IndraLab

Statements


UCHL3 is modified
5 | 1 48 22
UCHL3 is phosphorylated.
5 | 1 47 22
UCHL3 is phosphorylated. 10 / 59
| 1 43 15

sparser
"To further demonstrate a potential function of UCHL3 phosphorylation, we reconstituted UCHL3-deficient cells with wild-type UCHL3 or the UCHL3 S75A mutant."

sparser
"Because S75 is located in the catalytic domain of UCHL3, we next examined whether UCHL3 phosphorylation regulates its activation."

sparser
"Following the recruitment of XRCC4-LIG4 and ligation of the broken DNA ends, UCHL3 is phosphorylated and destabilized, which tilts the ubiquitylation balance of Ku80 in favour of ubiquitylation by exposing Ku80 to E3 ligases."

sparser
"As shown in xref C, S75 was phosphorylated after DNA damage, while the S75A mutation inhibited DNA damage-induced UCHL3 phosphorylation."

sparser
"Wild-type UCHL3 (UCHL3 WT ) and simulated phosphorylated UCHL3 (UCHL3 S75E ) were recombinantly expressed and purified using affinity chromatography, ion exchange chromatography, and gel filtration chromatography."

rlimsp
"Our data supports the idea that Ku80 ubiquitylation is induced by loss of UCHL3 triggered by ATM-mediated phosphorylation, and E3 ligases targeting Ku80 might be constitutively activated."

sparser
"To further establish that the signal detected with the anti-phospho S/TQ antibody indicates phosphorylated UCHL3, immunoblotting was performed with a mutant UCHL3 in which Ser75 was substituted with Ala (S75A)."

rlimsp
"Lastly, we provide evidence showing that DNA damage-induced UCHL3 phosphorylation (which is dependent on UCHL3 catalytic activity, ATM kinase and downstream NHEJ factors) promotes destabilization of UCHL3, suggesting that UCHL3 may dissociate from Ku80 to allow Ku ubiquitylation and subsequent removal from DNA."

rlimsp
"Following the recruitment of XRCC4-LIG4 and ligation of the broken DNA ends, UCHL3 is phosphorylated and destabilized, which tilts the ubiquitylation balance of Ku80 in favour of ubiquitylation by exposing Ku80 to E3 ligases."

sparser
"To test whether UCHL3 could be phosphorylated by ATM/ATR, we examined UCHL3 phosphorylation using an antibody against the consensus of ATM/ATR phosphorylation sites (anti-phospho-SQ/TQ [anti-pSQ/TQ]) following DNA damage."
UCHL3 is phosphorylated on S75. 10 / 13
2 | 4 7

rlimsp
"Altogether, these data suggest that phosphorylation of UCHL3 on S75, which is dependent on ATM, UCHL3′s catalytic activity and downstream NHEJ factors such as XRCC4 and LigaseIV, regulates Ku80 ubiquitylation by regulating UCHL3 stability rather than affecting its deubiquitylating activity after DSB induction."

rlimsp
"Since UCHL3 was previously listed as a potentially phosphorylated protein in response to IR, and harbours a single SQ site (Ser75), the consensus sequence of ATM, ATR and DNA-PKcs kinases, we tested whether UCHL3 is phosphorylated upon phleomycin treatment."

rlimsp
"To further establish that the signal detected with the anti-phospho S/TQ antibody indicates phosphorylated UCHL3, immunoblotting was performed with a mutant UCHL3 in which Ser75 was substituted with Ala (S75A)."

rlimsp
"The ensuing results demonstrated that the S75A mutation abolished the signal from the anti-phospho-S/TQ antibody, thus confirming that Ser75 of UCHL3 is phosphorylated in response to DSBs (Supplementary Fig."

rlimsp
"The ensuing results demonstrated that the S75A mutation abolished the signal from the anti-phospho-S/TQ antibody, thus confirming that Ser75 of UCHL3 is phosphorylated in response to DSBs (Supplementary Fig. S6C). We also sought to establish the function of this UCHL3 phosphorylation. In an in vitro deubiquitylating activity assay, purified UCHL3 harbouring either S75A (unphosphorylatable mutant) or S75E (phosphomimetic mutant) showed comparable levels of binding to HA-tagged ubiquitin vinyl sulfone (HA-Ub-VS), which covalently binds to DUB active sites, when compared with wild-type UCHL3, suggesting that Ser75 phosphorylation does not affect the intrinsic enzymatic activity of UCHL3 (Supplementary Fig."

sparser
"Altogether, these data suggest that phosphorylation of UCHL3 on S75, which is dependent on ATM, UCHL3′s catalytic activity and downstream NHEJ factors such as XRCC4 and LigaseIV, regulates Ku80 ubiquitylation by regulating UCHL3 stability rather than affecting its deubiquitylating activity after DSB induction."

rlimsp
"UCHL3 is phosphorylated on S75 in a manner dependent on its catalytic activity, ATM and XRCC4 (as well as LIG4). Although this phosphorylation neither affects the enzymatic activity of UCHL3 in vitro nor its interaction with Ku80 in cells, it facilitates destabilization of UCHL3 after DSB induction."

rlimsp
"UCHL3 is phosphorylated on S75 in a manner dependent on its catalytic activity, ATM and XRCC4 (as well as LIG4)."

sparser
"The ensuing results demonstrated that the S75A mutation abolished the signal from the anti-phospho-S/TQ antibody, thus confirming that Ser75 of UCHL3 is phosphorylated in response to DSBs (Supplementary Fig.  xref )."

sparser
"UCHL3 is phosphorylated on S75 in a manner dependent on its catalytic activity, ATM and XRCC4 (as well as LIG4)."
UCHL3 is phosphorylated on S130. 2 / 2
2 |

No evidence text available

No evidence text available
UCHL3 is phosphorylated on Y85. 1 / 1
1 |

No evidence text available
UCHL3 is methylated.
| 1
UCHL3 is methylated. 1 / 1
| 1

sparser
"Bag4 is a protein-coding gene that plays a role by interacting with various cell death and growth-related proteins, and regulates their activity by binding to hsp70/hsc70 family proteins ( xref ). syn2b is a member of the synaptotagmin gene family, encoding a neuronal phosphoprotein associated with synaptic vesicle cytoplasmic surfaces ( xref ). map3k4 is also a protein-coding gene, and studies have shown that high methylation of map3k4 kinase and low methylation of uchl3, observed in seawater-released marine sticklebacks, can potentially reduce protein ubiquitination levels and serve as an adaptation to osmotic stress ( xref )."
UCHL3 affects RAD51
5 5 1 | 35 10
UCHL3 deubiquitinates RAD51.
2 1 | 23
UCHL3 deubiquitinates RAD51. 10 / 24
1 | 23

"<span class="match term0">UCHL3</span>, in turn, deubiquitinates <span class="match term1">RAD51</span> and promotes the binding between <span class="match term1">RAD51</span> and BRCA2."

reach
"UCHL3 interacts with and deubiquitinates RAD51."

reach
"After DNA damages, UCHL3 is activated via phosphorylation by ATM and then deubiquitinates RAD51 promoting RAD51-BRCA2 interaction and proper HR repair [150]."

reach
"Reconstituting wild-type UCHL3 but not catalytically inactive UCHL3 (CA) reversed the increase in RAD51 ubiquitination induced by UCHL3 deficiency (Fig. 2E), suggesting that UCHL3 regulates RAD51 ubiquitination through its Ub protease activity.To determine whether UCHL3 directly deubiquitinates RAD51, we performed an in vitro deubiquitination assay."

reach
"Because UCHL3 promotes RAD51 deubiquitination following DNA damage, we further investigated whether and how UCHL3 itself is regulated following DNA damage."

reach
"Taken together, these results suggest that UCHL3 deubiquitinates RAD51 both in vitro and in vivo."

reach
"Collectively, these results suggest that UCHL3 promotes RAD51 deubiquitination following DNA damage."

reach
"Deubiquitination of RAD51 by UCHL3 is important for HR."

reach
"Following DNA damage, UCHL3 promotes RAD51 deubiquitination (Figs."

reach
"It is possible that RAD51 deubiquitination by UCHL3 is important for its recruitment to DSBs."
UCHL3 deubiquitinates RAD51 on K58. 1 / 1
1 |

"Here we report that ubiquitination of RAD51 hinders RAD51-BRCA2 interaction, while deubiquitination of RAD51 facilitates RAD51-BRCA2 binding and RAD51 recruitment and thus is critical for proper HR. | UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2.|Our results suggested that three lysine sites (56, 57, and 63) on RAD51 that are close to E59 are deubiquitinated by UCHL3."
UCHL3 deubiquitinates RAD51 on K64. 1 / 1
1 |

"Here we report that ubiquitination of RAD51 hinders RAD51-BRCA2 interaction, while deubiquitination of RAD51 facilitates RAD51-BRCA2 binding and RAD51 recruitment and thus is critical for proper HR. | UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2.|Our results suggested that three lysine sites (56, 57, and 63) on RAD51 that are close to E59 are deubiquitinated by UCHL3."
UCHL3 binds RAD51.
5 | 8 10
5 | 8 10

No evidence text available

sparser
"Furthermore, wild-type RAD51 and the 3KR mutant bound equally with UCHL3, suggesting that RAD51 deubiquitination does not affect RAD51/UCHL3 interaction ()."

No evidence text available

No evidence text available

No evidence text available

sparser
"We found that S75A mutation did not affect the interaction between UCHL3 and RAD51 ( xref G)."

sparser
"UCHL3 interacts with RAD51 and de-ubiquitinates residues that disrupt the RAD51–BRCA2 interaction."

sparser
"We also performed co-IP experiments to examine the changes in the interaction between RAD51 and UCHL3 in HEK293 cells that were treated with farrerol, and we excluded the possibility that farrerol promoted the deubiquitination of RAD51 by enhancing the UCHL3-RAD51 interaction (Supplementary Fig.  xref )."

sparser
"We performed in vitro and in vivo deubiquitination assay to establish that farrerol directly stimulates UCHL3 enzyme activity and promotes HR by UCHL3-RAD51 axis."

reach
"Here we carried out a systematic screen of DUBs for HR and established that UCHL3 interacts with and deubiquitinates RAD51 and promotes binding between RAD51 and BRCA2."
UCHL3 activates RAD51.
3 | 3
UCHL3 activates RAD51. 6 / 6
3 | 3

reach
"Another complimentary possibility is that, unlike BRCA2/RAD51 knockout, UCHL3 knockout only decreases Rad51 function."

"Here we report that ubiquitination of RAD51 hinders RAD51-BRCA2 interaction, while deubiquitination of RAD51 facilitates RAD51-BRCA2 binding and RAD51 recruitment and thus is critical for proper HR. | UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2.|Our results suggested that three lysine sites (56, 57, and 63) on RAD51 that are close to E59 are deubiquitinated by UCHL3."

"Here we report that ubiquitination of RAD51 hinders RAD51-BRCA2 interaction, while deubiquitination of RAD51 facilitates RAD51-BRCA2 binding and RAD51 recruitment and thus is critical for proper HR. | UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2.|Our results suggested that three lysine sites (56, 57, and 63) on RAD51 that are close to E59 are deubiquitinated by UCHL3."

reach
"Re-expression of wild-type UCHL3 restored RAD51 IRIF, while expression of the S75A mutant did not (Fig. 5D; Supplemental Fig. S9B)."

"Here we report that ubiquitination of RAD51 hinders RAD51-BRCA2 interaction, while deubiquitination of RAD51 facilitates RAD51-BRCA2 binding and RAD51 recruitment and thus is critical for proper HR. | UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2.|Our results suggested that three lysine sites (56, 57, and 63) on RAD51 that are close to E59 are deubiquitinated by UCHL3."

reach
"However, overexpression of UCHL3-mediated resistance to radiation and PARPi was abolished in RAD51-depleted cells (Fig. 6G,H)."
UCHL3 ubiquitinates RAD51.
| 1
UCHL3 ubiquitinates RAD51. 1 / 1
| 1

reach
"In previous studies, RING1,19 UCHL3,20 UAF1,21 FBH122, 23 UBL1,24, 25 UBC9,26 and BLM27 were reported to manipulate the ubiquitination of Rad51 by direct interaction."
Ubiquitin affects UCHL3
| 7 38
Ubiquitin binds UCHL3.
| 2 36
| 2 33

sparser
"One possibility for the enhanced discrimination of Gly over Val at P2 in the context of the natural substrate is that ubiquitin binding to UCH-L3 may help to position its P2 residue for optimal catalysis."

sparser
"Building on our previous work for computational development of UCH selective UbV-ABPs, we applied a rational design approach to incorporate mutations to the Ub-UCHL3 protein-protein interaction (PPI) interface that would improve binding affinity and impart selectivity over UCHL1 and other DUBs."

sparser
"Both ABPs were shown to interact with UCHL3 as indicated by the presence of the higher molecular weight band correlating to the UCHL3-Ub complex formation over the time course."

sparser
"To do this a binding analysis of the interactions at the PPI interface of both the Ub-UCHL3 (PDB: 1XD3 [ xref ]) and Ub-UCHL1 (PDB: 3KW5 [ xref ]) crystal structure complexes was carried out using BioLuminate (Schrödinger, LLC, New York, NY, USA) in an effort to identify interactions Ub makes that are different between the two DUBs."

sparser
"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 ( xref and xref ) and the yeast UCH Yuh1 xref , xref ."

sparser
"From a crystal structure of a complex between UCHL3 and synthetic K27 Ub 2 , we unexpectedly discover that free K27 Ub 2 and K27 Ub 2 -conjugated substrates are natural inhibitors of UCHL3."

reach
"We identified three inhibitors (IC 50 = 100–150 μM) of UCH-L3 by the DOCK–GOLD virtual screening of 32,799 compounds.In the 3D structure of the UCH-L3-ubiquitin complex, ubiquitin C-terminus is buried[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Isotopically labelled ubiquitin was bound to excess UCH-L3 and the chemical shift changes relative to free ubiquitin were determined."

sparser
"NMR assignments ( 1 HN, 15 N, 13 C α , 13 CO) were determined for 2 H, 13 C, 15 N-labeled human ubiquitin bound to UCH-L3."

sparser
"From the structure of UCHL3 WT -Ub-VME, we can see that Pro8 on UCHL3 interacts with Arg74 on Ub via hydrogen bonding."
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
Ubiquitin inhibits UCHL3.
| 3 2
| 3 2

sparser
"As an additional control experiment, we measured a K i of 0.27 μM for the inhibition of UCH-L3 by ubiquitin with the substrate ubiquitin-AMC."

reach
"The present data demonstrated that ubiquitin aldehyde can inhibit the activity of UCHL3."

sparser
"Using our Ub tools to profile UCHL3's activity, we generate a quantitative kinetic model of the inhibitory mechanism and we find that K27 Ub 2 can inhibit UCHL3 covalently, by binding to its catalytic cysteine, and allosterically, by locking its catalytic loop tightly in place."

reach
"(2) The kinetics of inhibition of UCH-L3 by ubiquitin C-terminal aldehyde (Ub-H) were determined and reveal a Ki that is less than 10 (-14) M. Several mechanisms are considered to account for the extreme inhibition."

reach
"As an additional control experiment, we measured a K of 0.27 μM for the inhibition of UCH-L3 by ubiquitin with the substrate ubiquitin-AMC."
Ubiquitin activates UCHL3.
| 2
| 2

reach
"This result was similar to that conducted by UCHL3, which produced more free coumarin over time (Fig. 2g) and was previously demonstrated to be activated by distal Ub ."

reach
"Regulating biological functions, UCHL3 promoted GSC proliferation, tumorigenesis, self-renewal, and radioresistance in a POLD4-dependent manner.The degradation of POLD4 by ubiquitin ligases in response to DNA damage and during cell cycle progression plays a less prominent role in normal cells [43], while with increasing insights into the complex mechanisms of DNA damage repair, POLD4 is projected to play an increasingly important role."
UCHL3 affects Ubiquitin
| 5 36
UCHL3 binds Ubiquitin.
| 2 36
| 2 33

sparser
"One possibility for the enhanced discrimination of Gly over Val at P2 in the context of the natural substrate is that ubiquitin binding to UCH-L3 may help to position its P2 residue for optimal catalysis."

sparser
"Building on our previous work for computational development of UCH selective UbV-ABPs, we applied a rational design approach to incorporate mutations to the Ub-UCHL3 protein-protein interaction (PPI) interface that would improve binding affinity and impart selectivity over UCHL1 and other DUBs."

sparser
"Both ABPs were shown to interact with UCHL3 as indicated by the presence of the higher molecular weight band correlating to the UCHL3-Ub complex formation over the time course."

sparser
"To do this a binding analysis of the interactions at the PPI interface of both the Ub-UCHL3 (PDB: 1XD3 [ xref ]) and Ub-UCHL1 (PDB: 3KW5 [ xref ]) crystal structure complexes was carried out using BioLuminate (Schrödinger, LLC, New York, NY, USA) in an effort to identify interactions Ub makes that are different between the two DUBs."

sparser
"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 ( xref and xref ) and the yeast UCH Yuh1 xref , xref ."

sparser
"From a crystal structure of a complex between UCHL3 and synthetic K27 Ub 2 , we unexpectedly discover that free K27 Ub 2 and K27 Ub 2 -conjugated substrates are natural inhibitors of UCHL3."

reach
"We identified three inhibitors (IC 50 = 100–150 μM) of UCH-L3 by the DOCK–GOLD virtual screening of 32,799 compounds.In the 3D structure of the UCH-L3-ubiquitin complex, ubiquitin C-terminus is buried[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Isotopically labelled ubiquitin was bound to excess UCH-L3 and the chemical shift changes relative to free ubiquitin were determined."

sparser
"NMR assignments ( 1 HN, 15 N, 13 C α , 13 CO) were determined for 2 H, 13 C, 15 N-labeled human ubiquitin bound to UCH-L3."

sparser
"From the structure of UCHL3 WT -Ub-VME, we can see that Pro8 on UCHL3 interacts with Arg74 on Ub via hydrogen bonding."
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
UCHL3 decreases the amount of Ubiquitin.
| 1
UCHL3 decreases the amount of Ubiquitin. 1 / 2
| 1

reach
"It has been proposed that the rs4885322 SNP may influence UCHL3 gene expression, disrupting cellular Ub levels, which in turn could impair insulin signaling, promoting insulin resistance and hyperglycemia [11]."
UCHL3 activates Ubiquitin.
| 2
| 2

reach
"In our study, we found that the ubiquitin–proteasome system (UPS) was modulated by the regulation of some ubiquitin enzyme and ligases genes (uchl3, trim 25, trim39, ubr7 and ube2i) and proteasome subunits (psma4, psma6, psmb10, psmc2 and rad23a), which act as protein binding signalling and protein degradation, respectively."

reach
"The relevant control exerted by these DUBs on ubiquitin homeostasis may account for these effects since pharmacological blocking of UCHL1 or UCHL3 reduced the monomeric ubiquitin pool, in turn restricting the ubiquitination of GlyT2 in neurons (XREF_FIG)."
UCHL3 affects CTNNB1
2 | 26 8
UCHL3 binds CTNNB1.
2 | 6 8
2 | 6 8

sparser
"These results suggest that UCHL3 might physically interact with CTNNB1 to trigger activation of the Wnt signaling pathway."

sparser
"The interaction between UCH-L3 and β-catenin proteins was confirmed, and it reduced the ubiquitination of β-catenin."

sparser
"Hence, we evaluated the interaction between UCHL3 and CTNNB1."

reach
"A Pierce Co-Immunoprecipitation Kit (Thermo Fisher, USA) was used to verify the physical interaction between UCHL3 and CTNNB1."

sparser
"Interestingly, we found that UCHL3 interacted with CTNNB1, as shown by co-IP and colocalization assays with T24 cells."

sparser
"To verify the interaction between UCHL3 and CTNNB1 in bladder cancer cells, the T24 cell line was selected for an endogenous immunoprecipitation (IP) analysis."

reach
"UCHL3 interacts with and stabilizes CTNNB1."

sparser
"The results showed that UCHL3 binds endogenous CTNNB1 under pathological conditions (Fig.  xref B)."

sparser
"The interaction between UCH-L3 and β-catenin proteins was observed, resulting in reduced ubiquitination and increased protein stability of β-catenin."

reach
"Hence, we evaluated the interaction between UCHL3 and CTNNB1."
UCHL3 activates CTNNB1.
| 8
UCHL3 activates CTNNB1. 8 / 8
| 8

reach
"Accordingly, UCHL3 deficiency decreased the CTNNB1 protein half-life (Fig. 5H)."

reach
"Taken together, UCHL3 promotes the stability of CTNNB1, and this effect depends on its deubiquitinating activity."

reach
"Our results suggest that UCH-L3 induces the stabilization of beta-catenin, which is required to promote stemness during autophagy activation."

reach
"We further speculated that ubiquitin C-terminal hydrolase-L3 (UCH-L3), the deubiquitinase formerly reported to maintain stem cell properties, could activate β-catenin signaling during autophagy (Lee et al., 2021; Ouyang et al., 2020)."

reach
"In addition, the knockdown of endogenous UCH-L3 abrogated the upregulation of β-catenin induced by glucose deprivation (Supplementary Fig. S2B)."

reach
"UCHL3 increases β-catenin protein stability via removing K48-specific poly-ubiquitin chains from β-catenin protein."

reach
"In general, our results indicates that UCHL3 increases the stability of β-catenin, which in turn facilitates tumorigenesis of HCC, suggesting that targeting UCHL3 may be a promising approach for the treatment of HCC."

reach
"However, it is possible that β-catenin signaling activated by UCH-L3 is required to maintain the stem cell properties of GCSCs during autophagy-activating conditions, such as nutrient deprivation.In summary, we suppose that the suppression of UCH-L3 could effectively limit gastric cancer recurrence after surgery by preventing GCSC induction."
UCHL3 increases the amount of CTNNB1.
| 4
UCHL3 increases the amount of CTNNB1. 4 / 4
| 4

reach
"UCHL3 deficiency also reduced CTNNB1 expression in xenograft tumors and mouse bladders."

reach
"We also observed that inhibition of UCH-L3 activity reduced beta-catenin protein levels."

reach
"Similarly, knockdown of UCHL3 decreased the expression level of Myc-beta-catenin, and overexpression of UCHL3 upregulated Myc-beta-catenin levels in a dose dependent manner."

reach
"UCHL3 knockdown markedly reduces the protein level of β-catenin in HCC cells."
UCHL3 deubiquitinates CTNNB1.
| 4
UCHL3 deubiquitinates CTNNB1. 4 / 4
| 4

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"However, whether UCHL3 is involved in the deubiquitination of CTNNB1 and its role in bladder cancer remain unknown.The aim of this study is to elucidate the mechanisms underlying the impact of UCHL3 on the initiation and progression of bladder cancer and to identify potential targets for its treatment."

reach
"Subsequently, we evaluated whether UCHL3 is able to catalyze the deubiquitination of CTNNB1."

reach
"Therefore, UCHL3 might suppress the ubiquitin-driven degradation of CTNNB1 and thereby trigger the Wnt signaling pathway, resulting in tumorigenic promotion.Although the stabilization of UCHL3 depends on its deubiquitinating activity, UCHL3 directly deubiquitinates CTNNB1, and the sites of CTNNB1 that are deubiquitinated by UCHL3 need to be identified.In summary, we showed that UCHL3 expression is amplified in bladder cancer and functions as a tumor promoter that enhances the proliferation and migration of bladder cancer cells in vitro and to bladder tumorigenesis and tumor progression in vivo."

reach
"Furthermore, β-catenin ubiquitination was reduced by the binding of wild-type UCH-L3 (Fig. 2J)."
UCHL3 inhibits CTNNB1.
| 2
UCHL3 inhibits CTNNB1. 2 / 2
| 2

reach
"UCH-L3 promotes stem cell properties by inhibiting beta-catenin degradation."

reach
"UCHL3 inhibits ferroptosis by stabilizing β-catenin and maintains stem-like properties of hepatocellular carcinoma cells."
UCHL3 ubiquitinates CTNNB1.
| 1
UCHL3 ubiquitinates CTNNB1. 1 / 1
| 1

reach
"Therefore, UCHL3 might suppress the ubiquitin-driven degradation of CTNNB1 and thereby trigger the Wnt signaling pathway, resulting in tumorigenic promotion.Although the stabilization of UCHL3 depends on its deubiquitinating activity, UCHL3 directly deubiquitinates CTNNB1, and the sites of CTNNB1 that are deubiquitinated by UCHL3 need to be identified.In summary, we showed that UCHL3 expression is amplified in bladder cancer and functions as a tumor promoter that enhances the proliferation and migration of bladder cancer cells in vitro and to bladder tumorigenesis and tumor progression in vivo."
UCHL3 decreases the amount of CTNNB1.
| 1
UCHL3 decreases the amount of ubiquitinated CTNNB1. 1 / 1
| 1

reach
"As shown in Fig. 4I, the level of ubiquitinated CTNNB1 was profoundly decreased by UCHL3 but not the UCHL3 mutant."
UCHL3 affects AHR
1 | 28 5
UCHL3 activates AHR.
| 13
UCHL3 activates AHR. 10 / 12
| 12

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"Taken together, experimental data from the present study delineated that LINC00665 acted as a miR-582-5p sponge to up-regulate UCHL3 levels, and enhanced the stability of AhR protein, thereby reducing the radiosensitivity and promoting the immune escape in NSCLC cells."

reach
"Generally, these findings indicated that as a DUB of AhR, UCHL3 increases AhR protein stabilization in a DUB activity-dependent manner."

reach
"Collectively, these findings indicate that inhibition of UCHL3 may effectively eliminate lung cancer stem cell properties by promoting AhR destabilization."

reach
"UCHL3 potentiates the resistance of NSCLC cells to radiotherapy by stabilizing AhR protein."

reach
"Together, these results indicated that UCHL3 enhanced the radioresistance of NSCLC cells by stabilizing AhR protein."

reach
"UCHL3 increased AhR stability and the binding of AhR to the promoter regions of the "stemness" genes ATP-binding cassette subfamily G member 2 (ABCG2), KLF4, and c-Myc."

reach
"In this paper, we found a new mechanism of AhR regulation and functional mechanism by which UCHL3 promotes lung cancer progression and stemness."

reach
"In summary, our experiments indicate that UCHL3 is a specific DUB for AhR and that UCHL3 elevates AhR protein stabilization through deubiquitination.We further detected mRNA levels in 35 paired ADC and SCC lung cancer and corresponding paracancerous normal tissues (Supplementary Fig. S4b, c) and noted no difference between histologic subtypes."

reach
"As a deubiquitination, UCHL3 has been suggested to maintain cancer stem-like properties and accelerate tumor cell growth by stabilizing AhR protein [11]."

reach
"In summary, our findings revealed that AhR plays a critical role in maintaining the stem-like properties of cells overexpressing UCHL3, confirming that UCHL3 promotes tumor stem-like properties through stabilizing AhR."
Mutated UCHL3 activates AHR. 1 / 1
| 1

reach
"Nevertheless, our results suggest that WT UCHL3, but not UCHL3 mutants, promotes tumor growth and stem-like properties through stabilizing AhR in a DUB activity-dependent manner."
UCHL3 binds AHR.
1 | 1 5
1 | 1 5

sparser
"Ouyang et al. revealed the direct interaction between AhR and UCH-L3 in which UCH-L3 exclusively deubiquitinates and thus, reduces AhR poly-ubiquitination which in turn, stabilizes AhR."

sparser
"First, UCHL3 and AhR directly interact."

sparser
"UCHL3 interacts with AhR and stabilizes the AhR protein through deubiquitination."

sparser
"Then, we explored the interaction between the UCHL3 and AhR proteins."

