IndraLab

Statements


OTULIN affects RNF31
20 | 52 81
OTULIN binds RNF31.
20 | 44 81
20 | 40 47

sparser
"Thus, we wondered whether biotin-PD of OTULIN ABP can be used to monitor association of active substrate-bound OTULIN to HOIP, and potentially the entire LUBAC complex ( xref )."

sparser
"Consistent with the structure, phosphorylation of Tyr56 prevents interaction between OTULIN and HOIP xref , xref (Fig. xref , bottom)."

sparser
"The HOIP-OTULIN interaction can be disrupted by mutation of a critical residue (N102) in HOIP’s PUB domain ( xref )."

reach
"Phosphorylation dependent regulation of the interaction between OTULIN and HOIP offers an elegant additional layer of control for Met1 signalling."

sparser
"However, while the Y56D mutation decreased OTULIN interaction with HOIP, it enhanced OTULIN association with β-catenin (Fig.  xref and Supplementary Fig.  xref ), suggesting that OTULIN Tyr56 phosphorylation may promote OTULIN interaction with β-catenin."

reach
"Several studies have shown that OTULIN binds directly to the LUBAC subunit HOIP (Elliott etal, 2014; Schaeffer etal, 2014; Takiuchi etal, 2014)."

No evidence text available

reach
"OTULIN interacts with HOIP via its PUB interacting motif (PIM), which binds to the N-terminal PUB domain of HOIP."

reach
"This posed the possibility for a very similar OTULIN binding mode to the HOIP PUB domain in comparison to p97 and the PNGase PUB domain.To further investigate the interaction between HOIP, OTULIN and [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"LUBAC seems to be constantly active in cells, but the accumulation of Met1-Ub is continuously counterbalanced by OTULIN bound to HOIP."
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
RNF31 binds OTULIN and PUB domain. 2 / 2
| 2

reach
"Although HOIP is able to interact with the AAA ATPase p97 and valosin containing protein (VCP) [XREF_BIBR, XREF_BIBR], which has recently been implicated in the recruitment of LUBAC to misfolded Huntingtin species [XREF_BIBR], the HOIP PUB domain binds OTULIN with a 40-fold higher affinity compared to p97 and VCP and NMR based in vitro studies indicate a more stable interaction between HOIP and OTULIN compared to p97 and VCP [XREF_BIBR]."

reach
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
RNF31 binds OTULIN and Met55. 1 / 1
| 1

reach
"A fluorescently labeled OTULIN peptide in which Met55 was exchanged to Leu (as in p97) bound HOIP with near-identical affinity (370nM), showing that the small change in the first hydrophobic residue did not account for the difference (XREF_FIG J)."
| 1

sparser
"They speculated that because both DVL2 and HOIP binds to gumby via the N-terminus, and the interaction with HOIP requires the catalytic activity of gumby, the interactions may regulate overall complex activity, substrate selection or specific signaling pathway ( xref )."
RNF31 binds OTULIN and Met1. 1 / 1
| 1

reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
OTULIN ubiquitinates RNF31.
| 4
OTULIN leads to the ubiquitination of RNF31. 2 / 2
| 2

reach
"Supporting this, ectopic expression of an inactive OTULIN with a mutation in the PIM (Y56A) did not lead to HOIP autoubiquitination, whereas inactive OTULIN with an intact PIM led to extensive HOIP ubiquitination (XREF_FIG B)."

reach
"This suggested that one role of OTULIN in the LUBAC complex (XREF_FIG E) is to prevent the autoubiquitination of HOIP."
Phosphorylated OTULIN leads to the ubiquitination of RNF31. 1 / 1
| 1

reach
"Recent studies reported that phosphorylation of the deubiquitinase OTULIN negatively regulates its interaction with HOIP and thus promotes HOIP ubiquitination XREF_BIBR, XREF_BIBR."
Modified OTULIN leads to the ubiquitination of RNF31. 1 / 1
| 1

reach
"Supporting this, ectopic expression of an inactive OTULIN with a mutation in the PIM (Y56A) did not lead to HOIP autoubiquitination, whereas inactive OTULIN with an intact PIM led to extensive HOIP ubiquitination (XREF_FIG B)."
OTULIN inhibits RNF31.
| 1
OTULIN inhibits RNF31. 1 / 2
| 1

reach
"Strikingly, while mice lacking OTULIN in B or Tcells do not display overt inflammatory phenotypes, further analysis indicates that these OTULIN deficient cells have downregulated LUBAC components HOIP and SHARPIN, but not HOIL-1."
OTULIN deubiquitinates RNF31.
| 2
Modified OTULIN-C129A leads to the deubiquitination of RNF31 on M1. 1 / 1
| 1

reach
"We have previously shown that knockdown of OTULIN or overexpression of a catalytically inactive OTULIN C129A mutant (CA) lead to the autoubiquitination of HOIP with Met1 linked polyUb chains (XREF_FIG A, compare lanes 1 and 4; XREF_FIG B, compare lanes 1 and 2)."
OTULIN leads to the deubiquitination of RNF31. 1 / 1
| 1

reach
"This observation suggests that, under basal conditions, the binding of OTULIN prevents HOIP autoubiquitination."
OTULIN activates RNF31.
| 1
OTULIN activates RNF31. 1 / 1
| 1

reach
"OTULIN hyper-phosphorylation promotes necroptosis by locking ring finger protein 31 (RNF31, also known as HOIP) away from the cylindromatosis (CYLD) complex, resulting in altered receptor interacting serine/threonine kinase 1 (RIPK1) ubiquitination."
RNF31 affects OTULIN
20 | 46 81
RNF31 binds OTULIN.
20 | 44 81
20 | 40 47

sparser
"Thus, we wondered whether biotin-PD of OTULIN ABP can be used to monitor association of active substrate-bound OTULIN to HOIP, and potentially the entire LUBAC complex ( xref )."

sparser
"Consistent with the structure, phosphorylation of Tyr56 prevents interaction between OTULIN and HOIP xref , xref (Fig. xref , bottom)."

sparser
"The HOIP-OTULIN interaction can be disrupted by mutation of a critical residue (N102) in HOIP’s PUB domain ( xref )."

reach
"Phosphorylation dependent regulation of the interaction between OTULIN and HOIP offers an elegant additional layer of control for Met1 signalling."

sparser
"However, while the Y56D mutation decreased OTULIN interaction with HOIP, it enhanced OTULIN association with β-catenin (Fig.  xref and Supplementary Fig.  xref ), suggesting that OTULIN Tyr56 phosphorylation may promote OTULIN interaction with β-catenin."

reach
"Several studies have shown that OTULIN binds directly to the LUBAC subunit HOIP (Elliott etal, 2014; Schaeffer etal, 2014; Takiuchi etal, 2014)."

No evidence text available

reach
"OTULIN interacts with HOIP via its PUB interacting motif (PIM), which binds to the N-terminal PUB domain of HOIP."

reach
"This posed the possibility for a very similar OTULIN binding mode to the HOIP PUB domain in comparison to p97 and the PNGase PUB domain.To further investigate the interaction between HOIP, OTULIN and [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"LUBAC seems to be constantly active in cells, but the accumulation of Met1-Ub is continuously counterbalanced by OTULIN bound to HOIP."
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
RNF31 binds OTULIN and PUB domain. 2 / 2
| 2

reach
"Although HOIP is able to interact with the AAA ATPase p97 and valosin containing protein (VCP) [XREF_BIBR, XREF_BIBR], which has recently been implicated in the recruitment of LUBAC to misfolded Huntingtin species [XREF_BIBR], the HOIP PUB domain binds OTULIN with a 40-fold higher affinity compared to p97 and VCP and NMR based in vitro studies indicate a more stable interaction between HOIP and OTULIN compared to p97 and VCP [XREF_BIBR]."

reach
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
RNF31 binds OTULIN and Met55. 1 / 1
| 1

reach
"A fluorescently labeled OTULIN peptide in which Met55 was exchanged to Leu (as in p97) bound HOIP with near-identical affinity (370nM), showing that the small change in the first hydrophobic residue did not account for the difference (XREF_FIG J)."
| 1

sparser
"They speculated that because both DVL2 and HOIP binds to gumby via the N-terminus, and the interaction with HOIP requires the catalytic activity of gumby, the interactions may regulate overall complex activity, substrate selection or specific signaling pathway ( xref )."
RNF31 binds OTULIN and Met1. 1 / 1
| 1

reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
RNF31 ubiquitinates OTULIN.
| 1
RNF31 ubiquitinates OTULIN. 1 / 1
| 1

reach
"Otulin interacted with LUBAC after virus infection and the E3-ubiquitin ligase, HOIP, a component of LUBAC, ubiquitinated Otulin to trigger its proteasome-mediated degradation."
RNF31 inhibits OTULIN.
| 1
Mutated RNF31 inhibits OTULIN. 1 / 1
| 1

reach
"Similarly, mutation of HOIP residues necessary for OTULIN binding prevented OTULIN to exert its inhibitory action on the NF-kappaB pathway."
OTULIN is modified
| 1 72 5
OTULIN is phosphorylated.
| 1 63 5
OTULIN is phosphorylated on Y56. 10 / 43
| 41 2

sparser
"OTULIN Tyr-56 phosphorylation is counteracted by the activity of dual-specificity phosphatase 14 (DUSP14), which we identify as an OTULIN phosphatase that limits necroptosis."

sparser
"Interestingly, OTULIN is hyper-phosphorylated at Tyr56 during necroptosis, pointing towards a role of OTULIN Tyr56 phosphorylation in the regulation of necroptotic cell death [ xref ] (see below)."

sparser
"These data suggest that c-Abl is activated in response to genotoxic treatment, which may be responsible for Tyr56 phosphorylation of OTULIN."

sparser
"Consistently, DNA-PKcs knockdown remarkably reduced OTULIN Tyr56 phosphorylation in cells treated with genotoxic drugs (Fig.  xref )."

sparser
"Supporting this notion, the phosphorylation of Y56 in OTULIN is reported to increase after cells were induced to undergo necroptosis (by treatment with TNF + cycloheximide+ the pan-caspase inhibitor zVAD-fmk), and, the phosphatase DUSP14 was found to counteract the phosphorylation [59]."

sparser
"Consistently, ABL1 deletion in MDA-MB-231 cells suppressed OTULIN Tyr56 phosphorylation by Dox, CBP, and CPT-11 (Supplementary Fig.  xref )."

rlimsp
"The conserved Tyr56 of OTULIN makes critical contacts with the HOIP PUB domain, and its phosphorylation negatively regulates this interaction."

sparser
"Supporting this notion, the phosphorylation of Y56 in OTULIN is reported to increase after cells were induced to undergo necroptosis (by treatment with TNF  +  cycloheximide+ the pan-caspase inhibitor zVAD-fmk), and, the phosphatase DUSP14 was found to counteract the phosphorylation [ xref ]."

sparser
"Such phosphorylation of OTULIN Tyr56 would regulate the general association of OTULIN with LUBAC before their recruitment to the TNFR complexes."

sparser
"In parallel, the decreased OTULIN association with HOIP correlated with increased OTULIN phosphorylation at Tyr56 upon genotoxic treatment (Supplementary Fig.  xref )."
OTULIN is phosphorylated. 10 / 26
| 1 22 3

rlimsp
"Moreover, LUBAC-OTULIN complex formation is regulated by OTULIN phosphorylation in the PIM."

sparser
"OTULIN is phosphorylated during necroptosis, which is inhibited by the phosphatase DUSP14 to suppress cell death [ xref ]."

sparser
"Wnt3a treatment induced a decrease of β-catenin linear ubiquitination with much-delayed kinetics (Supplementary Fig.  xref ), which may be due to, at least in part, the lack of OTULIN phosphorylation."

sparser
"Thus, stimulation-associated tyrosine phosphorylation of OTULIN, including at Tyr56, cannot be responsible for OTULIN’s absence from SCs."

trips
"ABL1-dependent OTULIN phosphorylation promotes genotoxic Wnt/beta-catenin activation to enhance drug resistance in breast cancers."

sparser
"Notably, neither mutation of Tyr56 nor phosphorylation of OTULIN affects the catalytic activity of OTULIN in vitro [ xref , xref ], suggesting that Tyr56 phosphorylation mainly regulates the OTULIN–LUBAC interaction."

rlimsp
"Interestingly, OTULIN directly interacts with LUBAC, and this interaction is dynamic and can be regulated by OTULIN phosphorylation."

sparser
"Additionally, DNA damage-induced ABL1/c-Abl (ABL proto-oncogene 1) activation can promote the phosphorylation of OTULIN, which enhances its interaction with β-catenin and promotes the activation of Wnt/β-catenin signalling ( xref ) This mechanism plays a critical role in the triple-negative breast cancer (TNBC) progression, metastasis, and drug resistance to cancer treatments ( xref ; xref ; xref )."

sparser
"Accordingly, we observed a decrease in linear ubiquitination of β-catenin with comparable kinetics in cells treated with Dox, which correlated with the increase of OTULIN phosphorylation (Fig.  xref )."

sparser
"DNA damage-activated c-Abl is required for OTULIN phosphorylation."
OTULIN is ubiquitinated.
| 7
OTULIN is ubiquitinated. 7 / 7
| 7

sparser
"Noticeably, HBV-induced ubiquitination of exogenous OTULIN was confirmed by immunoblotting (Fig.  xref e)."

sparser
"Taken together, we report a novel regulatory mechanism by which TRIM32-mediated non-proteolytic ubiquitination of OTULIN impedes the access of OTULIN to the LUBAC complex and promotes NF-κB activation."

sparser
"To further address the relevance of K66 lysine site for the up-regulation of OTULIN’s ubiquitination, we compared ubiquitination of wild-type and K66R mutant of OTULIN upon HBV replication."

sparser
"Whereas, ubiquitination of OTULIN (K66) was significantly up-regulated, resulting in a decline in its protein level (Fig.  xref d)."

sparser
"Noticeably, HBV-induced ubiquitination of exogenous OTULIN was confirmed by immunoblotting (Fig. 9e)."

sparser
"SNX27-driven membrane localisation of OTULIN antagonises linear ubiquitination and NF-κB signalling activation."

sparser
"Whereas, ubiquitination of OTULIN (K66) was significantly up-regulated, resulting in a decline in its protein level (Fig. 9d)."
OTULIN is dephosphorylated.
| 2
OTULIN is dephosphorylated. 2 / 2
| 2

sparser
"Furthermore, while OTULIN and the HOIP PUB domain form a gel filtration‐stable complex in vitro , the majority of OTULIN in cell lysates does not coelute with LUBAC indicating that OTULIN may be phosphorylated in cells, and dephosphorylation of OTULIN in cell extracts led to coelution of OTULIN with LUBAC xref ."

sparser
"Although the responsible kinase(s) have not yet been identified, the dual specificity protein phosphatase 14 (DUSP14) has been reported to dephosphorylate OTULIN [ xref ]."
OTULIN affects CYLD
| 1 10 41
OTULIN binds CYLD.
| 8 41
| 6 14

reach
"The interaction of CYLD and OTULIN with HOIP synergistically suppresses LUBAC mediated linear polyubiquitination and NF-kappaB activation."

reach
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP."

sparser
"Furthermore, we identified the endogenous association of CYLD and OTULIN with LUBAC and the CBM complex by the immunoprecipitation with anti-MALT1 antibody ( xref )."

reach
"Suppression of LUBAC mediated linear ubiquitination by a specific interaction between LUBAC and the deubiquitinases CYLD and OTULIN."

sparser
"Intriguingly, HOIP’s relatively small PUB domain mediates the interaction with both CYLD and OTULIN and even short deletions in this domain abolished interaction with both factors ( xref B–S3D)."

sparser
"The above question was answered by the discovery that OTULIN and CYLD interact with LUBAC [ xref – xref ] ( xref ) and further consolidated the role of OTULIN as a bona fide regulator of Met1 signalling."

sparser
"Given that, LUBAC may form a putative complex which includes both OTULIN and CYLD bound to two HOIP molecules [ xref , xref ]."

reach
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD Selective Recruitment to SCs."

sparser
"In addition, a mutant version of HOIP that can neither bind to OTULIN nor CYLD is also less capable of inducing canonical Wnt signaling than wildtype HOIP xref ."

sparser
"Furthermore, LUBAC constitutively interacts with the deubiquitinating enzymes (DUBs) OTULIN and CYLD, which cleave linear ubiquitin chains generated by LUBAC."
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
| 4

sparser
"LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"Fig. 2: LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [32]."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [ xref ]."
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
| 1

sparser
"Since the PUB domain mediates the interaction with CYLD and OTULIN (Schaeffer et al , xref ; Takiuchi et al , xref ; Draber et al , xref ) and the ZF and NZF1 are required for HOIP binding to ubiquitin and NEMO (Haas et al , xref ; Ikeda et al , xref ; Fujita et al , xref ), we hypothesised that cleavage of HOIP may modulate TRAIL‐induced gene activation."
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
OTULIN binds CYLD and Met1. 1 / 1
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
OTULIN phosphorylates CYLD.
| 1
OTULIN phosphorylates CYLD. 1 / 1
| 1

reach
"Interestingly, recent findings also suggest that OTULIN deficiency results in spontaneous inhibitory phosphorylation of CYLD, thereby limiting CYLD activity, which can be rescued by LUBAC inhibition [XREF_BIBR]."
OTULIN dephosphorylates CYLD.
| 1
OTULIN leads to the dephosphorylation of CYLD on S418. 1 / 1
| 1

reach
"Another NF-κB inhibitory DUB, OTULIN inhibits Ser418 phosphorylation of CYLD through a LUBAC-dependent mechanism, which sustains its catalytic activity, while hyperphosphorylation of OUTLIN at Tyr56 regulates necroptosis by modulating RIPK1 ubiquitin dynamics [30]."
OTULIN activates CYLD.
| 1
OTULIN activates CYLD. 1 / 1
| 1

eidos
"Interestingly , recent findings also suggest that OTULIN deficiency results in spontaneous inhibitory phosphorylation of CYLD , thereby limiting CYLD activity , which can be rescued by LUBAC inhibition [ 47 ] ."
OTULIN affects NFkappaB
| 3 24 1
OTULIN inhibits NFkappaB.
| 2 16 1
| 2 14 1

reach
"Nevertheless, the DUB OTULIN is identified to restrain the formation of M1 linked Ub chains on the bacterial coat, then sequester the recruitment of NEMO and ultimately inhibit NF-kappaB pathway."

reach
"It appears that OTULIN overexpression antagonizes NF-kappaB activation by removing Met1-polyUb, whereas OTULIN C129A acts as a high-affinity UBD that competes with other Met1 specific UBDs required for NF-kappaB signaling in a similar manner to the one recently reported for the overexpression of the NEMO UBAN domain."

reach
"As both CYLD and OTULIN negatively regulate the NF-kappaB signalling cascade, the impact of these deubiquitinases in different LUBAC dependent signalling cascades may be context dependent."

reach
"To directly address if OTULIN antagonizes NF-kappaB activation by disassembling Met1-Ub, we devised a system whereby a noncleavable (Gly76 --> Ser76; GS) Met1-Ub was docked to XIAP via an IAP binding [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"How the overexpression of OTULIN attenuated NF-kappaB activation at the ubiquitination level was not explored in the current study."

reach
"Formation of the secondary, receptor-free cytoplasmic complex II is largely dependent on the activity of DUBs, specifically cylindromatosis (CYLD), A20 (also known as TNFAIP3) and ubiquitin thioesterase OTULIN, which destabilize complex I, abrogate NF-kappaB activation and release RIPK1 from complex I, which then forms the cytosolic complex II XREF_BIBR, XREF_BIBR."

reach
"CYLD and OTULIN were not inhibiting NF-kappaB activity per se because the DUBs did not appreciably inhibit NF-kappaB activity induced by an engineered non cleavable Met1-Ub4 protein targeted to inactive XIAP."

reach
"Increased OTULIN expression was required for EA to inhibit the NF-kappaB signalling pathway in focal cerebral ischaemia and reperfusion rats."

eidos
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non-structural protein 11 ( Nsp11 ) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 ( IRF3 ) - mediated type-I IFN production [ 22 ] ."

