptosis pathway."
SMAD7 affects UCHL5
1 2 | 4 8
1 2 | 4 8

reach
"UCH37 binds strongly to Smad7 and weakly to Smad2 and Smad3."

sparser
"UCH37 also interacts with SMAD7 through the SMAD7 N-terminal domain (1–260 aa), and not via the PY motif, a region that mediates SMAD7’s binding to SMURF (Wicks et al., xref )."

sparser
"Using GST pull down assays, UCH37 was shown to specifically interact with Smad7 in vitro and in vivo via co-immunoprecipitation with HA-tagged UCH37 in transfected HEK-293 cells."

sparser
"However, ubiquitination is reversed by the deubiquitinating enzyme UCH37, which competitively binds Smad7 ( xref )."

reach
"The deubiquitinating enzyme (DUB) UCH37 can bind to Smad7."

sparser
"Endogenous Smad7 and UCH37 formed a stable complex in U4A/JAK1 cells, and FLAG-Smad7 co-immunoprecipitated with HA-UCH37 in transfected HEK-293 cells."

"Here, we report a novel interaction between smads and ubiquitin c-terminal hydrolase uch37, a deubiquitinating enzyme that could potentially reverse smurf-mediated ubiquitination. In gst pull down experiments, uch37 bound weakly to smad2 and smad3, and bound very strongly to smad7 in a region that is distinct from the -py- motif in smad7 that interacts with smurf ubiquitin ligases."

sparser
"UCH37 is associated with SMAD7 and influences TGF-β-mediated transcription during the early phase of TGF-β receptor activation."

sparser
"We have demonstrated specific interactions between UCH37 and inhibitory Smad7, as well as weaker associations with Smad2 and Smad3."

reach
"Wicks and colleagues reported UCH37 interacts with Smad7 to control TGF-beta and Smad signaling activity, suggesting that UCH37 mediated deubiquitination might contribute to tumorigenesis."
UCHL5 affects NLRP3
| 10 5
UCHL5 binds NLRP3.
| 3 5
| 3 4

sparser
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."

reach
"Mechanistically, NLRP3 directly bound to UCHL5 and maintained its stability through reducing ubiquitin-proteasome pathway degradation in mandibular MSCs."

reach
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation (XREF_FIG), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 (XREF_FIG)."

sparser
"Mechanistically, NLRP3 directly bound to UCHL5 and maintained its stability through reducing ubiquitin-proteasome pathway degradation in mandibular MSCs."

sparser
"S3D), suggesting that RACK1 could serve as a scaffold protein or a bridge between EST12 and UCHL5-NLRP3."

sparser
"However, UCHL5 did not interact with NLRP3 or with EST12 in RACK1-deficient macrophages upon EST12 stimulation (fig."

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
| 1

sparser
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation ( xref ), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 ( xref )."
UCHL5 deubiquitinates NLRP3.
| 6
UCHL5 deubiquitinates NLRP3. 5 / 5
| 5

reach
"Moreover, UCHL5 overexpression enhanced protein stability by deubiquitinating NLRP3."

reach
"At last, UCHL5 inhibition enhanced osteoblast differentiation by promoting NLRP3 ubiquitination and degradation."

reach
"Deubiquitination of NLRP3 by UCHL5 is required for inflammasome activation."

reach
"Rv1579c, secreted from MTB H37Rv RD3, interacts with the receptor for activated C kinase 1 (RACK1) via its amino acid Y80 at the C-terminus, then recruits ubiquitin C-terminal hydrolase L5 (UCHL5) to deubiquitinate NLRP3, and finally activate the NLRP3 inflammasome [XREF_BIBR]."

reach
"Deubiquitination and Activation of the NLRP3 Inflammasome by UCHL5 in HCV-Infected Cells."
UCHL5 leads to the deubiquitination of NLRP3 on K48. 1 / 1
| 1

reach
"S3E), suggesting that UCHL5 mediates K48 deubiquitination of NLRP3 after EST12 stimulation."
UCHL5 ubiquitinates NLRP3.
| 1
UCHL5 leads to the ubiquitination of NLRP3 on K48. 1 / 1
| 1

reach
"Knockdown of UCHL5 by its specific short hairpin RNA (shRNA) markedly decreased the K48 deubiquitination of NLRP3 after EST12 stimulation in RAW264.7 cells for 6 hours (XREF_FIG and fig."

reach
"UCHL5 knockdown markedly inhibited cell proliferation via regulating cell cycle proteins."

reach
"UCHL5 knockdown by two siRNA segments significantly inhibited cell proliferation in Hela cells."

eidos
"We also found that UCHL5 downregulation significantly suppressed both tumor growth in vivo and cell proliferation and migration in vitro ."

reach
"Deletion of UCH37 also down-regulated the transcription of c-Myc, a downstream effector of beta-catenin, and inhibited cell proliferation and motility."

eidos
"UCHL5 knockdown markedly inhibited cell proliferation via regulating cell cycle proteins ."

reach
"UCHL5 knockdown in LUAD cells significantly inhibited cell proliferation and reduced the expression of key cell cycle proteins."

reach
"UCHL5 knockdown inhibits LUAD cell proliferation via regulation of cell cycle proteins."

reach
"UCHL5 Promotes Proliferation and Migration of Bladder Cancer Cells by Activating c-Myc via AKT/mTOR Signaling."

reach
"Proteasomal deubiquitinase UCH37 inhibits degradation of beta-catenin and promotes cell proliferation and motility."

eidos
"UCH37 interacts with COPS5 to induce the ubiquitination and degradation of p53 and p21 , promoting cell proliferation ."

reach
"Furthermore, cyclin-dependent kinase 4/6 (CDK4/6) inhibition decreased UCHL5 expression, suppressed OTX015-R cell proliferation, and induced apoptosis."

reach
"We also found that UCHL5 downregulation significantly suppressed both tumor growth in vivo and cell proliferation and migration in vitro."

reach
"UCH37 depletion also decreases both cell proliferation and apoptosis induction in functional assays."
| 2 11
| 2 11

reach
"UCHL5 is also overexpressed in hepatocellular carcinoma, and was shown to promote cell migration and invasion 57 ."

eidos
"A high UCHL5 expression level predicts early recurrence and promotes cell migration and invasion in hepatocellular carcinoma ( HCC ) 12 ."

reach
"In vitro, we discovered that UCH37 could promote cell migration and invasion."

reach
"Furthermore, we discovered that UCH37 could promote cell migration and invasion in HCC cell lines through interacting and deubiquitinating PRP19, an essential RNA splicing factor.Tumor specimens used [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Fang Y et al. found UCH-L5 promotes cell migration and invasion via interacting and deubiquitinating splicing factor PRP19 in hepatocellular carcinoma [XREF_BIBR]."

eidos
"UCHL5 is also overexpressed in hepatocellular carcinoma , and was shown to promote cell migration and invasion57 ."

reach
"Collectively, this study demonstrated that UCH37 could promote cell migration and invasion in HCC cell lines through interacting and deubiquitinating PRP19, and suggested that UCH37 could be a novel predictor for HCC recurrence after curative resection."

reach
"Yet UCH37 knockdown significantly impairs cell migration in pancreatic cancer cell lines through the abolition of the TGF-beta-induced expression of MMP-2 and PAI-1, which are thought to play a key ro[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"All of the evidences suggest that the up-regulation of UCH37 may play an important role in oncogenesis through promoting some proto-oncogenes ' expression and stem cell like characteristics in the cel[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"A high UCHL5 expression level predicts early recurrence and promotes cell migration and invasion in hepatocellular carcinoma (HCC) XREF_BIBR."
UCHL5 affects TCF7
| 12 1
UCHL5 binds TCF7.
| 4 1
| 4 1

sparser
"UCH37 can specifically bind and deubiquitinate transcription factor 7 (Tcf7) to activate Wnt signaling in human liver cancer cells [ xref ]."

reach
"Having demonstrated that Uch37 is required for Wnt signalling and specifically interacts with Tcf7, we next hypothesized that Uch37 binds Tcf7 to serve as a bona fide DUB for Tcf7 protein during Wnt signalling."

reach
"Our Co-IP and GST pulldown analyses clearly demonstrated that Uch37 binds Tcf7 directly, and that the physical interaction requires the C-terminal region of Tcf7 protein that includes HMG domain and C-terminal extension."

reach
"Moreover, GST-pull down assay using purified recombinant Uch37 (His Uch37) and Tcf7 (GST-Tcf7) proteins demonstrated a direct interaction between Uch37 and Tcf7 (XREF_FIG)."

reach
"Uch37 specifically and directly interacts with Tcf7."
UCHL5 deubiquitinates TCF7.
| 4
UCHL5 deubiquitinates TCF7. 4 / 4
| 4

reach
"Because Uch37 mediates deubiquitination of Tcf7 protein without affecting its stability, we next sought to elucidate the non proteolytic role of Uch37 mediated deubiquitination of Tcf7 and subsequent activation of Wnt signalling."

reach
"Here, we report that Uch37 mediates the deubiquitination of Tcf7 without affecting protein stability."

reach
"To further examine if Uch37 directly deubiquitinates Tcf7, we conducted in vitro deubiquitination assays using immunopurified Tcf7-ubiquitin conjugates."

reach
"However, deubiquitination of TCF7 by UCH37 is involved in TCF7 binding to the promoters of c-Myc, and Cyclin D1 [XREF_BIBR, XREF_BIBR]."
UCHL5 inhibits TCF7.
| 2
UCHL5 inhibits TCF7. 2 / 2
| 2

reach
"The increased expression of UCH37 decreases the polyubiquitin of TCF7."

reach
"To test this hypothesis, we first depleted endogenous Uch37 to suppress Tcf7 mediated activation of target genes (XREF_FIG) and Tcf7 mediated reporter activity (XREF_FIG)."
UCHL5 activates TCF7.
| 2
UCHL5 activates TCF7. 2 / 2
| 2

reach
"Fractionation and immunostaining analyses in Xenopus gastrula showed that Uch37 co-localized with Tcf7 in the nucleus where Tcf7 protein exclusively resides despite both cytoplasmic and nuclear localization of Uch37 protein (XREF_SUPPLEMENTARY), raising the possibility that the nuclear pool of Uch37 regulates Tcf7 to activate Wnt signalling."

reach
"Importantly, we observed that knockdown of Uch37 by Uch37 MO inhibited binding of Tcf7 to target gene promoters (XREF_FIG lane 3)."
UCHL5 affects SMO
1 | 7 5
UCHL5 binds SMO.
| 3 5
| 3 5

reach
"Strikingly, we find that Hh enhances the interaction between UCHL5 and Smo, thereby stabilizing Smo."

sparser
"The deubiquitinating enzymes UBPY/Usp8 and Uchl5 interact with the C-tail of Smo and inhibits its ubiquitination [ xref – xref ]."

sparser
"Strikingly, we find that Hh enhances the interaction between UCHL5 and Smo, thereby stabilizing Smo."

sparser
"UCHL5 interacts with Smo to promote its deubiquitination and accumulation at the cell membrane, and the interaction between UCHL5 and Smo is enhanced by Hh [ xref ]."
| PMC

reach
"UCHL5 interacts with Smo to promote its deubiquitination and accumulation at the cell membrane, and the interaction between UCHL5 and Smo is enhanced by Hh [44]."
| PMC

sparser
"For example, UCH37’s binding to Smoothened (Smo) counteracts Smo ubiquitination and leads to the stabilization of the Smo protein and the activation of the Hedgehog signaling pathway [ xref ]."

sparser
"Interestingly, Hh enchances the interaction between Uchl5 and Smo [ xref ]."

reach
"We provide both genetic and biochemical evidence that UCHL5 interacts with and deubiquitinates Smo, increasing stability and promoting accumulation at the cell membrane."
UCHL5 deubiquitinates SMO.
1 | 2
UCHL5 deubiquitinates SMO. 3 / 3
1 | 2

reach
"UCHL5 could inhibit SMO ubiquitination and suppress PC-12 cell apoptosis during OGD/R."

reach
"We provide both genetic and biochemical evidence that UCHL5 interacts with and deubiquitinates Smo, increasing stability and promoting accumulation at the cell membrane."

"The deubiquitinase UCHL5/UCH37 positively regulates Hedgehog signaling by deubiquitinating Smoothened"
UCHL5 increases the amount of SMO.
| 1
UCHL5 increases the amount of SMO. 1 / 1
| 1

reach
"Furthermore, knockdown of UCH37 in NIH3T3 cells reduced human Smo level and Shh pathway activity, suggesting that UCHL5/UCH37 plays a conserved role in modulating Smo ubiquitination and turnover [44]."
| PMC
UCHL5 activates SMO.
| 1
UCHL5 activates SMO. 1 / 1
| 1

reach
"The human homologue UCH37 was reported to promote Smo localization to cilia."
UCHL5 affects PSMD14
8 | 5
8 | 1

No evidence text available

No evidence text available

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No evidence text available

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No evidence text available

No evidence text available

sparser
"USP14 is one of three human DUBs, along with UCH37 and RPN11, which associate with the 19S regulatory particle of the multi‐subunit 26S proteasome [103]."
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
WP1130 affects UCHL5
| 1 11
WP1130 inhibits UCHL5. 10 / 12
| 1 11

reach
"WP1130 acts as a partly selective deubiquitinase inhibitor, directly inhibiting deubiquitinase activity of USP9X, USP5, USP14, and UCH37, which are known to regulate survival protein stability and 26S proteasome function."

reach
"It was found that WP1130 can directly inhibit USP9X as well as DUBs of UCHL5, and USP14."

reach
"These data and the fact that UCH37 or USP14 may have a redundant DUB function and also that b-AP15 and WP1130 inhibit both UCH37 and USP14 suggests that a contribution from both enzymes may be needed."

reach
"For example, a general DUB inhibitor WP1130 has been known to inhibit the activities of USP9x, USP5, USP14 and UCH37 that regulate the survival proteins stability [XREF_BIBR]."

reach
"b-AP15 is a proteasomal DUBs inhibitor that blocks USP14 and UCHL5, which are also inhibited by WP1130."

reach
"WP1130 inhibits USP9X, USP5, USP14, UCH37, UCH-L1 in lymphoma cells, and potentially other DUBs as well XREF_BIBR."

reach
"WP1130 (Degrasyn) has been shown to inhibit USP14 and UCHL5 as well as other DUB enzymes (see below)."

reach
"WP1130, an inhibitor of DUBs, can suppress the activities of USP9X, USP5, USP14 and UCH37, deregulate anti-apoptotic protein MCL-1 and upregulate pro apoptotic protein p53."

eidos
"For example , 8-mercapto-N - ( ( tetrahydro-3-furanyl ) methyl ) -4 - quinoline carboxamide , LND-57444 , VLX1570 , ML323 , ( ADC-01 , ADC-03 , HBX41108 , HBX19818 , P5091 , P22077 ) , 9 - ( ethoxyimino ) -9 H-indeno ( 1,2 - b ) pyrazine-2 ,3 - dicarbonitrile , WP1130 , Mitoxantrone and GSK2643943A are able to inhibit PSMD14 , UCHL1 , UCHL5 and USP14 , USP1 , USP7 , USP8 , USP9X , USP11 and USP20 , respectively ."

reach
"WP1130, on the other hand, inhibits at least five DUBs : USP5, UCH-L1, USP9X, USP14, and UCH37, and induces the cellular accumulation of polyubiquitinated proteins [XREF_BIBR]."
UCHL5 affects TGFB1
| 5
UCHL5 activates TGFB1. 5 / 11
| 5

reach
"All these data show that UCHL5 stabilizes Smad2 and Smad3, and promotes TGFbeta-1 signaling."

reach
"b-AP15, an inhibitor of UCHL5, attenuates TGFbeta-1 signaling and diminishes pulmonary fibrosis in a bleomycin induced pulmonary fibrosis model."

reach
"UCHL5 stabilizes Smad2 and Smad3 and promotes TGFbeta-1 signaling."

reach
"In addition, UCH37 stabilizes Smad2/3 and promotes TGF-b1 signal transduction (39) ."

reach
"In this study, we reveal that b-AP15, an inhibitor of UCHL5, attenuates TGFbeta-1 signaling through inducing ubiquitination and degradation of Smad2 and Smad3."
UCHL5 affects proteolysis
| 10
UCHL5 inhibits proteolysis.
| 7
| 7

reach
"Importantly, whereas knockdown of POH1 interferes with the proteasome assembly, depletion of either USP14 or UCH37 alone does not affect or even slightly enhances protein degradation rates."

reach
"The RNAi of UCHL5 or USP14 alone does not affect cell growth and proteasome composition but accelerates cellular protein degradation; however, RNAi of both UCHL5 and USP14 can inhibit cellular protein degradation."

reach
"In contrast, RNAi of Uch37 or Usp14 had no detectable effect on cell growth, proteasome structure or proteolytic capacity, but accelerated cellular protein degradation."

reach
"It has been shown that UCHL5 controls proteasome function and inhibition of UCHL5 promotes protein degradation in autophagy XREF_BIBR."

reach
"Interestingly, b-AP15, a dual inhibitor of Usp14 and Uch37, was shown to prevent protein degradation and inhibit tumor progression in acute myeloid leukemia models [222]."

reach
"In contrast, RNAi of either UCHL5 or USP14 alone did not affect cell growth, proteasome structure, or proteolytic capacity, but increased the rate of protein degradation."

reach
"RNAi of either Uch37 or USP14 (the mammalian homologue of yeast Ubp6) accelerates protein degradation."
UCHL5 activates proteolysis.
| 3
| 3

reach
"RNAi of both Uch37 and Usp14 also had no effect on proteasome structure or proteolytic capacity, but inhibited cellular protein degradation."

reach
"The RNAi of UCHL5 or USP14 alone does not affect cell growth and proteasome composition but accelerates cellular protein degradation; however, RNAi of both UCHL5 and USP14 can inhibit cellular protein degradation."

reach
"It has been shown that UCHL5 controls proteasome function and inhibition of UCHL5 promotes protein degradation in autophagy XREF_BIBR."
SMAD3 affects UCHL5
2 | 1 2 5
2 | 1 2 4

sparser
"As shown in xref , UCHL5 was not associated with phosphorylated Smad3."

reach
"To examine whether TGFbeta-1 treatment enhances the association between phospho-Smad3 and UCHL5, HLF cells were treated with TGFbeta-1 for 30min, and co-IP were performed."

trips
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."

sparser
"As shown in xref , UCHL5 was associated with Smad3, not Smad2."

sparser
"In the study by Wicks and colleagues xref , the authors detected weak association between UCHL5 and Smad2/Smad3, while we show that UCHL5 associates with Smad3 on Thr66, not Smad2."

sparser
"We show that b-AP15 increased Smad2/3 poly-ubiquitination and that UCHL5 is associated with Smad3."

No evidence text available

reach
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY - motif in Smad7 that interacts with Smurf ubiquitin ligases."

No evidence text available
| 1

sparser
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."
RA190 affects UCHL5
| 8 2
RA190 inhibits UCHL5.
| 5
RA190 inhibits UCHL5. 5 / 5
| 5

reach
"Since Uch37 is expected to disassemble ubiquitin chains at hRpn13 in the proteasome, we hypothesized that RA190 inactivation of Uch37 could impair the disassembly and clearance of ubiquitin chains from the proteasome."

reach
"We propose that RA190 deactivation of Uch37 at the proteasome contributes to induction of apoptosis."

reach
"These data suggest that RA190 does not block UCH37 DUB activity in cellular microenvironment."

reach
"These findings indicate that RA190 inhibits Uch37 activity and, moreover, that it has a direct effect that is independent of hRpn13."

reach
"RA190 does not affect hRpn13 interaction with Uch37, but rather directly binds and inactivates Uch37."
RA190 binds UCHL5.
| 3 2
UCHL5 binds RA190. 5 / 5
| 3 2

reach
"We also report anti-cancer molecule RA190, which binds covalently to hRpn13 and UCHL5, to require hRpn13 Pru and not UCHL5 for cytotoxicity."

reach
"RA190 binds Uch37 and inhibits its catalytic activity."

sparser
"Thus, there are no compelling data from the isoTOP-ABPP experiment to indicate RA190 binding to Uch37 in cellulo ."

reach
"Biophysical analyses in combination with cell based assays indicate that RA190 directly binds and inactivates Uch37 [XREF_BIBR]."

sparser
"We also scrutinized the dataset for evidence of RA190 binding to the Rpn13-associated deubiquitylase Uch37 as well."
PSMD4 affects UCHL5
6 | 2 2
PSMD4 binds UCHL5.
6 | 1
6 | 1

No evidence text available

No evidence text available

sparser
"Biochemical data from fission yeast and mammalian cells suggested that the UCH37 also binds to the S5a, a mammalian homologue of Rpn10 [ xref , xref ]."