No evidence text available

sparser
"Then, Co-IP results confirmed that AhR interacted with UCHL3 (Fig.  xref E)."

reach
"Co-IP assay was performed to evaluate the interaction between UCHL3 and AhR."
UCHL3 deubiquitinates AHR.
| 6
UCHL3 deubiquitinates AHR. 6 / 6
| 6

reach
"In addition, UCHL3 promotes the proliferation and stem cell traits of NSCLC cells by deubiquitinating Aryl hydrocarbon receptor (AhR) (90)."

reach
"WT UCHL3, but not UCHL3-C95S, significantly reduced AhR protein polyubiquitination (Fig. 6f), whereas the other mutants did not decrease AhR polyubiquitination."

reach
"As expected, the ability of UCHL3 to deubiquitinate AhR was almost completely abrogated by TCID treatment (Fig. 7a)."

reach
"Second, UCHL3 reduces AhR polyubiquitination and enhances AhR protein stabilization in a DUB activity-dependent manner, and this ability could be inhibited by TCID, a UCHL3 inhibitor."

reach
"Moreover, UCHL3 in non-small cell lung cancers promotes stem-like characteristics and potent tumorigenic capacity by deubiquitinating the aryl hydrocarbon receptor [28]."

reach
"The results indicated that UCHL3 knockdown led to augmented ubiquitination of endogenous AhR protein while UCHL3 up-regulation led to the opposite (Fig. 2H)."
UCHL3 increases the amount of AHR.
| 4
UCHL3 increases the amount of AHR. 4 / 4
| 4

reach
"Silencing UCHL3 led to a reduced level of AhR protein, and UCHL3 overexpression led to an elevated AhR protein level (Fig. 2C)."

reach
"According to Western blot analysis results, protein levels of UCHL3, AhR and PD-L1 were all reduced in response to sh-UCHL3 treatment, while additional LV-AhR restored the protein levels of AhR and PD-L1 but showed on obvious effects on the UCHL3 protein level; consistently, LV-UCHL3 led to up-regulated protein levels of UCHL3, AhR and PD-L1, among which only the up-regulation of AhR and PD-L1 was reversed in response to additional sh-AhR treatment (Fig. 2M)."

reach
"Furthermore, overexpression of UCHL3 decreased the ubiquitination of endogenous AhR in A549 and H1299 cells (Fig. 4h, i), while depletion of endogenous UCHL3 elevated the ubiquitination level of endogenous AhR in H358 cells (Fig. 4j)."

reach
"WT UCHL3 obviously increased AhR and c-Myc protein levels, whereas the mutants did not (Fig. 6a)."
UCHL3 ubiquitinates AHR.
| 2
UCHL3 leads to the ubiquitination of AHR. 2 / 2
| 2

reach
"WT UCHL3, but not UCHL3-C95S, significantly reduced AhR protein polyubiquitination (Fig. 6f), whereas the other mutants did not decrease AhR polyubiquitination."

reach
"Correspondingly, TCID treatment reduced AhR protein levels and stemness-associated markers in UCHL3-overexpressing A549 cells without affecting AhR mRNA levels (Fig. 7b, c), suggesting that TCID, like UCHL3 knockdown, promotes AhR ubiquitination and degradation."
UCHL3 inhibits AHR.
| 2
UCHL3 inhibits AHR. 2 / 2
| 2

reach
"UCHL3 reduced the radiosensitivity of NSCLC cells by stabilizing AhR protein."

reach
"In agreement with these studies, our data illuminated that UCHL3 reduced the radiosensitivity of NSCLC cells by stabilizing AhR protein."
UCHL3 affects UBC
32 |
31 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

No evidence text available

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No evidence text available
1 |

No evidence text available
UCHL3 affects COPS5
3 | 17 8
UCHL3 binds COPS5.
3 | 7 8
3 | 7 8

reach
"Thus, we demonstrate that UCHL3 regulates Cullin1 neddylation dependently on COPS5.To further understand the relationship of UCHL3 and COPS5, we analyzed the interaction between UCHL3 and COPS5."

reach
"Furthermore, we found that endogenous UCHL3 can also interact with endogenous COPS5."

reach
"Given that we demonstrated that UCHL3 promoted activation of IFN signaling, we observed whether IFN-I could affect the interaction between UCHL3 and COPS5."

reach
"The result showed that IFNalpha promoted the binding of UCHL3 to COPS5 during the early stage of IFNalpha treatment, which is consistent with the effect of UCHL3 on TYK2 phosphorylation during the ear[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We demonstrate that UCHL3 physically interacts with COPS5, and increases COPS5 levels and protein stability by deubiquitinating COPS5."

sparser
"Furthermore, we found that endogenous UCHL3 can also interact with endogenous COPS5 ( Fig. 4 B)."

sparser
"Given that we demonstrated that UCHL3 promoted activation of IFN signaling, we observed whether IFN-I could affect the interaction between UCHL3 and COPS5."

sparser
"Next, we find that UCHL3 physically interacts with COPS5, and upregulates COPS5 levels and protein stability by deubiquitinating COPS5."

sparser
"The result showed that IFNα promoted the binding of UCHL3 to COPS5 during the early stage of IFNα treatment ( Fig. 4 B), which is consistent with the effect of UCHL3 on TYK2 phosphorylation during the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Given that UCHL3 interacts with COPS5 in IFN signaling and UCHL3 is a deubiquitinase, we question whether UCHL3 promotes COPS5 deubiquitination."
UCHL3 increases the amount of COPS5.
| 4
UCHL3 bound to COPS5 increases the amount of COPS5. 2 / 2
| 2

reach
"Next, we find that UCHL3 physically interacts with COPS5, and upregulates COPS5 levels and protein stability by deubiquitinating COPS5."

reach
"We demonstrate that UCHL3 physically interacts with COPS5, and increases COPS5 levels and protein stability by deubiquitinating COPS5."
UCHL3 increases the amount of COPS5. 2 / 2
| 2

reach
"In summary, our results demonstrate that UCHL3 upregulates the levels of SCF beta-TrCP substrates.We have demonstrated that UCHL3 positively regulated COPS5 levels and IFNAR1 levels."

reach
"Furthermore, we demonstrate that UCHL3 as a deubiquitinase can deubiquitinate COPS5, thereby upregulating COPS5 levels and protein stability (Figs. 4 and 5)."
UCHL3 deubiquitinates COPS5.
| 4
UCHL3 deubiquitinates COPS5. 4 / 4
| 4

reach
"These results imply that UCHL3 physically interacts with COPS5 and deubiquitinates COPS5.Given that UCHL3 can deubiquitinate COPS5, we speculate that UCHL3 could affect cellular levels of COPS5."

reach
"Given that UCHL3 interacts with COPS5 in IFN signaling and UCHL3 is a deubiquitinase, we question whether UCHL3 promotes COPS5 deubiquitination."

reach
"Furthermore, we demonstrate that UCHL3 as a deubiquitinase can deubiquitinate COPS5, thereby upregulating COPS5 levels and protein stability (Figs. 4 and 5)."

reach
"To address this question, we firstly performed an in vitro deubiquitination assay, which clearly demonstrated that UCHL3 can deubiquitinate COPS5 in vitro."
UCHL3 decreases the amount of COPS5.
| 2
UCHL3 decreases the amount of COPS5. 2 / 2
| 2

reach
"Since COPS5 have been demonstrated to downregulate Cullin1 neddylation and activity, we speculate that UCHL3 could downregulate the activity of SCF beta and TrCP complex by upregulating cellular COPS5[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"We found that UCHL3 knockdown noticeably upregulated ubiquitination levels of COPS5."
UCHL3 affects TDP1
1 | 14 10
UCHL3 binds TDP1.
1 | 4 10
1 | 3 10

sparser
"Immunoprecipitation of ectopic FLAG-TDP1 variants (WT, R361K, or R586K) showed a marked increase in the interaction of endogenous UCHL3 with ectopic R586K mutant TDP1, both in the presence and in the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Furthermore, compromised TDP1 proteostasis due to defective TDP1-R586 methylation impairs UCHL3-TDP1 binding and accumulates TDP1 in the cells (model)."

sparser
"Additionally, UCLH3 knockdown cells show enrichment of ubiquitylated-TDP1 R586K ( Figure 3 G), whereas overexpression of PRMT5 abrogates the binding of TDP1 to UCHL3 ( Figure 3 H), suggesting that met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To further test the PRMT5 dependence on the UCHL3-TDP1 binding, we overexpressed FLAG-PRMT5 and co-immunoprecipitated ectopic GFP-TDP1."

sparser
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we uncover the crosstalk between TDP1 arginine methylation at R586 and ubiquitylation, a determining factor for the association of TDP1 with UCHL3, thus acting as a switch for regulation of endo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report multiple evidence showing a direct role of UCHL3 in controlling TDP1 proteostasis: (1) reduction of K48-ubiquitylated TDP1 in cancer cells overexpressing UCHL3, (2) reduction of K48-uibiquitylated TDP1 upon addition of recombinant UCHL3, (3) enrichment of K48-ubiquitylated TDP1 upon MG132 treatment, which is dependent on UCHL3 levels, (4) epistatic relationship showing that co-depletion of TDP1 and UCHL3 does not further sensitize cells to TOP1 poisons than depletion of either enzyme alone, (5) physical interaction between TDP1 and UCHL3, and (6) changes in the ubiquitylation status and TDP1 turnover upon changes in UCHL3 levels induced by its ectopic overexpression, siRNA depletion, CRISPR editing, and pharmacologic inhibition and, also, in two physiologically relevant clinical settings."

sparser
"We detected a marked increase in the enrichment of ubiquitylated TDP1 R586K (Ub-TDP1 R586K ) in UCLH3 knockdown cells compared with UCHL3-proficient cells ( Figure 3 G, Ub-TDP1), consistent with methy[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Additionally, UCLH3 knockdown cells show enrichment of ubiquitylated-TDP1 R586K ( Figure 3 G), whereas overexpression of PRMT5 abrogates the binding of TDP1 to UCHL3 ( Figure 3 H), suggesting that met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 binds TDP1-R361K. 1 / 1
| 1

reach
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 ubiquitinates TDP1.
| 3
UCHL3 leads to the ubiquitination of TDP1. 3 / 3
| 3

reach
"Depletion of UCHL3 increases TDP1 ubiquitylation and turnover rate and sensitizes cells to TOP1 poisons."

reach
"In contrast, UCHL3 is downregulated in spinocerebellar ataxia with axonal neuropathy (SCAN1), causing elevated levels of TDP1 ubiquitylation and faster turnover rate."

reach
"To examine if the increased TDP1 ubiquitylation caused by UCHL3 depletion would lead to increased turnover, we monitored the TDP1 protein level following incubations with the protein synthesis inhibitor cycloheximide."
UCHL3 deubiquitinates TDP1.
| 3
UCHL3 deubiquitinates TDP1. 2 / 2
| 2

reach
"In addition, UCHL3 deubiquitinates and stabilizes TDP1 to promote the repair of TOP1-induced single-strand breaks , and UCHL3 also deubiquitinates KU80 to promote the retention of KU80 at DNA damage sites, thereby improving DNA repair by nonhomologous end joining (NHEJ) ."

reach
"In contrast, UCHL3 is downregulated in spinocerebellar ataxia with axonal neuropathy (SCAN1), causing elevated levels of TDP1 ubiquitylation and faster turnover rate."
Modified UCHL3 leads to the deubiquitination of TDP1. 1 / 1
| 1

reach
"Overexpression of UCHL3, but not a catalytically inactive mutant, suppresses TDP1 ubiquitylation and turnover rate."
UCHL3 decreases the amount of TDP1.
| 3
UCHL3 decreases the amount of TDP1. 3 / 3
| 3

reach
"Depletion of UCHL3 was sufficient to reduce TDP1 expression in rhabdomyosarcoma cells to levels comparable to control cells (XREF_FIG D, top) and led to hypersensitivity to the TOP1 poison CPT (XREF_FIG D, bottom)."

reach
"Depletion of UCHL3 in rhabdomyosarcoma cells reduced TDP1 levels and sensitized cells to TOP1 poisons."

reach
"UCHL3 decreases the ubiquitination level of TDP1 leading to the inhibition of TDP1 degradation."
UCHL3 increases the amount of TDP1.
| 1
Modified UCHL3 increases the amount of TDP1. 1 / 1
| 1

reach
"TDP1 overexpression in the topoisomerase therapy resistant rhabdomyosarcoma is driven by UCHL3 overexpression."
RAD51 affects UCHL3
5 | 8 10
5 | 8 10

No evidence text available

sparser
"Furthermore, wild-type RAD51 and the 3KR mutant bound equally with UCHL3, suggesting that RAD51 deubiquitination does not affect RAD51/UCHL3 interaction ()."

No evidence text available

No evidence text available

No evidence text available

sparser
"We found that S75A mutation did not affect the interaction between UCHL3 and RAD51 ( xref G)."

sparser
"UCHL3 interacts with RAD51 and de-ubiquitinates residues that disrupt the RAD51–BRCA2 interaction."

sparser
"We also performed co-IP experiments to examine the changes in the interaction between RAD51 and UCHL3 in HEK293 cells that were treated with farrerol, and we excluded the possibility that farrerol promoted the deubiquitination of RAD51 by enhancing the UCHL3-RAD51 interaction (Supplementary Fig.  xref )."

sparser
"We performed in vitro and in vivo deubiquitination assay to establish that farrerol directly stimulates UCHL3 enzyme activity and promotes HR by UCHL3-RAD51 axis."

reach
"Here we carried out a systematic screen of DUBs for HR and established that UCHL3 interacts with and deubiquitinates RAD51 and promotes binding between RAD51 and BRCA2."
ATM affects UCHL3
1 4 1 | 1 9 5
ATM phosphorylates UCHL3.
1 2 | 9 5
ATM phosphorylates UCHL3. 10 / 16
1 | 9 5

sparser
"These results suggest that UCHL3 is phosphorylated by ATM following DNA damage."

reach
"These results suggest that UCHL3 is phosphorylated by ATM following DNA damage."

reach
"Together, these results suggest that UCHL3 phosphorylation by ATM is important for RAD51 focus formation, HR, and radiosensitivity.Next, we further investigated how UCHL3 phosphorylation affects its function."

reach
"To test whether UCHL3 could be phosphorylated by ATM/ATR, we examined UCHL3 phosphorylation using an antibody against the consensus of ATM/ATR phosphorylation sites (anti-phospho-SQ/TQ [anti-pSQ/TQ]) following DNA damage."

reach
"ATM-dependent phosphorylation of the deubiquitinase UCHL3 enables UCHL3 to enhance the retention of Ku80 at damage sites by counteracting Ku80 ubiquitination (Nishi et al., 2018)."

sparser
"In addition, the polyubiquitin chains on the substrate RAD51 can be cleaved by ATM-phosphorylated UCHL3 instead of simulated phosphorylated UCHL3 in vivo."

reach
"In response to DNA damage, UCHL3 is phosphorylated and activated by ATM, and activated UCHL3 positively regulates HR by deubiquitinating RAD51 to promote the BRCA2-RAD51 interaction ."

"Mechanistically, in response to DNA damage, the deubiquitinase UCHL3 is phosphorylated and activated by ATM."

reach
"These results demonstrate that UCHL3 phosphorylation by ATM is important for UCHL3 activation following DNA damage."

sparser
"Additionally, ATM-induced phosphorylation of UCHL3 occurs after DNA damage, which in turn deubiquitinates RAD51 and promotes the binding between BRCA2 and RAD51 ( xref )."
ATM phosphorylates UCHL3 on S75. 2 / 2
1 1 |

"The deubiquitinase UCHL3 is phosphorylated and activated by ATM. UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2. |Mutation of S75 (S75A) abolished the pSQ/TQ signal, suggesting that S75 is a major ATM phosphorylation site following DNA damage"

No evidence text available
ATM activates UCHL3.
2 | 1
ATM activates UCHL3. 3 / 4
2 | 1

trips
"Mechanistically, in response to DNA damage, the deubiquitinase UCHL3 is phosphorylated and activated by ATM."

"The deubiquitinase UCHL3 is phosphorylated and activated by ATM. UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2. |Mutation of S75 (S75A) abolished the pSQ/TQ signal, suggesting that S75 is a major ATM phosphorylation site following DNA damage"

"Mechanistically, in response to DNA damage, the deubiquitinase UCHL3 is phosphorylated and activated by ATM."
ATM binds UCHL3.
1 |
1 |

No evidence text available
UCHL3 affects EEF1A1
| 15 7
UCHL3 binds EEF1A1.
| 6 7
| 4 7

reach
"Our findings revealed that UCHL3 binds and stabilizes EEF1A1 through deubiquitination."

reach
"Our above findings showed that UCHL3 could bind to EEF1A1 and mediated its deubiquitination."

reach
"Therefore, we aimed to elucidate the underlying molecular mechanism of UCHL3 binding to EEF1A1."

reach
"To further illustrated the effect of the binding between EEF1A1 and UCHL3 on the function of UCHL3, we conducted transwell and sphere formation assays."

sparser
"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

sparser
"Our findings revealed that UCHL3 binds and stabilizes EEF1A1 through deubiquitination."

sparser
"UCHL3 and EEF1A1 could bind with each other in both HEP3B and HUH7 cells."

sparser
"EEF1A1 binds to UCHL3 through the lysine site."

sparser
"Our above findings showed that UCHL3 could bind to EEF1A1 and mediated its deubiquitination."

sparser
"Therefore, we aimed to elucidate the underlying molecular mechanism of UCHL3 binding to EEF1A1."
UCHL3 binds EEF1A1 and lysine. 2 / 2
| 2

reach
"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

reach
"EEF1A1 binds to UCHL3 through the lysine site."
UCHL3 increases the amount of EEF1A1.
| 4
UCHL3 increases the amount of EEF1A1. 4 / 4
| 4

reach
"IHC analysis of the tumor in nude mouse tumors showed that UCHL3 overexpression increased EEF1A1 protein level, while UCHL3 inhibition downregulated EEF1A1 protein level (Fig. 2G-J)."

reach
"The experimental results showed that UCHL3 knockdown reduced the protein level of EEF1A1, and the migration ability and sphere formation ability were decreased, which was alleviated in mutated EEF1A1 group (Fig. 4G-H)."

reach
"The results showed that UCHL3 depletion decreased EEF1A1 protein expression, whereas UCHL3 overexpression upregulated EEF1A1 expression (Fig. 2D)."

reach
"UCHL3 upregulates the expression of EEF1A1 via deubiquitination, facilitating the migration of HCC cells through lysine-based interactions."
UCHL3 decreases the amount of EEF1A1.
| 3
UCHL3 decreases the amount of EEF1A1. 2 / 2
| 2

reach
"IHC analysis of the tumor in nude mouse tumors showed that UCHL3 overexpression increased EEF1A1 protein level, while UCHL3 inhibition downregulated EEF1A1 protein level (Fig. 2G-J)."

reach
"On the contrary, overexpression of UCHL3 reduce the ubiquitination level of EEF1A1(Fig. 3D)."
UCHL3 bound to lysine and EEF1A1 decreases the amount of EEF1A1. 1 / 1
| 1

reach
"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."
UCHL3 activates EEF1A1.
| 2
UCHL3 activates EEF1A1. 2 / 2
| 2

reach
"Knockdown of UCHL3 could promote the degradation of EEF1A1, and overexpression of UCHL3 increased the stability of EEF1A1 (Fig. 3F-I)."

reach
"UCHL3 promotes hepatocellular carcinoma progression by stabilizing EEF1A1 through deubiquitination."
UCHL3 affects SMAD1
2 1 | 1 14
UCHL3 binds SMAD1.
2 | 1 14
2 | 1 14

sparser
"It has been reported that UCHL3 can interact with SMAD1, a classical osteogenesis-related molecule involved in BMSC osteogenesis [ xref , xref , xref ]."

sparser
"The protein‒protein docking results from Alphafold 2 revealed that UCHL3 could directly bind with SMAD1 through hydrogen bonds (Fig.  xref i)."

sparser
"Further immunofluorescence and co-immunoprecipitation (Co-IP) verified the interaction between UCHL3 and SMAD1 in BMSCs (Fig.  xref j–l)."

sparser
"Consistently, our data showed that UCHL3 physically interacted with SMAD1 and that its expression in BMSCs was positively correlated with SMAD1."

sparser
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

sparser
"Furthermore, it wondered whether the endogenous Smad1 interacts with Uch-L3."

sparser
"Furthermore, the endogenous Uch-L3 was also interacted with the endogenous Smad1 in C2C12 cells as confirmed by co-IP against anti-Uch-L3 antibody and WB using anti-Smad1 antibody ( Fig. 1 E)."

sparser
"All of these results showed that Uch-L3 interacts with Smad1."

sparser
"After confirming the interaction between Smad1 and Uch-L3, we tested whether the deubiquitinating activity of Uch-L3 is able to affect the ubiquitination pattern of Smad1."

sparser
"UCH-L3 physically interacts with Smad1 and stabilizes it by deubiquitylating its polyubiquitin."
UCHL3 deubiquitinates SMAD1.
1 |
UCHL3 deubiquitinates SMAD1. 1 / 1
1 |

"<span class="match term0">Ubiquitin C-terminal hydrolase-L3</span> regulates <span class="match term1">Smad1</span> ubiquitination and osteoblast differentiation"
UCHL3 affects POLD4
| 12 6
UCHL3 binds POLD4.
| 4 6
| 4 6

sparser
"Tumorsphere formation and in vitro limiting dilution assay were performed to investigate the effect of UCHL3-POLD4 on GSC self-renewal."

sparser
"Apoptosis, TUNEL, cell cycle phase distribution, modification of the Single Cell Gel Electrophoresis (Comet), γ-H2AX immunofluorescence, and colony formation assays were conducted to evaluate the influence of UCHL3-POLD4 on GSC in ionizing radiation."

sparser
"Both in vitro and in vivo assays indicated that targeted depletion of the UCHL3-POLD4 axis reduced GSC self-renewal and tumorigenic capacity and resistance to IR treatment by impairing homologous recombination (HR) and nonhomologous end joining (NHEJ)."

sparser
"These findings reveal a new signaling axis for GSC PMT regulation and highlight UCHL3-POLD4 as a potential therapeutic target in GBM."

reach
"UCHL3 interacted with, depolyubiquitinated, and stabilized POLD4."

reach
"UCHL3 interacted with, stabilized and depolyubiquitylated POLD4."

reach
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig. 5D, E)."

sparser
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig.  xref D, E)."

sparser
"It has been established that the deubiquitinase UCHL3 binds and stabilizes p12 in GSCs."

reach
"It has been established that the deubiquitinase UCHL3 binds and stabilizes p12 in GSCs."
UCHL3 deubiquitinates POLD4.
| 5
UCHL3 deubiquitinates POLD4. 5 / 5
| 5

reach
"UCHL3 induces radiation resistance and acquisition of mesenchymal phenotypes by deubiquitinating POLD4 in glioma stem cells."

reach
"In view of the essential role of the enzyme activity of UCHL3 in driving POLD4 stabilization, we proposed that UCHL3 may deubiquitylate POLD4."

reach
"In MES GSCs, knockdown of UCHL3 increased the levels of POLD4 ubiquitination (Fig. 5J)."

reach
"Based on a DUB-siRNA profiling screen, UCHL3, as a unique DUB of POLD4, was confirmed to bind directly to, stabilizes, and deubiquitinates POLD4 in GSCs."

reach
"It has been shown that UCHL3 induces radiation resistance and the acquisition of mesenchymal phenotypes by deubiquitinating p12 in glioma stem cells."
UCHL3 increases the amount of POLD4.
| 3
UCHL3 increases the amount of POLD4. 3 / 3
| 3

reach
"Indeed, we found that silencing UCHL3 reduced POLD4 protein expression (not mRNA), which was reversed by the UPS inhibitor MG132 (Fig. 5F and Fig. S6K)."

reach
"Overexpression of wild-type UCHL3 (WT-UCHL3), but not the catalytically dead mutant S75A, induced elevated POLD4 levels in a dose-dependent manner in HEK293T cells (Fig. 5H), implying that the enzyme activity of UCHL3 was crucial for stable POLD4."

reach
"Stabilization of POLD4 assays revealed that ectopic expression of K48R (not WT) ubiquitin restored the POLD4 expression inhibited by UCHL3 depletion in MES GSCs (Fig. 5M)."
UCHL3 affects GARS1
| 18
UCHL3 binds GARS1.
| 5
| 5

reach
"The protein‒protein docking results from Alphafold 2 revealed that UCHL3 could directly bind with SMAD1 through hydrogen bonds (Fig. 7i)."

reach
"Further immunofluorescence and co-immunoprecipitation (Co-IP) verified the interaction between UCHL3 and SMAD1 in BMSCs (Fig. 7j–l)."

reach
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

reach
"All of these results showed that Uch-L3 interacts with Smad1.After confirming the interaction between Smad1 and Uch-L3, we tested whether the deubiquitinating activity of Uch-L3 is able to affect the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The physical interaction between Uch-L3 and Smad1 was revealed."
UCHL3 activates GARS1.
| 5
UCHL3 activates GARS1. 5 / 5
| 5

reach
"Furthermore, we wondered how UCHL3 modulated the function of SMAD1 in BMSCs."

reach
"These results suggest that the reduction of Uch-L3 decreases the active pSmad1 signal and thus reduces the osteoblast differentiation.This study showed a novel function for Uch-L3 in osteoblast differ[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Moreover, although we found that MS-BMDM-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1, additional questions, such as binding sites, still require clarification."

reach
"This study suggests that mechanical force-induced macrophage-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1, thereby promoting alveolar bone formation during OTM."

reach
"MS-BMDM-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1."
UCHL3 deubiquitinates GARS1.
| 3
UCHL3 deubiquitinates GARS1. 3 / 3
| 3

reach
"In this study, it was found Smad1 is physically interacted with Uch-L3, which deubiquitinates Smad1."

reach
"That is, Smad1 poly-ubiquitination degrees were decreased by the forced expression of Uch-L3."

reach
"Importantly, UCH-L3 deubiquitylates Smad1 and enhances osteoblast differentiation [73]."
UCHL3 decreases the amount of GARS1.
| 3
UCHL3 decreases the amount of ubiquitinated GARS1. 2 / 2
| 2

reach
"It has been reported that UCHL3, which decreases the amount of polyubiquitinated SMAD1, might be critical for osteogenesis [34]."

reach
"The activity of deubiquitinating enzyme Uch-L3 dramatically reduced the amount of ubiquitinated Smad1."
UCHL3 decreases the amount of GARS1. 1 / 1
| 1

reach
"In addition, inhibition of endogenous UCHL3 by TCID elevated the ubiquitination level of endogenous SMAD1 in BMSCs (Fig. 7m), which indicated that UCHL3 could elevate SMAD1 protein stabilization through deubiquitination."
UCHL3 ubiquitinates GARS1.
| 2
UCHL3 leads to the ubiquitination of GARS1. 2 / 2
| 2

reach
"This finding indicated that UCHL3 promotes BMSCs osteogenesis by decreasing SMAD1 ubiquitination degradation.Notably, we only characterized the protein profile in MS-BMDM-EXOs."

reach
"Ubiquitin C-terminal hydrolase-L3 (UCHL3) regulates BMP2-induced Smad1 polyubiquitination to promote osteoblast differentiation (65)."
SMAD1 affects UCHL3
2 | 1 14
2 | 1 14

sparser
"It has been reported that UCHL3 can interact with SMAD1, a classical osteogenesis-related molecule involved in BMSC osteogenesis [ xref , xref , xref ]."

sparser
"The protein‒protein docking results from Alphafold 2 revealed that UCHL3 could directly bind with SMAD1 through hydrogen bonds (Fig.  xref i)."

sparser
"Further immunofluorescence and co-immunoprecipitation (Co-IP) verified the interaction between UCHL3 and SMAD1 in BMSCs (Fig.  xref j–l)."

sparser
"Consistently, our data showed that UCHL3 physically interacted with SMAD1 and that its expression in BMSCs was positively correlated with SMAD1."

sparser
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

sparser
"Furthermore, it wondered whether the endogenous Smad1 interacts with Uch-L3."

sparser
"Furthermore, the endogenous Uch-L3 was also interacted with the endogenous Smad1 in C2C12 cells as confirmed by co-IP against anti-Uch-L3 antibody and WB using anti-Smad1 antibody ( Fig. 1 E)."

sparser
"All of these results showed that Uch-L3 interacts with Smad1."

sparser
"After confirming the interaction between Smad1 and Uch-L3, we tested whether the deubiquitinating activity of Uch-L3 is able to affect the ubiquitination pattern of Smad1."

sparser
"UCH-L3 physically interacts with Smad1 and stabilizes it by deubiquitylating its polyubiquitin."
COPS5 affects UCHL3
3 | 7 8
3 | 7 8

reach
"Thus, we demonstrate that UCHL3 regulates Cullin1 neddylation dependently on COPS5.To further understand the relationship of UCHL3 and COPS5, we analyzed the interaction between UCHL3 and COPS5."

reach
"Furthermore, we found that endogenous UCHL3 can also interact with endogenous COPS5."

reach
"Given that we demonstrated that UCHL3 promoted activation of IFN signaling, we observed whether IFN-I could affect the interaction between UCHL3 and COPS5."

reach
"The result showed that IFNalpha promoted the binding of UCHL3 to COPS5 during the early stage of IFNalpha treatment, which is consistent with the effect of UCHL3 on TYK2 phosphorylation during the ear[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We demonstrate that UCHL3 physically interacts with COPS5, and increases COPS5 levels and protein stability by deubiquitinating COPS5."