reach
"Overexpression of OTULIN inhibits TNF-alpha-induced NF-kappaB signaling by preventing the association of NEMO with polyubiquitinated RIP1 (Fiil et al., 2013; Keusekotten et al., 2013)."
OTULIN bound to Nsp11 inhibits NFkappaB. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
OTULIN-C129A inhibits NFkappaB. 1 / 1
| 1

reach
"Interestingly, overexpression of catalytically inactive OTULIN C129A also suppressed NF-kappaB signaling, in contrast to the Ub binding mutant OTULIN W96A, likely due to the ability of OTULIN C129A to compete with endogenous UBDs involved in NF-kappaB signaling, as it is the case for Ub binding domain in ABIN proteins and NEMO (UBAN)-based Met1 Ub sensors as well [XREF_BIBR, XREF_BIBR]."
OTULIN activates NFkappaB.
| 1 7
| 1 7

reach
"Small interfering RNA (siRNA) against OTULIN resulted in anincrease in LUBAC induced NF-kappaB activation in HEK 293ET and U2OS cells and also in T-REx 293 cells stably expressing a doxycycline inducible OTULIN targeting microRNA (miRNA) (XREF_FIG A and XREF_FIG A-S6C)."

reach
"In contrast, depletion of OTULIN enhanced the activation of NF-kappaB in response to NOD2 stimulation 64."

reach
"Primary fibroblast cultures from this patient reveal a surprising phenotype of deregulated M1 linked Ub signalling, whereby OTULIN loss coincides with LUBAC downregulation to prevent intrinsic NF-kappaB activation."

reach
"Immune cell specific depletion of OTULIN leads to cell-type specific effects : constitutive active NF-kappaB signalling and overproduction of cytokines in myeloid cells, and down-regulation of linear ubiquitination signal in the B and T cells based on LUBAC degradation [XREF_BIBR]."

reach
"Mutation of Tyr56 in the OTULIN PIM domain abrogates HOIP binding, resulting in reduced ability of OTULIN to inhibit NF-kappaB activity and reduced cell death signaling [XREF_BIBR, XREF_BIBR, XREF_BIBR, XREF_BIBR]."

reach
"The recently identified linear-chain-specific DUB OTULIN has been implicated in the regulation of TNF induced NF-kappaB activation and cell death XREF_BIBR."

reach
"In contrast, OTULIN deficient myeloid cells are hyper-inflammatory and spontaneously activate NF-kappaB and secrete TNF."

eidos
"OTULIN targets Met1-linked Ub and counteracts LUBAC auto-ubiquitination , thereby promoting NF-kappaB activation and MAPK signaling ."
OTULIN decreases the amount of NFkappaB.
| 1
Modified OTULIN decreases the amount of NFkappaB. 1 / 1
| 1

reach
"Overexpression of OTULIN, and hence removal of all Met1 linkages from cells, blocks NF-kappaB translocation to the nucleus upon TNF stimulation and reduces NF-kappaB target gene expression."
SNX27 affects OTULIN
| 12 19
SNX27 binds OTULIN.
| 10 19
| 8 19

sparser
"An X-ray crystal structure of OTULIN-SNX27 PDZ revealed a second interface in addition to the canonical PDZ-PDZbm interaction."

sparser
"Therefore, we concluded that the interaction between OTULIN and SNX27 has mutual effects on each other."
| PMC

sparser
"To further study the interaction between OTULIN and SNX27, we constituted the interaction assay using constructs containing deletions and mutations of SNX27 and OTULIN."
| PMC

sparser
"Here, we show that the interaction between SNX27 and Met1 linkage is mediated by OTULIN, consistent with the recent reported interaction of SNX27 and OTULIN [ xref ]."
| PMC

sparser
"By solving the structure of the OTU domain in complex with the PDZ domain, we demonstrate that a second interface contributes to the selective, high affinity interaction of OTULIN and SNX27."

reach
"By solving the structure of the OTU domain in complex with the PDZ domain, we demonstrate that a second interface contributes to the selective, high affinity interaction of OTULIN and SNX27."

sparser
"SNX27 interacts with OTULIN (OTU Deubiquitinase With Linear Linkage Specificity) that specifically hydrolyzes methionine1 (Met1)-linked ubiquitin chains."

reach
"Therefore, we concluded that the interaction between OTULIN and SNX27 has mutual effects on each other."
| PMC

sparser
"Mechanistic studies showed that, upon TNFα stimulation, OTULIN-SNX27 is localised to membrane-associated TNF receptor complex, where OTULIN deubiquitinates the linear polyubiquitin chain that formed by the LUBAC complex."

sparser
"Therefore, we speculated that when cells are undergoing TNFα stimulation, the interaction between OTULIN and SNX27 will be transiently increased, which subsequently decreases the loading of other SNX27 dependent cargo proteins [ xref ]."
| PMC
SNX27 binds OTULIN and Met1. 1 / 1
| 1

reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
SNX27 binds OTULIN and PDZ domain. 1 / 1
| 1

reach
"Immunoprecipitation in 293T cells and in vitro pull-down assays using bacterial recombinant proteins indicated that the direct interaction between SNX27 and OTULIN was mediated by the PDZ domain of SNX27 and the last 6 amino acids of OTULIN (Additional file 2 : Figure S2E-G), respectively."
| PMC
SNX27 activates OTULIN.
| 2
SNX27 activates OTULIN. 1 / 1
| 1

reach
"In this study, we combined immunoprecipitation, immunostaining and cell fractionation assay to show that SNX27 potentiates the localization of OTULIN to TNF receptor complex."
| PMC
SNX27 bound to OTULIN activates OTULIN. 1 / 1
| 1

reach
"In summary, we showed that the interaction of SNX27 and OTULIN potentiates the translocation of OTULIN to TNFalpha receptor complex."
| PMC
OTULIN affects SNX27
| 12 19
OTULIN binds SNX27.
| 10 19
| 8 19

sparser
"An X-ray crystal structure of OTULIN-SNX27 PDZ revealed a second interface in addition to the canonical PDZ-PDZbm interaction."

sparser
"Therefore, we concluded that the interaction between OTULIN and SNX27 has mutual effects on each other."
| PMC

sparser
"To further study the interaction between OTULIN and SNX27, we constituted the interaction assay using constructs containing deletions and mutations of SNX27 and OTULIN."
| PMC

sparser
"Here, we show that the interaction between SNX27 and Met1 linkage is mediated by OTULIN, consistent with the recent reported interaction of SNX27 and OTULIN [ xref ]."
| PMC

sparser
"By solving the structure of the OTU domain in complex with the PDZ domain, we demonstrate that a second interface contributes to the selective, high affinity interaction of OTULIN and SNX27."

reach
"By solving the structure of the OTU domain in complex with the PDZ domain, we demonstrate that a second interface contributes to the selective, high affinity interaction of OTULIN and SNX27."

sparser
"SNX27 interacts with OTULIN (OTU Deubiquitinase With Linear Linkage Specificity) that specifically hydrolyzes methionine1 (Met1)-linked ubiquitin chains."

reach
"Therefore, we concluded that the interaction between OTULIN and SNX27 has mutual effects on each other."
| PMC

sparser
"Mechanistic studies showed that, upon TNFα stimulation, OTULIN-SNX27 is localised to membrane-associated TNF receptor complex, where OTULIN deubiquitinates the linear polyubiquitin chain that formed by the LUBAC complex."

sparser
"Therefore, we speculated that when cells are undergoing TNFα stimulation, the interaction between OTULIN and SNX27 will be transiently increased, which subsequently decreases the loading of other SNX27 dependent cargo proteins [ xref ]."
| PMC
SNX27 binds OTULIN and Met1. 1 / 1
| 1

reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
SNX27 binds OTULIN and PDZ domain. 1 / 1
| 1

reach
"Immunoprecipitation in 293T cells and in vitro pull-down assays using bacterial recombinant proteins indicated that the direct interaction between SNX27 and OTULIN was mediated by the PDZ domain of SNX27 and the last 6 amino acids of OTULIN (Additional file 2 : Figure S2E-G), respectively."
| PMC
OTULIN inhibits SNX27.
| 2
OTULIN inhibits SNX27. 2 / 2
| 2

reach
"Previous study has shown that OTULIN negatively regulates SNX27 dependent cargo loading [XREF_BIBR]."
| PMC

reach
"However, via association, OTULIN antagonizes SNX27 dependent cargo loading, binding of SNX27 to the VPS26A-retromer subunit and endosome-to-plasma membrane trafficking."
OTULIN affects Ubiquitin
| 11 11
OTULIN binds Ubiquitin.
| 1 11

sparser
"On the molecular level, we find binding of OTULIN to linear ubiquitin to be compromised by both variants; however, protein stability and catalytic activity is most affected by OTULIN variant p."

sparser
"Using our OTULIN ABP we demonstrate that OTULIN is constitutively active in cells, and that substrate-bound OTULIN associates with the linear ubiquitin chain assembly complex (LUBAC)."

sparser
"Disease-associated mutations reside in the OTU domain and are predicted to affect binding of OTULIN to linear Ub chains."

sparser
"The interaction between Y91 and M86, which is lost in the mutant M86I, might, thus, also play a role in binding of OTULIN to linear ubiquitin (Fig  xref )."

sparser
"It is likely, therefore, that the W167S replacement affects the orientation of the helical arm's S1 site, made up of L259, A262, and R263 toward the I44 patch of distal ubiquitin, thereby indirectly interfering with OTULIN’s binding (K M ) to linear ubiquitin (Fig  xref )."

sparser
"In‐depth biochemical analysis revealed how both OTULIN variants contribute to diminished OTULIN protein expression and defective hydrolysis of linear ubiquitin linkages observed in patient‐derived cells: whereas binding of OTULIN to linear ubiquitin was compromised by both variants, protein stability and catalytic activity was most affected by the OTULIN variant p."

sparser
"Pull-downs demonstrate that substrate-bound OTULIN associates with the linear ubiquitin chain assembly complex (LUBAC)."

sparser
"In summary, we find binding of OTULIN to linear ubiquitin to be compromised by both variants; however, protein stability and catalytic activity is most affected by the OTULIN variant p."

sparser
"Leu272 is located in a helix of the catalytic OTU domain that forms part of the binding pocket for M1-linked diUb, and mutation to Pro would be predicted to disrupt OTULIN Ub binding ( xref F)."

sparser
"Collectively, this structural analysis suggests that both substitutions, M86I and W167S, may interfere with binding of OTULIN to linear ubiquitin (Fig  xref )."
Ubiquitin binds OTULIN and Ser65. 1 / 1
| 1

reach
"Ser20 and 57 in the proximal Ub are close to the distal Ub bound to OTULIN, while Ser65 on the proximal Ub is solvent exposed (XREF_SUPPLEMENTARY A)."
OTULIN inhibits Ubiquitin.
| 5
| 4

reach
"As expected, OTULIN treatment abolished the detection of ubiquitin chains (Fig XREF_FIG A, top panel), confirming that linear ubiquitination occurs in adult flies."

reach
"In this study, the ‘deubiquitinase’ OTULIN was shown to restrict bacterial growth by controlling the density of linear ubiquitin chains on the bacterial surface, which serves as a platform for NF-κB activation.Finally, lattice light-sheet microscopy (LLSM) is a newly developed super-resolution microscopy method, which capitalises on previous microscopy techniques (light-sheet microscopy, SIM) and provides gentle and fast super-resolution in a volumetric sample13."

reach
"In this study, the ‘deubiquitinase’ OTULIN was shown to restrict bacterial growth by controlling the density of linear ubiquitin chains on the bacterial surface, which serves as a platform for NF-κB activation.Finally, lattice light-sheet microscopy (LLSM) is a newly developed super-resolution microscopy method, which capitalises on previous microscopy techniques (light-sheet microscopy, SIM) and provides gentle and fast super-resolution in a volumetric sample ."

reach
"OTULIN prevents such promiscuity by requiring a properly positioned Ub residue, Glu16, for catalytic efficiency."
Mutated OTULIN inhibits Ubiquitin. 1 / 1
| 1

reach
"Consequently, mutant OTULIN proteins failed to prevent the accumulation of linear Ub chains on target substrates IKK and NEMO, RIPK1, TNFR1, and ASC."
OTULIN activates Ubiquitin.
| 3
| 3

reach
"Here, we discuss how OTULIN contributes to cell fate control and highlight novel ubiquitin dependent and -independent functions."

reach
"OTULIN haploinsufficiency causes an accumulation of linear ubiquitin in dermal fibroblasts, but TNF-receptor NF-kappaB-signaling remains intact."

reach
"OTULIN haploinsufficiency causes an accumulation of linear ubiquitin in dermal fibroblasts, but TNF-receptor NF-κB-signaling remains intact."
OTULIN increases the amount of Ubiquitin.
| 1
OTULIN increases the amount of Ubiquitin. 1 / 1
| 1

reach
"As HOIP associated OTULIN was not recruited to SCs, we next studied whether OTULIN deficiency may increase levels of linear ubiquitin found in the cytosol."
OTULIN decreases the amount of Ubiquitin.
| 1
OTULIN bound to Nsp11 decreases the amount of Ubiquitin. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
Ubiquitin affects OTULIN
| 6 11
Ubiquitin binds OTULIN.
| 1 11

sparser
"On the molecular level, we find binding of OTULIN to linear ubiquitin to be compromised by both variants; however, protein stability and catalytic activity is most affected by OTULIN variant p."

sparser
"Using our OTULIN ABP we demonstrate that OTULIN is constitutively active in cells, and that substrate-bound OTULIN associates with the linear ubiquitin chain assembly complex (LUBAC)."

sparser
"Disease-associated mutations reside in the OTU domain and are predicted to affect binding of OTULIN to linear Ub chains."

sparser
"The interaction between Y91 and M86, which is lost in the mutant M86I, might, thus, also play a role in binding of OTULIN to linear ubiquitin (Fig  xref )."

sparser
"It is likely, therefore, that the W167S replacement affects the orientation of the helical arm's S1 site, made up of L259, A262, and R263 toward the I44 patch of distal ubiquitin, thereby indirectly interfering with OTULIN’s binding (K M ) to linear ubiquitin (Fig  xref )."

sparser
"In‐depth biochemical analysis revealed how both OTULIN variants contribute to diminished OTULIN protein expression and defective hydrolysis of linear ubiquitin linkages observed in patient‐derived cells: whereas binding of OTULIN to linear ubiquitin was compromised by both variants, protein stability and catalytic activity was most affected by the OTULIN variant p."

sparser
"Pull-downs demonstrate that substrate-bound OTULIN associates with the linear ubiquitin chain assembly complex (LUBAC)."

sparser
"In summary, we find binding of OTULIN to linear ubiquitin to be compromised by both variants; however, protein stability and catalytic activity is most affected by the OTULIN variant p."

sparser
"Leu272 is located in a helix of the catalytic OTU domain that forms part of the binding pocket for M1-linked diUb, and mutation to Pro would be predicted to disrupt OTULIN Ub binding ( xref F)."

sparser
"Collectively, this structural analysis suggests that both substitutions, M86I and W167S, may interfere with binding of OTULIN to linear ubiquitin (Fig  xref )."
Ubiquitin binds OTULIN and Ser65. 1 / 1
| 1

reach
"Ser20 and 57 in the proximal Ub are close to the distal Ub bound to OTULIN, while Ser65 on the proximal Ub is solvent exposed (XREF_SUPPLEMENTARY A)."
Ubiquitin inhibits OTULIN.
| 4
| 2

reach
"Deficiency of the linear ubiquitin chain-specific deubiquitinase, OTULIN, causes OTULIN-related autoinflammatory syndrome (ORAS), a systemic inflammatory pathology affecting multiple organs including the skin."

reach
"Consistently, mutation of the proximal ubiquitin Glu16 to alanine (E16A) reduces the catalytic efficiency (k cat) of OTULIN by 240-fold without significantly affecting binding of the substrate (similar K D and only 3-fold reduction in K M) [XREF_BIBR]."
Ubiquitin-G76A inhibits OTULIN. 1 / 1
| 1

reach
"As expected, Ub G76A -Ub did not label or inhibit OTULIN and USP5, but it also induced the dose dependent assembly of Ub chains that could be detected with anti-M1-polyUb antibody (XREF_FIG G)."
Mutated Ubiquitin inhibits OTULIN. 1 / 1
| 1

reach
"Consistently, mutation of ubiquitin Glu16 to Ala reduced the catalytic efficiency of OTULIN by 240-fold without significantly affecting binding of the substrate to the enzyme (similar K d and only a threefold reduction in K m) 29."
Ubiquitin activates OTULIN.
| 1
| 1

reach
"Hence, we reveal a mechanism of substrate assisted catalysis in which Glu16 of the proximal Ub activates the catalytic triad of OTULIN by both restricting the movement of the catalytic His339 and introducing a negative charge, presumably for the proper coordination of Asn341 for catalysis."
Trim14 affects OTULIN
| 18
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
| 4

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"Mapping analysis showed that the N‐terminus of OTULIN bound to the PUB domain in HOIP."

sparser
"Full-length OTULIN bound the PUB domain with a similar affinity as the OTULIN stretch from aa 45 to aa 73 (Kd FL  = 294 nM versus Kd 45–73  = 440 nM), suggesting that this region contains the complete[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"While the interaction of the Rnf31 PUB domain with the p97 C-terminal peptide is only ∼50 μM, the DUB OTULIN binds this PUB domain with a dissociation constant ∼300 nM, indicating that this mode of binding can facilitate a wide range of specificities [ xref ]."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
RNF31 affects CYLD
| 18
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
OTULIN affects Trim14
| 18
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
| 4

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"Mapping analysis showed that the N‐terminus of OTULIN bound to the PUB domain in HOIP."

sparser
"Full-length OTULIN bound the PUB domain with a similar affinity as the OTULIN stretch from aa 45 to aa 73 (Kd FL  = 294 nM versus Kd 45–73  = 440 nM), suggesting that this region contains the complete[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"While the interaction of the Rnf31 PUB domain with the p97 C-terminal peptide is only ∼50 μM, the DUB OTULIN binds this PUB domain with a dissociation constant ∼300 nM, indicating that this mode of binding can facilitate a wide range of specificities [ xref ]."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
CYLD affects RNF31
| 18
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
| 10

reach
"Finally, the numerous other upregulated cytokines in LysMCre-OtulinLacZ and flox mice, which includes anti-inflammatory IL-10, will generate a delicate balance of responses, enabling these mice to deal with inflammation induced by loss of OTULIN."

reach
"Deletion of Otulin in Mouse Immune Cells Drives Acute Systemic Inflammation."

reach
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice."

reach
"In an acute model, induced loss of OTULIN in immune cells leads to multi-organ inflammation and deterioration of animals within a few days; this can be ameliorated by anti-TNF, but not by neutralization of other upregulated cytokines."

reach
"Interestingly, conditional deletion of otulin in myeloid cells, but not T or B cells, causes systemic inflammation, neutrophil infiltration, and multi-organ inflammation in mouse models."

reach
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase Mediated Hepatocyte Apoptosis."

reach
"OTULIN overexpression attenuated microglia activation in LPS induced brain inflammation."

reach
"While loss of OTULIN leads to excessive inflammation through accumulation of linear-Ub, germline deficiencies of LUBAC also result in autoinflammation."

reach
"Recent work suggested that OTULIN can limit inflammation by deubiquitinating LUBAC [9]."

reach
"OTULIN limits cell death and inflammation by deubiquitinating LUBAC."
| 5

reach
"Inactivation of OTULIN by expression of OTULIN C129A in adult mice has recently been reported to cause systemic inflammation, which can be prevented by combined ablation of cell death pathways by deletion of caspase-8, RIPK3 and RIPK1 (Heger etal, 2018)."

reach
"To exclude the possibility that the effect of OTULIN on reducing microglia activation is an indirect outcome of its neuroprotective role [XREF_BIBR], a commonly used LPS induced pure brain inflammation model without brain cell death served as a positive control."

reach
"OTULIN deficiency triggers metabolic alterations, apoptosis, and inflammation in the liver."

reach
"These cellular and molecular mediators are indicative of an inappropriate and damaging inflammatory response mediated by OTULIN deficient myeloid cells."

reach
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice."
CYLD affects OTULIN, and RNF31
| 15
| 15

sparser
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1 Ubiquitination at the TNF-RSC and Enhances TNF-Induced Gene Activation."

sparser
"HOIP interacts with both CYLD and OTULIN even in unstimulated cells."

sparser
"Mutually Exclusive Binding of CYLD and OTULIN to HOIP Causes CYLD-Selective Recruitment to SCs."