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PSMD4 inhibits UCHL5.
| 2 1
PSMD4 inhibits UCHL5. 3 / 3
| 2 1

reach
"Interestingly, we found that different from bortezomib, AF inhibits 19S proteasome associated DUBs UCHL5 and USP14 but not the 20S proteasome activity."

sparser
"Interestingly, we found that different from bortezomib, AF inhibits 19S proteasome-associated DUBs UCHL5 and USP14 but not the 20S proteasome activity."

reach
"This was also confirmed by computational molecular docking, K48 linked polyubiquitin disassembly, and HA-UbVS competitive binding assay; these experiments showed that AF might inhibit the proteasomal cysteine DUBs UCHL5 and USP14."
UCHL5 affects PSMD1
6 1 | 2
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
1 | 1

sparser
"We tested this possibility through pharmacological inhibition of the ADRM1-PSMD1-UCH37 interaction by using RA190 xref ."

No evidence text available
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
UCHL5 affects CTNNB1
| 1 8
UCHL5 increases the amount of CTNNB1.
| 4
UCHL5 increases the amount of CTNNB1. 4 / 4
| 4

reach
"Meanwhile, deletion of UCH37 decreased the levels of beta-catenin and the early endosomal protein Rab8."

reach
"In contrast, UCHL5 overexpression in AN3-CA cells elevated the expression of β-catenin and its effector proteins (CyclinD1, C-myc, Survivin) and decreased cleaved-caspase3 (Figures 5D–F)."

reach
"As shown in Figure 5A, UCHL5 knockdown decreased the expression of β-catenin."

reach
"In the study, UCHL5 overexpression elevated the levels of β-catenin and regulated its downstream genes (CyclinD1, C-myc, Survivin, and cleaved-caspase3), which were counteracted by the Wnt/β-catenin inhibitor XAV939."
UCHL5 activates CTNNB1.
| 1 2
UCHL5 activates CTNNB1. 3 / 3
| 1 2

reach
"Proteasomal deubiquitinase UCH37 inhibits degradation of beta-catenin and promotes cell proliferation and motility."

reach
"UCHL5 Activated Wnt/beta-Catenin Signaling and Affected the Expression of Its Target Genes."

eidos
"As shown in Figure 5A , UCHL5 knockdown decreased the expression of beta-catenin ."
UCHL5 decreases the amount of CTNNB1.
| 2
UCHL5 decreases the amount of ubiquitinated CTNNB1. 2 / 2
| 2

reach
"Previous researches documented that deletion of UCHL5 increased the level of the ubiquitinated β-catenin and accelerated the hydrogen peroxide–stimulated degradation of β-catenin in HeLa cells (36)."

reach
"We further found that deletion of UCH37 increased the levels of the ubiquitinated beta-catenin and accelerated the hydrogen peroxide stimulated degradation of beta-catenin."
PSMD1 affects UCHL5
6 1 | 2
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
1 | 1

sparser
"We tested this possibility through pharmacological inhibition of the ADRM1-PSMD1-UCH37 interaction by using RA190 xref ."

No evidence text available
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
UCHL5 affects Wnt
| 2 6
UCHL5 activates Wnt.
| 2 4
UCHL5 activates Wnt. 6 / 6
| 2 4

reach
"Moreover, Wnt reporter activity in Xenopus embryos was completely rescued by Uch37 WT, but not by Uch37 IN mRNA (XREF_FIG)."

eidos
"UCH37 activates Wnt signaling and affects the expression of its target genes , such as beta-catenin , cyclin D1 and c-Myc , thereby increasing the cell growth of EC ."

reach
"Next, either wild type Uch37 (WT) or catalytically inactive Uch37 (IN) XREF_BIBR was reintroduced to determine if Uch37 could rescue Wnt signalling."

reach
"Taken together, our results clearly demonstrate that Uch37 positively regulates Wnt signalling downstream of beta-catenin stabilization."

eidos
"UCH37 activates Wnt signaling and affects the expression of its target genes , such as beta-catenin , cyclin D1 and c-Myc , thereby increasing the cell growth of EC ( Liu et al. 2020 ) ."

reach
"UCHL5 Activated Wnt/beta-Catenin Signaling and Affected the Expression of Its Target Genes."
UCHL5 inhibits Wnt.
| 1
UCHL5 inhibits Wnt. 1 / 1
| 1

reach
"We also observed that downregulation of Uch37 inhibited ectopic Wnt activity that was induced by constitutively active LRP6 (LRP6DeltaN) and the GSK3beta inhibitor LiCl (XREF_SUPPLEMENTARY, g)."
UCHL5 increases the amount of Wnt.
| 1
UCHL5 increases the amount of Wnt. 1 / 1
| 1

reach
"We hypothesized that deubiquitinating activity of Uch37 is involved in the ability of Tcf7 to activate transcription of Wnt target genes."
UCHL5 affects GST
| 8
| 6

sparser
"However, FLAG-S14 (even at the 6×His-UIP1:FLAG-S14 molar ratio of 1:6) had no significant effect on the amount of 6×His-UIP1 bound to GST-UCH37 (lane 6 of Fig. 4B )."

sparser
"Only the fragment containing the N-terminal 101 residues of NFRKB (N101) bound selectively to GST-Uch37 ( Figure 2 B)."

sparser
"Almost no detectable FLAG-S14 was found bound to GST-UCH37 when the molar ratio of 6×His-UIP1:FLAG-S14 was 1:1 (lane 6 of Fig. 4A )."

sparser
"Only the fragment containing the N-terminal 101 residues of NFRKB (N101) bound selectively to GST-Uch37 ( xref )."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To define in more detail the interaction between Uch37 and NFRKB, we generated a series of fragments from the N-terminal third of NFRKB by in vitro translation using a bacterial cell-free system and t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
UCHL5 affects E2F1
4 1 | 3
UCHL5 deubiquitinates E2F1.
1 | 3
UCHL5 deubiquitinates E2F1. 4 / 4
1 | 3

"Here we identify UCH37 as the first, to our knowledge, deubiquitinating enzyme for E2F1."

reach
"In addition, Uch37 deubiquitinates E2 promoter binding factor 1 (E2F1), promoting transcription of pro apoptotic and cell cycle related genes XREF_BIBR."

reach
"Instead, UCH37, but not a catalytically dead mutant, decreases the Lys-63-linked ubiquitination of E2F1 and activates its transcriptional activity."

reach
"UCHL5 activates the E2F1 transcriptional activity by decreasing Lys-63-linked ubiquitination of E2F1."
UCHL5 binds E2F1.
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
SMO affects UCHL5
| 3 5
| 3 5

reach
"Strikingly, we find that Hh enhances the interaction between UCHL5 and Smo, thereby stabilizing Smo."

sparser
"The deubiquitinating enzymes UBPY/Usp8 and Uchl5 interact with the C-tail of Smo and inhibits its ubiquitination [ xref – xref ]."

sparser
"Strikingly, we find that Hh enhances the interaction between UCHL5 and Smo, thereby stabilizing Smo."

sparser
"UCHL5 interacts with Smo to promote its deubiquitination and accumulation at the cell membrane, and the interaction between UCHL5 and Smo is enhanced by Hh [ xref ]."
| PMC

reach
"UCHL5 interacts with Smo to promote its deubiquitination and accumulation at the cell membrane, and the interaction between UCHL5 and Smo is enhanced by Hh [44]."
| PMC

sparser
"For example, UCH37’s binding to Smoothened (Smo) counteracts Smo ubiquitination and leads to the stabilization of the Smo protein and the activation of the Hedgehog signaling pathway [ xref ]."

sparser
"Interestingly, Hh enchances the interaction between Uchl5 and Smo [ xref ]."

reach
"We provide both genetic and biochemical evidence that UCHL5 interacts with and deubiquitinates Smo, increasing stability and promoting accumulation at the cell membrane."
SMAD2 affects UCHL5
1 | 1 3 2
1 | 1 3 1

trips
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."

reach
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY - motif in Smad7 that interacts with Smurf ubiquitin ligases."

No evidence text available

sparser
"UCH37 also weakly binds to SMAD2 and 3, however it only deubiquitylates ALK5 and hence modifies TGFβ-induced transcription xref ."

reach
"UCH37 binds strongly to Smad7 and weakly to Smad2 and Smad3."

reach
"UCH37 also weakly binds to SMAD2 and 3, however it only deubiquitylates ALK5 and hence modifies TGFbeta induced transcription XREF_BIBR."
| 1

sparser
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."
NLRP3 affects UCHL5
| 3 5
| 3 4

sparser
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."

reach
"Mechanistically, NLRP3 directly bound to UCHL5 and maintained its stability through reducing ubiquitin-proteasome pathway degradation in mandibular MSCs."

reach
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation (XREF_FIG), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 (XREF_FIG)."

sparser
"Mechanistically, NLRP3 directly bound to UCHL5 and maintained its stability through reducing ubiquitin-proteasome pathway degradation in mandibular MSCs."

sparser
"S3D), suggesting that RACK1 could serve as a scaffold protein or a bridge between EST12 and UCHL5-NLRP3."

sparser
"However, UCHL5 did not interact with NLRP3 or with EST12 in RACK1-deficient macrophages upon EST12 stimulation (fig."

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
| 1

sparser
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation ( xref ), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 ( xref )."
GST affects UCHL5
| 8
| 6

sparser
"However, FLAG-S14 (even at the 6×His-UIP1:FLAG-S14 molar ratio of 1:6) had no significant effect on the amount of 6×His-UIP1 bound to GST-UCH37 (lane 6 of Fig. 4B )."

sparser
"Only the fragment containing the N-terminal 101 residues of NFRKB (N101) bound selectively to GST-Uch37 ( Figure 2 B)."

sparser
"Almost no detectable FLAG-S14 was found bound to GST-UCH37 when the molar ratio of 6×His-UIP1:FLAG-S14 was 1:1 (lane 6 of Fig. 4A )."

sparser
"Only the fragment containing the N-terminal 101 residues of NFRKB (N101) bound selectively to GST-Uch37 ( xref )."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To define in more detail the interaction between Uch37 and NFRKB, we generated a series of fragments from the N-terminal third of NFRKB by in vitro translation using a bacterial cell-free system and t[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
VLX1570 affects UCHL5
| 7
VLX1570 inhibits UCHL5.
| 5
VLX1570 inhibits UCHL5. 5 / 5
| 5

reach
"VLX1570 is known to induce apoptosis of myeloma by targeting USP4 and UCHL5 [XREF_BIBR]."

reach
"The small molecule VLX1570 and an analog b-AP15 specifically block the activity of DUBs USP14 and UCHL5 in the 19S regulatory subunit, which results in the rapid accumulation of high molecular weight ubiquitin conjugates, proteasome shutdown, and robust anti-tumor activity in well established orthotopic and xenograft models of MM, lymphoma, Ewing 's sarcoma, and other malignancies [XREF_BIBR - XREF_BIBR]."

reach
"Notwithstanding this success, the potency of pimozide (IC 50 ~ 2muM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC 50 ~ 100nM) (XREF_FIG and XREF_TABLE), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC 50 ~ 100nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240)."

reach
"We observed at 30min that VLX1570 inhibited the activity of both USP14 and UCHL5 in a manner similar to b-AP15, as noted by a decrease in UbVS labeled DUBs (XREF_FIG)."

reach
"VLX1570 and b-AP15 both inhibit USP14 and UCHL5 activity of 19S regulatory particles with the inhibition of USP14 being more pronounced (XREF_FIG)."
VLX1570 binds UCHL5.
| 2
UCHL5 binds VLX1570. 2 / 2
| 2

reach
"VLX1570 preferentially binds the Cys88 residue of UCHL5 and interfaces with a thiol of USP14 at residue Cys114 via a 1,4-Michael 's addition reaction, potentially forming a covalent bond."

reach
"VLX1570 binds and inhibits the activity of USP14 and UCHL5."
USP14 affects RP
| 7
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
5 | 1
5 | 1

No evidence text available

No evidence text available

No evidence text available

sparser
"The human UCH-L5 and yeast UCH37 (YUH1) associate with the 26S proteasome complex xref , xref ."

No evidence text available

No evidence text available
UCHL5 affects SMAD
| 1 6
| 1 6

sparser
"UCHL5 interacts with Smads and reverses Smurf-mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-β receptor in cells [ xref ]."

sparser
"Here, we report a novel interaction between Smads and ubiquitin C-terminal hydrolase UCH37, a deubiquitinating enzyme that could potentially reverse Smurf-mediated ubiquitination."

sparser
"UCHL5 interacts with Smads and potentially reverse Smurf-mediated degradation; it also stabilizes type 1 TGF-beta receptor and regulates TFG-beta signaling xref ."

sparser
"A novel specific interaction between Smad transcription factors and UCH37 was identified [ xref ]."

sparser
"The interaction between Smads and UCH37 could potentially counteract Smurf-mediated ubiquitination."

trips
"The deubiquitinating enzyme UCH37 interacts with Smads and regulates TGF-beta signalling."

sparser
"Recently, we have defined a novel interaction between Smads and UCH37 (ubiquitin C-terminal hydrolase 37), a DUB (de-ubiquitinating enzyme) that could potentially counteract Smurf-mediated ubiquitination."
UCHL5 affects RP
| 7
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
UCHL5 affects PSMD4
6 | 1
6 | 1

No evidence text available

No evidence text available

sparser
"Biochemical data from fission yeast and mammalian cells suggested that the UCH37 also binds to the S5a, a mammalian homologue of Rpn10 [ xref , xref ]."

No evidence text available

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMD2
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
SMAD affects UCHL5
| 1 6
| 1 6

sparser
"UCHL5 interacts with Smads and reverses Smurf-mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-β receptor in cells [ xref ]."

sparser
"Here, we report a novel interaction between Smads and ubiquitin C-terminal hydrolase UCH37, a deubiquitinating enzyme that could potentially reverse Smurf-mediated ubiquitination."

sparser
"UCHL5 interacts with Smads and potentially reverse Smurf-mediated degradation; it also stabilizes type 1 TGF-beta receptor and regulates TFG-beta signaling xref ."

sparser
"A novel specific interaction between Smad transcription factors and UCH37 was identified [ xref ]."

sparser
"The interaction between Smads and UCH37 could potentially counteract Smurf-mediated ubiquitination."

trips
"The deubiquitinating enzyme UCH37 interacts with Smads and regulates TGF-beta signalling."

sparser
"Recently, we have defined a novel interaction between Smads and UCH37 (ubiquitin C-terminal hydrolase 37), a DUB (de-ubiquitinating enzyme) that could potentially counteract Smurf-mediated ubiquitination."
RP affects USP14
| 7
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
PSMD2 affects UCHL5
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

No evidence text available

reach
"BITC and PEITC inhibit USP9x and UCH37."

reach
"BITC and PEITC inhibit USP9x and UCH37 in vitro."

reach
"Here we report that both BITC and PEITC inhibit USP9X and UCH37 and other DUBs at physiologically relevant concentrations and time scales."

reach
"Docking results suggest that benzyl isothiocyanate, phenethyl isothiocyanate, and DL-sulforaphane are more potent inhibitors of UCHL5 than USP14, and these ITCs could interact with the catalytic triads of UCHL5 and USP14."

reach
"Both PEITC and BITC inhibited UCH37 with values of EC 50 of 36 +/- 5 muM and 31 +/- 6 muM, in reasonable agreement with the lysate assays (XREF_FIG and XREF_SUPPLEMENTARY)."

sparser
"We observed a dose-dependent inhibition of UCHL5 by BITC ( xref , lower panel , lanes 3–5 vs ."
Aur affects UCHL5
| 3 3
Aur inhibits UCHL5. 6 / 6
| 3 3

reach
"However, we have recently unraveled that Aur inhibits 19S proteasome associated DUBs (mainly UCHL5 and USP14), accumulates ubiquitinated proteins (Ub-prs), and induces unfolded protein response (UPR) followed by cell apoptosis."

sparser
"Here we report that (i) Aur shows proteasome-inhibitory effect that is comparable to that of bortezomib/Velcade (Vel); (ii) different from bortezomib, Aur inhibits proteasome-associated deubiquitinases (DUBs) UCHL5 and USP14 rather than the 20S proteasome; (iii) inhibition of the proteasome-associated DUBs is required for Aur-induced cytotoxicity; and (iv) Aur selectively inhibits tumor growth in vivo and induces cytotoxicity in cancer cells from acute myeloid leukemia patients."

reach
"We found that the remaining active forms of both UCHL5 and USP14 (i.e., those can be covalently bound by HA-UbVS) were clearly reduced in the 26S proteasomes pre-treated with Aur at 2 muM and became completely undetectable in those pre-treated with 40 muM Aur, indicating that Aur inhibits both UCHL5 and USP14."

reach
"Here we report that (i) Aur shows proteasome-inhibitory effect that is comparable to that of bortezomib and Velcade (Vel); (ii) different from bortezomib, Aur inhibits proteasome associated deubiquitinases (DUBs) UCHL5 and USP14 rather than the 20S proteasome; (iii) inhibition of the proteasome associated DUBs is required for Aur induced cytotoxicity; and (iv) Aur selectively inhibits tumor growth in vivo and induces cytotoxicity in cancer cells from acute myeloid leukemia patients."

sparser
"Molecularly, Aur inhibits 19S-associated DUBs USP14 and UCHL5 [ xref , xref ]."

sparser
"We found that the remaining active forms of both UCHL5 and USP14 (i.e., those can be covalently bound by HA-UbVS) were clearly reduced in the 26S proteasomes pre-treated with Aur at 2 μM and became completely undetectable in those pre-treated with 40 μM Aur (Fig. xref ), indicating that Aur inhibits both UCHL5 and USP14."

reach
"UCHL5 promotes HR and extensive DNA-end resection."

reach
"Since INO80 chromatin remodeling complex had been implicated in DSB-end resection and DSB repair [97,98], UCHL5 may contribute HR via chromatin re-organization such as histone eviction."

reach
"Specifically, UCHL5 promotes DNA-end resection and HR through regulating the stability of the NFRKB protein that is a subunit of chromatin remodeling complex INO80 [XREF_BIBR, XREF_BIBR]."

reach
"In addition to identifying many DUBs with DDR roles, this work also lead to establish that one, UCHL5 promotes DSB end resection and HR through regulating the stability of the NFRKB protein that is a subunit of chromatin remodeling complex INO80."

reach
"The function of UCHL1 in DSB repair is still unclear, but it has been established that UCHL5 promotes repair by enhancing DNA-end resection, whereas BAP1 promotes HR via an unknown mechanism ."

reach
"UCHL5 (Ubiquitin carboxyl-terminal hydrolase isozyme L5) positively regulates DSB end resection and HR repair by deubiquitinating and stabilizing NFRKB protein (nuclear factor related to kappaB bindin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects TGFBR1
1 1 | 4
UCHL5 deubiquitinates TGFBR1.
1 | 4
UCHL5 deubiquitinates TGFBR1. 5 / 5
1 | 4

reach
"XREF_BIBR reported that UCHL5 de-ubiquitinates and stabilizes TbetaRI in human cancer cells."

reach
"One such DUB is UCH37, which was shown to bind to Smad7 and deubiquitinate TbetaRI."