sparser
"Furthermore, we found that endogenous UCHL3 can also interact with endogenous COPS5 ( Fig. 4 B)."

sparser
"Given that we demonstrated that UCHL3 promoted activation of IFN signaling, we observed whether IFN-I could affect the interaction between UCHL3 and COPS5."

sparser
"Next, we find that UCHL3 physically interacts with COPS5, and upregulates COPS5 levels and protein stability by deubiquitinating COPS5."

sparser
"The result showed that IFNα promoted the binding of UCHL3 to COPS5 during the early stage of IFNα treatment ( Fig. 4 B), which is consistent with the effect of UCHL3 on TYK2 phosphorylation during the[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Given that UCHL3 interacts with COPS5 in IFN signaling and UCHL3 is a deubiquitinase, we question whether UCHL3 promotes COPS5 deubiquitination."
Farrerol affects UCHL3
| 17
Farrerol binds UCHL3.
| 9

reach
"The results revealed a strong interaction between farrerol and UCHL3, and the dissociation constant (Kd) of farrerol/UCHL3 reached a low of 36.73 nM (Fig. 1e and Supplementary Fig. 1d)."

reach
"Farrerol binds to UCHL3 through two amino acid residues, Lys187 and Arg215."

reach
"To elucidate how farrerol specifically binds to UCHL3, we crystallized UCHL3 in the presence of farrerol and determined the structure of the complex at 1."

reach
"The alignment indicated that farrerol did not affect the dynamic structure of UCHL3.Then, we attempted to confirm whether the two amino acid residues were essential for the interaction between UCHL3 and farrerol through the use of CETSAs."

reach
"In contrast to the UCHL3 WT, an improvement in thermal stability was not observed for the two mutants (Fig. 2d–g), indicating that farrerol directly binds to UCHL3 through the two amino acid residues."

reach
"Further SPR assay with the purified recombinant UCHL3–K187A R215A mutant protein and farrerol confirmed that mutating the two amino acid sites abolished the binding of farrerol to UCHL3 (Supplementary Fig. 1f)."

reach
"Since both farrerol and UCHL3 participate in regulating HR by improving the recruitment of RAD51 to DNA damage sites and because we demonstrated that farrerol binds to UCHL3 with a high affinity, we wondered whether farrerol stimulated HR in a UCHL3-dependent fashion."

reach
"Taken together, these data indicated that farrerol promoted DNA repair by HR in a UCHL3-dependent manner.Since the cocrystal structure of the UCHL3farrerol complex revealed that farrerol directly binds to UCHL3 (Fig. 2a–c), a ubiquitin–AMC assay was then performed to determine whether farrerol activates UCHL3 enzymatic activity, and we found that farrerol affected both K and K , by slightly increasing K from 47.72 to 56.27 nM and significantly increasing K from 11.99 to 19.64 s (Fig. 3i), suggesting that farrerol activates UCHL3 deubiquitinase activity."

reach
"Since we demonstrated that farrerol binds to UCHL3 through two amino acid residues to activate its enzymatic activity, in order to treat HR-overactivated cancer or overcome radio/chemo-resistance, it is likely worthwhile to try generating farrerol derivatives that exhibit an inhibitory effect on UCHL3 while still retaining a high affinity to UCHL3.It was reported that genomic stability is critical in the dynamic process of somatic cells reprogramming into iPSCs and is essential for the pluripotency of ESCs and iPSCs, whereas chromosomal aneuploidy results in a sharp decrease in cell differentiation potentials in vivo ."
Farrerol activates UCHL3.
| 8
| 8

reach
"Further thorough studies on whether farrerol directly targets these proteins and the underlying mechanisms might open up new avenues for the application of farrerol when treating other types of diseases.Notably, several reports also indicated that UCHL3 is aberrantly overexpressed in different types of tumors, so caution should be taken when utilizing farrerol, as the farrerol-mediated activation of UCHL3 might be dangerous for cancer patients."

reach
"Experiments on both in vitro and in vivo functions indicate that farrerol promotes the catalytic activity of UCHL3, thereby leading to a reduced ubiquitination level of RAD51 and enhanced HR repair."

reach
"Farrerol activates the enzymatic activity of UCHL3 to promote HR repair."

reach
"Taken together, these data indicated that farrerol promoted DNA repair by HR in a UCHL3-dependent manner.Since the cocrystal structure of the UCHL3farrerol complex revealed that farrerol directly binds to UCHL3 (Fig. 2a–c), a ubiquitin–AMC assay was then performed to determine whether farrerol activates UCHL3 enzymatic activity, and we found that farrerol affected both K and K , by slightly increasing K from 47.72 to 56.27 nM and significantly increasing K from 11.99 to 19.64 s (Fig. 3i), suggesting that farrerol activates UCHL3 deubiquitinase activity."

reach
"Moreover, we also performed a Ub-CHOP2 assay to test whether farrerol directly regulates the enzymatic activity of UCHL3."

reach
"Farrerol directly activates the deubiqutinase UCHL3 to promote DNA repair and reprogramming when mediated by somatic cell nuclear transfer."

reach
"The data indicated that farrerol at a concentration of 0.1 μM significantly increased the deubiquitinase activity of UCHL3 (Supplementary Fig. 3a)."

reach
"We performed in vitro and in vivo deubiquitination assay to establish that farrerol directly stimulates UCHL3 enzyme activity and promotes HR by UCHL3-RAD51 axis."
UCHL3 affects XRCC5
3 | 11 3
UCHL3 binds XRCC5.
3 | 2 3
3 | 2 3

sparser
"UCHL3 interacts with Ku80 and is recruited to DNA damage sites in a manner dependent on its catalytic activity."

No evidence text available

No evidence text available

sparser
"Given the findings that UCHL3 interacted with Ku80 and was recruited to sites of damage in a catalytic activity dependent manner (Fig.  xref , Supplementary Fig.  xref ), we investigated whether catalytic activity is required for its phosphorylation."

No evidence text available

reach
"Accordingly, the interaction of Ku80 and UCHL3, which requires UCHL3 catalytic activity (Fig. 1E), was not markedly affected when FLAG-tagged phospho-mutants of UCHL3 were examined by immunoprecipitation (Supplementary Fig. S7B)."

reach
"Here we found that UCHL3, a deubiquitylating enzyme, interacts with and can directly deubiquitylate Ku80 in a catalytic activity dependent manner."

sparser
"Accordingly, the interaction of Ku80 and UCHL3, which requires UCHL3 catalytic activity (Fig.  xref ), was not markedly affected when FLAG-tagged phospho-mutants of UCHL3 were examined by immunoprecipitation (Supplementary Fig.  xref )."
UCHL3 deubiquitinates XRCC5.
| 6
UCHL3 deubiquitinates XRCC5. 6 / 6
| 6

reach
"UCHL3 de-ubiquitinates and stabilises Ku80 which is required for Ku80 retention at DSBs [10]."

reach
"In addition, UCHL3 deubiquitinates and stabilizes TDP1 to promote the repair of TOP1-induced single-strand breaks , and UCHL3 also deubiquitinates KU80 to promote the retention of KU80 at DNA damage sites, thereby improving DNA repair by nonhomologous end joining (NHEJ) ."

reach
"UCHL3 enhances Ku80 chromatin binding and directly deubiquitylates Ku80."

reach
"Taken together, these data suggest that UCHL3 could enhance Ku chromatin retention upon DSB induction by deubiquitylating Ku80."

reach
"Conversely, the ubiquitin carboxyl-terminal hydrolase L3 (UCHL3) can directly deubiquitylate Ku80."

reach
"Here we found that UCHL3, a deubiquitylating enzyme, interacts with and can directly deubiquitylate Ku80 in a catalytic activity dependent manner."
UCHL3 activates XRCC5.
| 2
UCHL3 activates XRCC5. 2 / 2
| 2

reach
"ATM-dependent phosphorylation of the deubiquitinase UCHL3 enables UCHL3 to enhance the retention of Ku80 at damage sites by counteracting Ku80 ubiquitination (Nishi et al., 2018)."

reach
"In accordance with reduced Ku80 IRIF, we observed that knockout and knockdown of UCHL3 reduced DSB-induced chromatin binding of Ku80 using chromatin fractionation until 3 hours after phleomycin removal (Fig. 4C and Supplementary Fig. S5)."
UCHL3 ubiquitinates XRCC5.
| 1
UCHL3 leads to the ubiquitination of XRCC5. 1 / 1
| 1

reach
"Since it was reported that Ku ubiquitylation is a prerequisite for Ku removal from chromatin, we hypothesized that UCHL3 may enhance Ku80 chromatin binding by antagonizing Ku80 ubiquitylation."
CTNNB1 affects UCHL3
2 | 7 8
CTNNB1 binds UCHL3.
2 | 6 8
2 | 6 8

sparser
"These results suggest that UCHL3 might physically interact with CTNNB1 to trigger activation of the Wnt signaling pathway."

sparser
"The interaction between UCH-L3 and β-catenin proteins was confirmed, and it reduced the ubiquitination of β-catenin."

sparser
"Hence, we evaluated the interaction between UCHL3 and CTNNB1."

reach
"A Pierce Co-Immunoprecipitation Kit (Thermo Fisher, USA) was used to verify the physical interaction between UCHL3 and CTNNB1."

sparser
"Interestingly, we found that UCHL3 interacted with CTNNB1, as shown by co-IP and colocalization assays with T24 cells."

sparser
"To verify the interaction between UCHL3 and CTNNB1 in bladder cancer cells, the T24 cell line was selected for an endogenous immunoprecipitation (IP) analysis."

reach
"UCHL3 interacts with and stabilizes CTNNB1."

sparser
"The results showed that UCHL3 binds endogenous CTNNB1 under pathological conditions (Fig.  xref B)."

sparser
"The interaction between UCH-L3 and β-catenin proteins was observed, resulting in reduced ubiquitination and increased protein stability of β-catenin."

reach
"Hence, we evaluated the interaction between UCHL3 and CTNNB1."
CTNNB1 increases the amount of UCHL3.
| 1
CTNNB1 increases the amount of UCHL3. 1 / 1
| 1

reach
"Western blot analyses revealed that HA-CTNNB1 expression gradually increased with the increased expression of FLAG-UCHL3, and this increase was dose dependent (Fig. 4D)."
UCHL3 affects Neoplasms
| 16
| 16

reach
"In this study, we showed that UCHL3 overexpression increased tumor stem-like properties."

reach
"Importantly, stable knockdown of UCHL3 suppressed tumor growth from H358 cells and inhibited tumor stem-like properties."

reach
"UCHL3 promoted tumor stem-like properties through stabilization of AhR."

reach
"Additionally, UCHL3 and EEF1A1 enhance the growth capacity of subcutaneous tumors in nude mice."

reach
"GSK3β and UCHL3 govern RIPK4 homeostasis via deubiquitination to enhance tumor metastasis in ovarian cancer."

reach
"Finally, UCHL3 functions to promote tumor growth and lung cancer stem-like properties through AhR.Disorder in the deubiquitination process is usually related to tumorigenesis."

reach
"Compared with the injection of vector-transfected A549 cells, injection of UCHL3-overexpressing A549 cells significantly increased tumor size, volume, and weight after 1 month of growth (Fig. 2l–n), while the overall body weight was maintained (Supplementary Fig. S2c)."

reach
"Our data demonstrated the tumor-promoting function of UCHL3 in bladder cancer, suggesting that targeting UCHL3 might be a potential approach for preventing bladder tumorigenesis and progression.Previously, UCHL3 has been found to be robustly expressed in epithelial ovarian cancer and related to intra-abdominal metastases [29]."

reach
"Ubiquitin carboxyl-terminal hydrolase L3 (UCHL3) has been reported to promote diverse tumors, but little is known about its role in HCC."

reach
"The mechanism by which UCHL3 promotes tumor stem-like properties through AhR requires further investigation.In conclusion, UCHL3 is an AhR DUB that promotes lung cancer proliferation, tumor growth and tumor stem-like properties through stabilizing AhR by its deubiquitination."
UCHL3 affects NEDD8
5 1 | 7 2
UCHL3 binds NEDD8.
5 1 | 3 2
5 1 | 3 2

No evidence text available

reach
"In vitro GST pull-down assay demonstrated that UCH-L3 bound to both NEDD8 and ubiquitin."

sparser
"18 These results suggest that Nedd8 may interact closely with UCH-L3 during testicular atrophy and germ cell loss."

reach
"While UCHL1 and UCHL3 both bind to NEDD8, the hydrolytic processing of NEDD8 is carried out by UCHL3 (Wada et al., 1998)."

reach
"UCH-L3, one of the ubiquitin C-terminal hydrolases, also binds to NEDD8 and functions as a NEDD8 C-terminal hydrolase (Wada et al., 1998)."

No evidence text available

sparser
"These observations suggest that UCH-L3 may function as a deneddylating enzyme 16 in vivo , although further studies are necessary to clarify whether UCH-L3 interacts with Nedd8 during spermatogenesis."

No evidence text available

No evidence text available

No evidence text available
UCHL3 increases the amount of NEDD8.
| 2
UCHL3 increases the amount of NEDD8. 2 / 2
| 2

reach
"The results of the present study demonstrated that UCHL3 knockdown in melanoma cells not only inhibited NEDD8 expression, but also decreased the ubiquitination of NEDD8."

reach
"RT-qPCR analysis demonstrated no significant differences in UCHL3, NEDD8 and LC3B expression levels between the NC and si-NC groups (P>0.05; Fig. 4A and B); however, UCHL3 knockdown significantly decreased the expression levels of UCHL3 and NEDD8, and significantly increased LC3B expression in the si-UCHL3 group compared with the NC group (all P<0.001; Fig. 4A and B)."
UCHL3 ubiquitinates NEDD8.
| 1
UCHL3 leads to the ubiquitination of NEDD8. 1 / 1
| 1

reach
"The results of the present study demonstrated that UCHL3 knockdown in melanoma cells not only inhibited NEDD8 expression, but also decreased the ubiquitination of NEDD8."
UCHL3 activates NEDD8.
| 1
UCHL3 activates NEDD8. 1 / 1
| 1

reach
"It was reported that UCHL3 enables to cleave the C-terminus of NEDD8, which is a ubiquitin like protein that exerts the function of Ub to be conjugated to a lysine residue of the substrate."

reach
"UCHL3 promotes proliferation of colorectal cancer cells by regulating SOX12 via AKT and mTOR signaling pathway."

reach
"Regulating biological functions, UCHL3 promoted GSC proliferation, tumorigenesis, self-renewal, and radioresistance in a POLD4-dependent manner.The degradation of POLD4 by ubiquitin ligases in response to DNA damage and during cell cycle progression plays a less prominent role in normal cells [43], while with increasing insights into the complex mechanisms of DNA damage repair, POLD4 is projected to play an increasingly important role."

reach
"UCH-L3 promotes non small cell lung cancer proliferation via accelerating cell cycle and inhibiting cell apoptosis."

reach
"Furthermore, autophagy was significantly enhanced along with the decrease in NEDD8 ubiquitination.In conclusion, the results of the present study demonstrated that UCHL3 knockdown decreased melanoma cell proliferation by increasing cell autophagy through regulating NEDD8 expression and autophagosome numbers in vitro."

reach
"UCHL3 promoted pancreatic cancer cell proliferation and migration by deubiquitinating FOXM1 (20)."

reach
"UCHL3 depletion repressed cell proliferation and migration both in vitro and in vivo."

reach
"UCH-L3 knockdown significantly inhibited the proliferation of NSCLC cells, whereas UCH-L3 overexpression had the opposite effect."

reach
"Moreover, UCH-L3 promoted NSCLC cells proliferation via accelerating cell cycle and inhibiting cell apoptosis."

reach
"Colony formation assays indicated that POLD4 overexpression restored the proliferation ability reduced by UCHL3 depletion in the case of IR (Fig. 6C)."

reach
"As expected, overexpressing UCHL3 favored the proliferative and migratory activity of bladder cancer cells, and UCHL3 depletion caused reduced bladder cancer cell proliferation and migration."

reach
"Genetic ablation of UCHL3 was found to markedly block ovarian cancer cell proliferation, viability and migration both in vitro and in vivo."

reach
"Abrogation of UCHL3 impairs the proliferation and migration abilities of bladder cancer cells."
TDP1 affects UCHL3
1 | 4 10
1 | 3 10

sparser
"Immunoprecipitation of ectopic FLAG-TDP1 variants (WT, R361K, or R586K) showed a marked increase in the interaction of endogenous UCHL3 with ectopic R586K mutant TDP1, both in the presence and in the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Furthermore, compromised TDP1 proteostasis due to defective TDP1-R586 methylation impairs UCHL3-TDP1 binding and accumulates TDP1 in the cells (model)."

sparser
"Additionally, UCLH3 knockdown cells show enrichment of ubiquitylated-TDP1 R586K ( Figure 3 G), whereas overexpression of PRMT5 abrogates the binding of TDP1 to UCHL3 ( Figure 3 H), suggesting that met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To further test the PRMT5 dependence on the UCHL3-TDP1 binding, we overexpressed FLAG-PRMT5 and co-immunoprecipitated ectopic GFP-TDP1."

sparser
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we uncover the crosstalk between TDP1 arginine methylation at R586 and ubiquitylation, a determining factor for the association of TDP1 with UCHL3, thus acting as a switch for regulation of endo[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Here, we report multiple evidence showing a direct role of UCHL3 in controlling TDP1 proteostasis: (1) reduction of K48-ubiquitylated TDP1 in cancer cells overexpressing UCHL3, (2) reduction of K48-uibiquitylated TDP1 upon addition of recombinant UCHL3, (3) enrichment of K48-ubiquitylated TDP1 upon MG132 treatment, which is dependent on UCHL3 levels, (4) epistatic relationship showing that co-depletion of TDP1 and UCHL3 does not further sensitize cells to TOP1 poisons than depletion of either enzyme alone, (5) physical interaction between TDP1 and UCHL3, and (6) changes in the ubiquitylation status and TDP1 turnover upon changes in UCHL3 levels induced by its ectopic overexpression, siRNA depletion, CRISPR editing, and pharmacologic inhibition and, also, in two physiologically relevant clinical settings."

sparser
"We detected a marked increase in the enrichment of ubiquitylated TDP1 R586K (Ub-TDP1 R586K ) in UCLH3 knockdown cells compared with UCHL3-proficient cells ( Figure 3 G, Ub-TDP1), consistent with methy[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Additionally, UCLH3 knockdown cells show enrichment of ubiquitylated-TDP1 R586K ( Figure 3 G), whereas overexpression of PRMT5 abrogates the binding of TDP1 to UCHL3 ( Figure 3 H), suggesting that met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 binds TDP1-R361K. 1 / 1
| 1

reach
"Unlike the methylation mutant TDP1 R586K , we detected reduced binding of UCHL3 with TDP1 WT or TDP1 R361K in the absence of CPT ( Figures 3 D and 3F), which corresponds to the enrichment of ubiquityl[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 1 12
| 1 11

reach
"The deubiquitinating enzyme UCHL3 promotes anaplastic thyroid cancer progression and metastasis through Hippo signaling pathway."

reach
"UCHL3 stimulates metastasis of gastric cancer through upregulating IGF2."

reach
"Collectively, UCHL3 can promote cancer progression and metastasis, indicating that UCHL3-specific inhibitors may have great therapeutic potential in HNSCC (152)."

reach
"GSK3β and UCHL3 govern RIPK4 homeostasis via deubiquitination to enhance tumor metastasis in ovarian cancer."

reach
"UCHL3 promotes cervical cancer development and metastasis by stabilizing NRF2 via deubiquitination."

eidos
"UCHL3 promotes cervical cancer development and metastasis by stabilizing NRF2 via deubiquitination ."

reach
"Our findings showed that knockdown of UCHL3 inhibited cell growth, migration and invasion in cervical cancer cells while UCHL3 knockdown inhibited cervical cancer development and metastasis in vivo in mouse models."

reach
"In cervical cancer studies, it has been found that UCHL3 promotes the development and metastasis of cervical cancer by stabilizing NRF2 through deubiquitination [120]."

reach
"Collectively, our findings suggest that UCHL3 promotes cervical cancer development and metastasis by stabilizing NRF2 via deubiquitination."

reach
"Therefore, we propose that the inhibition of UCH-L3 activity after anti-cancer therapy could suppress tumor remodeling and metastasis, providing a new strategy to improve the prognosis of gastric cancer patients."
| 1

reach
"Although the promotion of tumor progression by UCH-L3 has been identified in several reports (Miyoshi et al. 2006; Zhang et al. 2020), a previous study reported that UCH-L3 suppressed cancer metastasis through EMT inhibition in prostate cancer cells (Song et al. 2014)."
EEF1A1 affects UCHL3
| 7 7
EEF1A1 binds UCHL3.
| 6 7
| 4 7

reach
"Our findings revealed that UCHL3 binds and stabilizes EEF1A1 through deubiquitination."

reach
"Our above findings showed that UCHL3 could bind to EEF1A1 and mediated its deubiquitination."

reach
"Therefore, we aimed to elucidate the underlying molecular mechanism of UCHL3 binding to EEF1A1."

reach
"To further illustrated the effect of the binding between EEF1A1 and UCHL3 on the function of UCHL3, we conducted transwell and sphere formation assays."

sparser
"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

sparser
"Our findings revealed that UCHL3 binds and stabilizes EEF1A1 through deubiquitination."

sparser
"UCHL3 and EEF1A1 could bind with each other in both HEP3B and HUH7 cells."

sparser
"EEF1A1 binds to UCHL3 through the lysine site."

sparser
"Our above findings showed that UCHL3 could bind to EEF1A1 and mediated its deubiquitination."

sparser
"Therefore, we aimed to elucidate the underlying molecular mechanism of UCHL3 binding to EEF1A1."
UCHL3 binds EEF1A1 and lysine. 2 / 2
| 2

reach
"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

reach
"EEF1A1 binds to UCHL3 through the lysine site."
EEF1A1 activates UCHL3.
| 1
EEF1A1 activates UCHL3. 1 / 1
| 1

reach
"Notably, depleting EEF1A1 could reverse the impact of UCHL3 on the malignant behaviors of HCC cells and reduce the drug resistance of HCC to Lenvatinib."

reach
"On the other hand, UCH-L3 inhibited EMT and had the opposite effect of UCH-L1.The view that UCH-L1 and UCH-L3 regulate EMT suggests that these two DUBs might also be associated with the regulatory mechanism of the CSC-like characteristics in prostate cancer cells."

reach
"The reverse of the EMT event caused by UCH-L3 overexpression in PC3 coincided with a decrease of cell migration and invasion.UCH-L3 and UCH-L1 are very close relatives with high amino acid identity an[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In our study, ablation of the UCH-L3 deubiquitinating activity did not inhibit EMT and it did not reduce cancer cell migration and invasion in PC3."

sparser
"These results indicate that UCH-L3 inhibits EMT by repression of EMT promoting genes such as Snail , Slug , Twist , and MMPs in normal prostate cell lines."

eidos
"By contrast , UCHL3 is reduced in metastatic prostate cancer cell lines , and knockdown of UCHL3 promotes epithelial-to-mesenchymal transition ( EMT ) , contributing to cancer cell invasion and metastasis ( 102 ) ."

reach
"We found that knockdown of UCH-L3 induced EMT process."

reach
"Similarly, knockdown of UCHL3 promoted EMT in normal prostate cell lines and led to increased cell migration and invasion, whereas UCHL3 overexpression in prostate cancer cell lines reversed such processes [118]."

reach
"Contrarily, overexpression of UCH-L3 in the highly metastatic prostate cell line PC3 reverses EMT depending on the deubiquitinating activity of UCH-L3."

reach
"As shown in Fig. 2 B, knockdown of UCH-L3 induces the molecular alterations of EMT."

reach
"UCH-L3 was found to be downregulated in the highly metastatic prostate cell lines, and knockdown of UCH-L3 induced EMT process."

reach
"Knockdown of UCH-L3 in the normal prostate cell line RWPE1 induces morphological and molecular events of EMT."

reach
"We have already shown that UCH-L1, a close relative of UCH-L3, induces EMT and promotes metastasis in prostate cancer cells [11]."
UCHL3 affects FOXM1
3 1 | 5 4
UCHL3 binds FOXM1.
3 | 1 4
3 | 1 4

sparser
"Co-immunoprecipitation (co-IP) and in vitro deubiquitination assays were performed to investigate the interactions between UCHL3 and FOXM1."

sparser
"High UCHL3 expression was positively associated with FOXM1 expression level in pancreatic cancer patient samples."

No evidence text available

sparser
"For instance, the progression of pancreatic cancer is promoted by UCHL3-FOXM1 axis [ xref ]."

reach
"Co-immunoprecipitation (co-IP) and in vitro deubiquitination assays were performed to investigate the interactions between UCHL3 and FOXM1."

sparser
"UCH-L3-FOXM1 axis could be targeted for the treatment of pancreatic cancer and re-sensitizing gemcitabine resistant cells."

No evidence text available

No evidence text available
UCHL3 deubiquitinates FOXM1.
1 | 2
UCHL3 deubiquitinates FOXM1. 3 / 3
1 | 2

reach
"Furthermore, knockdown of UCHL3 increased FOXM1 ubiquitination, which enhanced FOXM1 turnover and promoted pancreatic cancer cells ' sensitivity to gemcitabine."

"Our results demonstrate that <span class="match term0">UCHL3</span> deubiquitinated and stabilized <span class="match term1">FOXM1</span>, thereby potentiating proliferation, migration, and invasion of pancreatic cancer cells."

reach
"Deubiquitination of FOXM1 by UCHL3 was also shown to promote pancreatic cancer progression and gemcitabine resistance."
UCHL3 ubiquitinates FOXM1.
| 2
UCHL3 leads to the ubiquitination of FOXM1. 2 / 2
| 2

reach
"Ubiquitin C-terminal hydrolase L3 (UCHL3) as a deubiquitinating enzyme is able to inhibit FOXM1 ubiquitination and degradation 270, while Human leukocyte antigen F-associated transcript 10 (FAT10) as a ubiquitin like protein is able to inhibit FOXM1 ubiquitination and stabilize FOXM1 expression by competing with ubiquitin for binding to FOXM1 271."

reach
"The down-regulation of UCHL3 and FAT10 increased FOXM1 ubiquitination, and also promoted the sensitivity of PC cells to gemcitabine 270, 271."
UCHL3 affects VIM
2 | 8 2
UCHL3 binds VIM.
2 | 1 2
2 | 1 2

sparser
"Co-immunoprecipitation experiments in HCC cells confirmed the interaction of UCHL3 with Vimentin ( xref )."

No evidence text available

reach
"co-immunoprecipitation, Immunofluorescence, and IHC were used to detect the interaction and expression association between UCHL3 and Vimentin in the cells."

sparser
"UCHL3-binding protein complex contained peptide fragments of Vimentin, indicating a VimentinUCHL3 interaction ( xref )."

No evidence text available
UCHL3 activates VIM.
| 3
UCHL3 activates VIM. 3 / 3
| 3

reach
"In conclusion, the role of UCHL3 as a DUB allowed stabilization of Vimentin."

reach
"UCHL3 overexpression enhanced the stability of Vimentin protein, while UCHL3 inhibition suppressed its stability ( Figure 4F )."

reach
"In conclusion, the findings suggest that UCHL3 promotes HCC cell migration through stabilizing Vimentin."
UCHL3 deubiquitinates VIM.
| 2
UCHL3 deubiquitinates VIM. 2 / 2
| 2

reach
"UCHL3 promoted migration of the liver cancer cells of the current study by deubiquitinating Vimentin."

reach
"Vimentin was identified as a potential de-ubiquitination substrate of UCHL3 and UCHL3 interacted with and promoted the de-ubiquitination of Vimentin, enhancing its stability."
UCHL3 decreases the amount of VIM.
| 2
UCHL3 decreases the amount of VIM. 2 / 2
| 2

reach
"UCHL3 promoted Vimentin protein expression, and UCHL3 knockdown inhibited Vimentin protein expression ( Figure 4E )."

reach
"Consistent with the role of UCHL3 as a DUB, UCHL3 overexpression decreased the ubiquitination level of endogenous Vimentin ( Figure 4G ), and Vimentin ubiquitination was increased when UCHL3 was knocked down ( Figure 4H )."
NEDD8 affects UCHL3
5 1 | 3 2
5 1 | 3 2

No evidence text available

reach
"In vitro GST pull-down assay demonstrated that UCH-L3 bound to both NEDD8 and ubiquitin."

sparser
"18 These results suggest that Nedd8 may interact closely with UCH-L3 during testicular atrophy and germ cell loss."

reach
"While UCHL1 and UCHL3 both bind to NEDD8, the hydrolytic processing of NEDD8 is carried out by UCHL3 (Wada et al., 1998)."

reach
"UCH-L3, one of the ubiquitin C-terminal hydrolases, also binds to NEDD8 and functions as a NEDD8 C-terminal hydrolase (Wada et al., 1998)."