sparser
"The HOIP missense mutation affects the conserved PUB domain of HOIP ( xref ), which has recently been shown to be important for the interaction of HOIP with OTULIN and CYLD, two deubiquitinases ( xref ; xref ; xref )."

sparser
"Additionally, we used a HOIP-PUB domain point mutant (N102A), which abolishes the interaction of both OTULIN and CYLD with HOIP ( xref , xref )."

sparser
"In line with recent reports ( xref , xref , xref , xref , xref ), our data showed that prior to stimulation, both CYLD and OTULIN interacted with HOIP ( xref B–S1D)."

sparser
"In contrast to these findings, Draber et al. demonstrated that, although both OTULIN and CYLD interact with HOIP under basal conditions, OTULIN is absent from RSCs [ xref ] (Fig.  xref ) and that knock-out of OTULIN resulted in an increase of Met1-linked poly-Ub chains in the cytosol but not at TNFR1 or NOD2 RSCs [ xref ]."

sparser
"The explanation was provided by our discovery that HOIP cannot simultaneously bind OTULIN and CYLD as both require HOIP-Asn102 for binding."

sparser
"An interaction between HOIP and OTULIN or CYLD at first inspection would seem contradictory, as a futile energy-consuming cycle would exist."

sparser
"This suggested that steric hindrance may prevent simultaneous interaction of HOIP with CYLD and OTULIN."
SPATA2 affects OTULIN
| 1 11
SPATA2 binds OTULIN.
| 11
| 4

sparser
"OTULIN and SPATA2 bind the same PIM pocket in HOIP, which excludes the simultaneous binding of both proteins to LUBAC and gives rise to two distinct LUBAC–DUB complexes; LUBAC–OTULIN and LUBAC–SPATA2–CYLD [11, 31] (Fig. 2)."

sparser
"OTULIN-released LUBAC interacts with SPATA2 and is recruited to the TNF-R1sc, facilitating SPATA2-CYLD interaction."

sparser
"OTULIN and SPATA2 bind the same PIM pocket in HOIP, which excludes the simultaneous binding of both proteins to LUBAC and gives rise to two distinct LUBAC–DUB complexes; LUBAC–OTULIN and LUBAC–SPATA2–CYLD [ xref , xref ] (Fig.  xref )."

sparser
"Because the N102A mutation in the PUB domain of HOIP abolishes the interaction with OTULIN and SPATA2, we assumed that, similar to OTULIN, SPATA2 might also contain a PUB-domain interacting motif (PIM)."
| 4

sparser
"LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"Fig. 2: LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [32]."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [ xref ]."
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
SPATA2 inhibits OTULIN.
| 1
| 1

reach
"SPATA2 restricts OTULIN-dependent LUBAC activity independently of CYLD."
Trim14 affects RNF31
| 12
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
TNF affects OTULIN
1 | 10 1
TNF activates OTULIN.
| 7
TNF activates OTULIN. 7 / 7
| 7

reach
"Importantly, similar effects were obtained when the OTULIN deficient patients were treated with the TNF antagonist infliximab."

reach
"XREF_FIG E, cholera toxin strongly inhibited SNX27 mediated TNFalpha stimulation mediated OTULIN membrane localization."
| PMC

reach
"XREF_FIG A, the interaction between endogenous SNX27 and OTULIN was upregulated when cells were treated with TNFalpha for 5min."
| PMC

reach
"However, whether this is due to changes in E3 ligase activity (cIAP and LUBAC) or deubiquitinating enzymes (CYLD, tumor necrosis factor alpha-induced protein 3 (Tnfip3 or A20), and OTU deubiquitinase with linear linkage specificity (OTULIN)) is unclear."

reach
"However, although A20 and CYLD are induced by TNFalpha for the establishment of a negative feedback loop, neither OTULIN (XREF_FIG F) nor LUBAC are induced by TNFalpha, again suggesting that they function as a differentially regulated signaling module."

reach
"Significantly, the fact that the here described OTULIN related autoinflammatory syndrome and ORAS can be treated with TNF neutralizing antibodies suggests a potential therapeutic strategy to treat conditions caused by excessive M1 linked polyUb signaling."

reach
"The deubiquitinase OTULIN removes methionine 1 (M1)-linked polyubiquitin signals conjugated by the linear ubiquitin chain assembly complex (LUBAC) and is critical for preventing TNF driven inflammation in OTULIN related autoinflammatory syndrome (ORAS)."
TNF binds OTULIN.
1 | 1 1
1 | 1

reach
"Interestingly, after caspase mediated cleavage during apoptosis, the N-terminal fragment of HOIP also binds to the DUBs OTULIN and CYLD, which are down-regulators of LUBAC mediated ubiquitination, providing a further regulatory feedback loop."

No evidence text available
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
TNF inhibits OTULIN.
| 2
TNF inhibits OTULIN. 2 / 2
| 2

reach
"XREF_FIG E, cholera toxin strongly inhibited SNX27 mediated TNFalpha stimulation mediated OTULIN membrane localization."
| PMC

reach
"Similarly, overexpression of OTULIN reduces the clonogenic survival of cells treated with TNF XREF_BIBR."
RNF31 affects Trim14
| 12
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
OTULIN affects SPATA2
| 11
| 4

sparser
"OTULIN and SPATA2 bind the same PIM pocket in HOIP, which excludes the simultaneous binding of both proteins to LUBAC and gives rise to two distinct LUBAC–DUB complexes; LUBAC–OTULIN and LUBAC–SPATA2–CYLD [11, 31] (Fig. 2)."

sparser
"OTULIN-released LUBAC interacts with SPATA2 and is recruited to the TNF-R1sc, facilitating SPATA2-CYLD interaction."

sparser
"OTULIN and SPATA2 bind the same PIM pocket in HOIP, which excludes the simultaneous binding of both proteins to LUBAC and gives rise to two distinct LUBAC–DUB complexes; LUBAC–OTULIN and LUBAC–SPATA2–CYLD [ xref , xref ] (Fig.  xref )."

sparser
"Because the N102A mutation in the PUB domain of HOIP abolishes the interaction with OTULIN and SPATA2, we assumed that, similar to OTULIN, SPATA2 might also contain a PUB-domain interacting motif (PIM)."
| 4

sparser
"LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"Fig. 2: LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [32]."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [ xref ]."
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
UBD affects OTULIN
| 4 6
UBD binds OTULIN.
| 1 5
| 1 5

reach
"OTULIN binding to di-ubiquitin is characterized by a substrate-assisted catalysis step, in which the catalytic triad is only activated upon tight substrate binding (Keusekotten et al., 2013)."

sparser
"OTULIN binds Met1-linked diubiquitin with a K D of 150 nM, 100-fold higher affinity than Lys63-linked diubiquitin [ xref ]."

sparser
"Nevertheless, simply mutating the catalytic Cys in DUBs to Ala or Ser, as used for A20 xref , xref , is problematic, as this potentially converts the DUB domain to a high‐affinity UBD; OTULIN C129A binds Met1‐diubiquitin with nanomolar affinity, and interferes with NF‐κB signalling in artificial ways xref ."

sparser
"The structure of OTULIN bound to Met1-linked diubiquitin revealed extensive contacts between both distal and proximal ubiquitin moieties ( xref )."

sparser
"Indeed, OTULIN C129A binds Met1‐linked diubiquitin with 150 n m affinity, a 100‐fold greater affinity compared to topologically similar Lys63‐linked diubiquitin."

sparser
"OTULIN binding to di-ubiquitin is characterized by a substrate-assisted catalysis step, in which the catalytic triad is only activated upon tight substrate binding ( xref )."
UBD activates OTULIN.
| 2 1
UBD activates OTULIN. 3 / 3
| 2 1

sparser
"More strikingly, we found that OTULIN was directly activated by the bound diubiquitin."

reach
"More strikingly, we found that OTULIN was directly activated by the bound diubiquitin."

reach
"More strikingly, the structure also revealed that OTULIN is directly activated by the bound diubiquitin : placement of the proximal ubiquitin inserts the ubiquitin side chain of Glu16 directly into the active site of OTULIN, thereby pushing His339 into an active conformation and additionally co-ordinating the third catalytic residue, Asn341 (XREF_FIG)."
UBD inhibits OTULIN.
| 1
UBD inhibits OTULIN. 1 / 1
| 1

reach
"We found that carbamylation of linear diubiquitin blocks the activity of the anti-inflammatory deubiquitinase OTULIN."
OTULIN affects cell death
| 1 10
OTULIN activates cell death.
| 6
| 6

reach
"OTULIN overexpression or knockdown affects NF-kappaB responses to LUBAC, TNFalpha, and poly (I : C) and sensitizes cells to TNFalpha induced cell death."

reach
"The recently identified linear-chain-specific DUB OTULIN has been implicated in the regulation of TNF induced NF-kappaB activation and cell death XREF_BIBR."

reach
"During necroptosis, OTULIN is hyper-phosphorylated at Tyr 56, which modulates RIPK1 ubiquitin dynamics and promotes cell death."

reach
"Moreover, stable overexpression of OTULIN sensitized cells to TNFalpha induced cell death (XREF_FIG D and 6E), consistent with observations in cpdm mice that lack Sharpin or in humans with mutations in HOIL1."

reach
"OTULIN deficiency sensitises fibroblasts and monocytes to TNF induced cell death."

reach
"Surprisingly, overexpression of wild-type OTULIN abrogates the RIPK1-NEMO interaction and sensitizes to TNFalpha induced cell death, as well [XREF_BIBR]."
OTULIN inhibits cell death.
| 1 4
| 1 4

eidos
"Furthermore , Otulin inactivation reduces TNFR1 complex formation and increases cell death [ 83 ] ."

reach
"However, both overexpression and knockdown of OTULIN sensitized cells to TNFalpha induced cell death, suggesting more complex roles for Met1 linkages in balancing signaling pathways downstream of the TNFalpha receptor."

reach
"OTULIN limits cell death and inflammation by deubiquitinating LUBAC."

reach
"This suggests that loss of OTULIN does not cause aberrant cell death in immune cells (XREF_FIG E) and that the high levels of cytokines and chemokines present activate the entire immune cell population."

reach
"Mutation of Tyr56 in the OTULIN PIM domain abrogates HOIP binding, resulting in reduced ability of OTULIN to inhibit NF-kappaB activity and reduced cell death signaling [XREF_BIBR, XREF_BIBR, XREF_BIBR, XREF_BIBR]."
OTULIN affects TNF
1 | 4 2
OTULIN inhibits TNF.
| 3
OTULIN inhibits TNF. 3 / 6
| 3

reach
"Consistent with the finding that OTULIN overexpression was shown to delay NF-kappaB activation of TNF, whereas its knockdown was reported to increase the output of NF-kappaB signaling [XREF_BIBR, XREF_BIBR, XREF_BIBR], we found that OTULIN overexpression strongly depressed NF-kappaB activity and reduced the release of TNF-alpha, IL-1beta, and IL-6, which are the primary pro inflammatory cytokines that aggravate brain injury."

reach
"Concomitant Loss of OTULIN and CYLD Interaction with HOIP Increases M1Ubiquitination at the TNF-RSC and Enhances TNF Induced Gene Activation."

reach
"OTULIN overexpression significantly suppressed the production of TNF-alpha, IL-1beta, and IL-6 in PM and N9 cells via inhibiting the nuclear translocation of NF-kappaB p65 but had no effect on the pro inflammatory cytokines secreted by un activated microglia, which directly indicates that OTULIN suppressed microglia mediated neuroinflammation by inhibiting the NF-kappaB pathway and attenuated the release of inflammatory mediators under the in vitro ischemic condition."
OTULIN binds TNF.
1 | 1 1
1 | 1

reach
"Interestingly, after caspase mediated cleavage during apoptosis, the N-terminal fragment of HOIP also binds to the DUBs OTULIN and CYLD, which are down-regulators of LUBAC mediated ubiquitination, providing a further regulatory feedback loop."

No evidence text available
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
OTULIN ubiquitinates TNF.
| 1
OTULIN ubiquitinates TNF. 1 / 1
| 1

sparser
"Thus, abolishing HOIP’s capacity to interact with both CYLD and OTULIN enhances M1 ubiquitination in the TNF-RSC and, consequently, TNF-induced gene activation."
OTULIN affects RNF31, and Trim14
| 10
| 10

sparser
"While OTULIN binds HOIP directly through its PUB-interacting motif (PIM) ( xref , xref ), CYLD is recruited via a PIM in the bridging factor SPATA2 ( xref , xref , xref , xref )."

sparser
"The specificity of the HOIP PUB-OTULIN interaction stems from a binding preference of the HOIP PIM pocket for internal PIMs in contrast to the C-terminal PIM of p97/VCP [ xref , xref ]."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif to remove the Met-1 Ub on IKKγ [ xref – xref ], thus leading to the inactivation of IKK."

sparser
"OTULIN interacts with the N-terminal PUB domain of HOIP via an evolutionarily conserved PUB-interacting motif ( xref )."

sparser
"To test that OTULIN indeed directly binds to the Ψ–Y pocket of HOIP, we introduced various point mutations and detected a decrease in OTULIN binding to the HOIP PUB domain with both single mutants Y82[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Now, we show that OTULIN binds via a conserved PUB-interacting motif (PIM) to the PUB domain of the LUBAC component HOIP."

sparser
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide: N-glycanase/UBA- or UBX-containing proteins) domain [ xref , xref ]."

sparser
"OTULIN contains a PIM sequence and is bound directly by HOIP-PUB [ xref , xref ]."

sparser
"OTULIN binds to the PUB domain of HOIP and loss of HOIP-OTULIN interaction reduces OTULIN’s capacity to restrict LUBAC-induced immune responses [ xref ]."

sparser
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
OTULIN affects LUBAC
| 1 9
OTULIN binds LUBAC.
| 1 5
OTULIN binds LUBAC. 6 / 6
| 1 5

reach
"Upon TNFalpha stimulation, the TNF-R1 was enriched in SHARPIN IPs (XREF_FIG E), further supporting the idea that a LUBAC and OTULIN complex is formed and that this complex may translocate to the TNF receptor signaling complex (TNF-RSC)."

reach
"We show that OTULIN binds LUBAC and that overexpression of OTULIN prevents TNFalpha induced NEMO association with ubiquitinated RIPK1."

reach
"Indeed, as discussed below, LUBAC and OTULIN form a complex and their activities are likely to be interdependent."

trips
"OTULIN allies with LUBAC to govern angiogenesis by editing ALK1 linear polyubiquitin."

reach
"Moreover, immunoprecipitation of SHARPIN copurified HOIP, HOIL-1 and OTULIN, and OTULIN interacted with HOIP in proteomic experiments, indicating that OTULIN may associate with HOIP and/or LUBAC."

reach
"Moreover, LUBAC and OTULIN complex formation is regulated by OTULIN phosphorylation in the PIM."
OTULIN inhibits LUBAC.
| 4
OTULIN inhibits LUBAC. 4 / 4
| 4

reach
"Ectopic expression of OTULIN in HEK293 and HeLa cells inhibits LUBAC dependent NF-kappaB signaling, suggesting that OTULIN is a negative regulator of this signaling pathway 18, 20."

reach
"OTULIN antagonizes LUBAC signaling by specifically hydrolyzing Met1 linked polyubiquitin."

reach
"Moreover, a linear ubiquitin specific DUB, OTULIN, antagonizes LUBAC function in the xenophagy of Salmonella [XREF_BIBR]."

reach
"LUBAC function is restricted by OTULIN through its docking to the PUB domain of HOIP."
OTULIN affects GZMM
| 10
OTULIN inhibits GZMM.
| 4
OTULIN inhibits GZMM. 4 / 4
| 4

reach
"Conversely, OTULIN overexpression suppressed Met1-Ub of RIPK2 in a manner dependent on its catalytic activity."

reach
"To directly address if OTULIN antagonizes NF-kappaB activation by disassembling Met1-Ub, we devised a system whereby a noncleavable (Gly76 --> Ser76; GS) Met1-Ub was docked to XIAP via an IAP binding [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"Contrary to this, OTULIN functions to restrict the accumulation of Met1-Ub at basal conditions and early during signaling, and OTULIN expression is not induced by stimulation of NF-kappaB activity."

reach
"In contrast, our data suggest that OTULIN functions to restrict Met1-Ub accumulation and regulate early signaling processes."
OTULIN activates GZMM.
| 4
OTULIN activates GZMM. 3 / 3
| 3

reach
"Accordingly, Met1-Ub on RIPK2 was largely inhibited by overexpressing OTULIN and was increased by OTULIN depletion."

reach
"Nonetheless, OTULIN overexpression strongly impairs coprecipitation of ubiquitinated RIPK1 with NEMO in TNF treated cells, whereas small interfering RNA mediated depletion of OTULIN increases the early accumulation of Met1-Ub on RIPK1 XREF_BIBR."

reach
"Activation of the MAP kinases p38 and JNK1/2 was only slightly increased by OTULIN depletion, consistent with the notion that OTULIN disassembles Met1-Ub primarily involved in IKK activation."
OTULIN-C129A activates GZMM. 1 / 1
| 1

reach
"Accordingly, ectopic expression of inactive OTULIN (C129A), but not inactive CYLD (C601A), caused extensive accumulation of Met1-Ub on HOIP (XREF_FIG H and XREF_SUPPLEMENTARY F)."
OTULIN decreases the amount of GZMM.
| 2
Modified OTULIN decreases the amount of GZMM. 2 / 2
| 2

reach
"Accordingly, the amount of Met1-Ub on RIPK2 was decreased by overexpression of OTULIN and strongly enhanced in OTULIN depleted cells."

reach
"Conversely, overexpression of OTULIN decreased the level of Met1-Ub in the TNF-RSC and reduced the apparent molecular weight of ubiquitinated TNFR1."
ABL1 affects OTULIN
| 2 8
ABL1 phosphorylates OTULIN on Y56. 4 / 4
| 4

sparser
"Consistently, phosphorylation of OTULIN at Tyr56 by c-Abl was also significantly increased in a time-dependent manner upon Dox treatment (Fig.  xref and Supplementary Fig.  xref )."

sparser
"C-Abl phosphorylates OTULIN at tyrosine 56, thereby disrupting its relationship with LUBAC."

sparser
"Upon genotoxic treatment, DNA damage apical kinase DNA-PK could activate c-Abl which subsequently phosphorylates OTULIN at Tyr56."

sparser
"All these results indicate that, upon DNA damage, DNA-PK-activated c-Abl phosphorylates OTULIN at Tyr56, which promotes OTULIN association with β-catenin, resulting in β-catenin stabilization and Wnt/β-catenin signaling activation."
ABL1 phosphorylates OTULIN. 4 / 4
| 4

sparser
"The c-Abl-dependent OTULIN phosphorylation in response to genotoxic treatments relies on DNA-PK but not ATM or ATR."

sparser
"ABL1-dependent OTULIN phosphorylation promotes genotoxic Wnt/β-catenin activation to enhance drug resistance in breast cancers."

sparser
"The potential role of c-Abl-dependent OTULIN phosphorylation in regulating genotoxic NF-κB signaling warrant further investigation."

sparser
"We showed that ABL1 (ABL proto-oncogene 1)-dependent phosphorylation of OTULIN (OTU deubiquitinase with linear linkage specificity) upon DNA damage drives β-catenin activation which promotes drug resistance in triple-negative breast cancer."
ABL1 phosphorylates OTULIN on tyrosine. 1 / 1
| 1

reach
"C-Abl phosphorylates OTULIN at tyrosine 56, thereby disrupting its relationship with LUBAC."
ABL1 phosphorylates OTULIN at position 56. 1 / 1
| 1

reach
"C-Abl phosphorylates OTULIN at tyrosine 56, thereby disrupting its relationship with LUBAC."
OTULIN affects RIPK1
| 7
OTULIN inhibits RIPK1.
| 4
OTULIN inhibits RIPK1. 2 / 4
| 2

reach
"Formation of the secondary, receptor-free cytoplasmic complex II is largely dependent on the activity of DUBs, specifically cylindromatosis (CYLD), A20 (also known as TNFAIP3) and ubiquitin thioesterase OTULIN, which destabilize complex I, abrogate NF-kappaB activation and release RIPK1 from complex I, which then forms the cytosolic complex II XREF_BIBR, XREF_BIBR."

reach
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice."
Mutated OTULIN inhibits RIPK1. 1 / 1
| 1

reach
"Mutant OTULIN proteins can not restrict the accumulation of Met1 Ub chains on target substrates IKK and NEMO, RIPK1, and ASC."
OTULIN inhibits ubiquitinated RIPK1. 1 / 1
| 1

reach
"Nonetheless, OTULIN overexpression strongly impairs coprecipitation of ubiquitinated RIPK1 with NEMO in TNF treated cells, whereas small interfering RNA mediated depletion of OTULIN increases the early accumulation of Met1-Ub on RIPK1 XREF_BIBR."
OTULIN ubiquitinates RIPK1.
| 1
OTULIN ubiquitinates RIPK1. 1 / 1
| 1

reach
"Mechanistically, OTULIN overexpression abrogated association between NEMO and polyubiquitinated RIPK1 29, a key target of ubiquitination in this pathway that is also modified with multiple chain types including Lys63 and Met1 linkages 22."
OTULIN deubiquitinates RIPK1.
| 1
OTULIN leads to the deubiquitination of RIPK1. 1 / 1
| 1

reach
"OTULIN reduced ubiquitination of both, caspase-8 and RIPK1, identifying RIPK1 and caspase-8 as linearly ubiquitinated proteins (Fig XREF_FIG A)."
OTULIN activates RIPK1.
| 1
OTULIN activates RIPK1. 1 / 1
| 1

reach
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase Mediated Hepatocyte Apoptosis."
VCP affects OTULIN
2 | 3 3
2 | 3 1

reach
"K99R mutation had similar marginal effects on OTULIN or p97 interaction (XREF_FIG D)."