"Via SMAD7, UCH37 can further be recruited to TβRI, where it removes polyubiquitin chains synthesized by SMURF"

reach
"It has been revealed that the DUBs, UCH37, USP11, and USP15, de-ubiquitinate and stabilize TbetaRI."

reach
"Similarly to USP15, another deubiquitylating enzyme, UCH37, forms a complex with Smad7 by binding to a sequence that is distinct from the PY motif with which Smurf1 or Smurf2 interact, and also deubiquitylates and stabilizes TbetaRI."
UCHL5 increases the amount of TGFBR1.
1 |
UCHL5 increases the amount of TGFBR1. 1 / 1
1 |

"Smad7 can act as an adaptor able to recruit uch37 to the type i tgf-beta receptor. Consequently, uch37 dramatically up-regulates tgf-beta-dependent gene expression by de-ubiquitinating and stabilizing the type i tgf-beta receptor."
UCHL5 affects SNRPF
| 6
UCHL5 decreases the amount of SNRPF. 4 / 4
| 4

reach
"Thus, the possible mechanism of UCH-L5 downregulates SNRPF expression may through interacting with NFRKB or other components of INO80 complex."

reach
"We also found that knockdown of UCH-L5 could upregulate the mRNA and protein level of SNRPF, while overexpression of UCH-L5 downregulated the mRNA and protein level of SNRPF in U87MG cells infected by Lentivirus."

reach
"Accordingly, we found UCH-L5 inhibited mRNA expression and protein level of SNRPF both in U87MG cells and U251 cells significantly."

reach
"To further confirm that UCH-L5 inhibits SNRPF expression, U87MG and U251 cells were subjected to analysis for lentivirus mediated gene knockdown and overexpression."
Modified UCHL5 decreases the amount of SNRPF. 2 / 2
| 2

reach
"We also found that knockdown of UCH-L5 could upregulate the mRNA and protein level of SNRPF, while overexpression of UCH-L5 downregulated the mRNA and protein level of SNRPF in U87MG cells infected by Lentivirus."

reach
"Considering the function of UCH-L5 regulates DNA transcription and mRNA expression of the spliceosome components, the possible reason is that knockdown of UCH-L5 expression upregulates mRNA level of SNRPF which promots the splicing of downstream oncogenes, causing a promotion of oncogenic genes and tumorigenesis."
UCHL5 affects PSMC2
6 |
6 |

No evidence text available

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No evidence text available

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No evidence text available
UCHL5 affects PSMC1
6 |
6 |

No evidence text available

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No evidence text available

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No evidence text available
UCHL5 affects PRPF19
2 | 1 3
2 | 1 3

reach
"Subsequently, the possibility that UCH37 could interact with PRP19 was confirmed by co-IP and confocal laser scanning microscopy analysis."

sparser
"Meanwhile, control experiments showed that PRP19 was exactly associated with UCH37, but not with Flag ( Supplementary Fig. 3 )."

sparser
"Subsequently, the possibility that UCH37 could interact with PRP19 was confirmed by co-IP and confocal laser scanning microscopy analysis."

No evidence text available

No evidence text available

sparser
"This interaction of UCH37 with PRP19 with endogenously UCH37 was confirmed in Huh7 cells ( Fig. 7 C, D)."
UCHL5 affects NOS2
2 | 1 2
UCHL5 binds NOS2.
2 | 2
2 | 1

sparser
"In this study, we show that Rpn13 is involved in iNOS degradation and is required for iNOS interaction with the deubiquitination protein UCH37."

No evidence text available

No evidence text available
ADRM1 binds UCHL5 and NOS2. 1 / 1
| 1

sparser
"We previously found that the cisplatin treatment induced Rpn13 transcription by p-ΔNp63α and subsequently increased the physical interaction of Rpn13, UCH37 and NOS2 proteins leading to an essential degradation of the latter through a proteasome-dependent mechanism in SCC cells [ xref ]."
UCHL5 ubiquitinates NOS2.
| 1
UCHL5 ubiquitinates NOS2. 1 / 1
| 1

reach
"Ubiquitinated NOS2 forms a ternary complex with the proteasome associated ubiquitin receptor ADRM1, and the ubiquitin hydrolase UCHL5, both of which are important mediators of NOS2 proteasomal degradation [XREF_BIBR]."
UCHL5 affects INO80E
6 |
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
UBE3A affects UCHL5
3 1 | 1 1
UBE3A ubiquitinates UCHL5.
1 | 1 1
UBE3A leads to the ubiquitination of UCHL5. 3 / 3
1 | 1 1

sparser
"It was reported that ubiquitination of Rpt5, Rpn10/S5A, Rpn13/ADRM1, and UCHL5 is induced by proteasome-associated UBE3A and UBE3C at the purified proteasome [ xref ]."

reach
"Ubiquitination of S5a and Uch37 on PA700 was strongly promoted by addition of UbE3A, whereas Rpt5 ubiquitination was only mildly increased (XREF_FIG, lane 6)."
UBE3A binds UCHL5.
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
SERPB12 affects UCHL5
| 5 1
SERPB12 increases the amount of UCHL5.
| 2
Modified SERPB12 increases the amount of UCHL5. 2 / 2
| 2

reach
"SLFN12 or SERPB12 overexpression increases expression of the complementary deubiquitylases USP14 and UCHL5 in vitro, and SERPB12 stimulates USP14 deubiquitylase activity."

reach
"Either SLFN12 overexpression (XREF_FIG and XREF_FIG) or SERPB12 overexpression (XREF_FIG and XREF_FIG) increased both USP14 (XREF_FIG and XREF_FIG) and UCHL5 (XREF_FIG and XREF_FIG) expression."
SERPB12 binds UCHL5.
| 1 1
UCHL5 binds SERPB12. 2 / 2
| 1 1

reach
"Alternatively, it is possible that, as reported by Fang in hepatocellular carcinoma tissues [XREF_BIBR], SERPB12 also interacts with UCHL5 directly in this pathway even though we were unable to co-precipitate the two proteins here."

sparser
"Alternatively, it is possible that, as reported by Fang in hepatocellular carcinoma tissues [ xref ], SERPB12 also interacts with UCHL5 directly in this pathway even though we were unable to co-precipitate the two proteins here."
SERPB12 activates UCHL5.
| 2
SERPB12 activates UCHL5. 2 / 2
| 2

reach
"SERPB12 stimulated USP14 but not UCHL5 activity."

reach
"Moreover, when we mixed purified SERPB12 with each of these two purified deubuiquitylases in vitro, SERPB12 stimulated the DUB activity of USP14 (XREF_FIG) but not UCHL5 (XREF_FIG)."
RP affects UCHL5, and USP14
| 6
USP14 binds UCHL5 and RP. 6 / 6
| 6

sparser
"Apart from the integral DUB — Rpn11, there are other pDUBs — USP14 and UCHL5, which bind transiently to the 19S RP during the degradation process [ xref , xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [ xref ]."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."

sparser
"It selectively inhibits the activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP while not affecting the non-proteasomal DUBs."

sparser
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [ xref ]."

sparser
"Similar to the mode of action exerted by b-AP15, this set of compounds inhibits activity of two DUBs, UCHL5 and USP14 that are associated with 19S RP, and do not affect non-proteasomal DUBs [75] ."
PTPN2 affects UCHL5
| 4 2
PTPN2 inhibits UCHL5.
| 2 1
PTPN2 inhibits UCHL5. 3 / 3
| 2 1

sparser
"PtPT significantly inhibited USP14 and UCHL5, thereby accumulating Ub-conjugates."

reach
"It is known that b-AP15 and PtPT can inhibit the activity of both USP14 and UCHL5 simultaneously, implying that the downregulation of ERα by b-AP15 and PtPT may attribute to the suppression of USP14 and UCHL5."

reach
"PtPT significantly inhibited USP14 and UCHL5, thereby accumulating Ub-conjugates."
PTPN2 activates UCHL5.
| 2
PTPN2 activates UCHL5. 2 / 2
| 2

reach
"PtPT inhibits the UPS by targeting DUBs USP14 and UCHL5 associated with 26S proteasomes."

reach
"These computational and experimental results indicate that PtPT can selectively target UCHL5 and USP14, two proteasome associated DUBs."
PTPN2 binds UCHL5.
| 1
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
PSMD8 affects HAUS7
| 6
| 6

sparser
"These results indicate that S14 and UIP1 could bind UCH37 specifically in vitro in a C-terminal extension dependent manner."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The Y187 yeast strain harbouring S14/pGAD-GH or UIP1/pGAD-GH was mated with PJ69-2A strain transformed with pU1, pU2 or pU3 respectively to test the interaction between S14 or UIP1 with UCH37, UCH37ΔC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report the identification of two proteins, S14 and UCH37 interacting protein 1 (UIP1), which interact with UCH37."

sparser
"The interaction of UCH37 with S14 or UIP1 was confirmed by in vitro binding assay and in vivo co-immunoprecipitation analysis."
PSMC2 affects UCHL5
6 |
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PSMC1 affects UCHL5
6 |
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PRPF19 affects UCHL5
2 | 1 3
2 | 1 3

reach
"Subsequently, the possibility that UCH37 could interact with PRP19 was confirmed by co-IP and confocal laser scanning microscopy analysis."

sparser
"Meanwhile, control experiments showed that PRP19 was exactly associated with UCH37, but not with Flag ( Supplementary Fig. 3 )."

sparser
"Subsequently, the possibility that UCH37 could interact with PRP19 was confirmed by co-IP and confocal laser scanning microscopy analysis."

No evidence text available

No evidence text available

sparser
"This interaction of UCH37 with PRP19 with endogenously UCH37 was confirmed in Huh7 cells ( Fig. 7 C, D)."
INO80E affects UCHL5
6 |
6 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
HAUS7 affects PSMD8, and UCHL5
| 6
| 6

sparser
"These results indicate that S14 and UIP1 could bind UCH37 specifically in vitro in a C-terminal extension dependent manner."

sparser
"We carried out in vitro binding assays to confirm that S14 or UIP1 could individually bind UCH37 in vitro under conditions other than that of the yeast two-hybrid system, As shown in Fig. 3A,B [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"The Y187 yeast strain harbouring S14/pGAD-GH or UIP1/pGAD-GH was mated with PJ69-2A strain transformed with pU1, pU2 or pU3 respectively to test the interaction between S14 or UIP1 with UCH37, UCH37ΔC[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report the identification of two proteins, S14 and UCH37 interacting protein 1 (UIP1), which interact with UCH37."

sparser
"The interaction of UCH37 with S14 or UIP1 was confirmed by in vitro binding assay and in vivo co-immunoprecipitation analysis."
E2F1 affects UCHL5
4 | 1
E2F1 binds UCHL5.
4 |
4 |

No evidence text available

No evidence text available

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No evidence text available
E2F1 increases the amount of UCHL5.
| 1
E2F1 increases the amount of UCHL5. 1 / 2
| 1

reach
"Interestingly, UCH37 expression is induced by E2F1, and its level rises in G1/S transition and S phase, suggesting a positive feedback loop between UCH37 and E2F1."
| 4 1
| 2 1

reach
"UBH-4 and the mammalian ortholog UCHL5 were revealed to interact with the RPN-13 proteasome subunit [XREF_BIBR, XREF_BIBR] and to negatively regulate UPS activity [XREF_BIBR]."

sparser
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."

reach
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."
| 2

reach
"We also find that proteasome subunit RPN13, an activator of UCHL5, could enhance the effect of UCHL5 on Smo protein level."

reach
"Interestingly, UCHL5 is activated by the proteasome subunit RPN13 and inhibited by the INO80G subunit ."

reach
"Both PEITC and BITC inhibited UCH37 with values of EC 50 of 36 +/- 5 muM and 31 +/- 6 muM, in reasonable agreement with the lysate assays (XREF_FIG and XREF_SUPPLEMENTARY)."

reach
"Here we report that both BITC and PEITC inhibit USP9X and UCH37 and other DUBs at physiologically relevant concentrations and time scales."

reach
"In a recent publication, PEITC has been shown to inhibit the activity of proteasomal cysteine deubiquitinases UCHL5 [62]."

reach
"BITC and PEITC inhibit USP9x and UCH37 in vitro."

reach
"BITC and PEITC inhibit USP9x and UCH37."
Auranofin affects UCHL5
| 5
Auranofin inhibits UCHL5.
| 4
| 4

reach
"Auranofin (Aur) inhibits proteasome associated deubiquitinases (DUBs) UCHL5 and USP14 rather than the 20S proteasome; inhibition of the proteasome associated DUBs is required for Aur induced cytotoxicity; and Aur selectively inhibits tumor growth in vivo and induces cytotoxicity in cancer cells from patients with acute myeloid leukemia [XREF_BIBR]."

reach
"The computational model also indicates that an active metabolite of auranofin can inhibit UCHL5 and USP14."

reach
"Auranofin was recently reported to inhibit proteasome activity at the level of the proteasome associated deubiquitinases (DUBs) UCHL5 and USP14."

reach
"HA-UbVS pretreatment of auranofin could bind the HA tagged UbVS in the purified 26S proteasome, supporting that auranofin inhibits UCHL5 and USP14."
Auranofin binds UCHL5.
| 1
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
ULD affects UCHL5
| 5
| 4

sparser
"Like Rpn13, NFRKB employs a DEUBAD domain (NFRKB DEU ) to interact with the ULD domain of Uch37 [ xref , xref ]."

sparser
"Both RPN13 DEU and INO80 DEU primarily bind the C-terminal ULD domain of UCH-L5, but are positioned radically differently relative to the UCH-L5 CD, which itself hardly changes conformation among all UCH-L5 structures."

sparser
"Rpn13 and NFRKB both employ DEUBAD (DEUBiquitinase Adaptor) domains to bind Uch37 via its unique C-terminal ULD (Uch37-Like Domain)."

sparser
"Like Rpn13, NFRKB employs a DEUBAD domain (NFRKB DEU ) to interact with the ULD domain of Uch37 [12,13] ."
UCHL5 binds BAP1 and ULD. 1 / 1
| 1

sparser
"UCH-L5 and BAP1 both interact with the DEUBAD domains of their binding partners via their ULDs."
UCHL5 affects UBC
5 |
5 |

No evidence text available

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No evidence text available

No evidence text available
UCHL5 affects TFPT
5 |
5 |

No evidence text available

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No evidence text available
UCHL5 affects SP
| 5
| 5

sparser
"Prediction of Sp-UCHL3 and Sp-UCHL5 Three-Dimensional (3D) Structure."

sparser
"In the present study, Sp-UCHL3 mRNA expression in T3 was significantly higher than in the T1 and T2 stages ( p < 0.05), while Sp-UCHL5 mRNA expressions in the three stages of testes development were not significantly different from each other ( p > 0.05), indicating that Sp- UCHL3 and Sp- UCHL5 have different aspects in regulating mechanism of testis development."

sparser
"Meanwhile, the full-length cDNA of Sp-UCHL5 is 1217 bp."

sparser
"Therefore, the expression patterns of Sp-UCHL3 and Sp-UCHL5 in different tissues and ovarian development suggest that they play important roles in cellular proteins’ degradation during oogenesis and ovarian maturation."

sparser
"Phylogenetic and Homology Analyses of Sp-UCHL3 and Sp-UCHL5."
UCHL5 affects RA190
| 3 2
UCHL5 binds RA190. 5 / 5
| 3 2

reach
"We also report anti-cancer molecule RA190, which binds covalently to hRpn13 and UCHL5, to require hRpn13 Pru and not UCHL5 for cytotoxicity."

reach
"RA190 binds Uch37 and inhibits its catalytic activity."

sparser
"Thus, there are no compelling data from the isoTOP-ABPP experiment to indicate RA190 binding to Uch37 in cellulo ."

reach
"Biophysical analyses in combination with cell based assays indicate that RA190 directly binds and inactivates Uch37 [XREF_BIBR]."

sparser
"We also scrutinized the dataset for evidence of RA190 binding to the Rpn13-associated deubiquitylase Uch37 as well."
UCHL5 affects PSMD7
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UCHL5 affects PSMD3
5 |
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UCHL5 affects PSMD13
5 |
5 |

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UCHL5 affects PSMD11
5 |
5 |

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UCHL5 affects PSMC6
5 |
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UCHL5 affects PSMC5
5 |
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UCHL5 affects PSMC4
5 |
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UCHL5 affects PSMC3
5 |
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UCHL5 affects PAAF1
3 | 2
3 | 2

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sparser
"Based on these clues, it is reasonable to propose that both UCHL5 and PAAF1 can bind the 19S regulatory complex to form a larger one."

No evidence text available

sparser
"The database contains no information about whether it can interact with PAAF1 and UCHL5, but the predicted “core_5” suggests this possibility."

No evidence text available
UCHL5 affects Neoplasms
| 5
UCHL5 inhibits Neoplasms.
| 3
| 3

reach
"We also found that UCHL5 downregulation significantly suppressed both tumor growth in vivo and cell proliferation and migration in vitro."

reach
"Interestingly, b-AP15, a dual inhibitor of Usp14 and Uch37, was shown to prevent protein degradation and inhibit tumor progression in acute myeloid leukemia models [222]."

reach
"As shown in Figures 7A–C, UCHL5 silence obviously repressed the size, volume, and weight of the xenograft tumors in the nude mice."
UCHL5 activates Neoplasms.
| 2
| 2

reach
"On the other hand, UCH37, the deubiquitinase activated by ADRM1, inhibits glioma cell migration and invasion [42], suggesting that ADRM1 inhibition could have a positive or negative effect on tumor progression, depending on the stage of the tumor.The discovery of a new role for HDAC8 and ADRM1 in MGMT regulation expands the possibilities of the development of new therapies to overcome TMZ resistance in GBM, although several questions about the mechanisms remain to be answered."

reach
"Treatment with b-AP15, a UCHL5 and USP14 deubiquitinating activity inhibitor in 19S regulatory subunit, induces tumor regression and prolong the survival period of tumor-loaded mice through down-regulation of COPS5 and its downstream AP-1 and E2F1, and up-regulation of the cell cycle-related proteins p27 and Cyclin E1."
UCHL5 affects HSPA5
5 |
5 |

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| 2 3
UCHL5 activates Cell Survival.
| 2 1
| 2 1

eidos
"UCHL5 knockdown decreased cell viability , increased apoptosis , and arrested cell cycle in HCE-1-A endometrial cancer cells ."

reach
"UCHL5 overexpression raised cell viability at different time points after lentiviral infection (Figure 4C)."

eidos
"Upregulation of UCHL5 promoted cell viability , attenuated apoptosis , and accelerated cell cycle in AN3-CA cells ."
UCHL5 inhibits Cell Survival.
| 2

reach
"USP14 and UCHL5 short interfering RNA knockdown decreases MM cell viability."

reach
"Knock-down of UCHL5 expression in non small cell lung carcinoma cells resulted in alterations in components of the mitochondrial apoptosis pathway including down-regulation of the anti-apoptotic Bcl-2[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects COPS5
| 2 3
| 1 3

sparser
"UCH37 interacts with COPS5 to induce the ubiquitination and degradation of p53 and p21, promoting cell proliferation."

sparser
"Moreover, the endogenous UCH37 interaction with endogenous COPS5 in U2OS cells (Supplementary Fig. xref ) was also demonstrated."

sparser
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA (Fig. xref ) or anti-Flag antibody (Fig. xref )."

reach
"Moreover, the endogenous UCH37 interaction with endogenous COPS5 in U2OS cells was also demonstrated."
UCHL5 binds COPS5 and 293T. 1 / 1
| 1

reach
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA or anti-Flag antibody."
UCHL5 affects AXIN1
| 5
| 5

sparser
"These results imply that the interaction between Axin1 and UCHL5 is most likely mediated by the coordination of multiple domains of Axin1 protein."

sparser
"In vitro binding assay using recombinant proteins demonstrated that UCHL5 directly interacts with Axin1 protein (Fig.  xref f)."

sparser
"Coincidentally, UCHL5 interacts with Axin1 through multiple domains (1–705; RGS, GSK3ß, ß-catenin, and PP2A binding domain) where intra-molecular interactions arise."

sparser
"Co-immunoprecipitation (Co-IP) analysis revealed that UCHL5 interacted with Axin1 in both ectopically and endogenously expressed conditions (Fig.  xref c,d)."

sparser
"We then examined if UCHL5 physically interacts with Axin1, which is a scaffolder of the ß-catenin destruction complex."
UCHL5 affects ACTR5
5 |
5 |

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UBC affects UCHL5
5 |
5 |

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TFPT affects UCHL5
5 |
5 |

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TCF7 affects UCHL5
| 4 1
| 4 1

sparser
"UCH37 can specifically bind and deubiquitinate transcription factor 7 (Tcf7) to activate Wnt signaling in human liver cancer cells [ xref ]."

reach
"Having demonstrated that Uch37 is required for Wnt signalling and specifically interacts with Tcf7, we next hypothesized that Uch37 binds Tcf7 to serve as a bona fide DUB for Tcf7 protein during Wnt signalling."

reach
"Our Co-IP and GST pulldown analyses clearly demonstrated that Uch37 binds Tcf7 directly, and that the physical interaction requires the C-terminal region of Tcf7 protein that includes HMG domain and C-terminal extension."

reach
"Moreover, GST-pull down assay using purified recombinant Uch37 (His Uch37) and Tcf7 (GST-Tcf7) proteins demonstrated a direct interaction between Uch37 and Tcf7 (XREF_FIG)."

reach
"Uch37 specifically and directly interacts with Tcf7."
SP affects UCHL5
| 5
| 5

sparser
"Prediction of Sp-UCHL3 and Sp-UCHL5 Three-Dimensional (3D) Structure."

sparser
"In the present study, Sp-UCHL3 mRNA expression in T3 was significantly higher than in the T1 and T2 stages ( p < 0.05), while Sp-UCHL5 mRNA expressions in the three stages of testes development were not significantly different from each other ( p > 0.05), indicating that Sp- UCHL3 and Sp- UCHL5 have different aspects in regulating mechanism of testis development."

sparser
"Meanwhile, the full-length cDNA of Sp-UCHL5 is 1217 bp."

sparser
"Therefore, the expression patterns of Sp-UCHL3 and Sp-UCHL5 in different tissues and ovarian development suggest that they play important roles in cellular proteins’ degradation during oogenesis and ovarian maturation."

sparser
"Phylogenetic and Homology Analyses of Sp-UCHL3 and Sp-UCHL5."
PSMD7 affects UCHL5
5 |
5 |

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PSMD3 affects UCHL5
5 |
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PSMD13 affects UCHL5
5 |
5 |

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PSMD11 affects UCHL5
5 |
5 |

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PSMC6 affects UCHL5
5 |
5 |

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PSMC5 affects UCHL5
5 |
5 |

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PSMC4 affects UCHL5
5 |
5 |

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PSMC3 affects UCHL5
5 |
5 |

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PAAF1 affects UCHL5
3 | 2
3 | 2

No evidence text available

sparser
"Based on these clues, it is reasonable to propose that both UCHL5 and PAAF1 can bind the 19S regulatory complex to form a larger one."