No evidence text available

sparser
"These observations suggest that UCH-L3 may function as a deneddylating enzyme 16 in vivo , although further studies are necessary to clarify whether UCH-L3 interacts with Nedd8 during spermatogenesis."

No evidence text available

No evidence text available

No evidence text available
| 11
UCHL3 activates cell migration.
| 9

reach
"In conclusion, the findings suggest that UCHL3 promotes HCC cell migration through stabilizing Vimentin."

reach
"UCHL3 promoted HCC cell migration and stem-like properties.Cell migration and stem-like properties contribute to the aggressiveness of tumors and poor patient prognosis (43)."

reach
"UCHL3 promotes hepatocellular carcinoma cell migration by de-ubiquitinating and stabilizing Vimentin."

reach
"Our results show that UCH-L3 overexpression induced the expression of stemness genes (OCT4, SOX2, ZEB2, and MMP2), and increased cell migration (Figs."

reach
"Vimentin attenuated the suppression of cell migration caused by knockdown of UCHL3."

reach
"UCHL3 overexpression promoted cell migration during wound healing ( Figure 2C ) and Transwell assays ( Figure 2D )."

reach
"UCHL3 knockdown inhibited cell migration and stem-like properties, as revealed by wound healing and Transwell assays ( Figures 3C, D ) and spheroid formation assays ( Figure 3E )."

reach
"In addition, UCHL3 overexpression promoted EJ cell migration, but no alteration in EJ cell migration was observed when the deubiquitination-catalyzing site in UCHL3 was mutated (Fig. 1E)."

reach
"Therefore, the biological function of UCHL5 requires more investigation.One recent study reported that UCHL3 promotes GC cell migration and invasion by upregulating IGF2 [85]."
UCHL3 inhibits cell migration.
| 2

reach
"In ovarian cancer, UCHL3 deubiquitinates TRAF2 to promote tumor progression [ 10 ], and in prostate cancer, overexpression of UCHL3 inhibits the progression of EMT, thereby significantly promoting cel[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In highly metastatic prostate PC3 cells, UCH-L3 inhibits the cell migration and invasion but UCH-L3 C95S had no effects, suggesting that the hydrolase activity of UCH-L3 is required to inhibit the met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 7

reach
"Our findings showed that knockdown of UCHL3 inhibited cell growth, migration and invasion in cervical cancer cells while UCHL3 knockdown inhibited cervical cancer development and metastasis in vivo in mouse models."

reach
"Moreover, a soft agar colony formation assay showed a similar result: stable UCHL3 overexpression, but not UCHL3 expression, notably increased the EJ cell invasion rate (Fig. 1C)."

reach
"UCHL3 induced the growth, invasion, and malignancy of CRC cells."

reach
"Finally, UCHL3 deubiquitinate and stabilize the proliferation-associated transcription factor FOXM1 and promote the invasiveness of pancreatic cancer cells [169]."

reach
"Therefore, UCHL3 increased EJ cell proliferation, viability, migration and invasion, and these effects depended on its deubiquitinating activity."

reach
"Therefore, the biological function of UCHL5 requires more investigation.One recent study reported that UCHL3 promotes GC cell migration and invasion by upregulating IGF2 [85]."

reach
"UCHL3 and UCHL5 slightly enhanced B and T cell infiltration, which may suggest the poor prognosis in RCC."

reach
"In ovarian cancer, UCHL3 deubiquitinates TRAF2 to promote tumor progression [ 10 ], and in prostate cancer, overexpression of UCHL3 inhibits the progression of EMT, thereby significantly promoting cel[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"A previous study reported that UCHL3 in Macrobrachium nipponense is involved in innate immunity to resist bacterial invasion by activating the NF-κB signaling cascade (Zhu et al., 2022; Cheng et al., 2021)."

reach
"In highly metastatic prostate PC3 cells, UCH-L3 inhibits the cell migration and invasion but UCH-L3 C95S had no effects, suggesting that the hydrolase activity of UCH-L3 is required to inhibit the met[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The invasion through the matrigel was increased by UCH-L3 knockdown."
POLD4 affects UCHL3
| 5 6
POLD4 binds UCHL3.
| 4 6
| 4 6

sparser
"Tumorsphere formation and in vitro limiting dilution assay were performed to investigate the effect of UCHL3-POLD4 on GSC self-renewal."

sparser
"Apoptosis, TUNEL, cell cycle phase distribution, modification of the Single Cell Gel Electrophoresis (Comet), γ-H2AX immunofluorescence, and colony formation assays were conducted to evaluate the influence of UCHL3-POLD4 on GSC in ionizing radiation."

sparser
"Both in vitro and in vivo assays indicated that targeted depletion of the UCHL3-POLD4 axis reduced GSC self-renewal and tumorigenic capacity and resistance to IR treatment by impairing homologous recombination (HR) and nonhomologous end joining (NHEJ)."

sparser
"These findings reveal a new signaling axis for GSC PMT regulation and highlight UCHL3-POLD4 as a potential therapeutic target in GBM."

reach
"UCHL3 interacted with, depolyubiquitinated, and stabilized POLD4."

reach
"UCHL3 interacted with, stabilized and depolyubiquitylated POLD4."

reach
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig. 5D, E)."

sparser
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig.  xref D, E)."

sparser
"It has been established that the deubiquitinase UCHL3 binds and stabilizes p12 in GSCs."

reach
"It has been established that the deubiquitinase UCHL3 binds and stabilizes p12 in GSCs."
POLD4 inhibits UCHL3.
| 1
POLD4 inhibits UCHL3. 1 / 1
| 1

reach
"Tumor sphere formation assays and limiting dilution assays showed that POLD4 overexpression reversed the effects of UCHL3 on the sphere-forming ability and self-renewal capacity of MES GSCs (Fig. 6A, B)."
UCHL3 affects farrerol
| 10
UCHL3 binds farrerol.
| 9

reach
"The results revealed a strong interaction between farrerol and UCHL3, and the dissociation constant (Kd) of farrerol/UCHL3 reached a low of 36.73 nM (Fig. 1e and Supplementary Fig. 1d)."

reach
"Farrerol binds to UCHL3 through two amino acid residues, Lys187 and Arg215."

reach
"To elucidate how farrerol specifically binds to UCHL3, we crystallized UCHL3 in the presence of farrerol and determined the structure of the complex at 1."

reach
"The alignment indicated that farrerol did not affect the dynamic structure of UCHL3.Then, we attempted to confirm whether the two amino acid residues were essential for the interaction between UCHL3 and farrerol through the use of CETSAs."

reach
"In contrast to the UCHL3 WT, an improvement in thermal stability was not observed for the two mutants (Fig. 2d–g), indicating that farrerol directly binds to UCHL3 through the two amino acid residues."

reach
"Further SPR assay with the purified recombinant UCHL3–K187A R215A mutant protein and farrerol confirmed that mutating the two amino acid sites abolished the binding of farrerol to UCHL3 (Supplementary Fig. 1f)."

reach
"Since both farrerol and UCHL3 participate in regulating HR by improving the recruitment of RAD51 to DNA damage sites and because we demonstrated that farrerol binds to UCHL3 with a high affinity, we wondered whether farrerol stimulated HR in a UCHL3-dependent fashion."

reach
"Taken together, these data indicated that farrerol promoted DNA repair by HR in a UCHL3-dependent manner.Since the cocrystal structure of the UCHL3farrerol complex revealed that farrerol directly binds to UCHL3 (Fig. 2a–c), a ubiquitin–AMC assay was then performed to determine whether farrerol activates UCHL3 enzymatic activity, and we found that farrerol affected both K and K , by slightly increasing K from 47.72 to 56.27 nM and significantly increasing K from 11.99 to 19.64 s (Fig. 3i), suggesting that farrerol activates UCHL3 deubiquitinase activity."

reach
"Since we demonstrated that farrerol binds to UCHL3 through two amino acid residues to activate its enzymatic activity, in order to treat HR-overactivated cancer or overcome radio/chemo-resistance, it is likely worthwhile to try generating farrerol derivatives that exhibit an inhibitory effect on UCHL3 while still retaining a high affinity to UCHL3.It was reported that genomic stability is critical in the dynamic process of somatic cells reprogramming into iPSCs and is essential for the pluripotency of ESCs and iPSCs, whereas chromosomal aneuploidy results in a sharp decrease in cell differentiation potentials in vivo ."
UCHL3 activates farrerol.
| 1
| 1

reach
"Strikingly, the depletion of Uchl3 completely diminished the positive effect of farrerol on producing cloned mice, as no 173 UCHL3-deficient farrerol-treated two-cell SCNT embryos could normally develop into a term E19.5 stage (Supplementary Table 3)."
UCHL3 affects UCHL1
| 1 9
UCHL3 binds UCHL1.
| 9
| 6

sparser
"To identify the proteins that interact with UCH-L1 or UCH-L3, which might represent the potential substrates or the functional effecters, we performed immunoaffinity chromatography."

sparser
"In addition, the increased levels of both UCH-L1 and UCH-L3 mRNA are associated with early tumor recurrence of invasive breast cancer and poor prognosis [55] ."

sparser
"Nedd8, an Ub-like protein with 57% identity to Ub, is a previously reported UCH-L3 and UCH-L1 interacting protein ( Wada et al., 1998; Hemelaar et al., 2004 )."

sparser
"Thus, the interactions of di-Ub with UCH-L3 and mono-Ub with UCH-L1 occupy the catalytic centers of UCH-L3, and UCH-L1, respectively."

sparser
"This resulted in UCHL3:UCHL1 selectivity for UbV Q40V/T66K/V70F -PRG of approximately 20,000-fold, almost doubling the selectivity ratio described above for UbV T66K/V70F -PRG (10,700-fold)."

sparser
"First, the authors identified multiple DUBs besides UCHL1 and UCHL3 that were associated with SARS-CoV-2 spike particle uptake in their siRNA screen."
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
UCHL3 activates UCHL1.
| 1
UCHL3 activates UCHL1. 1 / 1
| 1

reach
"UCHL1 and UCHL3 are 73% similar but display different patterns of tissue specific gene expression : an upstream neuron-restrictive silencing element (NRSE) drives neuron specific expression in the former and renders it a critical player in neuronal homeostasis (UCHL1 deficiency in neurodegeneration is not physiologically rescued by UCHL3)."
UCHL3 affects NFE2L2
2 | 4 4
UCHL3 binds NFE2L2.
2 | 4
2 | 4

sparser
"In the current study, co-IP assay suggested that UCHL3 directly interacted with NRF2 and stabilized NRF2 protein via deubiquitination."

sparser
"Moreover, rescue experiments were performed and the results showed overexpression of NRF2 antagonized the effect of UCHL3 knockdown, further confirmed the interaction between NRF2 and UCHL3."

No evidence text available

No evidence text available

sparser
"We found that UCHL3 directly interacted with NRF2 and stabilized NRF2 via deubiquitination."

sparser
"To investigate the relationship between UCHL3 and NRF2, co-immunoprecipitation assay was done in C3AA and SiHa cells and the results confirmed that UCHL3 directly interacted with NRF2 ( Fig. 3 A and B[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 activates NFE2L2.
| 3
UCHL3 activates NFE2L2. 3 / 3
| 3

reach
"UCHL3 promotes cervical cancer development and metastasis by stabilizing NRF2 via deubiquitination."

reach
"In cervical cancer studies, it has been found that UCHL3 promotes the development and metastasis of cervical cancer by stabilizing NRF2 through deubiquitination [120]."

reach
"Collectively, our findings suggest that UCHL3 promotes cervical cancer development and metastasis by stabilizing NRF2 via deubiquitination."
UCHL3 increases the amount of NFE2L2.
| 1
UCHL3 increases the amount of NFE2L2. 1 / 1
| 1

reach
"Knockdown of UCHL3 decreased NRF2 expression while overexpression of UCHL3 stabilized NRF2 via deubiquitination."
UCHL3 affects Flag
| 10
| 10

sparser
"Consistently, we co-overexpressed Flag-UCHL3, Myc-NRF2, and HA-ubiquitin (HA-Ub) in HEK293 and the ubiquitination assay also demonstrated that UCHL3 could stabilize NRF2 protein via deubiquitination ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

sparser
"Immunoblotting of cell lysates showed that UCHL3 coimmunoprecipitated with CTNNB1 when Flag-UCHL3 and HA-CTNNB1 were coexpressed in 293 T cells (Fig.  xref A)."

sparser
"Uch-L3 protein (Flag-Uch-L3) was detected in the eluent of the immuno-complex by western blot using anti-Flag antibody ( Fig. 1 A)."

sparser
"1 Experimental Models: Cell Lines HEK293T Laboratory of Michiel Vermeulen N/A HeLa Cells Spruijt et al., 2013 N/A IB10 Cells Spruijt et al., 2013 N/A NPCs Spruijt et al., 2013 N/A HEK293T-EGFP-TAB-NZF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Reciprocally, the co-IP against Flag-Uch-L3 demonstrated that the eluent contained HA-Smad1 ( Fig. 1 B)."

sparser
"As shown in Fig. 1 D, the endogenous Smad1 was also detected in the immunoprecipitated complex of Uch-L3 (Flag-Uch-L3)."

sparser
"We first cotransfected different amounts of Flag-UCHL3 (0, 200, 400, 600 and 800 ng) into 293 T cells with HA-CTNNB1."

sparser
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig.  xref D, E)."

sparser
"For Co-IP and ubiquitination assay in HEK293T cells, HEK293T cells were co-transfected with Myc-NRF2, HA-Ub, and Flag-UCHL3."
UCHL3 affects EP300
| 7 3
UCHL3 binds EP300.
| 3 3
| 3 3

sparser
"The interaction between UCHL3 and p300 was significantly increased by TGF treatment (Fig. xref )."

reach
"However, co-IP analysis showed that p300 could bind to UCHL3 but not UCHL5 or USP14, indicating that UCHL3 specifically binds to and stabilizes p300 in the context of TGF-β signaling (Fig. 3b)."

reach
"Mapping analysis showed that UCHL3 directly interacted with the bromodomain of p300 (Supplementary Fig. 12b–d)."

reach
"The interaction between UCHL3 and p300 was significantly increased by TGF treatment (Fig. 3c)."

sparser
"However, co-IP analysis showed that p300 could bind to UCHL3 but not UCHL5 or USP14, indicating that UCHL3 specifically binds to and stabilizes p300 in the context of TGF-β 1 signaling (Fig. xref )."

sparser
"Mapping analysis showed that UCHL3 directly interacted with the bromodomain of p300 (Supplementary Fig. xref )."
UCHL3 activates EP300.
| 2
UCHL3 activates EP300. 2 / 2
| 2

reach
"The deubiquitinase UCHL3 mediates p300-dependent chemokine signaling in alveolar type II cells to promote pulmonary fibrosis."

reach
"As expected, overexpression of wild-type UCHL3 increased the half-life of p300 compared with that of the inactive mutant UCHL3 (Fig. 3h and Supplementary Fig. 13c)."
UCHL3 ubiquitinates EP300.
| 1
UCHL3 leads to the ubiquitination of EP300. 1 / 1
| 1

reach
"Importantly, wild-type UCHL3 but not inactive mutant UCHL3 efficiently reduced p300 ubiquitination (Fig. 3e)."
UCHL3 increases the amount of EP300.
| 1
UCHL3 increases the amount of EP300. 1 / 1
| 1

reach
"TCID, a selective inhibitor of ubiquitin carboxyl-terminal esterase L3 (UCHL3), robustly inhibited the increase in p300 protein levels induced by TGF-β treatment (Fig. 3a and Supplementary Fig. 12a)."
Flag affects UCHL3
| 10
| 10

sparser
"Consistently, we co-overexpressed Flag-UCHL3, Myc-NRF2, and HA-ubiquitin (HA-Ub) in HEK293 and the ubiquitination assay also demonstrated that UCHL3 could stabilize NRF2 protein via deubiquitination ([MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

sparser
"Immunoblotting of cell lysates showed that UCHL3 coimmunoprecipitated with CTNNB1 when Flag-UCHL3 and HA-CTNNB1 were coexpressed in 293 T cells (Fig.  xref A)."

sparser
"Uch-L3 protein (Flag-Uch-L3) was detected in the eluent of the immuno-complex by western blot using anti-Flag antibody ( Fig. 1 A)."

sparser
"1 Experimental Models: Cell Lines HEK293T Laboratory of Michiel Vermeulen N/A HeLa Cells Spruijt et al., 2013 N/A IB10 Cells Spruijt et al., 2013 N/A NPCs Spruijt et al., 2013 N/A HEK293T-EGFP-TAB-NZF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Reciprocally, the co-IP against Flag-Uch-L3 demonstrated that the eluent contained HA-Smad1 ( Fig. 1 B)."

sparser
"As shown in Fig. 1 D, the endogenous Smad1 was also detected in the immunoprecipitated complex of Uch-L3 (Flag-Uch-L3)."

sparser
"We first cotransfected different amounts of Flag-UCHL3 (0, 200, 400, 600 and 800 ng) into 293 T cells with HA-CTNNB1."

sparser
"Then, we confirmed these interactions by co-IP assay and GST pull-down assay and found that inactive UCHL3 (Flag-UCHL3 S75A) can bind POLD4 directly, indicating that the enzyme activity of UCHL3 was not crucial for their interaction (Fig.  xref D, E)."

sparser
"For Co-IP and ubiquitination assay in HEK293T cells, HEK293T cells were co-transfected with Myc-NRF2, HA-Ub, and Flag-UCHL3."
UCHL3 affects HYCC1
| 9
UCHL3 activates HYCC1. 9 / 9
| 9

reach
"The observation of the growth volume and weight of tumors in nude mice indicates that UCHL3 significantly promoted the tumor growth of HCC cells (Fig. 1I-J)."

reach
"Knockdown of Vimentin inhibited HCC cell migration and suppressed UCHL3-induced HCC cell migration, as illustrated by wound healing and Transwell assays ( Figures 6A–C )."

reach
"Firstly, siRNA was used to knockdown the protein expression of EEF1A1 in UCHL3 overexpressing HCC cells, and the knockdown level was detected by western blot (Fig. 5A)."

reach
"Transwell assay suggested that knockdown of EEF1A1 significantly attenuated the HCC cells migration enhancement induced by UCHL3 overexpression (Fig. 5B)."

reach
"In conclusion, the findings suggest that UCHL3 promotes HCC cell migration through stabilizing Vimentin."

reach
"Together, the abovementioned results show that the DUB activity of UCHL3 stabilizes the expression of Vimentin and promotes the migration of HCC."

reach
"UCHL3 promoted HCC cell migration and stem-like properties.Cell migration and stem-like properties contribute to the aggressiveness of tumors and poor patient prognosis (43)."

reach
"Functional rescue experiments indicated the dependence of UCHL3-mediated HCC cell migration on Vimentin regulation.Previous reports have indicated the deubiquitinating and protein-stabilizing effect of UCHL3 in several human cancer types (19, 20, 22) and demonstrated its contribution to carcinogenesis (14)."

reach
"The DUB activity of UCHL3 stabilized Vimentin and promoted HCC malignant progression."
UCHL1 affects UCHL3
| 9
| 6

sparser
"To identify the proteins that interact with UCH-L1 or UCH-L3, which might represent the potential substrates or the functional effecters, we performed immunoaffinity chromatography."

sparser
"In addition, the increased levels of both UCH-L1 and UCH-L3 mRNA are associated with early tumor recurrence of invasive breast cancer and poor prognosis [55] ."

sparser
"Nedd8, an Ub-like protein with 57% identity to Ub, is a previously reported UCH-L3 and UCH-L1 interacting protein ( Wada et al., 1998; Hemelaar et al., 2004 )."

sparser
"Thus, the interactions of di-Ub with UCH-L3 and mono-Ub with UCH-L1 occupy the catalytic centers of UCH-L3, and UCH-L1, respectively."

sparser
"This resulted in UCHL3:UCHL1 selectivity for UbV Q40V/T66K/V70F -PRG of approximately 20,000-fold, almost doubling the selectivity ratio described above for UbV T66K/V70F -PRG (10,700-fold)."

sparser
"First, the authors identified multiple DUBs besides UCHL1 and UCHL3 that were associated with SARS-CoV-2 spike particle uptake in their siRNA screen."
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
FOXM1 affects UCHL3
3 | 2 4
FOXM1 binds UCHL3.
3 | 1 4
3 | 1 4

sparser
"Co-immunoprecipitation (co-IP) and in vitro deubiquitination assays were performed to investigate the interactions between UCHL3 and FOXM1."

sparser
"High UCHL3 expression was positively associated with FOXM1 expression level in pancreatic cancer patient samples."

No evidence text available

sparser
"For instance, the progression of pancreatic cancer is promoted by UCHL3-FOXM1 axis [ xref ]."

reach
"Co-immunoprecipitation (co-IP) and in vitro deubiquitination assays were performed to investigate the interactions between UCHL3 and FOXM1."

sparser
"UCH-L3-FOXM1 axis could be targeted for the treatment of pancreatic cancer and re-sensitizing gemcitabine resistant cells."

No evidence text available

No evidence text available
FOXM1 activates UCHL3.
| 1
FOXM1 activates UCHL3. 1 / 1
| 1

reach
"UCHL3 promoted pancreatic cancer cell proliferation and migration by deubiquitinating FOXM1 (20)."
8 |
Valproic acid increases the amount of UCHL3. 8 / 8
8 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
XRCC5 affects UCHL3
3 | 2 3
3 | 2 3

sparser
"UCHL3 interacts with Ku80 and is recruited to DNA damage sites in a manner dependent on its catalytic activity."

No evidence text available

No evidence text available

sparser
"Given the findings that UCHL3 interacted with Ku80 and was recruited to sites of damage in a catalytic activity dependent manner (Fig.  xref , Supplementary Fig.  xref ), we investigated whether catalytic activity is required for its phosphorylation."

No evidence text available

reach
"Accordingly, the interaction of Ku80 and UCHL3, which requires UCHL3 catalytic activity (Fig. 1E), was not markedly affected when FLAG-tagged phospho-mutants of UCHL3 were examined by immunoprecipitation (Supplementary Fig. S7B)."

reach
"Here we found that UCHL3, a deubiquitylating enzyme, interacts with and can directly deubiquitylate Ku80 in a catalytic activity dependent manner."

sparser
"Accordingly, the interaction of Ku80 and UCHL3, which requires UCHL3 catalytic activity (Fig.  xref ), was not markedly affected when FLAG-tagged phospho-mutants of UCHL3 were examined by immunoprecipitation (Supplementary Fig.  xref )."
UCHL3 affects TRAF2
| 8
UCHL3 deubiquitinates TRAF2.
| 5
UCHL3 deubiquitinates TRAF2. 5 / 5
| 5

reach
"The elevated UCHL3 facilitates carcinogenesis and enhances inflammation by deubiquitinating and stabilizing TNF Receptor Associated Factor 2 (TRAF2)."

reach
"In ovarian cancer, UCHL3 activates NF-κB signaling by deubiquitinating and stabilizing TRAF2 [9]."

reach
"In ovarian cancer, UCHL3 deubiquitinates TRAF2 to promote tumor progression [ 10 ], and in prostate cancer, overexpression of UCHL3 inhibits the progression of EMT, thereby significantly promoting cel[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"In addition, UCHL3 induced tumorigenesis in ovarian cancer by deubiquitinating and stabilizing TRAF2 (19)."

reach
"UCHL3 was found to activate NF-kappaB signalling by deubiquitinating and stabilizing TRAF2, leading to tumourigenesis."
UCHL3 activates TRAF2.
| 3
UCHL3 activates TRAF2. 3 / 3
| 3

reach
"For example, UCHL3 contributes to non-small cell lung cancer cell growth by stabilizing TRAF2 (TNF receptor-associated factor 2) [22]."

reach
"Zhang et al. found that UCHL3 promotes ovarian cancer progression by stabilizing TRAF2 to activate the NF-κB pathway [51]."

reach
"UCHL3 promotes ovarian cancer progression by stabilizing TRAF2 to activate the NF-kappaB pathway."
LINC00665 affects UCHL3
| 8
LINC00665 increases the amount of UCHL3.
| 5
LINC00665 increases the amount of UCHL3. 5 / 5
| 5

reach
"LINC00665 sponges miR-582-5p to up-regulate UCHL3 expression."

reach
"Meanwhile, Western blot analysis unraveled that LINC00665 overexpression alone led to up-regulated protein levels of UCHL3, AhR and PD-L1 in tumor tissues from irradiation-treated tumor-bearing mice, and its combination with AhR silencing led to relatively decreased protein levels of AhR and PD-L1 and almost unchanged UCHL3 expression (Fig. 7D)."

reach
"Mechanistic investigations further manifested that LINC00665 functioned as a sponge of miR-582-5p and thus up-regulated UCHL3 levels."

reach
"Taken together, experimental data from the present study delineated that LINC00665 acted as a miR-582-5p sponge to up-regulate UCHL3 levels, and enhanced the stability of AhR protein, thereby reducing the radiosensitivity and promoting the immune escape in NSCLC cells."

reach
"LINC00665 functioned as a sponge of miR-582-5p and thus up-regulated the expression of the miR-582-5p target UCHL3."
LINC00665 activates UCHL3.
| 3
LINC00665 activates UCHL3. 3 / 3
| 3

reach
"Collectively, the aforementioned results revealed that LINC00665 functioned as a sponge of miR-582-5p and thus up-regulated UCHL3."

reach
"To explore the up-restream mechanism of the potential UCHL3/AhR/PD-L1 axis, we conducted bioinformatics analysis and predicted that LINC00665 may sponge microRNA (miR)-582-5p to mediate UCHL3."

reach
"Overexpression of LINC00665 and silencing of miR-582-5p enhanced the resistance of NSCLC cells to radiotherapy by upregulating UCHL3 and PD-L1 and stabilizing AhR to promote immune escape (113)."
EP300 affects UCHL3
| 3 5
EP300 binds UCHL3.
| 3 3
| 3 3

sparser
"The interaction between UCHL3 and p300 was significantly increased by TGF treatment (Fig. xref )."

reach
"However, co-IP analysis showed that p300 could bind to UCHL3 but not UCHL5 or USP14, indicating that UCHL3 specifically binds to and stabilizes p300 in the context of TGF-β signaling (Fig. 3b)."

reach
"Mapping analysis showed that UCHL3 directly interacted with the bromodomain of p300 (Supplementary Fig. 12b–d)."

reach
"The interaction between UCHL3 and p300 was significantly increased by TGF treatment (Fig. 3c)."

sparser
"However, co-IP analysis showed that p300 could bind to UCHL3 but not UCHL5 or USP14, indicating that UCHL3 specifically binds to and stabilizes p300 in the context of TGF-β 1 signaling (Fig. xref )."

sparser
"Mapping analysis showed that UCHL3 directly interacted with the bromodomain of p300 (Supplementary Fig. xref )."
EP300 ubiquitinates UCHL3.
| 2
EP300 ubiquitinates UCHL3. 2 / 2
| 2

sparser
"Further studies are required to identify the E3 ligase that mediates the ubiquitination of p300 and examine the detailed mechanisms underlying the removal of p300 ubiquitination of UCHL3."

sparser
"Importantly, wild-type UCHL3 but not inactive mutant UCHL3 C95A efficiently reduced p300 ubiquitination (Fig. xref )."
UCHL3 affects IFN-I
| 7
UCHL3 activates IFN-I. 7 / 7
| 7

reach
"Interestingly, overexpression of UCHL3 robustly promoted IFN-I mediated antiviral activity, as shown by a significantly decreased VSV-GFP signaling."

reach
"Taken all together, our data suggest that UCHL3 positively regulates IFN-I mediated signaling pathway and antiviral activity.Deubiquitinases play important roles in controlling signaling pathways rela[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"However, the roles of most of deubiquitinases in regulating IFN-I antiviral signaling are still largely unexplored.Here, we report that UCHL3 positively regulates IFN-I antiviral signaling pathway."

reach
"Our data showed that knockdown of endogenous UCHL3 inhibited IFN-I induced transcription activation of all three ISGs, strongly suggesting that UCHL3 positively regulates IFN-I signaling pathway.Given[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"If doing so, one could understand that UCHL3 promotes IFN-I signaling by deneddylation of Cullin1 and inhibition of SCF beta and TrCP complex activity.To test this hypothesis, cells were transfected w[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, we reveal that UCHL3 increases levels of SCF beta-TrCP substrates and enhances IFN-I mediated signaling and antiviral activity."

reach
"This finding suggests that UCHL3 could target either TYK2 itself or IFN-I receptors (IFNAR1 and IFNAR2), which are the direct proteins upstream of TYK2 as well as the first signaling proteins in IFN-I[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
NFE2L2 affects UCHL3
2 | 1 4
NFE2L2 binds UCHL3.
2 | 4
2 | 4

sparser
"In the current study, co-IP assay suggested that UCHL3 directly interacted with NRF2 and stabilized NRF2 protein via deubiquitination."

sparser
"Moreover, rescue experiments were performed and the results showed overexpression of NRF2 antagonized the effect of UCHL3 knockdown, further confirmed the interaction between NRF2 and UCHL3."