No evidence text available

No evidence text available

reach
"Therefore, the unique combination of amino acids of HOIP Lys99, Asn101, and Arg106 would specifically expand its PUB domain to interact with internal PIMs, as opposed to the PUB domains of UBXD1 or PN[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"Based on the structural data showing a tight fit of the central tyrosines and the ITC measurements of the phosphorylated OTULIN-peptide, the phosphorylation of these tyrosines inhibits the interaction[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

reach
"This posed the possibility for a very similar OTULIN binding mode to the HOIP PUB domain in comparison to p97 and the PNGase PUB domain.To further investigate the interaction between HOIP, OTULIN and [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTULIN affects CTNNB1
| 1 7
OTULIN binds CTNNB1.
| 7
| 7

sparser
"Interestingly, OTULIN-Y56D mutant overexpression induced more robust Wnt/β-catenin activation (Fig.  xref ), which may be a consequence of the increased interaction between β-catenin and OTULIN-Y56D mutant (Fig.  xref )."

sparser
"In contrast, the interaction of OTULIN with β-catenin was substantially increased upon Dox treatment, which may be responsible for decreased linear ubiquitination and stabilization of β-catenin in cells exposed to genotoxic treatments (Supplementary Fig.  xref , Fig.  xref , lanes 1 and 2)."

sparser
"Further characterization of domains required for the interaction between OTULIN and β-catenin revealed that the armadillo-repeats of β-catenin and the N-terminal region of OTULIN are required for their interaction (Fig.  xref and Supplementary Fig.  xref )."

sparser
"However, while the Y56D mutation decreased OTULIN interaction with HOIP, it enhanced OTULIN association with β-catenin (Fig.  xref and Supplementary Fig.  xref ), suggesting that OTULIN Tyr56 phosphorylation may promote OTULIN interaction with β-catenin."

sparser
"We speculate that the genotoxic stress-induced OTULIN phosphorylation at Tyr56 may not only increase the interaction between OTULIN and β-catenin but also enhance the LUBAC autoubiquitination and self-inhibition through dissociation from HOIP xref , which render OTULIN highly efficient in promoting β-catenin deubiquitination and stabilization upon DNA damage."

sparser
"Inhibiting either c-Abl or DNA-PK with imatinib or Nu-7441 blocked Dox-enhanced OTULIN interaction with β-catenin while stabilizing OTULIN/HOIP interaction (Fig.  xref )."

sparser
"We found genotoxic treatment enhanced the interaction between OTULIN and β-catenin."
OTULIN increases the amount of CTNNB1.
| 1
OTULIN increases the amount of CTNNB1. 1 / 1
| 1

reach
"Mechanically, OTULIN from M2-Exos could activate the Wnt/β-catenin signaling by increasing the protein level of β-catenin via inhibiting its ubiquitination and trigger the expression of angiogenesis-related genes that are reported to be the downstream targets of Wnt/β-catenin signaling."
UBC affects OTULIN
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

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No evidence text available

No evidence text available
OTULIN affects UBD
| 1 5
| 1 5

reach
"OTULIN binding to di-ubiquitin is characterized by a substrate-assisted catalysis step, in which the catalytic triad is only activated upon tight substrate binding (Keusekotten et al., 2013)."

sparser
"OTULIN binds Met1-linked diubiquitin with a K D of 150 nM, 100-fold higher affinity than Lys63-linked diubiquitin [ xref ]."

sparser
"Nevertheless, simply mutating the catalytic Cys in DUBs to Ala or Ser, as used for A20 xref , xref , is problematic, as this potentially converts the DUB domain to a high‐affinity UBD; OTULIN C129A binds Met1‐diubiquitin with nanomolar affinity, and interferes with NF‐κB signalling in artificial ways xref ."

sparser
"The structure of OTULIN bound to Met1-linked diubiquitin revealed extensive contacts between both distal and proximal ubiquitin moieties ( xref )."

sparser
"Indeed, OTULIN C129A binds Met1‐linked diubiquitin with 150 n m affinity, a 100‐fold greater affinity compared to topologically similar Lys63‐linked diubiquitin."

sparser
"OTULIN binding to di-ubiquitin is characterized by a substrate-assisted catalysis step, in which the catalytic triad is only activated upon tight substrate binding ( xref )."
OTULIN affects UBC
7 |
7 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available

No evidence text available
OTULIN affects STAT1
| 6 1
OTULIN ubiquitinates STAT1.
| 4
OTULIN ubiquitinates STAT1. 3 / 3
| 3

reach
"When OTULIN was knocked down, IFN-I was no longer able to induce downregulation of STAT1 linear ubiquitination (Fig. 5f), suggesting that IFN-I-mediated removal of STAT1 linear ubiquitination is dependent on OTULIN.Given that IFN-I signaling utilized OTULIN to remove linear ubiquitination of STAT1, we speculated that overexpression of OTULIN could promote the binding of STAT1 to IFNAR2 in IFN-I signaling."

reach
"OTULIN knockout inhibited downregulation of STAT1 linear ubiquitination induced by IFN-I, which further inhibited the binding of STAT1 to IFNAR2 and subsequent STAT1 activation."

reach
"Mutation of Lys511 and Lys652 of STAT1 abolished OTULIN-mediated regulation of STAT1 linear ubiquitination (Fig. 5d), suggesting that OTULIN could be a deubiquitinase that removes STAT1 linear ubiquitination."
OTULIN ubiquitinates ubiquitinated STAT1. 1 / 1
| 1

reach
"Taken together, these findings demonstrated that IFN-I signaling utilizes OTULIN to downregulate linear ubiquitination of STAT1 for activation of IFN-I antiviral signaling."
OTULIN binds STAT1.
| 1 1
| 1 1

sparser
"Interestingly, IFNɑ treatment significantly promoted the binding of OTULIN to STAT1 in a time-dependent manner (Fig.  xref and Supplementary Fig.  xref )."

reach
"Interestingly, IFNɑ treatment significantly promoted the binding of OTULIN to STAT1 in a time-dependent manner (Fig. 5e and Supplementary Fig. 5g, h)."
OTULIN deubiquitinates STAT1.
| 1
OTULIN leads to the deubiquitination of STAT1. 1 / 1
| 1

reach
"Knockdown of OTULIN upregulated linear ubiquitination of cellular STAT1 (Fig. 5c)."
CTNNB1 affects OTULIN
| 7
| 7

sparser
"Interestingly, OTULIN-Y56D mutant overexpression induced more robust Wnt/β-catenin activation (Fig.  xref ), which may be a consequence of the increased interaction between β-catenin and OTULIN-Y56D mutant (Fig.  xref )."

sparser
"In contrast, the interaction of OTULIN with β-catenin was substantially increased upon Dox treatment, which may be responsible for decreased linear ubiquitination and stabilization of β-catenin in cells exposed to genotoxic treatments (Supplementary Fig.  xref , Fig.  xref , lanes 1 and 2)."

sparser
"Further characterization of domains required for the interaction between OTULIN and β-catenin revealed that the armadillo-repeats of β-catenin and the N-terminal region of OTULIN are required for their interaction (Fig.  xref and Supplementary Fig.  xref )."

sparser
"However, while the Y56D mutation decreased OTULIN interaction with HOIP, it enhanced OTULIN association with β-catenin (Fig.  xref and Supplementary Fig.  xref ), suggesting that OTULIN Tyr56 phosphorylation may promote OTULIN interaction with β-catenin."

sparser
"We speculate that the genotoxic stress-induced OTULIN phosphorylation at Tyr56 may not only increase the interaction between OTULIN and β-catenin but also enhance the LUBAC autoubiquitination and self-inhibition through dissociation from HOIP xref , which render OTULIN highly efficient in promoting β-catenin deubiquitination and stabilization upon DNA damage."

sparser
"Inhibiting either c-Abl or DNA-PK with imatinib or Nu-7441 blocked Dox-enhanced OTULIN interaction with β-catenin while stabilizing OTULIN/HOIP interaction (Fig.  xref )."

sparser
"We found genotoxic treatment enhanced the interaction between OTULIN and β-catenin."
OTULIN affects SHARPIN
4 | 1 1
OTULIN binds SHARPIN.
4 | 1

No evidence text available

No evidence text available

No evidence text available

No evidence text available
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
OTULIN inhibits SHARPIN.
| 1
| 1

reach
"Strikingly, while mice lacking OTULIN in B or Tcells do not display overt inflammatory phenotypes, further analysis indicates that these OTULIN deficient cells have downregulated LUBAC components HOIP and SHARPIN, but not HOIL-1."
OTULIN affects RBCK1
4 | 1
OTULIN binds RBCK1.
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
OTULIN inhibits RBCK1.
| 1
OTULIN inhibits RBCK1. 1 / 1
| 1

reach
"Strikingly, while mice lacking OTULIN in B or Tcells do not display overt inflammatory phenotypes, further analysis indicates that these OTULIN deficient cells have downregulated LUBAC components HOIP and SHARPIN, but not HOIL-1."
OTULIN affects IKBKG
1 | 1 3
OTULIN inhibits IKBKG.
| 1 3
OTULIN inhibits IKBKG. 3 / 4
| 1 2

reach
"Accordingly, depletion of OTULIN enhanced the recruitment of NEMO to Met1 ubiquitinated RIPK2 in response to L18-MDP, whereas NEMO recruitment was inhibited by OTULIN overexpression."

eidos
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non-structural protein 11 ( Nsp11 ) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 ( IRF3 ) - mediated type-I IFN production [ 22 ] ."

reach
"Overexpression of OTULIN inhibits TNF-alpha-induced NF-kappaB signaling by preventing the association of NEMO with polyubiquitinated RIP1 (Fiil et al., 2013; Keusekotten et al., 2013)."
Mutated OTULIN inhibits IKBKG. 1 / 1
| 1

reach
"Mutant OTULIN proteins can not restrict the accumulation of Met1 Ub chains on target substrates IKK and NEMO, RIPK1, and ASC."
OTULIN deubiquitinates IKBKG.
1 |
OTULIN deubiquitinates IKBKG. 1 / 1
1 |

"Deubiquitination effects of swine OTULIN were identified. The interaction between porcine OTULIN and PRRSV Nsp11 is dependent on the OTU domain. PRRSV Nsp11 recruits OTULIN through a nonenzymatic combination to promote removal of linear ubiquitination targeting NEMO, resulting in a superimposed effect that inhibits the production of type I IFNs."
OTULIN affects Glu-Ala
| 6
OTULIN inhibits Glu-Ala.
| 3
| 3

reach
"Consistent with our hypothesis, OTULIN silencing reversed the inhibitory effect of EA on the secretion of pro inflammatory factors in the ischaemic cerebral cortex."

reach
"EA also enhanced microglial OTULIN expression in the present study, and OTULIN silencing effectively reversed the inhibitory effect of EA on microglial activation, as shown by the increased mean Iba-1 immunofluorescence intensity, which strongly indicates that enhanced OTULIN expression is necessary for EA to alleviate microglial activation."

reach
"OTULIN silencing reversed EA suppression of microglial activation after focal cerebral ischaemia and reperfusion injury."
OTULIN activates Glu-Ala.
| 3
| 3

reach
"Notably, the silencing of OTULIN obviously reversed the neuroprotective effects of EA, which indicated that OTULIN was involved in the anti-preventive role of EA following acute ischemic stroke."

reach
"OTULIN knockdown reversed the neuroprotective effects of EA against ischaemic brain injury."

reach
"OTULIN silencing obviously reversed EA neuroprotection, which was demonstrated by worsened neurobehavioural performance, cerebral infarct volume and neuronal injury."
LUBAC affects OTULIN
| 1 5
OTULIN binds LUBAC. 6 / 6
| 1 5

reach
"Upon TNFalpha stimulation, the TNF-R1 was enriched in SHARPIN IPs (XREF_FIG E), further supporting the idea that a LUBAC and OTULIN complex is formed and that this complex may translocate to the TNF receptor signaling complex (TNF-RSC)."

reach
"We show that OTULIN binds LUBAC and that overexpression of OTULIN prevents TNFalpha induced NEMO association with ubiquitinated RIPK1."

reach
"Indeed, as discussed below, LUBAC and OTULIN form a complex and their activities are likely to be interdependent."

trips
"OTULIN allies with LUBAC to govern angiogenesis by editing ALK1 linear polyubiquitin."

reach
"Moreover, immunoprecipitation of SHARPIN copurified HOIP, HOIL-1 and OTULIN, and OTULIN interacted with HOIP in proteomic experiments, indicating that OTULIN may associate with HOIP and/or LUBAC."

reach
"Moreover, LUBAC and OTULIN complex formation is regulated by OTULIN phosphorylation in the PIM."
TNFAIP3 affects OTULIN
| 1 4
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"Negative regulators of linear ubiquitination-dependent NF-κB activation have recently been reported, including the FAM105B (otulin) and CYLD linear ubiquitin-specific deubiquitinases (DUBs) and A20, which specifically binds the linear ubiquitin moiety and edits Rip1 ubiquitination to inhibit its activation ( xref ; xref ; xref ; xref ; xref )."
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
SHARPIN affects OTULIN
4 | 1

No evidence text available

No evidence text available

No evidence text available

No evidence text available
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
RBCK1 affects OTULIN
4 |
4 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
OTULIN affects TNFAIP3
| 1 4
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"Negative regulators of linear ubiquitination-dependent NF-κB activation have recently been reported, including the FAM105B (otulin) and CYLD linear ubiquitin-specific deubiquitinases (DUBs) and A20, which specifically binds the linear ubiquitin moiety and edits Rip1 ubiquitination to inhibit its activation ( xref ; xref ; xref ; xref ; xref )."
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
OTULIN affects DVL2
3 1 | 1
3 1 |

No evidence text available

No evidence text available

No evidence text available

No evidence text available
| 1

sparser
"They speculated that because both DVL2 and HOIP binds to gumby via the N-terminus, and the interaction with HOIP requires the catalytic activity of gumby, the interactions may regulate overall complex activity, substrate selection or specific signaling pathway ( xref )."
TNFAIP3 affects CYLD
| 1 3
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
SPATA2 affects CYLD
| 4
| 4

sparser
"LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"Fig. 2: LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [32]."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [ xref ]."
OTULIN affects linear Ub chains
| 4
OTULIN binds linear Ub chains.
| 2
OTULIN binds linear Ub chains. 2 / 2
| 2

reach
"Disease associated mutations affect the OTU domain and binding of OTULIN to linear Ub chains."

reach
"Disease associated mutations reside in the OTU domain and are predicted to affect binding of OTULIN to linear Ub chains."
OTULIN inhibits linear Ub chains.
| 1
Mutated OTULIN inhibits linear Ub chains. 1 / 1
| 1

reach
"Consequently, mutant OTULIN proteins failed to prevent the accumulation of linear Ub chains on target substrates IKK and NEMO, RIPK1, TNFR1, and ASC."
OTULIN activates linear Ub chains.
| 1
OTULIN activates linear Ub chains. 1 / 1
| 1

reach
"Further, ablation of OTULIN but not CYLD was reported to promote significant accumulation of linear Ub chains in the absence of any stimulation."
OTULIN affects Syndrome
| 4
OTULIN inhibits Syndrome.
| 2
| 2

reach
"For the negative role of OTULIN in immune responses, OTULIN deficiency might cause auto-inflammatory syndrome [128]."

reach
"Deficiency of the linear ubiquitin chain-specific deubiquitinase, OTULIN, causes OTULIN-related autoinflammatory syndrome (ORAS), a systemic inflammatory pathology affecting multiple organs including the skin."
OTULIN activates Syndrome.
| 2
Mutated OTULIN activates Syndrome. 1 / 1
| 1

reach
"Finally, expression of a hypomorphic mutant Otulin allele, previously shown to cause OTULIN-related autoinflammatory syndrome in humans, induces a similar inflammatory phenotype, thus supporting the importance of OTULIN for restraining skin inflammation and maintaining immune homeostasis."
| 1

reach
"Biallelic hypomorphic mutation in human OTULIN gene causes a potentially fatal autoinflammatory condition variably termed Otulipenia, OTULIN-related autoinflammatory syndrome (ORAS), or Autoinflammation, panniculitis, and dermatosis syndrome (AIPDS)."
OTULIN affects RIPK2
| 4
OTULIN ubiquitinates RIPK2.
| 2
OTULIN ubiquitinates RIPK2 on M1. 1 / 1
| 1

reach
"OTULIN was shown to limit M1-linked ubiquitination of RIPK2 and antagonize LUBAC after NOD2 activation and subsequent NF-κB and MAPK signaling (Fiil et al., 2013)."
OTULIN leads to the ubiquitination of RIPK2. 1 / 1
| 1

reach
"By deubiquitinating RIPK2, OTULIN prevents NEMO binding and hence decreases its downstream signalling."
OTULIN inhibits RIPK2.
| 1
OTULIN inhibits RIPK2. 1 / 1
| 1

reach
"Conversely, OTULIN overexpression suppressed Met1-Ub of RIPK2 in a manner dependent on its catalytic activity."
OTULIN deubiquitinates RIPK2.
| 1
OTULIN deubiquitinates RIPK2 on M1. 1 / 1
| 1

reach
"OTULIN was shown to limit M1-linked ubiquitination of RIPK2 and antagonize LUBAC after NOD2 activation and subsequent NF-κB and MAPK signaling (Fiil et al., 2013)."
CYLD affects TNFAIP3
| 1 3
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
CYLD affects SPATA2
| 4
| 4

sparser
"LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"Fig. 2: LUBAC exists in distinct complexes with the DUBs OTULIN (1) and CYLD-SPATA2 (2)."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [32]."

sparser
"The mechanisms regulating the interaction of LUBAC with OTULIN and SPATA2-CYLD in cells are not well-understood but in vitro studies show that phosphorylation of the tyrosine (Y56) in the OTULIN PIM abrogates its interaction with HOIP, suggesting that this may be a mechanism to regulate the LUBAC–OTULIN complex [ xref ]."
Tamoxifen affects OTULIN
| 3
| 3

reach
"Tamoxifen induced OTULIN deficiency in immune cells led to an acute, serious multi-organ inflammatory phenotype [XREF_BIBR]."

reach
"Tamoxifen induced Otulin deficiency in immune cells results in an acute severe multiorgan inflammatory phenotype."