No evidence text available

sparser
"The database contains no information about whether it can interact with PAAF1 and UCHL5, but the predicted “core_5” suggests this possibility."

No evidence text available
NOS2 affects UCHL5
2 | 2
2 | 1

sparser
"In this study, we show that Rpn13 is involved in iNOS degradation and is required for iNOS interaction with the deubiquitination protein UCH37."

No evidence text available

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ADRM1 binds UCHL5 and NOS2. 1 / 1
| 1

sparser
"We previously found that the cisplatin treatment induced Rpn13 transcription by p-ΔNp63α and subsequently increased the physical interaction of Rpn13, UCH37 and NOS2 proteins leading to an essential degradation of the latter through a proteasome-dependent mechanism in SCC cells [ xref ]."
HSPA5 affects UCHL5
5 |
5 |

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COPS5 affects UCHL5
| 2 3
| 1 3

sparser
"UCH37 interacts with COPS5 to induce the ubiquitination and degradation of p53 and p21, promoting cell proliferation."

sparser
"Moreover, the endogenous UCH37 interaction with endogenous COPS5 in U2OS cells (Supplementary Fig. xref ) was also demonstrated."

sparser
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA (Fig. xref ) or anti-Flag antibody (Fig. xref )."

reach
"Moreover, the endogenous UCH37 interaction with endogenous COPS5 in U2OS cells was also demonstrated."
UCHL5 binds COPS5 and 293T. 1 / 1
| 1

reach
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA or anti-Flag antibody."
AXIN1 affects UCHL5
| 5
| 5

sparser
"These results imply that the interaction between Axin1 and UCHL5 is most likely mediated by the coordination of multiple domains of Axin1 protein."

sparser
"In vitro binding assay using recombinant proteins demonstrated that UCHL5 directly interacts with Axin1 protein (Fig.  xref f)."

sparser
"Coincidentally, UCHL5 interacts with Axin1 through multiple domains (1–705; RGS, GSK3ß, ß-catenin, and PP2A binding domain) where intra-molecular interactions arise."

sparser
"Co-immunoprecipitation (Co-IP) analysis revealed that UCHL5 interacted with Axin1 in both ectopically and endogenously expressed conditions (Fig.  xref c,d)."

sparser
"We then examined if UCHL5 physically interacts with Axin1, which is a scaffolder of the ß-catenin destruction complex."
ACTR5 affects UCHL5
5 |
5 |

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4 |
Valproic acid increases the amount of UCHL5. 4 / 4
4 |

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YY1 affects UCHL5
4 |
4 |

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Ubiquitin affects USP14
| 4
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
USP14 affects Proteasome
| 4
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
USP14 affects PSMD14
| 4
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
UCHL5 affects YY1
4 |
4 |

No evidence text available

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No evidence text available
UCHL5 affects TGFBR
| 4
UCHL5 deubiquitinates TGFBR.
| 3
UCHL5 deubiquitinates TGFBR. 2 / 2
| 2

reach
"In addition, we show that UCH37 can deubiquitinate and stabilize the type I TGF-beta receptor."

reach
"The study hypothesized that Smad7 could act as an adaptor to recruit UCH37 to the type I TGF-beta receptor and showed that UCH37 dramatically up-regulates TGF-beta-dependent gene expression by deubiquitinating and stabilizing the type I TGF-beta receptor [XREF_BIBR]."
UCHL5 deubiquitinates TGFBR on R1. 1 / 1
| 1

reach
"UCHL5 can deubiquitinate and stabilize Smads as well as TGF-beta receptor 1 (TGF-beta R1) and therefore activate TGF-beta signaling [XREF_BIBR]."
UCHL5 binds TGFBR.
| 1
| 1

reach
"UCHL5 interacts with Smads and reverses Smurf mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-beta receptor in cells [XREF_BIBR]."
UCHL5 affects RUVBL2
4 |
4 |

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UCHL5 affects PSMD6
4 |
4 |

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UCHL5 affects PSMD12
4 |
4 |

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UCHL5 affects MCRS1
4 |
4 |

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UCHL5 affects INO80B
4 |
4 |

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UCHL5 affects IL1B
| 4
UCHL5 inhibits IL1B.
| 2
UCHL5 inhibits IL1B. 2 / 2
| 2

reach
"It is possible that the level of UCH37 silencing we achieved with siRNA may not have been sufficient to observe a measurable effect on nigericin induced IL-1beta release."

reach
"However, inhibition of USP14 activity with IU1, silencing of UCH37 with siRNA, or a combination of both approaches to inhibit both USP14 and UCH37 simultaneously did not inhibit the release of IL-1beta (XREF_FIG)."
UCHL5 activates IL1B.
| 2
UCHL5 activates IL1B. 2 / 2
| 2

reach
"We also showed that overexpression of UCH-L5 was associated with a significant increase in caspase-1 activity, while inhibition of UCH-L5 by selective inhibitor [XREF_BIBR] or UCH-L5 knock-down led to decrease in inflammasome dependent IL-1beta release in chicken as well as in human macrophages during infection with Salmonella and during inflammasome activation by lipopolysaccharide (LPS) and nigericin."

reach
"UCH-L5 inhibition down-regulates IL-1beta release during inflammasome activation in macrophages."
UCHL5 affects ACTR8
4 |
4 |

No evidence text available

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No evidence text available
UBE2W affects UCHL5
| 3 1
UBE2W ubiquitinates UCHL5. 4 / 4
| 3 1

reach
"Strikingly, we observed that UCHL5 is N-terminally ubiquitinated by UBE2W, and this modification significantly enhanced its catalytic activity in vitro (Figs."

sparser
"In addition, UbE2L3 and UbE2W slightly enhanced ubiquitination of Uch37 ( xref )."

reach
"In addition, UbE2L3 and UbE2W slightly enhanced ubiquitination of Uch37 (XREF_FIG)."

reach
"To validate that these two DUBs were indeed N-terminally ubiquitinated, we established in vitro ubiquitination assays with purified proteins and observed that UBE2W can monoubiquitinate lysine-less versions of both UCHL1 and UCHL5 (Fig. 7a)."
RUVBL2 affects UCHL5
4 |
4 |

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RPN13DEUBAD affects UCHL5
| 2 2
RPN13DEUBAD activates UCHL5. 4 / 4
| 2 2

eidos
"On the basis of multiple intermediate structures , it was found that RPN13DEUBAD activates UCH-L5 by positioning its domains while INO80DEUBAD inhibits UCH-L5 by blocking Ub binding.18 Finally , monoUb probes can also be employed to study the mechanism for activation and thioester bond formation in E1s ."

eidos
"To explain regulatory mechanisms , Ub-Prg was used to capture the catalytic conformation by which RPN13DEUBAD activates UCH-L5 ; and the catalytic conformation that truncated INO80DEUBAD lacks to inhibit UCH-L5 ."

reach
"To explain regulatory mechanisms, Ub-Prg was used to capture the catalytic conformation by which RPN13DEUBAD activates UCH-L5; and the catalytic conformation that truncated INO80DEUBAD lacks to inhibit UCH-L5."

reach
"On the basis of multiple intermediate structures, it was found that RPN13DEUBAD activates UCH-L5 by positioning its domains while INO80DEUBAD inhibits UCH-L5 by blocking Ub binding.18Finally, monoUb probes can also be employed to study the mechanism for activation and thioester bond formation in E1s."
Proteasome affects USP14
| 4
| 4

sparser
"USP14 is one of the three proteasome-associated deubiquitinase (DUB) enzymes, USP14 and UCHL5 being the two that are associated reversibly with the proteasome [ xref ]."

sparser
"USP14 and UCHL5 associate with the proteasome through different ubiquitin receptors, USP14 to Rpn1 at the base of the 19S RP and UCHL5 to Rpn13/ADRM1 at the top of the 19S RP [ xref ] ( xref )."

sparser
"Later, it was determined that b-AP15 targeted the proteasome-bound USP14 and UCHL5, which belong to the USP and UCH families, respectively [ xref ]."

sparser
"The HA-UbVS pull-down also isolated known DUBs interactors ( xref ) such as proteasome subunits, consistent with the known interaction of UCHL5 and USP14 with the proteasome ( xref )."
PSMD6 affects UCHL5
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
PSMD14 affects USP14
| 4
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
PSMD12 affects UCHL5
4 |
4 |

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MCRS1 affects UCHL5
4 |
4 |

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INO80B affects UCHL5
4 |
4 |

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ACTR8 affects UCHL5
4 |
4 |

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Pyrithione affects UCHL5
| 2 1
Pyrithione activates UCHL5.
| 2
| 2

reach
"Another compound, copper pyrithione (CuPT), was reported to target both 19S proteasome specific DUBs, UCH37 and USP14, as well as 20S proteolytic peptidases."

reach
"Here we report that zinc pyrithione (ZnPT) targets the proteasome associated DUBs (USP14 and UCHL5) and inhibits their activities, resulting in a rapid accumulation of protein-ubiquitin conjugates, but without inhibiting the proteolytic activities of 20S proteasomes."
Pyrithione inhibits UCHL5.
| 1
| 1

sparser
"And only recently we have definitely confirmed that nickel pyrithione (NiPT) inhibits the 19S proteasome-associated deubiquitinases (DUBs) USP14 and UCHL5, but not the 20S proteasome peptidases, and the inhibition of proteasome-associated DUBs induces NiPT-mediated cytotoxicity, revealing a novel mechanism for the anti-cancer effects of nickel-containing compounds [ xref ]."
Methylmercury chloride increases the amount of UCHL5. 3 / 3
3 |

No evidence text available

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No evidence text available
BAP15 affects UCHL5
| 3
BAP15 inhibits UCHL5. 3 / 3
| 3

reach
"Taken together, these data reveal that the inhibition of UCHL5 by bAP15 stabilizes Smad2 and induces downregulation of TGF-β signaling."

reach
"In contrast to 20S proteasome inhibitors, bAP15 blocks the deubiquitylating activity of both USP14 and UCHL5 to induce strong anti-tumor activity without affecting proteolytic activities of the 20S proteasome [25, 26, 28, 29]."

reach
"In the present study, we revealed, for the first time, evidence of the clinical significance of cytoplasmic UCHL5 expression in ovarian cancer, and demonstrated that bAP15 significantly suppressed UCHL5 in TP53-mutant ovarian cancer cell lines in a dose-dependent manner through downregulation of the TGF-β/Smad signaling pathway (Figure 7)."

reach
"In addition, UCH37 stabilizes Smad2/3 and promotes TGF-b1 signal transduction (39) ."

reach
"In conclusion, our results that UCHL5 promoted the growth of EC in vivo and vitro via activating the Wnt/β-catenin signaling pathway may provide potential targets for EC control in the future."

reach
"In addition, UCH37 stabilizes Smad2/3 and promotes TGF-β1 signal transduction (39)."
| 2 1

reach
"UBH-4 and the mammalian ortholog UCHL5 were revealed to interact with the RPN-13 proteasome subunit [XREF_BIBR, XREF_BIBR] and to negatively regulate UPS activity [XREF_BIBR]."

sparser
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."

reach
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."
UCHL5 affects paaD
| 3
UCHL5 activates paaD. 3 / 3
| 3

reach
"In contrast, UCHL5 deficiency could suppress PAAD stemness features, and ectopic expression of ELK3 could rescue this effect."

reach
"UCHL5 could also promote PAAD migration in vitro and in vivo."

reach
"UCHL5 could thus promote self-renewal abilities of PAAD and targeting ELK3 could inhibit the stemness features."
UCHL5 affects cell cycle
| 1 2
| 1 2

eidos
"As shown in Figure 3E , UCHL5 silent cells showed a remarkable increase in the sub-G1 population compared to negative control , indicating that UCHL5 deficiency induced cell cycle arrest ."

reach
"UCHL5 knockdown in LUAD cells significantly inhibited cell proliferation and reduced the expression of key cell cycle proteins."

reach
"Additionally, XAV939 treatment showed a smaller sub-G1 population when compared with the untreated group and prevented the cell cycle acceleration caused by UCHL5 overexpression (Figure 6D)."
UCHL5 affects Wnt/β-catenin
| 3
UCHL5 activates Wnt/β-catenin. 3 / 3
| 3

reach
"In addition, UCHL5 overexpression mediated by lentivirus vectors activated Wnt/β-catenin signaling in endometrial cancer cells and resulted in a decrease in apoptosis and an increase in proliferation."

reach
"To further confirm whether the UCHL5-regulated cell phenotype in EC cells was mediated by the Wnt/β-catenin signaling pathway, Wnt/β-catenin inhibitor XAV939 was used in the present study."

reach
"In conclusion, our results that UCHL5 promoted the growth of EC in vivo and vitro via activating the Wnt/β-catenin signaling pathway may provide potential targets for EC control in the future."
UCHL5 affects USP14, and Ubiquitin
| 3
| 3

sparser
"Indeed, Ub-VS assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"Indeed, ubiquitin vinyl sulfone assay confirmed the inhibitory activity of each ITC on the ubiquitin-binding activity of UCHL5 and USP14."

sparser
"We found that FB-28 and FB-71 at 2.5 μM were able to completely inhibit the ubiquitin-binding activity of both UCHL5 and USP14, while DAST at the same concentration had only a partial inhibitory effect ( xref , top and bottom panels, lanes 3–5 vs. 1–2)."
UCHL5 affects UBE3A
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects TGF-beta-dependent gene
| 3
UCHL5 increases the amount of TGF-beta-dependent gene. 3 / 3
| 3

reach
"Consequently, UCH37 dramatically up-regulates TGF-beta-dependent gene expression by de-ubiquitinating and stabilizing the type I TGF-beta receptor."

reach
"The study hypothesized that Smad7 could act as an adaptor to recruit UCH37 to the type I TGF-beta receptor and showed that UCH37 dramatically up-regulates TGF-beta-dependent gene expression by deubiquitinating and stabilizing the type I TGF-beta receptor [XREF_BIBR]."

reach
"For example, Smad7 acts as an adaptor molecule to recruit Uch37 to the type I TGF-beta (transforming growth factor-beta) receptor, where Uch37 up-regulates TGF-beta-dependent gene expression by de-ubiquitinating and stabilizing the type I TGF-beta receptor."
UCHL5 affects RUVBL1
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMD10
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMB6
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMB5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMB3
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects PSMA2
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects MYC
| 3
UCHL5 activates MYC. 3 / 3
| 3

reach
"As expected, UCHL5 overexpression induced the increase in β-catenin and its target genes CyclinD1, C-myc, and Survivin and the decrease in cleaved-caspase3 in AN3-CA cells, which were reversed by excessive XAV939 treatment (Figures 6A–C)."

reach
"In contrast, UCHL5 overexpression in AN3-CA cells elevated the expression of β-catenin and its effector proteins (CyclinD1, C-myc, Survivin) and decreased cleaved-caspase3 (Figures 5D–F)."

reach
"UCHL5 Promotes Proliferation and Migration of Bladder Cancer Cells by Activating c-Myc via AKT/mTOR Signaling."
UCHL5 affects INO80C
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL5 affects DMAC2L
| 3
| 2

sparser
"In addition, hydrogen bonding interactions with GLN82 and HIS164 further stabilize interaction between FB-71 and the UCHL5 active site."

sparser
"xref displays the docking results of predicted interactions between FB-28 and the active site of UCHL5."
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."

reach
"This study firstly reported that UCHL5 was more highly expressed on EC tissues and cell lines and UCHL5 overexpression accelerated EC growth."

reach
"In conclusion, UCHL5 accelerated the growth of EC via the Wnt/beta-catenin pathway and was expected to be an attractive target for EC treatment."

reach
"In conclusion, our results that UCHL5 promoted the growth of EC in vivo and vitro via activating the Wnt/β-catenin signaling pathway may provide potential targets for EC control in the future."
UCHL5 affects BAP1
| 1 2
| 1 1

sparser
"These differences can rationalize why GK13S does not bind to UCHL5 and BAP1."

reach
"This suggests that these residues play functionally important roles that have yet to be defined but may include binding of specific substrates or regions of partner proteins within their respective UCH37 and BAP1 complexes."
UCHL5 binds BAP1 and ULD. 1 / 1
| 1

sparser
"UCH-L5 and BAP1 both interact with the DEUBAD domains of their binding partners via their ULDs."
UCHL5 affects AR
| 1 2
| 1 1

reach
"We found that USP14, but not UCHL5, directly binds AR protein."

sparser
"We further explored the effect of UCHL5 knockdown on AR protein level and protein interaction between UCHL5 and AR."
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
UCHL5 affects 19S
| 3
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
UCHL5 binds Bloc1s3 and 19S. 1 / 1
| 1

reach
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
TGFB affects UCHL5
| 1 2
TGFB increases the amount of UCHL5.
| 1
TGFB increases the amount of UCHL5. 1 / 1
| 1

reach
"UCH37 especially influences TGFbeta mediated transcription during the early phase of TGFbeta receptor activation."
TGFB binds UCHL5.
| 1
| 1

reach
"Smad6 is thought to function preferentially as an inhibitor of BMP, while Smad7 is a general inhibitor of TGFbeta family induced signals (mainly by binding to the TGF-beta type I receptors and preventing phosphorylation of Smads2 and 3, or recruiting ligases that degrade the TGF-beta type I and 2 complexes); however, Smad7 also may enhance TGF-beta, when binding Smad7 to the deubiquitinating enzyme UCH37 [XREF_BIBR])."
TGFB activates UCHL5.
| 1
TGFB activates UCHL5. 1 / 1
| 1

eidos
"In contrast , C-CBL , heat shock protein 90 ( Hsp90 ) , transforming growth factor-beta stimulated clone 22 ( TSC-22 ) , tumor necrosis factor receptor-associated factor 4 ( TRAF4 ) , ubiquitin-specific protease 4 ( USP4 ) , ubiquitin-specific protease 11 ( USP11 ) , ubiquitin-specific protease 15 ( USP15 ) , and UCH37 can stabilize the receptor by blocking the ubiquitination of the receptor [ 103-110 ] , thereby activating the TGF-beta signaling pathway ."
RUVBL1 affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
PSMD14 affects UCHL5, and USP14
| 3
| 3

sparser
"Among these DUBs, POH1, UCHL5 and USP14 are associated with the 19S proteasome; they are often overexpressed in several carcinoma cells, which renders them potentially new therapeutic targets in these cancer cells [ xref – xref ]."

sparser
"In addition to RPN11, UCHL5 and USP14 are also associated with cell survival and cancer progession [ xref , xref ]."

sparser
"There are three important DUBs associated with the 19S proteasome, the JAMM family member POH1 (also known as RPN11/pda1/S13/mpr1), the USP family member USP14 and the UCHs family member UCHL5 (also known as UCH37)."
PSMD10 affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
PSMB6 affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
PSMB5 affects UCHL5
3 |
3 |

No evidence text available

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No evidence text available
PSMB3 affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
PSMA2 affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
INO80C affects UCHL5
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
DMAC2L affects UCHL5
| 3
| 2

sparser
"In addition, hydrogen bonding interactions with GLN82 and HIS164 further stabilize interaction between FB-71 and the UCHL5 active site."

sparser
"xref displays the docking results of predicted interactions between FB-28 and the active site of UCHL5."
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."
BAP1 affects UCHL5
| 1 2
| 1 1

sparser
"These differences can rationalize why GK13S does not bind to UCHL5 and BAP1."

reach
"This suggests that these residues play functionally important roles that have yet to be defined but may include binding of specific substrates or regions of partner proteins within their respective UCH37 and BAP1 complexes."
UCHL5 binds BAP1 and ULD. 1 / 1
| 1

sparser
"UCH-L5 and BAP1 both interact with the DEUBAD domains of their binding partners via their ULDs."
AR affects UCHL5
| 1 2
| 1 1

reach
"We found that USP14, but not UCHL5, directly binds AR protein."

sparser
"We further explored the effect of UCHL5 knockdown on AR protein level and protein interaction between UCHL5 and AR."
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
19S affects UCHL5
| 3
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
UCHL5 binds Bloc1s3 and 19S. 1 / 1
| 1

reach
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
| 1 1
| 1 1

reach
"Docking results suggest that benzyl isothiocyanate, phenethyl isothiocyanate, and DL-sulforaphane are more potent inhibitors of UCHL5 than USP14, and these ITCs could interact with the catalytic triads of UCHL5 and USP14."

sparser
"When lysates of MDA-MB-231 cells treated with 25 µM of SFN were used in the Ub-VS assay, inhibition of both USP14 and UCHL5 by SFN was observed ( xref , lanes 3 vs ."
Pru affects UCHL5
| 2
| 2

sparser
"The two domains may become constrained by the simultaneous binding of Uch37 and hRpn13’s Pru domain to different ubiquitin moieties within a chain."

sparser
"hRpn13’s PRU binds its C-terminal Uch37-binding domain, which reduces the exposure of its ubiquitin binding surface and in turn, its affinity for ubiquitin ( xref )."
Paraquat affects UCHL5
2 |
Paraquat increases the amount of UCHL5. 2 / 2
2 |

No evidence text available

No evidence text available
Ethanol affects UCHL5
1 | 1
Ethanol decreases the amount of UCHL5. 2 / 2
1 | 1

reach
"Consistent with previous findings, we discovered that ethanol feeding decreased the levels of Ecm29 and UCHL5."