No evidence text available

No evidence text available

sparser
"We found that UCHL3 directly interacted with NRF2 and stabilized NRF2 via deubiquitination."

sparser
"To investigate the relationship between UCHL3 and NRF2, co-immunoprecipitation assay was done in C3AA and SiHa cells and the results confirmed that UCHL3 directly interacted with NRF2 ( Fig. 3 A and B[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
NFE2L2 activates UCHL3.
| 1
NFE2L2 activates UCHL3. 1 / 1
| 1

reach
"Moreover, we revealed that overexpression of NRF2 could antagonize the function of UCHL3 knockdown in cervical cancer cells."
AHR affects UCHL3
1 | 1 5
1 | 1 5

sparser
"Ouyang et al. revealed the direct interaction between AhR and UCH-L3 in which UCH-L3 exclusively deubiquitinates and thus, reduces AhR poly-ubiquitination which in turn, stabilizes AhR."

sparser
"First, UCHL3 and AhR directly interact."

sparser
"UCHL3 interacts with AhR and stabilizes the AhR protein through deubiquitination."

sparser
"Then, we explored the interaction between the UCHL3 and AhR proteins."

No evidence text available

sparser
"Then, Co-IP results confirmed that AhR interacted with UCHL3 (Fig.  xref E)."

reach
"Co-IP assay was performed to evaluate the interaction between UCHL3 and AhR."
VIM affects UCHL3
2 | 2 2
VIM binds UCHL3.
2 | 1 2
2 | 1 2

sparser
"Co-immunoprecipitation experiments in HCC cells confirmed the interaction of UCHL3 with Vimentin ( xref )."

No evidence text available

reach
"co-immunoprecipitation, Immunofluorescence, and IHC were used to detect the interaction and expression association between UCHL3 and Vimentin in the cells."

sparser
"UCHL3-binding protein complex contained peptide fragments of Vimentin, indicating a VimentinUCHL3 interaction ( xref )."

No evidence text available
VIM activates UCHL3.
| 1
VIM activates UCHL3. 1 / 1
| 1

reach
"Vimentin overexpression promoted HCC cell migration and attenuated the inhibition of cell migration due to UCHL3 knockdown ( Figures 6D–F )."

eidos
"Importantly , UCH-L3 deubiquitylates Smad1 and enhances osteoblast differentiation [ 73 ] ."

reach
"These results of increased expression of differentiation markers and down-stream target genes suggest that Uch-L3 may enhance osteoblast differentiation by elevating the stability of Smad1.To determin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These results suggest that the reduction of Uch-L3 decreases the active pSmad1 signal and thus reduces the osteoblast differentiation.This study showed a novel function for Uch-L3 in osteoblast differ[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"These results suggest that Uch-L3 plays a role as a modulator in the BMP2-mediated osteoblastogenesis and thus enhancing the differentiation of osteoblasts.In our previous study, we reported elevated [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, the forced expression of Uch-L3 promoted the osteoblast differentiation, and Uch-L3 knock-down showed reduced differentiation.This study shows a novel finding that osteoblast differentiat[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

eidos
"UCH-L3 activates osteoblast differentiation by deubiquitinating and stabilizing Smad1 ( Kim et al ., 2011 ) ."
| 6
| 6

reach
"MS-BMDM-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1."

reach
"We next explored the mechanism by which MS-BMDM-derived exosomal UCHL3 modulates BMSCs osteogenesis."

reach
"Thus, we hypothesized that MS-BMDM-derived exosomal UCHL3 could mediate BMSC osteogenesis through the regulation of SMAD1."

reach
"Moreover, although we found that MS-BMDM-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1, additional questions, such as binding sites, still require clarification."

reach
"The ubiquitin carboxyl-terminal hydrolase isozyme L3 (UCHL3) in cell-derived exosomes induced by mechanical stimulation promoted osteogenesis in MSCs, thereby facilitating alveolar bone remodeling during orthodontic tooth movement [77]."

reach
"This study suggests that mechanical force-induced macrophage-derived exosomal UCHL3 promotes BMSCs osteogenesis by targeting SMAD1, thereby promoting alveolar bone formation during OTM."

reach
"These results suggest that Uch-L3 can deubiquitinate the poly-ubiquitinated Smad1.We then tested whether Uch-L3, a possible DUB enzyme of Smad1, is able to accelerate the differentiation of osteoblast[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Consequently, it was suggested that UCHL3 stabilizes Smad1 signaling, which may explain the enhanced osteoblast differentiation caused by UCHL3 overexpression."

reach
"UCH-L3 promotes the differentiation of osteoblast from C2C12 cells, while knockdown of Uch-l3 delays osteoblast differentiation [73]."

reach
"Even though the function of UCH-L3 in neurons has not been elucidated to the extent of our knowledge, it may be plausible that there can be reciprocal or differential functions of UCH-L3 to the UCH-L1[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects YAP1
5 | 1
UCHL3 binds YAP1.
5 |
5 |

No evidence text available

No evidence text available

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UCHL3 increases the amount of YAP1.
| 1
UCHL3 increases the amount of YAP1. 1 / 1
| 1

reach
"Depletion of UCHL3 decreased the YAP protein level and the expression of YAP/TEAD target genes in ATC."
| 3

reach
"In addition, since the increased sensitivity to IR caused by UCHL3 depletion was moderate and the human genome encodes around 100 DUBs, it would be interesting to examine if any other DUBs can deubiquitylate Ku."

reach
"To exclude the possibility that the increased IR sensitivity caused by UCHL3 depletion was due to decreased proliferation or cell cycle issues, cell proliferation and cell cycle profile of these cell lines were examined by 3-(4,5-dimethylthiazol-2-yl)-2,5-diphenyltetrazolium bromide (MTT)-based assay and bromodeoxyuridine (BrdU) incorporation assay, respectively (Supplementary Fig. S2A,B)."

reach
"Finally, re-expression of wild-type UCHL3 but not the S75A mutant was able to reverse hypersensitivity to IR induced by UCHL3 deficiency (Fig. 5F)."
| 3

reach
"UCHL3 promotes PMT in GSCs and IR resistance via POLD4.Currently, small-molecule drugs targeted specifically against POLD4 have not been developed clinically."

reach
"UCHL3 promotes GSC progression and IR resistance via POLD4."

reach
"Colony formation assays indicated that POLD4 overexpression restored the proliferation ability reduced by UCHL3 depletion in the case of IR (Fig. 6C)."
UCHL3 affects MTA2
| 2 4
UCHL3 binds MTA2.
| 1 4
| 1 4

sparser
"In vitro binding assays indicated that the C1 box domain (90–100 amino acids), but not other domains, of UCHL3 protein specifically bound to MTA2 ( xref G)."

sparser
"Moreover, the Co-IP assays indicated that UCHL3 and MTA2 could bind to each other in AGS and MKN-45 cells ( xref D)."

sparser
"Moreover, overexpression of circ MTA2 significantly enhanced the interaction between UCHL3 and MTA2 ( xref E) and increased the ubiquitination level of MTA2 ( xref F)."

sparser
"Of note, circ MTA2 could interact with UCHL3, which ultimately reduced the ubiquitination degradation of MTA2 , thus promoting GC progression."

reach
"Moreover, overexpression of circMTA2 significantly enhanced the interaction between UCHL3 and MTA2 (Figure 6E) and increased the ubiquitination level of MTA2 (Figure 6F)."
UCHL3 increases the amount of MTA2.
| 1
UCHL3 increases the amount of MTA2. 1 / 1
| 1

reach
"Knockdown of UCHL3 in AGS and MKN-45 cells could further decrease MTA2 expression under CHX treatment, while UCHL3 overexpression partially mitigated MTA2 decline (Figure 5F)."
UCHL3 affects FlaG
| 6
| 6

sparser
"These data were in line with observed c-NHEJ activities being reduced in UCHL3 KO#2 cells, which was largely rescued by exogenously expressed FLAG-UCHL3 (Fig.  xref )."

sparser
"Western blot analyses revealed that HA-CTNNB1 expression gradually increased with the increased expression of FLAG-UCHL3, and this increase was dose dependent (Fig.  xref D)."

sparser
"Our results demonstrate that there was an interaction between FLAG-UCH-L3 and GFP-β-catenin; however, we did not observe this interaction in cells transfected with the inactive mutant of UCH-L3 (UCH-L3 C95S) ( xref )."

sparser
"To evaluate the effect of UCHL3 DUB activity on CTNNB1 stability, we overexpressed FLAG-UCHL3 or the FLAG-UCHL3 C94A mutant and found that the stability of FLAG-UCHL3 binding to HA-CTNNB1 depended on its ubiquitinase activity (Fig.  xref E)."

sparser
"We observed that FLAG-BAP1 ( xref B, upper panel), but not FLAG-UCHL3 ( xref B, lower panel), directly binds to HA-KEAP1."

sparser
"As shown in Fig.  xref G, the protein half-life of HA-CTNNB1 was obviously extended in the presence of FLAG-UCHL3 compared to that observed when an empty vector was used."
| 6
UCHL3 activates Carcinogenesis.
| 5

reach
"In conclusion, our findings suggest that UCHL3 promotion of RCC tumorigenesis may be achieved through inactivating CD8 T cells."

reach
"Regulating biological functions, UCHL3 promoted GSC proliferation, tumorigenesis, self-renewal, and radioresistance in a POLD4-dependent manner.The degradation of POLD4 by ubiquitin ligases in response to DNA damage and during cell cycle progression plays a less prominent role in normal cells [43], while with increasing insights into the complex mechanisms of DNA damage repair, POLD4 is projected to play an increasingly important role."

reach
"Our data suggested that UCHL3 depletion reduced bladder tumorigenesis in a xenograft model and that this effect was at least partially dependent on CTNNB1 stabilization."

reach
"Uchl3 deficiency leads to inhibited BBN-induced bladder tumorigenesis in mice."

reach
"In summary, it is suggested that UCHL3 may contribute to liver tumorigenesis and may be a useful prognostic biomarker in HCC."
UCHL3 inhibits Carcinogenesis.
| 1

reach
"These results suggested that Uchl3 knockout alleviated BBN-induced bladder tumorigenesis in vivo."
UCHL3 affects CUL1
| 6
UCHL3 inhibits CUL1.
| 4
UCHL3 inhibits CUL1. 4 / 4
| 4

reach
"We find that in cells UCHL3 can decrease neddylation of Cullin1."

reach
"We found that overexpression of increased amounts of UCHL3 gradually inhibited endogenous Cullin1 neddylation, indicating that cellular Cullin1 neddylation are regulated by UCHL3 levels."

reach
"We reveal that UCHL3 can inhibit Cullin1 neddylation in cells dependently on COPS5."

reach
"Our results showed that UCHL3 overexpression inhibits Cullin1 neddylation."
UCHL3 activates CUL1.
| 2
UCHL3 activates CUL1. 2 / 2
| 2

reach
"However, we found that knockout of COPS5 abolished the effect of UCHL3 on Cullin1 deneddylation, indicating that COPS5 is required for UCHL3 mediated deneddylation of Cullin1 in cells."

reach
"Subsequently, UCHL3 mediated upregulation of COPS5 promotes Cullin1 deneddylation."
UCHL3 affects BRD4
2 | 4
UCHL3 binds BRD4.
2 | 2
2 | 2

reach
"Further investigation also showed the endogenous BRD4 binding with UCHL3 which was enhanced by rIL6 but inhibited by pacritinib (Figs."

No evidence text available

reach
"Of note, our experiments were performed in the presence of IL6/8. whether other deubiquitinases are preferential in other contexts need to be further investigated.To investigate if UCHL3 binds to BRD4, we performed reciprocal co-immunoprecipitation assays in cells transfected with BRD4, alone or with UCHL3 and JAK2-V617F."

No evidence text available
UCHL3 ubiquitinates BRD4.
| 1
UCHL3 leads to the ubiquitination of BRD4. 1 / 1
| 1

reach
"In contrast, deubiquitinases DUB3 and UCHL3 were reported to reverse BRD4 ubiquitination in prostate cancer and colorectal cancer, respectively (11, 15)."
UCHL3 activates BRD4.
| 1
UCHL3 activates BRD4. 1 / 1
| 1

reach
"Conversely, ectopic expression of UCHL3 prolonged the half-life of BRD4 (Fig. S4j); It also increased the protein stability of WT BRD4 but not BRD4-Y97/98A mutant (Fig. 3f)."
FlaG affects UCHL3
| 6
| 6

sparser
"These data were in line with observed c-NHEJ activities being reduced in UCHL3 KO#2 cells, which was largely rescued by exogenously expressed FLAG-UCHL3 (Fig.  xref )."

sparser
"Western blot analyses revealed that HA-CTNNB1 expression gradually increased with the increased expression of FLAG-UCHL3, and this increase was dose dependent (Fig.  xref D)."

sparser
"Our results demonstrate that there was an interaction between FLAG-UCH-L3 and GFP-β-catenin; however, we did not observe this interaction in cells transfected with the inactive mutant of UCH-L3 (UCH-L3 C95S) ( xref )."

sparser
"To evaluate the effect of UCHL3 DUB activity on CTNNB1 stability, we overexpressed FLAG-UCHL3 or the FLAG-UCHL3 C94A mutant and found that the stability of FLAG-UCHL3 binding to HA-CTNNB1 depended on its ubiquitinase activity (Fig.  xref E)."

sparser
"We observed that FLAG-BAP1 ( xref B, upper panel), but not FLAG-UCHL3 ( xref B, lower panel), directly binds to HA-KEAP1."

sparser
"As shown in Fig.  xref G, the protein half-life of HA-CTNNB1 was obviously extended in the presence of FLAG-UCHL3 compared to that observed when an empty vector was used."
FOS affects UCHL3
| 5 1
FOS increases the amount of UCHL3.
| 5
FOS increases the amount of UCHL3. 5 / 5
| 5

reach
"Using ChIP-qPCR, we confirmed that this increased UCH-L3 gene expression is caused by direct binding of c-Fos to the UCH-L3 promoter (Fig. 3D)."

reach
"In GCSCs, c-Fos translocated into the nucleus and induced UCH-L3 expression under stress conditions, resulting in the inhibition of β-catenin degradation."

reach
"In conclusion, we demonstrated that the expression of UCH-L3 in GCSCs under stress conditions is induced by c-Fos, resulting in nuclear accumulation of β-catenin, and conferring high resistance to external stimuli by expressing stemness genes."

reach
"UCH-L3 expression was induced by the transcription factor c-Fos, and the levels of c-Fos increased in response to autophagy activation."

reach
"UCH-L3 expression in GCSCs is induced directly by c-Fos."
FOS binds UCHL3.
| 1
| 1

sparser
"Using ChIP-qPCR, we confirmed that this increased UCH-L3 gene expression is caused by direct binding of c-Fos to the UCH-L3 promoter ( xref )."
YAP1 affects UCHL3
5 |
5 |

No evidence text available

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USP5 affects UCHL3
1 | 1 3
1 | 1 1

No evidence text available

sparser
"His-PD showed that OTULIN ABPΔG76 still bound to USP5 and UCHL3 ( xref I), confirming our MS results that these DUBs are associating largely in a non-covalent manner with the probes (compare xref B and 6C)."

reach
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non covalent and reversible."
USP5 binds UCHL3 and SHBG. 2 / 2
| 2

sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."

reach
"Ablating UCHL3 significantly attenuates farrerol-mediated stimulation in HR and SCNT embryo development."

reach
"Furthermore, UCHL3 was recently suggested to facilitate HR by deubiquitylating RAD51 in a DSB-dependent manner via phosphorylation on the same site that we identified (S75) , however, we did not observe any defects in HR caused by UCHL3 depletion, nor did we detect any interaction with RAD51 in our mass spectrometry analyses."

reach
"Our results also suggest that UCHL3 deficiency causes HR defect but does not affect cell proliferation (Fig. 1D; Supplemental Fig. S1A)."

reach
"We found that depleting UCHL3 abolished the stimulation of HR by farrerol (Fig. 3a and Supplementary Fig. 2c)."

reach
"We first confirmed that overexpressing UCHL3 promoted HR using the HR reporter cell line (Supplementary Fig. 2a, b)."
UCHL3 affects cell growth
| 5
UCHL3 activates cell growth.
| 4
| 4

reach
"Moreover, CRY2 silencing abolished UCHL3 deficiency-mediated inhibition in cell growth and migration."

reach
"UCHL3 overexpression enhanced A549 and H1299 cell growth in vitro (Fig. 2a, b)."

reach
"Our findings showed that knockdown of UCHL3 inhibited cell growth, migration and invasion in cervical cancer cells while UCHL3 knockdown inhibited cervical cancer development and metastasis in vivo in mouse models."

reach
"For example, UCHL3 contributes to non-small cell lung cancer cell growth by stabilizing TRAF2 (TNF receptor-associated factor 2) [22]."
UCHL3 inhibits cell growth.
| 1
Mutated UCHL3 inhibits cell growth. 1 / 1
| 1

reach
"Interestingly, stable overexpression of UCHL3 mutants in A549 cells significantly attenuated cell growth compared to that in cells expressing WT UCHL3 (Fig. 6b)."
| 4

reach
"Furthermore, UCHL3 stabilized CTNNB1 and subsequently triggered the oncogenic Wnt signaling pathway (Fig. 8)."

reach
"UCHL3 activates the Wnt signaling pathway."

reach
"UCHL3 was highly associated with the Wnt signaling pathway and triggered the activation of the Wnt signaling pathway, which showed that its functions depend on its deubiquitination activity."

reach
"All the data suggested that UCHL3 triggered the activation of the Wnt signaling pathway."

reach
"Western blot analysis showed that the protein levels of Wnt/β-Catenin downstream gene-encoded products were reduced in UCHL3-deficient tumor tissues, including LEF1, Axin2, C-myc and Cyclin D1, confirming that the abolition of UCHL3 inhibited WNT signaling pathway activation, which is consistent with our aforementioned experiments."
UCHL3 affects USP5
1 | 1 3
1 | 1 1

No evidence text available

sparser
"His-PD showed that OTULIN ABPΔG76 still bound to USP5 and UCHL3 ( xref I), confirming our MS results that these DUBs are associating largely in a non-covalent manner with the probes (compare xref B and 6C)."

reach
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non covalent and reversible."
USP5 binds UCHL3 and SHBG. 2 / 2
| 2

sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."
UCHL3 affects UCHL3
| 5
UCHL3 increases the amount of UCHL3.
| 3
UCHL3 increases the amount of UCHL3. 3 / 3
| 3

reach
"UCHL3 promoter analysis revealed that TEAD4, through which YAP bind to DNA, activated UCHL3 transcription by binding to the promoter of UCHL3."

reach
"RT-qPCR analysis demonstrated no significant differences in UCHL3, NEDD8 and LC3B expression levels between the NC and si-NC groups (P>0.05; Fig. 4A and B); however, UCHL3 knockdown significantly decreased the expression levels of UCHL3 and NEDD8, and significantly increased LC3B expression in the si-UCHL3 group compared with the NC group (all P<0.001; Fig. 4A and B)."

reach
"According to Western blot analysis results, protein levels of UCHL3, AhR and PD-L1 were all reduced in response to sh-UCHL3 treatment, while additional LV-AhR restored the protein levels of AhR and PD-L1 but showed on obvious effects on the UCHL3 protein level; consistently, LV-UCHL3 led to up-regulated protein levels of UCHL3, AhR and PD-L1, among which only the up-regulation of AhR and PD-L1 was reversed in response to additional sh-AhR treatment (Fig. 2M)."
UCHL3 activates UCHL3.
| 2
UCHL3 activates UCHL3. 2 / 2
| 2

reach
"ATM-dependent phosphorylation of the deubiquitinase UCHL3 enables UCHL3 to enhance the retention of Ku80 at damage sites by counteracting Ku80 ubiquitination (Nishi et al., 2018)."

reach
"To further test whether UCHL3 phosphorylation is able to activate UCHL3, we performed a two-step assay."
UCHL3 affects SLC11A1
| 5
UCHL3 binds SLC11A1.
| 2
| 2

reach
"At the same time, the lncRNA GIAT4RA counteracts the deubiquitinating effects of LSH by interfering with interaction between LSH and UCH-L3, reducing levels of LSH protein, and playing the role in inhibiting the invasion and migration of LUAD cells83."

reach
"However, HOTAIR was also reported to reduce the ubiquitination degradation of AR by preventing its combination with MDM2, and GIAT4RA was recently reported to interfere the interaction between Uchl3 and LSH to repress ubiquitin-proteasome pathway [30], which indicate multiple roles of lncRNA in ubiquitination."
UCHL3 activates SLC11A1.
| 2
UCHL3 activates SLC11A1. 2 / 2
| 2

reach
"GIAT4RA interfered with ubiquitin hydrolase Uchl3 mediated interaction and stabilization of LSH."

reach
"Ubiquitin C-Terminal Hydrolase L3 (UCHL3) stimulates LSH stabilization, and lncRNA GIAT4RA interferes with UCHL3-mediated LSH deubiquitination in a UCHL3-dependent manner [5]."
UCHL3 deubiquitinates SLC11A1.
| 1
UCHL3 deubiquitinates SLC11A1. 1 / 1
| 1

reach
"UCHL3 deubiquitinates chromatin modifier lymphoid-specific helicase (LSH)."
UCHL3 affects K27
| 5
K27 binds UCHL3. 5 / 5
| 5

sparser
"Furthermore, we showed that UCHL3 preferentially binds to K27 diUb and that its catalytic site is important for regulating K27 polyUb chains in cells."

sparser
"Further studies on the interaction between UCHL3 and K27 diUb by Pan et al. revealed an unique inhibition mechanism where the crossover loop of UCHL3 binds between the two Ub moieties putting UCHL3 in[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We decided to further investigate the detected selective interaction between UCHL3 and K27 diUb."

sparser
"We confirmed a direct interaction between UCHL3 and both non-hydrolyzable and native K27 diUb ( Figures 5 E and 5F)."

sparser
"Previous studies have analyzed the crystal structure of UCHL3-K27 di-Ub, in which one ubiquitin occupies the S1 site of UCHL3 and the other ubiquitin occupies the S2 site of UCHL3 ( xref A) [ xref ]."
UCHL3 affects CRY2
| 5
UCHL3 leads to the methylation of CRY2. 5 / 5
| 5

reach
"In summary, this study reveals that knockdown of UCHL3 inhibits ESCC growth and migration by reducing CRY2 methylation through downregulation of FOXM1 expression."

reach
"UCHL3 promotes the malignant progression of esophageal carcinoma by influencing CRY2 methylation [26]."

reach
"In the current study, we aimed to investigate the role of UCHL3 in ESCC growth and migration, and whether UCHL3 could modulate CRY2 methylation through FOXM1."

reach
"In addition, UCHL3 knockdown decreased CRY2 methylation through downregulating FOXM1, leading to an increase in the expression of CRY2."

reach
"Knockdown of UCHL3 inhibits esophageal squamous cell carcinoma progression by reducing CRY2 methylation."
SLC4A1 affects UCHL3
| 5
SLC4A1 binds UCHL3.
| 3
| 3

reach
"To confirm the direct interaction of di-Ub and UCH-L3, we performed pull-down assays using recombinant mono-Ub or di-Ub and UCH-L3."

reach
"Likewise, UCH-L3 might interact with di-Ub to protect it from degradation."

reach
"Thus, we can not exclude the possibility that, under some conditions, UCH-L3 may interact with di-Ub to protect it from degradation.Previous biochemical analysis indicated that UCH-L3 is unable to hyd[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
SLC4A1 inhibits UCHL3.
| 1
SLC4A1 inhibits UCHL3. 1 / 1
| 1

reach
"Thus, we speculated that di-Ub might inhibit the enzymatic activity of UCH-L3."
SLC4A1 activates UCHL3.
| 1
SLC4A1 activates UCHL3. 1 / 1
| 1

reach
"To explore the cleavage preference of UCHL3 for different types of ubiquitin chains and whether phosphorylation modification gives UCHL3 the ability to cleave ubiquitin chains in vitro, different linkages (K6, K11, K27, K48, and K63) of di-Ub were synthesized and tested for their activity of UCHL3 and UCHL3 ."
MTA2 affects UCHL3
| 1 4
| 1 4

sparser
"In vitro binding assays indicated that the C1 box domain (90–100 amino acids), but not other domains, of UCHL3 protein specifically bound to MTA2 ( xref G)."

sparser
"Moreover, the Co-IP assays indicated that UCHL3 and MTA2 could bind to each other in AGS and MKN-45 cells ( xref D)."

sparser
"Moreover, overexpression of circ MTA2 significantly enhanced the interaction between UCHL3 and MTA2 ( xref E) and increased the ubiquitination level of MTA2 ( xref F)."

sparser
"Of note, circ MTA2 could interact with UCHL3, which ultimately reduced the ubiquitination degradation of MTA2 , thus promoting GC progression."

reach
"Moreover, overexpression of circMTA2 significantly enhanced the interaction between UCHL3 and MTA2 (Figure 6E) and increased the ubiquitination level of MTA2 (Figure 6F)."
K27 affects UCHL3
| 5
K27 binds UCHL3. 5 / 5
| 5

sparser
"Furthermore, we showed that UCHL3 preferentially binds to K27 diUb and that its catalytic site is important for regulating K27 polyUb chains in cells."

sparser
"Further studies on the interaction between UCHL3 and K27 diUb by Pan et al. revealed an unique inhibition mechanism where the crossover loop of UCHL3 binds between the two Ub moieties putting UCHL3 in[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"We decided to further investigate the detected selective interaction between UCHL3 and K27 diUb."

sparser
"We confirmed a direct interaction between UCHL3 and both non-hydrolyzable and native K27 diUb ( Figures 5 E and 5F)."

sparser
"Previous studies have analyzed the crystal structure of UCHL3-K27 di-Ub, in which one ubiquitin occupies the S1 site of UCHL3 and the other ubiquitin occupies the S2 site of UCHL3 ( xref A) [ xref ]."
GARS1 affects UCHL3
| 5
| 5

reach
"The protein‒protein docking results from Alphafold 2 revealed that UCHL3 could directly bind with SMAD1 through hydrogen bonds (Fig. 7i)."

reach
"Further immunofluorescence and co-immunoprecipitation (Co-IP) verified the interaction between UCHL3 and SMAD1 in BMSCs (Fig. 7j–l)."

reach
"To investigate the interaction between Uch-L3 and Smad1, HA-Smad1 and Flag-Uch-L3 plasmids were transfected into HEK 293 cells."

reach
"All of these results showed that Uch-L3 interacts with Smad1.After confirming the interaction between Smad1 and Uch-L3, we tested whether the deubiquitinating activity of Uch-L3 is able to affect the [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"The physical interaction between Uch-L3 and Smad1 was revealed."

reach
"The EEF1A1/UCHL3 axis promotes HCC cell malignant and drug resistance."

reach
"Functional experiments and nude mouse models have demonstrated that the UCHL3/EEF1A1 axis promotes the migration, stemness, and drug resistance of HCC cells."

reach
"However, the effects of the UCHL3/EEF1A1 axis on HCC tumors as revealed by patient-derived xenograft (PDX) models remains to be explored.Therefore, UCHL3 and EEF1A1 jointly promote the migration, stemness, and drug resistance of HCC cells, as well as influencing the growth of mouse tumor models."

reach
"UCHL3 enhances drug resistance and glycolysis in pancreatic cancer through the stabilization of FOXM1 and upregulation of LDHA [6, 7]."
UCHL3 affects SP
| 4
| 4

sparser
"Prediction of Sp-UCHL3 and Sp-UCHL5 Three-Dimensional (3D) Structure."

sparser
"Therefore, the expression patterns of Sp-UCHL3 and Sp-UCHL5 in different tissues and ovarian development suggest that they play important roles in cellular proteins’ degradation during oogenesis and ovarian maturation."