reach
"Tamoxifen induced Otulin ablation in murine immune cells (CreERT2-OtulinLacZ and flox chimeras) resembles the phenotype described in patients with otulipenia and ORAS including responsiveness to therapy with TNF inhibitors."
Trim14 affects CYLD
| 3
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
S1 affects OTULIN
| 3
S1-E314R inhibits OTULIN. 1 / 1
| 1

reach
"Point mutations in the S1 (L259E and E314R) or S1 ' (W96A and TQK100-102AAA) Ub binding sites reduced OTULIN activity toward Met1-diUb (XREF_FIG I) by decreasing Met1-diUb affinity (XREF_FIG J)."
S1-L259E inhibits OTULIN. 1 / 1
| 1

reach
"Point mutations in the S1 (L259E and E314R) or S1 ' (W96A and TQK100-102AAA) Ub binding sites reduced OTULIN activity toward Met1-diUb (XREF_FIG I) by decreasing Met1-diUb affinity (XREF_FIG J)."
S1 inhibits OTULIN. 1 / 1
| 1

reach
"Point mutations in the S1 (L259E and E314R) or S1 ' (W96A and TQK100-102AAA) Ub binding sites reduced OTULIN activity toward Met1-diUb (XREF_FIG I) by decreasing Met1-diUb affinity (XREF_FIG J)."
PUB affects OTULIN
| 3
| 3

reach
"52 Interestingly, Otulin binds to the PUB (N-terminal peptide : N-glycanase and UBA- or UBX containing protein) domain of HOIP, while CYLD is also recruited to the PUB domain of HOIP, but indirectly via the protein SPATA2 (spermatogenesis associated protein 2)."

reach
"LUBAC activity is counterbalanced by the Met1 specific deubiquitinase (DUB) OTULIN, which binds to the catalytic subunit HOIP via interactions between the HOIP peptide : N-glycanase and UBA- or UBX containing proteins (PUB) domain and a PUB interacting motif (PIM) in OTULIN."

reach
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide : N-glycanase and UBA- or UBX containing proteins) domain [XREF_BIBR, XREF_BIBR]."
PIM affects OTULIN
| 1 2
PIM binds OTULIN.
| 2
| 1

sparser
"OTULIN selectively interacts with LUBAC through the N-terminal PIM, covering Asp54-Met55-Tyr56-Arg57-Ala58 [ xref , xref ], but is not able to interact with the PUB domains of UBX domain-containing protein 1 (UBXD1) or PNGase."
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
PIM inhibits OTULIN.
| 1
PIM inhibits OTULIN. 1 / 1
| 1

reach
"Importantly, mutation of the cornerstone residue Asn102 to Asp (N102D) or a mutation that affect the hydrophobic PIM pocket (Y82A) reduced the ability of OTULIN to antagonize LUBAC induced NFkappaB activity in comparison to wild-type HOIP (XREF_FIG D)."

reach
"Increased OTULIN expression was required for EA to inhibit the NF-kappaB signalling pathway in focal cerebral ischaemia and reperfusion rats."

reach
"As both CYLD and OTULIN negatively regulate the NF-kappaB signalling cascade, the impact of these deubiquitinases in different LUBAC dependent signalling cascades may be context dependent."

reach
"To further investigate whether enhanced OTULIN was required for EA to inhibit the NF-kappaB signalling pathway, OTULIN, p-IkappaBalpha, IkappaBalpha, cytoplasm-p65 and nucleus-p65 proteins from brain tissues in the ischaemic penumbra of the cerebral cortex were detected 24h after reperfusion."
| 2

reach
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice."

reach
"OTULIN Tyr 56 phosphorylation is counteracted by the activity of dual-specificity phosphatase 14 (DUSP14), which we identify as an OTULIN phosphatase that limits necroptosis."
| 1

reach
"OTULIN hyper-phosphorylation promotes necroptosis by locking ring finger protein 31 (RNF31, also known as HOIP) away from the cylindromatosis (CYLD) complex, resulting in altered receptor interacting serine/threonine kinase 1 (RIPK1) ubiquitination."
OTULIN affects monoubiquitin
| 2 1
OTULIN binds monoubiquitin.
| 1 1
OTULIN binds monoubiquitin. 2 / 2
| 1 1

reach
"Interestingly, monoubiquitin binding to OTULIN did not induce rearrangement of the active site into a catalytically competent configuration 30, and monoubiquitin based suicide probes, where the C terminus of ubiquitin is replaced with a Cys reactive group 44, were unable to react with OTULIN, confirming that the catalytic triad in OTULIN was incompetent 29."

sparser
"Interestingly, monoubiquitin binding to OTULIN did not induce rearrangement of the active site into a catalytically competent configuration xref , and monoubiquitin‐based suicide probes, where the C terminus of ubiquitin is replaced with a Cys‐reactive group xref , were unable to react with OTULIN, confirming that the catalytic triad in OTULIN was incompetent xref ."
OTULIN activates monoubiquitin.
| 1
OTULIN activates monoubiquitin. 1 / 1
| 1

reach
"Recent evidence suggests that monoubiquitin release from its precursors is mediated predominantly by five main DUBs : ubiquitin thioesterase otulin, UCHL3, USP5, USP7 and USP9X [XREF_BIBR], although each has multiple substrates that can be targeted for deubiquitination to maintain the free ubiquitin pool."
| 3

reach
"OTULIN haploinsufficiency impairs intrinsic immunity to alpha-toxin."

reach
"Human OTULIN haploinsufficiency impairs cell-intrinsic immunity to staphylococcal α-toxin."

reach
"Human OTULIN haploinsufficiency impairs cell-intrinsic immunity to staphylococcal alpha-toxin."
| 1 2
| 1 1

reach
"The OTULIN mediated suppression of hepatocyte apoptosis plays a crucial role in liver pathogeneses, such as hepatitis, fibrosis, and hepatocellular carcinoma [XREF_BIBR]."

trips
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase-Mediated Hepatocyte Apoptosis."
| 1

reach
"OTULIN deficiency triggers metabolic alterations, apoptosis, and inflammation in the liver."
OTULIN affects activity
| 3
OTULIN inhibits activity. 3 / 3
| 3

sparser
"Indeed, at the cellular level, OTULIN inhibits LUBAC-induced NF-κB activity dependently on the catalytic activity of OTULIN ( xref )."

sparser
"CYLD and OTULIN were not inhibiting NF-κB activity per se because the DUBs did not appreciably inhibit NF-κB activity induced by an engineered non-cleavable Met1-Ub4 protein targeted to inactive XIAP ( xref ; xref E and S3F)."

sparser
"XIAP-induced NF-κB activity relies not exclusively on Met1-Ub ( xref ) but was also dependent on Ubc13-mediated formation of Lys63-Ub ( xref E and xref D), which might explain why OTULIN only partially inhibited XIAP-induced NF-κB activity ( xref D)."
OTULIN affects Wnt
| 2
OTULIN inhibits Wnt.
| 1
OTULIN inhibits Wnt. 1 / 2
| 1

reach
"The interaction of HOIP with OTULIN is also involved in OTULIN suppressing the canonical Wnt signaling pathway activation by LUBAC [XREF_BIBR]."
OTULIN activates Wnt.
| 1
OTULIN activates Wnt. 1 / 1
| 1

reach
"Thus, OTULIN mediated Wnt and beta-catenin activation upon genotoxic treatments promotes drug resistance and metastasis in breast cancers."
OTULIN affects RELA
| 3
OTULIN phosphorylates RELA.
| 2
OTULIN leads to the phosphorylation of RELA. 2 / 2
| 2

reach
"Importantly, OTULIN deficiency in THP-1 monocytes led to spontaneous phosphorylation of p65 and RelA (Fig XREF_FIG A) and drastically increased turnover of IkappaBalpha (Fig XREF_FIG B), indicative of NF-kappaB activation in shOTULIN cells."

reach
"Importantly, OTULIN deficiency in THP-1 monocytes led to spontaneous phosphorylation of p65 and RelA (Fig XREF_FIG A) and drastically increased turnover of IkappaBalpha (Fig XREF_FIG B), indicative of NF-kappaB activation in shOTULIN cells."
OTULIN inhibits RELA.
| 1
OTULIN inhibits RELA. 1 / 1
| 1

reach
"Another study found that OTULIN overexpression inhibited TNF-α-induced nuclear translocation of p65 in HEK293 T cells [83]."
OTULIN affects PUB
| 3
| 3

reach
"52 Interestingly, Otulin binds to the PUB (N-terminal peptide : N-glycanase and UBA- or UBX containing protein) domain of HOIP, while CYLD is also recruited to the PUB domain of HOIP, but indirectly via the protein SPATA2 (spermatogenesis associated protein 2)."

reach
"LUBAC activity is counterbalanced by the Met1 specific deubiquitinase (DUB) OTULIN, which binds to the catalytic subunit HOIP via interactions between the HOIP peptide : N-glycanase and UBA- or UBX containing proteins (PUB) domain and a PUB interacting motif (PIM) in OTULIN."

reach
"Mass spectrometry and biochemical analysis showed that OTULIN binds to the HOIP PUB (peptide : N-glycanase and UBA- or UBX containing proteins) domain [XREF_BIBR, XREF_BIBR]."
OTULIN affects PIM
| 3
OTULIN binds PIM.
| 2
| 1

sparser
"OTULIN selectively interacts with LUBAC through the N-terminal PIM, covering Asp54-Met55-Tyr56-Arg57-Ala58 [ xref , xref ], but is not able to interact with the PUB domains of UBX domain-containing protein 1 (UBXD1) or PNGase."
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
OTULIN phosphorylates PIM.
| 1
OTULIN phosphorylates PIM. 1 / 1
| 1

sparser
"Moreover, LUBAC-OTULIN complex formation is regulated by OTULIN phosphorylation in the PIM."
OTULIN affects OTULIN
| 3
OTULIN activates OTULIN.
| 2
Mutated OTULIN activates OTULIN. 1 / 1
| 1

reach
"Finally, expression of a hypomorphic mutant Otulin allele, previously shown to cause OTULIN-related autoinflammatory syndrome in humans, induces a similar inflammatory phenotype, thus supporting the importance of OTULIN for restraining skin inflammation and maintaining immune homeostasis."
OTULIN activates OTULIN. 1 / 1
| 1

reach
"This is in stark contrast to the interaction of HOIP with OTULIN, which was substantially increased by OTULIN overexpression (XREF_FIG C)."
OTULIN inhibits OTULIN.
| 1
OTULIN-C129A inhibits OTULIN. 1 / 1
| 1

reach
"Inactivation of OTULIN by expression of OTULIN C129A in adult mice has recently been reported to cause systemic inflammation, which can be prevented by combined ablation of cell death pathways by deletion of caspase-8, RIPK3 and RIPK1 (Heger etal, 2018)."
OTULIN affects NGLY1
| 1
| 1

reach
"OTULIN interacts directly with a peptide : N-glycanase and UBA-containing or UBX containing protein (PUB) domain in HOIP via an evolutionarily conserved PUB interacting motif (PIM) in the N-terminal region preceding the OTU domain XREF_BIBR - XREF_BIBR."
OTULIN affects Met1-polyUb
| 3
OTULIN activates Met1-polyUb.
| 2
OTULIN activates Met1-polyUb. 2 / 2
| 2

reach
"Interestingly, both WT and DUB domain mutant OTULIN disabled LUBAC induced NF-kappaB activation, indicating that OTULIN mediated Met1-polyUb is not the only factor influencing NF-kappaB activation [XREF_BIBR]."

reach
"Interestingly, both WT and DUB domain mutant OTULIN disabled LUBAC-induced NF-κB activation, indicating that OTULIN-mediated Met1-polyUb is not the only factor influencing NF-κB activation [83]."
OTULIN inhibits Met1-polyUb.
| 1
OTULIN inhibits Met1-polyUb. 1 / 1
| 1

reach
"It appears that OTULIN overexpression antagonizes NF-kappaB activation by removing Met1-polyUb, whereas OTULIN C129A acts as a high-affinity UBD that competes with other Met1 specific UBDs required for NF-kappaB signaling in a similar manner to the one recently reported for the overexpression of the NEMO UBAN domain."
OTULIN affects Met1
| 3
SNX27 binds OTULIN and Met1. 1 / 1
| 1

reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
RNF31 binds OTULIN and Met1. 1 / 1
| 1

reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
OTULIN binds CYLD and Met1. 1 / 1
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
OTULIN affects MTOR
| 1 2
OTULIN binds MTOR.
| 2
| 2

sparser
"Instead, we find that steatohepatitis in OTULIN-deficient livers is associated with aberrant mTOR activation, and inhibition of mTOR by rapamycin administration significantly reduces the liver pathology."

sparser
"Treated Otulin ∆hep mice displayed reduced weight gain when compared with vehicle-treated mice and even rapamycin-treated controls (Fig.  xref ), demonstrating a pharmacogenetic interaction between OTULIN deficiency and mTOR inhibition."
OTULIN inhibits MTOR.
| 1
OTULIN inhibits MTOR. 1 / 1
| 1

reach
"Loss of the deubiquitinase OTULIN promotes hepatocellular carcinoma (HCC) in an mTOR dependent manner."
OTULIN affects MALT1
| 3
| 3

sparser
"Moreover, neither MALT1 nor OTULIN could bind the PUB domain-deleted mutant of HOIP."

sparser
"These results indicated that MALT1 and OTULIN bind the PUB domain of HOIP in independent manners."

sparser
"Both MALT1 and OTULIN simultaneously bound the WT-HOIP."
OTULIN affects IRF3
| 3
OTULIN ubiquitinates IRF3.
| 1
OTULIN ubiquitinates IRF3. 1 / 1
| 1

reach
"Here, we report that Otulin, the deubiquitinase that removes linear polyubiquitin chains, inhibits RIPA by deubiquitinating IRF3."
OTULIN inhibits IRF3.
| 1
OTULIN bound to Nsp11 inhibits IRF3. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
OTULIN deubiquitinates IRF3.
| 1
OTULIN deubiquitinates IRF3. 1 / 1
| 1

reach
"The antiviral action of the RIG-I induced pathway of apoptosis (RIPA) is enhanced by its ability to degrade Otulin, which deubiquitinates IRF3."
OTULIN affects IL6
| 3
OTULIN inhibits IL6.
| 2
OTULIN inhibits IL6. 2 / 2
| 2

reach
"Consistent with the finding that OTULIN overexpression was shown to delay NF-kappaB activation of TNF, whereas its knockdown was reported to increase the output of NF-kappaB signaling [XREF_BIBR, XREF_BIBR, XREF_BIBR], we found that OTULIN overexpression strongly depressed NF-kappaB activity and reduced the release of TNF-alpha, IL-1beta, and IL-6, which are the primary pro inflammatory cytokines that aggravate brain injury."

reach
"OTULIN overexpression significantly suppressed the production of TNF-alpha, IL-1beta, and IL-6 in PM and N9 cells via inhibiting the nuclear translocation of NF-kappaB p65 but had no effect on the pro inflammatory cytokines secreted by un activated microglia, which directly indicates that OTULIN suppressed microglia mediated neuroinflammation by inhibiting the NF-kappaB pathway and attenuated the release of inflammatory mediators under the in vitro ischemic condition."
OTULIN increases the amount of IL6.
| 1
OTULIN increases the amount of IL6. 1 / 1
| 1

reach
"By contrast, OTULIN haploinsufficiency did not lead to a loss of LUBAC expression in PDFs (Figure 3A), or to impaired TNF-receptor signaling (Figure S3F) or IL-6 and IL-8 secretion (Figure S3G)."
| 1 2
| 2

sparser
"Nsp11 was reported to interact with OTULIN and the IKK complex, and to inhibit NF-κB activation in response to SeV infection but the mechanism is unclear [88]."

sparser
"Nsp11 was reported to interact with OTULIN and the IKK complex, and to inhibit NF-κB activation in response to SeV infection but the mechanism is unclear [ xref ]."
OTULIN inhibits IKK_complex.
| 1
Mutated OTULIN inhibits IKK_complex. 1 / 1
| 1

reach
"Mutant OTULIN proteins can not restrict the accumulation of Met1 Ub chains on target substrates IKK and NEMO, RIPK1, and ASC."
OTULIN affects GS26575
| 3
OTULIN inhibits GS26575.
| 2
| 2

reach
"The deubiquitinase OTULIN negatively regulates M1 linked polyubiquitin signalling by removing the chains conjugated by LUBAC, and OTULIN deficiency causes OTULIN related autoinflammatory syndrome (ORAS) in humans."

reach
"OTULIN directly binds the LUBAC component HOIP, and knockdown of OTULIN in human cell lines increases M1 linked chains on LUBAC and its substrates."
OTULIN activates GS26575.
| 1
| 1

reach
"While absence of OTULIN induces spontaneous M1 linked polyubiquitination and NF-kappaB activation in myeloid cells, OTULIN deficiency in T and B cells results in loss of LUBAC."
OTULIN affects DVL
| 3
| 3

sparser
"The observations that OTULIN mutation negatively impacts vessel development and that OTULIN interacts with dishevelled (DVL) 2, a Wnt signaling adapter, suggested that linear ubiquitination could be involved in Wnt signaling."

sparser
"The interaction of OTULIN with a key component Dishevelled (Dvl/Dsh) mediates this regulation through the Wnt signalling cascade."

sparser
"Moreover, a reported interaction of OTULIN and dishevelled‐2 (DVL2) could be confirmed, suggesting a role of Met1‐linked ubiquitination in Wnt signalling xref , xref ."
NGLY1 affects OTULIN
| 1
| 1

reach
"OTULIN interacts directly with a peptide : N-glycanase and UBA-containing or UBX containing protein (PUB) domain in HOIP via an evolutionarily conserved PUB interacting motif (PIM) in the N-terminal region preceding the OTU domain XREF_BIBR - XREF_BIBR."
NFkappaB affects OTULIN
| 3
NFkappaB activates OTULIN.
| 2
| 2

reach
"In line with these observations, TNFalpha stimulation results in reduced Met1 ubiquitination of RIPK1 and NEMO, and decreased NF-kappaB signaling in OTULIN deficient keratinocytes [XREF_BIBR]."

reach
"NF-kappaB activation and TNF secretion in human OTULIN deficient monocytes."
NFkappaB increases the amount of OTULIN.
| 1
NFkappaB increases the amount of OTULIN. 1 / 1
| 1

reach
"Contrary to this, OTULIN functions to restrict the accumulation of Met1-Ub at basal conditions and early during signaling, and OTULIN expression is not induced by stimulation of NF-kappaB activity."
Met1 affects OTULIN
| 3
SNX27 binds OTULIN and Met1. 1 / 1
| 1

reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
RNF31 binds OTULIN and Met1. 1 / 1
| 1

reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
OTULIN binds CYLD and Met1. 1 / 1
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
MALT1 affects OTULIN
| 3
| 3

sparser
"Moreover, neither MALT1 nor OTULIN could bind the PUB domain-deleted mutant of HOIP."

sparser
"These results indicated that MALT1 and OTULIN bind the PUB domain of HOIP in independent manners."

sparser
"Both MALT1 and OTULIN simultaneously bound the WT-HOIP."
Glu-Ala affects OTULIN
| 3
Glu-Ala increases the amount of OTULIN. 3 / 3
| 3

reach
"EA treatment promotes OTULIN gene transcription and protein synthesis in the brain, and enhanced OTULIN expression was primarily located in neurons."

reach
"EA enhanced OTULIN expression following focal cerebral ischaemia and reperfusion injury."

reach
"EA also enhanced microglial OTULIN expression in the present study, and OTULIN silencing effectively reversed the inhibitory effect of EA on microglial activation, as shown by the increased mean Iba-1 immunofluorescence intensity, which strongly indicates that enhanced OTULIN expression is necessary for EA to alleviate microglial activation."
DVL affects OTULIN
| 3
| 3

sparser
"The observations that OTULIN mutation negatively impacts vessel development and that OTULIN interacts with dishevelled (DVL) 2, a Wnt signaling adapter, suggested that linear ubiquitination could be involved in Wnt signaling."

sparser
"The interaction of OTULIN with a key component Dishevelled (Dvl/Dsh) mediates this regulation through the Wnt signalling cascade."

sparser
"Moreover, a reported interaction of OTULIN and dishevelled‐2 (DVL2) could be confirmed, suggesting a role of Met1‐linked ubiquitination in Wnt signalling xref , xref ."
CYLD affects Trim14
| 3
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
2 |
Valproic acid demethylates OTULIN.
1 |
1 |

No evidence text available
Valproic acid decreases the amount of OTULIN.
1 |
Valproic acid decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Monoubiquitin affects OTULIN
| 1 1
OTULIN binds monoubiquitin. 2 / 2
| 1 1

reach
"Interestingly, monoubiquitin binding to OTULIN did not induce rearrangement of the active site into a catalytically competent configuration 30, and monoubiquitin based suicide probes, where the C terminus of ubiquitin is replaced with a Cys reactive group 44, were unable to react with OTULIN, confirming that the catalytic triad in OTULIN was incompetent 29."