No evidence text available
2 |
Dibutyl phthalate decreases the amount of UCHL5. 2 / 2
2 |

No evidence text available

No evidence text available
2 |
Cyclosporin A increases the amount of UCHL5. 2 / 2
2 |

No evidence text available

No evidence text available
| 1 1
| 1 1

eidos
"Taken together , our data suggest that the cyclic peptide permeates the cell membrane , inhibits UCH-L5 by possibly blocking its deubiquitinating function , and contributes to the accumulation of polyubiquitinated substrates ."

reach
"Taken together, our data suggest that the cyclic peptide permeates the cell membrane, inhibits UCH-L5 by possibly blocking its deubiquitinating function, and contributes to the accumulation of polyubiquitinated substrates."
Bortezomib affects UCHL5
| 2
| 2

reach
"Notwithstanding this success, the potency of pimozide (IC 50 ~ 2muM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC 50 ~ 100nM) (XREF_FIG and XREF_TABLE), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC 50 ~ 100nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240)."

reach
"Notwithstanding this success, the potency of pimozide (IC50 ~2 μM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC50 ~100 nM) (Fig. 1 and Table 2), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC50 ~100 nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240) (ClinicalTrials.gov, 2018; Wang et al., 2016)."
Bisphenol A affects UCHL5
2 |
Bisphenol A increases the amount of UCHL5. 2 / 2
2 |

No evidence text available

No evidence text available
2 |
Benzo[a]pyrene increases the amount of UCHL5. 2 / 2
2 |

No evidence text available

No evidence text available
XPO5 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
Ubiquitin affects ADRM1
| 2
| 2

sparser
"Structures of UCH-L5-Rpn13 bound to ubiquitin are available."

sparser
"RPN13 binding to ubiquitin chains and UCH37 might simultaneously facilitate UCH37's distal end xref deubiquitylating activity xref - xref by orienting the ubiquitin moieties in a configuration favorable to hydrolysis."
UPS inhibitor affects UCHL5
| 2
UPS inhibitor inhibits UCHL5. 2 / 2
| 2

reach
"Notwithstanding this success, the potency of pimozide (IC 50~2 μM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC 50~1 00 nM) ( Fig. 1 and Table 2 ), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC 50~1 00 nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240) (ClinicalTrials.gov, 2018; Wang et al., 2016) ."

reach
"Notwithstanding this success, the potency of pimozide (IC 50 ~ 2muM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC 50 ~ 100nM) (XREF_FIG and XREF_TABLE), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC 50 ~ 100nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240)."
UCHL5 affects wnt8
| 2
UCHL5 activates wnt8. 2 / 2
| 2

reach
"Ectopic expression of Uch37 mRNA enhanced the ability of Wnt8 to form a secondary axis, whereas depletion of Uch37 by Uch37 MO efficiently inhibited it (XREF_FIG, XREF_SUPPLEMENTARY)."

reach
"Importantly, reintroduction of Uch37 WT mRNA restored the ability of Wnt8 to induce Xpo, Vent1 and Vent2 expression, whereas Uch37 IN mRNA did not (XREF_FIG lane 7 and 8)."

reach
"Furthermore, overexpression of UCH37 upregulates TGF-beta-dependent transcription, and this effect is reversed in cells subject to RNAi mediated knockdown of endogenous UCH37."

eidos
"* p < 0.05 USP14 and UCHL5 inhibitors increase the ubiquitination of ERalpha and decrease ERalpha-mediated transcription activity ."
UCHL5 affects pru
| 2
| 2

sparser
"The two domains may become constrained by the simultaneous binding of Uch37 and hRpn13’s Pru domain to different ubiquitin moieties within a chain."

sparser
"hRpn13’s PRU binds its C-terminal Uch37-binding domain, which reduces the exposure of its ubiquitin binding surface and in turn, its affinity for ubiquitin ( xref )."
| 2
| 2

reach
"The human homologue UCH37 was reported to promote Smo localization to cilia."

reach
"Uchl5 stabilizes Drosophila Smo and promotes its localization on the cell surface."
UCHL5 affects expression
| 2
UCHL5 inhibits expression. 2 / 2
| 2

sparser
"Accordingly, we found UCH-L5 inhibited mRNA expression (Figure xref and xref ) and protein level (Figure xref and xref ) of SNRPF both in U87MG cells and U251 cells significantly."

sparser
"To further confirm that UCH-L5 inhibits SNRPF expression, U87MG and U251 cells were subjected to analysis for lentivirus-mediated gene knockdown and overexpression."
UCHL5 ubiquitinates endoplasmic reticulum.
| 1
UCHL5 leads to the ubiquitination of endoplasmic reticulum. 1 / 1
| 1

reach
"We found that inhibitors of USP14 and UCHL5 dramatically increased the ubiquitinated ERα (Fig. 7d)."

reach
"Inhibition of USP14 and UCHL5 activates caspase and triggers apoptosis of ER + BCa cells."
UCHL5 affects cell death
| 2
| 2

reach
"Overexpression of UCH-L5 (but not overexpression of UCH-L3) for two days prior to the infection with Salmonella for one hour also lead to significant cell death (XREF_FIG), while cell treatment with b-AP15 inhibitor prior to Salmonella infection led to slight increase in cell viability in comparison to vehicle treated cells (XREF_SUPPLEMENTARY)."

reach
"In this report, we found that PdPT is an inhibitor of multiple DUBs including USP7, USP10, USP14, USP15, USP25 and UCHL5, which contributes to the accumulation of ubiquitinated proteins and subsequent cell death in NSCLC cell lines.Regulated cell death requires activation of various regulators and effectors (23)."
UCHL5 affects XPO5
2 |
2 |

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No evidence text available
UCHL5 affects VLX1570
| 2
UCHL5 binds VLX1570. 2 / 2
| 2

reach
"VLX1570 preferentially binds the Cys88 residue of UCHL5 and interfaces with a thiol of USP14 at residue Cys114 via a 1,4-Michael 's addition reaction, potentially forming a covalent bond."

reach
"VLX1570 binds and inhibits the activity of USP14 and UCHL5."
UCHL5 affects U2AF2
| 2
| 2

sparser
"For instance, depending on the shared target gene bound by the splicing factor U2AF2 and the protease UCHL5, completely opposite effects on TE could be observed and independently replicated (Figs xref , xref )."

sparser
"“For instance, depending on the shared target gene bound by the splicing factor U2AF2 and the protease UCHL5, completely opposite effects on TE could be observed and independently replicated (Figure 2C, Figure S2B and S2C).”"
UCHL5 affects TXNL1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects TGF-betaR1 protein
| 2
UCHL5 ubiquitinates TGF-betaR1 protein.
| 1
UCHL5 leads to the ubiquitination of TGF-betaR1 protein. 1 / 1
| 1

reach
"High glucose induced TGF-betaR1 protein expression and TGF-betaR1 protein deubiquitination were attenuated by UCHL5 shRNA."
UCHL5 decreases the amount of TGF-betaR1 protein.
| 1
UCHL5 decreases the amount of TGF-betaR1 protein. 1 / 1
| 1

reach
"High glucose induced TGF-betaR1 protein expression and TGF-betaR1 protein deubiquitination were attenuated by UCHL5 shRNA."
UCHL5 affects Smads
| 2
UCHL5 binds Smads. 2 / 2
| 2

reach
"Recently, we have defined a novel interaction between Smads and UCH37 (ubiquitin C-terminal hydrolase 37), a DUB (de-ubiquitinating enzyme) that could potentially counteract Smurf mediated ubiquitination."

reach
"Here, we report a novel interaction between Smads and ubiquitin C-terminal hydrolase UCH37, a deubiquitinating enzyme that could potentially reverse Smurf mediated ubiquitination."
UCHL5 affects Ser-Met
| 2
| 2

reach
"In stable UCH-L5 knockdown and stable UCH-L5 overexpressing U87MG cells, we found that knockdown UCH-L5 expression upregulated mRNA level of Sm genes except for SNRPN (XREF_SUPPLEMENTARY), while UCH-L5 overexpression downregulates mRNA level of Sm genes in U87MG cells (XREF_SUPPLEMENTARY)."

reach
"And we found UCH-L5 downregulated mRNA level of other Sm genes."
UCHL5 affects SKP2
| 2
| 2

reach
"We found that SKP2 interacted with USP10, USP13, UCHL5, USP14, and USP7 in KBM5-T315I cells."

reach
"GST-pulldown and co-immunoprecipitation (co-IP) assays have been applied to investigate the interactions between USP14, UCHL5, and Skp2."
UCHL5 affects SERPB12
| 1 1
UCHL5 binds SERPB12. 2 / 2
| 1 1

reach
"Alternatively, it is possible that, as reported by Fang in hepatocellular carcinoma tissues [XREF_BIBR], SERPB12 also interacts with UCHL5 directly in this pathway even though we were unable to co-precipitate the two proteins here."

sparser
"Alternatively, it is possible that, as reported by Fang in hepatocellular carcinoma tissues [ xref ], SERPB12 also interacts with UCHL5 directly in this pathway even though we were unable to co-precipitate the two proteins here."
UCHL5 affects RGPD8
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects RGPD5
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMB7
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMB4
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMB2
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMB1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMA7
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMA5
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMA4
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMA3
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PSMA1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects PRDX1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects MGRN1
| 1 1
| 1 1

reach
"In further support of this possibility, co-immunoprecipitation experiment demonstrated a physical interaction between UCHL5 and MGRN1, and a UCHL5 overexpression experiment confirmed a role of UCHL5 i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In further support of this possibility, co-immunoprecipitation experiment demonstrated a physical interaction between UCHL5 and MGRN1 (G), and a UCHL5 overexpression experiment confirmed a role of UCH[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects MBP
| 2
| 2

sparser
"Equimolar amounts of a Maltose Binding Protein (MBP)-Uch37 fusion protein and 6His-Rpn13 (1 μM) were incubated in PBS supplemented with 50 μM RA190 or vehicle control (0.5% DMSO) for 1h at 4°C. Amylose resin was added to the solution to capture MBP-Uch37."

sparser
"To assess the effect of the peptoid on these binding events, His6-Rpn13 (1 μM) was pre-incubated with KDT-11 (50 μM) or vehicle (0.5% DMSO) for one hour followed by addition of either ` or MBP-Uch37."
UCHL5 affects INO80D
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects HRPN
| 2
| 2

sparser
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."

sparser
"Thus, hRpn13 activates hINO80-associated Uch37 without displacing it from the hINO80 complex, and activation is a consequence of transient interactions between Uch37 and hRpn13."
UCHL5 affects Growth Endometrial Cancer
| 2
UCHL5 activates Growth Endometrial Cancer. 2 / 2
| 2

eidos
"Ubiquitin C-Terminal Hydrolase L5 ( UCHL5 ) Accelerates the Growth of Endometrial Cancer via Activating the Wnt / beta-Catenin Signaling Pathway Endometrial cancer ( EC ) is the most prevalent gynecological malignancy with high mortality ."

eidos
"Ubiquitin C-Terminal Hydrolase L5 ( UCHL5 ) Accelerates the Growth of Endometrial Cancer via Activating the Wnt / beta-Catenin Signaling Pathway ."
UCHL5 affects FlaG
| 2
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
UCHL5 affects FHOD1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects DRAIC
| 2
UCHL5 binds DRAIC. 2 / 2
| 2

reach
"DRAIC could increase NFRKB protein significantly instead of NFRKB mRNA and UCHL5, and bind to UCHL5."

reach
"So we predicted potential interacting molecules of DRAIC by bioinformatics analysis, and the results revealed that UCHL5 may interact with DRAIC as a binding protein."

reach
"In addition to identifying many DUBs with DDR roles, this work also lead to establish that one, UCHL5 promotes DSB end resection and HR through regulating the stability of the NFRKB protein that is a subunit of chromatin remodeling complex INO80."

reach
"UCHL5 (Ubiquitin carboxyl-terminal hydrolase isozyme L5) positively regulates DSB end resection and HR repair by deubiquitinating and stabilizing NFRKB protein (nuclear factor related to kappaB bindin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects CTSD
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects CCND1
| 2
UCHL5 activates CCND1. 2 / 2
| 2

reach
"As expected, UCHL5 overexpression induced the increase in β-catenin and its target genes CyclinD1, C-myc, and Survivin and the decrease in cleaved-caspase3 in AN3-CA cells, which were reversed by excessive XAV939 treatment (Figures 6A–C)."

reach
"In contrast, UCHL5 overexpression in AN3-CA cells elevated the expression of β-catenin and its effector proteins (CyclinD1, C-myc, Survivin) and decreased cleaved-caspase3 (Figures 5D–F)."
UCHL5 affects CCDC92
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects CCDC74B
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects CASP1
| 1 1
UCHL5 increases the amount of CASP1.
| 1
Modified UCHL5 increases the amount of CASP1. 1 / 1
| 1

reach
"The authors went on to demonstrate that UCHL5 expression increases IL-1b secretion, and increases the levels of caspase-1, a key member of the inflammasome that processes pro-IL-1b into its active form."
UCHL5 binds CASP1.
| 1
| 1

sparser
"We also showed that overexpression of UCH-L5 was associated with a significant increase in caspase-1 activity, while inhibition of UCH-L5 by selective inhibitor [ xref ] or UCH-L5 knock-down led to decrease in inflammasome-dependent IL-1β release in chicken as well as in human macrophages during infection with Salmonella and during inflammasome activation by lipopolysaccharide (LPS) and nigericin."
UCHL5 affects BIRC5
| 2
UCHL5 activates BIRC5. 2 / 2
| 2

reach
"In contrast, UCHL5 overexpression in AN3-CA cells elevated the expression of β-catenin and its effector proteins (CyclinD1, C-myc, Survivin) and decreased cleaved-caspase3 (Figures 5D–F)."

reach
"As expected, UCHL5 overexpression induced the increase in β-catenin and its target genes CyclinD1, C-myc, and Survivin and the decrease in cleaved-caspase3 in AN3-CA cells, which were reversed by excessive XAV939 treatment (Figures 6A–C)."
UCHL5 affects ASPM
| 2
| 2

sparser
"Indeed, the docking modes suggested that the ITC could interact with at least one or two amino acid residues of the catalytic triad of UCHL5, CYS-88, HIS-164, and ASP-179."

sparser
"SFN seems to only interact with ASP-179 of UCHL5 through a similar addition reaction ( xref )."
UCHL5 affects ARG1
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects AKT
| 2
UCHL5 activates AKT. 2 / 2
| 2

reach
"All things considered, our findings show that increased UCHL5 expression stimulates AKT/mTOR signaling, subsequently triggering the expression of c-Myc, SLC25A19, and ICAM5, which in turn promotes carcinogenesis in bladder cancer."

reach
"According to RNA-Seq analyses and Western blotting experiments, the expression of c-Myc, SLC25A19, and ICAM5 was modified as a result of UCHL5 activating AKT/mTOR signaling in bladder cancer cells."
UCHL5 affects ACTL6A
2 |
2 |

No evidence text available

No evidence text available
UCHL5 affects 6xHis-UIP1
| 2
UCHL5 binds 6xHis-UIP1. 2 / 2
| 2

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, FLAG-S14 (even at the 6xHis-UIP1 : FLAG-S14 molar ratio of 1:6) had no significant effect on the amount of 6xHis-UIP1 bound to GST-UCH37 (lane 6 of Fig. 4B)."
UBE2L3 affects UCHL5
| 1 1
UBE2L3 leads to the ubiquitination of UCHL5. 2 / 2
| 1 1

reach
"In addition, UbE2L3 and UbE2W slightly enhanced ubiquitination of Uch37 (XREF_FIG)."

sparser
"In addition, UbE2L3 and UbE2W slightly enhanced ubiquitination of Uch37 ( xref )."
U2AF2 affects UCHL5
| 2
| 2

sparser
"For instance, depending on the shared target gene bound by the splicing factor U2AF2 and the protease UCHL5, completely opposite effects on TE could be observed and independently replicated (Figs xref , xref )."

sparser
"“For instance, depending on the shared target gene bound by the splicing factor U2AF2 and the protease UCHL5, completely opposite effects on TE could be observed and independently replicated (Figure 2C, Figure S2B and S2C).”"
TXNL1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
Smads affects UCHL5
| 2
UCHL5 binds Smads. 2 / 2
| 2

reach
"Recently, we have defined a novel interaction between Smads and UCH37 (ubiquitin C-terminal hydrolase 37), a DUB (de-ubiquitinating enzyme) that could potentially counteract Smurf mediated ubiquitination."

reach
"Here, we report a novel interaction between Smads and ubiquitin C-terminal hydrolase UCH37, a deubiquitinating enzyme that could potentially reverse Smurf mediated ubiquitination."
SLFN12 affects UCHL5
| 2
Modified SLFN12 increases the amount of UCHL5. 2 / 2
| 2

reach
"SLFN12 or SERPB12 overexpression increases expression of the complementary deubiquitylases USP14 and UCHL5 in vitro, and SERPB12 stimulates USP14 deubiquitylase activity."

reach
"Either SLFN12 overexpression (XREF_FIG and XREF_FIG) or SERPB12 overexpression (XREF_FIG and XREF_FIG) increased both USP14 (XREF_FIG and XREF_FIG) and UCHL5 (XREF_FIG and XREF_FIG) expression."
SKP2 affects UCHL5
| 2
| 2

reach
"We found that SKP2 interacted with USP10, USP13, UCHL5, USP14, and USP7 in KBM5-T315I cells."

reach
"GST-pulldown and co-immunoprecipitation (co-IP) assays have been applied to investigate the interactions between USP14, UCHL5, and Skp2."
RPN10 affects UCHL5
| 2
RPN10 deubiquitinates UCHL5.
| 1
RPN10 deubiquitinates UCHL5. 1 / 1
| 1

reach
"RP ubiquitin receptors S5a and Rpn10 and Rpn13 capture substrates by recognizing their covalently attached ubiquitin chains, which are removed and disassembled by three deubiquitinating enzymes Rpn11, Ubp6 and Usp14 and Uch37 and UCHL5."
RPN10 binds UCHL5.
| 1
UCHL5 binds RPN10-S5A. 1 / 1
| 1

reach
"For instance, Uch37 can also associate with Rpn10 (S5a) at the proteasome complex XREF_BIBR, XREF_BIBR, as well as with the human Ino80 chromatin remodeling complex XREF_BIBR and with Smad proteins, particularly with Smad7 XREF_BIBR."
RGPD8 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
RGPD5 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
RA-190 affects UCHL5
| 2
RA-190 inhibits UCHL5. 2 / 2
| 2

reach
"XREF_BIBR, XREF_BIBR, XREF_BIBR Both G5 and RA-190 do, however, inhibit the activity of recombinant UCHL5."

reach
"96 Rpn13 is known to bind and to activate the DUB UCHL5, and it was subsequently shown that RA-190 does not affect the interaction of Rpn13 with UCHL5 but directly inactivates UCHL5."
PSMD8 affects GST
| 2
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
PSMD1 affects ADRM1
1 | 1
1 | 1

sparser
"We tested this possibility through pharmacological inhibition of the ADRM1-PSMD1-UCH37 interaction by using RA190 xref ."