sparser
"Phylogenetic and Homology Analyses of Sp-UCHL3 and Sp-UCHL5."

sparser
"In the present study, Sp-UCHL3 mRNA expression in T3 was significantly higher than in the T1 and T2 stages ( p < 0.05), while Sp-UCHL5 mRNA expressions in the three stages of testes development were not significantly different from each other ( p > 0.05), indicating that Sp- UCHL3 and Sp- UCHL5 have different aspects in regulating mechanism of testis development."
UCHL3 affects Proteasome
| 4
UCHL3 increases the amount of Proteasome.
| 2
UCHL3-C95S increases the amount of Proteasome. 1 / 1
| 1

reach
"To address whether UCHL3 C95S impedes Ub(G76V)‐GFP turnover at the level of p97 or the proteasome, we employed a variant Ub(G76V)‐GFP protein containing a C‐terminal 20‐amino acid extension (Ub(G76V)‐GFP‐20AA), which has been shown to override the requirement for p97‐mediated unfolding to enable its proteasomal degradation (Beskow et al, 2009; Godderz et al, 2015) (Fig 5E)."
UCHL3 increases the amount of Proteasome. 1 / 1
| 1

reach
"Since mild inhibition of 26S proteasomes by the addition of inhibitors is reported to induce the expression of 26S proteasome subunits ( Meiners et al., 2003 ), partial inhibition of 26S proteasome mi[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 inhibits Proteasome.
| 1
| 1

reach
"Inhibiting the proteasome using MG132 led to marked enrichment of K48 ubiquitylated TDP1 in control cells but no impact in rhabdomyosarcoma cells, due to high UCHL3 levels (XREF_FIG C)."
UCHL3 activates Proteasome.
| 1
| 1

reach
"In our study, we found that the ubiquitin–proteasome system (UPS) was modulated by the regulation of some ubiquitin enzyme and ligases genes (uchl3, trim 25, trim39, ubr7 and ube2i) and proteasome subunits (psma4, psma6, psmb10, psmc2 and rad23a), which act as protein binding signalling and protein degradation, respectively."
UCHL3 affects Ku80 chromatin binding
| 4
UCHL3 activates Ku80 chromatin binding. 4 / 4
| 4

eidos
"To gain insight into how UCHL3 contributes to Ku80 chromatin binding after DSB induction , firstly , we examined the effect of UCHL3 on the DNA-binding activity of Ku by electrophoretic mobility shift assay with purified these proteins ( Fig. 5A-C ) ."

eidos
"Since it was reported that Ku ubiquitylation is a prerequisite for Ku removal from chromatin , we hypothesized that UCHL3 may enhance Ku80 chromatin binding by antagonizing Ku80 ubiquitylation ."

eidos
"UCHL3 enhances Ku80 chromatin binding and directly deubiquitylates Ku80 To examine the effect of UCHL3 on Ku regulation , we investigated IR-induced foci ( IRIF ) formation of Ku80 , which indicates loading of Ku onto DSB sites ."

eidos
"UCHL3 enhances Ku80 chromatin binding upon DSB induction ."
UCHL3 affects KITLG
| 4
UCHL3 activates KITLG. 4 / 4
| 4

reach
"Furthermore, we reveal that UCHL3 increases levels of SCF beta-TrCP substrates and enhances IFN-I mediated signaling and antiviral activity."

reach
"In addition, studies have confirmed that UCHL3 can upregulate the degree of SCF β−TrCP substrates, enhance the IFN-I-mediated signaling pathway and antiviral activity, and participate in immune regula[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Thus, our findings identify a novel in vivo substrate of UCHL3, and reveal a novel deneddylation regulation by UCHL3.It is interesting that UCHL3 controls the activity of cellular SCF beta and TrCP co[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Moreover, we find that UCHL3 upregulates levels of SCF beta-TrCP substrates including IFN-I receptor IFNAR1, thereby enhancing IFN-I mediated signaling and antiviral activity."
UCHL3 affects INS
| 4
UCHL3 activates INS.
| 3
UCHL3 activates INS. 3 / 3
| 3

reach
"Furthermore, UCHL3 has deneddylating activity essential for various cellular functions and promotes insulin signaling and adipogenesis, hence contributing to retinal maintenance in stress conditions [15]."

reach
"These results suggest that UCH-L3 promotes adipogenesis by enhancing insulin signaling in a hydrolase activity dependent manner."

reach
"Suzuki et al. have shown that UCHL3 enhances insulin signalling in (pre) adipocytes, but UCHL3 is unlikely to target critical components of insulin signalling like the insulin receptor, the IGF-I receptor, IRS-1 and Grb10 XREF_BIBR."
UCHL3 phosphorylates INS.
| 1
Modified UCHL3 leads to the phosphorylation of INS. 1 / 1
| 1

reach
"Moreover, ectopic expression of wild-type UCH-L3 restored the phosphorylation of insulin and IGF-I receptor and adipocyte differentiation in Uchl3 (-/-) MEFs."
UCHL3 affects HELLS
3 | 1
UCHL3 binds HELLS.
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL3 deubiquitinates HELLS.
| 1
UCHL3 deubiquitinates HELLS. 1 / 1
| 1

reach
"UCHL3 deubiquitinates chromatin modifier lymphoid-specific helicase (LSH)."
UCHL3 affects GST
| 4
| 4

sparser
"We observed a reduced ubiquitin signal with GST-UCHL3 but not with GST, suggesting that UCHL3 directly removed ubiquitin from Ku80."

sparser
"Purified His-RAD51 was able to interact with GST-UCHL3 under cell-free conditions even with a relatively high salt concentration in the washing buffer ( xref C), suggesting a direct interaction between UCHL3 and RAD51."

sparser
"The cDNA of UCH-L3 was subcloned from the library plasmid with GenBank N27190 by PCR into the Eco RI/ Xho I sites of pGEX-4T1 (Amersham) encoding for a GST-UCH-L3 fusion protein."

sparser
"1 Experimental Models: Cell Lines HEK293T Laboratory of Michiel Vermeulen N/A HeLa Cells Spruijt et al., 2013 N/A IB10 Cells Spruijt et al., 2013 N/A NPCs Spruijt et al., 2013 N/A HEK293T-EGFP-TAB-NZF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects CLPB
4 |
4 |

No evidence text available

No evidence text available

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No evidence text available
UCHL3 affects CD274
| 4
UCHL3 increases the amount of CD274. 4 / 4
| 4

reach
"UCHL3 promotes stabilization of AhR protein through deubiquitination, thereby elevating PD-L1 expression."

reach
"In summary, these results unraveled that UCHL3 augmented the stability of AhR protein through deubiquitination, thereby promoting PD-L1 expression."

reach
"According to Western blot analysis results, protein levels of UCHL3, AhR and PD-L1 were all reduced in response to sh-UCHL3 treatment, while additional LV-AhR restored the protein levels of AhR and PD-L1 but showed on obvious effects on the UCHL3 protein level; consistently, LV-UCHL3 led to up-regulated protein levels of UCHL3, AhR and PD-L1, among which only the up-regulation of AhR and PD-L1 was reversed in response to additional sh-AhR treatment (Fig. 2M)."

reach
"Furthermore, our data unraveled that UCHL3 augmented the stability of AhR protein through deubiquitination, thereby promoting PD-L1 expression."
UCHL3 affects ABCG2
| 4
UCHL3 increases the amount of ABCG2.
| 2
UCHL3 increases the amount of ABCG2. 2 / 2
| 2

reach
"FACS analysis showed that inhibition of UCHL3 using TCID reduced CD338 expression and the percentage of SP cells among A549 cells overexpressing UCHL3 (Fig. 7e, f)."

reach
"As expected, overexpression of UCHL3 increased the expression of ABCG2, c-Myc and KLF4, genes associated with stemness, at the protein level in A549 and H1299 cells (Fig. 5b, c), while depletion of UCHL3 decreased their protein expression (Fig. 5d)."
UCHL3 activates ABCG2.
| 2
UCHL3 activates ABCG2. 2 / 2
| 2

reach
"Ubiquitin C-terminal hydrolase L3 (UCHL3) stabilizes aryl hydrocarbon receptor (AhR) by its deubiquitylation and subsequently increases ABCG2 to promote stem-like properties in lung cancer [71]."

reach
"Flow cytometry analysis indicated that knockdown of UCHL3 reduced the proportion of CD338 H358 cells (Fig. 3d)."
SP affects UCHL3
| 4
| 4

sparser
"Prediction of Sp-UCHL3 and Sp-UCHL5 Three-Dimensional (3D) Structure."

sparser
"Therefore, the expression patterns of Sp-UCHL3 and Sp-UCHL5 in different tissues and ovarian development suggest that they play important roles in cellular proteins’ degradation during oogenesis and ovarian maturation."

sparser
"Phylogenetic and Homology Analyses of Sp-UCHL3 and Sp-UCHL5."

sparser
"In the present study, Sp-UCHL3 mRNA expression in T3 was significantly higher than in the T1 and T2 stages ( p < 0.05), while Sp-UCHL5 mRNA expressions in the three stages of testes development were not significantly different from each other ( p > 0.05), indicating that Sp- UCHL3 and Sp- UCHL5 have different aspects in regulating mechanism of testis development."
RA-9 affects UCHL3
| 4
RA-9 inhibits UCHL3. 4 / 4
| 4

reach
"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"RA-9, a chalcone derivative with a structure similar to b-AP15, was reported to inhibit proteasomal DUBs [XREF_BIBR] as well as UCHL1, UCHL3, USP2, USP5, and USP8 [XREF_BIBR]."

reach
"Indeed, it was found that RA-9 inhibited in vitro UCHL1, UCHL3, USP8 and USP9x, besides USP14 and UCHL5."
GST affects UCHL3
| 4
| 4

sparser
"We observed a reduced ubiquitin signal with GST-UCHL3 but not with GST, suggesting that UCHL3 directly removed ubiquitin from Ku80."

sparser
"Purified His-RAD51 was able to interact with GST-UCHL3 under cell-free conditions even with a relatively high salt concentration in the washing buffer ( xref C), suggesting a direct interaction between UCHL3 and RAD51."

sparser
"The cDNA of UCH-L3 was subcloned from the library plasmid with GenBank N27190 by PCR into the Eco RI/ Xho I sites of pGEX-4T1 (Amersham) encoding for a GST-UCH-L3 fusion protein."

sparser
"1 Experimental Models: Cell Lines HEK293T Laboratory of Michiel Vermeulen N/A HeLa Cells Spruijt et al., 2013 N/A IB10 Cells Spruijt et al., 2013 N/A NPCs Spruijt et al., 2013 N/A HEK293T-EGFP-TAB-NZF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
CLPB affects UCHL3
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
BRD4 affects UCHL3
2 | 2
2 | 2

reach
"Further investigation also showed the endogenous BRD4 binding with UCHL3 which was enhanced by rIL6 but inhibited by pacritinib (Figs."

No evidence text available

reach
"Of note, our experiments were performed in the presence of IL6/8. whether other deubiquitinases are preferential in other contexts need to be further investigated.To investigate if UCHL3 binds to BRD4, we performed reciprocal co-immunoprecipitation assays in cells transfected with BRD4, alone or with UCHL3 and JAK2-V617F."

No evidence text available
4,4'-sulfonyldiphenol increases the amount of UCHL3.
2 |
4,4'-sulfonyldiphenol increases the amount of UCHL3. 2 / 2
2 |

No evidence text available

No evidence text available
4,4'-sulfonyldiphenol decreases the amount of UCHL3.
2 |
4,4'-sulfonyldiphenol decreases the amount of UCHL3. 2 / 2
2 |

No evidence text available

No evidence text available
UbpA affects UCHL3
| 3
| 2

reach
"Therefore, the co-crystal structure of UCHL3 bound to Ub-PA (UCHL3 -Ub-PA) at high resolution (2."

reach
"The results for UCHL3 bound to Ub-PA are shown in Supplementary Figure S2."
UCHL3-S75E binds ubpA. 1 / 1
| 1

reach
"UCHL3 S75E Bound to Ub-PA."
3 |
Trichostatin A increases the amount of UCHL3. 3 / 3
3 |

No evidence text available

No evidence text available

No evidence text available
3 |
Sodium arsenite increases the amount of UCHL3. 3 / 3
3 |

No evidence text available

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3 |
Cobalt dichloride decreases the amount of UCHL3. 3 / 3
3 |

No evidence text available

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No evidence text available
ZMYM6 affects UCHL3
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
Ubiquitin affects UCHL1
| 3
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
1 2 |
UCHL3 binds.
2 |
2 |

No evidence text available

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UCHL3 is active.
1 |
UCHL3 phosphorylated on S75 is active. 1 / 1
1 |

"The deubiquitinase UCHL3 is phosphorylated and activated by ATM. UCHL3, in turn, deubiquitinates RAD51 and promotes the binding between RAD51 and BRCA2. |Mutation of S75 (S75A) abolished the pSQ/TQ signal, suggesting that S75 is a major ATM phosphorylation site following DNA damage"
UCHL3 affects ubpA
| 3
| 2

reach
"Therefore, the co-crystal structure of UCHL3 bound to Ub-PA (UCHL3 -Ub-PA) at high resolution (2."

reach
"The results for UCHL3 bound to Ub-PA are shown in Supplementary Figure S2."
UCHL3-S75E binds ubpA. 1 / 1
| 1

reach
"UCHL3 S75E Bound to Ub-PA."
UCHL3 affects glioblastoma multiforme cancer stem cell
| 3
UCHL3 activates glioblastoma multiforme cancer stem cell. 3 / 3
| 3

reach
"UCHL3 promotes GSC progression and IR resistance via POLD4."

reach
"At the molecular level, under IR exposure UCHL3 knockdown reduced the protein expression of well-known MES GSC markers (CD44, TAZ, etc.), and POLD4 overexpression restored the expression of these markers (Fig. 6H)."

reach
"Regulating biological functions, UCHL3 promoted GSC proliferation, tumorigenesis, self-renewal, and radioresistance in a POLD4-dependent manner.The degradation of POLD4 by ubiquitin ligases in response to DNA damage and during cell cycle progression plays a less prominent role in normal cells [43], while with increasing insights into the complex mechanisms of DNA damage repair, POLD4 is projected to play an increasingly important role."
| 3

reach
"Results of flow cytometry then showed that UCHL3 knockdown augmented cell apoptosis, along with increased cells arrested in the G0/G1 phase, decreased cells arrested in the S phase, and repressed cell division, while the restoration of UCHL3 caused opposite results (Fig. 3E, F)."

reach
"Programmed cell death is implicated in a number of human diseases, including neurodegenerative disease, 3 autoimmune disease, 4 cancers 5,6 , etc. Loss of UCH-L3 leads to programmed cell death by apop[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Furthermore, under conditions of IR, the level of apoptosis increased by silencing UCHL3 was overcome by POLD4 overexpression in MES GSCs (Fig. 6D and Fig. S7A)."
UCHL3 affects ZMYM6
3 |
3 |

No evidence text available

No evidence text available

No evidence text available
UCHL3 affects XRCC1
| 3
UCHL3 activates XRCC1.
| 2
UCHL3 activates XRCC1. 2 / 2
| 2

reach
"In conclusion, our findings suggest that UCHL3 promotion of RCC tumorigenesis may be achieved through inactivating CD8 T cells."

reach
"The remodeling of the tumor microenvironment may be one of the factors contributing to the promotion of RCC by UCHL3 and UCHL5."
UCHL3 inhibits XRCC1.
| 1
UCHL3 inhibits XRCC1. 1 / 1
| 1

reach
"According to the GO analysis, UCHL3 expressed in CD8 T cells could inhibit the activation of RCC-infiltrating T cells, whereas UCHL5 may impede vacuolar transport instead of antigen processing and presentation in B cells infiltrating RCC."
UCHL3 affects TGFB
| 3
UCHL3 activates TGFB.
| 2
UCHL3 activates TGFB. 2 / 2
| 2

reach
"Overexpression of UCH-L1, but not of UCH-L3 (the other human homolog of UBH1) or of the catalytic mutant UCH L1C90A, enhanced TGF-beta and SMAD-induced transcriptional activity, indicating that the deubiquitination activity of UCH-L1 is indispensable for enhancing TGF-beta and SMAD signaling."

reach
"Overexpression of UCH-L1, but not of UCH-L3 (the other human homolog of UBH1) or of the catalytic mutant UCH-L1 C90A, enhanced TGF-beta and SMAD-induced transcriptional activity, indicating that the deubiquitination activity of UCH-L1 is indispensable for enhancing TGF-beta and SMAD signaling."
UCHL3 deubiquitinates TGFB.
| 1
UCHL3 deubiquitinates TGFB. 1 / 1
| 1

reach
"UCHL3 deubiquitinates p300 in response to TGF-beta 1 signaling activation."
UCHL3 affects SLC4A1
| 3
| 3

reach
"To confirm the direct interaction of di-Ub and UCH-L3, we performed pull-down assays using recombinant mono-Ub or di-Ub and UCH-L3."

reach
"Likewise, UCH-L3 might interact with di-Ub to protect it from degradation."

reach
"Thus, we can not exclude the possibility that, under some conditions, UCH-L3 may interact with di-Ub to protect it from degradation.Previous biochemical analysis indicated that UCH-L3 is unable to hyd[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects SHBG
| 3
USP5 binds UCHL3 and SHBG. 2 / 2
| 2

sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."
| 1

sparser
"The non-covalent binding of OTULIN ABP to UCHL3 is most likely due to the high abundance of this enzyme."
UCHL3 affects PRKDC
1 | 2
1 | 1

sparser
"Our mass spectrometry and ensuing immunoprecipitation analysis indicated that UCHL3 could interact with DNA-PKcs, possibly suggesting context specific functions of UCHL3 for maintaining genome integrity."

No evidence text available
UCHL3 binds Ku and PRKDC. 1 / 1
| 1

sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
UCHL3 affects OTULIN ABP
| 3
UCHL3 binds OTULIN ABP. 3 / 3
| 3

reach
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non covalent and reversible manner."

reach
"The non covalent binding of OTULIN ABP to UCHL3 is most likely due to the high abundance of this enzyme."

reach
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non covalent and reversible."
UCHL3 affects Ku
| 1 2
UCHL3 binds Ku.
| 2
UCHL3 binds Ku and PRKDC. 1 / 1
| 1

sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
| 1

sparser
"The immunoprecipitation of endogenous UCHL3 using an anti-UCHL3 antibody followed by immunoblotting analysis detected the interaction with endogenous Ku80, providing further support that UCHL3 interacts with Ku (Fig.  xref )."
UCHL3 ubiquitinates Ku.
| 1
UCHL3 leads to the ubiquitination of Ku. 1 / 1
| 1

reach
"Lastly, we provide evidence showing that DNA damage-induced UCHL3 phosphorylation (which is dependent on UCHL3 catalytic activity, ATM kinase and downstream NHEJ factors) promotes destabilization of UCHL3, suggesting that UCHL3 may dissociate from Ku80 to allow Ku ubiquitylation and subsequent removal from DNA."
UCHL3 affects IFNAR1
| 3
UCHL3 increases the amount of IFNAR1.
| 2
UCHL3 increases the amount of IFNAR1. 2 / 2
| 2

reach
"In summary, our results demonstrate that UCHL3 upregulates the levels of SCF beta-TrCP substrates.We have demonstrated that UCHL3 positively regulated COPS5 levels and IFNAR1 levels."

reach
"These findings suggest that UCHL3 could positively regulate IFNAR1 levels via COPS5 mediated deneddylation of Cullin1, which is consistent with our previous findings that UCHL3 positively regulates IF[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 activates IFNAR1.
| 1
UCHL3 activates IFNAR1. 1 / 1
| 1

reach
"This finding suggests that UCHL3 could target either TYK2 itself or IFN-I receptors (IFNAR1 and IFNAR2), which are the direct proteins upstream of TYK2 as well as the first signaling proteins in IFN-I[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL1 affects Ubiquitin
| 3
| 3

sparser
"T218, R227, F228, and L230 residues of BAP1N were selected based on ubiquitin interaction with the corresponding residues of UCHL1 and UCHL3."

sparser
"In contrast with BAP1, the thermodynamics of ubiquitin binding to UCHL1 and UCHL3 is dominated by enthalpy factors and hydrogen bonding and ionic interactions that stabilize the complex."

sparser
"Unlike UCH-L1, UCH-L3 can interact with Lys48-linked Ub dimers (di-Ub) to protect it from degradation and meanwhile to inhibit its hydrolase activity."
UBD affects UCHL3
| 1 1 1
UBD inhibits UCHL3. 3 / 3
| 1 1 1

sparser
"K27-Linked Diubiquitin Inhibits UCHL3 via an Unusual Kinetic Trap."

reach
"K27 Linked Diubiquitin Inhibits UCHL3 via an Unusual Kinetic Trap."

trips
"K27-Linked Diubiquitin Inhibits UCHL3 via an Unusual Kinetic Trap."
SHBG affects UCHL3
| 3
USP5 binds UCHL3 and SHBG. 2 / 2
| 2

sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."
| 1

sparser
"The non-covalent binding of OTULIN ABP to UCHL3 is most likely due to the high abundance of this enzyme."
PRKDC affects UCHL3
1 | 2
1 | 1

sparser
"Our mass spectrometry and ensuing immunoprecipitation analysis indicated that UCHL3 could interact with DNA-PKcs, possibly suggesting context specific functions of UCHL3 for maintaining genome integrity."

No evidence text available
UCHL3 binds Ku and PRKDC. 1 / 1
| 1

sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
OTULIN ABP affects UCHL3
| 3
UCHL3 binds OTULIN ABP. 3 / 3
| 3

reach
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non covalent and reversible manner."

reach
"The non covalent binding of OTULIN ABP to UCHL3 is most likely due to the high abundance of this enzyme."

reach
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non covalent and reversible."
JUN affects UCHL3
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JUN decreases the amount of UCHL3. 3 / 3
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HELLS affects UCHL3
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Delta12-PGJ2 affects UCHL3
| 3
Delta12-PGJ2 inhibits UCHL3. 3 / 3
| 3

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"As shown in Fig. 5, Delta12-PGJ2 inhibited the activities of UCH-L1 (K i = 3.49 muM +/- 0.42, Fig. 5A) and UCH-L3 (K i = 8.1 muM +/- 0.8, Fig. 5B), indicating that this PGJ2 metabolite is an efficient[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"Delta12-PGJ2 and 15Delta-PGJ2 are potent cyclopentone PGs and have been reported to inhibit UCH-L3 and UCH-L1 respectively [282,283]."

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"Since Delta12-PGJ2 inhibited UCH-L1 and UCH-L3 activities, we postulated that this cyclopentenone prostaglandin would affect the endogenous levels of ubiquitinated proteins."
Δ affects UCHL3
| 2
Δ inhibits UCHL3. 2 / 2
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"Other investigators have reported that cyclopentone PGs inhibit cellular DUB activities: UCHL3 was found to be inhibited by Δ12-PGJ2 ( Li et al., 2004 ) and UCHL1 by 15Δ-PGJ2 ( Liu et al., 2011c )."

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"Treatment with Δ -PGJ in cells also inhibited UCH-L1 and UCH-L3 without alteration of proteasomal activity, indicating that prostaglandins can be suitable for neural disorder therapy [95]."
| PMC
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Titanium dioxide increases the amount of UCHL3.
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Titanium dioxide increases the amount of UCHL3. 1 / 1
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Titanium dioxide demethylates UCHL3.
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"Sodium citrate, at high concentrations, activated OTU-1, UCH-L3 and PLpro, and this could be due to an ionic effect or a Hofmeister effect."

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"Sodium citrate at 0.8 M enhanced activity of UCH-L3 by 12-fold versus 0.8 M NaCl for Ac-LRGG-AFC and 20-fold for Ac-ALRGG-AFC (XREF_FIG)."
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Pirinixic acid increases the amount of UCHL3. 2 / 2
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Phenylmercury acetate increases the amount of UCHL3. 2 / 2
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Peptide affects UCHL3
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Peptide activates UCHL3. 2 / 2
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"The largest stabilization was seen upon the addition of peptide 60297, which increased the melting temperature of PfUCHL3 by 5.1 °C, followed by peptide 60296 which increased the melting temperature by 3.1 °C at 25 μM. Peptides 58474 and 58474(2) were the only other two peptides to cause a significant increase in the melting temperature (1.2 °C and 1.3 °C, respectively)."

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"Peptide 60297 induced similar changes in PfUCHL3, albeit to a slightly lesser extent."
Paracetamol affects UCHL3
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Paracetamol increases the amount of UCHL3. 2 / 2
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| 2

sparser
"Pull-down analysis using recombinant UCH-L3 demonstrated that wild-type UCH-L3 interacted with polyubiquitinated lysozyme but not with free lysozyme (data not shown) or mono- and di-ubiquitinated lyso[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"UCH-L3 interacted with polyubiquitinated lysozyme in a pull-down analysis that was abrogated in D33A (lacking affinity for Ub) mutants ( Fig. 4 E)."
Lysine affects UCHL3
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UCHL3 binds EEF1A1 and lysine. 2 / 2
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"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

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"EEF1A1 binds to UCHL3 through the lysine site."
Lsh affects UCHL3
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UCHL3 binds lsh. 2 / 2
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sparser
"In addition, the disturbance of the interaction between UCH-L3 and LSH by the long non-coding RNA GIAT4RA affects this progression [ 116 ]."

sparser
"However, HOTAIR was also reported to reduce the ubiquitination degradation of AR by preventing its combination with MDM2, and GIAT4RA was recently reported to interfere the interaction between Uchl3 and LSH to repress ubiquitin-proteasome pathway [ xref ], which indicate multiple roles of lncRNA in ubiquitination."
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Hsa-mir-3941 decreases the amount of UCHL3. 2 / 2
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Hsa-miR-875-5p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-8485 decreases the amount of UCHL3. 2 / 2
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Hsa-miR-5580-3p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-5011-5p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-4789-3p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-362-3p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-329-3p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-3144-3p decreases the amount of UCHL3. 2 / 2
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Hsa-miR-190a-3p decreases the amount of UCHL3. 2 / 2
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Cyclosporin A increases the amount of UCHL3. 2 / 2
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| 2

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"These points highlight the ubiquitin pathway as a promising target for the development of novel antimalarials.In this study, we used the RaPID selection technique to identify cyclic peptide inhibitors of PfUCHL3, a dual activity DUB/deneddylating enzyme of P. falciparum, the deadliest malaria parasite."

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"The cyclic peptide inhibits UCH-L3 ten times less efficiently (IC = 22.5 µM), while BAP1 is inhibited three times less efficiently than UCH-L5 (IC = 6.8 µM)."
Bisphenol A affects UCHL3
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Bisphenol A decreases the amount of UCHL3. 2 / 2
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ZEB1 affects UCHL3
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USP5 affects SHBG
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USP5 binds UCHL3 and SHBG. 2 / 2
| 2

sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."
USP48 affects UCHL3
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"(E, F) Effects of UCHL3 overexpression (E; n =  4) and knockdown (F; n =  6) on gefitinib-induced cell cycle arrest."

reach
"Additionally, UCHL3 overexpression mitigated gefitinib-induced G0/G1 cell cycle arrest in PC9 cells (Fig 2E)."
| 2

sparser
"Pull-down analysis using recombinant UCH-L3 demonstrated that wild-type UCH-L3 interacted with polyubiquitinated lysozyme but not with free lysozyme (data not shown) or mono- and di-ubiquitinated lyso[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"UCH-L3 interacted with polyubiquitinated lysozyme in a pull-down analysis that was abrogated in D33A (lacking affinity for Ub) mutants ( Fig. 4 E)."
UCHL3 affects lysine
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UCHL3 binds EEF1A1 and lysine. 2 / 2
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"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

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"EEF1A1 binds to UCHL3 through the lysine site."
UCHL3 affects lsh
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UCHL3 binds lsh. 2 / 2
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sparser
"In addition, the disturbance of the interaction between UCH-L3 and LSH by the long non-coding RNA GIAT4RA affects this progression [ 116 ]."

sparser
"However, HOTAIR was also reported to reduce the ubiquitination degradation of AR by preventing its combination with MDM2, and GIAT4RA was recently reported to interfere the interaction between Uchl3 and LSH to repress ubiquitin-proteasome pathway [ xref ], which indicate multiple roles of lncRNA in ubiquitination."

reach
"Our results indicate that highly expressed UCHL3 enhances inflammation by stabilizing TRAF2, which in turn facilitates tumourigenesis in ovarian cancer, and that UCHL3 is a potential target for ovarian cancer patients with increased inflammation."

eidos
"The elevated UCHL3 facilitates carcinogenesis and enhances inflammation by deubiquitinating and stabilizing TNF Receptor Associated Factor 2 ( TRAF2 ) ( 74 ) ."