sparser
"Interestingly, monoubiquitin binding to OTULIN did not induce rearrangement of the active site into a catalytically competent configuration xref , and monoubiquitin‐based suicide probes, where the C terminus of ubiquitin is replaced with a Cys‐reactive group xref , were unable to react with OTULIN, confirming that the catalytic triad in OTULIN was incompetent xref ."
Linear Ub chains affects OTULIN
| 2
OTULIN binds linear Ub chains. 2 / 2
| 2

reach
"Disease associated mutations affect the OTU domain and binding of OTULIN to linear Ub chains."

reach
"Disease associated mutations reside in the OTU domain and are predicted to affect binding of OTULIN to linear Ub chains."
2 |
Hsa-miR-6849-3p decreases the amount of OTULIN. 2 / 2
2 |

No evidence text available

No evidence text available
VPS26A affects OTULIN
| 2
| 2

sparser
"It appears SNX27 cannot undertake simultaneous binding to both VPS26A and OTULIN (Figure 5), because OTULIN and VPS26A use a partially overlapping binding site located in the SNX27 β3–β4 hairpin loop and would experience clashes between atoms."

sparser
"It appears SNX27 cannot undertake simultaneous binding to both VPS26A and OTULIN ( xref ), because OTULIN and VPS26A use a partially overlapping binding site located in the SNX27 β3–β4 hairpin loop and would experience clashes between atoms."
VCP affects RNF31
| 2
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
UBE2L3 affects TNFAIP3
| 2
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
STAT1 affects OTULIN
| 1 1
| 1 1

sparser
"Interestingly, IFNɑ treatment significantly promoted the binding of OTULIN to STAT1 in a time-dependent manner (Fig.  xref and Supplementary Fig.  xref )."

reach
"Interestingly, IFNɑ treatment significantly promoted the binding of OTULIN to STAT1 in a time-dependent manner (Fig. 5e and Supplementary Fig. 5g, h)."
RNF31 affects VCP
| 2
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
RNF31 affects PUB domain
| 2
RNF31 binds OTULIN and PUB domain. 2 / 2
| 2

reach
"Although HOIP is able to interact with the AAA ATPase p97 and valosin containing protein (VCP) [XREF_BIBR, XREF_BIBR], which has recently been implicated in the recruitment of LUBAC to misfolded Huntingtin species [XREF_BIBR], the HOIP PUB domain binds OTULIN with a 40-fold higher affinity compared to p97 and VCP and NMR based in vitro studies indicate a more stable interaction between HOIP and OTULIN compared to p97 and VCP [XREF_BIBR]."

reach
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
OTULIN affects polyubiquitin chain
| 2
OTULIN deubiquitinates polyubiquitin chain. 2 / 2
| 2

reach
"Mechanistic studies showed that, upon TNFalpha stimulation, OTULIN-SNX27 is localised to membrane associated TNF receptor complex, where OTULIN deubiquitinates the linear polyubiquitin chain that formed by the LUBAC complex."

reach
"Mechanistic studies showed that, upon TNFalpha stimulation, OTULIN-SNX27 is localised to membrane associated TNF receptor complex, where OTULIN deubiquitinates the linear polyubiquitin chain that formed by the LUBAC complex."
| PMC

reach
"OTULIN is critical for restraining life threatening spontaneous inflammation, maintaining immune homeostasis and activating NF-kappaB to promote the secretion of pro inflammatory cytokines and restricts bacterial proliferation in infection [XREF_BIBR, XREF_BIBR]."

reach
"PRRSV infection promotes the expression of the OTULIN gene; in turn, overexpression of OTULIN contributes to PRRSV proliferation."
OTULIN affects auto-linear ubiquitination
| 2
OTULIN inhibits auto-linear ubiquitination. 2 / 2
| 2

eidos
"Therefore , under physiological conditions , OTULIN prevents the auto-linear ubiquitination of LUBAC , and functions to maintain the LUBAC activity ."

eidos
"Because OTULIN prevents auto-linear ubiquitination of LUBAC and maintains the LUBAC activity [ 23,63 ] , OTULIN deficiency induces deterioration of LUBAC ."
| PMC
OTULIN affects VPS26A
| 2
| 2

sparser
"It appears SNX27 cannot undertake simultaneous binding to both VPS26A and OTULIN (Figure 5), because OTULIN and VPS26A use a partially overlapping binding site located in the SNX27 β3–β4 hairpin loop and would experience clashes between atoms."

sparser
"It appears SNX27 cannot undertake simultaneous binding to both VPS26A and OTULIN ( xref ), because OTULIN and VPS26A use a partially overlapping binding site located in the SNX27 β3–β4 hairpin loop and would experience clashes between atoms."
OTULIN affects UBE2L3
| 2
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
OTULIN affects TNFRSF1A
| 2
OTULIN ubiquitinates TNFRSF1A.
| 1
OTULIN leads to the ubiquitination of TNFRSF1A. 1 / 1
| 1

reach
"Analysis of the TNF-RSC in TRex-293 cells expressing an OTULIN targeting miRNA showed that OTULIN depletion led to both accelerated and increased accumulation of Met1-Ub, and increased ubiquitination [MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTULIN inhibits TNFRSF1A.
| 1
Modified OTULIN inhibits ubiquitinated TNFRSF1A. 1 / 1
| 1

reach
"Conversely, overexpression of OTULIN decreased the level of Met1-Ub in the TNF-RSC and reduced the apparent molecular weight of ubiquitinated TNFR1."
OTULIN affects RNF31, and VCP
| 2
VCP binds RNF31 and OTULIN. 2 / 2
| 2

sparser
"To further investigate the interaction between HOIP, OTULIN and p97, we solved the structure of the HOIP PUB domain in complex with OTULIN (aa 52–61) and p97 (aa 796–806) peptides by molecular replace[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"

sparser
"In conclusion, both OTULIN and p97 directly bind to HOIP via a conserved Ψ–Y pocket in the PUB domain, but the affinity of OTULIN for HOIP PUB domain was much higher than the affinity of p97 for the P[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTULIN affects Proteasome
| 2
OTULIN deubiquitinates Proteasome.
| 1
OTULIN deubiquitinates Proteasome. 1 / 1
| 1

reach
"Deubiquitination of proteasome subunits by OTULIN regulates type I IFN production."
OTULIN activates Proteasome.
| 1
| 1

reach
"Otulin interacted with LUBAC after virus infection and the E3-ubiquitin ligase, HOIP, a component of LUBAC, ubiquitinated Otulin to trigger its proteasome-mediated degradation."
OTULIN affects PYCARD
| 2
| 1

reach
"Only the Otulin treatment reduced high molecular weight ASC, whereas the AMSH treatment had no effect (XREF_FIG)."
Mutated OTULIN inhibits PYCARD. 1 / 1
| 1

reach
"Mutant OTULIN proteins can not restrict the accumulation of Met1 Ub chains on target substrates IKK and NEMO, RIPK1, and ASC."
OTULIN affects PUB domain, and RNF31
| 2
RNF31 binds OTULIN and PUB domain. 2 / 2
| 2

reach
"Although HOIP is able to interact with the AAA ATPase p97 and valosin containing protein (VCP) [XREF_BIBR, XREF_BIBR], which has recently been implicated in the recruitment of LUBAC to misfolded Huntingtin species [XREF_BIBR], the HOIP PUB domain binds OTULIN with a 40-fold higher affinity compared to p97 and VCP and NMR based in vitro studies indicate a more stable interaction between HOIP and OTULIN compared to p97 and VCP [XREF_BIBR]."

reach
"Crystal structures and nuclear magnetic resonance experiments reveal the molecular basis for the high-affinity interaction and explain why OTULIN binds the HOIP PUB domain specifically."
OTULIN affects Nsp11
| 2
OTULIN binds Nsp11 and OTU domain. 1 / 1
| 1

reach
"The interaction between porcine OTULIN and PRRSV Nsp11 is dependent on the OTU domain."
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
OTULIN affects Non-structural protein 4A
2 |
OTULIN affects NOD2
| 2
OTULIN inhibits NOD2.
| 1
OTULIN inhibits NOD2. 1 / 1
| 1

reach
"Similarly, the recently identified DUB OTULIN XREF_BIBR, XREF_BIBR, XREF_BIBR negatively regulates NOD2 mediated signalling by preventing LUBAC autoubiquitination under basal conditions, as well as restricting the accumulation of linear ubiquitin chains on RIP2 and LUBAC upon stimulation (Fig XREF_FIG)."
OTULIN activates NOD2.
| 1
OTULIN activates NOD2. 1 / 1
| 1

reach
"Here, we report that OTULIN is critical for limiting Met1-Ub accumulation after nucleotide oligomerization domain containing protein 2 (NOD2) stimulation, and that OTULIN depletion augments signaling downstream of NOD2."
| 2
OTULIN ubiquitinates NCIT:C19802.
| 1
OTULIN ubiquitinates NCIT:C19802. 1 / 1
| 1

sparser
"Thus, abolishing HOIP’s capacity to interact with both CYLD and OTULIN enhances M1 ubiquitination in the TNF-RSC and, consequently, TNF-induced gene activation."
| 1

sparser
"Strikingly, while LUBAC translocates to receptor signaling complexes (RSCs), OTULIN is not stably associated with TNF or NOD2 RSCs ( xref ), and how it regulates signaling complexes, e.g., TNF signaling, is unclear ( xref )."
OTULIN affects IL1B
| 2
OTULIN inhibits IL1B. 2 / 2
| 2

reach
"OTULIN overexpression significantly suppressed the production of TNF-alpha, IL-1beta, and IL-6 in PM and N9 cells via inhibiting the nuclear translocation of NF-kappaB p65 but had no effect on the pro inflammatory cytokines secreted by un activated microglia, which directly indicates that OTULIN suppressed microglia mediated neuroinflammation by inhibiting the NF-kappaB pathway and attenuated the release of inflammatory mediators under the in vitro ischemic condition."

reach
"Consistent with the finding that OTULIN overexpression was shown to delay NF-kappaB activation of TNF, whereas its knockdown was reported to increase the output of NF-kappaB signaling [XREF_BIBR, XREF_BIBR, XREF_BIBR], we found that OTULIN overexpression strongly depressed NF-kappaB activity and reduced the release of TNF-alpha, IL-1beta, and IL-6, which are the primary pro inflammatory cytokines that aggravate brain injury."
OTULIN affects Flag
| 2
| 2

sparser
"e, f Co-immunoprecipitation and immunoblot analysis of extracts of HepG2.2.15 & HepG2 cells transfected with plasmid encoding Flag-OTULIN, Flag-OTULIN-K66-R and Hepatitis B virus (HBV) 1.3-fold genome plasmid (pHBV1.3) and treated with MG132, are shown"

sparser
"e, f Co-immunoprecipitation and immunoblot analysis of extracts of HepG2.2.15 & HepG2 cells transfected with plasmid encoding Flag-OTULIN, Flag-OTULIN-K66-R and Hepatitis B virus (HBV) 1.3-fold genome plasmid (pHBV1.3) and treated with MG132, are shown E3 ubiquitin ligases and deubiquitinases found to be significantly altered in response to HBV integration"
OTULIN affects DUSP14
2 |
2 |

No evidence text available

No evidence text available
Nsp11 affects OTULIN
| 2
OTULIN binds Nsp11 and OTU domain. 1 / 1
| 1

reach
"The interaction between porcine OTULIN and PRRSV Nsp11 is dependent on the OTU domain."
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
Non-structural protein 4A affects OTULIN
2 |
MTOR affects OTULIN
| 2
| 2

sparser
"Instead, we find that steatohepatitis in OTULIN-deficient livers is associated with aberrant mTOR activation, and inhibition of mTOR by rapamycin administration significantly reduces the liver pathology."

sparser
"Treated Otulin ∆hep mice displayed reduced weight gain when compared with vehicle-treated mice and even rapamycin-treated controls (Fig.  xref ), demonstrating a pharmacogenetic interaction between OTULIN deficiency and mTOR inhibition."
L18-MDP affects OTULIN
| 2
L18-MDP activates OTULIN. 2 / 2
| 2

reach
"Notably, ubiquitinated RIPK2 was readily detected in the lysates from OTULIN depleted cells treated with L18-MDP, illustrating extensive ubiquitination of RIPK2 when OTULIN is depleted."

reach
"In line with this, Met1-Ub on RIPK2 was barely detectable in control cells subjected to the low concentration of L18-MDP but accumulated in the OTULIN depleted cells to levels comparable with those in[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 2

sparser
"Nsp11 was reported to interact with OTULIN and the IKK complex, and to inhibit NF-κB activation in response to SeV infection but the mechanism is unclear [88]."

sparser
"Nsp11 was reported to interact with OTULIN and the IKK complex, and to inhibit NF-κB activation in response to SeV infection but the mechanism is unclear [ xref ]."
Flag affects OTULIN
| 2
| 2

sparser
"e, f Co-immunoprecipitation and immunoblot analysis of extracts of HepG2.2.15 & HepG2 cells transfected with plasmid encoding Flag-OTULIN, Flag-OTULIN-K66-R and Hepatitis B virus (HBV) 1.3-fold genome plasmid (pHBV1.3) and treated with MG132, are shown"

sparser
"e, f Co-immunoprecipitation and immunoblot analysis of extracts of HepG2.2.15 & HepG2 cells transfected with plasmid encoding Flag-OTULIN, Flag-OTULIN-K66-R and Hepatitis B virus (HBV) 1.3-fold genome plasmid (pHBV1.3) and treated with MG132, are shown E3 ubiquitin ligases and deubiquitinases found to be significantly altered in response to HBV integration"
| 2
E3_Ub_ligase ubiquitinates OTULIN.
| 1
E3_Ub_ligase ubiquitinates OTULIN. 1 / 1
| 1

sparser
"The OTULIN–HOIP interaction is also modulated by ubiquitination of OTULIN by the E3 ligase tripartite motif-containing protein 32 (TRIM32)."
| 1

sparser
"In this study, we showed that the DUB OTULIN interacts with the E3 ligase LUBAC via the HOIP PUB domain and that this binding is crucial for their ability to control the NF-κB signaling."
DUSP14 affects OTULIN
2 |
2 |

No evidence text available

No evidence text available
CYLD affects UBE2L3
| 2
| 2

sparser
"DUBs can directly regulate E2 enzymes, xref which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."

sparser
"DUBs can directly regulate E2 enzymes, 38 which raises the question as to whether OTULIN, CYLD, or A20 interact with UBE2L3."
CYLD affects OTULIN, RNF31, and Trim14
| 2
| 2

sparser
"A HOIP PUB mutant, which cannot interact with CYLD or OTULIN, activates NF-κB more prominently than HOIP wild type, confirming a critical role of CYLD and OTULIN as negative regulators."

sparser
"Our analysis of LUBAC obtained from non-stimulated cells confirmed previous reports that CYLD and OTULIN bind to the PUB domain of HOIP ( xref )."
Trimellitic anhydride increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Rpain-a affects OTULIN
| 1
| 1

reach
"In virus-infected cells, to overcome Otulin-mediated inhibition, RIPA actively degrades Otulin."
1 |

No evidence text available
1 |
Naphthoquinone increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Ionomycin affects OTULIN
1 |
Ionomycin decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-96-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-939-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-766-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6860 decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6771-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6720-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-661 decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6512-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-6501-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-616-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-612 decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548t-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548s decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548c-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548ay-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548at-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548as-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548ap-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-548aa decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-5189-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-508-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4696 decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-4684-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-451b decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3926 decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-373-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-372-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-371b-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-371a-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3675-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3667-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-32-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3190-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-3187-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-29c-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-196a-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-16-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-155-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-miR-1285-3p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Hsa-let-7a-5p decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Hexabromocyclododecane increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |

No evidence text available
Disheveled 2 affects OTULIN
| 1
OTULIN binds disheveled 2. 1 / 1
| 1

reach
"In addition, OTULIN interacts with disheveled 2 (DVL2), a signal transducer of canonical and non canonical Wnt receptor signaling [XREF_BIBR], via its N-terminal region [XREF_BIBR]; but how exactly OTULIN affects Wnt signaling has yet to be determined."
DiUb affects OTULIN
| 1
OTULIN binds diUb. 1 / 1
| 1

reach
"Owing to the lower resolution of the OTULIN and diUb complex external restraints were applied from the higher resolution OTULIN (C129A) Met1diUb structure in the early stages of refinement using PROSMART followed by separate JellyBody refinement according to using Refmac5."
Calcitriol affects OTULIN
1 |
Calcitriol increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
1 |
Bisphenol A decreases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Autosomal recessive mutations affects OTULIN
| 1
Autosomal recessive mutations inhibits OTULIN. 1 / 1
| 1

eidos
"Patients with reduced OTULIN expression due to autosomal recessive mutations develop fevers , dermatitis , and panniculitis ( Damgaard et al ., 2016 , 2019 ; Zhou et al ., 2016b ; Nabavi et al ., 2019 ) ."
Atrazine affects OTULIN
1 |
Atrazine increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
Anti-TNF/Infliximab affects OTULIN
| 1
Anti-TNF/Infliximab activates OTULIN. 1 / 1
| 1

reach
"We provide genetic evidence that OTULIN is a crucial invivo regulator of inflammation in mice and humans, identify TNF as a key driver of the phenotypes caused by OTULIN deficiency, and show that OTULIN deficiency and ORAS can be treated with anti-TNF and Infliximab in mice and humans."
Anti-TNF neutralizing antibodies affects OTULIN
| 1
Anti-TNF neutralizing antibodies activates OTULIN. 1 / 1
| 1

reach
"In accordance with the effective treatment of ORAS patients with Infliximab, we found that anti-TNF neutralizing antibodies, administered in parallel with tamoxifen induced Otulin deletion, completely ameliorated weight loss (XREF_FIG A)."
Acrylamide affects OTULIN
1 |
Acrylamide increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
WT-, S165A-, affects OTULIN
| 1
OTULIN binds WT-, S165A-,. 1 / 1
| 1

reach
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN, and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN (XREF_SUPPLEMENTARY H)."
1 |

No evidence text available
VIPR2 affects OTULIN
1 |
1 |

No evidence text available
Ubiquitin affects Ser65
| 1
Ubiquitin binds OTULIN and Ser65. 1 / 1
| 1

reach
"Ser20 and 57 in the proximal Ub are close to the distal Ub bound to OTULIN, while Ser65 on the proximal Ub is solvent exposed (XREF_SUPPLEMENTARY A)."
USP5 affects OTULIN
| 1
| 1

reach
"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."
UCHL3 affects OTULIN
| 1
| 1

reach
"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."
UBE2N affects RNF31
| 1
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
Trim14 affects SPATA2
| 1
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
Tobacco Smoke Pollution increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
TRIM32 affects OTULIN
| 1
| 1

sparser
"Importantly, stimulation with TNFα increased the OTULINTRIM32 interaction, indicating that TRIM32-mediated ubiquitination contributes to the control of inflammatory signaling [ xref ]."
TNFRSF1A affects OTULIN
| 1
| 1

reach
"While LUBAC appears to be active under basal conditions, Met1-Ub formation on LUBAC components was increased by NOD2 and TNFR1 stimulation in OTULIN depleted cells, suggesting that the propensity of L[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
TNFRSF affects OTULIN
| 1
| 1

sparser
"Such phosphorylation of OTULIN Tyr56 would regulate the general association of OTULIN with LUBAC before their recruitment to the TNFR complexes."
TNFAIP3 affects RNF31
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
TNF affects RNF31
| 1
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
TES affects OTULIN
1 |
1 |

No evidence text available
Ser65 affects Ubiquitin
| 1
Ubiquitin binds OTULIN and Ser65. 1 / 1
| 1

reach
"Ser20 and 57 in the proximal Ub are close to the distal Ub bound to OTULIN, while Ser65 on the proximal Ub is solvent exposed (XREF_SUPPLEMENTARY A)."
SPATA2 affects Trim14
| 1
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
SPATA2 affects RNF31
| 1
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
SPATA2 affects PIM
| 1
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
SNX27 affects PDZ domain
| 1
SNX27 binds OTULIN and PDZ domain. 1 / 1
| 1

reach
"Immunoprecipitation in 293T cells and in vitro pull-down assays using bacterial recombinant proteins indicated that the direct interaction between SNX27 and OTULIN was mediated by the PDZ domain of SNX27 and the last 6 amino acids of OTULIN (Additional file 2 : Figure S2E-G), respectively."
| PMC
SNX27 affects Met1
| 1
SNX27 binds OTULIN and Met1. 1 / 1
| 1

reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
SNW1 affects OTULIN
1 |
1 |