No evidence text available
PSMB7 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMB4 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMB2 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMB1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMA7 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMA5 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMA4 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMA3 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PSMA1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
PRDX1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
NFRKB affects ADRM1
| 2
| 2

sparser
"Because hRpn13 and NFRKB interact with Uch37 in the proteasome and hINO80, respectively, we asked whether the isolated proteins bind to Uch37 in a mutually exclusive fashion."

sparser
"Biological information indicated that both RPN13 and INO80G bind to a long fragment at the C-terminal of the UCH-L5."

reach
"As HA-UCHL5 was expressed under a constitutive CMV promoter in a plasmid containing only the coding sequence, MG132 dependent reduction of UCHL5 was most likely a post-transcriptional regulatory effec[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
MGRN1 affects UCHL5
| 1 1
| 1 1

reach
"In further support of this possibility, co-immunoprecipitation experiment demonstrated a physical interaction between UCHL5 and MGRN1, and a UCHL5 overexpression experiment confirmed a role of UCHL5 i[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In further support of this possibility, co-immunoprecipitation experiment demonstrated a physical interaction between UCHL5 and MGRN1 (G), and a UCHL5 overexpression experiment confirmed a role of UCH[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
MBP affects UCHL5
| 2
| 2

sparser
"Equimolar amounts of a Maltose Binding Protein (MBP)-Uch37 fusion protein and 6His-Rpn13 (1 μM) were incubated in PBS supplemented with 50 μM RA190 or vehicle control (0.5% DMSO) for 1h at 4°C. Amylose resin was added to the solution to capture MBP-Uch37."

sparser
"To assess the effect of the peptoid on these binding events, His6-Rpn13 (1 μM) was pre-incubated with KDT-11 (50 μM) or vehicle (0.5% DMSO) for one hour followed by addition of either ` or MBP-Uch37."
KDT-11 affects UCHL5
| 2
KDT-11 inhibits UCHL5. 2 / 2
| 2

sparser
"We did not compare unscrambled KDT-11 given the demonstrated lack of activity in the recombinant protein assay shown in xref , however the calculated p-value for KDT-11 inhibition of Uch37 is p=0.0003."

sparser
"We used Rpn13 binding as our optimization assay because Uch37 inhibition by KDT-11 is not extremely strong."
INO80D affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
HRPN affects UCHL5
| 2
| 2

sparser
"This is similar to what was previously observed for the interaction between Uch37 and hRpn13, the proteasome subunit that binds Uch37 via the C-terminal tail to recruit it to the 19S RP."

sparser
"Thus, hRpn13 activates hINO80-associated Uch37 without displacing it from the hINO80 complex, and activation is a consequence of transient interactions between Uch37 and hRpn13."
GST affects PSMD8
| 2
GST binds UCHL5, PSMD8, and FlaG. 2 / 2
| 2

sparser
"Furthermore, the interaction of FLAG-S14 with GST-UCH37 was inhibited by of 6×His-UIP1 in a concentration-dependent manner."

sparser
"As shown in Fig. 4A , the interaction of FLAG-S14 with GST-UCH37 was inhibited by 6×His-UIP1 but not by 6×His-NonO."
FHOD1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
| 2

reach
"WP1130 is a partially selective deubiquitinase (DUB) inhibitor that inhibits the deubiquitinating activities of USP9X, USP5, USP14, and UCHL5 (UCH37) [16]."

reach
"WP1130 is a partially selective deubiquitinase (DUB) inhibitor that inhibits the deubiquitinating activities of USP9X, USP5, USP14, and UCHL5 (UCH37) [16]."
DRAIC affects UCHL5
| 2
UCHL5 binds DRAIC. 2 / 2
| 2

reach
"DRAIC could increase NFRKB protein significantly instead of NFRKB mRNA and UCHL5, and bind to UCHL5."

reach
"So we predicted potential interacting molecules of DRAIC by bioinformatics analysis, and the results revealed that UCHL5 may interact with DRAIC as a binding protein."
CTSD affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
CCDC92 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
CCDC74B affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
ASPM affects UCHL5
| 2
| 2

sparser
"Indeed, the docking modes suggested that the ITC could interact with at least one or two amino acid residues of the catalytic triad of UCHL5, CYS-88, HIS-164, and ASP-179."

sparser
"SFN seems to only interact with ASP-179 of UCHL5 through a similar addition reaction ( xref )."
ARG1 affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
ADRM1 affects Ubiquitin
| 2
| 2

sparser
"Structures of UCH-L5-Rpn13 bound to ubiquitin are available."

sparser
"RPN13 binding to ubiquitin chains and UCH37 might simultaneously facilitate UCH37's distal end xref deubiquitylating activity xref - xref by orienting the ubiquitin moieties in a configuration favorable to hydrolysis."
ADRM1 affects PSMD1, and UCHL5
1 | 1
1 | 1

sparser
"We tested this possibility through pharmacological inhibition of the ADRM1-PSMD1-UCH37 interaction by using RA190 xref ."

No evidence text available
ADRM1 affects NFRKB, and UCHL5
| 2
| 2

sparser
"Because hRpn13 and NFRKB interact with Uch37 in the proteasome and hINO80, respectively, we asked whether the isolated proteins bind to Uch37 in a mutually exclusive fashion."

sparser
"Biological information indicated that both RPN13 and INO80G bind to a long fragment at the C-terminal of the UCH-L5."
ACTL6A affects UCHL5
2 |
2 |

No evidence text available

No evidence text available
6xHis-UIP1 affects UCHL5
| 2
UCHL5 binds 6xHis-UIP1. 2 / 2
| 2

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, FLAG-S14 (even at the 6xHis-UIP1 : FLAG-S14 molar ratio of 1:6) had no significant effect on the amount of 6xHis-UIP1 bound to GST-UCH37 (lane 6 of Fig. 4B)."
Vs affects Ubiquitin
| 1
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
Unknown mechanism affects UCHL5
| 1
Unknown mechanism activates UCHL5. 1 / 1
| 1

eidos
"S. Typhimurium upregulates the expression of UCH-L5 by an unknown mechanism and causes increased catalytic activity of caspase-1 , which limits the IL-1beta level ."
| PMC
Ubiquitin receptors affects UCHL5
| 1
Ubiquitin receptors-S5A deubiquitinates UCHL5. 1 / 1
| 1

reach
"RP ubiquitin receptors S5a and Rpn10 and Rpn13 capture substrates by recognizing their covalently attached ubiquitin chains, which are removed and disassembled by three deubiquitinating enzymes Rpn11, Ubp6 and Usp14 and Uch37 and UCHL5."
Trimellitic anhydride increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Trichostatin A increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Sodium(1+) affects UCHL5
| 1
| 1

reach
"Additional deleted genes in the region include multiple members of the Regulator of G protein Signaling Gene family (RGS1, RGS2, RGS13, and RGS18) as well as TROVE Domain Family Member 2, Ubiquitin C-terminal Hydrolase L5, and portions of Potassium Sodium Activated Channel Subfamily Member 2 (TROVE2, UCHL5, and KCNT2)."
1 |
Sodium arsenite increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
ShRNA interference technology affects UCHL5
| 1
ShRNA interference technology inhibits UCHL5. 1 / 1
| 1

eidos
"Lentiviral Vector Construction and Cell Transfection Lentiviral-mediated shRNA interference technology was used to downregulate the UCHL5 gene in HCE-1-A cells ."
Rpn12 affects UCHL5
| 1
| 1

reach
"Previously Uch2 and Uch37 has been shown to interact with Mts3 and Rpn12 26 and to localise to the " hinge region ", between the base and lid subcomplexes of the 19 S regulatory complex of the 26 S pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Proteasome ubiquitin affects UCHL5
| 1
Proteasome ubiquitin activates UCHL5. 1 / 1
| 1

reach
"The essential cysteine protease UCH-L5 is activated by proteasome ubiquitin receptor RPN13 (ADRM1) or inhibited by chromatin remodeling complex component INO80 (NFRKB) [XREF_BIBR]."
Proteasome inhibitor affects UCHL5
| 1
Proteasome inhibitor activates UCHL5. 1 / 1
| 1

reach
"Notwithstanding this success, the potency of pimozide (IC 50 ~ 2muM) is lower than those of clinically approved UPS inhibitor 26S proteasome inhibitor Bortezomib (IC 50 ~ 100nM) (XREF_FIG and XREF_TABLE), and another UPS inhibitor VLX1570, which inhibited the DUBs UCHL5 and USP14 (IC 50 ~ 100nM) and was previously studied in clinical trials but terminated due to limiting toxicities (study identifier NCT02372240)."
Proteasomal DEUBAD-containing protein Rpn13 affects UCHL5
| 1
Proteasomal DEUBAD-containing protein Rpn13 activates UCHL5. 1 / 1
| 1

eidos
"UCHL5 carries out deubiquitination at the 26S proteasome [ 19,20,21,22 ] when activated by the proteasomal DEUBAD-containing protein Rpn13 ."
Piperidones affects UCHL5
| 1

reach
"From these data, we can predict that the piperidones being investigated bind to deubiquitinases UCHL5 and USP14."
| 1

reach
"We here explored the phenotypic response of colon cancer cells to b-AP15, a bis-benzylidine piperidone previously shown to inhibit the proteasome deubiquitinases (DUBs) USP14 and UCHL5."
1 |

No evidence text available
Phlorizin affects UCHL5
1 |
Phlorizin decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Paclitaxel affects UCHL5
1 |
Paclitaxel increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
P2 affects USP14
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
Nucleus affects UCHL5
| 1
| 1

eidos
"In addition , inhibiting hnRNPA1 prevented apoptosis in PC-12 cells subjected to OGD / R. hnRNPA1 bound to CRNDE and remained in the nucleus , which inhibited UCHL5 expression through the formation of CRNDE-hnRNPA1-mRNA complex ."
Nickel(2+) affects UCHL5
| 1
| 1

reach
"This novel nickel complex can also target proteasomal DUBs (UCHL5 and USP14) and induce apoptosis in cancer cells [XREF_BIBR]."
Monoubiquitin affects UCHL5
| 1
ADRM1 binds UCHL5 and monoubiquitin. 1 / 1
| 1

sparser
"Additionally, there has been recent development of another competitive FP assay that includes fluorescently-labeled monoubiquitin bound to the Uch37:Rpn13 complex and then competed off with unlabeled ubiquitin chains."
Methylmercury compound increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
| 1

sparser
"We previously reported that phenethyl isothiocyanate (PEITC) inhibits two deubiquitinases (DUBs), USP9x and UCH37."
1 |
Hydroperoxide increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6872-3p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Hsa-miR-657 affects UCHL5
1 |
Hsa-miR-657 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6512-5p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-5584-5p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-5581-3p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-539-3p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-485-3p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4755-5p decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3665 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Glucose affects UCHL5
| 1
Glucose increases the amount of UCHL5. 1 / 1
| 1

reach
"High glucose also increased UCHL5 protein expression, which was attenuated by LY294002, the dominant negative p85 and the dominant negative CREB."
Genistein affects UCHL5
1 |
Genistein increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Finasteride affects UCHL5
1 |
Finasteride increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Ethyl methanesulfonate decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Dorsomorphin increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1 |
Disodium selenite increases the amount of UCHL5. 1 / 1
1 |

No evidence text available

reach
"Control cells treated with b-AP15 inhibitor of UCH-L5 had lower caspase-1 activity than control cells treated with vehicle control (DMSO; XREF_FIG)."
Dimerization.9 USP14 affects UCHL5
| 1
Dimerization.9 USP14 inhibits UCHL5. 1 / 1
| 1

eidos
"Rpn13 , the proteasomal receptor for Uch37 in the proteasome 19S regulatory particle , can activate UCH37 by disrupting dimerization.9 USP14 binds to the regulatory particle Rpn1 to release its catalytic USP domain and polyubiquitin chains of substrate protein.10 Overexpression of Rpn13 or Rpn1 upregulated the COPS5 protein levels and downregulated the p53 protein levels ( Supplementary Fig. S2p ) ."
Dicrotophos affects UCHL5
1 |
Dicrotophos decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Diazinon affects UCHL5
1 |
Diazinon methylates UCHL5. 1 / 1
1 |

No evidence text available
Diarsenic trioxide increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Deubiquitinating function affects UCHL5
| 1
Deubiquitinating function activates UCHL5. 1 / 1
| 1

eidos
"Taken together , our data suggest that the cyclic peptide permeates the cell membrane , inhibits UCH-L5 by possibly blocking its deubiquitinating function , and contributes to the accumulation of polyubiquitinated substrates ."
Deubiquitinase adaptor domain affects UCHL5
| 1
UCHL5 binds deubiquitinase adaptor domain. 1 / 1
| 1

reach
"Uch37 and UCHL5 binds to a deubiquitinase adaptor domain (DEUBAD) of Rpn13."
Curcumin affects UCHL5
| 1
| 1

reach
"USP14 and UCH37 are also inhibited by curcumin analogue AC17."
Coumestrol affects UCHL5
1 |
Coumestrol increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Catalytic conformation affects UCHL5
| 1
Catalytic conformation activates UCHL5. 1 / 1
| 1

eidos
"To explain regulatory mechanisms , Ub-Prg was used to capture the catalytic conformation by which RPN13DEUBAD activates UCH-L5 ; and the catalytic conformation that truncated INO80DEUBAD lacks to inhibit UCH-L5 ."
1 |
Carbon nanotube increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
B-AP15 is therapeutic molecule affects UCHL5
| 1
B-AP15 is therapeutic molecule inhibits UCHL5. 1 / 1
| 1

eidos
"B-AP15 is a small therapeutic molecule that inhibits USP14 and UCHl5 [ 75 ] ."
Auranofin affects 19S
| 1
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
Aumdubin affects UCHL5
| 1
Aumdubin inhibits UCHL5. 1 / 1
| 1

reach
"One potential shortcoming of aumdubin is that it also inhibits other DUBs, including USP14, USP10, and UCHL5."
| 1
| 1

reach
"To determine whether DUBs of the 26S proteasome regulate proteasomal ubiquitination, we set up in vitro ubiquitination reactions in which Ub aldehyde (an inhibitor of both Usp14 and Uch37) or 1,10-phenanthroline (an inhibitor of Rpn11) was added in the reactions."
ZEB1 affects UCHL5
1 |
ZEB1 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
ZBED1 affects UCHL5
1 |
1 |

No evidence text available
YTHDF1 affects UCHL5
| 1
| 1

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
YP_009227200 affects UCHL5
1 |
UCHL5 binds YP_009227200. 1 / 1
1 |

No evidence text available
Ubiquitin affects vs
| 1
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
Ubiquitin affects TYMS
| 1
| 1

sparser
"Out of these ten proposed residues S10, E31, Y33, I214, F228, and L230, positions were chosen based on Ts UCHL5ubiquitin interactions."
Ubiquitin affects PSMD1
| 1
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
USP5 affects USP14
| 1
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
USP28 affects UCHL5
1 |
1 |

No evidence text available
USP14-3 affects UCHL5
| 1
USP14-3 activates UCHL5. 1 / 1
| 1

reach
"Inhibition of deubiquitinating activity by USP14 aptamers was specific to USP14, with the USP14-3 aptamer failing to inhibit the DUB activity of UCHL3, USP47, USP5, and UCHL5 and UCH37 (XREF_FIG)."
USP14 affects vs
| 1
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
USP14 affects p2
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
USP14 affects USP5
| 1
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
USP14 affects RPN1
| 1
RPN1 binds USP14 and UCHL5. 1 / 1
| 1

sparser
"Interestingly, USP14 and UCH37 bind to RPN1 and to RPN13, respectively, which are, in addition to the RPN10, substrate receptors of the 19S regulatory particle [ xref ], which provide a versatile binding platform for various ubiquitin chains [ xref ]."
USP14 affects PTPN2
| 1
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
USP14 affects PRF1
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
USP14 affects DMAC2L
| 1
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."
USP14 affects AR
| 1
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
ULD affects BAP1
| 1
UCHL5 binds BAP1 and ULD. 1 / 1
| 1

sparser
"UCH-L5 and BAP1 both interact with the DEUBAD domains of their binding partners via their ULDs."
UCHL5 affects vs
| 1
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
UCHL5 affects translation
| 1
| 1

reach
"In order to understand the endogenous function of Uch37 in Xenopus, as suggested by its local enrichment in the mesodermal region of the embryo, we knocked down endogenous Uch37 by injecting antisense morpholino oligonucleotides (Uch37 MO) that specifically and efficiently blocked translation of Uch37 protein (XREF_SUPPLEMENTARY)."

trips
"In addition, we show that UCH37 can deubiquitinate and stabilize the type I TGF-beta receptor."
UCHL5 affects rpn12
| 1
| 1

reach
"Previously Uch2 and Uch37 has been shown to interact with Mts3 and Rpn12 26 and to localise to the " hinge region ", between the base and lid subcomplexes of the 19 S regulatory complex of the 26 S pr[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects repair
| 1
UCHL5 activates repair. 1 / 1
| 1

eidos
"The function of UCHL1 in DSB repair is still unclear , but it has been established that UCHL5 promotes repair by enhancing DNA-end resection , whereas BAP1 promotes HR via an unknown mechanism27 ,28 ."
UCHL5 affects protein COPS5
| 1
UCHL5 activates protein COPS5. 1 / 1
| 1

eidos
"We further found that UCH37 significantly upregulated the protein level of COPS5 , but there is no obvious change for other candidate interacting protein , including RPN10 , HAUS7 , and RPN13 ( Supplementary Fig. S1h , i ) ."
UCHL5 affects prenatal lethality mice
| 1
UCHL5 inhibits prenatal lethality mice. 1 / 1
| 1

eidos
"Depletion of the UCH family member , UCHL5 , causes prenatal lethality in mice , with disorganized neuronal growth most evident in the areas of the forebrain , cerebellum , and midbrain65 ."
UCHL5 affects polyUb chain
| 1
UCHL5 activates polyUb chain. 1 / 1
| 1

reach
"For instance, the two proteasome associating DUBs Usp14 and Uch37 catalyze polyUb chain trimming by shortening polyUb chains, which could either promote or inhibit proteasomal degradation."
UCHL5 affects piperidones
| 1

reach
"From these data, we can predict that the piperidones being investigated bind to deubiquitinases UCHL5 and USP14."
UCHL5 affects p2
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
UCHL5 affects monoubiquitin
| 1
ADRM1 binds UCHL5 and monoubiquitin. 1 / 1
| 1

sparser
"Additionally, there has been recent development of another competitive FP assay that includes fluorescently-labeled monoubiquitin bound to the Uch37:Rpn13 complex and then competed off with unlabeled ubiquitin chains."
UCHL5 affects hydrolysis model proteasome substrates
| 1
UCHL5 inhibits hydrolysis model proteasome substrates. 1 / 1
| 1

eidos
"Depletion of UCH37 leads to accelerated hydrolysis of model proteasome substrates with a concomitant decrease in the levels of cellular polyubiquitinated proteins ( 59 ) ."
UCHL5 affects growth endometrial cancer cells
| 1
UCHL5 activates growth endometrial cancer cells. 1 / 1
| 1

eidos
"These results indicated that upregulation of UCHL5 expression contributed to the growth of endometrial cancer cells ."
UCHL5 affects deubiquitinase adaptor domain
| 1
UCHL5 binds deubiquitinase adaptor domain. 1 / 1
| 1

reach
"Uch37 and UCHL5 binds to a deubiquitinase adaptor domain (DEUBAD) of Rpn13."
UCHL5 affects beta-Catenin Signaling Pathway
| 1
UCHL5 activates beta-Catenin Signaling Pathway. 1 / 1
| 1

eidos
"Ubiquitin C-Terminal Hydrolase L5 ( UCHL5 ) Accelerates the Growth of Endometrial Cancer via Activating the Wnt / beta-Catenin Signaling Pathway ."
UCHL5 affects b-AP15
| 1
UCHL5 activates b-AP15. 1 / 1
| 1

reach
"We confirmed platelets contained both USP14 and UCHL5 targets of b-AP15 by recovering platelet proteasomes by high speed centrifugation, and then western blotting proteins of the isolated proteasome (XREF_FIG)."
UCHL5 affects auranofin
| 1
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
UCHL5 affects alpha-SMA
| 1
UCHL5 increases the amount of alpha-SMA. 1 / 1
| 1

reach
"As shown in XREF_FIG, UCHL5 knockdown attenuated expression of FN and alpha-SMA induced by TGFbeta-1, which is consistent with b-AP15 treatment in XREF_FIG."
UCHL5 affects ZBED1
1 |
1 |