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"UCHL3 promotes aerobic glycolysis of pancreatic cancer through upregulating LDHA expression."

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"UCHL3 enhances drug resistance and glycolysis in pancreatic cancer through the stabilization of FOXM1 and upregulation of LDHA [6, 7]."
UCHL3 affects ferroptosis
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"Furthermore, the depletion of UCHL3 induces ferroptosis and hinders the growth, invasion, and stem cell properties of HCC cells."

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"UCHL3 inhibits ferroptosis by stabilizing β-catenin and maintains stem-like properties of hepatocellular carcinoma cells."

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"The results of this investigation are consistent with a work by Suzuki et al. (39), who demonstrated that the reduction of UCHL3 induced an impairment in the adipogenesis that resulted in the prevention of lipid accumulation in adipocytes derived from UCHL3-/- mice."

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"Furthermore, UCHL3 has deneddylating activity essential for various cellular functions and promotes insulin signaling and adipogenesis, hence contributing to retinal maintenance in stress conditions [15]."

eidos
"Together , these data support the idea that UCHL3 promotes c-NHEJ , not HR , in a catalytic activity-dependent manner ."

reach
"Together, these data support the idea that UCHL3 promotes c-NHEJ, not HR, in a catalytic activity-dependent manner."
UCHL3 affects cell death
| 2
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"29 Therefore, loss of UCH-L3 may mildly promote the cell death of these cells.After 3w of age, prominent and progressive photoreceptor cell apoptosis was disclosed in the outer nuclear layer of Uchl3 [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

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"6 On the other hand, loss of UCH-L3 is known to induce cell death in knock-out studies."
UCHL3 affects autophagy
| 2
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"Taken together, the results of the present study suggest that UCHL3 knockdown decreases melanoma cell proliferation by increasing cell autophagy through regulating NEDD8 expression and autophagosome numbers."

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"Furthermore, autophagy was significantly enhanced along with the decrease in NEDD8 ubiquitination.In conclusion, the results of the present study demonstrated that UCHL3 knockdown decreased melanoma cell proliferation by increasing cell autophagy through regulating NEDD8 expression and autophagosome numbers in vitro."
UCHL3 affects ZEB1
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UCHL3 affects USP48
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UCHL3 affects UBQLN1
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UCHL3 affects UBA6
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"The deubiquitinating enzyme UCHL3 promotes anaplastic thyroid cancer progression and metastasis through Hippo signaling pathway."

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"UCHL3 increases the stability of the YAP protein in a deubiquitinating activity-dependent manner, thereby promoting the progression and metastasis of anaplastic thyroid cancer [27]."
UCHL3 affects TRIM63
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UCHL3 affects TRIM55
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UCHL3 affects Recurrence
| 2
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"Overall, these results showed that UCH-L1 and UCH-L3 could be effective therapeutic targets to eliminate the recurrence caused by CSCs in prostate cancers."

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"Since the differences in stress responses between GCSCs and GCICs appeared to be due to UCH-L3 expression levels, we stipulated that the regulation of UCH-L3 expression in response to autophagy could suppress cancer recurrence induced by GCSCs."
UCHL3 affects PARP1
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UCHL3 inhibits PARP1.
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UCHL3 inhibits PARP1. 1 / 1
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"(A, B) Effects of UCHL3 overexpression (A) and knockdown (B) on gefitinib-induced EGFR degradation and PARP cleavage."
UCHL3 activates PARP1.
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UCHL3 activates PARP1. 1 / 1
| 1

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"Overexpression of UCHL3 attenuated gefitinib-induced EGFR downregulation and increased PARP cleavage, while UCHL3 knockdown intensified these effects (Fig 2B)."
UCHL3 affects OTULIN
1 | 1
1 | 1

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"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."

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"The CSC-like properties were enhanced by UCH-L1 but inhibited by UCH-L3."

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"In DU145 cells, the CSC-like properties were inhibited by UCH-L1 knockdown or UCH-L3 overexpression."
UCHL3 affects Mice
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UCHL3 activates Mice. 2 / 2
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"Uchl3 deficiency leads to inhibited BBN-induced bladder tumorigenesis in mice."

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"Nevertheless, according to Fig. 6F, we successfully established a BBN-induced bladder cancer model and found that the deletion of UCHL3 reduced the rate of BBN-induced bladder cancerogenesis in the Uchl3 ;Upk2-Cre mice compared with the control group (Fig. 6G)."
UCHL3 affects Melanoma
| 2
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"Taken together, the results of the present study suggest that UCHL3 knockdown decreases melanoma cell proliferation by increasing cell autophagy through regulating NEDD8 expression and autophagosome numbers."

reach
"Furthermore, autophagy was significantly enhanced along with the decrease in NEDD8 ubiquitination.In conclusion, the results of the present study demonstrated that UCHL3 knockdown decreased melanoma cell proliferation by increasing cell autophagy through regulating NEDD8 expression and autophagosome numbers in vitro."
UCHL3 affects MYC
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UCHL3 increases the amount of MYC. 2 / 2
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"As expected, overexpression of UCHL3 increased the expression of ABCG2, c-Myc and KLF4, genes associated with stemness, at the protein level in A549 and H1299 cells (Fig. 5b, c), while depletion of UCHL3 decreased their protein expression (Fig. 5d)."

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"WT UCHL3 obviously increased AhR and c-Myc protein levels, whereas the mutants did not (Fig. 6a)."
UCHL3 affects MES
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UCHL3 activates MES. 2 / 2
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"At the molecular level, under IR exposure UCHL3 knockdown reduced the protein expression of well-known MES GSC markers (CD44, TAZ, etc.), and POLD4 overexpression restored the expression of these markers (Fig. 6H)."

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"For the HDR and NHEJ reporter assay, silencing UCHL3 reduced HDR and NHEJ efficiency in MES GSCs followed by IR, and ectopic expression of POLD4 overcame the inhibitory effects of UCHL3 depletion on MES GSCs (Fig. 6F, G and Fig. S7B, C)."
UCHL3 affects ENO1
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UCHL3 affects DNA repair
| 2
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"Silencing UCHL3 enhances radio-sensitivity of non small cell lung cancer cells by inhibiting DNA repair."

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"We hypothesized that increased UCHL3 expression in breast cancer cells would increase DNA repair capability and render cells resistant to cancer therapy."
UCHL3 affects DNA Damage
| 2
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"Regulating biological functions, UCHL3 promoted GSC proliferation, tumorigenesis, self-renewal, and radioresistance in a POLD4-dependent manner.The degradation of POLD4 by ubiquitin ligases in response to DNA damage and during cell cycle progression plays a less prominent role in normal cells [43], while with increasing insights into the complex mechanisms of DNA damage repair, POLD4 is projected to play an increasingly important role."

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"UCHL3 enhanced the DNA damage repair mediated by radiotherapy and chemotherapy in cancer cells via binding to tyrosyl DNA phosphodiesterase 1 (TDP1), a chromosome breakage repair related enzyme, and mediating TDP1 deubiquitination [XREF_BIBR - XREF_BIBR]."
UCHL3 affects Chemokine
| 2
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"These data indicate that UCHL3 mediates p300/C/EBPβ-dependent chemokine signaling in ATII cells."

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"The deubiquitinase UCHL3 mediates p300-dependent chemokine signaling in alveolar type II cells to promote pulmonary fibrosis."
| 1 1
| 1 1

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"UCHL3 promotes cell survival after DSB induction and is epistatic with XRCC4."

eidos
"UCHL3 promotes cell survival after DSB induction and is epistatic with XRCC4 To further explore whether UCHL3 plays a role in cellular responses to DSBs , clonogenic survival assays were performed upon treating cells with various short interfering RNAs ( siRNAs ) targeting UCHL3 ."

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"UCHL3 promotes hepatocellular carcinoma progression by stabilizing EEF1A1 through deubiquitination."

reach
"UCHL3 promotes hepatocellular carcinoma cell migration by de-ubiquitinating and stabilizing Vimentin."
UCHL3 affects CRY2 methylation
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UCHL3 activates CRY2 methylation. 2 / 2
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eidos
"In addition , UCHL3 knockdown decreased CRY2 methylation through downregulating FOXM1 , leading to an increase in the expression of CRY2 ."

eidos
"In the current study , we aimed to investigate the role of UCHL3 in ESCC growth and migration , and whether UCHL3 could modulate CRY2 methylation through FOXM1 ."
UCHL3 affects ATXN7L3
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UCHL3 affects ACAA2
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UBQLN1 affects UCHL3
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UBA6 affects UCHL3
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TRIM63 affects UCHL3
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TRIM55 affects UCHL3
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No evidence text available
TEAD4 affects UCHL3
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TEAD4 increases the amount of UCHL3. 2 / 2
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"YAP/TEAD4 promotes progression and metastasis of anaplastic thyroid cancer by binding to the UCHL3 promoter and activating UCHL3 transcription [40]."

reach
"UCHL3 promoter analysis revealed that TEAD4, through which YAP bind to DNA, activated UCHL3 transcription by binding to the promoter of UCHL3."
TCID affects UCHL3
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TCID inhibits UCHL3. 2 / 2
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"To further demonstrate our findings, we used TCID, a small-molecule inhibitor of UCHL3; under our experimental conditions, we first detected whether TCID inhibits the deubiquitinase activity of UCHL3 against AhR."

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"Moreover, TCID attenuated UCHL3 activity without affecting its protein level."
SLC11A1 affects UCHL3
| 2
| 2

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"At the same time, the lncRNA GIAT4RA counteracts the deubiquitinating effects of LSH by interfering with interaction between LSH and UCH-L3, reducing levels of LSH protein, and playing the role in inhibiting the invasion and migration of LUAD cells83."

reach
"However, HOTAIR was also reported to reduce the ubiquitination degradation of AR by preventing its combination with MDM2, and GIAT4RA was recently reported to interfere the interaction between Uchl3 and LSH to repress ubiquitin-proteasome pathway [30], which indicate multiple roles of lncRNA in ubiquitination."
SHBG affects USP5
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USP5 binds UCHL3 and SHBG. 2 / 2
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sparser
"These results suggest that despite the partial labeling of USP5 in cell extracts, USP5 and UCHL3 are primarily bound to the OTULIN ABP in a non-covalent and reversible manner."

sparser
"Thus, the binding of USP5 and UCHL3 to OTULIN ABP is largely non-covalent and reversible."
RAG2 affects UCHL3
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RAG2 inhibits UCHL3. 2 / 2
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"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTULIN affects UCHL3
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"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."
MIR7-3HG affects UCHL3
| 2
| 2

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"Firstly, collect HUH7 cell lines overexpressing UCHL3 and extract the total protein."

reach
"Huh7 and Hep3B cells stably overexpressing UCHL3 were generated ( Figures 2A, B )."
MIR603 affects UCHL3
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MIR603 decreases the amount of UCHL3. 2 / 2
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Ku affects UCHL3
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UCHL3 binds Ku and PRKDC. 1 / 1
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sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
| 1

sparser
"The immunoprecipitation of endogenous UCHL3 using an anti-UCHL3 antibody followed by immunoblotting analysis detected the interaction with endogenous Ku80, providing further support that UCHL3 interacts with Ku (Fig.  xref )."
Gentamicins affects UCHL3
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Gentamicins increases the amount of UCHL3.
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Gentamicins increases the amount of UCHL3. 1 / 1
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Gentamicins decreases the amount of UCHL3.
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Gentamicins decreases the amount of UCHL3. 1 / 1
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No evidence text available
ENO1 affects UCHL3
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EEF1A1 affects lysine
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UCHL3 binds EEF1A1 and lysine. 2 / 2
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"UCHL3 binds to EEF1A1 through the lysine site, which reduces the ubiquitination level of EEF1A1."

reach
"EEF1A1 binds to UCHL3 through the lysine site."
ATXN7L3 affects UCHL3
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No evidence text available
AM146 affects UCHL3
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AM146 inhibits UCHL3. 2 / 2
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"AM146, RA-9, and RA-14 were found to inhibit UCHL1, UCHL3, USP2 and USP8, but did not suppress the activity of Ataxin-3, A20, BAP1, Otubain 1 or USP7 ( Issaenko & Amerik, 2012 )."

reach
"Additionally, chalcone-based derivatives AM146, RA-9 and RA-14, which contain α,β-unsaturated carbonyl groups, directly target and suppress the activity of UCH-L1, UCH-L3, USP2, USP5 and USP8 without [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
ACAA2 affects UCHL3
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Undecane affects UCHL3
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Undecane decreases the amount of UCHL3. 1 / 1
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No evidence text available
Trimellitic anhydride increases the amount of UCHL3. 1 / 1
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No evidence text available
| 1

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"Pimozide, flupenthixol and trifluoperazine were poor inhibitors of UCH-L1 and UCH-L3 (IC 50> 500 muM)."
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Sorafenib affects UCHL3
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Sorafenib decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Sodium chloride decreases the amount of UCHL3. 1 / 1
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No evidence text available
Sirolimus affects UCHL3
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Sirolimus increases the amount of UCHL3. 1 / 1
| 1

reach
"The aim of this study was to explore whether rapamycin can induce the autophagy of HL-60 cells and UCH-L3 expression."
1 |
Silicon dioxide decreases the amount of UCHL3. 1 / 1
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No evidence text available
Rotenone affects UCHL3
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Rotenone increases the amount of UCHL3. 1 / 1
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No evidence text available
Reported Akt affects UCHL3
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Reported Akt activates UCHL3. 1 / 1
| 1

eidos
"Here we report that perifosine , a previously reported Akt inhibitor , can inhibit UCHL3 in vitro and in vivo ."
Quercetin affects UCHL3
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Quercetin decreases the amount of UCHL3. 1 / 1
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No evidence text available
Polyubiquitin affects UCHL3
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UCHL3 binds polyubiquitin. 1 / 1
| 1

sparser
"UCH-L3 binds to K48-linked polyubiquitin to protect against degradation and inhibit hydrolytic enzyme activity [ xref ]."
Pimozide affects UCHL3
| 1
| 1

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"Pimozide, flupenthixol and trifluoperazine were poor inhibitors of UCH-L1 and UCH-L3 (IC 50> 500 muM)."
| 1

eidos
"Furthermore , this phosphorylation destabilizes UCHL3 , despite having no effect on its catalytic activity ."
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No evidence text available
Perifosine affects UCHL3
| 1
| 1

eidos
"Here we report that perifosine , a previously reported Akt inhibitor , can inhibit UCHL3 in vitro and in vivo ."
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No evidence text available
Ozone affects UCHL3
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Ozone increases the amount of UCHL3. 1 / 1
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No evidence text available
MiR-582-5p affects UCHL3
| 1
MiR-582-5p increases the amount of UCHL3. 1 / 1
| 1

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"LINC00665 sponges miR-582-5p to up-regulate UCHL3 expression."
Methylmercury chloride decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Methapyrilene increases the amount of UCHL3. 1 / 1
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No evidence text available
Menadione affects UCHL3
| 1
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"Moreover, the addition of the chemical inhibitor menadione 11 (4,5,6,7-tetrachloroindan-1,3-dione; IC50 > 0.6 μM ( Liu et al., 2003 )) along with UCH-L3, also abrogated the de-ubiquitinating function [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
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Lead(II) chloride increases the amount of UCHL3. 1 / 1
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Lead atom affects UCHL3
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Lead atom increases the amount of UCHL3. 1 / 1
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No evidence text available
Jinfukang affects UCHL3
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Jinfukang decreases the amount of UCHL3. 1 / 1
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No evidence text available
Ivermectin affects UCHL3
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Ivermectin decreases the amount of UCHL3. 1 / 1
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No evidence text available
Hydrolase affects UCHL3
| 1
Hydrolase decreases the amount of UCHL3. 1 / 1
| 1

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"We also found that polyubiquitinated protein accumulation can be reduced by the expression of wild-type UCH-L3 but not hydrolase activity-deficient (C95S) UCH-L3 in MEFs derived from Uchl3 −/− mice ( [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
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Hsa-mir-4534 decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-mir-3134 decreases the amount of UCHL3. 1 / 1
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No evidence text available
1 |
Hsa-miR-8082 decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-7156-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-532-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
1 |
Hsa-miR-514b-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-5093 decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-4633-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-372-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-3688-3p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-3124-3p decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Hsa-miR-130b-5p decreases the amount of UCHL3. 1 / 1
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No evidence text available
Hexabromocyclododecane decreases the amount of UCHL3. 1 / 1
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No evidence text available
Gefitinib affects UCHL3
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"These results demonstrate that extended gefitinib exposure induces sustained UCHL3 downregulation and USP54 upregulation in gefitinib-resistant sublines, suggesting a potential interplay between these genes in NSCLC cells under prolonged drug exposure.Total RNA was extracted from PC9 cells treated with 1 μM gefitinib for 1, 6, and 24 h and analyzed by QuantSeq 3’ mRNA sequencing."
Furfural affects UCHL3
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Furfural decreases the amount of UCHL3. 1 / 1
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No evidence text available
Folic acid affects UCHL3
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Folic acid decreases the amount of UCHL3. 1 / 1
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No evidence text available
Fenthion affects UCHL3
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Fenthion increases the amount of UCHL3. 1 / 1
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No evidence text available

sparser
"These endogenous electrophiles (1) inhibit ubiquitin isopeptidase activity [ xref ] as well as ubiquitin hydrolases UCH-L1 and UCH-L3 [ xref ], (2) induce the formation of cysteine-targeted thyolation of UCH-L1 [ xref ] and of PGJ2/proteasome conjugates [ xref ], (3) trigger the oxidation of the S6 ATPase subunit of the 26S proteasome [ xref ], and disrupt 26S proteasome assembly [ xref ]."

No evidence text available
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Dibutyl phthalate decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Dibenzothiophene decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Deltamethrin increases the amount of UCHL3. 1 / 1
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No evidence text available
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Cyanoginosin LR decreases the amount of UCHL3. 1 / 1
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No evidence text available
Chalcone affects UCHL3
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"RA-9, a chalcone derivative with a structure similar to b-AP15, was reported to inhibit proteasomal DUBs [XREF_BIBR] as well as UCHL1, UCHL3, USP2, USP5, and USP8 [XREF_BIBR]."
Cannabidiol affects UCHL3
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Cannabidiol decreases the amount of UCHL3. 1 / 1
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No evidence text available
Bisphenol F affects UCHL3
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Bisphenol F decreases the amount of UCHL3. 1 / 1
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No evidence text available
Bisphenol B affects UCHL3
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Bisphenol B increases the amount of UCHL3. 1 / 1
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No evidence text available
Bis(2-ethylhexyl) phthalate increases the amount of UCHL3. 1 / 1
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No evidence text available
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Beta-lapachone increases the amount of UCHL3. 1 / 1
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No evidence text available
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Beta-D-glucose increases the amount of UCHL3. 1 / 1
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No evidence text available
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Benzatropine decreases the amount of UCHL3. 1 / 1
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No evidence text available
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Arsenite(3-) decreases the amount of UCHL3. 1 / 1
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No evidence text available
Aldehyde affects UCHL3
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sparser
"We found complex viscosity dependencies and interpreted these in the context of a model in which association and acylation are viscosity-dependent but deacylation is viscosity-independent. (2) The kinetics of inhibition of UCH-L3 by ubiquitin C-terminal aldehyde (Ub-H) were determined and reveal a Ki that is less than 10(-14) M. Several mechanisms are considered to account for the extreme inhibition. (3) The IPaseT-catalyzed hydrolysis of Ub-AMC is modulated by Ub with activation at low [Ub] and inhibition at high [Ub]. (4) Finally, we compare kc/Km values for deubiquitinating enzyme-catalyzed hydrolysis of Ub-AMC and Z-Leu-Arg-Gly-Gly-AMC."
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"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 (XREF_FIG and XREF_FIG) and the yeast UCH Yuh1 XREF_BIBR, XREF_BIBR."
ZRANB2 affects UCHL3
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No evidence text available
ZNRD2 affects UCHL3
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No evidence text available
ZFYVE19 affects UCHL3
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No evidence text available
XRCC6 affects UCHL3
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No evidence text available
VEA-260 affects UCHL3
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VEA-260 inhibits UCHL3. 1 / 1
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"In contrast, hSENP6 (a homolog of PfSENP2) and UCH-L3 (a DUB) were not inhibited by VEA-260."
UbVME affects UCHL3
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UbVME binds UCHL3. 1 / 1
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"In contrast, when UbVME binds to UCH-L3, the S shaped loop (153-167) is reordered and helix-6 ' (142-152) forms and covers the active site bound to the C-terminus of ubiquitin [13]."
Ub dimers affects UCHL3
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Ub dimers inhibits UCHL3. 1 / 1
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"These different phenotypes of mutant mice indicate that although both enzymes share similar properties in terms of biochemical identity (Johnston et al., 1997; Larsen et al., 1998; Misaghi et al., 200[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
USP2cd affects UCHL3
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USP2cd inhibits UCHL3. 1 / 1
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"At 2% (v/v) DMSO, the proteolytic activity of USP2cd was reduced by ∼20% and that of UCH-L3 by ∼5% in comparison to the DMSO-free conditions."
UCHL3 affects ubiquitination TDP1 leading TDP1
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UCHL3 inhibits ubiquitination TDP1 leading TDP1. 1 / 1
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eidos
"UCHL3 decreases the ubiquitination level of TDP1 leading to the inhibition of TDP1 degradation ."

reach
"The researchers were surprised to discover that their C-terminally propargylated Ub (Ub-Prg), originally synthesized for site specific ubiquitination of peptides, inhibited the human DUB ubiquitin carboxyl-terminal hydrolase isoenzyme L3 (UCHL3)."
UCHL3 affects subsequent removal DNA
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UCHL3 activates subsequent removal DNA. 1 / 1
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eidos
"Lastly , we provide evidence showing that DNA damage-induced UCHL3 phosphorylation ( which is dependent on UCHL3 catalytic activity , ATM kinase and downstream NHEJ factors ) promotes destabilization of UCHL3 , suggesting that UCHL3 may dissociate from Ku80 to allow Ku ubiquitylation and subsequent removal from DNA ."
UCHL3 affects repair
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UCHL3 inhibits repair. 1 / 1
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eidos
"Taken together , these data suggested that the AKT inhibitor perifosine has an off-target effect at low dose by targeting UCHL3 , which in turn impairs RAD51 foci formation and HR repair ."
UCHL3 affects radiosensitivity NSCLC cells
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UCHL3 inhibits radiosensitivity NSCLC cells. 1 / 1
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eidos
"UCHL3 reduced the radiosensitivity of NSCLC cells by stabilizing AhR protein ."
UCHL3 affects proteolysis
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"UCHL3 was responsible for the disassembly of the ubiquitin protein complex or polyubiquitin chains during the substrate priming for proteasomal proteolysis to prevent inappropriate protein degradation [XREF_BIBR]."

reach
"map3k4 is also a protein-coding gene, and studies have shown that high methylation of map3k4 kinase and low methylation of uchl3, observed in seawater-released marine sticklebacks, can potentially reduce protein ubiquitination levels and serve as an adaptation to osmotic stress (Artemov et al., 2017)."
UCHL3 affects polyubiquitin
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UCHL3 binds polyubiquitin. 1 / 1
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"UCH-L3 binds to K48-linked polyubiquitin to protect against degradation and inhibit hydrolytic enzyme activity [ xref ]."
UCHL3 affects polyubiquitin chains
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UCHL3 inhibits polyubiquitin chains. 1 / 1
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"25,26 Mechanistically, USP13 deubiquitinates RAP80 and promotes BRCA1 recruitment to DSBs, whereas UCH-L3 eliminates the polyubiquitin chains of RAD51 and facilitates RAD51–BRCA2 binding."
UCHL3 affects polyubiquitin chain
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UCHL3 inhibits polyubiquitin chain. 1 / 1
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"The polyubiquitin chain is degraded by UCHL3 and further processed by de-ubiquitinating enzymes (DUBs) into monomeric ubiquitin."
UCHL3 affects pY701-STAT1
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UCHL3 increases the amount of pY701-STAT1. 1 / 1
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"The result showed that UCHL3 overexpression robustly increased the level of pY701-STAT1 (p-STAT1) induced by IFNalpha in a time dependent manner."
UCHL3 affects monoubiquitin
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UCHL3 activates monoubiquitin. 1 / 1
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"Recent evidence suggests that monoubiquitin release from its precursors is mediated predominantly by five main DUBs : ubiquitin thioesterase otulin, UCHL3, USP5, USP7 and USP9X [XREF_BIBR], although each has multiple substrates that can be targeted for deubiquitination to maintain the free ubiquitin pool."
UCHL3 affects mono-Ub
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UCHL3 activates mono-Ub. 1 / 1
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"After incubation for 24 h, UCH-L3 did not produce mono-Ub from these three forms of di-Ub (data not shown), which is consistent with previous reports (Larsen et al., 1998)."
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"ALP staining and Alizarin red staining further confirmed a similar rescue effect of SMAD1 overexpression in promoting the decreased ALP level and extracellular matrix mineralization deposition caused by UCHL3 inhibition in MS-BMDM-EXOs (Fig. 7g, h)."
UCHL3 affects migration
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UCHL3 activates migration. 1 / 1
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eidos
"In summary , this study reveals that knockdown of UCHL3 inhibits ESCC growth and migration by reducing CRY2 methylation through downregulation of FOXM1 expression ."
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"Instead, UCHL3 promotes localization of Aurora B to kinetochores, suggesting its role in the error correction mechanism monitoring bi-orientation of chromosomes during metaphase."
UCHL3 affects localization Aurora B kinetochores
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UCHL3 activates localization Aurora B kinetochores. 1 / 1
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eidos
"Instead , UCHL3 promotes localization of Aurora B to kinetochores , suggesting its role in the error correction mechanism monitoring bi-orientation of chromosomes during metaphase ."

reach
"Ablating UCHL3 significantly attenuates farrerol-mediated stimulation in HR and SCNT embryo development."
UCHL3 affects differentiation osteoblast C2C12 cells
| 1
UCHL3 activates differentiation osteoblast C2C12 cells. 1 / 1
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eidos
"UCH-L3 promotes the differentiation of osteoblast from C2C12 cells , while knockdown of Uch-l3 delays osteoblast differentiation [ 73 ] ."
UCHL3 affects cleave
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UCHL3 activates cleave. 1 / 1
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eidos
"It was reported that UCHL3 enables to cleave the C-terminus of NEDD8 , which is a ubiquitin-like protein that exerts the function of Ub to be conjugated to a lysine residue of the substrate ( 115 ) ."
UCHL3 affects chromosome alignment defects
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UCHL3 inhibits chromosome alignment defects. 1 / 1
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eidos
"Downregulation or inhibition of UCHL3 leads to chromosome alignment defects during metaphase ."
UCHL3 affects chromatin retention
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UCHL3 activates chromatin retention. 1 / 1
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eidos
"Taken together , these data suggest that UCHL3 could enhance Ku chromatin retention upon DSB induction by deubiquitylating Ku80 ."
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"Results of flow cytometry then showed that UCHL3 knockdown augmented cell apoptosis, along with increased cells arrested in the G0/G1 phase, decreased cells arrested in the S phase, and repressed cell division, while the restoration of UCHL3 caused opposite results (Fig. 3E, F)."
| 1

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"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 (XREF_FIG and XREF_FIG) and the yeast UCH Yuh1 XREF_BIBR, XREF_BIBR."
UCHL3 affects ZRANB2
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No evidence text available
UCHL3 affects ZNRD2
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No evidence text available
UCHL3 affects ZFYVE19
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No evidence text available
UCHL3 affects XRCC6
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No evidence text available
UCHL3 affects VSV-G
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Modified UCHL3 decreases the amount of VSV-G. 1 / 1
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"The result showed that UCHL3 overexpression decreased the levels of VSV-G to some extent, suggesting that UCHL3 promotes cellular antiviral ability under a basal IFN secreting environment."
UCHL3 affects UbVME
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UbVME binds UCHL3. 1 / 1
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"In contrast, when UbVME binds to UCH-L3, the S shaped loop (153-167) is reordered and helix-6 ' (142-152) forms and covers the active site bound to the C-terminus of ubiquitin [13]."
UCHL3 affects USP9X
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UCHL3 deubiquitinates USP9X. 1 / 1
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"Deubiquitination of FAM-H2B(117-127)-K120Ub by UCH-L3."
UCHL3 affects UCHL3 KO
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UCHL3 activates UCHL3 KO. 1 / 1
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"These data were in line with observed c-NHEJ activities being reduced in UCHL3 KO#2 cells, which was largely rescued by exogenously expressed FLAG-UCHL3 (Fig. 3C)."
UCHL3 affects UCH
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UCHL3 activates UCH. 1 / 1
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"This suggests that UCH functions associated with the egg cortex and oolemma may be at least partially restored by UCHL3 function, if opportunity for this is presented during oogenesis."
UCHL3 affects UCH Yuh1
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UCH Yuh1 binds UCHL3. 1 / 1
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"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 (XREF_FIG and XREF_FIG) and the yeast UCH Yuh1 XREF_BIBR, XREF_BIBR."
UCHL3 affects UBE2V1
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No evidence text available
UCHL3 affects UBE2L3
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No evidence text available
UCHL3 affects UBE2A
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No evidence text available
UCHL3 affects UBA52
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UCHL3 increases the amount of UBA52. 1 / 1
1 |