No evidence text available
SLC18A1 affects OTULIN
1 |

No evidence text available
SHBG affects OTULIN
| 1
| 1

sparser
"Only three DUBs were identified and the statistical comparison of sample 1 (no probe) and sample 2 (OTULIN ABP) showed that OTULIN, USP5, and UCHL3 were significantly enriched in the OTULIN ABP-bound sample (volcano plot, xref B)."
SHARPIN affects CYLD
| 1
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
SELENBP1 affects OTULIN
1 |

No evidence text available
S165D-SHARPIN affects OTULIN
| 1
OTULIN binds S165D-SHARPIN. 1 / 1
| 1

reach
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN, and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN (XREF_SUPPLEMENTARY H)."
Results EA affects OTULIN
| 1
Results EA activates OTULIN. 1 / 1
| 1

eidos
"Results EA enhanced OTULIN expression following focal cerebral ischaemia / reperfusion injury To detect the effects of EA on the time course of OTULIN expression , OTULIN mRNA and protein in the ischaemic penumbra of the cerebral cortex ( Fig. 2a ) were measured from 2 h to 7 days after reperfusion using RT-qPCR and Western blotting , respectively ."

sparser
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non-structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-κB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [ xref ]."
RNF31 affects UBE2N
| 1
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
RNF31 affects TNFAIP3
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
RNF31 affects TNF
| 1
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
RNF31 affects SPATA2
| 1
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
RNF31 affects Met55
| 1
RNF31 binds OTULIN and Met55. 1 / 1
| 1

reach
"A fluorescently labeled OTULIN peptide in which Met55 was exchanged to Leu (as in p97) bound HOIP with near-identical affinity (370nM), showing that the small change in the first hydrophobic residue did not account for the difference (XREF_FIG J)."
RNF31 affects Met1
| 1
RNF31 binds OTULIN and Met1. 1 / 1
| 1

reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
RNF31 affects MYSM1
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
RNF31 affects DVL2
| 1
| 1

sparser
"They speculated that because both DVL2 and HOIP binds to gumby via the N-terminus, and the interaction with HOIP requires the catalytic activity of gumby, the interactions may regulate overall complex activity, substrate selection or specific signaling pathway ( xref )."

reach
"A recent work showed that porcine reproductive and respiratory syndrome virus infection upregulates the expression of OTULIN [41]."
PRRSV non-structural protein 11 affects OTULIN
| 1
OTULIN binds PRRSV non-structural protein 11. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
PRRSV infection affects OTULIN
| 1
PRRSV infection activates OTULIN. 1 / 1
| 1

eidos
"PRRSV infection promotes the expression of the OTULIN gene ; in turn , overexpression of OTULIN contributes to PRRSV proliferation ."
PR-619 affects OTULIN
| 1
PR-619 activates OTULIN. 1 / 1
| 1

reach
"For inhibition, OTULIN was treated with OTULIN ABP or PR-619 at the indicated concentrations (1h, RT), prior to addition of the tetraUb chains."
PINK1 affects OTULIN
| 1
PINK1 activates OTULIN. 1 / 1
| 1

reach
"Notably, TNF-induced stabilization of the mitochondrial kinase PINK1 furthermore contributes to signal amplification by antagonizing the M1-ubiquitin-specific deubiquitinase OTULIN."
PIM affects SPATA2
| 1
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
PDZK1 affects OTULIN
1 |
1 |

No evidence text available
PDZ domain affects SNX27
| 1
SNX27 binds OTULIN and PDZ domain. 1 / 1
| 1

reach
"Immunoprecipitation in 293T cells and in vitro pull-down assays using bacterial recombinant proteins indicated that the direct interaction between SNX27 and OTULIN was mediated by the PDZ domain of SNX27 and the last 6 amino acids of OTULIN (Additional file 2 : Figure S2E-G), respectively."
| PMC
| 1
OTULIN translocates to the membrane. 1 / 1
| 1

reach
"In the meantime, SNX27 transports OTULIN to the membrane associated TNFalpha receptor complex, where OTULIN limits LUBAC induced linear polyubiquitination."
| PMC
OTULIN affects signaling
| 1
OTULIN inhibits signaling. 1 / 1
| 1

sparser
"It was shown that OTULIN inhibits both NF-κB and NOD2 signaling by limiting LUBAC-mediated Met1 ubiquitination of NEMO and RIPK2."
OTULIN affects rpain-a
| 1
| 1

reach
"Ablation of Otulin expression enhanced RIPA and its overexpression inhibited RIPA."
| 1

reach
"OTULIN overexpression significantly suppressed the production of TNF-alpha, IL-1beta, and IL-6 in PM and N9 cells via inhibiting the nuclear translocation of NF-kappaB p65 but had no effect on the pro inflammatory cytokines secreted by un activated microglia, which directly indicates that OTULIN suppressed microglia mediated neuroinflammation by inhibiting the NF-kappaB pathway and attenuated the release of inflammatory mediators under the in vitro ischemic condition."
OTULIN affects polyubiquitin chains
| 1
OTULIN activates polyubiquitin chains. 1 / 1
| 1

reach
"Similarly, OTULIN produced no detectable mono- or polyubiquitin chains, demonstrating that TAK-1 was not ubiquitinated by linear (Met1) linkages in response to injury."
OTULIN affects phosphorylation
| 1
OTULIN inhibits phosphorylation. 1 / 1
| 1

sparser
"AMSH, OTULIN and USP2, but not Otubain-1, inhibited the phosphorylation of IκBα induced by Tax ( xref ), indicating that generation of both K63 and M1 chains is essential for Tax-induced IKK activation."
OTULIN affects oraS
| 1
OTULIN activates oraS. 1 / 1
| 1

reach
"Mutations in OTULIN have recently been described to cause ORAS, an autosomal recessive autoinflammatory disease (Damgaard etal, 2016; Zhou etal, 2016)."
OTULIN affects linear ubiquitination RIP2
| 1
OTULIN inhibits linear ubiquitination RIP2. 1 / 1
| 1

eidos
"Similarly , the DUB Ovarian Tumor Deubiquitinase With Linear Linkage Specificity ( OTULIN ) which hydrolyses linear ubiquitin chains synthesized on RIP1 by LUBAC in TNF-signaling pathways [ 188 ] , was identified to restrict linear ubiquitination of RIP2 , attenuating signaling downstream of NOD2 [ 185 ] ."

eidos
"Therefore , OTULIN downregulates the LUBAC-mediated innate immune responses ."
OTULIN affects independent functions
| 1
OTULIN activates independent functions. 1 / 1
| 1

eidos
"Here , we discuss how OTULIN contributes to cell fate control and highlight novel ubiquitin-dependent and - independent functions ."
OTULIN affects highlight novel ubiquitin-dependent
| 1
OTULIN activates highlight novel ubiquitin-dependent. 1 / 1
| 1

eidos
"Here , we discuss how OTULIN contributes to cell fate control and highlight novel ubiquitin-dependent and - independent functions ."

eidos
"OTULIN inhibits liver inflammation and hepatocellular carcinoma via restraining FADD - and RIPK1 kinase-mediated hepatocyte apoptosis [ 159 ] ."
| 1
| 1

reach
"Small interfering RNA (siRNA) against OTULIN resulted in anincrease in LUBAC induced NF-kappaB activation in HEK 293ET and U2OS cells and also in T-REx 293 cells stably expressing a doxycycline inducible OTULIN targeting microRNA (miRNA) (XREF_FIG A and XREF_FIG A-S6C)."
OTULIN affects disulfide
| 1
| 1

reach
"Oxidative stress induces OTULIN dimerization via cysteine-mediated covalent disulfide bonds."
OTULIN affects disheveled 2
| 1
OTULIN binds disheveled 2. 1 / 1
| 1

reach
"In addition, OTULIN interacts with disheveled 2 (DVL2), a signal transducer of canonical and non canonical Wnt receptor signaling [XREF_BIBR], via its N-terminal region [XREF_BIBR]; but how exactly OTULIN affects Wnt signaling has yet to be determined."
OTULIN affects diUb
| 1
OTULIN binds diUb. 1 / 1
| 1

reach
"Owing to the lower resolution of the OTULIN and diUb complex external restraints were applied from the higher resolution OTULIN (C129A) Met1diUb structure in the early stages of refinement using PROSMART followed by separate JellyBody refinement according to using Refmac5."
OTULIN affects cytokine
| 1
| 1

eidos
"Loss of OTULIN expression induces increased cytokine production upon treatment with LPS or staphylococcal enterotoxin B in ex vivo experiments [ 9 ] ."
OTULIN affects cellular Met1-linked Ub
| 1
OTULIN inhibits cellular Met1-linked Ub. 1 / 1
| 1

eidos
"Loss of OTULIN expression remarkably increases cellular Met1-linked Ub levels , suggesting that OTULIN regulates cytosolic Met1 ubiquitination-dependent processes ."
OTULIN affects cell fate control
| 1
OTULIN activates cell fate control. 1 / 1
| 1

eidos
"Here , we discuss how OTULIN contributes to cell fate control and highlight novel ubiquitin-dependent and - independent functions ."
OTULIN affects autoubiquitylation
| 1
OTULIN inhibits autoubiquitylation. 1 / 1
| 1

eidos
"In the mouse model , OTULIN was shown to prevent autoubiquitylation of LUBAC , thereby promoting its E3 ligase function106 ( Fig. 2a ) ."
OTULIN affects autophagy maturation
| 1
OTULIN activates autophagy maturation. 1 / 1
| 1

eidos
"OTULIN knockdown , however , promotes autophagy initiation but blocks autophagy maturation ."
OTULIN affects autophagy initiation
| 1
OTULIN inhibits autophagy initiation. 1 / 1
| 1

eidos
"OTULIN knockdown , however , promotes autophagy initiation but blocks autophagy maturation ."
OTULIN affects auto-inflammatory syndrome
| 1
OTULIN inhibits auto-inflammatory syndrome. 1 / 1
| 1

eidos
"For the negative role of OTULIN in immune responses , OTULIN deficiency might cause auto-inflammatory syndrome [ 128 ] ."
OTULIN affects astrogliosis
| 1
OTULIN inhibits astrogliosis. 1 / 1
| 1

eidos
"OTULIN reversed the suppression of astrogliosis by EA in focal cerebral ischaemia / reperfusion rats Although OTULIN protein was not detected in astrocytes , we examined whether the attenuated neuroinflammation adversely affected the activation of astrocytes ."
OTULIN affects anti-TNF neutralizing antibody
| 1
Mutated OTULIN activates anti-TNF neutralizing antibody. 1 / 1
| 1

reach
"Here, we describe that a homozygous hypomorphic OTULIN mutation in a consanguineous family causes a potentially fatal autoinflammatory disorder termed OTULIN related autoinflammatory syndrome (ORAS), which can be managed by Infliximab (anti-TNF neutralizing antibody)."
OTULIN affects abundance linear chains
| 1
OTULIN inhibits abundance linear chains. 1 / 1
| 1

eidos
"Although the precise functions of CYLD in linear ubiquitination remain unknown , the absence of CYLD does not overtly increase the amount of linear ubiquitin chains ; by contrast , the absence of OTULIN drastically increases the abundance of linear chains [ 67 ] ."
| PMC
OTULIN affects WT-, S165A-,
| 1
OTULIN binds WT-, S165A-,. 1 / 1
| 1

reach
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN, and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN (XREF_SUPPLEMENTARY H)."
OTULIN affects VIPR2
1 |
1 |

No evidence text available
OTULIN affects Ub chains attached IRAK1
| 1
OTULIN inhibits Ub chains attached IRAK1. 1 / 1
| 1

eidos
"Otulin reduced the size of the Ub chains attached to IRAK1 ( Fig. 4 C and D ) without generating monoubiquitylated - or diubiquitylated-IRAK1 , similar to our earlier findings in human IL-1R and THP1 cells ( 32 , 33 ) ."
OTULIN affects USP5
| 1
| 1

reach
"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."
OTULIN affects UCHL3
| 1
| 1

reach
"His PD showed that OTULIN ABPDeltaG76 still bound to USP5 and UCHL3 (XREF_FIG I), confirming our MS results that these DUBs are associating largely in a non covalent manner with the probes (compare XREF_FIG B and 6C)."
OTULIN affects UBE2N
| 1
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
OTULIN affects TSPAN33
| 1
| 1

reach
"Taken together, these results indicate that OTULIN overexpression limits the accumulation of microglia in the ischemic penumbra after cerebral ischemia, but this had no effect on the microglial cells in the contralateral cortex."
OTULIN affects TRIM32
| 1
| 1

sparser
"Importantly, stimulation with TNFα increased the OTULINTRIM32 interaction, indicating that TRIM32-mediated ubiquitination contributes to the control of inflammatory signaling [ xref ]."
OTULIN affects TREM2
| 1
Modified OTULIN increases the amount of TREM2. 1 / 1
| 1

reach
"The expression of TREM2 mRNA and protein was significantly upregulated by OTULIN overexpression before the induction of tMCAO (Additionalfile1 : Figure S1)."
OTULIN affects TNFRSF
| 1
| 1

sparser
"Such phosphorylation of OTULIN Tyr56 would regulate the general association of OTULIN with LUBAC before their recruitment to the TNFR complexes."
OTULIN affects TNFR1 complex
| 1
OTULIN activates TNFR1 complex. 1 / 1
| 1

eidos
"Furthermore , Otulin inactivation reduces TNFR1 complex formation and increases cell death [ 83 ] ."
OTULIN affects TES
1 |
1 |

No evidence text available
OTULIN affects TCR-mediated NF-kappaB
| 1
OTULIN inhibits TCR-mediated NF-kappaB. 1 / 1
| 1

eidos
"OTULIN predominantly downregulates TCR-mediated NF-kappaB activation in Jurkat cells ."
OTULIN affects TCF3
| 1
OTULIN bound to Nsp11 inhibits TCF3. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."

reach
"Exosomal OTULIN from M2 macrophages promotes the recovery of spinal cord injuries via stimulating Wnt/β-catenin pathway-mediated vascular regeneration."
| 1

reach
"Biallelic hypomorphic mutation in human OTULIN gene causes a potentially fatal autoinflammatory condition variably termed Otulipenia, OTULIN-related autoinflammatory syndrome (ORAS), or Autoinflammation, panniculitis, and dermatosis syndrome (AIPDS)."
OTULIN affects Ser65, and Ubiquitin
| 1
Ubiquitin binds OTULIN and Ser65. 1 / 1
| 1

reach
"Ser20 and 57 in the proximal Ub are close to the distal Ub bound to OTULIN, while Ser65 on the proximal Ub is solvent exposed (XREF_SUPPLEMENTARY A)."
OTULIN affects Sepsis
| 1
OTULIN activates Sepsis. 1 / 1
| 1

reach
"In addition, the expression of Semaphorin-7A, OTULIN, and MAP3K1 increased in sepsis, whereas that of transferrin decreased."
OTULIN affects STAT3
| 1
OTULIN inhibits STAT3. 1 / 1
| 1

reach
"Furthermore, we demonstrated that preferential expression of OTULIN in GSCs restricts linear ubiquitination on STAT3 and drives persistent STAT3 signaling, and thereby maintains the stemness and self-renewal of GSCs."
OTULIN affects STAT1-IFNAR2
| 1
OTULIN activates STAT1-IFNAR2. 1 / 1
| 1

reach
"In line with this hypothesis, we observed that in IFN-I signaling, the STAT1-IFNAR2 interaction was significantly enhanced by OTULIN overexpression (Fig. 5g)."
OTULIN affects STAT1-IFNAR2 interaction
| 1
OTULIN activates STAT1-IFNAR2 interaction. 1 / 1
| 1

eidos
"Consistently , OTULIN knockdown blocks STAT1-IFNAR2 interaction and IFN-I-mediated STAT1 activation , demonstrating that OTULIN regulates STAT1 Met1-linked ubiquitination during IFN-I-induced antiviral signaling [ 21 ] ."
OTULIN affects SPATA2, and Trim14
| 1
| 1

sparser
"The N-terminal portion of HOIP contains a PUB domain, which mediates the interaction with the PUB interaction motifs (PIMs) of OTULIN and SPATA2 ( xref )."
OTULIN affects SNW1
1 |
1 |

No evidence text available
OTULIN affects SLC18A1
1 |

No evidence text available
OTULIN affects SHBG
| 1
| 1

sparser
"Only three DUBs were identified and the statistical comparison of sample 1 (no probe) and sample 2 (OTULIN ABP) showed that OTULIN, USP5, and UCHL3 were significantly enriched in the OTULIN ABP-bound sample (volcano plot, xref B)."
OTULIN affects SELENBP1
1 |

No evidence text available
OTULIN affects SDS
| 1
OTULIN inhibits SDS. 1 / 1
| 1

reach
"OTULIN coupled resin was washed eight times with buffer B (PBS, 0.1% BSA, 5% glycerol, 0.5% Triton-X and protease inhibitors) to get rid of loose protein interactions and subsequently boiled in 25mul of 2x reducing SDS sample buffer."
OTULIN affects S165D-SHARPIN
| 1
OTULIN binds S165D-SHARPIN. 1 / 1
| 1

reach
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN, and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN (XREF_SUPPLEMENTARY H)."

sparser
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non-structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-κB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [ xref ]."
OTULIN affects RNF31, and UBE2N
| 1
| 1

sparser
"It is not feasible to control interactions for all proteins in such a panel, but in xref we show some examples, e.g. expected interaction of cIAPs with UBC13 or HOIP with OTULIN."
OTULIN affects RNF31, and TNF
| 1
TNF binds RNF31 and OTULIN. 1 / 1
| 1

sparser
"In addition to the effect of OTULIN Y56F on LUBAC-induced NF-κB signalization, we also investigated the role of OTULIN-HOIP interaction in the TNFα-induced pathway."
OTULIN affects RNF31, and SPATA2
| 1
| 1

sparser
"Importantly, binding of OTULIN and SPATA2 to HOIP is mutually exclusive, since both proteins compete for binding to the PUB domain (Draber et al., xref ; Elliott et al., xref )."
OTULIN affects RIPK1-mediated keratinocyte necroptosis
| 1
OTULIN inhibits RIPK1-mediated keratinocyte necroptosis. 1 / 1
| 1

eidos
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice ."
OTULIN affects RIPK1 ubiquitin
| 1
OTULIN activates RIPK1 ubiquitin. 1 / 1
| 1

reach
"During necroptosis, OTULIN is hyper-phosphorylated at Tyr 56, which modulates RIPK1 ubiquitin dynamics and promotes cell death."
| 1
| 1

reach
"Biallelic hypomorphic mutation in human OTULIN gene causes a potentially fatal autoinflammatory condition variably termed Otulipenia, OTULIN-related autoinflammatory syndrome (ORAS), or Autoinflammation, panniculitis, and dermatosis syndrome (AIPDS)."
OTULIN affects PRRSV non-structural protein 11
| 1
OTULIN binds PRRSV non-structural protein 11. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
OTULIN affects PIM, and SPATA2
| 1
| 1

sparser
"HOIP, however, can also bind the OTULIN- and SPATA2-PIMs, which are situated within the protein sequence."
OTULIN affects PDZK1
1 |
1 |