No evidence text available
UCHL5 affects YTHDF1
| 1
| 1

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
UCHL5 affects YP_009227200
1 |
UCHL5 binds YP_009227200. 1 / 1
1 |

No evidence text available
UCHL5 affects Wnt beta-catenin signaling
| 1
UCHL5 activates Wnt beta-catenin signaling. 1 / 1
| 1

eidos
"UCHL5 activated Wnt / beta-catenin signaling and affected the expression of its target genes ."
UCHL5 affects USP5
| 1
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
UCHL5 affects USP28
1 |
1 |

No evidence text available
UCHL5 affects USP14, Ubiquitin, and vs
| 1
| 1

sparser
"Ub-VS, a potent and irreversible inhibitor of UCHL5 and USP14 (and other DUBs), is able to bind to the active site, shifting UCHL5 protein from 37 to 45 kD, or USP14 protein from 60 to 70 kD; in the presence of an inhibitor to such a DUB, the super-shifts can be inhibited ( xref )."
UCHL5 affects USP14, and USP5
| 1
USP14 binds USP5 and UCHL5. 1 / 1
| 1

sparser
"In S63del, increased amounts of the deubiquitinating enzymes USP14, UCH37, and USP5 were associated with proteasomes, the first time this has been reported in a human disease model."
UCHL5 affects USP10, USP14, USP15, USP25, and USP7
| 1
| 1

sparser
"PdPT (2.5 and 25 μM) was able to compete with HA-Ub-VS for binding to USP14, UCHL5, USP15, USP10, USP7, and USP25 like b-AP15, but not OTUB1 and OTUD1 ( xref )."
UCHL5 affects UCHL5
| 1
UCHL5 inhibits UCHL5. 1 / 1
| 1

reach
"Overexpression of UCH37 upregulates TGF-β-dependent transcription which is reversed in cells subject to RNAi-mediated knockdown of UCH37 [21]."
UCHL5 affects UCH domain
| 1
ADRM1 binds UCHL5 and UCH domain. 1 / 1
| 1

reach
"Binding of Adrm1 to UCH37 relieved auto-inhibition by the UCH domain and activated its deubiquitination activity [XREF_BIBR]."
UCHL5 affects UBP6
| 1
| 1

sparser
"Two other deubiquitinating enzymes associated with the proteasome, named Ubp6 and Uch37 in yeast, trim ubiquitin chains sequentially from the distal end xref , xref ."
UCHL5 affects TbetaR complex
| 1
UCHL5 activates TbetaR complex. 1 / 1
| 1

reach
"In contrast, UCH37 demonstrated a minimal increase in CAGA-Luc activity suggesting that UCH37 may also target the TGF-beta pathway downstream of the TbetaR complex to regulate overall TGF-beta signaling (XREF_SUPPLEMENTARY)."
UCHL5 affects TYMS
| 1
| 1

sparser
"Out of these ten proposed residues S10, E31, Y33, I214, F228, and L230, positions were chosen based on Ts UCHL5ubiquitin interactions."
UCHL5 affects TXN2
1 |
1 |

No evidence text available
UCHL5 affects TRIM63
1 |
1 |

No evidence text available
UCHL5 affects TRIM55
1 |
1 |

No evidence text available
UCHL5 affects TRIB2
| 1
UCHL5 inhibits TRIB2. 1 / 1
| 1

reach
"However, neither the DUB inhibitor VLX1570 or EOAI nor the depletion of key DUBs, USP14, and UCH37 30, abolished the effect of TRIB2 in Bel-7402 and SMMC-7721 cells, excluding the possibility that TRIB2 regulates Ub via these DUBs."
UCHL5 affects TP53
| 1
Modified UCHL5 increases the amount of TP53. 1 / 1
| 1

reach
"We found that p53 protein level was rescued after knockdown of COPS5 and induced by UCH37 and USP14 overexpression."
UCHL5 affects TIPRL
1 |
1 |

No evidence text available
UCHL5 affects THRSP
| 1
| 1

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects TGM3
1 |
1 |

No evidence text available
UCHL5 affects TDP2
| 1
| 1

reach
"Thus, we observe strong association between UCH37 and TDP2 : molecules targeting one protein have a reasonable chance to target the other as an off target."
UCHL5 affects Smurf
| 1
| 1

reach
"UCHL5 interacts with Smads and reverses Smurf mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-beta receptor in cells [XREF_BIBR]."
UCHL5 affects SMURF
| 1
UCHL5 leads to the deubiquitination of SMURF. 1 / 1
| 1

reach
"Likely, UCH37 is found to connect to Smad7 and reverse Smurf-mediated ubiquitination [114]."
UCHL5 affects SLFN12
| 1
UCHL5 activates SLFN12. 1 / 1
| 1

reach
"This, in turn, represents a key mechanism by which these proteins influence enterocytic differentiation since the SLFN12 effects are blocked by pharmacologic inhibition or molecular reduction of the deubuiquitylases USP14 and UCHL5."
UCHL5 affects SERPINB3
1 |

No evidence text available
UCHL5 affects SERPINB12
1 |

No evidence text available
UCHL5 affects S14
| 1
UCHL5 binds HAUS7 and S14. 1 / 1
| 1

reach
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL5 affects S100A7
1 |
1 |

No evidence text available
UCHL5 affects S100A16
1 |
1 |

No evidence text available
UCHL5 affects RPS21
1 |
1 |

No evidence text available
UCHL5 affects RPN13 subunit whose C-terminal domain
| 1
UCHL5 binds RPN13 subunit whose C-terminal domain. 1 / 1
| 1

reach
"This activity requires the RPN13 subunit whose C-terminal domain binds UCH-L5."
UCHL5 affects RPN10
| 1
UCHL5 binds RPN10-S5A. 1 / 1
| 1

reach
"For instance, Uch37 can also associate with Rpn10 (S5a) at the proteasome complex XREF_BIBR, XREF_BIBR, as well as with the human Ino80 chromatin remodeling complex XREF_BIBR and with Smad proteins, particularly with Smad7 XREF_BIBR."
UCHL5 affects RPN1
| 1
RPN1 binds USP14 and UCHL5. 1 / 1
| 1

sparser
"Interestingly, USP14 and UCH37 bind to RPN1 and to RPN13, respectively, which are, in addition to the RPN10, substrate receptors of the 19S regulatory particle [ xref ], which provide a versatile binding platform for various ubiquitin chains [ xref ]."
UCHL5 affects RNF20
1 |
1 |

No evidence text available
UCHL5 affects RI
| 1
UCHL5 deubiquitinates RI. 1 / 1
| 1

reach
"UCH37, a member of DUBs, can deubiquitinate and stabilize the TβRI which is targeted for ubiquitylation-mediated degradation."
UCHL5 affects RFFL
1 |
1 |

No evidence text available
UCHL5 affects RAD23B
1 |
1 |

No evidence text available
UCHL5 affects RACK1
| 1
| 1

sparser
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation ( xref ), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 ( xref )."
UCHL5 affects RAB7A
1 |
1 |

No evidence text available
UCHL5 affects Q1K9H5
1 |
1 |

No evidence text available
| 1

reach
"The increased expression of UCH37 decreases the polyubiquitin of TCF7."
UCHL5 affects PTPN2
| 1
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
UCHL5 affects PSMB9
1 |
1 |

No evidence text available
UCHL5 affects PSMB8
1 |
1 |

No evidence text available
UCHL5 affects PRF1
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
UCHL5 affects POTEF
1 |
1 |

No evidence text available
UCHL5 affects POTEE
1 |
1 |

No evidence text available
UCHL5 affects POF1B
1 |
1 |

No evidence text available
UCHL5 affects PNP
1 |
1 |

No evidence text available
UCHL5 affects PLRG1
1 |
1 |

No evidence text available
UCHL5 affects PLA2G2A
1 |
1 |

No evidence text available
UCHL5 affects PKP1
1 |
1 |

No evidence text available
UCHL5 affects PJA1
1 |
1 |

No evidence text available
UCHL5 affects OTX015-R
| 1
UCHL5 inhibits OTX015-R. 1 / 1
| 1

reach
"Furthermore, cyclin-dependent kinase 4/6 (CDK4/6) inhibition decreased UCHL5 expression, suppressed OTX015-R cell proliferation, and induced apoptosis."

reach
"And the above findings suggest that UCH-L5 may inhibit occurrence and metastasis of gliomas."
UCHL5 affects NFKBIA
1 |
UCHL5 deubiquitinates NFKBIA. 1 / 1
1 |

"Furthermore, we discovered that IkappaB-alpha, a protein whose proteasomal degradation activates the transcription factor NF-kappaB, is also a substrate for the Rpn13/UCH37 complex."
UCHL5 affects NACA
1 |
1 |

No evidence text available
UCHL5 affects Mice
| 1
UCHL5 activates Mice. 1 / 1
| 1

reach
"Treatment with b-AP15, a UCHL5 and USP14 deubiquitinating activity inhibitor in 19S regulatory subunit, induces tumor regression and prolong the survival period of tumor-loaded mice through down-regulation of COPS5 and its downstream AP-1 and E2F1, and up-regulation of the cell cycle-related proteins p27 and Cyclin E1."
UCHL5 affects MTOR
| 1
UCHL5 activates MTOR. 1 / 1
| 1

reach
"All things considered, our findings show that increased UCHL5 expression stimulates AKT/mTOR signaling, subsequently triggering the expression of c-Myc, SLC25A19, and ICAM5, which in turn promotes carcinogenesis in bladder cancer."
UCHL5 affects METTL14
| 1
| 1

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
UCHL5 affects LMNA
1 |
1 |

No evidence text available
UCHL5 affects KRT9
1 |
1 |

No evidence text available
UCHL5 affects KRT73
1 |
1 |

No evidence text available
UCHL5 affects KRT6B
1 |
1 |

No evidence text available
UCHL5 affects KRT28
1 |
1 |

No evidence text available
UCHL5 affects KRT12
1 |
1 |

No evidence text available
UCHL5 affects KRT10
1 |
1 |

No evidence text available
UCHL5 affects KDM1A
1 |
1 |

No evidence text available
UCHL5 affects KDELR1
1 |
1 |

No evidence text available
UCHL5 affects K48- linked polyubiquitin chains
| 1
UCHL5 activates K48- linked polyubiquitin chains. 1 / 1
| 1

reach
"Because our results suggested the possibility that Uch37 enhances resistance of Tcf7 to proteasomal degradation by removing K48- linked polyubiquitin chains from Tcf7, despite its removal activity on K63- linked polyubiquitin chains, we next examined if Uch37 stabilizes Tcf7 protein."
UCHL5 affects Ile-Leu
| 1
UCHL5 increases the amount of Ile-Leu. 1 / 1
| 1

reach
"Whether USP7 and USP47 directly interact with NLRP3 receptor or with other inflammasome component was not investigated.Enzymatic activity of UCHL5 (from the ubiquitin C‐terminal hydrolase family) is also increased after Salmonella enterica serovar Typhimurium infection, and UCHL5 silencing leads to impaired release of IL‐1β after infection 57."
UCHL5 affects IVL
1 |
1 |

No evidence text available
UCHL5 affects IGHG1
1 |
1 |

No evidence text available
UCHL5 affects Herc4-mediated ubiquitination Smo
| 1
UCHL5 inhibits Herc4-mediated ubiquitination Smo. 1 / 1
| 1

eidos
"To test whether USP8 , UCHL5 attenuate Herc4-mediated ubiquitination of Smo , we did the western blot experiments ."
UCHL5 affects Hedgehog
| 1
| 1

reach
"The deubiquitinase UCHL5 and UCH37 positively regulates Hedgehog signaling by deubiquitinating Smoothened."
UCHL5 affects HSPB1
1 |
1 |

No evidence text available
UCHL5 affects HMBS
| 1
UCHL5 inhibits HMBS. 1 / 1
| 1

reach
"Most importantly, it was shown that loss of uchl5, the human orthologue of ubh-4, also increases UPS activity and the degradation of proteotoxic proteins in mammalian cells XREF_BIBR, thus further verifying the value of C. elegans as a model organism to evaluate the consequences of UPS modulation."
UCHL5 affects HERC4
| 1
| 1

reach
"Herc4 interacts with USP8, UCHL5, and Hrs to regulate Smo ubiquitination."
UCHL5 affects HBA2
1 |
1 |

No evidence text available
UCHL5 affects H4C6
1 |
1 |

No evidence text available
UCHL5 affects H4C4
1 |
1 |

No evidence text available
UCHL5 affects H4C3
1 |
1 |

No evidence text available
UCHL5 affects H4C15
1 |
1 |

No evidence text available
UCHL5 affects H4C13
1 |
1 |

No evidence text available
UCHL5 affects H4C11
1 |
1 |

No evidence text available
UCHL5 affects H4C1
1 |
1 |

No evidence text available
UCHL5 affects Glioma
| 1
UCHL5 inhibits Glioma. 1 / 1
| 1

reach
"In vitro analysis revealed that UCHL5 can inhibit migration and invasion of glioma cells mediated via a downregulation of SNRPF [142]."
UCHL5 affects Flag
| 1
| 1

sparser
"On the one hand, the introduction of either Flag-UCHL5 or non-tagged UCHL5 into HeLa cells remarkably enhanced myc-Axin1 and endogenous Axin1 (Fig.  xref a,b)."
UCHL5 affects FTL
1 |
1 |

No evidence text available
UCHL5 affects FN1
| 1
UCHL5 decreases the amount of FN1. 1 / 1
| 1

reach
"Additionally, high glucose induced p21 (WAF1), fibronectin protein expression and cell hypertrophy were attenuated by UCHL5 shRNA."
UCHL5 affects FLG
1 |
1 |

No evidence text available

reach
"In conclusion, UCHL5 accelerated the growth of EC via the Wnt and beta-catenin pathway and was expected to be an attractive target for EC treatment."
UCHL5 affects Endometrial Cancer Cells
| 1
UCHL5 activates Endometrial Cancer Cells. 1 / 1
| 1

eidos
"Upregulation of UCHL5 Promoted the Growth of Endometrial Cancer Cells Since the expression of UCHL5 was relatively lower in AN3-CA endometrial cancer cells , we increased the expression of UCHL5 by lentiviral infection in AN3-CA cells ."
UCHL5 affects ER
| 1
UCHL5 inhibits ER. 1 / 1
| 1

reach
"To further investigate whether the ESIs previously reported effect on protein translocation contributed to its inhibitory effect on IL-1 release or whether it was due to an inhibition of DUBs, we investigated the effects of a selective DUB inhibitor b-AP15 and of the protein translocation inhibitor cpd A. b-AP15 is a small molecule DUB inhibitor of the proteasome associated DUBs UCH37 and USP14, whereas cpd A is a selective inhibitor of the ER translocon with no effect on DUB activity."
UCHL5 affects EEF1G
1 |
1 |

No evidence text available
UCHL5 affects ECPAS
1 |
1 |

No evidence text available
UCHL5 affects DUSP14
1 |
1 |

No evidence text available
UCHL5 affects DSP
1 |
1 |

No evidence text available
UCHL5 affects DSC3
1 |
1 |

No evidence text available
UCHL5 affects DSC1
1 |
1 |

No evidence text available
UCHL5 affects DNA resection
| 1
UCHL5 inhibits DNA resection. 1 / 1
| 1

eidos
"UCHL5 contributes to DNA end resection by recruiting EXO1 to DSBs [ 21 ] ."
UCHL5 affects Cyclin
| 1
UCHL5 increases the amount of Cyclin. 1 / 1
| 1

reach
"However, UCHL5 knockdown only blocked the expression of Cyclin D1 and CDK4 in A549 cells, suggesting that the effects of UCHL5 on cell cycle proteins is cell-type specific."
UCHL5 affects CuPT
| 1
UCHL5 inhibits CuPT. 1 / 1
| 1

reach
"However, in HepG2 cancer cells, either UCHL5 or USP14 knockdown did not abrogate CuPT mediated ubiquitinated protein accumulation."
UCHL5 affects Caspase
| 1
| 1

reach
"Inhibition of USP14 and UCHL5 activates caspase and triggers apoptosis of ER + BCa cells."
UCHL5 affects CYS1
| 1
| 1

sparser
"The ITC compounds may interact stronger with UCHL5 than to USP14 since more favorable bonding interactions were found upon analyzing the general decrease of free energy across the three compounds corresponding to xref . xref suggests interaction of BITC with the active site CYS-88 of UCHL5 through two formed bonds."
UCHL5 affects CYB5A
1 |
1 |

No evidence text available
UCHL5 affects CSTA
1 |
1 |

No evidence text available
UCHL5 affects COMMD4
1 |
1 |

No evidence text available
UCHL5 affects CHI3L1
| 1
| 1

sparser
"Uch37 could form oligomers in solution."
UCHL5 affects CFL1
1 |
1 |

No evidence text available
UCHL5 affects CDSN
1 |
1 |

No evidence text available
UCHL5 affects Bloc1s3
| 1
UCHL5 binds Bloc1s3 and 19S. 1 / 1
| 1

reach
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."
UCHL5 affects BAX
| 1
UCHL5 inhibits BAX. 1 / 1
| 1

reach
"Similarly, knockdown of UCH37 and USP14 or b-AP15 treatment rescued p53 protein, induced by COPS5 overexpression, and enhanced the activity of transfected luciferase reporter plasmids for p53, Bax, and p21 expression and protein levels of p53 downstream target genes BAX and p21."
UCHL5 affects ARMT1
1 |
1 |

No evidence text available
UCHL5 affects APOB
1 |
1 |

No evidence text available
UCHL5 affects APOA1
1 |
1 |

No evidence text available
UCHL5 affects ANXA8
1 |
1 |

No evidence text available
UCHL5 affects ANXA1
1 |
1 |

No evidence text available
UCHL5 affects ANP32B
1 |
1 |

No evidence text available
UCHL5 affects ACTC1
1 |
1 |

No evidence text available
UCHL5 affects 2c-biotin
| 1
UCHL5 binds 2c-biotin. 1 / 1
| 1

reach
"Figure XREF_FIG confirms that in vivo 2c-biotin binds with comparable intensity USP14 and UCH-L5."
UCHL5 affects 293T
| 1
UCHL5 binds COPS5 and 293T. 1 / 1
| 1

reach
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA or anti-Flag antibody."
UCHL5 affects (ULD) domain
| 1
UCHL5 binds RPN13 and (ULD) domain. 1 / 1
| 1

reach
"Rpn13 also binds to the C-terminal UCH37 like domain (ULD) domain of the UCH, locking the ULD into a favorable conformation for ubiquitin binding."
UCH domain affects UCHL5
| 1
ADRM1 binds UCHL5 and UCH domain. 1 / 1
| 1

reach
"Binding of Adrm1 to UCH37 relieved auto-inhibition by the UCH domain and activated its deubiquitination activity [XREF_BIBR]."
UBP6 affects UCHL5
| 1
| 1

sparser
"Two other deubiquitinating enzymes associated with the proteasome, named Ubp6 and Uch37 in yeast, trim ubiquitin chains sequentially from the distal end xref , xref ."
UBE3C affects UCHL5
| 1
UBE3C ubiquitinates UCHL5. 1 / 1
| 1

sparser
"It was reported that ubiquitination of Rpt5, Rpn10/S5A, Rpn13/ADRM1, and UCHL5 is induced by proteasome-associated UBE3A and UBE3C at the purified proteasome [ xref ]."
TYMS affects Ubiquitin
| 1
| 1

sparser
"Out of these ten proposed residues S10, E31, Y33, I214, F228, and L230, positions were chosen based on Ts UCHL5ubiquitin interactions."
TXN2 affects UCHL5
1 |
1 |

No evidence text available
TRIM63 affects UCHL5
1 |
1 |

No evidence text available
TRIM55 affects UCHL5
1 |
1 |

No evidence text available
TIPRL affects UCHL5
1 |
1 |

No evidence text available
THRSP affects UCHL5
| 1
| 1

reach
"Bands of ~ 1000 bp with approximately equal intensity were amplified from multiple tissues when G3PDH was used as a control.We carried out in vitro binding assays to confirm that S14 or UIP1 could ind[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
TGM3 affects UCHL5
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1 |