"Here we provide data suggesting that two of the four mammalian ubiquitin precursors, UBA52 and UBA80, are processed mostly post-translationally whereas the other two, UBB and UBC, probably undergo a combination of co- and post-translational processing. Using an unbiased biochemical approach we found that UCHL3, USP9X, USP7, USP5 and Otulin/Gumby/FAM105b are by far the most active DUBs acting on these precursors."
UCHL3 affects UBA1
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No evidence text available
UCHL3 affects TWIST1
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UCHL3 inhibits TWIST1. 1 / 1
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"We found that UCH-L3 knockdown in RWPE1 upregulates Snail, Slug, and Twist."
UCHL3 affects TTC9C
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No evidence text available
UCHL3 affects TSN
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No evidence text available
UCHL3 affects TRM1
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No evidence text available
UCHL3 affects TRIM54
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No evidence text available
UCHL3 affects TEAD
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UCHL3 increases the amount of TEAD. 1 / 1
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"Depletion of UCHL3 decreased the YAP protein level and the expression of YAP/TEAD target genes in ATC."
UCHL3 affects TAGLN2
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No evidence text available
UCHL3 affects TAGLN
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No evidence text available
UCHL3 affects Smad1.To
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UCHL3 activates Smad1.To. 1 / 1
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"These results of increased expression of differentiation markers and down-stream target genes suggest that Uch-L3 may enhance osteoblast differentiation by elevating the stability of Smad1.To determin[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects SUMO2
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sparser
"In contrast, UCH-L3 did not bind to sentrin-1, sentrin-2, or sentrin-3."
UCHL3 affects STUB1
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UCHL3 deubiquitinates STUB1. 1 / 1
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"No other tested DUB behaved in a similar manner : USP5 deubiquitinated Ub-CHIP in an unregulated manner, while UCHL-1 and UCHL-3 did not deubiquitinate Ub-CHIP under any conditions (XREF_FIG)."
UCHL3 affects STIP1
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No evidence text available
UCHL3 affects SSRP1
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No evidence text available
UCHL3 affects SPOP
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UCHL3 inhibits SPOP. 1 / 1
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"Interestingly, we found that the concomitant knockdown of SPOP with UCHL3 rescued the effect of UCHL3 knockdown on BRD4 protein (Fig. S4i), indicating that UCHL3 antagonizes SPOP to maintain BRD4 stability."
UCHL3 affects SOX2
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No evidence text available
UCHL3 affects SOX12
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UCHL3 activates SOX12. 1 / 1
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"UCHL3 promotes proliferation of colorectal cancer cells by regulating SOX12 via AKT and mTOR signaling pathway."
UCHL3 affects SOD2
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No evidence text available
UCHL3 affects SNRPD1
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No evidence text available
UCHL3 affects SNAI1
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UCHL3 inhibits SNAI1. 1 / 1
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"We found that UCH-L3 knockdown in RWPE1 upregulates Snail, Slug, and Twist."
UCHL3 affects SLC25A6
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No evidence text available
UCHL3 affects SLC25A13
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No evidence text available
UCHL3 affects SERBP1
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No evidence text available
UCHL3 affects SDHA
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No evidence text available
UCHL3 affects Rho-Ub-PA
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UCHL3 binds Rho-Ub-PA. 1 / 1
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"The activity of UCHL3 was also monitored using the fluorescent Ub-propargylamide (PA) DUB ABP Rho-Ub-PA.3 d Fluorescent bands corresponding to UCHL3 bound to Rho-Ub-PA were observed for both unlabeled UCHL3 and DTT treated Ub SS -UCHL3 complexes, but not for untreated Ub SS -UCHL3 complexes."
UCHL3 affects RUNX2
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UCHL3 decreases the amount of RUNX2. 1 / 1
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"Overexpression of SMAD1 significantly reversed the downregulated osteogenic gene expression (Alp, Runx2, Col1, and Ocn) and protein level of RUNX2 caused by UCHL3 inhibition in MS-BMDM-EXOs (Fig. 7d–f)."
UCHL3 affects RPN1
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No evidence text available
UCHL3 affects RPL10A
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No evidence text available
UCHL3 affects RIPK4
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UCHL3 binds RIPK4 and K469. 1 / 1
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"We elucidate the mechanism that UCHL3 interacts with and deubiquitinates RIPK4 at the K469 site, removing the K48-linked ubiquitin chain and thus enhancing RIPK4 stabilization."
UCHL3 affects RAB1B
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No evidence text available
UCHL3 affects Paralysis
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"The losses of UchL1 and UchL3 in mice lead to neurodegeneration and paralysis [45] ; thus, their up-regulation could be connected to the attempt to protect neurons from the disruption of neuronal func[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects PYCR1
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No evidence text available
UCHL3 affects PPP1CA
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No evidence text available
UCHL3 affects PPIB
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No evidence text available
UCHL3 affects POU5F1
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UCHL3 activates POU5F1. 1 / 1
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"Our results show that UCH-L3 overexpression induced the expression of stemness genes (OCT4, SOX2, ZEB2, and MMP2), and increased cell migration (Figs."
UCHL3 affects POGK
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No evidence text available
UCHL3 affects PGK1
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No evidence text available
UCHL3 affects PGD
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No evidence text available
UCHL3 affects PEBP1
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No evidence text available
UCHL3 affects PCMT1
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No evidence text available
UCHL3 affects P4HB
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No evidence text available
UCHL3 affects NSG2
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No evidence text available
UCHL3 affects NOP56
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No evidence text available
UCHL3 affects NONO
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No evidence text available
UCHL3 affects NFkappaB
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eidos
"Moreover , UCH-L3 was reported to activate NF-kappaB signaling by deubiquitinating and stabilizing the ubiquitin ligase TRAF2 ( Zhang et al ., 2020 ) ."
UCHL3 affects NFKBIA
| 1
UCHL3 increases the amount of NFKBIA. 1 / 1
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"The results showed that UCHL3 upregulated protein levels of IkappaBalpha."
UCHL3 affects MKI67
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No evidence text available
UCHL3 affects MDH2
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No evidence text available
UCHL3 affects MCL1
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UCHL3 deubiquitinates MCL1. 1 / 1
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"Cdu1, but not the control enzyme UCH-L3 strongly deubiquitinated Mcl-1 (XREF_FIG)."
UCHL3 affects MAPRE1
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No evidence text available
UCHL3 affects LMNB1
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No evidence text available
UCHL3 affects LDHA
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UCHL3 increases the amount of LDHA. 1 / 1
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"Further investigations revealed that LDHA expression was promoted by UCHL3 and could be reduced by shFOXM1, and that low expressed LDHA partly reversed the inhibition of aerobic glycolysis induced by overexpressed UCHL3."
UCHL3 affects LC3II
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UCHL3 inhibits LC3II. 1 / 1
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"However, UCHL3 knockdown significantly decreased NEDD8 protein expression and markedly increased the LC3II/LC3I ratio in the si-UCHL3 group compared with the NC group (all P<0.001; Fig. 5A), accompanied by notably decreased NEDD8 ubiquitination (Fig. 5B)."
UCHL3 affects LC3B
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UCHL3 decreases the amount of LC3B. 1 / 1
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"RT-qPCR analysis demonstrated no significant differences in UCHL3, NEDD8 and LC3B expression levels between the NC and si-NC groups (P>0.05; Fig. 4A and B); however, UCHL3 knockdown significantly decreased the expression levels of UCHL3 and NEDD8, and significantly increased LC3B expression in the si-UCHL3 group compared with the NC group (all P<0.001; Fig. 4A and B)."
UCHL3 affects Ku ubiquitylation
| 1
UCHL3 activates Ku ubiquitylation. 1 / 1
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eidos
"Lastly , we provide evidence showing that DNA damage-induced UCHL3 phosphorylation ( which is dependent on UCHL3 catalytic activity , ATM kinase and downstream NHEJ factors ) promotes destabilization of UCHL3 , suggesting that UCHL3 may dissociate from Ku80 to allow Ku ubiquitylation and subsequent removal from DNA ."
UCHL3 affects KRT6A
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No evidence text available
UCHL3 affects KRT18
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No evidence text available
UCHL3 affects KLF4
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UCHL3 increases the amount of KLF4. 1 / 1
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"As expected, overexpression of UCHL3 increased the expression of ABCG2, c-Myc and KLF4, genes associated with stemness, at the protein level in A549 and H1299 cells (Fig. 5b, c), while depletion of UCHL3 decreased their protein expression (Fig. 5d)."
UCHL3 affects KEAP1
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BAP1 binds UCHL3 and KEAP1. 1 / 1
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sparser
"To examine whether BAP1 and UCHL3 directly interacts with KEAP1, we cotransfected pcDNA3-FLAG-BAP1 or pcDNA3-FLAG-UCHL3 with pcDNA3-HA-KEAP1 in 293T cells and immunoprecipitated cell lysates with FLAG-agarose beads followed by Western blot analysis with HA-HRP antibody."
UCHL3 affects K469
| 1
UCHL3 binds RIPK4 and K469. 1 / 1
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"We elucidate the mechanism that UCHL3 interacts with and deubiquitinates RIPK4 at the K469 site, removing the K48-linked ubiquitin chain and thus enhancing RIPK4 stabilization."
UCHL3 affects Interferon
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"Given that we demonstrated that UCHL3 promoted activation of IFN signaling, we observed whether IFN-I could affect the interaction between UCHL3 and COPS5."
UCHL3 affects ISYNA1
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No evidence text available
UCHL3 affects ILF3
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No evidence text available
UCHL3 affects ILF2
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No evidence text available
UCHL3 affects IGF1R
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Modified UCHL3 leads to the phosphorylation of IGF1R. 1 / 1
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"Moreover, ectopic expression of wild-type UCH-L3 restored the phosphorylation of insulin and IGF-I receptor and adipocyte differentiation in Uchl3 (-/-) MEFs."
UCHL3 affects IFN-I receptors
| 1
UCHL3 activates IFN-I receptors. 1 / 1
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"This finding suggests that UCHL3 could target either TYK2 itself or IFN-I receptors (IFNAR1 and IFNAR2), which are the direct proteins upstream of TYK2 as well as the first signaling proteins in IFN-I[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UCHL3 affects HSD17B10
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No evidence text available
UCHL3 affects HNRNPL
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No evidence text available
UCHL3 affects HNRNPD
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No evidence text available
UCHL3 affects HINT1
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No evidence text available
UCHL3 affects HADHB
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No evidence text available
UCHL3 affects GPC1
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No evidence text available
UCHL3 affects GFP-β-catenin
| 1
UCHL3 binds GFP-β-catenin. 1 / 1
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"Our results demonstrate that there was an interaction between FLAG-UCH-L3 and GFP-β-catenin; however, we did not observe this interaction in cells transfected with the inactive mutant of UCH-L3 (UCH-L3 C95S) (Fig. 2I)."
UCHL3 affects G0/G1
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UCHL3 activates G0/G1. 1 / 1
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"While UCHL3 knockdown slightly reduced G0/G1 arrest, it significantly increased the number of cells in the sub-G0 phase, indicative of enhanced apoptotic cell death."
UCHL3 affects Fig. 3F-I
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UCHL3 activates Fig. 3F-I. 1 / 1
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"Knockdown of UCHL3 could promote the degradation of EEF1A1, and overexpression of UCHL3 increased the stability of EEF1A1 (Fig. 3F-I)."
UCHL3 affects FOS
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sparser
"Using ChIP-qPCR, we confirmed that this increased UCH-L3 gene expression is caused by direct binding of c-Fos to the UCH-L3 promoter ( xref )."
UCHL3 affects FDXR
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No evidence text available
UCHL3 affects FASN
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No evidence text available
UCHL3 affects EIF4A1
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No evidence text available
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"These results suggest that UCHL3 might physically interact with CTNNB1 to trigger activation of the Wnt signaling pathway.Since UCHL3 is a DUB [18], its deubiquitinating activity is crucial to its function."
UCHL3 affects DDX39B
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No evidence text available
UCHL3 affects DDX39A
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No evidence text available
UCHL3 affects DDX21
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No evidence text available
UCHL3 affects Collagen
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"Selective inhibition of UCHL3 by TCID significantly reversed collagen deposition and the increase in M2 macrophage markers induced by BLM injection."
UCHL3 affects CLTC
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No evidence text available
UCHL3 affects CEBPA
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No evidence text available
UCHL3 affects CDH1
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No evidence text available
UCHL3 affects CD44
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UCHL3 activates CD44. 1 / 1
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"At the molecular level, under IR exposure UCHL3 knockdown reduced the protein expression of well-known MES GSC markers (CD44, TAZ, etc.), and POLD4 overexpression restored the expression of these markers (Fig. 6H)."
UCHL3 affects CCT7
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No evidence text available
UCHL3 affects CCT6A
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No evidence text available
UCHL3 affects CCT4
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No evidence text available
UCHL3 affects CCNB1
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No evidence text available
UCHL3 affects BRCA2
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"Furthermore, the 3KR mutant enhanced the RAD51/BRCA2 interaction in UCHL3-deficient cells (Supplemental Fig. S7C)."
UCHL3 affects BMP
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UCHL3 activates BMP. 1 / 1
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eidos
"UCH-L3 can deubiquitinate and stabilize Smadl , and enhance BMP signaling ."
UCHL3 affects BAP1
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BAP1 binds UCHL3 and KEAP1. 1 / 1
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sparser
"To examine whether BAP1 and UCHL3 directly interacts with KEAP1, we cotransfected pcDNA3-FLAG-BAP1 or pcDNA3-FLAG-UCHL3 with pcDNA3-HA-KEAP1 in 293T cells and immunoprecipitated cell lysates with FLAG-agarose beads followed by Western blot analysis with HA-HRP antibody."
UCHL3 affects AURKA
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No evidence text available
UCHL3 affects ATP6V1F
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No evidence text available
UCHL3 affects ATM
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No evidence text available
UCHL3 affects ARHGDIA
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No evidence text available
UCHL3 affects ARHGAP35
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No evidence text available
UCHL3 affects ARHGAP11B
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No evidence text available
UCHL3 affects APP
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UCHL3 inhibits APP. 1 / 1
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"This may be mechanistically attributed to the fact that loss of UCHL1 and UCHL3 accelerates the degradation of APP, thus accounting for the overproduction of Aβ (Zhang, Cai, Zhang, Zhang, & Song, 2014) (Table 2).2.5 Therapeutic implications."
UCHL3 affects AMPK
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UCHL3 activates AMPK. 1 / 1
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"Genetic ablation of UCHL3 deubiquitinating enzyme in mice specifically potentiates AMPK activity in skeletal muscle without impact on AMPK expression level, indicative of the casual linkage between ubiquitination modification and AMPK activation [155]."
UCHL3 affects ALDH18A1
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No evidence text available
UCHL3 affects ACSL3
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No evidence text available
UCHL3 inhibitors affects UCHL3
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UCHL3 inhibitors inhibits UCHL3. 1 / 1
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"From the cancer point of view, UCHL3 is an attractive " druggable " target and previous efforts focused on developing UCHL3 inhibitors that specifically target UCHL3 in plasmodium falciparum parasite, offering promising selectivity due to structural differences compared to human UCHL3."
UCH Yuh1 affects UCHL3
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UCH Yuh1 binds UCHL3. 1 / 1
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"Binding of ubiquitin leads to apparent stabilization of the crossover loop, as it was seen straddling the active site in co-crystal structures of ubiquitin aldehyde bound to UCHL3 (XREF_FIG and XREF_FIG) and the yeast UCH Yuh1 XREF_BIBR, XREF_BIBR."
UBE2V1 affects UCHL3
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No evidence text available
UBE2L3 affects UCHL3
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No evidence text available
UBE2A affects UCHL3
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No evidence text available
UBA1 affects UCHL3
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No evidence text available
TTC9C affects UCHL3
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No evidence text available
TSN affects UCHL3
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No evidence text available
TRM1 affects UCHL3
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No evidence text available
TRIM54 affects UCHL3
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No evidence text available
TAGLN2 affects UCHL3
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No evidence text available
TAGLN affects UCHL3
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No evidence text available
Soot affects UCHL3
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Soot increases the amount of UCHL3. 1 / 1
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No evidence text available
Senkirkine affects UCHL3
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Senkirkine increases the amount of UCHL3. 1 / 1
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No evidence text available
SUMO2 affects UCHL3
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"In contrast, UCH-L3 did not bind to sentrin-1, sentrin-2, or sentrin-3."
STIP1 affects UCHL3
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No evidence text available
SSRP1 affects UCHL3
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No evidence text available
SOX2 affects UCHL3
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No evidence text available
SOD2 affects UCHL3
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No evidence text available
SNRPD1 affects UCHL3
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No evidence text available
SLPI affects UCHL3
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SLPI activates UCHL3. 1 / 1
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"The enhanced ALP activity may be caused by the deubiquitinating activity of Uch-L3 stabilizing Smad1."
SLC25A6 affects UCHL3
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No evidence text available
SLC25A13 affects UCHL3
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No evidence text available
SERBP1 affects UCHL3
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No evidence text available
SDHA affects UCHL3
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No evidence text available
Rho-Ub-PA affects UCHL3
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UCHL3 binds Rho-Ub-PA. 1 / 1
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"The activity of UCHL3 was also monitored using the fluorescent Ub-propargylamide (PA) DUB ABP Rho-Ub-PA.3 d Fluorescent bands corresponding to UCHL3 bound to Rho-Ub-PA were observed for both unlabeled UCHL3 and DTT treated Ub SS -UCHL3 complexes, but not for untreated Ub SS -UCHL3 complexes."
Radiation, Ionizing leads to the phosphorylation of UCHL3. 1 / 1
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"The phosphorylation of UCHL3 was also induced by IR treatment (data not shown)."
RPN1 affects UCHL3
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No evidence text available
RPL10A affects UCHL3
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No evidence text available
RIPK4 affects UCHL3
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UCHL3 binds RIPK4 and K469. 1 / 1
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"We elucidate the mechanism that UCHL3 interacts with and deubiquitinates RIPK4 at the K469 site, removing the K48-linked ubiquitin chain and thus enhancing RIPK4 stabilization."
RAB1B affects UCHL3
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No evidence text available
Proteasome affects UCHL3
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"In addition, several ubiquitin-proteasome-related proteins (ubiquitin-activating enzyme E1, ubiquitin C-terminal hydrolase UCHL3 and proteasome 26S regulatory subunit) were crotonylated and 2-hydroxyisobutyrylated in the phenotypically different T. gondii parasites (supplemental Data S21)."
PYCR1 affects UCHL3
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No evidence text available
PRKDC affects Ku
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UCHL3 binds Ku and PRKDC. 1 / 1
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sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
PR-619 affects UCHL3
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"Remarkably, only OTULIN binding was lost after treatment with the broad-spectrum DUB inhibitor PR-619, which also inhibits USP5 and UCHL3."
PPP1CA affects UCHL3
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No evidence text available
PPIB affects UCHL3
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No evidence text available
POGK affects UCHL3
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No evidence text available
PKA affects UCHL3
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PKA activates UCHL3. 1 / 1
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"UCH-L3, for example, is apparently stimulated by cAMP and PKA so as to enhance ENaC recycling."
PGK1 affects UCHL3
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No evidence text available
PGD affects UCHL3
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No evidence text available
PEBP1 affects UCHL3
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No evidence text available
PCMT1 affects UCHL3
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No evidence text available
P4HB affects UCHL3
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No evidence text available
NSG2 affects UCHL3
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No evidence text available
NOP56 affects UCHL3
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No evidence text available
NONO affects UCHL3
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No evidence text available

sparser
"As LDN can also inhibit UCHL3, we determined the UCHL3 protein level in the sham group and MI group 14 days after operation and found that no significant differences between the two groups (Supplemental Fig.  xref )."
Mice affects UCHL3
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Mice activates UCHL3. 1 / 1
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"The generation of UCHL3-overexpressing mice and studies with these mice on aging or tumorigenesis might be necessary before further testing the roles of farrerol in alleviating the syndromes of these age-related diseases, delaying the onset of aging, and preventing tumorigenesis."
MMV676603 affects UCHL3
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MMV676603 inhibits UCHL3. 1 / 1
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"Consequently, we can infer that MMV676603 inhibits PfUCHL3 activity by altering the key interaction of cysteine residue present in the active site pocket of the enzyme."
MKI67 affects UCHL3
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No evidence text available
MIR375 affects UCHL3
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MIR375 decreases the amount of UCHL3. 1 / 1
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No evidence text available
MDH2 affects UCHL3
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No evidence text available
MAPRE1 affects UCHL3
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No evidence text available
LMNB1 affects UCHL3
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No evidence text available
LDN57444 affects UCHL3
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LDN57444 inhibits UCHL3. 1 / 1
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"Despite being more sensitive to UCHL1 inhibition, LDN57444 also suppresses the activity of UCHL3 [34]."
LDN-57444 affects UCHL3
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LDN-57444 inhibits UCHL3. 1 / 1
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"Many DUB inhibitors, such as P5091, inhibits USP7, Sputin1, which inhibits USP10 and USP13, and LDN-57444, which inhibits UCHL1 and UCHL3 [167]."
LDN affects UCHL3
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LDN inhibits UCHL3. 1 / 1
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"As LDN can also inhibit UCHL3, we determined the UCHL3 protein level in the sham group and MI group 14days after operation and found that no significant differences between the two groups (Supplemental Fig. 1)."
Ku affects PRKDC, and UCHL3
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UCHL3 binds Ku and PRKDC. 1 / 1
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sparser
"These data thus suggested that UCHL3 interacts with Ku and/or DNA-PKcs in cells and could have a role at sites of DSBs in a catalytic-activity-dependent manner."
KRT6A affects UCHL3
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No evidence text available
KRT18 affects UCHL3
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No evidence text available
KEAP1 affects UCHL3
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BAP1 binds UCHL3 and KEAP1. 1 / 1
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sparser
"To examine whether BAP1 and UCHL3 directly interacts with KEAP1, we cotransfected pcDNA3-FLAG-BAP1 or pcDNA3-FLAG-UCHL3 with pcDNA3-HA-KEAP1 in 293T cells and immunoprecipitated cell lysates with FLAG-agarose beads followed by Western blot analysis with HA-HRP antibody."
K469 affects UCHL3
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UCHL3 binds RIPK4 and K469. 1 / 1
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"We elucidate the mechanism that UCHL3 interacts with and deubiquitinates RIPK4 at the K469 site, removing the K48-linked ubiquitin chain and thus enhancing RIPK4 stabilization."
Infections affects UCHL3
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"Collectively, this study unveiled that P. gingivalis infection up-regulated UCHL3 and stabilized its substrate protein GNG12 to activate the NF-κB signal pathway to promote colon cancer progression."
ISYNA1 affects UCHL3
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No evidence text available
INS affects UCHL3
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INS inhibits UCHL3. 1 / 1
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"The phosphorylation of insulin and IGF-I receptor, Akt, glycogen synthase kinase-3beta, and FoxO1 was decreased in Uchl3 (-/-) MEFs treated with insulin."
ILF3 affects UCHL3
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No evidence text available
ILF2 affects UCHL3
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No evidence text available
ICG 001 affects UCHL3
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ICG 001 decreases the amount of UCHL3. 1 / 1
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No evidence text available
HSD17B10 affects UCHL3
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No evidence text available
HNRNPL affects UCHL3
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No evidence text available
HNRNPD affects UCHL3
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No evidence text available
HINT1 affects UCHL3
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No evidence text available
HADHB affects UCHL3
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No evidence text available
GSK-J4 affects UCHL3
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GSK-J4 increases the amount of UCHL3. 1 / 1
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No evidence text available
GPC1 affects UCHL3
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No evidence text available
GNG12 affects UCHL3
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GNG12 inhibits UCHL3. 1 / 1
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"The protein level of GNG12 was significantly reduced when knock out UCHL3."
GFP-β-catenin affects UCHL3
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UCHL3 binds GFP-β-catenin. 1 / 1
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reach
"Our results demonstrate that there was an interaction between FLAG-UCH-L3 and GFP-β-catenin; however, we did not observe this interaction in cells transfected with the inactive mutant of UCH-L3 (UCH-L3 C95S) (Fig. 2I)."
FOXJ2 affects UCHL3
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FOXJ2 decreases the amount of UCHL3. 1 / 1
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No evidence text available
FDXR affects UCHL3
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No evidence text available
FASN affects UCHL3
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No evidence text available
Environmental Pollutants increases the amount of UCHL3. 1 / 1
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No evidence text available
EIF4A1 affects UCHL3
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No evidence text available
EGR1 affects UCHL3
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EGR1 decreases the amount of UCHL3. 1 / 1
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No evidence text available
DNA UCHL3 phosphorylation affects UCHL3
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DNA UCHL3 phosphorylation activates UCHL3. 1 / 1
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eidos
"Lastly , we provide evidence showing that DNA damage-induced UCHL3 phosphorylation ( which is dependent on UCHL3 catalytic activity , ATM kinase and downstream NHEJ factors ) promotes destabilization of UCHL3 , suggesting that UCHL3 may dissociate from Ku80 to allow Ku ubiquitylation and subsequent removal from DNA ."
DDX39B affects UCHL3
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No evidence text available
DDX39A affects UCHL3
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No evidence text available
DDX21 affects UCHL3
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No evidence text available
CLTC affects UCHL3
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No evidence text available
CEBPA affects UCHL3
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No evidence text available
CDH1 affects UCHL3
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No evidence text available
CCT7 affects UCHL3
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No evidence text available
CCT6A affects UCHL3
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No evidence text available
CCT4 affects UCHL3
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No evidence text available
CCNB1 affects UCHL3
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No evidence text available
BRCA2 affects UCHL3
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reach
"Furthermore, the 3KR mutant enhanced the RAD51/BRCA2 interaction in UCHL3-deficient cells (Supplemental Fig. S7C)."
BMP2 affects UCHL3
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BMP2 activates UCHL3. 1 / 1
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"Moreover, BMP2-induced stimulation further increased ALP activity in Uch-L3 over-expressing cells compared to control cells ( Fig. 3 B, lower panels)."
BAP1 affects UCHL3
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BAP1 binds UCHL3 and KEAP1. 1 / 1
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sparser
"To examine whether BAP1 and UCHL3 directly interacts with KEAP1, we cotransfected pcDNA3-FLAG-BAP1 or pcDNA3-FLAG-UCHL3 with pcDNA3-HA-KEAP1 in 293T cells and immunoprecipitated cell lysates with FLAG-agarose beads followed by Western blot analysis with HA-HRP antibody."
AURKA affects UCHL3
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No evidence text available
ATP6V1F affects UCHL3
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No evidence text available
ARHGDIA affects UCHL3
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1 |

No evidence text available
ARHGAP35 affects UCHL3
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No evidence text available
ARHGAP11B affects UCHL3
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No evidence text available
ALDH18A1 affects UCHL3
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No evidence text available
ACSL3 affects UCHL3
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No evidence text available
2-hydroxypropanoic acid decreases the amount of UCHL3. 1 / 1
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No evidence text available
2,6-dimethoxyphenol decreases the amount of UCHL3. 1 / 1
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No evidence text available
1 |

No evidence text available
1H-pyrazole affects UCHL3
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1H-pyrazole increases the amount of UCHL3. 1 / 1
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No evidence text available
15Delta-PGJ2 affects UCHL3
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15Delta-PGJ2 inhibits UCHL3. 1 / 1
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"Delta12-PGJ2 and 15Delta-PGJ2 are potent cyclopentone PGs and have been reported to inhibit UCH-L3 and UCH-L1 respectively [282,283]."
1,2-dichloroethane decreases the amount of UCHL3. 1 / 1
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No evidence text available