No evidence text available
OTULIN affects PDZ domain, and SNX27
| 1
SNX27 binds OTULIN and PDZ domain. 1 / 1
| 1

reach
"Immunoprecipitation in 293T cells and in vitro pull-down assays using bacterial recombinant proteins indicated that the direct interaction between SNX27 and OTULIN was mediated by the PDZ domain of SNX27 and the last 6 amino acids of OTULIN (Additional file 2 : Figure S2E-G), respectively."
| PMC
OTULIN affects OTULINL
| 1
OTULIN deubiquitinates OTULINL. 1 / 1
| 1

reach
"Intriguingly, the OTULIN pseudo deubiquitinating enzyme FAM105A (OTULINL) shares ~ 33% amino acid sequence identity with OTULIN, but lacks Met1 Ub catalytic activity or the ability to interact with Ub or LUBAC [XREF_BIBR, XREF_BIBR]."
OTULIN affects OTULIN-HOIP
| 1
OTULIN activates OTULIN-HOIP. 1 / 1
| 1

reach
"The OTULIN-HOIP interaction is also modulated by ubiquitination of OTULIN by the E3 ligase tripartite motif containing protein 32 (TRIM32)."
OTULIN affects OTUD7B
| 1
OTULIN ubiquitinates OTUD7B. 1 / 1
| 1

sparser
"Multiple DUBs from the human ovarian tumor (OTU) DUB family that display various degrees of specificities towards different ubiquitin linkage types have been identified and can be used for UbiCRest: For instance, OTUB1 specifically cleaves K48-, OTUD1 K63-, Cezanne K11-, and OTULIN M1-linked ubiquitylation, whereas OTUD3 displays specificity towards K6- and K11-linked ubiquitylation ( xref )."
OTULIN affects OTU domain
| 1
OTULIN binds Nsp11 and OTU domain. 1 / 1
| 1

reach
"The interaction between porcine OTULIN and PRRSV Nsp11 is dependent on the OTU domain."

reach
"Inhibiting OTULIN or Wnt and beta-catenin sensitizes triple negative breast cancer xenograft tumors to chemotherapeutics and reduces metastasis."

reach
"Interestingly, conditional deletion of otulin in myeloid cells, but not T or B cells, causes systemic inflammation, neutrophil infiltration, and multi-organ inflammation in mouse models."
OTULIN affects NLRP6
| 1
OTULIN deubiquitinates NLRP6. 1 / 1
| 1

reach
"Cyld cleaved Ub chains from NLRP6 (XREF_FIG, top blot, lane 2) and released Ub (XREF_FIG, bottom blot, lane 2), whereas otulin failed to deubiquitinate NLRP6 (XREF_FIG, top blot, lane 3)."
OTULIN affects NFKBIA
| 1
OTULIN leads to the dephosphorylation of NFKBIA. 1 / 1
| 1

reach
"AMSH, OTULIN and USP2, but not Otubain-1, inhibited the phosphorylation of IkappaBalpha induced by Tax (XREF_FIG), indicating that generation of both K63 and M1 chains is essential for Tax induced IKK activation."
OTULIN affects NF-kappaB-responsive genes
| 1
OTULIN increases the amount of NF-kappaB-responsive genes. 1 / 1
| 1

reach
"RNAi mediated depletion of OTULIN increased L18-MDP-induced transcription of NF-kappaB-responsive genes and led to more pronounced degradation of IkappaBalpha compared with cells transfected with mism[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
OTULIN affects NF-kappaB activation pathway
| 1
OTULIN inhibits NF-kappaB activation pathway. 1 / 1
| 1

eidos
"These results clearly indicated that OTULIN down-regulates the TCR-mediated NF-kappaB activation pathway ."
OTULIN affects Mice
| 1
OTULIN inhibits Mice. 1 / 1
| 1

reach
"OTULIN inhibits RIPK1-mediated keratinocyte necroptosis to prevent skin inflammation in mice."
OTULIN affects Met55
| 1
RNF31 binds OTULIN and Met55. 1 / 1
| 1

reach
"A fluorescently labeled OTULIN peptide in which Met55 was exchanged to Leu (as in p97) bound HOIP with near-identical affinity (370nM), showing that the small change in the first hydrophobic residue did not account for the difference (XREF_FIG J)."
OTULIN affects Met1-Ub
| 1
OTULIN inhibits Met1-Ub. 1 / 1
| 1

eidos
"Despite not being stably present at receptor complexes , OTULIN restricts Met1-Ub at the NOD2 receptor complex to limit inflammatory signalling [ 58 ] , suggesting that it can regulate LUBAC-mediated signalling independently of LUBAC-binding ."
OTULIN affects MYSM1
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
OTULIN affects MRLN
| 1
OTULIN inhibits MRLN. 1 / 1
| 1

reach
"OTULIN deficiency causes a gene dosage-dependent accumulation of M1-Ub, but OTULIN haploinsufficiency is both clinically and biochemically silent for TNF-induced signaling events, instead causing a predisposition to infection."
| 1
| 1

eidos
"OTULIN targets Met1-linked Ub and counteracts LUBAC auto-ubiquitination , thereby promoting NF-kappaB activation and MAPK signaling ."
OTULIN affects M1-ubiquitin
| 1
OTULIN inhibits M1-ubiquitin. 1 / 1
| 1

reach
"The function of OTULIN is therefore likely to prevent accumulation of M1-ubiquitin linkages outside of SCs (XREF_FIG G)."
OTULIN affects M1-Ubiquitin
| 1
OTULIN inhibits M1-Ubiquitin. 1 / 1
| 1

reach
"OTULIN Antagonizes Basal LUBAC Activity, but Not M1-Ubiquitin in SCs."
OTULIN affects M1 polyubiquitination
| 1
OTULIN inhibits M1 polyubiquitination. 1 / 1
| 1

eidos
"M1 polyubiquitination can be reversed by the DUBs CYLD and OTULIN ( Komander et al ., 2009 ; Keusekotten et al ., 2013 ) ."
OTULIN affects LUBAC-
| 1
OTULIN binds LUBAC-. 1 / 1
| 1

reach
"Since immune receptor signaling networks need to be tightly controlled, LUBAC- and OTULIN centered interaction networks are dynamic and carefully regulated as well."
OTULIN affects LRRC15
1 |
1 |

No evidence text available
OTULIN affects LIVER-INFLAMMATION
| 1
OTULIN inhibits LIVER-INFLAMMATION. 1 / 1
| 1

trips
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase-Mediated Hepatocyte Apoptosis."
OTULIN affects LAMP2
1 |
1 |

No evidence text available
| 1

eidos
"OTULIN predominantly downregulates TCR-mediated NF-kappaB activation in Jurkat cells ."
OTULIN affects JUN
| 1
Modified OTULIN leads to the phosphorylation of JUN. 1 / 1
| 1

reach
"OTULIN overexpression promoted enhanced and persistent JNK activation and c-Jun phosphorylation at later time points, leading to the cleavage of PARP and caspase-3, two key players in the regulation of cell-death induction (XREF_FIG F)."
OTULIN affects JNK
| 1
Modified OTULIN leads to the phosphorylation of JNK. 1 / 1
| 1

reach
"OTULIN overexpression promoted enhanced and persistent JNK activation and c-Jun phosphorylation at later time points, leading to the cleavage of PARP and caspase-3, two key players in the regulation of cell-death induction (XREF_FIG F)."
OTULIN affects JM8A3.N1
| 1
OTULIN activates JM8A3.N1. 1 / 1
| 1

reach
"Otulin and Fam105b targeted JM8A3.N1 (C57BL/6 background strain) ES cells (Fam105b tm1a (EUCOMM) Hmgu) were obtained from EUCOMM and used to generate mice bearing neomycin selection and LacZ cassettes and a loxP flanked exon 3 of Otulin, termed the targeted " LacZ " or " LZ " allele (see also XREF_FIG)."
OTULIN affects Interferon
| 1
OTULIN bound to Nsp11 inhibits Interferon. 1 / 1
| 1

reach
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non structural protein 11 (Nsp11) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 (IRF3)-mediated type-I IFN production [XREF_BIBR]."
OTULIN affects IFN-regulatory transcription
| 1
OTULIN inhibits IFN-regulatory transcription. 1 / 1
| 1

eidos
"OTULIN is upregulated in a PRRSV infection model and interacts with the PRRSV non-structural protein 11 ( Nsp11 ) to decrease Ub levels on NEMO and to block NF-kappaB activation and IFN-regulatory transcription factor 3 ( IRF3 ) - mediated type-I IFN production [ 22 ] ."
OTULIN affects IFN-I-mediated STAT1 activation
| 1
OTULIN activates IFN-I-mediated STAT1 activation. 1 / 1
| 1

eidos
"Consistently , OTULIN knockdown blocks STAT1-IFNAR2 interaction and IFN-I-mediated STAT1 activation , demonstrating that OTULIN regulates STAT1 Met1-linked ubiquitination during IFN-I-induced antiviral signaling [ 21 ] ."
OTULIN affects IFN-I
| 1
OTULIN inhibits IFN-I. 1 / 1
| 1

reach
"Interestingly, liver inflammation observed in OTULIN deficient mice is accompanied by increased expression of IFN response genes [XREF_BIBR] and murine Otulin C129A and C129A Ripk3 -/- Casp8 -/- BMDMs show spontaneous activation of IKKepsilon and TANK binding kinase 1 (TBK1) [XREF_BIBR], indicating that OTULIN suppresses IFN-I production pathways."
OTULIN affects IFN-I pathways
| 1
OTULIN inhibits IFN-I pathways. 1 / 1
| 1

eidos
"Interestingly , liver inflammation observed in OTULIN-deficient mice is accompanied by increased expression of IFN response genes [ 68 ] and murine OtulinC129A / C129A Ripk3 - / - Casp8 - / - BMDMs show spontaneous activation of IKKepsilon and TANK-binding kinase 1 ( TBK1 ) [ 23 ] , indicating that OTULIN suppresses IFN-I production pathways ."
OTULIN affects ICAM1
1 |
1 |

No evidence text available
OTULIN affects Hepatitis
| 1
| 1

eidos
"OTULIN inhibits liver inflammation and hepatocellular carcinoma via restraining FADD - and RIPK1 kinase-mediated hepatocyte apoptosis [ 159 ] ."

trips
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase-Mediated Hepatocyte Apoptosis."
OTULIN affects FAM168A
1 |

No evidence text available
OTULIN affects FAM126A
| 1
| 1

reach
"Indeed, hepatocyte-specific deletion of OTULIN induced HCC through RIPK1 kinase activity and FADD-dependent apoptosis [53]."
OTULIN affects FADD
| 1
OTULIN activates FADD. 1 / 1
| 1

reach
"OTULIN Prevents Liver Inflammation and Hepatocellular Carcinoma by Inhibiting FADD- and RIPK1 Kinase Mediated Hepatocyte Apoptosis."
OTULIN affects ELAVL1
1 |
1 |

No evidence text available
| 1

sparser
"In this study, we showed that the DUB OTULIN interacts with the E3 ligase LUBAC via the HOIP PUB domain and that this binding is crucial for their ability to control the NF-κB signaling."
OTULIN affects E3 ligase function106
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OTULIN activates E3 ligase function106. 1 / 1
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eidos
"In the mouse model , OTULIN was shown to prevent autoubiquitylation of LUBAC , thereby promoting its E3 ligase function106 ( Fig. 2a ) ."
OTULIN affects Death
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OTULIN activates Death. 1 / 1
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reach
"Inactivation of OTULIN in adult mice also caused pro inflammatory cell death."
OTULIN affects DUSP9
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1 |

No evidence text available
OTULIN affects DUSP23
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1 |

No evidence text available
OTULIN affects DSTN
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1 |

No evidence text available
OTULIN affects DPEP1
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OTULIN inhibits DPEP1. 1 / 1
| 1

reach
"OTULIN negatively regulates muramyl dipeptide (MDP)-induced activation of the cytosolic pattern recognition receptor NOD2 [XREF_BIBR, XREF_BIBR]."
OTULIN affects DAZAP2
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1 |

No evidence text available
OTULIN affects CUL4A
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No evidence text available
OTULIN affects CREBZF
| 1
| 1

sparser
"Since the PUB domain mediates the interaction with CYLD and OTULIN (Schaeffer et al , xref ; Takiuchi et al , xref ; Draber et al , xref ) and the ZF and NZF1 are required for HOIP binding to ubiquitin and NEMO (Haas et al , xref ; Ikeda et al , xref ; Fujita et al , xref ), we hypothesised that cleavage of HOIP may modulate TRAIL‐induced gene activation."
OTULIN affects CASP8
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OTULIN leads to the deubiquitination of CASP8. 1 / 1
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reach
"OTULIN reduced ubiquitination of both, caspase-8 and RIPK1, identifying RIPK1 and caspase-8 as linearly ubiquitinated proteins (Fig XREF_FIG A)."
OTULIN affects BOP1
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No evidence text available
OTULIN affects BCL10
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No evidence text available
OTULIN affects ATG13
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OTULIN ubiquitinates ATG13. 1 / 1
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reach
"A more recent report showed that the linear ubiquitination of ATG13 by LUBAC and the deubiquitination by OTULIN are associated with autophagy initiation."
OTULIN affects AIF1
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OTULIN inhibits AIF1. 1 / 1
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reach
"OTULIN overexpression significantly suppressed the activation of Iba-1 + microglial cells located in the cortical penumbra."
OTULIN affects ACVRL1
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OTULIN deubiquitinates ACVRL1. 1 / 1
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reach
"Conversely, OTULIN deubiquitinates ALK1 to promote Smad1/5 activation."
OTULIN ABP affects OTULIN
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OTULIN ABP activates OTULIN. 1 / 1
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reach
"The pre-cleared cell extracts were then incubated at with 2 - 4mug OTULIN ABP (15 - 60min), enabling the covalent coupling of active OTULIN and the probe."
OTU domain affects OTULIN
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OTULIN binds Nsp11 and OTU domain. 1 / 1
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reach
"The interaction between porcine OTULIN and PRRSV Nsp11 is dependent on the OTU domain."
NOD2 affects OTULIN
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NOD2 activates OTULIN. 1 / 1
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reach
"This was supported by the observation that OTULIN was not transcriptionally activated by NOD2 stimulation and that OTULIN levels were not elevated, consistent with data obtained using TNF (Keusekotten[MISSING/INVALID CREDENTIALS: limited to 200 char for Elsevier]"
| 1

sparser
"Strikingly, while LUBAC translocates to receptor signaling complexes (RSCs), OTULIN is not stably associated with TNF or NOD2 RSCs ( xref ), and how it regulates signaling complexes, e.g., TNF signaling, is unclear ( xref )."
Met55 affects RNF31
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RNF31 binds OTULIN and Met55. 1 / 1
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reach
"A fluorescently labeled OTULIN peptide in which Met55 was exchanged to Leu (as in p97) bound HOIP with near-identical affinity (370nM), showing that the small change in the first hydrophobic residue did not account for the difference (XREF_FIG J)."
Met1 affects SNX27
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SNX27 binds OTULIN and Met1. 1 / 1
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reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
Met1 affects RNF31
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RNF31 binds OTULIN and Met1. 1 / 1
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reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
Met1 affects OTULIN, and SNX27
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SNX27 binds OTULIN and Met1. 1 / 1
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reach
"To study whether the interaction of SNX27, OTULIN and Met1 linkage had any impact on NF-kappaB signalling activation, we first analysed TNFalpha induced NF-kappaB signalling activation in cells stably overexpressing SNX27 by checking the phosphorylation of key NF-kappaB signalling regulators."
| PMC
Met1 affects OTULIN, and RNF31
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RNF31 binds OTULIN and Met1. 1 / 1
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reach
"OTULIN interacts with the LUBAC component HOIP and controls Met1 ubiquitination levels, NF-kappaB activation and cell fate in cell- and context dependent cellular settings."
Met1 affects CYLD
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OTULIN binds CYLD and Met1. 1 / 1
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reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
MYSM1 affects TNFAIP3
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
MYD88 affects OTULIN
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MYD88 activates OTULIN. 1 / 1
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reach
"Finally, MyD88 deficiency suppresses skin lesion development in OTULIN E-KO mice, suggesting that toll-like receptor and/or IL-1 signalling are involved in mediating skin inflammation."
LUBAC- affects OTULIN
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OTULIN binds LUBAC-. 1 / 1
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reach
"Since immune receptor signaling networks need to be tightly controlled, LUBAC- and OTULIN centered interaction networks are dynamic and carefully regulated as well."
LRRC15 affects OTULIN
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1 |

No evidence text available
LAMP2 affects OTULIN
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1 |

No evidence text available
K66R be key ubiquitination site K48-linked poly-ubiquitination affects OTULIN
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K66R be key ubiquitination site K48-linked poly-ubiquitination inhibits OTULIN. 1 / 1
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eidos
"Single-mutant variants displayed a significant reduction in ubiquitination compared to wild-type OTULIN ( Fig. 9f ) , implying that K66R might be a key ubiquitination site for K48-linked poly-ubiquitination which could lead to OTULIN degradation ."
Infections affects OTULIN
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Infections increases the amount of OTULIN. 1 / 1
| 1

reach
"PRRSV infection promotes the expression of the OTULIN gene; in turn, overexpression of OTULIN contributes to PRRSV proliferation."
ICAM1 affects OTULIN
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1 |

No evidence text available
GS26575 affects OTULIN
| 1
| 1

reach
"Decreased OTULIN function increases M1 linked Ub conjugation and NF-kappaB activation in response to TNF stimulation in LUBAC-proficient cells (Fiil etal, 2013; Keusekotten etal, 2013; Damgaard etal, 2016; Hrdinka etal, 2016)."
FAM168A affects OTULIN
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No evidence text available
ELAVL1 affects OTULIN
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1 |

No evidence text available
Drugs, Chinese Herbal increases the amount of OTULIN. 1 / 1
1 |

No evidence text available
DVL2 affects RNF31
| 1
| 1

sparser
"They speculated that because both DVL2 and HOIP binds to gumby via the N-terminus, and the interaction with HOIP requires the catalytic activity of gumby, the interactions may regulate overall complex activity, substrate selection or specific signaling pathway ( xref )."
DUSP9 affects OTULIN
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1 |

No evidence text available
DUSP23 affects OTULIN
1 |
1 |

No evidence text available
DSTN affects OTULIN
1 |
1 |

No evidence text available
DAZAP2 affects OTULIN
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1 |

No evidence text available
CYLD affects SHARPIN
| 1
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
CYLD affects OTULIN, and Trim14
| 1
| 1

sparser
"Indeed, OTULIN and CYLD bind to LUBAC via a PUB (peptide:N-glycanase/UBA- or UBX-containing proteins) domain in HOIP."
CYLD affects OTULIN, and TNFAIP3
| 1
| 1

reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
CYLD affects OTULIN, and SHARPIN
| 1
| 1

sparser
"The LUBAC exists in mutually exclusive complexes, with the deubiquitinating enzymes CYLD or OTULIN ( xref , xref ), and WT-, S165A-, or S165D-SHARPIN evenly bound to CYLD and OTULIN ( xref H)."
CYLD affects Met1, and OTULIN
| 1
OTULIN binds CYLD and Met1. 1 / 1
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reach
"Indeed it is intriguing that OTULIN, A20 and CYLD either bind or hydrolyse Met1 linkages with some degree of specificity."
CYLD affects MYSM1, OTULIN, RNF31, and TNFAIP3
| 1
| 1

sparser
"Upon immunoprecipitation, we observed no direct interaction between MYSM1 with HOIP, NEMO or the negative regulators CYLD, OTULIN or A20 (Fig.  xref )."
CYLD affects ERMAP, NCIT:C19802, NOD2, OTULIN, and TNF
| 1

sparser
"To our surprise, we found that, unlike CYLD, OTULIN formed part of neither the native TNF-RSC nor the NOD2-SC."
CYLD affects CREBZF
| 1
| 1

sparser
"Since the PUB domain mediates the interaction with CYLD and OTULIN (Schaeffer et al , xref ; Takiuchi et al , xref ; Draber et al , xref ) and the ZF and NZF1 are required for HOIP binding to ubiquitin and NEMO (Haas et al , xref ; Ikeda et al , xref ; Fujita et al , xref ), we hypothesised that cleavage of HOIP may modulate TRAIL‐induced gene activation."
CUL4A affects OTULIN
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1 |

No evidence text available
CREBZF affects OTULIN
| 1
| 1

sparser
"Since the PUB domain mediates the interaction with CYLD and OTULIN (Schaeffer et al , xref ; Takiuchi et al , xref ; Draber et al , xref ) and the ZF and NZF1 are required for HOIP binding to ubiquitin and NEMO (Haas et al , xref ; Ikeda et al , xref ; Fujita et al , xref ), we hypothesised that cleavage of HOIP may modulate TRAIL‐induced gene activation."
1 |
C646 compound increases the amount of OTULIN. 1 / 1
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No evidence text available
BOP1 affects OTULIN
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1 |

No evidence text available
BCL10 affects OTULIN
1 |
1 |

No evidence text available