No evidence text available
TGFBR affects UCHL5
| 1
| 1

reach
"UCHL5 interacts with Smads and reverses Smurf mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-beta receptor in cells [XREF_BIBR]."
TDP2 affects UCHL5
| 1
| 1

reach
"Thus, we observe strong association between UCH37 and TDP2 : molecules targeting one protein have a reasonable chance to target the other as an off target."
Smurf affects UCHL5
| 1
| 1

reach
"UCHL5 interacts with Smads and reverses Smurf mediated ubiqutination, and it can also deubiquitinate and stabilize the type I TGF-beta receptor in cells [XREF_BIBR]."
SMAD3 affects SMAD2
| 1
| 1

sparser
"In GST pull down experiments, UCH37 bound weakly to Smad2 and Smad3, and bound very strongly to Smad7 in a region that is distinct from the -PY- motif in Smad7 that interacts with Smurf ubiquitin ligases."
SKP2 affects UCHL5, USP10, USP13, USP14, and USP7
| 1
| 1

sparser
"We found that SKP2 interacted with USP10, USP13, UCHL5, USP14, and USP7 in KBM5-T315I cells (Fig.  xref )."
SERPINB3 affects UCHL5
1 |

No evidence text available
SERPINB12 affects UCHL5
1 |

No evidence text available
S14 affects UCHL5
| 1
UCHL5 binds HAUS7 and S14. 1 / 1
| 1

reach
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
S100A7 affects UCHL5
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1 |

No evidence text available
S100A16 affects UCHL5
1 |
1 |

No evidence text available
RPS21 affects UCHL5
1 |
1 |

No evidence text available
RPN13 affects (ULD) domain
| 1
UCHL5 binds RPN13 and (ULD) domain. 1 / 1
| 1

reach
"Rpn13 also binds to the C-terminal UCH37 like domain (ULD) domain of the UCH, locking the ULD into a favorable conformation for ubiquitin binding."
RPN13 subunit whose C-terminal domain affects UCHL5
| 1
UCHL5 binds RPN13 subunit whose C-terminal domain. 1 / 1
| 1

reach
"This activity requires the RPN13 subunit whose C-terminal domain binds UCH-L5."
RPN1 affects USP14
| 1
RPN1 binds USP14 and UCHL5. 1 / 1
| 1

sparser
"Interestingly, USP14 and UCH37 bind to RPN1 and to RPN13, respectively, which are, in addition to the RPN10, substrate receptors of the 19S regulatory particle [ xref ], which provide a versatile binding platform for various ubiquitin chains [ xref ]."
RP affects PSMD14
| 1
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
RNF20 affects UCHL5
1 |
1 |

No evidence text available
RFFL affects UCHL5
1 |
1 |

No evidence text available
RAD23B affects UCHL5
1 |
1 |

No evidence text available
RACK1 affects UCHL5
| 1
| 1

sparser
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation ( xref ), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 ( xref )."
RAB7A affects UCHL5
1 |
1 |

No evidence text available
Q1K9H5 affects UCHL5
1 |
1 |

No evidence text available
Proteasome affects INO80
| 1
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
Proteasome affects 19S
| 1
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
PTPN2 affects USP14
| 1
| 1

sparser
"The docking analyses predict that PtPT could bind to the active sites of USP14 and UCHL5, with CDOCKER Interaction Energy of −15.99 and −16.78 kcal mol −1 , and the binding modes are dis-played in xref ."
PSMD14 affects RP, UCHL5, and USP14
| 1
USP14 binds PSMD14, UCHL5, and RP. 1 / 1
| 1

sparser
"There are three essential DUBs: RPN11, UCH37, and USP14 that are associated with the 19S RP of human proteasome [ xref , xref , xref ]."
| PMC
PSMD1 affects Ubiquitin
| 1
| 1

sparser
"In vitro, we found that KDT-11 has no measurable effect on any of the protein-protein interactions in which Rpn13 engages, including interactions with Uch37, Ub and Rpn2 ( xref )."
PSMB9 affects UCHL5
1 |
1 |

No evidence text available
PSMB8 affects UCHL5
1 |
1 |

No evidence text available
PRF1 affects p2
| 1
USP14 binds UCHL5, PRF1, and p2. 1 / 1
| 1

sparser
"Given the fact that also P1 and P2 have a α,β-unsaturated keto structure, we believe that P1 and P2 are also interacting with UCHL5 and USP14."
POTEF affects UCHL5
1 |
1 |

No evidence text available
POTEE affects UCHL5
1 |
1 |

No evidence text available
POF1B affects UCHL5
1 |
1 |

No evidence text available
PNP affects UCHL5
1 |
1 |

No evidence text available
PLRG1 affects UCHL5
1 |
1 |

No evidence text available
PLA2G2A affects UCHL5
1 |
1 |

No evidence text available
PKP1 affects UCHL5
1 |
1 |

No evidence text available
PJA1 affects UCHL5
1 |
1 |

No evidence text available
PI3K_p85 affects UCHL5
| 1
PI3K_p85 increases the amount of UCHL5. 1 / 1
| 1

reach
"High glucose also increased UCHL5 protein expression, which was attenuated by LY294002, the dominant negative p85 and the dominant negative CREB."
NOS2 affects ADRM1
| 1
ADRM1 binds UCHL5 and NOS2. 1 / 1
| 1

sparser
"We previously found that the cisplatin treatment induced Rpn13 transcription by p-ΔNp63α and subsequently increased the physical interaction of Rpn13, UCH37 and NOS2 proteins leading to an essential degradation of the latter through a proteasome-dependent mechanism in SCC cells [ xref ]."
NLRP3 affects RACK1, and UCHL5
| 1
| 1

sparser
"Furthermore, through co-immunoprecipitation (IP) analysis, we observed that both NLRP3 and RACK1 directly interacted with UCHL5 after EST12 stimulation ( xref ), and EST12-Y80A stimulation could not promote the interaction of RACK1 with UCHL5 and NLRP3 ( xref )."
NACA affects UCHL5
1 |
1 |

No evidence text available
Myc-Ub affects UCHL5
| 1
Modified Myc-Ub leads to the ubiquitination of UCHL5. 1 / 1
| 1

reach
"In the control assays, overexpression of Myc-Ub promoted ubiquitination of Adrm1, S5a, Rpt5, and Uch37 but not Rpn2 (compare lanes 1 and 2 in XREF_FIG)."
METTL14 affects UCHL5
| 1
| 1

reach
"The interactions between METTL14/YTHDF1, UCHL5 and NLRP3 were analyzed using RIP and/or dual-luciferase reporter gene and/or Co-IP assays."
LMNA affects UCHL5
1 |
1 |

No evidence text available
KRT9 affects UCHL5
1 |
1 |

No evidence text available
KRT73 affects UCHL5
1 |
1 |

No evidence text available
KRT6B affects UCHL5
1 |
1 |

No evidence text available
KRT28 affects UCHL5
1 |
1 |

No evidence text available
KRT12 affects UCHL5
1 |
1 |

No evidence text available
KRT10 affects UCHL5
1 |
1 |

No evidence text available
KDM1A affects UCHL5
1 |
1 |

No evidence text available
KDELR1 affects UCHL5
1 |
1 |

No evidence text available
K 7174 affects UCHL5
1 |
K 7174 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
Infections affects UCHL5
| 1
Infections increases the amount of UCHL5. 1 / 1
| 1

reach
"Since the expression of UCHL5 was relatively lower in AN3-CA endometrial cancer cells, we increased the expression of UCHL5 by lentiviral infection in AN3-CA cells."
IVL affects UCHL5
1 |
1 |

No evidence text available
INO80 affects Proteasome, and UCHL5
| 1
| 1

sparser
"UCHL5 is associated with both the proteasome and the INO80 chromatin remodeling complex, and functions in both contexts [ xref ]."
IGHG1 affects UCHL5
1 |
1 |

No evidence text available
ICG 001 affects UCHL5
1 |
ICG 001 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
HSPB1 affects UCHL5
1 |
1 |

No evidence text available
HSP90 affects UCHL5
| 1
HSP90 activates UCHL5. 1 / 1
| 1

eidos
"In contrast , C-CBL , heat shock protein 90 ( Hsp90 ) , transforming growth factor-beta stimulated clone 22 ( TSC-22 ) , tumor necrosis factor receptor-associated factor 4 ( TRAF4 ) , ubiquitin-specific protease 4 ( USP4 ) , ubiquitin-specific protease 11 ( USP11 ) , ubiquitin-specific protease 15 ( USP15 ) , and UCH37 can stabilize the receptor by blocking the ubiquitination of the receptor [ 103-110 ] , thereby activating the TGF-beta signaling pathway ."
HNF1A affects UCHL5
1 |
HNF1A decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
HERC4 affects UCHL5
| 1
| 1

reach
"Herc4 interacts with USP8, UCHL5, and Hrs to regulate Smo ubiquitination."
HBA2 affects UCHL5
1 |
1 |

No evidence text available
HAUS7 affects S14, and UCHL5
| 1
UCHL5 binds HAUS7 and S14. 1 / 1
| 1

reach
"In the present study we found that UCH37 interact with UIP1 and S14 in a C-terminal extension dependent manner, and UIP1 was found to interact with the C-terminal extension of UCH37 alone by yeast two[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
H4C6 affects UCHL5
1 |
1 |

No evidence text available
H4C4 affects UCHL5
1 |
1 |

No evidence text available
H4C3 affects UCHL5
1 |
1 |

No evidence text available
H4C15 affects UCHL5
1 |
1 |

No evidence text available
H4C13 affects UCHL5
1 |
1 |

No evidence text available
H4C11 affects UCHL5
1 |
1 |

No evidence text available
H4C1 affects UCHL5
1 |
1 |

No evidence text available
GAS2DN affects UCHL5
| 1
GAS2DN decreases the amount of UCHL5. 1 / 1
| 1

reach
"Lastly, we generated microarray data to identify the differentially expressed genes upon GAS2DN and validated that the expression of HNRPDL, PTK7 and UCHL5 was suppressed by GAS2DN."
Flag affects UCHL5
| 1
| 1

sparser
"On the one hand, the introduction of either Flag-UCHL5 or non-tagged UCHL5 into HeLa cells remarkably enhanced myc-Axin1 and endogenous Axin1 (Fig.  xref a,b)."
FTL affects UCHL5
1 |
1 |

No evidence text available
FOXN1 affects UCHL5
1 |
FOXN1 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
FLG affects UCHL5
1 |
1 |

No evidence text available
EEF1G affects UCHL5
1 |
1 |

No evidence text available
ECPAS affects UCHL5
1 |
1 |

No evidence text available
| 1
E3_Ub_ligase ubiquitinates UCHL5-C88A. 1 / 1
| 1

reach
"Since catalytically dead UCH37 C88A is also strongly ubiquitinated by an unknown E3 ligase, we reasoned that both BAP1 and UCH37 might use their respective CTD regions for interaction with the UCH dom[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
Degrasyn affects UCHL5
| 1
Degrasyn inhibits UCHL5. 1 / 1
| 1

reach
"WP1130 (Degrasyn) has been shown to inhibit USP14 and UCHL5 as well as other DUB enzymes (see below)."
DUSP14 affects UCHL5
1 |
1 |

No evidence text available
DUB inhibitor affects UCHL5
| 1
DUB inhibitor activates UCHL5. 1 / 1
| 1

reach
"Instead, it was found to be a partially selective DUB inhibitor, directly inhibiting DUB activity of USP9x, USP5, USP14 and UCH37, which are known to regulate survival protein stability and 26S proteasome function [XREF_BIBR]."
DSP affects UCHL5
1 |
1 |

No evidence text available
DSC3 affects UCHL5
1 |
1 |

No evidence text available
DSC1 affects UCHL5
1 |
1 |

No evidence text available
DMAC2L affects USP14
| 1
| 1

sparser
"In the current study we hypothesize that electrophilic OB compounds, such as 4,4'-diamino-2,2'-stilbenedisulfonic acid(DAST), fluorescent brightener 28 (FB-28) and FB-71, can interact with the catalytic triads (CYS, HIS, and ASP) of UCHL5 and USP14 and inhibit their enzymatic activities, leading to cell growth suppression."
Cyclin affects UCHL5
| 1
Cyclin decreases the amount of UCHL5. 1 / 1
| 1

reach
"Furthermore, cyclin-dependent kinase 4/6 (CDK4/6) inhibition decreased UCHL5 expression, suppressed OTX015-R cell proliferation, and induced apoptosis."
CuPT affects UCHL5
| 1
CuPT activates UCHL5. 1 / 1
| 1

reach
"These results suggest that CuPT could target 19S DUBs, UCHL5 and USP14."
CdPT affects UCHL5
| 1
CdPT inhibits UCHL5. 1 / 1
| 1

reach
"CdPT potently inhibited the activity of proteasomal DUBs (USP14 and UCHL5), but slightly inhibited 20S proteasome activity."

reach
"These results were opposite in UCHL5 knockdown EC cells."
CYS1 affects UCHL5
| 1
| 1

sparser
"The ITC compounds may interact stronger with UCHL5 than to USP14 since more favorable bonding interactions were found upon analyzing the general decrease of free energy across the three compounds corresponding to xref . xref suggests interaction of BITC with the active site CYS-88 of UCHL5 through two formed bonds."
CYLD affects OTUB1, SMAD, UCHL5, USP11, USP15, and USP4
| 1
| 1

sparser
"Numerous studies have implicated the potential of DUBs to regulate the TGF-β signaling pathway including USP4, USP11, USP15, CYLD, OTUB1, and UCHL5, which interact with Smads to regulate TGF-β signaling [ xref , xref – xref ]."
CYB5A affects UCHL5
1 |
1 |

No evidence text available
CSTA affects UCHL5
1 |
1 |

No evidence text available
CRNDE affects UCHL5
| 1
CRNDE activates UCHL5. 1 / 1
| 1

eidos
"Knockdown of CRNDE reduced cell damage and elevated UCHL5 and SMO expressions in OGD / R-treated PC-12 cells ."
COPS5 affects 293T
| 1
UCHL5 binds COPS5 and 293T. 1 / 1
| 1

reach
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA or anti-Flag antibody."
COMMD4 affects UCHL5
1 |
1 |

No evidence text available
CNTLN affects UCHL5
| 1
CNTLN inhibits UCHL5. 1 / 1
| 1

reach
"This discrepancy in centrosomal protein stability may result from the ability of WP1130 to inhibit USP5, USP14, and UCH37 in addition to USP9X."
CHI3L1 affects UCHL5
| 1
| 1

sparser
"Uch37 could form oligomers in solution."
CFL1 affects UCHL5
1 |
1 |

No evidence text available
CDSN affects UCHL5
1 |
1 |

No evidence text available
CASP1 affects UCHL5
| 1
| 1

sparser
"We also showed that overexpression of UCH-L5 was associated with a significant increase in caspase-1 activity, while inhibition of UCH-L5 by selective inhibitor [ xref ] or UCH-L5 knock-down led to decrease in inflammasome-dependent IL-1β release in chicken as well as in human macrophages during infection with Salmonella and during inflammasome activation by lipopolysaccharide (LPS) and nigericin."
Bloc1s3 affects UCHL5
| 1
UCHL5 binds Bloc1s3 and 19S. 1 / 1
| 1

reach
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."
BAP1 affects ULD
| 1
UCHL5 binds BAP1 and ULD. 1 / 1
| 1

sparser
"UCH-L5 and BAP1 both interact with the DEUBAD domains of their binding partners via their ULDs."
1 |
Aroclor 1254 decreases the amount of UCHL5. 1 / 1
1 |

No evidence text available
AgDT affects UCHL5
| 1
AgDT inhibits UCHL5. 1 / 1
| 1

reach
"Mechanistically, AgDT potently inhibited the activities of proteasomal DUBs USP14 and UCHL5, without altering the 20S proteasome peptidases."
ARMT1 affects UCHL5
1 |
1 |

No evidence text available
AR affects USP14
| 1
USP14 binds UCHL5 and AR. 1 / 1
| 1

sparser
"To determine if there is any interaction between AR protein and USP14 or UCHL5 protein, we performed co-immunoprecipitation (co-IP) for AR, USP14, and UCHL5."
APOB affects UCHL5
1 |
1 |

No evidence text available
APOA1 affects UCHL5
1 |
1 |

No evidence text available
ANXA8 affects UCHL5
1 |
1 |

No evidence text available
ANXA1 affects UCHL5
1 |
1 |

No evidence text available
ANP32B affects UCHL5
1 |
1 |

No evidence text available
| 1

sparser
"AIG1, ATP6V0B, BLOC1S1, IYD, NDUFB2, RPL23A, and UCHL5 were significantly associated with 11 splicing factors, and SLC27A5 was associated with up to 12 splicing factors."
ADRM1 affects monoubiquitin
| 1
ADRM1 binds UCHL5 and monoubiquitin. 1 / 1
| 1

sparser
"Additionally, there has been recent development of another competitive FP assay that includes fluorescently-labeled monoubiquitin bound to the Uch37:Rpn13 complex and then competed off with unlabeled ubiquitin chains."
ADRM1 affects UCH domain, and UCHL5
| 1
ADRM1 binds UCHL5 and UCH domain. 1 / 1
| 1

reach
"Binding of Adrm1 to UCH37 relieved auto-inhibition by the UCH domain and activated its deubiquitination activity [XREF_BIBR]."
ADRM1 affects NOS2, and UCHL5
| 1
ADRM1 binds UCHL5 and NOS2. 1 / 1
| 1

sparser
"We previously found that the cisplatin treatment induced Rpn13 transcription by p-ΔNp63α and subsequently increased the physical interaction of Rpn13, UCH37 and NOS2 proteins leading to an essential degradation of the latter through a proteasome-dependent mechanism in SCC cells [ xref ]."
ACTC1 affects UCHL5
1 |
1 |

No evidence text available
4-phenylbutyric acid increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
2c-biotin affects UCHL5
| 1
UCHL5 binds 2c-biotin. 1 / 1
| 1

reach
"Figure XREF_FIG confirms that in vivo 2c-biotin binds with comparable intensity USP14 and UCH-L5."
293T affects UCHL5
| 1
UCHL5 binds COPS5 and 293T. 1 / 1
| 1

reach
"The interaction of UCH37 and COPS5 in 293T cells, in which both proteins were overexpressed, was confirmed using immunoprecipitation analysis by either anti-HA or anti-Flag antibody."
19S affects auranofin
| 1
Auranofin binds UCHL5 and 19S. 1 / 1
| 1

reach
"CuPT and auranofin on 19S proteasome associated UCHL5 and USP14."
19S affects Proteasome, and UCHL5
| 1
Proteasome binds UCHL5 and 19S. 1 / 1
| 1

reach
"Rpn11 is an intrinsic subunit of 19S regulatory particle, whereas USP14 and UCHL5 reversibly associate with 19S proteasome, indicative of attractive and versatile roles for these DUBs [XREF_BIBR - XREF_BIBR]."
19S affects Bloc1s3, and UCHL5
| 1
UCHL5 binds Bloc1s3 and 19S. 1 / 1
| 1

reach
"Both USP14 and UCHL5 bind reversibly to the 19S RP and are implicated in cancer [48] ."
17alpha-ethynylestradiol increases the amount of UCHL5. 1 / 1
1 |

No evidence text available
1.171 affects UCHL5
| 1
1.171 increases the amount of UCHL5. 1 / 1
| 1

isi
"The multivariate analysis indicated that the UCHL5 expression level was an independent prognostic factor for OS (HR=1.171, 95% CI=1.052-1.303) and DFS (HR=1.143, 95% CI=1.031-1.267) in these patients."

reach
"They further showed that some DUBs that have been exposed to ROS can have different responses to reduction through DTT treatment, with some being activated by DTT (e.g., USP8, USP10, and USP19), some having only enhancement of activity in the presence of DTT (e.g., USP7, CYLD, and UCHL5), and others exhibiting no activity, despite the presence of a reducing agent (e.g., USP1, USP22, and A20) (35)."
(ULD) domain affects UCHL5
| 1
UCHL5 binds RPN13 and (ULD) domain. 1 / 1
| 1

reach
"Rpn13 also binds to the C-terminal UCH37 like domain (ULD) domain of the UCH, locking the ULD into a favorable conformation for ubiquitin binding."
| 1

reach
"Analysis of the associated and non associated DUBs, such as UCHL2, USP14, UCH37, and UBC9 and DUB associated with core 20S proteasome (PSMD13) in RA-FLS showed that in the presence of IL-1beta, EGCG increased the expression of UCH37 and USP14, DUBs known to preferentially hydrolyze K48 linked polyubiquitin chains, with a marginal effect on unassociated DUB such as UCHL2 (XREF_FIG)."
1 |

No evidence text available